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<document id="E7715E855704717B382EA73A670F8ED4" ID-DOI="10.11646/zootaxa.5375.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10200036" ID-ZooBank="27D02F09-01B7-457A-8A99-D8644B7B6ADE" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700728652862" checkinUser="plazi" docAuthor="Arimoto, Kôichi" docDate="2023" docId="03AE8A2DFFA0CF6FFF47ED34FC43FCF8" docLanguage="en" docName="zootaxa.5375.3.1.pdf" docOrigin="Zootaxa 5375 (3)" docSource="https://mapress.com/zt/article/download/zootaxa.5375.3.1/52332" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Penia babai Kishii 1994" docType="treatment" docVersion="2" lastPageNumber="312" masterDocId="FF97F255FFA6CF64FFD0EB1BFFFFFFED" masterDocTitle="Taxonomic revision of the genus Penia Laporte (Coleoptera, Elateridae, Dendrometrinae) from Taiwan" masterLastPageNumber="335" masterPageNumber="301" pageNumber="307" updateTime="1700729251563" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="600C21E2358CBDFD69A7B208AA48AF51">Taxonomic revision of the genus Penia Laporte (Coleoptera, Elateridae, Dendrometrinae) from Taiwan</mods:title>
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<treatment id="03AE8A2DFFA0CF6FFF47ED34FC43FCF8" LSID="urn:lsid:plazi:treatment:03AE8A2DFFA0CF6FFF47ED34FC43FCF8" httpUri="http://treatment.plazi.org/id/03AE8A2DFFA0CF6FFF47ED34FC43FCF8" lastPageId="11" lastPageNumber="312" pageId="6" pageNumber="307">
<subSubSection id="C31D68B0FFA0CF62FF47ED34FE3CF9A4" box="[151,451,1583,1610]" pageId="6" pageNumber="307" type="nomenclature">
<paragraph id="8BB83B3BFFA0CF62FF47ED34FE3CF9A4" blockId="6.[151,451,1583,1646]" box="[151,451,1583,1610]" pageId="6" pageNumber="307">
<heading id="D0F08C57FFA0CF62FF47ED34FE3CF9A4" bold="true" box="[151,451,1583,1610]" fontSize="11" level="1" pageId="6" pageNumber="307" reason="1">
<taxonomicName id="4C0740B8FFA0CF62FF47ED34FE3CF9A4" authority="Kishii, 1994" authorityName="Kishii" authorityYear="1994" box="[151,451,1583,1610]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="babai">
<emphasis id="B973E729FFA0CF62FF47ED34FE3CF9A4" bold="true" box="[151,451,1583,1610]" pageId="6" pageNumber="307">
<emphasis id="B973E729FFA0CF62FF47ED34FED7F9A7" bold="true" box="[151,296,1583,1610]" italics="true" pageId="6" pageNumber="307">Penia babai</emphasis>
<bibRefCitation id="EF9646CAFFA0CF62FEFFED34FE3CF9A4" author="Kishii, T." box="[303,451,1583,1609]" pageId="6" pageNumber="307" pagination="211 - 216" refId="ref22955" refString="Kishii, T. (1994) Two new Elaterid-species from Taiwan collected by the late Dr. Kintaro Baba, with a remainder of the deceased (Coleoptera, Elateridae). Special Bulletin of the Essa Entomological Society, 2, 211 - 216." type="journal article" year="1994">Kishii, 1994</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31D68B0FFA0CF62FF47ED48FECFF983" box="[151,304,1619,1646]" pageId="6" pageNumber="307" type="description">
<paragraph id="8BB83B3BFFA0CF62FF47ED48FECFF983" blockId="6.[151,451,1583,1646]" box="[151,304,1619,1646]" pageId="6" pageNumber="307">
(
<figureCitation id="133C27BEFFA0CF62FF4FED48FED7F983" box="[159,296,1619,1646]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStart-3="FIGURE 7" captionStartId-0="7.[151,250,857,881]" captionStartId-1="8.[151,250,1618,1642]" captionStartId-2="9.[151,250,1405,1429]" captionStartId-3="10.[151,250,1852,1876]" captionTargetBox-0="[151,1434,165,833]" captionTargetBox-1="[151,1436,381,1595]" captionTargetBox-2="[151,1431,381,1377]" captionTargetBox-3="[151,1436,669,1829]" captionTargetId-0="figure-16@7.[151,1436,159,841]" captionTargetId-1="figure-115@8.[151,1436,381,1595]" captionTargetId-2="figure-122@9.[151,1436,381,1382]" captionTargetId-3="figure-240@10.[151,1436,669,1829]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionText-0="FIGURE 4. Penia babai. AC: holotype, female (5481); D: male (PBK02). A: labels; B, D: habitus, dorsal side; C: habitus, ventral side." captionText-1="FIGURE 5. Penia babai. AC, EM: holotype, female (5481); D: male (PBK01). A: head, anterior view; B: head, dorsal view; C, D: basal antennomeres of left antenna; E: right maxillary palp, ventral side; F: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); G: left hypomeron (arrow: hypomeron with distinct mesial projection); H: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 5F); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 5F); K: left mid tarsus." captionText-2="FIGURE 6. Penia babai, male (PBK01). A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal side; F: aedeagus, ventral side; G: apical part of left paramera, ventral side." captionText-3="FIGURE 7. Penia babai, females.A, B, E, F: non-type (PBK03); C, D: holotype (5481); G: non-type (PBK04). A: tergite VIII; B: sternite VIII; C: slide; D, E: pregenital segments and genitalia; F: coxites, ventral side; G: vagina and bursa copulatrix (arrow 1: bursa copulatrix posteriorly with a short sac, arrow 2: bursa copulatrix with a sac at the middle)." figureDoi-0="http://doi.org/10.5281/zenodo.10200044" figureDoi-1="http://doi.org/10.5281/zenodo.10200047" figureDoi-2="http://doi.org/10.5281/zenodo.10200049" figureDoi-3="http://doi.org/10.5281/zenodo.10200051" httpUri-0="https://zenodo.org/record/10200044/files/figure.png" httpUri-1="https://zenodo.org/record/10200047/files/figure.png" httpUri-2="https://zenodo.org/record/10200049/files/figure.png" httpUri-3="https://zenodo.org/record/10200051/files/figure.png" pageId="6" pageNumber="307">Figures 47</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31D68B0FFA0CF62FF47ED83FD5BF8E1" pageId="6" pageNumber="307" type="reference_group">
<paragraph id="8BB83B3BFFA0CF62FF47ED83FA9CF95D" blockId="6.[151,1437,1688,1804]" box="[151,1379,1688,1712]" pageId="6" pageNumber="307">
<treatmentCitationGroup id="AB171C15FFA0CF62FF47ED83FA9CF95D" box="[151,1379,1688,1712]" pageId="6" pageNumber="307">
<taxonomicName id="4C0740B8FFA0CF62FF47ED83FDF8F95D" authority="Szombathy, 1910" authorityName="Szombathy" authorityYear="1910" box="[151,519,1688,1712]" class="Insecta" family="Elateridae" genus="Csikia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="dimatoides">
<emphasis id="B973E729FFA0CF62FF47ED83FEB2F95D" box="[151,333,1688,1712]" italics="true" pageId="6" pageNumber="307">Csikia dimatoides</emphasis>
<bibRefCitation id="EF9646CAFFA0CF62FE84ED83FDF8F95D" author="Szombathy, C." box="[340,519,1688,1712]" pageId="6" pageNumber="307" pagination="353 - 360" refId="ref23734" refString="Szombathy, C. (1910) Elaterides nouveaux ou peu connus appartenant au Musee National Hongrois. Annales Musei Nationalis Hungarici, 8, 353 - 360." type="journal article" year="1910">Szombathy, 1910</bibRefCitation>
</taxonomicName>
;
<treatmentCitation id="0AA61D2AFFA0CF62FDC1ED83FD57F95D" author="Kishii, T." box="[529,680,1688,1712]" page="3" pageId="6" pageNumber="307" year="1991">
<bibRefCitation id="EF9646CAFFA0CF62FDC1ED83FD57F95D" author="Kishii, T." box="[529,680,1688,1712]" pageId="6" pageNumber="307" pagination="3 - 32" refId="ref22903" refString="Kishii, T. (1991) Elateridae from Taiwan, with descriptions of some new taxa (5) (Coleoptera). A study of the materials collected by Dr. Kintaro Baba from 1986 to 1989. Transactions of Essa Entomological Society, 71, 3 - 32." type="journal article" year="1991">Kishii, 1991: 3</bibRefCitation>
</treatmentCitation>
(record of female from
<collectingRegion id="49C3F5D9FFA0CF62FC4CED83FBC2F95D" box="[924,1085,1688,1712]" country="Taiwan" name="Kaohsiung" pageId="6" pageNumber="307">Kaohsiung City</collectingRegion>
,
<collectingRegion id="49C3F5D9FFA0CF62FB97ED83FB66F95D" box="[1095,1177,1688,1712]" country="China" name="Taiwan)" pageId="6" pageNumber="307">Taiwan)</collectingRegion>
[misidentification].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB83B3BFFA0CF62FF47EDACFD5BF8E1" blockId="6.[151,1437,1688,1804]" pageId="6" pageNumber="307">
<treatmentCitationGroup id="AB171C15FFA0CF62FF47EDACFD5BF8E1" pageId="6" pageNumber="307">
<treatmentCitation id="0AA61D2AFFA0CF62FF47EDACFE28F922" author="Kishii, T." box="[151,471,1718,1743]" page="211" pageId="6" pageNumber="307" year="1994">
<taxonomicName id="4C0740B8FFA0CF62FF47EDACFE28F922" authority="Kishii, 1994: 211" authorityName="Kishii" authorityPageNumber="211" authorityYear="1994" box="[151,471,1718,1743]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="babai">
<emphasis id="B973E729FFA0CF62FF47EDACFEE9F923" box="[151,278,1718,1742]" italics="true" pageId="6" pageNumber="307">Penia babai</emphasis>
<bibRefCitation id="EF9646CAFFA0CF62FEF0EDADFE28F922" author="Kishii, T." box="[288,471,1718,1743]" pageId="6" pageNumber="307" pagination="211 - 216" refId="ref22955" refString="Kishii, T. (1994) Two new Elaterid-species from Taiwan collected by the late Dr. Kintaro Baba, with a remainder of the deceased (Coleoptera, Elateridae). Special Bulletin of the Essa Entomological Society, 2, 211 - 216." type="journal article" year="1994">Kishii, 1994: 211</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description;
<typeStatus id="54BC8599FFA0CF62FD14EDACFD0FF922" box="[708,752,1719,1743]" pageId="6" pageNumber="307">type</typeStatus>
locality:
<collectingCountry id="F3107BABFFA0CF62FC87EDADFC5BF923" box="[855,932,1718,1742]" name="Taiwan" pageId="6" pageNumber="307">Taiwan</collectingCountry>
,
<collectingRegion id="49C3F5D9FFA0CF62FC61EDADFBA9F923" box="[945,1110,1718,1742]" country="Taiwan" name="Kaohsiung" pageId="6" pageNumber="307">Kaohsiung City</collectingRegion>
, Liouguei District, Mt. Wukon Shan);
<treatmentCitation id="0AA61D2AFFA0CF62FEDFEDCEFE38F900" author="Suzuki, W." box="[271,455,1749,1773]" page="121" pageId="6" pageNumber="307" year="1999">
<bibRefCitation id="EF9646CAFFA0CF62FEDFEDCEFE38F900" author="Suzuki, W." box="[271,455,1749,1773]" pageId="6" pageNumber="307" pagination="1 - 348" refId="ref23700" refString="Suzuki, W. (1999) Catalogue of the family Elateridae (Coleoptera) of Taiwan. Miscellaneous Reports of the Hiwa Museum for Natural History, 38, 1 - 348." type="journal article" year="1999">Suzuki, 1999: 121</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="0AA61D2AFFA0CF62FD9BEDCEFD14F900" author="Cate, P. C." box="[587,747,1749,1773]" page="185" pageId="6" pageNumber="307" year="2007">
<bibRefCitation id="EF9646CAFFA0CF62FD9BEDCEFD14F900" author="Cate, P. C." box="[587,747,1749,1773]" pageId="6" pageNumber="307" refId="ref22578" refString="Cate, P. C. (2007) Family Elateridae. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Vol. 4. Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 89 - 209." type="book" year="2007">Cate, 2007: 185</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="0AA61D2AFFA0CF62FCA0EDCEFBE4F900" author="Platia, G." box="[880,1051,1749,1773]" page="193" pageId="6" pageNumber="307" year="2008">
<bibRefCitation id="EF9646CAFFA0CF62FCA0EDCEFBE4F900" author="Platia, G." box="[880,1051,1749,1773]" pageId="6" pageNumber="307" pagination="187 - 215" refId="ref23321" refString="Platia, G. (2008) Descriptions of new species of click-beetles from the Palearctic region and Taiwan with chorological and synonymical notes (Insecta Coleoptera Elateridae). Quaderno di Studi e Notizie di Storia Naturale della Romagna, 27, 187 - 215." type="journal article" year="2008">Platia, 2008: 193</bibRefCitation>
</treatmentCitation>
(record of male from
<collectingRegion id="49C3F5D9FFA0CF62FB28EDCEFA66F900" box="[1272,1433,1749,1773]" country="Taiwan" name="Kaohsiung" pageId="6" pageNumber="307">Kaohsiung City</collectingRegion>
, Taiwan);
<treatmentCitation id="0AA61D2AFFA0CF62FEF7EDEFFDDBF8E1" ID-DOI="http://doi.org/10.5281/zenodo.5985708" ID-Zenodo-Dep="5985708" author="Kundrata, R. &amp; Musalkova, M. &amp; Kubaczkova, M." box="[295,548,1780,1804]" httpUri="http://treatment.plazi.org/id/A648DA7791619B1CFF5112D0FC4CDC94" page="36" pageId="6" pageNumber="307" year="2018">
<bibRefCitation id="EF9646CAFFA0CF62FEF7EDEFFDDBF8E1" author="Kundrata, R. &amp; Musalkova, M. &amp; Kubaczkova, M." box="[295,548,1780,1804]" pageId="6" pageNumber="307" pagination="1 - 75" refId="ref23051" refString="Kundrata, R., Musalkova, M. &amp; Kubaczkova, M. (2018) Annotated catalogue of the click-beetle tribe Dimini (Coleoptera: Elateridae: Dendrometrinae). Zootaxa, 4412 (1), 1 - 75. https: // doi. org / 10.11646 / zootaxa. 4412.1" type="journal article" year="2018">
Kundrata
<emphasis id="B973E729FFA0CF62FE5CEDEEFE40F8E1" box="[396,447,1780,1804]" italics="true" pageId="6" pageNumber="307">et al.</emphasis>
, 2018: 36
</bibRefCitation>
</treatmentCitation>
(catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31D68B0FFA0CF63FF47EC21FA6DFB7C" lastPageId="7" lastPageNumber="308" pageId="6" pageNumber="307" type="materials_examined">
<paragraph id="8BB83B3BFFA0CF62FF47EC21FF2FF82D" blockId="6.[151,1437,1850,1984]" pageId="6" pageNumber="307">
<emphasis id="B973E729FFA0CF62FF47EC21FEB9F8B9" bold="true" box="[151,326,1850,1876]" pageId="6" pageNumber="307">Type material.</emphasis>
<materialsCitation id="3B6F3166FFA0CF62FE9EEC21FF2FF82D" collectingDate="1989-06-03" collectionCode="OMNH" collectorName="Kintaro Baba &amp; Col. K. Baba" country="Taiwan" county="Female" location="Mt. Wukon Shan" municipality="Liouguei District" pageId="6" pageNumber="307" specimenCount="2" stateProvince="Kaohsiung" typeStatus="holotype">
<emphasis id="B973E729FFA0CF62FE9EEC21FE3DF8B9" bold="true" box="[334,450,1850,1876]" pageId="6" pageNumber="307">
<typeStatus id="54BC8599FFA0CF62FE9EEC21FE42F8B9" box="[334,445,1850,1876]" pageId="6" pageNumber="307" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62D943B7FFA0CF62FE1AEC21FDE0F8B9" box="[458,543,1850,1876]" pageId="6" pageNumber="307">Female</collectingCounty>
,
<collectingCountry id="F3107BABFFA0CF62FDFBEC21FD7FF8B9" box="[555,640,1850,1876]" name="Taiwan" pageId="6" pageNumber="307">Taiwan</collectingCountry>
,
<collectingRegion id="49C3F5D9FFA0CF62FD5CEC21FCC1F8B9" box="[652,830,1850,1876]" country="Taiwan" name="Kaohsiung" pageId="6" pageNumber="307">Kaohsiung City</collectingRegion>
, near
<collectingMunicipality id="6BDCA141FFA0CF62FC53EC21FBB8F8B9" box="[899,1095,1850,1876]" pageId="6" pageNumber="307">Liouguei District</collectingMunicipality>
,
<location id="8ED86DE0FFA0CF62FB83EC20FAE9F8B9" LSID="urn:lsid:plazi:treatment:03AE8A2DFFA0CF6FFF47ED34FC43FCF8:8ED86DE0FFA0CF62FB83EC20FAE9F8B9" box="[1107,1302,1850,1877]" country="Taiwan" county="Female" municipality="Liouguei District" name="Mt. Wukon Shan" pageId="6" pageNumber="307" stateProvince="Kaohsiung">Mt. Wukon Shan</location>
,
<date id="FFB91DFBFFA0CF62FAF2EC21FA67F8B9" box="[1314,1432,1850,1877]" pageId="6" pageNumber="307" value="1989-06-03">
<collectingDate id="EFFDE413FFA0CF62FAF2EC21FA67F8B9" box="[1314,1432,1850,1877]" pageId="6" pageNumber="307" value="1989-06-03">3 VI 1989</collectingDate>
</date>
,
<collectorName id="26F25EEDFFA0CF62FF47EC45FED1F895" box="[151,302,1886,1912]" pageId="6" pageNumber="307">Kintarô Baba</collectorName>
leg. [
<collectionCode id="ED16A3FEFFA0CF62FEBBEC45FE38F895" box="[363,455,1886,1912]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="6" pageNumber="307" type="Museum">OMNH</collectionCode>
; 5481]. Verbatim label data (
<figureCitation id="133C27BEFFA0CF62FCDBEC45FC99F895" box="[779,870,1886,1912]" captionStart="FIGURE 4" captionStartId="7.[151,250,857,881]" captionTargetBox="[151,1434,165,833]" captionTargetId="figure-16@7.[151,1436,159,841]" captionTargetPageId="7" captionText="FIGURE 4. Penia babai. AC: holotype, female (5481); D: male (PBK02). A: labels; B, D: habitus, dorsal side; C: habitus, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200044" httpUri="https://zenodo.org/record/10200044/files/figure.png" pageId="6" pageNumber="307">Fig. 4A</figureCitation>
). “Female symbol”; “
<typeStatus id="54BC8599FFA0CF62FB87EC45FB0CF895" box="[1111,1267,1886,1912]" pageId="6" pageNumber="307" type="holotype">HOLOTYPE</typeStatus>
”; “5481”; “Mt. Wu Kon Shan,/ near Liu kuei,/ S-Taiwan/
<date id="FFB91DFBFFA0CF62FDA5EC99FD0BF871" box="[629,756,1922,1949]" pageId="6" pageNumber="307" value="1989-06-03">
<collectingDate id="EFFDE413FFA0CF62FDA5EC99FD0BF871" box="[629,756,1922,1949]" pageId="6" pageNumber="307" value="1989-06-03">3. VI. 1989</collectingDate>
</date>
/
<collectorName id="26F25EEDFFA0CF62FCD5EC99FC61F871" box="[773,926,1922,1949]" pageId="6" pageNumber="307">Col. K. Baba</collectorName>
”; “
<taxonomicName id="4C0740B8FFA0CF62FC6EEC99FC01F871" box="[958,1022,1922,1948]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="335" phylum="Arthropoda" rank="genus">Penia</taxonomicName>
/ babai/ KISHII,
<taxonomicNameLabel id="A2405A52FFA0CF62FB6EEC98FB06F871" box="[1214,1273,1922,1949]" pageId="6" pageNumber="307" rank="species">sp. n.</taxonomicNameLabel>
/ Det. KISHII, 93”.
</materialsCitation>
</paragraph>
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<emphasis id="B973E729FFA1CF63FF47E842FEEBFC9C" bold="true" box="[151,276,857,881]" pageId="7" pageNumber="308">FIGURE 4.</emphasis>
<taxonomicName id="4C0740B8FFA1CF63FECCE842FE66FC9C" authorityName="Kishii" authorityYear="1994" box="[284,409,857,881]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="babai">
<emphasis id="B973E729FFA1CF63FECCE842FE66FC9C" box="[284,409,857,881]" italics="true" pageId="7" pageNumber="308">Penia babai</emphasis>
</taxonomicName>
. AC: holotype, female (5481); D: male (PBK02). A: labels; B, D: habitus, dorsal side; C: habitus, ventral side.
</paragraph>
</caption>
<paragraph id="8BB83B3BFFA1CF63FF17E8D8FA6DFB7C" blockId="7.[151,1437,963,2034]" pageId="7" pageNumber="308">
<emphasis id="B973E729FFA1CF63FF17E8D8FE53FC30" bold="true" box="[199,428,963,989]" pageId="7" pageNumber="308">Non-type material.</emphasis>
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<collectingCountry id="F3107BABFFA1CF63FE66E8D8FDF3FC30" box="[438,524,963,989]" name="Taiwan" pageId="7" pageNumber="308">Taiwan</collectingCountry>
.
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,
<collectingRegion id="49C3F5D9FFA1CF63FDA8E8DFFCD6FC30" box="[632,809,963,990]" country="Taiwan" name="Nantou" pageId="7" pageNumber="308">Nantou County</collectingRegion>
,
<collectingMunicipality id="6BDCA141FFA1CF63FCE7E8D8FC03FC30" box="[823,1020,963,989]" pageId="7" pageNumber="308">Renai Township</collectingMunicipality>
,
<location id="8ED86DE0FFA1CF63FBDAE8D8FB69FC30" LSID="urn:lsid:plazi:treatment:03AE8A2DFFA0CF6FFF47ED34FC43FCF8:8ED86DE0FFA1CF63FBDAE8D8FB69FC30" box="[1034,1174,963,989]" country="Taiwan" municipality="Ren'ai Township" name="Nanshan Xi" pageId="7" pageNumber="308" stateProvince="Nantou">Nanshan Xi</location>
,
<date id="FFB91DFBFFA1CF63FB74E8D8FB18FC33" box="[1188,1255,963,990]" pageId="7" pageNumber="308" value="1985-04-19">
19
<collectionCode id="ED16A3FEFFA1CF63FB1AE8DFFB18FC33" box="[1226,1255,964,990]" pageId="7" pageNumber="308">IV</collectionCode>
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<materialsCitation id="3B6F3166FFA1CF63FB21E8D8FE26FBEC" collectionCode="OMNH" country="Taiwan" pageId="7" pageNumber="308" specimenCode="PBK01" specimenCount="1">
1985, collector unknown [
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;
<specimenCode id="DBA19340FFA1CF63FEAAE8FCFE2DFBEC" box="[378,466,999,1025]" collectionCode="PBK" pageId="7" pageNumber="308">PBK01</specimenCode>
]
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;
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<specimenCount id="9D01F0B2FFA1CF63FE37E8FCFDC7FBEC" box="[487,568,999,1025]" count="1" pageId="7" pageNumber="308" type="male">1 male</specimenCount>
,
<collectingRegion id="49C3F5D9FFA1CF63FD96E8FCFD05FBEC" box="[582,762,999,1025]" country="Taiwan" name="Kaohsiung" pageId="7" pageNumber="308">Kaohsiung City</collectingRegion>
,
<collectingRegion id="49C3F5D9FFA1CF63FCD8E8F3FC36FBEC" box="[776,969,999,1026]" country="Taiwan" name="Taoyuan" pageId="7" pageNumber="308">Taoyuan District</collectingRegion>
,
<location id="8ED86DE0FFA1CF63FC06E8FCFBC3FBEC" LSID="urn:lsid:plazi:treatment:03AE8A2DFFA0CF6FFF47ED34FC43FCF8:8ED86DE0FFA1CF63FC06E8FCFBC3FBEC" box="[982,1084,999,1025]" country="Taiwan" name="Tengjhih" pageId="7" pageNumber="308" stateProvince="Kaohsiung">Tengjhih</location>
,
<date id="FFB91DFBFFA1CF63FB9AE8FCFB89FBEF" box="[1098,1142,999,1026]" pageId="7" pageNumber="308" value="1985-05-05">
5
<collectionCode id="ED16A3FEFFA1CF63FBB2E8F3FB89FBEF" box="[1122,1142,1000,1026]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="308" type="Museum">V</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="3B6F3166FFA1CF63FBAFE8FCFE99FBC8" collectionCode="OMNH" country="Taiwan" pageId="7" pageNumber="308" specimenCode="PBK02" specimenCount="1">
1985, collector unknown [
<collectionCode id="ED16A3FEFFA1CF63FF4EEF10FF05FBC8" box="[158,250,1035,1061]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="7" pageNumber="308" type="Museum">OMNH</collectionCode>
;
<specimenCode id="DBA19340FFA1CF63FED6EF10FEA0FBC8" box="[262,351,1035,1061]" collectionCode="PBK" pageId="7" pageNumber="308">PBK02</specimenCode>
]
</materialsCitation>
;
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<specimenCount id="9D01F0B2FFA1CF63FEA2EF10FE27FBC8" box="[370,472,1035,1061]" count="1" pageId="7" pageNumber="308" type="female">1 female</specimenCount>
,
<collectingRegion id="49C3F5D9FFA1CF63FE35EF10FD67FBC8" box="[485,664,1035,1061]" country="Taiwan" name="Kaohsiung" pageId="7" pageNumber="308">Kaohsiung City</collectingRegion>
,
<location id="8ED86DE0FFA1CF63FD75EF10FC96FBC8" LSID="urn:lsid:plazi:treatment:03AE8A2DFFA0CF6FFF47ED34FC43FCF8:8ED86DE0FFA1CF63FD75EF10FC96FBC8" box="[677,873,1035,1061]" country="Taiwan" name="Liouguei District" pageId="7" pageNumber="308" stateProvince="Kaohsiung">Liouguei District</location>
,
<date id="FFB91DFBFFA1CF63FCA6EF10FC0AFBC8" box="[886,1013,1035,1062]" pageId="7" pageNumber="308" value="1981-07-04">
<collectingDate id="EFFDE413FFA1CF63FCA6EF10FC0AFBC8" box="[886,1013,1035,1062]" pageId="7" pageNumber="308" value="1981-07-04">4 VII 1981</collectingDate>
</date>
,
<collectionCode id="ED16A3FEFFA1CF63FBD2EF17FBEBFBCB" box="[1026,1044,1036,1062]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="308">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B6F3166FFA1CF63FBF1EF10FA67FBC8" box="[1057,1432,1035,1061]" collectionCode="OMNH" collectorName="Kamakari" country="Taiwan" pageId="7" pageNumber="308" specimenCode="PBK03" specimenCount="1">
<collectorName id="26F25EEDFFA1CF63FBF1EF10FB70FBC8" box="[1057,1167,1035,1061]" pageId="7" pageNumber="308">Kamakari</collectorName>
leg. [
<collectionCode id="ED16A3FEFFA1CF63FB00EF10FAD3FBC8" box="[1232,1324,1035,1061]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="7" pageNumber="308" type="Museum">OMNH</collectionCode>
;
<specimenCode id="DBA19340FFA1CF63FAE9EF10FA6EFBC8" box="[1337,1425,1035,1061]" collectionCode="PBK" pageId="7" pageNumber="308">PBK03</specimenCode>
]
</materialsCitation>
;
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<specimenCount id="9D01F0B2FFA1CF63FF47EF34FF04FBA4" box="[151,251,1071,1097]" count="1" pageId="7" pageNumber="308" type="female">1 female</specimenCount>
,
<collectingRegion id="49C3F5D9FFA1CF63FED7EF34FE47FBA4" box="[263,440,1071,1097]" country="Taiwan" name="Kaohsiung" pageId="7" pageNumber="308">Kaohsiung City</collectingRegion>
,
<collectingMunicipality id="6BDCA141FFA1CF63FE14EF34FD78FBA4" box="[452,647,1071,1097]" pageId="7" pageNumber="308">Liouguei District</collectingMunicipality>
,
<location id="8ED86DE0FFA1CF63FD42EF2BFCB1FBA4" LSID="urn:lsid:plazi:treatment:03AE8A2DFFA0CF6FFF47ED34FC43FCF8:8ED86DE0FFA1CF63FD42EF2BFCB1FBA4" box="[658,846,1071,1098]" country="Taiwan" municipality="Liouguei District" name="Mt. Nanho Shan" pageId="7" pageNumber="308" stateProvince="Kaohsiung">Mt. Nanho Shan</location>
,
<date id="FFB91DFBFFA1CF63FC89EF34FC6FFBA7" box="[857,912,1071,1098]" pageId="7" pageNumber="308" value="1985-05-18">
18
<collectionCode id="ED16A3FEFFA1CF63FCACEF2BFC6FFBA7" box="[892,912,1072,1098]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="308" type="Museum">V</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="3B6F3166FFA1CF63FC47EF34FA7CFBA4" box="[919,1411,1071,1098]" collectionCode="OMNH" country="Taiwan" pageId="7" pageNumber="308" specimenCode="PBK04" specimenCount="1">
1985, collector unknown [
<collectionCode id="ED16A3FEFFA1CF63FB6CEF34FAE7FBA4" box="[1212,1304,1071,1097]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="7" pageNumber="308" type="Museum">OMNH</collectionCode>
;
<specimenCode id="DBA19340FFA1CF63FAF3EF34FA82FBA4" box="[1315,1405,1071,1097]" collectionCode="PBK" pageId="7" pageNumber="308">PBK04</specimenCode>
]
</materialsCitation>
;
<materialsCitation id="3B6F3166FFA1CF63FA5EEF34FB61FB83" collectionCode="V" country="Taiwan" location="Chungshinrun" municipality="Mt. Pao-shan" pageId="7" pageNumber="308" specimenCount="1" specimenCount-male="1" stateProvince="Kaohsiung">
<specimenCount id="9D01F0B2FFA1CF63FA5EEF34FF2FFB80" count="1" pageId="7" pageNumber="308" type="male">1 male</specimenCount>
,
<collectingRegion id="49C3F5D9FFA1CF63FF0BEF48FE73FB80" box="[219,396,1107,1133]" country="Taiwan" name="Kaohsiung" pageId="7" pageNumber="308">Kaohsiung City</collectingRegion>
,
<collectingRegion id="49C3F5D9FFA1CF63FE47EF4FFDAAFB80" box="[407,597,1107,1134]" country="Taiwan" name="Taoyuan" pageId="7" pageNumber="308">Taoyuan District</collectingRegion>
,
<collectingMunicipality id="6BDCA141FFA1CF63FDB0EF4FFD0AFB80" box="[608,757,1107,1134]" pageId="7" pageNumber="308">Mt. Pao-shan</collectingMunicipality>
(as
<location id="8ED86DE0FFA1CF63FCF3EF48FC3DFB80" LSID="urn:lsid:plazi:treatment:03AE8A2DFFA0CF6FFF47ED34FC43FCF8:8ED86DE0FFA1CF63FCF3EF48FC3DFB80" box="[803,962,1107,1133]" country="Taiwan" municipality="Mt. Pao-shan" name="Chungshinrun" pageId="7" pageNumber="308" stateProvince="Kaohsiung">Chungshinrun</location>
in data label),
<date id="FFB91DFBFFA1CF63FBB8EF48FB61FB83" box="[1128,1182,1107,1134]" pageId="7" pageNumber="308" value="1975-05-20">
20
<collectionCode id="ED16A3FEFFA1CF63FB5AEF4FFB61FB83" box="[1162,1182,1108,1134]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="308" type="Museum">V</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="3B6F3166FFA1CF63FB74EF48FE9BFB7F" collectionCode="OMNH" collectorName="K. Matsuda" country="Taiwan" pageId="7" pageNumber="308" specimenCode="PBK05" specimenCount="1">
1975,
<collectorName id="26F25EEDFFA1CF63FB3AEF4FFA93FB80" box="[1258,1388,1107,1134]" pageId="7" pageNumber="308">K. Matsuda</collectorName>
leg. [
<collectionCode id="ED16A3FEFFA1CF63FF4EEF6CFF05FB7C" box="[158,250,1143,1169]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="7" pageNumber="308" type="Museum">OMNH</collectionCode>
;
<specimenCode id="DBA19340FFA1CF63FED5EF6CFEA1FB7F" box="[261,350,1143,1170]" collectionCode="PBK" pageId="7" pageNumber="308">PBK05</specimenCode>
]
</materialsCitation>
;
<materialsCitation id="3B6F3166FFA1CF63FEBFEF6CFC01FB7F" box="[367,1022,1143,1170]" collectionCode="V" country="Taiwan" location="Gokang Shan" municipality="Liouguei" pageId="7" pageNumber="308" specimenCount="1" specimenCount-male="1" stateProvince="Kaohsiung">
<specimenCount id="9D01F0B2FFA1CF63FEBFEF6CFE42FB7C" box="[367,445,1143,1169]" count="1" pageId="7" pageNumber="308" type="male">1 male</specimenCount>
,
<collectingRegion id="49C3F5D9FFA1CF63FE19EF6CFD85FB7C" box="[457,634,1143,1169]" country="Taiwan" name="Kaohsiung" pageId="7" pageNumber="308">Kaohsiung City</collectingRegion>
, near
<collectingMunicipality id="6BDCA141FFA1CF63FD6DEF6CFCDAFB7C" box="[701,805,1143,1169]" pageId="7" pageNumber="308">Liouguei</collectingMunicipality>
,
<location id="8ED86DE0FFA1CF63FCE0EF6CFC34FB7C" LSID="urn:lsid:plazi:treatment:03AE8A2DFFA0CF6FFF47ED34FC43FCF8:8ED86DE0FFA1CF63FCE0EF6CFC34FB7C" box="[816,971,1143,1169]" country="Taiwan" municipality="Liouguei" name="Gokang Shan" pageId="7" pageNumber="308" stateProvince="Kaohsiung">Gokang Shan</location>
,
<date id="FFB91DFBFFA1CF63FC06EF63FC01FB7F" box="[982,1022,1144,1170]" pageId="7" pageNumber="308" value="1991-05-07">
7
<collectionCode id="ED16A3FEFFA1CF63FC3AEF63FC01FB7F" box="[1002,1022,1144,1170]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="308" type="Museum">V</collectionCode>
</date>
</materialsCitation>
1991,
<collectorName id="26F25EEDFFA1CF63FB9BEF63FB5AFB7C" box="[1099,1189,1143,1170]" pageId="7" pageNumber="308">M. Yagi</collectorName>
leg. [
<collectionCode id="ED16A3FEFFA1CF63FB32EF6CFAC1FB7C" box="[1250,1342,1143,1169]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="7" pageNumber="308" type="Museum">OMNH</collectionCode>
; 6689].
</paragraph>
</subSubSection>
<subSubSection id="C31D68B0FFA1CF63FF17EF80FDC0F8F4" pageId="7" pageNumber="308" type="diagnosis">
<paragraph id="8BB83B3BFFA1CF63FF17EF80FCF8F988" blockId="7.[151,1437,963,2034]" pageId="7" pageNumber="308">
<emphasis id="B973E729FFA1CF63FF17EF80FEBDFB58" bold="true" box="[199,322,1179,1205]" pageId="7" pageNumber="308">Diagnosis.</emphasis>
This species is characterized by the following features: eyes 0.3 x longer than interocular distance in dorsal view; antennae extending beyond pronotum posterior lateral apices by antennomere VI or VII, surpassing elytral half by antennomere XI; antennomeres III distinctly longer than II; IV 1.31.4 x longer than III, 0.80.9 x longer than IIIII combined; apical maxillary palpomere 1.92.4 x longer than wide, shorter than maximum length of eye; pronotum straightly and slightly narrowed ahead of hind angles; posterior edge of pronotum with sublateral incisions; hind angles of pronotum broad, weakly protruding posterolaterad; hypomeron with distinct mesial projection; anterior angle of hypomeron almost right angle; hind angle of hypomeron broadly triangular; scutellar shield almost as long as wide; mesosternal process between mesocoxae distinctly higher than mesocoxae, visible in lateral view; posterior edge of mesosternal process 0.20.25 x wider than total width of mesosternum; elytron 3.13.6 x longer than wide, 2.52.7 x longer than pronotum length; abdominal ventrite V semicircular, rounded apically; phallobase 0.9 x longer than wide; apex of parameres beyond preapical expansions large triangular; apex length 0.40.6 x width of parameres at expansions in ventral side; spiculum ventrale 5.6 x longer than length of sternite VIII; ovipositor longer than length of abdomen.
</paragraph>
<paragraph id="8BB83B3BFFA1CF63FF17ED74FDC0F8F4" blockId="7.[151,1437,963,2034]" pageId="7" pageNumber="308">
This species is similar to
<taxonomicName id="4C0740B8FFA1CF63FE0DED6BFDD5F964" authorityName="Arimoto" authorityYear="2023" box="[477,554,1648,1674]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="tsou">
<emphasis id="B973E729FFA1CF63FE0DED6BFE11F964" box="[477,494,1648,1673]" italics="true" pageId="7" pageNumber="308">P</emphasis>
.
<emphasis id="B973E729FFA1CF63FE2BED6BFDD5F964" box="[507,554,1648,1673]" italics="true" pageId="7" pageNumber="308">tsou</emphasis>
</taxonomicName>
in features of the eyes, the basal antennomeres, the apical maxillary palpomere, the hind angles of the pronotum, the anterior and hind angles of the hypomeron, the scutellar shield, the mesosternal process between mesocoxae, the elytron, and the abdominal ventrite V. It is distinguished by antenna length, the shapes of the pronotum and apex of parameres, and the degree of development of the mesial projection of the hypomeron (see diagnosis of
<taxonomicName id="4C0740B8FFA1CF63FE31EC1BFDD0F8F4" authorityName="Arimoto" authorityYear="2023" box="[481,559,1792,1818]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="tsou">
<emphasis id="B973E729FFA1CF63FE31EC1BFE0DF8F4" box="[481,498,1792,1817]" italics="true" pageId="7" pageNumber="308">P</emphasis>
.
<emphasis id="B973E729FFA1CF63FDD0EC1BFDD0F8F4" box="[512,559,1792,1817]" italics="true" pageId="7" pageNumber="308">tsou</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C31D68B0FFA1CF6FFF17EC38FD9CFD1C" lastPageId="11" lastPageNumber="312" pageId="7" pageNumber="308" type="description">
<paragraph id="8BB83B3BFFA1CF63FF17EC38FBE9F81C" blockId="7.[151,1437,963,2034]" pageId="7" pageNumber="308">
<emphasis id="B973E729FFA1CF63FF17EC38FE3FF8D0" bold="true" box="[199,448,1827,1853]" pageId="7" pageNumber="308">Measurements. Male</emphasis>
(n=4). BL: 8.6210.6, BW: 3.324.24, MAE: 1.341.67, MBE: 0.831.10, OI: 151162, PL: 2.242.75, PML: 1.752.13, PW: 2.693.52, PAW: 1.461.84, PLI: 78.283.3, PWI: 178191, EL: 5.677.43, EW: 1.672.06, EI: 253270, BI: 253270.
<emphasis id="B973E729FFA1CF63FD44EC70FD14F868" bold="true" box="[660,747,1899,1925]" pageId="7" pageNumber="308">Female</emphasis>
(n=3;
<typeStatus id="54BC8599FFA1CF63FCECEC70FC61F868" box="[828,926,1899,1925]" pageId="7" pageNumber="308" type="holotype">holotype</typeStatus>
in parentheses). BL: 7.839.23 (8.30), BW: 3.293.86 (3.48), MAE: 1.321.48 (1.40), MBE: 0.830.97 (0.91), OI: 152160 (154), PL: 2.202.58 (2.32), PML: 1.722.06 (1.83), PW: 2.693.15 (2.80), PAW: 1.451.73 (1.54), PLI: 81.682.9 (82.9), PWI: 182186 (182), EL: 5.406.49 (5.75), EW: 1.711.94 (1.77), EI: 315334 (325), BI: 246251 (247).
</paragraph>
<paragraph id="8BB83B3BFFAECF6CFF17EB8DFD6BFE88" blockId="8.[151,1436,150,357]" pageId="8" pageNumber="309">
<emphasis id="B973E729FFAECF6CFF17EB8DFE8AFF5D" bold="true" box="[199,373,150,176]" pageId="8" pageNumber="309">Redescription.</emphasis>
Body broad, widest ahead of elytral midlength (
<figureCitation id="133C27BEFFAECF6CFC5EEB8CFC1AFF5C" box="[910,997,151,177]" captionStart="FIGURE 4" captionStartId="7.[151,250,857,881]" captionTargetBox="[151,1434,165,833]" captionTargetId="figure-16@7.[151,1436,159,841]" captionTargetPageId="7" captionText="FIGURE 4. Penia babai. AC: holotype, female (5481); D: male (PBK02). A: labels; B, D: habitus, dorsal side; C: habitus, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200044" httpUri="https://zenodo.org/record/10200044/files/figure.png" pageId="8" pageNumber="309">Fig. 4B</figureCitation>
); surface generally smooth; interspaces between punctures distinctly larger than fine puncture diameter (
<figureCitation id="133C27BEFFAECF6CFCA1EBA0FC13FF38" box="[881,1004,187,213]" captionStart="FIGURE 4" captionStartId="7.[151,250,857,881]" captionTargetBox="[151,1434,165,833]" captionTargetId="figure-16@7.[151,1436,159,841]" captionTargetPageId="7" captionText="FIGURE 4. Penia babai. AC: holotype, female (5481); D: male (PBK02). A: labels; B, D: habitus, dorsal side; C: habitus, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200044" httpUri="https://zenodo.org/record/10200044/files/figure.png" pageId="8" pageNumber="309">Fig. 4B, C</figureCitation>
). Color. Body light orange to dark red (
<figureCitation id="133C27BEFFAECF6CFF4FEBC4FEE0FF14" box="[159,287,223,249]" captionStart="FIGURE 4" captionStartId="7.[151,250,857,881]" captionTargetBox="[151,1434,165,833]" captionTargetId="figure-16@7.[151,1436,159,841]" captionTargetPageId="7" captionText="FIGURE 4. Penia babai. AC: holotype, female (5481); D: male (PBK02). A: labels; B, D: habitus, dorsal side; C: habitus, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200044" httpUri="https://zenodo.org/record/10200044/files/figure.png" pageId="8" pageNumber="309">Fig. 4B, D</figureCitation>
). Lateral margin of elytra paler. External edge of mandible, lateral and posterior edges of pronotum, posterior edge of prosternum, outer and posterior edges of hypomere, external edge of scutellum, posterior edge of mesosternum and anterior edge of elytra black. Antennae reddish brown or black. Legs light orange to reddish brown. Body covered with long yellow setae.
</paragraph>
<caption id="DF786BB3FFAECF6CFF47ED49FEB0F8AF" ID-DOI="http://doi.org/10.5281/zenodo.10200047" ID-Zenodo-Dep="10200047" httpUri="https://zenodo.org/record/10200047/files/figure.png" pageId="8" pageNumber="309" startId="8.[151,250,1618,1642]" targetBox="[151,1436,381,1595]" targetPageId="8" targetType="figure">
<paragraph id="8BB83B3BFFAECF6CFF47ED49FEB0F8AF" blockId="8.[151,1436,1618,1858]" pageId="8" pageNumber="309">
<emphasis id="B973E729FFAECF6CFF47ED49FEEDF986" bold="true" box="[151,274,1618,1643]" pageId="8" pageNumber="309">FIGURE 5.</emphasis>
<taxonomicName id="4C0740B8FFAECF6CFEC9ED48FE6BF987" authorityName="Kishii" authorityYear="1994" box="[281,404,1618,1642]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="babai">
<emphasis id="B973E729FFAECF6CFEC9ED48FE6BF987" box="[281,404,1618,1642]" italics="true" pageId="8" pageNumber="309">Penia babai</emphasis>
</taxonomicName>
. AC, EM: holotype, female (5481); D: male (PBK01). A: head, anterior view; B: head, dorsal view; C, D: basal antennomeres of left antenna; E: right maxillary palp, ventral side; F: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); G: left hypomeron (arrow: hypomeron with distinct mesial projection); H: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 5F); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 5F); K: left mid tarsus.
</paragraph>
</caption>
<paragraph id="8BB83B3BFFAECF6DFF17EC6AFBF0FE88" blockId="8.[151,1437,1904,2039]" lastBlockId="9.[151,1437,151,357]" lastPageId="9" lastPageNumber="310" pageId="8" pageNumber="309">
<materialsCitation id="3B6F3166FFAECF6DFF17EC6AFBF4FE88" collectionCode="VI, II, III, IV, V" collectorName="X. Mandible &amp; Apical" lastPageId="9" lastPageNumber="310" pageId="8" pageNumber="309" specimenCode="IV 2.6, 3.5, XI 4.1, 5.6, PBK04" specimenCount="1" typeStatus="holotype">
Head. Frons flatted medially (
<figureCitation id="133C27BEFFAECF6CFDC5EC6AFD71F866" box="[533,654,1904,1931]" captionStart="FIGURE 5" captionStartId="8.[151,250,1618,1642]" captionTargetBox="[151,1436,381,1595]" captionTargetId="figure-115@8.[151,1436,381,1595]" captionTargetPageId="8" captionText="FIGURE 5. Penia babai. AC, EM: holotype, female (5481); D: male (PBK01). A: head, anterior view; B: head, dorsal view; C, D: basal antennomeres of left antenna; E: right maxillary palp, ventral side; F: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); G: left hypomeron (arrow: hypomeron with distinct mesial projection); H: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 5F); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 5F); K: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200047" httpUri="https://zenodo.org/record/10200047/files/figure.png" pageId="8" pageNumber="309">Fig. 5A, B</figureCitation>
); frontal carina not complete (
<figureCitation id="133C27BEFFAECF6CFC0FEC6AFBC6F866" box="[991,1081,1904,1931]" captionStart="FIGURE 5" captionStartId="8.[151,250,1618,1642]" captionTargetBox="[151,1436,381,1595]" captionTargetId="figure-115@8.[151,1436,381,1595]" captionTargetPageId="8" captionText="FIGURE 5. Penia babai. AC, EM: holotype, female (5481); D: male (PBK01). A: head, anterior view; B: head, dorsal view; C, D: basal antennomeres of left antenna; E: right maxillary palp, ventral side; F: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); G: left hypomeron (arrow: hypomeron with distinct mesial projection); H: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 5F); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 5F); K: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200047" httpUri="https://zenodo.org/record/10200047/files/figure.png" pageId="8" pageNumber="309">Fig. 5A</figureCitation>
); frontal margin rectangular but broadly rounded apically in dorsal view (
<figureCitation id="133C27BEFFAECF6CFDA0EC8EFD35F842" box="[624,714,1940,1967]" captionStart="FIGURE 5" captionStartId="8.[151,250,1618,1642]" captionTargetBox="[151,1436,381,1595]" captionTargetId="figure-115@8.[151,1436,381,1595]" captionTargetPageId="8" captionText="FIGURE 5. Penia babai. AC, EM: holotype, female (5481); D: male (PBK01). A: head, anterior view; B: head, dorsal view; C, D: basal antennomeres of left antenna; E: right maxillary palp, ventral side; F: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); G: left hypomeron (arrow: hypomeron with distinct mesial projection); H: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 5F); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 5F); K: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200047" httpUri="https://zenodo.org/record/10200047/files/figure.png" pageId="8" pageNumber="309">Fig. 5B</figureCitation>
); frontoclypeal region protruding beyond base of labrum. Eyes relatively normal in convexity, 0.3 x longer than interocular distance in dorsal view (
<figureCitation id="133C27BEFFAECF6CFB80ECA2FB57F83E" box="[1104,1192,1976,2003]" captionStart="FIGURE 5" captionStartId="8.[151,250,1618,1642]" captionTargetBox="[151,1436,381,1595]" captionTargetId="figure-115@8.[151,1436,381,1595]" captionTargetPageId="8" captionText="FIGURE 5. Penia babai. AC, EM: holotype, female (5481); D: male (PBK01). A: head, anterior view; B: head, dorsal view; C, D: basal antennomeres of left antenna; E: right maxillary palp, ventral side; F: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); G: left hypomeron (arrow: hypomeron with distinct mesial projection); H: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 5F); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 5F); K: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200047" httpUri="https://zenodo.org/record/10200047/files/figure.png" pageId="8" pageNumber="309">Fig. 5B</figureCitation>
). Antennae extending beyond pronotum posterior lateral apices by antennomere
<collectionCode id="ED16A3FEFFAECF6CFCF6ECC6FCBBF81A" box="[806,836,2013,2039]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="8" pageNumber="309">VI</collectionCode>
in
<emphasis id="B973E729FFAECF6CFCB8ECC6FC5DF81A" bold="true" box="[872,930,2013,2039]" pageId="8" pageNumber="309">male</emphasis>
and by VII in
<emphasis id="B973E729FFAECF6CFB97ECC7FB69F81B" bold="true" box="[1095,1174,2012,2038]" pageId="8" pageNumber="309">female</emphasis>
, surpassing elytral half by antennomere XI; antennomeres longer than wide;
<collectionCode id="ED16A3FEFFAFCF6DFD3EEB8CFCFEFF5C" box="[750,769,151,177]" pageId="9" pageNumber="310">II</collectionCode>
obconical, shortest, 1.21.7 x longer than wide;
<collectionCode id="ED16A3FEFFAFCF6DFAF8EB8CFABBFF5C" box="[1320,1348,151,177]" pageId="9" pageNumber="310">III</collectionCode>
weakly serrated, 2.12.4 x longer than wide, 1.72.0 x longer than
<collectionCode id="ED16A3FEFFAFCF6DFC81EBA0FC98FF38" box="[849,871,187,213]" pageId="9" pageNumber="310">II</collectionCode>
;
<collectionCode id="ED16A3FEFFAFCF6DFCA5EBA0FC6CFF38" box="[885,915,187,213]" pageId="9" pageNumber="310">IV</collectionCode>
XI filiform;
<specimenCode id="DBA19340FFAFCF6DFBE3EBA0FB80FF38" box="[1075,1151,186,213]" collectionCode="IV" pageId="9" pageNumber="310">IV 2.6</specimenCode>
<specimenCode id="DBA19340FFAFCF6DFB5EEBA1FB4EFF38" box="[1166,1201,186,213]" collectionCode="IV" pageId="9" pageNumber="310">3.5</specimenCode>
x longer than wide, 1.31.4 x longer than
<collectionCode id="ED16A3FEFFAFCF6DFE5FEBC4FE51FF14" box="[399,430,223,249]" pageId="9" pageNumber="310">III</collectionCode>
, 0.80.9 x longer than
<collectionCode id="ED16A3FEFFAFCF6DFD7AEBC4FD43FF14" box="[682,700,223,249]" pageId="9" pageNumber="310">II</collectionCode>
<collectionCode id="ED16A3FEFFAFCF6DFD1BEBC4FD18FF14" box="[715,743,223,249]" pageId="9" pageNumber="310">III</collectionCode>
combined (
<figureCitation id="133C27BEFFAFCF6DFCB9EBC4FC1FFF14" box="[873,992,222,249]" captionStart="FIGURE 5" captionStartId="8.[151,250,1618,1642]" captionTargetBox="[151,1436,381,1595]" captionTargetId="figure-115@8.[151,1436,381,1595]" captionTargetPageId="8" captionText="FIGURE 5. Penia babai. AC, EM: holotype, female (5481); D: male (PBK01). A: head, anterior view; B: head, dorsal view; C, D: basal antennomeres of left antenna; E: right maxillary palp, ventral side; F: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); G: left hypomeron (arrow: hypomeron with distinct mesial projection); H: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 5F); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 5F); K: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200047" httpUri="https://zenodo.org/record/10200047/files/figure.png" pageId="9" pageNumber="310">Fig. 5C, D</figureCitation>
);
<collectionCode id="ED16A3FEFFAFCF6DFC24EBC4FBF7FF14" box="[1012,1032,223,249]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="9" pageNumber="310" type="Museum">V</collectionCode>
3.03.6 x longer than wide, 1.01.1 x longer than
<collectionCode id="ED16A3FEFFAFCF6DFEE7EA18FEA9FEF0" box="[311,342,259,285]" pageId="9" pageNumber="310">IV</collectionCode>
;
<specimenCode id="DBA19340FFAFCF6DFEB3EA18FE53FEF0" box="[355,428,258,285]" collectionCode="IV" pageId="9" pageNumber="310">XI 4.1</specimenCode>
<specimenCode id="DBA19340FFAFCF6DFE6BEA19FE20FEF0" box="[443,479,258,285]" collectionCode="IV" pageId="9" pageNumber="310">5.6</specimenCode>
x longer than wide, 1.01.2 x longer than
<collectorName id="26F25EEDFFAFCF6DFC02EA18FB9EFEF0" box="[978,1121,259,285]" pageId="9" pageNumber="310">X. Mandible</collectorName>
bidentate (
<figureCitation id="133C27BEFFAFCF6DFB32EA18FAC1FEF0" box="[1250,1342,258,285]" captionStart="FIGURE 5" captionStartId="8.[151,250,1618,1642]" captionTargetBox="[151,1436,381,1595]" captionTargetId="figure-115@8.[151,1436,381,1595]" captionTargetPageId="8" captionText="FIGURE 5. Penia babai. AC, EM: holotype, female (5481); D: male (PBK01). A: head, anterior view; B: head, dorsal view; C, D: basal antennomeres of left antenna; E: right maxillary palp, ventral side; F: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); G: left hypomeron (arrow: hypomeron with distinct mesial projection); H: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 5F); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 5F); K: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200047" httpUri="https://zenodo.org/record/10200047/files/figure.png" pageId="9" pageNumber="310">Fig. 5A</figureCitation>
).
<collectorName id="26F25EEDFFAFCF6DFA82EA18FA62FEF0" box="[1362,1437,259,285]" pageId="9" pageNumber="310">Apical</collectorName>
maxillary palpomere semicircular or triangular in
<typeStatus id="54BC8599FFAFCF6DFD19EA3CFCD4FEAC" box="[713,811,295,321]" pageId="9" pageNumber="310" type="holotype">holotype</typeStatus>
(
<figureCitation id="133C27BEFFAFCF6DFCEAEA3CFC70FEAC" box="[826,911,294,321]" captionStart="FIGURE 5" captionStartId="8.[151,250,1618,1642]" captionTargetBox="[151,1436,381,1595]" captionTargetId="figure-115@8.[151,1436,381,1595]" captionTargetPageId="8" captionText="FIGURE 5. Penia babai. AC, EM: holotype, female (5481); D: male (PBK01). A: head, anterior view; B: head, dorsal view; C, D: basal antennomeres of left antenna; E: right maxillary palp, ventral side; F: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); G: left hypomeron (arrow: hypomeron with distinct mesial projection); H: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 5F); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 5F); K: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200047" httpUri="https://zenodo.org/record/10200047/files/figure.png" pageId="9" pageNumber="310">Fig. 5E</figureCitation>
) and left side of non-type (
<specimenCode id="DBA19340FFAFCF6DFB11EA3CFAE6FEAC" box="[1217,1305,295,321]" collectionCode="PBK" pageId="9" pageNumber="310">PBK04</specimenCode>
), 1.92.4 x longer than wide, shorter than maximum length of eyes; anterior edge rounded
</materialsCitation>
.
</paragraph>
<caption id="DF786BB3FFAFCF6DFF47EE66FC6EFA54" ID-DOI="http://doi.org/10.5281/zenodo.10200049" ID-Zenodo-Dep="10200049" httpUri="https://zenodo.org/record/10200049/files/figure.png" pageId="9" pageNumber="310" startId="9.[151,250,1405,1429]" targetBox="[151,1431,381,1377]" targetPageId="9" targetType="figure">
<paragraph id="8BB83B3BFFAFCF6DFF47EE66FC6EFA54" blockId="9.[151,1436,1405,1465]" pageId="9" pageNumber="310">
<emphasis id="B973E729FFAFCF6DFF47EE66FEECFA78" bold="true" box="[151,275,1405,1429]" pageId="9" pageNumber="310">FIGURE 6.</emphasis>
<taxonomicName id="4C0740B8FFAFCF6DFECAEE65FE6AFA78" authorityName="Kishii" authorityYear="1994" box="[282,405,1405,1429]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="babai">
<emphasis id="B973E729FFAFCF6DFECAEE65FE6AFA78" box="[282,405,1405,1429]" italics="true" pageId="9" pageNumber="310">Penia babai</emphasis>
</taxonomicName>
, male (PBK01). A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal side; F: aedeagus, ventral side; G: apical part of left paramera, ventral side.
</paragraph>
</caption>
<paragraph id="8BB83B3BFFAFCF6EFF17EEFCFE8EFD68" blockId="9.[151,1437,1511,2042]" lastBlockId="10.[151,1437,150,645]" lastPageId="10" lastPageNumber="311" pageId="9" pageNumber="310">
Prothorax. Pronotum hexagonal, 0.8 x longer than wide, roundly widening anteriorly, straightly and moderately narrowed ahead of hind angles, widest just ahead of posterior lateral apices (
<figureCitation id="133C27BEFFAFCF6DFBC0ED10FB94F9CB" box="[1040,1131,1547,1574]" captionStart="FIGURE 4" captionStartId="7.[151,250,857,881]" captionTargetBox="[151,1434,165,833]" captionTargetId="figure-16@7.[151,1436,159,841]" captionTargetPageId="7" captionText="FIGURE 4. Penia babai. AC: holotype, female (5481); D: male (PBK02). A: labels; B, D: habitus, dorsal side; C: habitus, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200044" httpUri="https://zenodo.org/record/10200044/files/figure.png" pageId="9" pageNumber="310">Fig. 4D</figureCitation>
) but in
<typeStatus id="54BC8599FFAFCF6DFB19ED10FAD4F9C8" box="[1225,1323,1547,1573]" pageId="9" pageNumber="310" type="holotype">holotype</typeStatus>
widest at midlength and just ahead of posterior lateral apices (
<figureCitation id="133C27BEFFAFCF6DFD3DED34FCB8F9A7" box="[749,839,1583,1610]" captionStart="FIGURE 4" captionStartId="7.[151,250,857,881]" captionTargetBox="[151,1434,165,833]" captionTargetId="figure-16@7.[151,1436,159,841]" captionTargetPageId="7" captionText="FIGURE 4. Penia babai. AC: holotype, female (5481); D: male (PBK02). A: labels; B, D: habitus, dorsal side; C: habitus, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200044" httpUri="https://zenodo.org/record/10200044/files/figure.png" pageId="9" pageNumber="310">Fig. 4B</figureCitation>
), tallest around midlength (
<figureCitation id="133C27BEFFAFCF6DFB50ED34FB2AF9A7" box="[1152,1237,1583,1610]" captionStart="FIGURE 5" captionStartId="8.[151,250,1618,1642]" captionTargetBox="[151,1436,381,1595]" captionTargetId="figure-115@8.[151,1436,381,1595]" captionTargetPageId="8" captionText="FIGURE 5. Penia babai. AC, EM: holotype, female (5481); D: male (PBK01). A: head, anterior view; B: head, dorsal view; C, D: basal antennomeres of left antenna; E: right maxillary palp, ventral side; F: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); G: left hypomeron (arrow: hypomeron with distinct mesial projection); H: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 5F); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 5F); K: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200047" httpUri="https://zenodo.org/record/10200047/files/figure.png" pageId="9" pageNumber="310">Fig. 5F</figureCitation>
), without median longitudinal depression posteriorly; anterior edge strongly concave; anterior angles simple, nearly right angle; punctate lateral ridge extending from anterior angles to hind angles (
<figureCitation id="133C27BEFFAFCF6DFC77ED6CFBFEF97F" box="[935,1025,1655,1682]" captionStart="FIGURE 4" captionStartId="7.[151,250,857,881]" captionTargetBox="[151,1434,165,833]" captionTargetId="figure-16@7.[151,1436,159,841]" captionTargetPageId="7" captionText="FIGURE 4. Penia babai. AC: holotype, female (5481); D: male (PBK02). A: labels; B, D: habitus, dorsal side; C: habitus, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200044" httpUri="https://zenodo.org/record/10200044/files/figure.png" pageId="9" pageNumber="310">Fig. 4B</figureCitation>
); hind angles simple, broad, weakly protruding posterolaterad; posterior edge with a sublateral incision near each hind angle, without carinae next to sublateral incisions (
<figureCitation id="133C27BEFFAFCF6DFE51EDA4FE24F937" box="[385,475,1727,1754]" captionStart="FIGURE 4" captionStartId="7.[151,250,857,881]" captionTargetBox="[151,1434,165,833]" captionTargetId="figure-16@7.[151,1436,159,841]" captionTargetPageId="7" captionText="FIGURE 4. Penia babai. AC: holotype, female (5481); D: male (PBK02). A: labels; B, D: habitus, dorsal side; C: habitus, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200044" httpUri="https://zenodo.org/record/10200044/files/figure.png" pageId="9" pageNumber="310">Fig. 4B</figureCitation>
). Hypomeron with distinct mesial projection (
<figureCitation id="133C27BEFFAFCF6DFC39EDA4FBBAF937" box="[1001,1093,1727,1754]" captionStart="FIGURE 5" captionStartId="8.[151,250,1618,1642]" captionTargetBox="[151,1436,381,1595]" captionTargetId="figure-115@8.[151,1436,381,1595]" captionTargetPageId="8" captionText="FIGURE 5. Penia babai. AC, EM: holotype, female (5481); D: male (PBK01). A: head, anterior view; B: head, dorsal view; C, D: basal antennomeres of left antenna; E: right maxillary palp, ventral side; F: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); G: left hypomeron (arrow: hypomeron with distinct mesial projection); H: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 5F); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 5F); K: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200047" httpUri="https://zenodo.org/record/10200047/files/figure.png" pageId="9" pageNumber="310">Fig. 5G</figureCitation>
: arrow); anterior angle almost right angle; mesial edge almost straight; mesial and posterior margins with impunctate ridge (
<figureCitation id="133C27BEFFAFCF6DFB16EDF8FADCF913" box="[1222,1315,1763,1790]" captionStart="FIGURE 5" captionStartId="8.[151,250,1618,1642]" captionTargetBox="[151,1436,381,1595]" captionTargetId="figure-115@8.[151,1436,381,1595]" captionTargetPageId="8" captionText="FIGURE 5. Penia babai. AC, EM: holotype, female (5481); D: male (PBK01). A: head, anterior view; B: head, dorsal view; C, D: basal antennomeres of left antenna; E: right maxillary palp, ventral side; F: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); G: left hypomeron (arrow: hypomeron with distinct mesial projection); H: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 5F); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 5F); K: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200047" httpUri="https://zenodo.org/record/10200047/files/figure.png" pageId="9" pageNumber="310">Fig. 5G</figureCitation>
); posterior margin with rectangular projection between two large emarginations; hind angle broadly triangular. Prosternum nearly straight ventrally in lateral view; anterior lobe not protruding beyond prosternal ventral line in lateral view (
<figureCitation id="133C27BEFFAFCF6DFF4FEC54FF09F887" box="[159,246,1871,1898]" captionStart="FIGURE 5" captionStartId="8.[151,250,1618,1642]" captionTargetBox="[151,1436,381,1595]" captionTargetId="figure-115@8.[151,1436,381,1595]" captionTargetPageId="8" captionText="FIGURE 5. Penia babai. AC, EM: holotype, female (5481); D: male (PBK01). A: head, anterior view; B: head, dorsal view; C, D: basal antennomeres of left antenna; E: right maxillary palp, ventral side; F: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); G: left hypomeron (arrow: hypomeron with distinct mesial projection); H: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 5F); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 5F); K: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200047" httpUri="https://zenodo.org/record/10200047/files/figure.png" pageId="9" pageNumber="310">Fig. 5F</figureCitation>
); anterior edge broadly rounded but nearly straight apically in ventral view (
<figureCitation id="133C27BEFFAFCF6DFB8FEC54FB45F887" box="[1119,1210,1871,1898]" captionStart="FIGURE 4" captionStartId="7.[151,250,857,881]" captionTargetBox="[151,1434,165,833]" captionTargetId="figure-16@7.[151,1436,159,841]" captionTargetPageId="7" captionText="FIGURE 4. Penia babai. AC: holotype, female (5481); D: male (PBK02). A: labels; B, D: habitus, dorsal side; C: habitus, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200044" httpUri="https://zenodo.org/record/10200044/files/figure.png" pageId="9" pageNumber="310">Fig. 4C</figureCitation>
). Prosternal process broad, 1.82.1 x longer than procoxal cavity length, concave between procoxae, strongly curved dorsad from the middle of procoxal cavities in lateral view (
<figureCitation id="133C27BEFFAFCF6DFD54EC8CFD27F85F" box="[644,728,1943,1970]" captionStart="FIGURE 5" captionStartId="8.[151,250,1618,1642]" captionTargetBox="[151,1436,381,1595]" captionTargetId="figure-115@8.[151,1436,381,1595]" captionTargetPageId="8" captionText="FIGURE 5. Penia babai. AC, EM: holotype, female (5481); D: male (PBK01). A: head, anterior view; B: head, dorsal view; C, D: basal antennomeres of left antenna; E: right maxillary palp, ventral side; F: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); G: left hypomeron (arrow: hypomeron with distinct mesial projection); H: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 5F); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 5F); K: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200047" httpUri="https://zenodo.org/record/10200047/files/figure.png" pageId="9" pageNumber="310">Fig. 5F</figureCitation>
), without subapical tooth; dorsal lobe roundly expanded ahead of apex in ventral view (
<figureCitation id="133C27BEFFAFCF6DFE77ECA0FDFEF83B" box="[423,513,1979,2006]" captionStart="FIGURE 5" captionStartId="8.[151,250,1618,1642]" captionTargetBox="[151,1436,381,1595]" captionTargetId="figure-115@8.[151,1436,381,1595]" captionTargetPageId="8" captionText="FIGURE 5. Penia babai. AC, EM: holotype, female (5481); D: male (PBK01). A: head, anterior view; B: head, dorsal view; C, D: basal antennomeres of left antenna; E: right maxillary palp, ventral side; F: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); G: left hypomeron (arrow: hypomeron with distinct mesial projection); H: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 5F); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 5F); K: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200047" httpUri="https://zenodo.org/record/10200047/files/figure.png" pageId="9" pageNumber="310">Fig. 5H</figureCitation>
); ventral lobe roundly expanded near base and then abruptly narrowed posteriad in ventral view (
<figureCitation id="133C27BEFFAFCF6DFEE3ECC4FE71F817" box="[307,398,2015,2042]" captionStart="FIGURE 5" captionStartId="8.[151,250,1618,1642]" captionTargetBox="[151,1436,381,1595]" captionTargetId="figure-115@8.[151,1436,381,1595]" captionTargetPageId="8" captionText="FIGURE 5. Penia babai. AC, EM: holotype, female (5481); D: male (PBK01). A: head, anterior view; B: head, dorsal view; C, D: basal antennomeres of left antenna; E: right maxillary palp, ventral side; F: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); G: left hypomeron (arrow: hypomeron with distinct mesial projection); H: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 5F); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 5F); K: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200047" httpUri="https://zenodo.org/record/10200047/files/figure.png" pageId="9" pageNumber="310">Fig. 5H</figureCitation>
); ventral margin strongly and triangularly expanded medially in lateral view (
<figureCitation id="133C27BEFFAFCF6DFB2CECC4FAADF817" box="[1276,1362,2015,2042]" captionStart="FIGURE 5" captionStartId="8.[151,250,1618,1642]" captionTargetBox="[151,1436,381,1595]" captionTargetId="figure-115@8.[151,1436,381,1595]" captionTargetPageId="8" captionText="FIGURE 5. Penia babai. AC, EM: holotype, female (5481); D: male (PBK01). A: head, anterior view; B: head, dorsal view; C, D: basal antennomeres of left antenna; E: right maxillary palp, ventral side; F: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); G: left hypomeron (arrow: hypomeron with distinct mesial projection); H: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 5F); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 5F); K: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200047" httpUri="https://zenodo.org/record/10200047/files/figure.png" pageId="9" pageNumber="310">Fig. 5F</figureCitation>
); apex rounded in lateral and ventral views (
<figureCitation id="133C27BEFFACCF6EFD90EB8CFD47FF5C" box="[576,696,150,177]" captionStart="FIGURE 5" captionStartId="8.[151,250,1618,1642]" captionTargetBox="[151,1436,381,1595]" captionTargetId="figure-115@8.[151,1436,381,1595]" captionTargetPageId="8" captionText="FIGURE 5. Penia babai. AC, EM: holotype, female (5481); D: male (PBK01). A: head, anterior view; B: head, dorsal view; C, D: basal antennomeres of left antenna; E: right maxillary palp, ventral side; F: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); G: left hypomeron (arrow: hypomeron with distinct mesial projection); H: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 5F); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 5F); K: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200047" httpUri="https://zenodo.org/record/10200047/files/figure.png" pageId="10" pageNumber="311">Fig. 5F, H</figureCitation>
). Pronotosternal sutures not grooved (
<figureCitation id="133C27BEFFACCF6EFBB9EB8CFB3CFF5C" box="[1129,1219,150,177]" captionStart="FIGURE 5" captionStartId="8.[151,250,1618,1642]" captionTargetBox="[151,1436,381,1595]" captionTargetId="figure-115@8.[151,1436,381,1595]" captionTargetPageId="8" captionText="FIGURE 5. Penia babai. AC, EM: holotype, female (5481); D: male (PBK01). A: head, anterior view; B: head, dorsal view; C, D: basal antennomeres of left antenna; E: right maxillary palp, ventral side; F: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); G: left hypomeron (arrow: hypomeron with distinct mesial projection); H: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 5F); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 5F); K: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200047" httpUri="https://zenodo.org/record/10200047/files/figure.png" pageId="10" pageNumber="311">Fig. 5G</figureCitation>
), sinuate in ventral view (
<figureCitation id="133C27BEFFACCF6EFF0DEBA0FECAFF38" box="[221,309,187,213]" captionStart="FIGURE 4" captionStartId="7.[151,250,857,881]" captionTargetBox="[151,1434,165,833]" captionTargetId="figure-16@7.[151,1436,159,841]" captionTargetPageId="7" captionText="FIGURE 4. Penia babai. AC: holotype, female (5481); D: male (PBK02). A: labels; B, D: habitus, dorsal side; C: habitus, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200044" httpUri="https://zenodo.org/record/10200044/files/figure.png" pageId="10" pageNumber="311">Fig. 4C</figureCitation>
), moderately opened anteriorly. Scutellar shield tongue-shaped (
<figureCitation id="133C27BEFFACCF6EFBD5EBA0FBACFF38" box="[1029,1107,186,213]" captionStart="FIGURE 5" captionStartId="8.[151,250,1618,1642]" captionTargetBox="[151,1436,381,1595]" captionTargetId="figure-115@8.[151,1436,381,1595]" captionTargetPageId="8" captionText="FIGURE 5. Penia babai. AC, EM: holotype, female (5481); D: male (PBK01). A: head, anterior view; B: head, dorsal view; C, D: basal antennomeres of left antenna; E: right maxillary palp, ventral side; F: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); G: left hypomeron (arrow: hypomeron with distinct mesial projection); H: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 5F); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 5F); K: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200047" httpUri="https://zenodo.org/record/10200047/files/figure.png" pageId="10" pageNumber="311">Fig. 5I</figureCitation>
), 0.91.0 x longer than wide, widest anteriorly, weakly narrowed behind anterior ridge, almost parallel-sided in anterior half and then narrowed posterad, flat, inclined anterior-downwards, visible in lateral view (
<figureCitation id="133C27BEFFACCF6EFC76EA18FC00FEF0" box="[934,1023,258,285]" captionStart="FIGURE 5" captionStartId="8.[151,250,1618,1642]" captionTargetBox="[151,1436,381,1595]" captionTargetId="figure-115@8.[151,1436,381,1595]" captionTargetPageId="8" captionText="FIGURE 5. Penia babai. AC, EM: holotype, female (5481); D: male (PBK01). A: head, anterior view; B: head, dorsal view; C, D: basal antennomeres of left antenna; E: right maxillary palp, ventral side; F: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); G: left hypomeron (arrow: hypomeron with distinct mesial projection); H: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 5F); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 5F); K: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200047" httpUri="https://zenodo.org/record/10200047/files/figure.png" pageId="10" pageNumber="311">Fig. 5F</figureCitation>
); anterior edge broadly rounded, in some slightly protruding medially; posterior edge rounded. Mesosternum: borders of mesosternal cavity straight anteriorly and then curved ventrad in right angle in lateral view (
<figureCitation id="133C27BEFFACCF6EFCBAEA50FC3FFE88" box="[874,960,330,357]" captionStart="FIGURE 5" captionStartId="8.[151,250,1618,1642]" captionTargetBox="[151,1436,381,1595]" captionTargetId="figure-115@8.[151,1436,381,1595]" captionTargetPageId="8" captionText="FIGURE 5. Penia babai. AC, EM: holotype, female (5481); D: male (PBK01). A: head, anterior view; B: head, dorsal view; C, D: basal antennomeres of left antenna; E: right maxillary palp, ventral side; F: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); G: left hypomeron (arrow: hypomeron with distinct mesial projection); H: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 5F); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 5F); K: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200047" httpUri="https://zenodo.org/record/10200047/files/figure.png" pageId="10" pageNumber="311">Fig. 5F</figureCitation>
); mesosternal process between mesocoxae distinctly higher than mesocoxae, visible in lateral view (
<figureCitation id="133C27BEFFACCF6EFCF7EA74FC80FE64" box="[807,895,366,393]" captionStart="FIGURE 5" captionStartId="8.[151,250,1618,1642]" captionTargetBox="[151,1436,381,1595]" captionTargetId="figure-115@8.[151,1436,381,1595]" captionTargetPageId="8" captionText="FIGURE 5. Penia babai. AC, EM: holotype, female (5481); D: male (PBK01). A: head, anterior view; B: head, dorsal view; C, D: basal antennomeres of left antenna; E: right maxillary palp, ventral side; F: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); G: left hypomeron (arrow: hypomeron with distinct mesial projection); H: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 5F); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 5F); K: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200047" httpUri="https://zenodo.org/record/10200047/files/figure.png" pageId="10" pageNumber="311">Fig. 5F</figureCitation>
: white arrow); posterior edge 0.20.25 x wider than total width of mesosternum, weakly emarginate (
<figureCitation id="133C27BEFFACCF6EFCD9EA88FCA2FE40" box="[777,861,402,429]" captionStart="FIGURE 5" captionStartId="8.[151,250,1618,1642]" captionTargetBox="[151,1436,381,1595]" captionTargetId="figure-115@8.[151,1436,381,1595]" captionTargetPageId="8" captionText="FIGURE 5. Penia babai. AC, EM: holotype, female (5481); D: male (PBK01). A: head, anterior view; B: head, dorsal view; C, D: basal antennomeres of left antenna; E: right maxillary palp, ventral side; F: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); G: left hypomeron (arrow: hypomeron with distinct mesial projection); H: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 5F); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 5F); K: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200047" httpUri="https://zenodo.org/record/10200047/files/figure.png" pageId="10" pageNumber="311">Fig. 5J</figureCitation>
). Mesepisternum reaching mesocoxal cavity (
<figureCitation id="133C27BEFFACCF6EFABFEA88FF4CFE3C" captionStart="FIGURE 5" captionStartId="8.[151,250,1618,1642]" captionTargetBox="[151,1436,381,1595]" captionTargetId="figure-115@8.[151,1436,381,1595]" captionTargetPageId="8" captionText="FIGURE 5. Penia babai. AC, EM: holotype, female (5481); D: male (PBK01). A: head, anterior view; B: head, dorsal view; C, D: basal antennomeres of left antenna; E: right maxillary palp, ventral side; F: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); G: left hypomeron (arrow: hypomeron with distinct mesial projection); H: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 5F); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 5F); K: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200047" httpUri="https://zenodo.org/record/10200047/files/figure.png" pageId="10" pageNumber="311">Fig. 5J</figureCitation>
). Metasternum sulcate medially and ahead of metacoxal cavities (
<figureCitation id="133C27BEFFACCF6EFC7BEAACFBF9FE3C" box="[939,1030,439,465]" captionStart="FIGURE 4" captionStartId="7.[151,250,857,881]" captionTargetBox="[151,1434,165,833]" captionTargetId="figure-16@7.[151,1436,159,841]" captionTargetPageId="7" captionText="FIGURE 4. Penia babai. AC: holotype, female (5481); D: male (PBK02). A: labels; B, D: habitus, dorsal side; C: habitus, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200044" httpUri="https://zenodo.org/record/10200044/files/figure.png" pageId="10" pageNumber="311">Fig. 4C</figureCitation>
). Metacoxal plate narrowed toward outer side, becoming like a parallel-sided bar at its outer 2/
<quantity id="4CFF96DEFFACCF6EFC90EAC1FC93FE18" box="[832,876,474,501]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="10" pageNumber="311" unit="in" value="5.0">5 in</quantity>
ventral view (
<figureCitation id="133C27BEFFACCF6EFBC4EAC0FB90FE18" box="[1044,1135,475,501]" captionStart="FIGURE 4" captionStartId="7.[151,250,857,881]" captionTargetBox="[151,1434,165,833]" captionTargetId="figure-16@7.[151,1436,159,841]" captionTargetPageId="7" captionText="FIGURE 4. Penia babai. AC: holotype, female (5481); D: male (PBK02). A: labels; B, D: habitus, dorsal side; C: habitus, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200044" httpUri="https://zenodo.org/record/10200044/files/figure.png" pageId="10" pageNumber="311">Fig. 4C</figureCitation>
). Elytron broadly strongly convex, but with outer margin widely depressed, widest ahead of midlength, 3.13.6 x longer than wide, 2.52.7 x longer than pronotum length; apex rounded; elytral striae defined by lines of elongated punctures. Hind wings fully developed. Tibiae with paired spurs; relative tarsomere lengths: IV&lt;III II&lt;V&lt;I; tarsomeres III and IV with lobe ventrally (
<figureCitation id="133C27BEFFACCF6EFEDAE970FE9BFD68" box="[266,356,618,645]" captionStart="FIGURE 5" captionStartId="8.[151,250,1618,1642]" captionTargetBox="[151,1436,381,1595]" captionTargetId="figure-115@8.[151,1436,381,1595]" captionTargetPageId="8" captionText="FIGURE 5. Penia babai. AC, EM: holotype, female (5481); D: male (PBK01). A: head, anterior view; B: head, dorsal view; C, D: basal antennomeres of left antenna; E: right maxillary palp, ventral side; F: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); G: left hypomeron (arrow: hypomeron with distinct mesial projection); H: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 5F); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 5F); K: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200047" httpUri="https://zenodo.org/record/10200047/files/figure.png" pageId="10" pageNumber="311">Fig. 5K</figureCitation>
).
</paragraph>
<caption id="DF786BB3FFACCF6EFF47EC27FB7FF871" ID-DOI="http://doi.org/10.5281/zenodo.10200051" ID-Zenodo-Dep="10200051" httpUri="https://zenodo.org/record/10200051/files/figure.png" pageId="10" pageNumber="311" startId="10.[151,250,1852,1876]" targetBox="[151,1436,669,1829]" targetPageId="10" targetType="figure">
<paragraph id="8BB83B3BFFACCF6EFF47EC27FB7FF871" blockId="10.[151,1437,1852,1949]" pageId="10" pageNumber="311">
<emphasis id="B973E729FFACCF6EFF47EC27FEECF8B8" bold="true" box="[151,275,1852,1877]" pageId="10" pageNumber="311">FIGURE 7.</emphasis>
<taxonomicName id="4C0740B8FFACCF6EFEC9EC26FE6CF8B9" authorityName="Kishii" authorityYear="1994" box="[281,403,1852,1876]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="babai">
<emphasis id="B973E729FFACCF6EFEC9EC26FE6CF8B9" box="[281,403,1852,1876]" italics="true" pageId="10" pageNumber="311">Penia babai</emphasis>
</taxonomicName>
, females. A, B, E, F: non-type (PBK03); C, D: holotype (5481); G: non-type (PBK04). A: tergite VIII; B: sternite VIII; C: slide; D, E: pregenital segments and genitalia; F: coxites, ventral side; G: vagina and bursa copulatrix (arrow 1: bursa copulatrix posteriorly with a short sac, arrow 2: bursa copulatrix with a sac at the middle).
</paragraph>
</caption>
<paragraph id="8BB83B3BFFADCF6FFF17EB8CFA62FE3C" blockId="11.[151,1437,150,789]" pageId="11" pageNumber="312">
Abdomen. Ventrite V semicircular, rounded apically (
<figureCitation id="133C27BEFFADCF6FFCCAEB8CFC8EFF5C" box="[794,881,151,177]" captionStart="FIGURE 4" captionStartId="7.[151,250,857,881]" captionTargetBox="[151,1434,165,833]" captionTargetId="figure-16@7.[151,1436,159,841]" captionTargetPageId="7" captionText="FIGURE 4. Penia babai. AC: holotype, female (5481); D: male (PBK02). A: labels; B, D: habitus, dorsal side; C: habitus, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200044" httpUri="https://zenodo.org/record/10200044/files/figure.png" pageId="11" pageNumber="312">Fig. 4C</figureCitation>
), 0.450.6 x longer than wide.
<emphasis id="B973E729FFADCF6FFB1BEB8CFAF0FF5C" bold="true" box="[1227,1295,151,177]" pageId="11" pageNumber="312">Male.</emphasis>
Tergites and sternites VIIIX yellow. Tergite VIII 0.8 x longer than wide, trapezoidal, narrowed posterad; posterior margin nearly straight (
<figureCitation id="133C27BEFFADCF6FFF2AEBC4FEABFF14" box="[250,340,223,249]" captionStart="FIGURE 6" captionStartId="9.[151,250,1405,1429]" captionTargetBox="[151,1431,381,1377]" captionTargetId="figure-122@9.[151,1436,381,1382]" captionTargetPageId="9" captionText="FIGURE 6. Penia babai, male (PBK01). A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal side; F: aedeagus, ventral side; G: apical part of left paramera, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200049" httpUri="https://zenodo.org/record/10200049/files/figure.png" pageId="11" pageNumber="312">Fig. 6A</figureCitation>
). Sternite VIII posteriorly widely concave between two projections (
<figureCitation id="133C27BEFFADCF6FFB80EBC4FB57FF14" box="[1104,1192,223,249]" captionStart="FIGURE 6" captionStartId="9.[151,250,1405,1429]" captionTargetBox="[151,1431,381,1377]" captionTargetId="figure-122@9.[151,1436,381,1382]" captionTargetPageId="9" captionText="FIGURE 6. Penia babai, male (PBK01). A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal side; F: aedeagus, ventral side; G: apical part of left paramera, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200049" httpUri="https://zenodo.org/record/10200049/files/figure.png" pageId="11" pageNumber="312">Fig. 6B</figureCitation>
). Tergite IX almost as long as wide; median notch 0.7 x total length of tergite IX (
<figureCitation id="133C27BEFFADCF6FFCE6EA18FC70FEF0" box="[822,911,259,285]" captionStart="FIGURE 6" captionStartId="9.[151,250,1405,1429]" captionTargetBox="[151,1431,381,1377]" captionTargetId="figure-122@9.[151,1436,381,1382]" captionTargetPageId="9" captionText="FIGURE 6. Penia babai, male (PBK01). A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal side; F: aedeagus, ventral side; G: apical part of left paramera, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200049" httpUri="https://zenodo.org/record/10200049/files/figure.png" pageId="11" pageNumber="312">Fig. 6C</figureCitation>
). Tergite X longer than wide, rounded apically (
<figureCitation id="133C27BEFFADCF6FFF4FEA3CFF07FEAC" box="[159,248,295,321]" captionStart="FIGURE 6" captionStartId="9.[151,250,1405,1429]" captionTargetBox="[151,1431,381,1377]" captionTargetId="figure-122@9.[151,1436,381,1382]" captionTargetPageId="9" captionText="FIGURE 6. Penia babai, male (PBK01). A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal side; F: aedeagus, ventral side; G: apical part of left paramera, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200049" httpUri="https://zenodo.org/record/10200049/files/figure.png" pageId="11" pageNumber="312">Fig. 6C</figureCitation>
). Sternite IX 2.3 x longer than wide, constricted ahead of midlength (
<figureCitation id="133C27BEFFADCF6FFC2BEA3CFBAAFEAC" box="[1019,1109,295,321]" captionStart="FIGURE 6" captionStartId="9.[151,250,1405,1429]" captionTargetBox="[151,1431,381,1377]" captionTargetId="figure-122@9.[151,1436,381,1382]" captionTargetPageId="9" captionText="FIGURE 6. Penia babai, male (PBK01). A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal side; F: aedeagus, ventral side; G: apical part of left paramera, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200049" httpUri="https://zenodo.org/record/10200049/files/figure.png" pageId="11" pageNumber="312">Fig. 6D</figureCitation>
), rounded apically. Aedeagus yellow (
<figureCitation id="133C27BEFFADCF6FFF22EA50FE9BFE88" box="[242,356,331,357]" captionStart="FIGURE 6" captionStartId="9.[151,250,1405,1429]" captionTargetBox="[151,1431,381,1377]" captionTargetId="figure-122@9.[151,1436,381,1382]" captionTargetPageId="9" captionText="FIGURE 6. Penia babai, male (PBK01). A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal side; F: aedeagus, ventral side; G: apical part of left paramera, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200049" httpUri="https://zenodo.org/record/10200049/files/figure.png" pageId="11" pageNumber="312">Fig. 6E, F</figureCitation>
). Phallobase 0.3 x total length of aedeagus, 0.9 x longer than wide. Median lobe exceeding apices of parameres by apical 1/10; basal struts 0.3 x total length of median lobe. Parameres broad, not fused ventrally (
<figureCitation id="133C27BEFFADCF6FFABFEA74FF45FE40" captionStart="FIGURE 6" captionStartId="9.[151,250,1405,1429]" captionTargetBox="[151,1431,381,1377]" captionTargetId="figure-122@9.[151,1436,381,1382]" captionTargetPageId="9" captionText="FIGURE 6. Penia babai, male (PBK01). A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal side; F: aedeagus, ventral side; G: apical part of left paramera, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200049" httpUri="https://zenodo.org/record/10200049/files/figure.png" pageId="11" pageNumber="312">Fig. 6E</figureCitation>
); preapical expansions protruding laterad (
<figureCitation id="133C27BEFFADCF6FFD42EA88FD15FE40" box="[658,746,403,429]" captionStart="FIGURE 6" captionStartId="9.[151,250,1405,1429]" captionTargetBox="[151,1431,381,1377]" captionTargetId="figure-122@9.[151,1436,381,1382]" captionTargetPageId="9" captionText="FIGURE 6. Penia babai, male (PBK01). A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal side; F: aedeagus, ventral side; G: apical part of left paramera, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200049" httpUri="https://zenodo.org/record/10200049/files/figure.png" pageId="11" pageNumber="312">Fig. 6G</figureCitation>
); apex beyond preapical expansions large triangular (
<figureCitation id="133C27BEFFADCF6FFAE7EA88FA70FE40" box="[1335,1423,403,429]" captionStart="FIGURE 6" captionStartId="9.[151,250,1405,1429]" captionTargetBox="[151,1431,381,1377]" captionTargetId="figure-122@9.[151,1436,381,1382]" captionTargetPageId="9" captionText="FIGURE 6. Penia babai, male (PBK01). A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal side; F: aedeagus, ventral side; G: apical part of left paramera, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200049" httpUri="https://zenodo.org/record/10200049/files/figure.png" pageId="11" pageNumber="312">Fig. 6G</figureCitation>
), with a seta dorsally, with a seta ventrally; apex length 0.40.6 x width of parameres at expansions in ventral side.
</paragraph>
<paragraph id="8BB83B3BFFADCF6FFF47EAC0FD9CFD1C" blockId="11.[151,1437,150,789]" pageId="11" pageNumber="312">
<emphasis id="B973E729FFADCF6FFF47EAC0FF0AFE18" bold="true" box="[151,245,475,501]" pageId="11" pageNumber="312">Female.</emphasis>
Tergite VIII and sternite VIII yellow. Terigite VIII semicircular, 1.1 x longer than wide (
<figureCitation id="133C27BEFFADCF6FFB0AEAC0FACAFE18" box="[1242,1333,475,501]" captionStart="FIGURE 7" captionStartId="10.[151,250,1852,1876]" captionTargetBox="[151,1436,669,1829]" captionTargetId="figure-240@10.[151,1436,669,1829]" captionTargetPageId="10" captionText="FIGURE 7. Penia babai, females.A, B, E, F: non-type (PBK03); C, D: holotype (5481); G: non-type (PBK04). A: tergite VIII; B: sternite VIII; C: slide; D, E: pregenital segments and genitalia; F: coxites, ventral side; G: vagina and bursa copulatrix (arrow 1: bursa copulatrix posteriorly with a short sac, arrow 2: bursa copulatrix with a sac at the middle)." figureDoi="http://doi.org/10.5281/zenodo.10200051" httpUri="https://zenodo.org/record/10200051/files/figure.png" pageId="11" pageNumber="312">Fig. 7A</figureCitation>
); sternite VIII (between base of spiculum ventrale and apex) semicircular, 0.9 x longer than wide (
<figureCitation id="133C27BEFFADCF6FFB48EAE4FB0DFDF4" box="[1176,1266,511,537]" captionStart="FIGURE 7" captionStartId="10.[151,250,1852,1876]" captionTargetBox="[151,1436,669,1829]" captionTargetId="figure-240@10.[151,1436,669,1829]" captionTargetPageId="10" captionText="FIGURE 7. Penia babai, females.A, B, E, F: non-type (PBK03); C, D: holotype (5481); G: non-type (PBK04). A: tergite VIII; B: sternite VIII; C: slide; D, E: pregenital segments and genitalia; F: coxites, ventral side; G: vagina and bursa copulatrix (arrow 1: bursa copulatrix posteriorly with a short sac, arrow 2: bursa copulatrix with a sac at the middle)." figureDoi="http://doi.org/10.5281/zenodo.10200051" httpUri="https://zenodo.org/record/10200051/files/figure.png" pageId="11" pageNumber="312">Fig. 7B</figureCitation>
; the pregenital segments and genitalia of
<typeStatus id="54BC8599FFADCF6FFE68E938FDE5FDD0" box="[440,538,547,573]" pageId="11" pageNumber="312" type="holotype">holotype</typeStatus>
had been mounted in balsam on slides and were distorted by the pressure exerted by the coverslip,
<figureCitation id="133C27BEFFADCF6FFE87E95CFE30FD8C" box="[343,463,583,609]" captionStart="FIGURE 7" captionStartId="10.[151,250,1852,1876]" captionTargetBox="[151,1436,669,1829]" captionTargetId="figure-240@10.[151,1436,669,1829]" captionTargetPageId="10" captionText="FIGURE 7. Penia babai, females.A, B, E, F: non-type (PBK03); C, D: holotype (5481); G: non-type (PBK04). A: tergite VIII; B: sternite VIII; C: slide; D, E: pregenital segments and genitalia; F: coxites, ventral side; G: vagina and bursa copulatrix (arrow 1: bursa copulatrix posteriorly with a short sac, arrow 2: bursa copulatrix with a sac at the middle)." figureDoi="http://doi.org/10.5281/zenodo.10200051" httpUri="https://zenodo.org/record/10200051/files/figure.png" pageId="11" pageNumber="312">Fig. 7C, D</figureCitation>
); spiculum ventrale 5.6 (in
<typeStatus id="54BC8599FFADCF6FFCD1E95CFC9CFD8C" box="[769,867,583,609]" pageId="11" pageNumber="312" type="holotype">holotype</typeStatus>
4.5) x longer than length of sternite VIII (
<figureCitation id="133C27BEFFADCF6FFAE9E95CFA70FD8C" box="[1337,1423,583,609]" captionStart="FIGURE 7" captionStartId="10.[151,250,1852,1876]" captionTargetBox="[151,1436,669,1829]" captionTargetId="figure-240@10.[151,1436,669,1829]" captionTargetPageId="10" captionText="FIGURE 7. Penia babai, females.A, B, E, F: non-type (PBK03); C, D: holotype (5481); G: non-type (PBK04). A: tergite VIII; B: sternite VIII; C: slide; D, E: pregenital segments and genitalia; F: coxites, ventral side; G: vagina and bursa copulatrix (arrow 1: bursa copulatrix posteriorly with a short sac, arrow 2: bursa copulatrix with a sac at the middle)." figureDoi="http://doi.org/10.5281/zenodo.10200051" httpUri="https://zenodo.org/record/10200051/files/figure.png" pageId="11" pageNumber="312">Fig. 7E</figureCitation>
). Ovipositor 1.2 x longer than length of abdomen; coxites two segmented at ventral side (
<figureCitation id="133C27BEFFADCF6FFBBCE970FB3EFD68" box="[1132,1217,619,645]" captionStart="FIGURE 7" captionStartId="10.[151,250,1852,1876]" captionTargetBox="[151,1436,669,1829]" captionTargetId="figure-240@10.[151,1436,669,1829]" captionTargetPageId="10" captionText="FIGURE 7. Penia babai, females.A, B, E, F: non-type (PBK03); C, D: holotype (5481); G: non-type (PBK04). A: tergite VIII; B: sternite VIII; C: slide; D, E: pregenital segments and genitalia; F: coxites, ventral side; G: vagina and bursa copulatrix (arrow 1: bursa copulatrix posteriorly with a short sac, arrow 2: bursa copulatrix with a sac at the middle)." figureDoi="http://doi.org/10.5281/zenodo.10200051" httpUri="https://zenodo.org/record/10200051/files/figure.png" pageId="11" pageNumber="312">Fig. 7F</figureCitation>
), with several setae each dorsally, ventrally, and apically; stylus with several setae apically (
<figureCitation id="133C27BEFFADCF6FFC06E994FBD1FD44" box="[982,1070,655,681]" captionStart="FIGURE 7" captionStartId="10.[151,250,1852,1876]" captionTargetBox="[151,1436,669,1829]" captionTargetId="figure-240@10.[151,1436,669,1829]" captionTargetPageId="10" captionText="FIGURE 7. Penia babai, females.A, B, E, F: non-type (PBK03); C, D: holotype (5481); G: non-type (PBK04). A: tergite VIII; B: sternite VIII; C: slide; D, E: pregenital segments and genitalia; F: coxites, ventral side; G: vagina and bursa copulatrix (arrow 1: bursa copulatrix posteriorly with a short sac, arrow 2: bursa copulatrix with a sac at the middle)." figureDoi="http://doi.org/10.5281/zenodo.10200051" httpUri="https://zenodo.org/record/10200051/files/figure.png" pageId="11" pageNumber="312">Fig. 7F</figureCitation>
). Vagina short; bursa copulatrix elongated spheroid, without sclerotized structures (
<figureCitation id="133C27BEFFADCF6FFD04E9A8FCD1FD20" box="[724,814,691,717]" captionStart="FIGURE 7" captionStartId="10.[151,250,1852,1876]" captionTargetBox="[151,1436,669,1829]" captionTargetId="figure-240@10.[151,1436,669,1829]" captionTargetPageId="10" captionText="FIGURE 7. Penia babai, females.A, B, E, F: non-type (PBK03); C, D: holotype (5481); G: non-type (PBK04). A: tergite VIII; B: sternite VIII; C: slide; D, E: pregenital segments and genitalia; F: coxites, ventral side; G: vagina and bursa copulatrix (arrow 1: bursa copulatrix posteriorly with a short sac, arrow 2: bursa copulatrix with a sac at the middle)." figureDoi="http://doi.org/10.5281/zenodo.10200051" httpUri="https://zenodo.org/record/10200051/files/figure.png" pageId="11" pageNumber="312">Fig. 7G</figureCitation>
), with a short sac posteriorly (
<figureCitation id="133C27BEFFADCF6FFB53E9A8FB22FD20" box="[1155,1245,691,717]" captionStart="FIGURE 7" captionStartId="10.[151,250,1852,1876]" captionTargetBox="[151,1436,669,1829]" captionTargetId="figure-240@10.[151,1436,669,1829]" captionTargetPageId="10" captionText="FIGURE 7. Penia babai, females.A, B, E, F: non-type (PBK03); C, D: holotype (5481); G: non-type (PBK04). A: tergite VIII; B: sternite VIII; C: slide; D, E: pregenital segments and genitalia; F: coxites, ventral side; G: vagina and bursa copulatrix (arrow 1: bursa copulatrix posteriorly with a short sac, arrow 2: bursa copulatrix with a sac at the middle)." figureDoi="http://doi.org/10.5281/zenodo.10200051" httpUri="https://zenodo.org/record/10200051/files/figure.png" pageId="11" pageNumber="312">Fig. 7G</figureCitation>
: arrow 1), with a sac around midlength (
<figureCitation id="133C27BEFFADCF6FFE48E9CCFE0DFD1C" box="[408,498,727,753]" captionStart="FIGURE 7" captionStartId="10.[151,250,1852,1876]" captionTargetBox="[151,1436,669,1829]" captionTargetId="figure-240@10.[151,1436,669,1829]" captionTargetPageId="10" captionText="FIGURE 7. Penia babai, females.A, B, E, F: non-type (PBK03); C, D: holotype (5481); G: non-type (PBK04). A: tergite VIII; B: sternite VIII; C: slide; D, E: pregenital segments and genitalia; F: coxites, ventral side; G: vagina and bursa copulatrix (arrow 1: bursa copulatrix posteriorly with a short sac, arrow 2: bursa copulatrix with a sac at the middle)." figureDoi="http://doi.org/10.5281/zenodo.10200051" httpUri="https://zenodo.org/record/10200051/files/figure.png" pageId="11" pageNumber="312">Fig. 7G</figureCitation>
: arrow 2).
</paragraph>
</subSubSection>
<subSubSection id="C31D68B0FFADCF6FFF17E9E1FC43FCF8" box="[199,956,762,789]" pageId="11" pageNumber="312" type="distribution">
<paragraph id="8BB83B3BFFADCF6FFF17E9E1FC43FCF8" blockId="11.[151,1437,150,789]" box="[199,956,762,789]" pageId="11" pageNumber="312">
<emphasis id="B973E729FFADCF6FFF17E9E1FE9EFCF9" bold="true" box="[199,353,762,788]" pageId="11" pageNumber="312">Distribution.</emphasis>
<collectingCountry id="F3107BABFFADCF6FFEB7E9E0FE42FCF8" box="[359,445,763,789]" name="Taiwan" pageId="11" pageNumber="312">Taiwan</collectingCountry>
:
<collectingRegion id="49C3F5D9FFADCF6FFE18E9E0FD8CFCF8" box="[456,627,763,789]" country="Taiwan" name="Nantou" pageId="11" pageNumber="312">Nantou County</collectingRegion>
and
<collectingRegion id="49C3F5D9FFADCF6FFD7AE9E0FCA5FCF8" box="[682,858,763,789]" country="Taiwan" name="Kaohsiung" pageId="11" pageNumber="312">Kaohsiung City</collectingRegion>
(
<figureCitation id="133C27BEFFADCF6FFCBAE9E0FC54FCF8" box="[874,939,763,789]" captionStart="FIGURE 1" captionStartId="3.[151,250,902,926]" captionTargetBox="[158,1429,188,872]" captionTargetId="figure-16@3.[151,1436,181,879]" captionTargetPageId="3" captionText="FIGURE 1. Map of Taiwan. A: location of Taiwan; B: the collection sites of the material used in this study." figureDoi="http://doi.org/10.5281/zenodo.10200038" httpUri="https://zenodo.org/record/10200038/files/figure.png" pageId="11" pageNumber="312">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>