990 lines
114 KiB
XML
990 lines
114 KiB
XML
<document id="BD62C76FDD04FF92FCCF5BBBE48FA3AE" ID="10.11646/zootaxa.4204.1.1" ID-DOI="10.11646/zootaxa.4204.1.1" ID-GBIF-Dataset="3f8866d2-529e-43ad-b971-29fc52a13858" ID-ISSN="1175-5326" ID-Zenodo-Dep="192302" ID-ZooBank="C050058A-774D-49C0-93F9-7A055B51C2A0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1480667017343" checkinUser="plazi" docAuthor="Williams, Paul H., Huang, Jiaxing, Rasmont, Pierre & An, Jiandong" docDate="2016" docId="03AE67547C76332CB090A079A1E0FBFC" docLanguage="en" docName="zootaxa.4204.1.1.pdf" docOrigin="Zootaxa 4204 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Bombus waltoni Cockerell" docType="treatment" docVersion="9" lastPageNumber="33" masterDocId="FF971F2C7C6D330CB007A424A336FFA4" masterDocTitle="Early-diverging bumblebees from across the roof of the world: the high-mountain subgenus Mendacibombus revised from species’ gene coalescents and morphology (Hymenoptera, Apidae)" masterLastPageNumber="72" masterPageNumber="1" pageNumber="28" updateTime="1698395689147" updateUser="plazi">
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<mods:mods id="861EC803268F787A29E96ABDE80C6B4D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B07920519A20D99A0A1918CC5E9B3209">
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<mods:title id="596A53A87B7F4D527446CAA4A2B2C181">Early-diverging bumblebees from across the roof of the world: the high-mountain subgenus Mendacibombus revised from species’ gene coalescents and morphology (Hymenoptera, Apidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="AB46D76498519219F6A56D68A141FA9C" type="personal">
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<mods:roleTerm id="6F78DA1C86C08963790FAACE1C8DAF7C">Author</mods:roleTerm>
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<mods:namePart id="AFC12FCF3E62C849DD877BF63BDC4BEF">Williams, Paul H.</mods:namePart>
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<mods:name id="723F0807ABD3C00EC00055070D4645E0" type="personal">
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<mods:roleTerm id="DC344D35C9002606C71DB44701690662">Author</mods:roleTerm>
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<mods:namePart id="40BF6CF12632D8887593C10338737F50">Huang, Jiaxing</mods:namePart>
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</mods:name>
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<mods:name id="ACD6BB14E6CB4B651B18F7D7FFF7B590" type="personal">
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<mods:roleTerm id="0B2DCAD266054074081816B1954A39DA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5EA9C1513CB56211689DA49FB664380F">Rasmont, Pierre</mods:namePart>
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</mods:name>
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<mods:name id="306E32370B2DB5D16F7066F3C8CADFE8" type="personal">
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<mods:roleTerm id="99C7F410F918ADB6C1B3250B41299E56">Author</mods:roleTerm>
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<mods:namePart id="C4F2081EDFC118D962E773EF568C44CE">An, Jiandong</mods:namePart>
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<mods:typeOfResource id="6E2B9519CFB84F63DA1363A61C06F9CA">text</mods:typeOfResource>
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<mods:title id="547182243903FDC89697183F27247184">Zootaxa</mods:title>
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<mods:part id="BC6CD659C6FC91C554B0617A2072641D">
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<mods:date id="F75D15E9902B5235EE644771B3459567">2016</mods:date>
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<mods:detail id="86C1D9148D759AE90F86D39A611D6112" type="volume">
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<mods:number id="749D577B62C684BDB694C1D10B28CBA0">4204</mods:number>
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</mods:detail>
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<mods:detail id="E548F9759363448471B9F94F99F99232" type="issue">
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<mods:number id="B14EF392FB420069EB1DFF6F31069D3E">1</mods:number>
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<mods:classification id="7EC7D4957E38C38A88149FA329F6C136">journal article</mods:classification>
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<mods:identifier id="263D6513949309BD6F610B244FA63F47" type="DOI">10.11646/zootaxa.4204.1.1</mods:identifier>
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<mods:identifier id="6B8D925FD7D4F7285421DE82D8CE3843" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="22F88049EC309D39725AC3A08CBDF590" type="Zenodo-Dep">192302</mods:identifier>
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<treatment id="03AE67547C76332CB090A079A1E0FBFC" ID-DOI="http://doi.org/10.5281/zenodo.5625287" ID-GBIF-Taxon="124940392" ID-Zenodo-Dep="5625287" LSID="urn:lsid:plazi:treatment:03AE67547C76332CB090A079A1E0FBFC" httpUri="http://treatment.plazi.org/id/03AE67547C76332CB090A079A1E0FBFC" lastPageId="32" lastPageNumber="33" pageId="27" pageNumber="28">
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<subSubSection id="C31D85C97C763317B090A079A136FBD3" box="[151,512,1117,1143]" pageId="27" pageNumber="28" type="nomenclature">
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<paragraph id="8BB8D6427C763317B090A079A136FBD3" blockId="27.[151,512,1117,1177]" box="[151,512,1117,1143]" pageId="27" pageNumber="28">
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<heading id="D0F0612E7C763317B090A079A136FBD3" bold="true" box="[151,512,1117,1143]" fontSize="11" level="1" pageId="27" pageNumber="28" reason="1">
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||
<emphasis id="B9730A507C763317B090A079A136FBD3" bold="true" box="[151,512,1117,1143]" pageId="27" pageNumber="28">
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||
2.
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||
<taxonomicName id="4C07ADC17C763317B0B2A079A136FBD3" ID-CoL="MFY7" authority="Cockerell" authorityName="Cockerell" box="[181,512,1117,1143]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="waltoni">
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<emphasis id="B9730A507C763317B0B2A079A248FBD3" bold="true" box="[181,382,1117,1143]" italics="true" pageId="27" pageNumber="28">Bombus waltoni</emphasis>
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Cockerell
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</taxonomicName>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31D85C97C763317B090A0A4A045FB7D" pageId="27" pageNumber="28" type="description">
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<paragraph id="8BB8D6427C763317B090A0A4A2ABFB3D" blockId="27.[151,512,1117,1177]" box="[151,413,1152,1177]" pageId="27" pageNumber="28">
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(
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||
<figureCitation id="133CCAC77C763317B098A0A4A3DEFB3D" box="[159,232,1152,1177]" captionStart="FIGURES 1 ‒ 6" captionStartId="5.[151,264,1819,1841]" captionTargetBox="[168,1389,193,1768]" captionTargetId="figure@5.[168,1412,193,1784]" captionTargetPageId="5" captionText="FIGURES 1 ‒ 6. Views of perching males of species of the subgenus Mendacibombus, showing the anteriorly extended antennae and laterally extended wings that are characteristic of their form of male mate-searching behaviour: 1, B. waltoni at Diebu ca 3910 m, Minshan mountains, Gansu, China, 29. viii. 2009; 2, B. convexus at Diebu ca 3340 m, Minshan mountains, Gansu, China, 29. viii. 2009; 3, B. avinoviellus at Gulmarg ca 2700 m, Pir Panjal mountains, Kashmir, India, 16. viii. 1985; 4, B. himalayanus at Apharwat ca 4100 m, Pir Panjal mountains, Kashmir, India, 30. viii. 1986; 5 ‒ 6, B. margreiteri near Dulan ca 3520 m, Burhan Budai mountains, Qinghai, China, 31. viii. 2010. Photos by PW (Figs 3 – 4 are the best available for these species)." httpUri="https://zenodo.org/record/192303/files/figure.png" pageId="27" pageNumber="28">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="133CCAC77C763317B0F1A0A4A223FB3D" box="[246,277,1152,1177]" captionStart="FIGURES 24 ‒ 35" captionStartId="26.[151,264,1867,1889]" captionTargetBox="[233,1387,193,1789]" captionTargetId="figure@26.[233,1390,193,1894]" captionTargetPageId="26" captionText="FIGURES 24 ‒ 35. Morphology of the male genitalia for species of the subgenus Mendacibombus from the dorsal aspect, anterior to the left of the image, posterior to the right: 24, B. superbus, China MD # 339; 25, B. waltoni, China MD # 216; 26, B. convexus, China MD # 215; 27, B. makarjini, Kazakhstan MD # 1240; 28, B. marussinus, Pakistan MD # 408; 29, B. avinoviellus, India MD # 404; 30, B. himalayanus, India MD # 426; 31, B. turkestanicus, China MD # 351; 32, B. defector, Kyrgyzstan MD # 367; 33, B. margreiteri, China MD # 294; 34, B. handlirschianus, Russia MD # 835; 35, B. mendax, Spain MD # 689." httpUri="https://zenodo.org/record/192312/files/figure.png" pageId="27" pageNumber="28">25</figureCitation>
|
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,
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||
<figureCitation id="133CCAC77C763317B126A0A4A25DFB3D" box="[289,363,1152,1177]" captionStart="FIGURES 36 ‒ 55" captionStartId="29.[151,264,1875,1897]" captionTargetBox="[151,1415,299,1829]" captionTargetId="figure@29.[151,1436,193,1853]" captionTargetPageId="29" captionText="FIGURES 36 ‒ 55. Morphology of parts of the male genitalia from the dorsal aspect of the left side, anterior to the left of the image, posterior to the right, white bars indicating morphological characters referred to in the key to males: 36, B. waltoni; 37, B. mendax; 38, B. superbus; 39, B. convexus; 40, B. waltoni; 41, B. convexus; 42, B. defector; 43, B. mendax; 44, B. mendax; 45, B. turkestanicus; 46, B. avinoviellus; 47, B. mendax; 48, B. mendax; 49, B. makarjini; 50, B. handlirschianus; 51, B. himalayanus; 52, B. turkestanicus; 53, B. himalayanus; 54, B. himalayanus; 55, B. marussinus." httpUri="https://zenodo.org/record/192314/files/figure.png" pageId="27" pageNumber="28">36, 40</figureCitation>
|
||
,
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||
<figureCitation id="133CCAC77C763317B170A0A4A2A3FB3D" box="[375,405,1152,1177]" captionStart="FIGURES 56 ‒ 67" captionStartId="35.[151,264,1891,1913]" captionTargetBox="[320,1245,193,1835]" captionTargetId="figure@35.[320,1245,193,1843]" captionTargetPageId="35" captionText="FIGURES 56 ‒ 67. Maps of the global distributions of species of the subgenus Mendacibombus from the material examined, with grey spots for specimen records and black spots for the samples with COI sequences (each species is unknown from regions outside the map boundaries shown); grey crosses for records of all species of Mendacibombus combined. Relief map with hill shading, Cartesian orthonormal projection, grey lines show borders to national administration according to UN maps, north at the top of the map. Image created in ArcGIS using World _ Shaded _ Relief basemap © 2014 Esri." httpUri="https://zenodo.org/record/192315/files/figure.png" pageId="27" pageNumber="28">57</figureCitation>
|
||
)
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||
</paragraph>
|
||
<paragraph id="8BB8D6427C763317B090A0E0A045FB7D" blockId="27.[151,1437,1219,1915]" box="[151,883,1219,1241]" pageId="27" pageNumber="28">
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||
<treatmentCitationGroup id="AB17F16C7C763317B090A0E0A045FB7D" box="[151,883,1219,1241]" pageId="27" pageNumber="28">
|
||
[
|
||
<taxonomicName id="4C07ADC17C763317B098A0E7A2F9FB7D" ID-CoL="MFPJ" authority="Gerstaecker" authorityName="Gerstaecker" box="[159,463,1219,1241]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="mendax">
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||
<emphasis id="B9730A507C763317B098A0E7A27FFB7D" box="[159,329,1219,1241]" italics="true" pageId="27" pageNumber="28">Bombus mendax</emphasis>
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Gerstaecker
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||
</taxonomicName>
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;
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<treatmentCitation id="0AA6F0537C763317B1DDA0E7A19FFB7D" author="Morawitz" box="[474,681,1219,1241]" page="199" pageId="27" pageNumber="28" year="1887">
|
||
<bibRefCitation id="EF96ABB37C763317B1DDA0E7A14AFB7D" author="Morawitz" box="[474,636,1219,1241]" pageId="27" pageNumber="28" refString="Morawitz, F. F. (1887) Insecta in itinere cl. N. Przewalskii in Asia centrali novissime lecta. I. Apidae. Trudy Russkago entomologicheskago obshchestva, 20 (1886), 195 - 229." type="journal article" year="1887">Morawitz 1887</bibRefCitation>
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:199
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||
</treatmentCitation>
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||
, misidentification.]
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D85C97C763317B090A0C6A0A0F8DF" pageId="27" pageNumber="28" type="reference_group">
|
||
<paragraph id="8BB8D6427C763317B090A0C6A2C8FAD6" blockId="27.[151,1437,1219,1915]" pageId="27" pageNumber="28">
|
||
<materialsCitation id="3B6FDC1F7C763317B090A0C6A2C8FAD6" ID-GBIF-Occurrence="1413110420" collectionCode="ZISP" country="China" county="Cockerell" location="Cyrillic" municipality="Skorikov" pageId="27" pageNumber="28" specimenCount="1" stateProvince="Qinghai" typeStatus="lectotype">
|
||
<
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||
<taxonomicName id="4C07ADC17C763317B0A2A0C5A278FB53" ID-CoL="MFPJ" box="[165,334,1249,1271]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="mendax">
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||
<emphasis id="B9730A507C763317B0A2A0C5A278FB53" box="[165,334,1249,1271]" italics="true" pageId="27" pageNumber="28">Bombus mendax</emphasis>
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</taxonomicName>
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||
> subsp.
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<taxonomicName id="4C07ADC17C763317B1ACA0C5A01FFB53" authority="Skorikov 1910b" authorityName="Skorikov" authorityYear="1910" box="[427,809,1249,1271]" class="Insecta" family="Apidae" genus="Mendacibombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="chinensis">
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||
<emphasis id="B9730A507C763317B1ACA0C5A13AFB53" box="[427,524,1249,1271]" italics="true" pageId="27" pageNumber="28">chinensis</emphasis>
|
||
<bibRefCitation id="EF96ABB37C763317B215A0C5A18EFB53" author="Skorikov" box="[530,696,1249,1271]" pageId="27" pageNumber="28" refString="Skorikov, A. S. (1910 b) [Intraspecific forms of Bombus mendax Gerst.] (Hymenoptera, Bombidae). Russkoe entomologicheskoe Obozrenie, 9 (1909), 328 - 330." type="journal article" year="1910" yearSuffix="b">Skorikov 1910b</bibRefCitation>
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||
(February)
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</taxonomicName>
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||
:330 (not of
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<bibRefCitation id="EF96ABB37C763317B3A1A0C5A77EFB53" author="Morawitz" box="[934,1096,1249,1271]" pageId="27" pageNumber="28" refString="Morawitz, F. F. (1890) Insecta a cl. G. N. Potanin in China et in Mongolia novissime lecta. XIV. Hymenoptera Aculeata. II). III. Apidae. Trudy Russkago entomologicheskago obshchestva, 24, 349 - 385." type="journal article" year="1890">Morawitz 1890</bibRefCitation>
|
||
:352, =
|
||
<taxonomicName id="4C07ADC17C763317B493A0C6A6A1FB53" ID-CoL="MFG2" authority="Morawitz" authorityName="Morawitz" box="[1172,1431,1249,1271]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="chinensis">
|
||
<emphasis id="B9730A507C763317B493A0C6A626FB53" box="[1172,1296,1249,1271]" italics="true" pageId="27" pageNumber="28">B. chinensis</emphasis>
|
||
(Morawitz))
|
||
</taxonomicName>
|
||
, type-locality citation (Cyrillic) ‘[Mountains of Sining and Burkhan Budda]’.
|
||
<emphasis id="B9730A507C763317B3F3A124A757FAB2" bold="true" box="[1012,1121,1280,1302]" pageId="27" pageNumber="28">
|
||
<typeStatus id="54BC68E07C763317B3F3A124A757FAB2" box="[1012,1121,1280,1302]" pageId="27" pageNumber="28" type="lectotype">Lectotype</typeStatus>
|
||
</emphasis>
|
||
queen by present designation
|
||
<collectionCode id="ED164E877C763317B0C0A13BA3CAFA91" box="[199,252,1311,1333]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="27" pageNumber="28">ZISP</collectionCode>
|
||
examined, (
|
||
<location id="8ED880997C763317B183A13BA2E0FA91" LSID="urn:lsid:plazi:treatment:03AE67547C76332CB090A079A1E0FBFC:8ED880997C763317B183A13BA2E0FA91" box="[388,470,1311,1333]" country="China" county="Cockerell" municipality="Skorikov" name="Cyrillic" pageId="27" pageNumber="28" stateProvince="Qinghai">Cyrillic</location>
|
||
) ‘[
|
||
<location id="8ED880997C763317B1F4A13BA106FA91" LSID="urn:lsid:plazi:treatment:03AE67547C76332CB090A079A1E0FBFC:8ED880997C763317B1F4A13BA106FA91" box="[499,560,1311,1333]" country="China" county="Cockerell" municipality="Skorikov" name="Sinin" pageId="27" pageNumber="28" stateProvince="Qinghai">Sinin</location>
|
||
mts]’ (
|
||
<location id="8ED880997C763317B283A13BA1FBFA91" LSID="urn:lsid:plazi:treatment:03AE67547C76332CB090A079A1E0FBFC:8ED880997C763317B283A13BA1FBFA91" box="[644,717,1311,1333]" country="China" county="Cockerell" municipality="Skorikov" name="Xining" pageId="27" pageNumber="28" stateProvince="Qinghai">Xining</location>
|
||
range,
|
||
<collectingRegion id="49C318A07C763317B323A13BA04CFA91" box="[804,890,1311,1333]" country="China" name="Qinghai" pageId="27" pageNumber="28">Qinghai</collectingRegion>
|
||
,
|
||
<collectingCountry id="F31096D27C763317B38EA13BA0FAFA91" box="[905,972,1311,1333]" name="China" pageId="27" pageNumber="28">China</collectingCountry>
|
||
).
|
||
<location id="8ED880997C763317B3E7A104A724FA91" LSID="urn:lsid:plazi:treatment:03AE67547C76332CB090A079A1E0FBFC:8ED880997C763317B3E7A104A724FA91" box="[992,1042,1312,1333]" country="China" county="Cockerell" municipality="Skorikov" name="Note" pageId="27" pageNumber="28" stateProvince="Qinghai">Note</location>
|
||
1.
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||
<location id="8ED880997C763317B43BA13BA79AFA91" LSID="urn:lsid:plazi:treatment:03AE67547C76332CB090A079A1E0FBFC:8ED880997C763317B43BA13BA79AFA91" box="[1084,1196,1311,1333]" country="China" county="Cockerell" municipality="Skorikov" name="Synonymy" pageId="27" pageNumber="28" stateProvince="Qinghai">Synonymy</location>
|
||
with
|
||
<taxonomicName id="4C07ADC17C763317B4F7A13BA21AFAF7" ID-CoL="MFY7" authority="Cockerell" authorityName="Cockerell" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="waltoni">
|
||
<emphasis id="B9730A507C763317B4F7A13BA6ADFA91" box="[1264,1435,1311,1333]" italics="true" pageId="27" pageNumber="28">Bombus waltoni</emphasis>
|
||
Cockerell
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||
</taxonomicName>
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||
implied when
|
||
<taxonomicName id="4C07ADC17C763317B1C0A119A1ECFAF7" ID-CoL="MFY7" authority="Cockerell" authorityName="Cockerell" box="[455,730,1341,1363]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="waltoni">
|
||
<emphasis id="B9730A507C763317B1C0A119A159FAF7" box="[455,623,1341,1363]" italics="true" pageId="27" pageNumber="28">Bombus waltoni</emphasis>
|
||
Cockerell
|
||
</taxonomicName>
|
||
was synonymised with
|
||
<taxonomicName id="4C07ADC17C763317B3D2A119A64DFAF7" ID-CoL="MFG2" authority="Skorikov" authorityName="Skorikov" box="[981,1403,1341,1363]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="subSpecies" species="mendax" subSpecies="chinensis">
|
||
<emphasis id="B9730A507C763317B3D2A119A749FAF7" box="[981,1151,1341,1363]" italics="true" pageId="27" pageNumber="28">Bombus mendax</emphasis>
|
||
ssp.
|
||
<emphasis id="B9730A507C763317B4B2A119A620FAF7" box="[1205,1302,1341,1363]" italics="true" pageId="27" pageNumber="28">chinensis</emphasis>
|
||
Skorikov
|
||
</taxonomicName>
|
||
by
|
||
<collectingMunicipality id="6BDC4C387C763317B0C0A178A210FAD6" box="[199,294,1372,1394]" pageId="27" pageNumber="28">Skorikov</collectingMunicipality>
|
||
in
|
||
<bibRefCitation id="EF96ABB37C763317B140A178A2CCFAD6" author="Cockerell" box="[327,506,1372,1394]" pageId="27" pageNumber="28" refString="Cockerell, T. D. A. (1911) Bees from the Himalaya mountains. The Entomologist, 44, 176 - 177." type="journal article" year="1911">
|
||
<collectingCounty id="62D9AECE7C763317B140A178A29DFAD6" box="[327,427,1372,1394]" pageId="27" pageNumber="28">Cockerell</collectingCounty>
|
||
(1911)
|
||
</bibRefCitation>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BB8D6427C763317B090A15FA702FA0B" blockId="27.[151,1437,1219,1915]" pageId="27" pageNumber="28">
|
||
<materialsCitation id="3B6FDC1F7C763317B090A15FA702FA0B" ID-GBIF-Occurrence="1413110417" collectionCode="NHM" country="China" county="Note" location="Khamba Jong'" municipality="Gamba Dzong" pageId="27" pageNumber="28" specimenCount="1" stateProvince="Xizang" typeStatus="holotype">
|
||
<taxonomicName id="4C07ADC17C763317B090A15FA1B6FA35" ID-CoL="MFY7" authority="Cockerell 1910" authorityName="Cockerell" authorityYear="1910" box="[151,640,1403,1425]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="waltoni">
|
||
<emphasis id="B9730A507C763317B090A15FA270FA35" box="[151,326,1403,1425]" italics="true" pageId="27" pageNumber="28">Bombus waltoni</emphasis>
|
||
<bibRefCitation id="EF96ABB37C763317B153A15FA2CDFA35" author="Cockerell" box="[340,507,1403,1425]" pageId="27" pageNumber="28" refString="Cockerell, T. D. A. (1910) Some bees from high altitudes in the Himalaya mountains. Entomologist, 43, 238 - 239." type="journal article" year="1910">Cockerell 1910</bibRefCitation>
|
||
(September)
|
||
</taxonomicName>
|
||
:239, type-locality citation ‘Khamba Jong, Sikkim’.
|
||
<typeStatus id="54BC68E07C763317B4CFA15FA61EFA35" box="[1224,1320,1403,1425]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
|
||
queen by monotypy
|
||
<collectionCode id="ED164E877C763317B131A1BEA244FA0B" box="[310,370,1434,1455]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="27" pageNumber="28">NHM</collectionCode>
|
||
examined, ‘
|
||
<location id="8ED880997C763317B1E9A1BDA1BCFA0B" LSID="urn:lsid:plazi:treatment:03AE67547C76332CB090A079A1E0FBFC:8ED880997C763317B1E9A1BDA1BCFA0B" box="[494,650,1433,1455]" country="China" county="Note" municipality="Gamba Dzong" name="Khamba Jong'" pageId="27" pageNumber="28" stateProvince="Xizang">Khamba Jong’</location>
|
||
(
|
||
<collectingMunicipality id="6BDC4C387C763317B290A1BDA004FA0B" box="[663,818,1433,1455]" pageId="27" pageNumber="28">Gamba Dzong</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49C318A07C763317B33BA1BDA0BFFA0B" box="[828,905,1433,1455]" country="China" name="Xizang" pageId="27" pageNumber="28">Xizang</collectingRegion>
|
||
,
|
||
<collectingCountry id="F31096D27C763317B393A1BDA0E1FA0B" box="[916,983,1433,1455]" name="China" pageId="27" pageNumber="28">China</collectingCountry>
|
||
).
|
||
<collectingCounty id="62D9AECE7C763317B3E0A1BEA72FFA0B" box="[999,1049,1434,1455]" pageId="27" pageNumber="28">Note</collectingCounty>
|
||
2.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BB8D6427C763317B090A19CA012FA6A" blockId="27.[151,1437,1219,1915]" box="[151,804,1464,1486]" pageId="27" pageNumber="28">
|
||
<treatmentCitationGroup id="AB17F16C7C763317B090A19CA012FA6A" box="[151,804,1464,1486]" pageId="27" pageNumber="28">
|
||
<treatmentCitation id="0AA6F0537C763317B090A19CA1B1FA6A" author="Cockerell" box="[151,647,1464,1486]" page="176" pageId="27" pageNumber="28" year="1911">
|
||
<taxonomicName id="4C07ADC17C763317B090A19CA16FFA6A" ID-CoL="MFY7" authority="Cockerell" authorityName="Cockerell" authorityYear="1911" box="[151,601,1464,1486]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="waltoni">
|
||
<emphasis id="B9730A507C763317B090A19CA208FA6A" box="[151,318,1464,1486]" italics="true" pageId="27" pageNumber="28">Bombus waltoni</emphasis>
|
||
<bibRefCitation id="EF96ABB37C763317B142A19CA16FFA6A" author="Cockerell" box="[325,601,1464,1486]" pageId="27" pageNumber="28" refString="Cockerell, T. D. A. (1911) Bees from the Himalaya mountains. The Entomologist, 44, 176 - 177." type="journal article" year="1911">Cockerell; Cockerell 1911</bibRefCitation>
|
||
</taxonomicName>
|
||
:176
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA6F0537C763317B296A19DA016FA6A" author="Wu" box="[657,800,1464,1486]" page="281" pageId="27" pageNumber="28" year="1941">
|
||
<bibRefCitation id="EF96ABB37C763317B296A19DA1C5FA6A" author="Wu" box="[657,755,1464,1486]" pageId="27" pageNumber="28" refString="Wu, C. - F. (1941) Catalogus insectorum sinensium (Vol. 6). Peiping, Department of Biology, Yenching University." type="journal volume" year="1941">Wu 1941</bibRefCitation>
|
||
:281
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BB8D6427C763317B090A1FCA0B3FA49" blockId="27.[151,1437,1219,1915]" box="[151,901,1495,1517]" pageId="27" pageNumber="28">
|
||
<treatmentCitationGroup id="AB17F16C7C763317B090A1FCA0B3FA49" box="[151,901,1495,1517]" pageId="27" pageNumber="28">
|
||
[
|
||
<taxonomicName id="4C07ADC17C763317B098A1F3A1F3FA49" ID-CoL="5FZ5X" authority="Skorikov 1912:608" authorityName="Skorikov" authorityPageNumber="608" authorityYear="1912" box="[159,709,1495,1517]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="variety" species="waltoni" variety="kozloviellus">
|
||
<emphasis id="B9730A507C763317B098A1F3A271FA49" box="[159,327,1495,1517]" italics="true" pageId="27" pageNumber="28">Bombus waltoni</emphasis>
|
||
var.
|
||
<emphasis id="B9730A507C763317B17DA1F3A2C3FA49" box="[378,501,1495,1517]" italics="true" pageId="27" pageNumber="28">kozloviellus</emphasis>
|
||
<treatmentCitation id="0AA6F0537C763317B1FBA1F3A1F3FA49" author="Skorikov" box="[508,709,1495,1517]" page="608" pageId="27" pageNumber="28" year="1912">
|
||
<bibRefCitation id="EF96ABB37C763317B1FBA1F3A1AEFA49" author="Skorikov" box="[508,664,1495,1517]" pageId="27" pageNumber="28" refString="Skorikov, A. S. (1912) Neue Hummelformen (Hymenoptera, Bombidae). IV. Russkoe entomologicheskoe Obozrenie, 12, 606 - 610." type="journal article" year="1912">Skorikov 1912</bibRefCitation>
|
||
:608
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
, infrasubspecific.]
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BB8D6427C763317B090A1D1A132F9ED" blockId="27.[151,1437,1219,1915]" pageId="27" pageNumber="28">
|
||
<materialsCitation id="3B6FDC1F7C763317B090A1D1A132F9ED" ID-GBIF-Occurrence="1413110445" collectionCode="MNHU" country="China" county="Skorikov" location="Mongolei" municipality="Synonymised" pageId="27" pageNumber="28" specimenCount="1" stateProvince="Qinghai" typeStatus="lectotype">
|
||
<treatmentCitationGroup id="AB17F16C7C763317B090A1D1A132F9ED" pageId="27" pageNumber="28">
|
||
<taxonomicName id="4C07ADC17C763317B090A1D1A2C0F9AF" ID-CoL="MFTX" authority="Friese 1913:85" authorityName="Friese" authorityPageNumber="85" authorityYear="1913" box="[151,502,1525,1547]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="rufitarsis">
|
||
<emphasis id="B9730A507C763317B090A1D1A267F9AF" box="[151,337,1525,1547]" italics="true" pageId="27" pageNumber="28">Bombus rufitarsis</emphasis>
|
||
<treatmentCitation id="0AA6F0537C763317B15EA1D1A2C0F9AF" author="Friese" box="[345,502,1525,1547]" page="85" pageId="27" pageNumber="28" year="1913">
|
||
<bibRefCitation id="EF96ABB37C763317B15EA1D1A2EEF9AF" author="Friese" box="[345,472,1525,1547]" pageId="27" pageNumber="28" refString="Friese, H. (1913) Uber einige neue Apiden (Hym.). Archiv fur Naturgeschichte, 78, 85 - 89." type="journal article" year="1913">Friese 1913</bibRefCitation>
|
||
:85
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
, type-locality citation ‘Zentralasien’.
|
||
<emphasis id="B9730A507C763317B384A1D1A0C6F9AF" bold="true" box="[899,1008,1525,1547]" pageId="27" pageNumber="28">
|
||
<typeStatus id="54BC68E07C763317B384A1D1A0C6F9AF" box="[899,1008,1525,1547]" pageId="27" pageNumber="28" type="lectotype">Lectotype</typeStatus>
|
||
</emphasis>
|
||
queen (not a worker)
|
||
<collectionCode id="ED164E877C763317B4DCA1D2A61FF9AF" box="[1243,1321,1526,1547]" pageId="27" pageNumber="28">MNHU</collectionCode>
|
||
examined, ‘
|
||
<location id="8ED880997C763317B0CCA230A201F98E" LSID="urn:lsid:plazi:treatment:03AE67547C76332CB090A079A1E0FBFC:8ED880997C763317B0CCA230A201F98E" box="[203,311,1556,1578]" country="China" county="Skorikov" municipality="Synonymised" name="Mongolei" pageId="27" pageNumber="28" stateProvince="Qinghai">Mongolei</location>
|
||
,
|
||
<location id="8ED880997C763317B14EA230A2ADF98E" LSID="urn:lsid:plazi:treatment:03AE67547C76332CB090A079A1E0FBFC:8ED880997C763317B14EA230A2ADF98E" box="[329,411,1556,1578]" country="China" county="Skorikov" municipality="Synonymised" name="Monda'" pageId="27" pageNumber="28" stateProvince="Qinghai">Monda’</location>
|
||
believed incorrect (probably
|
||
<collectingRegion id="49C318A07C763317B2E9A230A072F98E" box="[750,836,1556,1578]" country="China" name="Qinghai" pageId="27" pageNumber="28">Qinghai</collectingRegion>
|
||
,
|
||
<collectingCountry id="F31096D27C763317B351A230A0AEF98E" box="[854,920,1556,1578]" name="China" pageId="27" pageNumber="28">China</collectingCountry>
|
||
).
|
||
<location id="8ED880997C763317B3B5A231A0D2F98E" LSID="urn:lsid:plazi:treatment:03AE67547C76332CB090A079A1E0FBFC:8ED880997C763317B3B5A231A0D2F98E" box="[946,996,1557,1578]" country="China" county="Skorikov" municipality="Synonymised" name="Note" pageId="27" pageNumber="28" stateProvince="Qinghai">Note</location>
|
||
3.
|
||
<collectingMunicipality id="6BDC4C387C763317B412A230A797F98E" box="[1045,1185,1556,1578]" pageId="27" pageNumber="28">Synonymised</collectingMunicipality>
|
||
with
|
||
<taxonomicName id="4C07ADC17C763317B4EBA230A21AF9ED" ID-CoL="MFY7" authority="Cockerell" authorityName="Cockerell" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="waltoni">
|
||
<emphasis id="B9730A507C763317B4EBA230A6AAF98E" box="[1260,1436,1556,1578]" italics="true" pageId="27" pageNumber="28">Bombus waltoni</emphasis>
|
||
Cockerell
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EF96ABB37C763317B154A217A136F9ED" author="Skorikov" box="[339,512,1587,1609]" pageId="27" pageNumber="28" refString="Skorikov, A. S. (1914) Les formes nouvelles des bourdons (Hymenoptera, Bombidae). VI. Russkoe entomologicheskoe Obozrenie, 14, 119 - 129." type="journal article" year="1914">
|
||
<collectingCounty id="62D9AECE7C763317B154A217A284F9ED" box="[339,434,1587,1609]" pageId="27" pageNumber="28">Skorikov</collectingCounty>
|
||
(1914)
|
||
</bibRefCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BB8D6427C763317B090A275A0F5F9C3" blockId="27.[151,1437,1219,1915]" box="[151,963,1617,1639]" pageId="27" pageNumber="28">
|
||
<treatmentCitationGroup id="AB17F16C7C763317B090A275A0F5F9C3" box="[151,963,1617,1639]" pageId="27" pageNumber="28">
|
||
<taxonomicName id="4C07ADC17C763317B090A275A123F9C3" authority="Cockerell" authorityName="Cockerell" box="[151,533,1617,1639]" class="Insecta" family="Apidae" genus="Mendacibombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="waltoni">
|
||
<emphasis id="B9730A507C763317B090A275A2A0F9C3" box="[151,406,1617,1639]" italics="true" pageId="27" pageNumber="28">Mendacibombus waltoni</emphasis>
|
||
(Cockerell)
|
||
</taxonomicName>
|
||
;
|
||
<treatmentCitation id="0AA6F0537C763317B227A275A1DCF9C3" author="Skorikov" box="[544,746,1617,1639]" page="125" pageId="27" pageNumber="28" year="1914">
|
||
<bibRefCitation id="EF96ABB37C763317B227A275A189F9C3" author="Skorikov" box="[544,703,1617,1639]" pageId="27" pageNumber="28" refString="Skorikov, A. S. (1914) Les formes nouvelles des bourdons (Hymenoptera, Bombidae). VI. Russkoe entomologicheskoe Obozrenie, 14, 119 - 129." type="journal article" year="1914">Skorikov 1914</bibRefCitation>
|
||
:125
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA6F0537C763317B2F2A275A089F9C3" author="Skorikov" box="[757,959,1617,1639]" page="149" pageId="27" pageNumber="28" year="1923">
|
||
<bibRefCitation id="EF96ABB37C763317B2F2A275A0A5F9C3" author="Skorikov" box="[757,915,1617,1639]" pageId="27" pageNumber="28" refString="Skorikov, A. S. (1923) [Palaearctic bumblebees. Part I. General biology (including zoogeography)]. Izvestiya Severnoi oblastnoi stantsii zashchity rastenii ot vreditelei, 4 (1922), 1 - 160." type="journal article" year="1923">Skorikov 1923</bibRefCitation>
|
||
:149
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BB8D6427C763317B090A254A06BF967" blockId="27.[151,1437,1219,1915]" pageId="27" pageNumber="28">
|
||
<materialsCitation id="3B6FDC1F7C763317B090A254A06BF967" ID-GBIF-Occurrence="1413110435" collectionCode="MNHU" collectorName="F. Koch & Regarded & Bischoff" country="China" location="Mongolei" municipality="Tippeti'" pageId="27" pageNumber="28" specimenCount="1" stateProvince="Qinghai" typeStatus="syntype">
|
||
<treatmentCitationGroup id="AB17F16C7C763317B090A254A06BF967" infered="true" pageId="27" pageNumber="28">
|
||
<taxonomicName id="4C07ADC17C763317B090A254A2DEF922" ID-CoL="68NP8" authority="Friese 1924:438" authorityName="Friese" authorityPageNumber="438" authorityYear="1924" box="[151,488,1648,1670]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="asellus">
|
||
<emphasis id="B9730A507C763317B090A254A20EF922" box="[151,312,1648,1670]" italics="true" pageId="27" pageNumber="28">Bombus asellus</emphasis>
|
||
<treatmentCitation id="0AA6F0537C763317B138A254A2DEF922" author="Friese" box="[319,488,1648,1670]" page="438" pageId="27" pageNumber="28" year="1924">
|
||
<bibRefCitation id="EF96ABB37C763317B138A254A28DF922" author="Friese" box="[319,443,1648,1670]" pageId="27" pageNumber="28" refString="Friese, H. (1924) Uber auffallende Hummelformen. (Hym. Apid.). Deutsche Entomologische Zeitschrift, 1924, 437 - 439. http: // dx. doi. org / 10.1002 / mmnd. 192419240508" type="journal article" year="1924">Friese 1924</bibRefCitation>
|
||
:438
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
, type-locality citation ‘
|
||
<location id="8ED880997C763317B2D2A254A008F922" LSID="urn:lsid:plazi:treatment:03AE67547C76332CB090A079A1E0FBFC:8ED880997C763317B2D2A254A008F922" box="[725,830,1648,1670]" country="China" municipality="Tippeti'" name="Mongolei" pageId="27" pageNumber="28" stateProvince="Qinghai">Mongolei</location>
|
||
bei
|
||
<collectingMunicipality id="6BDC4C387C763317B36DA254A08BF922" box="[874,957,1648,1670]" pageId="27" pageNumber="28">Tippeti’</collectingMunicipality>
|
||
believed incorrect (probably
|
||
<collectingRegion id="49C318A07C763317B4E8A254A673F922" box="[1263,1349,1648,1670]" country="China" name="Qinghai" pageId="27" pageNumber="28">Qinghai</collectingRegion>
|
||
,
|
||
<collectingCountry id="F31096D27C763317B548A254A6A7F922" box="[1359,1425,1648,1670]" name="China" pageId="27" pageNumber="28">China</collectingCountry>
|
||
).
|
||
<typeStatus id="54BC68E07C763317B0C0A2ABA22AF901" box="[199,284,1679,1701]" pageId="27" pageNumber="28" type="syntype">Syntype</typeStatus>
|
||
workers and male
|
||
<collectionCode id="ED164E877C763317B1EFA2B4A100F901" box="[488,566,1680,1701]" pageId="27" pageNumber="28">MNHU</collectionCode>
|
||
not found by
|
||
<collectorName id="26F2B3947C763317B2C9A2B4A014F901" box="[718,802,1679,1701]" pageId="27" pageNumber="28">F. Koch</collectorName>
|
||
and not seen (but taxon identity not in doubt).
|
||
<collectorName id="26F2B3947C763317B51CA2ABA648F901" box="[1307,1406,1679,1701]" pageId="27" pageNumber="28">Regarded</collectorName>
|
||
as conspecific with
|
||
<taxonomicName id="4C07ADC17C763317B17FA289A1BCF967" ID-CoL="MFY7" authority="Cockerell" authorityName="Cockerell" box="[376,650,1709,1731]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="waltoni">
|
||
<emphasis id="B9730A507C763317B17FA289A116F967" box="[376,544,1709,1731]" italics="true" pageId="27" pageNumber="28">Bombus waltoni</emphasis>
|
||
Cockerell
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EF96ABB37C763317B2B6A289A06FF967" author="Bischoff" box="[689,857,1709,1731]" pageId="27" pageNumber="28" refString="Bischoff, H. (1936) Schwedisch - chinesische wissenschaftliche Expedition nach den nordwestlichen Provinzen Chinas, unter Leitung von Dr. Sven Hedin und Prof. Su Ping - chang. Insekten gesammelt vom schwedischen Arzt der Expedition Dr. David Hummel 1927 - 1930. 56. Hymenoptera. 10. Bombinae. Arkiv for zoologi, 27 A, 1 - 27." type="journal article" year="1936">
|
||
<collectorName id="26F2B3947C763317B2B6A289A03CF967" box="[689,778,1709,1731]" pageId="27" pageNumber="28">Bischoff</collectorName>
|
||
(1936)
|
||
</bibRefCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BB8D6427C763317B090A2E8A2EFF89A" blockId="27.[151,1437,1219,1915]" pageId="27" pageNumber="28">
|
||
<treatmentCitationGroup id="AB17F16C7C763317B090A2E8A2EFF89A" pageId="27" pageNumber="28">
|
||
<taxonomicName id="4C07ADC17C763317B090A2E8A143F946" ID-CoL="MFY7" authority="Cockerell" authorityName="Cockerell" box="[151,629,1740,1762]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="waltoni" subGenus="Mendacibombus">
|
||
<emphasis id="B9730A507C763317B090A2E8A132F946" box="[151,516,1740,1762]" italics="true" pageId="27" pageNumber="28">Bombus (Mendacibombus) waltoni</emphasis>
|
||
Cockerell
|
||
</taxonomicName>
|
||
;
|
||
<treatmentCitation id="0AA6F0537C763317B285A2E8A07AF946" author="Richards" box="[642,844,1740,1762]" page="633" pageId="27" pageNumber="28" year="1930">
|
||
<bibRefCitation id="EF96ABB37C763317B285A2E8A029F946" author="Richards" box="[642,799,1740,1762]" pageId="27" pageNumber="28" refString="Richards, O. W. (1930) The humble - bees captured on the expeditions to Mt. Everest (Hymenoptera, Bombidae). Annals and Magazine of Natural History (10), 5, 633 - 658." type="journal article" year="1930">Richards 1930</bibRefCitation>
|
||
:633
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA6F0537C763317B35EA2E8A738F946" author="Tkalcu" box="[857,1038,1740,1762]" page="369" pageId="27" pageNumber="28" year="1961">
|
||
<bibRefCitation id="EF96ABB37C763317B35EA2E8A0E9F946" author="Tkalcu" box="[857,991,1740,1762]" pageId="27" pageNumber="28" refString="Tkalcu, B. (1961) Zur Hummelfauna der Umgebung Kuku - Nors (Hymenoptera, Bombinae). Casopis Ceskoslovenske spolecnosti entomologicke, 58, 344 - 379." type="journal article" year="1961">Tkalců 1961</bibRefCitation>
|
||
:369
|
||
</treatmentCitation>
|
||
; S.-
|
||
<treatmentCitation id="0AA6F0537C763317B431A2E8A7CBF946" author="Wang" box="[1078,1277,1740,1762]" page="430" pageId="27" pageNumber="28" year="1982">
|
||
<bibRefCitation id="EF96ABB37C763317B431A2E8A7F9F946" author="Wang" box="[1078,1231,1740,1762]" pageId="27" pageNumber="28" refString="Wang, S. - F. (1982) Hymenoptera: Apidae - Bombus. In: Q. - T. P. C. S. I. T. o. t. C. A. o. Sciences (Ed), Insects of Xizang. Science Press, Beijing, pp. 427 - 447." type="book chapter" year="1982">F. Wang 1982</bibRefCitation>
|
||
:430
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA6F0537C763317B50DA2E9A229F8A5" author="Williams" page="42" pageId="27" pageNumber="28" year="1991">
|
||
<bibRefCitation id="EF96ABB37C763317B50DA2E9A3C8F8A5" author="Williams" pageId="27" pageNumber="28" refString="Williams, P. H. (1991) The bumble bees of the Kashmir Himalaya (Hymenoptera: Apidae, Bombini). Bulletin of the British Museum (Natural History) (Entomology), 60, 1 - 204." type="journal article" year="1991">P.H. Williams 1991</bibRefCitation>
|
||
:42
|
||
</treatmentCitation>
|
||
; S.-
|
||
<treatmentCitation id="0AA6F0537C763317B140A2CFA12BF8A5" author="Wang" box="[327,541,1771,1793]" page="1424" pageId="27" pageNumber="28" year="1992">
|
||
<bibRefCitation id="EF96ABB37C763317B140A2CFA2D4F8A5" author="Wang" box="[327,482,1771,1793]" pageId="27" pageNumber="28" refString="Wang, S. - F. (1992) Hymenoptera: Apoidae (II) - Bombus. In: The series of the scientific expedition to the Hengduan Mountains region of Qinghai - Xizang Plateau. Chinese Academy of Sciences, pp. 1422 - 1430." type="book chapter" year="1992">F. Wang 1992</bibRefCitation>
|
||
:1424
|
||
</treatmentCitation>
|
||
; S.-
|
||
<treatmentCitation id="0AA6F0537C763317B241A2CFA068F8A5" author="Wang" box="[582,862,1771,1793]" page="303" pageId="27" pageNumber="28" year="1996">
|
||
<bibRefCitation id="EF96ABB37C763317B241A2CFA007F8A5" author="Wang" box="[582,817,1771,1793]" pageId="27" pageNumber="28" refString="Wang, S. - F. & Yao, J. (1996) Hymenoptera: Apodae [sic] - Bombini. In: F. - S. Huang (Ed), Insects of the Karakorum - Kunlun mountains. Science Press, Beijing, pp. 303 - 309." type="book chapter" year="1996">F. Wang & Yao 1996</bibRefCitation>
|
||
:303
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA6F0537C763317B36CA2C8A75AF8A5" author="Williams" box="[875,1132,1771,1793]" page="100" pageId="27" pageNumber="28" year="1998">
|
||
<bibRefCitation id="EF96ABB37C763317B36CA2C8A709F8A5" author="Williams" box="[875,1087,1771,1793]" pageId="27" pageNumber="28" refString="Williams, P. H. (1998) An annotated checklist of bumble bees with an analysis of patterns of description (Hymenoptera: Apidae, Bombini). Bulletin of The Natural History Museum (Entomology), 67, 79 - 152. [updated at www. nhm. ac. uk / bombus / accessed 2015]." type="journal article" year="1998">P.H. Williams 1998</bibRefCitation>
|
||
:100
|
||
</treatmentCitation>
|
||
;
|
||
<bibRefCitation id="EF96ABB37C763317B47DA2C8A679F8A5" author="Williams" box="[1146,1359,1771,1793]" pageId="27" pageNumber="28" refString="Williams, P. H. (2004) Genus Bombus Latreille. In: R. K. Gupta (Ed), An annotated catalogue of the bee species of the Indian Region. Jai Narain Vyas University, Jodhpur, p. Web page providing a list of bumblebees of India and adjacent countries with common synonyms and a summary of their distributions within the region." type="book" year="2004">P.H. Williams 2004</bibRefCitation>
|
||
:no. 30;
|
||
<treatmentCitation id="0AA6F0537C763317B0C0A32EA281F8BB" author="Burger" box="[199,439,1801,1823]" page="462" pageId="27" pageNumber="28" year="2009">
|
||
<bibRefCitation id="EF96ABB37C763317B0C0A32EA2BDF8BB" author="Burger" box="[199,395,1801,1823]" pageId="27" pageNumber="28" refString="Burger, F., Creutzburg, F., Hartmann, J. & Hartmann, M. (2009) Die Hummeln von Nepal (Insecta: Hymenoptera: Apidae: Bombus). In: M. Hartmann & J. Weipert (Eds), Biodiversitat und Naturausstattung im Himalaya III. Verein der Freunde und Forderer des Naturkundemuseums Erfurt e. V., Erfurt, pp. 455 - 462." type="book chapter" year="2009">
|
||
Burger
|
||
<emphasis id="B9730A507C763317B111A32EA27AF8BB" box="[278,332,1801,1823]" italics="true" pageId="27" pageNumber="28">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
:462
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA6F0537C763317B1C4A32DA1FDF8BB" author="Cameron" box="[451,715,1801,1823]" page="165" pageId="27" pageNumber="28" year="2007">
|
||
<bibRefCitation id="EF96ABB37C763317B1C4A32DA196F8BB" author="Cameron" box="[451,672,1801,1823]" pageId="27" pageNumber="28" refString="Cameron, S. A., Hines, H. M. & Williams, P. H. (2007) A comprehensive phylogeny of the bumble bees (Bombus). Biological Journal of the Linnean Society, 91, 161 - 188. http: // dx. doi. org / 10.1111 / j. 1095 - 8312.2007.00784. x" type="journal article" year="2007">
|
||
Cameron
|
||
<emphasis id="B9730A507C763317B22EA32EA169F8BB" box="[553,607,1801,1823]" italics="true" pageId="27" pageNumber="28">et al.</emphasis>
|
||
2007
|
||
</bibRefCitation>
|
||
:165
|
||
</treatmentCitation>
|
||
; P.H.
|
||
<treatmentCitation id="0AA6F0537C763317B30DA32DA727F8BB" author="Williams" box="[778,1041,1801,1823]" page="129" pageId="27" pageNumber="28" year="2009">
|
||
<bibRefCitation id="EF96ABB37C763317B30DA32DA0D3F8BB" author="Williams" box="[778,997,1801,1823]" pageId="27" pageNumber="28" refString="Williams, P. H., Tang, Y., Yao, J. & Cameron, S. (2009) The bumblebees of Sichuan (Hymenoptera: Apidae, Bombini). Systematics and Biodiversity, 7, 101 - 190." type="journal article" year="2009">
|
||
Williams
|
||
<emphasis id="B9730A507C763317B368A32EA093F8BB" box="[879,933,1801,1823]" italics="true" pageId="27" pageNumber="28">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
:129
|
||
</treatmentCitation>
|
||
; P.H. Williams
|
||
<emphasis id="B9730A507C763317B4B2A32EA7DCF8BB" box="[1205,1258,1801,1823]" italics="true" pageId="27" pageNumber="28">et al.</emphasis>
|
||
2010:124; An
|
||
<emphasis id="B9730A507C763317B58EA32EA3D4F89A" italics="true" pageId="27" pageNumber="28">et al.</emphasis>
|
||
2011:6;
|
||
<bibRefCitation id="EF96ABB37C763317B13AA30DA2E3F89A" author="An" box="[317,469,1832,1854]" pageId="27" pageNumber="28" refString="An, J. - D., Huang, J. - X., Shao, Y. - Q., Zhang, S. - W., Wang, B., Liu, X. - Y., Wu, J. & Williams, P. H. (2014) The bumblebees of North China (Apidae, Bombus Latreille). Zootaxa, 3830 (1), 1 - 89. http: // dx. doi. org / 10.11646 / zootaxa. 3830.1.1" type="journal article" year="2014">
|
||
An
|
||
<emphasis id="B9730A507C763317B164A30DA2AEF89A" box="[355,408,1832,1854]" italics="true" pageId="27" pageNumber="28">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BB8D6427C763317B090A363A02BF8F9" blockId="27.[151,1437,1219,1915]" box="[151,797,1863,1885]" pageId="27" pageNumber="28">
|
||
<taxonomicName id="4C07ADC17C763317B090A363A1FBF8F9" authority="Skorikov" authorityName="Skorikov" authorityYear="1931" box="[151,717,1863,1885]" class="Insecta" family="Apidae" genus="Mendacibombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="chinensis">
|
||
<emphasis id="B9730A507C763317B090A363A29EF8F9" box="[151,424,1863,1885]" italics="true" pageId="27" pageNumber="28">Mendacibombus chinensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EF96ABB37C763317B1B1A363A1FBF8F9" author="Skorikov" box="[438,717,1863,1885]" pageId="27" pageNumber="28" refString="Skorikov, A. S. (1931) Die Hummelfauna Turkestans und ihre Beziehungen zur zentralasiatischen Fauna (Hymenoptera, Bombidae). In: V. A. Lindholm (Ed.), Abhandlungen der Pamir - Expedition 1928. Academy of Sciences of the USSR, Leningrad, pp. 175 - 247." type="journal article" year="1931">Skorikov); Skorikov, 1931</bibRefCitation>
|
||
</taxonomicName>
|
||
:fig. 20.
|
||
</paragraph>
|
||
<paragraph id="8BB8D6427C763317B090A341A0A0F8DF" blockId="27.[151,1437,1219,1915]" box="[151,918,1893,1915]" pageId="27" pageNumber="28">
|
||
<treatmentCitationGroup id="AB17F16C7C763317B090A341A0A0F8DF" box="[151,918,1893,1915]" pageId="27" pageNumber="28">
|
||
<taxonomicName id="4C07ADC17C763317B090A341A1E6F8DF" ID-CoL="MFG2" authority="Skorikov" authorityName="Skorikov" box="[151,720,1893,1915]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="subSpecies" species="waltoni" subGenus="Mendacibombus" subSpecies="chinensis">
|
||
<emphasis id="B9730A507C763317B090A341A151F8DF" box="[151,615,1893,1915]" italics="true" pageId="27" pageNumber="28">Bombus (Mendacibombus) waltoni chinensis</emphasis>
|
||
Skorikov
|
||
</taxonomicName>
|
||
;
|
||
<treatmentCitation id="0AA6F0537C763317B2DCA341A0A4F8DF" author="Bischoff" box="[731,914,1893,1915]" page="17" pageId="27" pageNumber="28" year="1936">
|
||
<bibRefCitation id="EF96ABB37C763317B2DCA341A042F8DF" author="Bischoff" box="[731,884,1893,1915]" pageId="27" pageNumber="28" refString="Bischoff, H. (1936) Schwedisch - chinesische wissenschaftliche Expedition nach den nordwestlichen Provinzen Chinas, unter Leitung von Dr. Sven Hedin und Prof. Su Ping - chang. Insekten gesammelt vom schwedischen Arzt der Expedition Dr. David Hummel 1927 - 1930. 56. Hymenoptera. 10. Bombinae. Arkiv for zoologi, 27 A, 1 - 27." type="journal article" year="1936">Bischoff 1936</bibRefCitation>
|
||
:17
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D85C97C763310B090A382A6AAFC4B" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="description">
|
||
<paragraph id="8BB8D6427C763310B090A382A0E9FE1E" blockId="27.[151,1436,1957,2016]" lastBlockId="28.[151,1437,151,1981]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
|
||
<materialsCitation id="3B6FDC1F7C763310B090A382A714FF14" ID-GBIF-Occurrence="1413110439" county="Burhan Budai mountains" lastPageId="28" lastPageNumber="29" location="Note" municipality="Skorikov's" pageId="27" pageNumber="28" specimenCount="2" typeStatus="syntype">
|
||
<location id="8ED880997C763317B090A382A3F9F819" LSID="urn:lsid:plazi:treatment:03AE67547C76332CB090A079A1E0FBFC:8ED880997C763317B090A382A3F9F819" box="[151,207,1958,1981]" county="Burhan Budai mountains" municipality="Skorikov's" name="Note" pageId="27" pageNumber="28">Note</location>
|
||
1 (
|
||
<taxonomicName id="4C07ADC17C763317B0F3A381A269F81A" box="[244,351,1957,1982]" class="Insecta" family="Apidae" genus="Mendacibombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="chinensis">
|
||
<emphasis id="B9730A507C763317B0F3A381A269F81A" box="[244,351,1957,1982]" italics="true" pageId="27" pageNumber="28">chinensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<collectingMunicipality id="6BDC4C387C763317B171A381A2C4F81A" box="[374,498,1957,1982]" pageId="27" pageNumber="28">Skorikov’s</collectingMunicipality>
|
||
original description of the taxon
|
||
<taxonomicName id="4C07ADC17C763317B368A381A0EFF81A" box="[879,985,1957,1982]" class="Insecta" family="Apidae" genus="Mendacibombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="chinensis">
|
||
<emphasis id="B9730A507C763317B368A381A0EFF81A" box="[879,985,1957,1982]" italics="true" pageId="27" pageNumber="28">chinensis</emphasis>
|
||
</taxonomicName>
|
||
cites two mountain ranges (Xining and
|
||
<collectingCounty id="62D9AECE7C763317B090A3E3A2F2F844" box="[151,452,1991,2016]" pageId="27" pageNumber="28">Burhan Budai mountains</collectingCounty>
|
||
) distant from one another (and both credible) as the
|
||
<typeStatus id="54BC68E07C763317B465A3ECA7A5F844" box="[1122,1171,1992,2016]" pageId="27" pageNumber="28">type</typeStatus>
|
||
locality of the taxon
|
||
<taxonomicName id="4C07ADC17C713310B090A4B3A234FF14" box="[151,258,151,176]" class="Insecta" family="Apidae" genus="Mendacibombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="chinensis">
|
||
<emphasis id="B9730A507C713310B090A4B3A234FF14" box="[151,258,151,176]" italics="true" pageId="28" pageNumber="29">chinensis</emphasis>
|
||
</taxonomicName>
|
||
, so that there is likely to have been more than one original
|
||
<typeStatus id="54BC68E07C713310B3C2A4BCA714FF14" box="[965,1058,152,176]" pageId="28" pageNumber="29" type="syntype">syntype</typeStatus>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B6FDC1F7C713310B428A4B3A76CFEDC" ID-GBIF-Occurrence="1413110433" collectingDate="1990-05-30" collectionCode="ZISP" collectorName="Gr. Grzmailo & Cyrillic & Skorikova & Podbolotsk. & M. Podbolotskaya & Skorikov" county="Sinin" determinerName="PH Williams" location="The" municipality="Cryrillic" pageId="28" pageNumber="29" specimenCount="2" typeStatus="lectotype">
|
||
<location id="8ED880997C713310B428A4B3A76DFF14" LSID="urn:lsid:plazi:treatment:03AE67547C76332CB090A079A1E0FBFC:8ED880997C713310B428A4B3A76DFF14" box="[1071,1115,151,176]" county="Sinin" municipality="Cryrillic" name="The" pageId="28" pageNumber="29">The</location>
|
||
<collectionCode id="ED164E877C713310B462A4BCA796FF14" box="[1125,1184,152,176]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="28" pageNumber="29">ZISP</collectionCode>
|
||
collection studied by
|
||
<location id="8ED880997C713310B090A49DA236FF76" LSID="urn:lsid:plazi:treatment:03AE67547C76332CB090A079A1E0FBFC:8ED880997C713310B090A49DA236FF76" box="[151,256,185,210]" county="Sinin" municipality="Cryrillic" name="Skorikov" pageId="28" pageNumber="29">Skorikov</location>
|
||
contains a queen that agrees with the original description and carries the labels: (1) gold disc; (2) white, handwritten (
|
||
<collectingMunicipality id="6BDC4C387C713310B182A4FEA2DCFF57" box="[389,490,218,243]" pageId="28" pageNumber="29">Cryrillic</collectingMunicipality>
|
||
) ‘[[illegible]
|
||
<collectingCounty id="62D9AECE7C713310B28AA4FEA1FFFF57" box="[653,713,218,243]" pageId="28" pageNumber="29">Sinin</collectingCounty>
|
||
mts / [before]]
|
||
<date id="FFB9F0827C713310B389A4FFA0D4FF57" box="[910,994,219,243]" pageId="28" pageNumber="29" value="1990-05-30">
|
||
<collectingDate id="EFFD096A7C713310B389A4FFA0D4FF57" box="[910,994,219,243]" pageId="28" pageNumber="29" value="1990-05-30">30.v.90</collectingDate>
|
||
</date>
|
||
/ [
|
||
<collectorName id="26F2B3947C713310B40FA4FFA786FF57" box="[1032,1200,218,243]" pageId="28" pageNumber="29">Gr. Grzmailo</collectorName>
|
||
]’; (3) white, printed (
|
||
<collectorName id="26F2B3947C713310B098A4D8A3CCFEB1" box="[159,250,252,277]" pageId="28" pageNumber="29">Cyrillic</collectorName>
|
||
) ‘[
|
||
<emphasis id="B9730A507C713310B11AA4D8A29AFEB1" box="[285,428,252,277]" italics="true" pageId="28" pageNumber="29">
|
||
k.
|
||
<collectorName id="26F2B3947C713310B13DA4D8A29AFEB1" box="[314,428,252,277]" pageId="28" pageNumber="29">Skorikova</collectorName>
|
||
</emphasis>
|
||
]’; (4) red, handwritten ‘
|
||
<emphasis id="B9730A507C713310B2CEA4D9A09FFEB1" box="[713,937,252,277]" italics="true" pageId="28" pageNumber="29">
|
||
<typeStatus id="54BC68E07C713310B2CEA4D9A073FEB0" box="[713,837,253,276]" pageId="28" pageNumber="29" type="lectotype">Lectotypus</typeStatus>
|
||
<taxonomicName id="4C07ADC17C713310B34AA4D8A09FFEB1" box="[845,937,252,277]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">Bombus</taxonomicName>
|
||
</emphasis>
|
||
/
|
||
<taxonomicName id="4C07ADC17C713310B3C4A4D8A75BFEB1" authority="Subsp." authorityName="Subsp." box="[963,1133,252,277]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="44" phylum="Arthropoda" rank="species" species="mendax">
|
||
<emphasis id="B9730A507C713310B3C4A4D8A75BFEB1" box="[963,1133,252,277]" italics="true" pageId="28" pageNumber="29">mendax subsp.</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C07ADC17C713310B482A4D8A607FEB1" authority="Skor." authorityName="Skor." box="[1157,1329,252,277]" class="Insecta" family="Apidae" genus="Mendacibombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="chinensis">
|
||
<emphasis id="B9730A507C713310B482A4D8A607FEB1" box="[1157,1329,252,277]" italics="true" pageId="28" pageNumber="29">chinensis Skor.</emphasis>
|
||
</taxonomicName>
|
||
/ design.
|
||
<collectorName id="26F2B3947C713310B090A539A228FE92" box="[151,286,285,310]" pageId="28" pageNumber="29">
|
||
<emphasis id="B9730A507C713310B090A539A22EFE92" box="[151,280,285,310]" italics="true" pageId="28" pageNumber="29">Podbolotsk</emphasis>
|
||
.
|
||
</collectorName>
|
||
’ (
|
||
<collectorName id="26F2B3947C713310B13FA53AA13BFE92" box="[312,525,285,310]" pageId="28" pageNumber="29">M. Podbolotskaya</collectorName>
|
||
, unpublished); (5) green, printed ‘
|
||
<taxonomicName id="4C07ADC17C713310B39FA539A763FE92" box="[920,1109,285,310]" class="Insecta" family="Apidae" genus="Mendacibombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9730A507C713310B39FA539A763FE92" box="[920,1109,285,310]" italics="true" pageId="28" pageNumber="29">Mendacibombus</emphasis>
|
||
</taxonomicName>
|
||
/ MD# 3521 det. PHW’; (6) red, printed ‘
|
||
<typeStatus id="54BC68E07C713310B129A51BA2E0FEF3" box="[302,470,319,343]" pageId="28" pageNumber="29" type="lectotype">LECTOTYPE</typeStatus>
|
||
[female] /
|
||
<taxonomicName id="4C07ADC17C713310B267A51AA016FEF3" box="[608,800,318,343]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="mendax">
|
||
<emphasis id="B9730A507C713310B267A51AA016FEF3" box="[608,800,318,343]" italics="true" pageId="28" pageNumber="29">Bombus mendax</emphasis>
|
||
</taxonomicName>
|
||
ssp. /
|
||
<taxonomicName id="4C07ADC17C713310B372A51AA0D6FEF3" box="[885,992,318,343]" class="Insecta" family="Apidae" genus="Mendacibombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="chinensis">
|
||
<emphasis id="B9730A507C713310B372A51AA0D6FEF3" box="[885,992,318,343]" italics="true" pageId="28" pageNumber="29">chinensis</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<collectorName id="26F2B3947C713310B3F9A51AA75EFEF3" box="[1022,1128,318,343]" pageId="28" pageNumber="29">Skorikov</collectorName>
|
||
, 1910 / det.
|
||
<determinerName id="7E643A867C713310B502A51BA6AAFEF3" box="[1285,1436,318,343]" pageId="28" pageNumber="29">PH Williams</determinerName>
|
||
2012’; (7) white, printed ‘[female]
|
||
<taxonomicName id="4C07ADC17C713310B24EA57BA193FEDC" box="[585,677,351,376]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9730A507C713310B24EA57BA193FEDC" box="[585,677,351,376]" italics="true" pageId="28" pageNumber="29">Bombus</emphasis>
|
||
</taxonomicName>
|
||
/ (
|
||
<taxonomicName id="4C07ADC17C713310B2D6A57BA0B8FEDC" box="[721,910,351,376]" class="Insecta" family="Apidae" genus="Mendacibombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9730A507C713310B2D6A57BA0B8FEDC" box="[721,910,351,376]" italics="true" pageId="28" pageNumber="29">Mendacibombus</emphasis>
|
||
</taxonomicName>
|
||
) /
|
||
<taxonomicName id="4C07ADC17C713310B3BDA57BA726FEDC" box="[954,1040,351,376]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="waltoni">
|
||
<emphasis id="B9730A507C713310B3BDA57BA726FEDC" box="[954,1040,351,376]" italics="true" pageId="28" pageNumber="29">waltoni</emphasis>
|
||
</taxonomicName>
|
||
/ det
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B6FDC1F7C713310B46CA544A0E9FE1E" ID-GBIF-Occurrence="1413110443" country="Philippines" location="This" municipality="Skorikov's" pageId="28" pageNumber="29" specimenCount="2" typeStatus="syntype">
|
||
<collectingCountry id="F31096D27C713310B46CA544A7B9FED3" box="[1131,1167,352,375]" name="Philippines" pageId="28" pageNumber="29">PH</collectingCountry>
|
||
Williams 2012’.
|
||
<location id="8ED880997C713310B56EA57BA6AAFEDC" LSID="urn:lsid:plazi:treatment:03AE67547C76332CB090A079A1E0FBFC:8ED880997C713310B56EA57BA6AAFEDC" box="[1385,1436,351,376]" country="Philippines" municipality="Skorikov's" name="This" pageId="28" pageNumber="29">This</location>
|
||
specimen, which is complete, is regarded as one of
|
||
<collectingMunicipality id="6BDC4C387C713310B2E9A5A5A05CFE3E" box="[750,874,385,410]" pageId="28" pageNumber="29">Skorikov’s</collectingMunicipality>
|
||
<typeStatus id="54BC68E07C713310B375A5A6A0E0FE3E" box="[882,982,386,410]" pageId="28" pageNumber="29" type="syntype">syntypes</typeStatus>
|
||
and is designated here as the
|
||
<typeStatus id="54BC68E07C713310B534A5A5A6AAFE3E" box="[1331,1436,385,410]" pageId="28" pageNumber="29" type="lectotype">lectotype</typeStatus>
|
||
in order to reduce uncertainty in the identity and application of the name.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BB8D6427C713310B0C0A5E1A153FDBB" blockId="28.[151,1437,151,1981]" pageId="28" pageNumber="29">
|
||
Note 2 (
|
||
<taxonomicName id="4C07ADC17C713310B12DA5E0A2B6FE79" box="[298,384,452,477]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="waltoni">
|
||
<emphasis id="B9730A507C713310B12DA5E0A2B6FE79" box="[298,384,452,477]" italics="true" pageId="28" pageNumber="29">waltoni</emphasis>
|
||
</taxonomicName>
|
||
). The original publication specifies that there was only one type specimen of
|
||
<taxonomicName id="4C07ADC17C713310B518A5E0A643FE79" box="[1311,1397,452,477]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="waltoni">
|
||
<emphasis id="B9730A507C713310B518A5E0A643FE79" box="[1311,1397,452,477]" italics="true" pageId="28" pageNumber="29">waltoni</emphasis>
|
||
</taxonomicName>
|
||
by Cockerell, so the single specimen in the NHM collection labeled ‘Khamba Jong’ is regarded as the holotype by monotypy (
|
||
<bibRefCitation id="EF96ABB37C713310B11DA623A16FFDBB" author="ICZN" box="[282,601,518,543]" pageId="28" pageNumber="29" refString="ICZN (1999) International code of zoological nomenclature (4 ed.). London, International Commission on Zoological Nomenclature." type="book chapter" year="1999">ICZN, 1999: Article 73.1.2</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB8D6427C713310B0C0A60CA6AAFC4B" blockId="28.[151,1437,151,1981]" pageId="28" pageNumber="29">
|
||
<materialsCitation id="3B6FDC1F7C713310B0C0A60CA194FCB3" ID-GBIF-Occurrence="1413110446" collectionCode="MNHU" collectorName="B. Tkalcu & Zool." country="Mongolia" county="Friese" determinerName="Tkalcu" location="Weiske'" municipality="Monda" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Mus" typeStatus="lectotype">
|
||
Note 3 (
|
||
<taxonomicName id="4C07ADC17C713310B123A60CA2BBFDE4" box="[292,397,552,576]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="rufitarsis">
|
||
<emphasis id="B9730A507C713310B123A60CA2BBFDE4" box="[292,397,552,576]" italics="true" pageId="28" pageNumber="29">rufitarsis</emphasis>
|
||
</taxonomicName>
|
||
). Friese’s original description of the taxon
|
||
<taxonomicName id="4C07ADC17C713310B37BA60CA0D0FDE4" box="[892,998,552,576]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="rufitarsis">
|
||
<emphasis id="B9730A507C713310B37BA60CA0D0FDE4" box="[892,998,552,576]" italics="true" pageId="28" pageNumber="29">rufitarsis</emphasis>
|
||
</taxonomicName>
|
||
lists four workers, although the queen described here is labelled as a worker by Friese. The
|
||
<collectionCode id="ED164E877C713310B305A669A06FFDC0" box="[770,857,589,612]" pageId="28" pageNumber="29">MNHU</collectionCode>
|
||
collection studied by
|
||
<collectingCounty id="62D9AECE7C713310B45DA668A796FDC1" box="[1114,1184,588,613]" pageId="28" pageNumber="29">Friese</collectingCounty>
|
||
contains a queen that agrees with the original description and carries the labels: (1) white, printed ‘
|
||
<collectingCountry id="F31096D27C713310B444A64BA78EFD2C" box="[1091,1208,623,648]" name="Mongolia" pageId="28" pageNumber="29">Mongolei</collectingCountry>
|
||
/
|
||
<collectingMunicipality id="6BDC4C387C713310B4DFA64BA61FFD2C" box="[1240,1321,623,648]" pageId="28" pageNumber="29">Monda</collectingMunicipality>
|
||
/ 6. 0 8 /
|
||
<location id="8ED880997C713310B090A6B0A3CEFD09" LSID="urn:lsid:plazi:treatment:03AE67547C76332CB090A079A1E0FBFC:8ED880997C713310B090A6B0A3CEFD09" box="[151,248,660,685]" country="Mongolia" county="Friese" municipality="Monda" name="Weiske'" pageId="28" pageNumber="29" stateProvince="Mus">Weiske’</location>
|
||
; (2) white, handwritten by
|
||
<location id="8ED880997C713310B23DA6B0A1B6FD09" LSID="urn:lsid:plazi:treatment:03AE67547C76332CB090A079A1E0FBFC:8ED880997C713310B23DA6B0A1B6FD09" box="[570,640,660,685]" country="Mongolia" county="Friese" municipality="Monda" name="Friese" pageId="28" pageNumber="29" stateProvince="Mus">Friese</location>
|
||
‘
|
||
<taxonomicName id="4C07ADC17C713310B294A6B0A1D9FD09" box="[659,751,660,685]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9730A507C713310B294A6B0A1D9FD09" box="[659,751,660,685]" italics="true" pageId="28" pageNumber="29">Bombus</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C07ADC17C713310B30DA6B0A042FD08" box="[778,884,660,684]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="rufitarsis">
|
||
<emphasis id="B9730A507C713310B30DA6B0A042FD08" box="[778,884,660,684]" italics="true" pageId="28" pageNumber="29">rufitarsis</emphasis>
|
||
</taxonomicName>
|
||
/ [worker] 1909
|
||
<emphasis id="B9730A507C713310B447A6B1A76DFD08" box="[1088,1115,661,684]" italics="true" pageId="28" pageNumber="29">Fr</emphasis>
|
||
. Det.’; (3) maroon, printed ‘
|
||
<location id="8ED880997C713310B09CA69CA3D0FD74" LSID="urn:lsid:plazi:treatment:03AE67547C76332CB090A079A1E0FBFC:8ED880997C713310B09CA69CA3D0FD74" box="[155,230,696,720]" country="Mongolia" county="Friese" municipality="Monda" name="Type'" pageId="28" pageNumber="29" stateProvince="Mus">Type’</location>
|
||
; (4) white, handwritten by
|
||
<location id="8ED880997C713310B244A693A1BFFD74" LSID="urn:lsid:plazi:treatment:03AE67547C76332CB090A079A1E0FBFC:8ED880997C713310B244A693A1BFFD74" box="[579,649,695,720]" country="Mongolia" county="Friese" municipality="Monda" name="Friese" pageId="28" pageNumber="29" stateProvince="Mus">Friese</location>
|
||
‘
|
||
<emphasis id="B9730A507C713310B2A6A69DA1EEFD6B" box="[673,728,696,720]" italics="true" pageId="28" pageNumber="29">B.?</emphasis>
|
||
/
|
||
<emphasis id="B9730A507C713310B2FAA69DA05AFD74" box="[765,876,697,720]" italics="true" pageId="28" pageNumber="29">pyrosoma</emphasis>
|
||
/ [worker] 1900 Friese
|
||
<emphasis id="B9730A507C713310B4A5A69DA78BFD74" box="[1186,1213,697,720]" italics="true" pageId="28" pageNumber="29">Fr</emphasis>
|
||
. Det.’; (5) white, handwritten by
|
||
<location id="8ED880997C713310B15AA6F8A29AFD51" LSID="urn:lsid:plazi:treatment:03AE67547C76332CB090A079A1E0FBFC:8ED880997C713310B15AA6F8A29AFD51" box="[349,428,732,757]" country="Mongolia" county="Friese" municipality="Monda" name="Tkalcu" pageId="28" pageNumber="29" stateProvince="Mus">Tkalců</location>
|
||
‘
|
||
<emphasis id="B9730A507C713310B1C0A6F9A157FD50" box="[455,609,733,756]" italics="true" pageId="28" pageNumber="29">
|
||
<typeStatus id="54BC68E07C713310B1C0A6F9A157FD50" box="[455,609,733,756]" pageId="28" pageNumber="29" type="lectotype">LECTOTYPE</typeStatus>
|
||
</emphasis>
|
||
/
|
||
<taxonomicName id="4C07ADC17C713310B28CA6F8A1D1FD51" box="[651,743,732,757]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9730A507C713310B28CA6F8A1D1FD51" box="[651,743,732,757]" italics="true" pageId="28" pageNumber="29">Bombus</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C07ADC17C713310B317A6F8A0E2FD50" authority="Friese" authorityName="Friese" box="[784,980,732,756]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="rufitarsis">
|
||
<emphasis id="B9730A507C713310B317A6F8A0E2FD50" box="[784,980,732,756]" italics="true" pageId="28" pageNumber="29">rufitarsis Friese</emphasis>
|
||
</taxonomicName>
|
||
/ [female]
|
||
<emphasis id="B9730A507C713310B469A6F8A7D9FD51" box="[1134,1263,732,757]" italics="true" pageId="28" pageNumber="29">
|
||
<determinerName id="7E643A867C713310B469A6F8A78AFD51" box="[1134,1212,732,757]" pageId="28" pageNumber="29">Tkalců</determinerName>
|
||
det
|
||
</emphasis>
|
||
.’ (
|
||
<collectorName id="26F2B3947C713310B51DA6F8A6AEFD51" box="[1306,1432,732,757]" pageId="28" pageNumber="29">B. Tkalců</collectorName>
|
||
, unpublished); (6) white, printed ‘
|
||
<collectorName id="26F2B3947C713310B218A6DBA157FCBC" box="[543,609,767,792]" pageId="28" pageNumber="29">Zool.</collectorName>
|
||
<collectingRegion id="49C318A07C713310B26BA724A194FCB3" box="[620,674,768,791]" country="Turkey" name="Mus" pageId="28" pageNumber="29">Mus</collectingRegion>
|
||
</materialsCitation>
|
||
. / Berlin’; (7) green, printed ‘
|
||
<taxonomicName id="4C07ADC17C713310B40BA6DBA7FFFCBC" box="[1036,1225,767,792]" class="Insecta" family="Apidae" genus="Mendacibombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9730A507C713310B40BA6DBA7FFFCBC" box="[1036,1225,767,792]" italics="true" pageId="28" pageNumber="29">Mendacibombus</emphasis>
|
||
</taxonomicName>
|
||
/ MD# 3538 det. PHW’; (7) red, printed ‘LECTOTYPE [female] /
|
||
<taxonomicName id="4C07ADC17C713310B2CCA700A011FC99" box="[715,807,804,829]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9730A507C713310B2CCA700A011FC99" box="[715,807,804,829]" italics="true" pageId="28" pageNumber="29">Bombus</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C07ADC17C713310B33BA700A093FC98" box="[828,933,804,828]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="rufitarsis">
|
||
<emphasis id="B9730A507C713310B33BA700A093FC98" box="[828,933,804,828]" italics="true" pageId="28" pageNumber="29">rufitarsis</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<bibRefCitation id="EF96ABB37C713310B3BCA700A77EFC98" author="Friese" box="[955,1096,804,829]" pageId="28" pageNumber="29" refString="Friese, H. (1913) Uber einige neue Apiden (Hym.). Archiv fur Naturgeschichte, 78, 85 - 89." type="journal article" year="1913">Friese, 1913</bibRefCitation>
|
||
/ det. PH Williams 2012’; (8) white, printed ‘[female]
|
||
<taxonomicName id="4C07ADC17C713310B1B5A763A138FCC4" box="[434,526,839,864]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9730A507C713310B1B5A763A138FCC4" box="[434,526,839,864]" italics="true" pageId="28" pageNumber="29">Bombus</emphasis>
|
||
</taxonomicName>
|
||
/ (
|
||
<taxonomicName id="4C07ADC17C713310B237A763A1DBFCC4" box="[560,749,839,864]" class="Insecta" family="Apidae" genus="Mendacibombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9730A507C713310B237A763A1DBFCC4" box="[560,749,839,864]" italics="true" pageId="28" pageNumber="29">Mendacibombus</emphasis>
|
||
</taxonomicName>
|
||
) /
|
||
<taxonomicName id="4C07ADC17C713310B308A763A053FCC4" box="[783,869,839,864]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="waltoni">
|
||
<emphasis id="B9730A507C713310B308A763A053FCC4" box="[783,869,839,864]" italics="true" pageId="28" pageNumber="29">waltoni</emphasis>
|
||
</taxonomicName>
|
||
/ det.
|
||
<materialsCitation id="3B6FDC1F7C713310B3B7A76CA633FC69" ID-GBIF-Occurrence="1413110441" country="Philippines" location="Qinghai" pageId="28" pageNumber="29" specimenCount="4" stateProvince="Qinghai" typeStatus="syntype">
|
||
<collectingCountry id="F31096D27C713310B3B7A76CA0E2FCFB" box="[944,980,840,863]" name="Philippines" pageId="28" pageNumber="29">PH</collectingCountry>
|
||
Williams 2012’. This specimen, which is complete, is regarded as one of Friese’s
|
||
<typeStatus id="54BC68E07C713310B288A749A1C2FC21" box="[655,756,877,901]" pageId="28" pageNumber="29" type="syntype">syntypes</typeStatus>
|
||
and is designated here as the
|
||
<typeStatus id="54BC68E07C713310B45FA748A7F7FC21" box="[1112,1217,876,901]" pageId="28" pageNumber="29" type="lectotype">lectotype</typeStatus>
|
||
in order to reduce uncertainty in the identity and application of the name. For the same reasons as given in note 1
|
||
<collectedFrom id="381B95BF7C713310B4EEA7B4A6A2FC0C" box="[1257,1428,911,936]" pageId="28" pageNumber="29">
|
||
on
|
||
<taxonomicName id="4C07ADC17C713310B50AA7B5A6A2FC0C" box="[1293,1428,911,936]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="superbus">
|
||
<emphasis id="B9730A507C713310B50AA7B5A6A2FC0C" box="[1293,1428,911,936]" italics="true" pageId="28" pageNumber="29">B. superbus</emphasis>
|
||
</taxonomicName>
|
||
</collectedFrom>
|
||
, we interpret the origin of the
|
||
<typeStatus id="54BC68E07C713310B1E0A790A166FC69" box="[487,592,948,973]" pageId="28" pageNumber="29" type="lectotype">lectotype</typeStatus>
|
||
and the
|
||
<typeStatus id="54BC68E07C713310B2B5A791A1D5FC69" box="[690,739,949,973]" pageId="28" pageNumber="29">type</typeStatus>
|
||
locality as most likely to have been in
|
||
<collectingRegion id="49C318A07C713310B4A2A790A633FC69" box="[1189,1285,948,973]" country="China" name="Qinghai" pageId="28" pageNumber="29">Qinghai</collectingRegion>
|
||
</materialsCitation>
|
||
. Even so, the precise type locality remains very uncertain and consequently no location for the lectotype is shown on our map.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D85C97C713310B0C0A7D8A674F964" pageId="28" pageNumber="29" type="etymology">
|
||
<paragraph id="8BB8D6427C713310B0C0A7D8A106FB6C" blockId="28.[151,1437,151,1981]" pageId="28" pageNumber="29">
|
||
<emphasis id="B9730A507C713310B0C0A7D8A266FBB1" bold="true" box="[199,336,1020,1045]" pageId="28" pageNumber="29">Etymology.</emphasis>
|
||
The species is named after H. Walton, the medical officer during the 1904 British expedition known as the
|
||
<collectingRegion id="49C318A07C713310B13CA03BA241FB9C" box="[315,375,1055,1080]" country="China" name="Xizang" pageId="28" pageNumber="29">Tibet</collectingRegion>
|
||
Frontier Commission with F. Younghusband as Commissioner. According to the specimen labels, in 1904 Walton collected at least two specimens of the species between Phari and Gyangzê (NHM). In
|
||
<date id="FFB9F0827C713310B090A043A23CFB24" box="[151,266,1127,1152]" pageId="28" pageNumber="29" value="1903-07">July 1903</date>
|
||
the previous expedition of the Commission, which was also led by Younghusband, visited Gamba Dzong (
|
||
<bibRefCitation id="EF96ABB37C713310B0F6A0A8A2A1FB00" author="Hopkirk" box="[241,407,1164,1189]" pageId="28" pageNumber="29" refString="Hopkirk, P. (1982) Tresspassers on the roof of the world: the race for Lhasa. London, John Murray." type="book" year="1982">Hopkirk 1982</bibRefCitation>
|
||
), just north of
|
||
<collectingRegion id="49C318A07C713310B239A0A8A1A3FB01" box="[574,661,1164,1189]" country="India" name="Sikkim" pageId="28" pageNumber="29">Sikkim</collectingRegion>
|
||
. This is the locality where the
|
||
<typeStatus id="54BC68E07C713310B3F6A0A9A714FB01" box="[1009,1058,1165,1189]" pageId="28" pageNumber="29">type</typeStatus>
|
||
specimen was collected, but it is unclear who collected the
|
||
<typeStatus id="54BC68E07C713310B1C3A08BA11AFB6C" box="[452,556,1199,1224]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB8D6427C713310B0C0A0F0A674F964" blockId="28.[151,1437,151,1981]" pageId="28" pageNumber="29">
|
||
<emphasis id="B9730A507C713310B0C0A0F0A2CAFB49" bold="true" box="[199,508,1236,1261]" pageId="28" pageNumber="29">Taxonomy and variation.</emphasis>
|
||
This species shows two principal colour patterns of the hair: extensively black, but with either an extensive orange tail, or with a black tail. The orange-tailed pattern matches the original description of the taxon
|
||
<taxonomicName id="4C07ADC17C713310B1BCA138A127FA91" box="[443,529,1308,1333]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="waltoni">
|
||
<emphasis id="B9730A507C713310B1BCA138A127FA91" box="[443,529,1308,1333]" italics="true" pageId="28" pageNumber="29">waltoni</emphasis>
|
||
</taxonomicName>
|
||
. Both colour patterns are unique within the subgenus. No specimens of this species have distinct pale bands on the thorax and none has yellow hair anywhere on the body. Specimens of the species from the Kunlun (MD#3868, 3869) and Tanggula (MD#4132‒4137) mountains have the hair of the thoracic dorsum and T1 black and of T2‒5 orange, and are unusual for the species because the hairs are mostly not white-tipped. A specimen from the far west of the range in Ladakh (MD#4023) has no white hairs at all and T2 is predominantly orange. In the east and south, individuals of this species usually have many white-tipped hairs and often have intermixed white hairs (e.g. MD#193). Sometimes T1 has many white hairs intermixed and T2‒
|
||
<date id="FFB9F0827C713310B0C1A233A23DF994" box="[198,267,1559,1584]" pageId="28" pageNumber="29">5 may</date>
|
||
have white hairs in the posterior quarter of each tergum (e.g. MD#1284). Some males (and according to Friese’s description, some workers, although we have not seen examples of these) in the northern and eastern areas of the distribution have little or no orange hair on the metasomal terga (Friese’s taxon
|
||
<taxonomicName id="4C07ADC17C713310B4AAA27BA7CBF9DC" box="[1197,1277,1631,1656]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="asellus">
|
||
<emphasis id="B9730A507C713310B4AAA27BA7CBF9DC" box="[1197,1277,1631,1656]" italics="true" pageId="28" pageNumber="29">asellus</emphasis>
|
||
</taxonomicName>
|
||
), although all males share the same form of the genitalia and similar COI sequences (
|
||
<figureCitation id="133CCAC77C713310B3B8A2A0A722F938" box="[959,1044,1668,1693]" captionStart="FIGURE 13" captionStartId="16.[151,250,1733,1755]" captionTargetBox="[314,1260,194,1721]" captionTargetId="figure@16.[314,1261,193,1722]" captionTargetPageId="16" captionText="FIGURE 13. Assigning names to species from an estimate of phylogeny for 130 COI sequences of the subgenus Mendacibombus. Based on a BEAST analysis of the 30 unique haplotypes (Fig. 11), with the remaining sequences interpolated back into the tree with 0 branch lengths from their matching samples (where a sequence matches more than one longer sequence within a species in the Collapse results, it is added after the first matching sequence, with the addition of the two short-sequence yellow unbanded specimens from Pakistan, MD # 4058, 4059). The sequence labels follow the format used in Figs 11, 12. Lineages accepted as species from the GMYC models (Fig. 11) are shown as thick branches, the most recent common ancestors of each species are shown with a black spot, and branches within species are shown as thin branches. Asterisks mark samples used as informal proxies for the type specimens of each of the taxon names (Table 1). The proxy sample for the type specimen for the oldest available name (the valid name) for each species (from Fig. 11) is marked in bold. The interpretations of the species with their valid names are shown in grey to the right." httpUri="https://zenodo.org/record/192309/files/figure.png" pageId="28" pageNumber="29">Fig. 13</figureCitation>
|
||
: the orange-tailed taxon
|
||
<emphasis id="B9730A507C713310B52AA2A0A382F964" italics="true" pageId="28" pageNumber="29">
|
||
<taxonomicName id="4C07ADC17C713310B52AA2A0A6B5F939" box="[1325,1411,1668,1693]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" sensu="stricto" species="waltoni">waltoni</taxonomicName>
|
||
<taxonomicNameLabel id="A240B72B7C713310B58EA2A1A382F964" pageId="28" pageNumber="29" sensu="stricto">s. str</taxonomicNameLabel>
|
||
</emphasis>
|
||
. MD#1482 and the black-tailed taxon
|
||
<taxonomicName id="4C07ADC17C713310B268A283A189F964" box="[623,703,1703,1728]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="asellus">
|
||
<emphasis id="B9730A507C713310B268A283A189F964" box="[623,703,1703,1728]" italics="true" pageId="28" pageNumber="29">asellus</emphasis>
|
||
</taxonomicName>
|
||
MD#300). The form of the male genitalia is diagnostic.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D85C97C71332CB0C0A2E8A258FE09" lastPageId="32" lastPageNumber="33" pageId="28" pageNumber="29" type="description">
|
||
<paragraph id="8BB8D6427C713310B0C0A2E8A7FBF819" blockId="28.[151,1437,151,1981]" pageId="28" pageNumber="29">
|
||
COI sequences show two principal groups of haplotypes, although these do not coincide with known morphological or colour-pattern differences. The majority of individuals are in one group. The second divergent group (differing in at least 12 nucleotides) of three individuals (MD#267, 1482, 1483) is from a narrow region in the south, associated with the Himalaya (from Nepal and from the Yadong region of Xizang, the region of the type of the taxon
|
||
<emphasis id="B9730A507C713310B123A378A285F8D0" box="[292,435,1884,1909]" italics="true" pageId="28" pageNumber="29">
|
||
<taxonomicName id="4C07ADC17C713310B123A378A24EF8D1" box="[292,376,1884,1909]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" sensu="stricto" species="waltoni">waltoni</taxonomicName>
|
||
<taxonomicNameLabel id="A240B72B7C713310B179A379A285F8D0" box="[382,435,1885,1908]" pageId="28" pageNumber="29" sensu="stricto">s. str</taxonomicNameLabel>
|
||
</emphasis>
|
||
.). The latter sequences are all short, perhaps because there are also nucleotide changes in the primer region. When fresh material becomes available from the Himalayan population and a specific primer can be developed, this group needs to be checked in case it represents a separate cryptic species.
|
||
</paragraph>
|
||
<caption id="DF7886CA7C703311B090A377A76FF847" httpUri="https://zenodo.org/record/192314/files/figure.png" pageId="29" pageNumber="30" targetBox="[151,1415,299,1829]" targetPageId="29">
|
||
<paragraph id="8BB8D6427C703311B090A377A76FF847" blockId="29.[151,1436,1875,2019]" pageId="29" pageNumber="30">
|
||
<emphasis id="B9730A507C703311B090A377A278F8CD" bold="true" box="[151,334,1875,1897]" pageId="29" pageNumber="30">FIGURES 36‒55</emphasis>
|
||
. Morphology of parts of the male genitalia from the dorsal aspect of the left side, anterior to the left of the image, posterior to the right, white bars indicating morphological characters referred to in the key to males: 36,
|
||
<taxonomicName id="4C07ADC17C703311B501A356A659F823" box="[1286,1391,1905,1927]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="waltoni">
|
||
<emphasis id="B9730A507C703311B501A356A659F823" box="[1286,1391,1905,1927]" italics="true" pageId="29" pageNumber="30">B. waltoni</emphasis>
|
||
</taxonomicName>
|
||
; 37,
|
||
<taxonomicName id="4C07ADC17C703311B090A3B5A235F802" box="[151,259,1936,1958]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="mendax">
|
||
<emphasis id="B9730A507C703311B090A3B5A235F802" box="[151,259,1936,1958]" italics="true" pageId="29" pageNumber="30">B. mendax</emphasis>
|
||
</taxonomicName>
|
||
; 38,
|
||
<taxonomicName id="4C07ADC17C703311B13FA3B5A286F802" box="[312,432,1936,1958]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="superbus">
|
||
<emphasis id="B9730A507C703311B13FA3B5A286F802" box="[312,432,1936,1958]" italics="true" pageId="29" pageNumber="30">B. superbus</emphasis>
|
||
</taxonomicName>
|
||
; 39,
|
||
<taxonomicName id="4C07ADC17C703311B1E1A3B5A156F802" box="[486,608,1937,1958]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="convexus">
|
||
<emphasis id="B9730A507C703311B1E1A3B5A156F802" box="[486,608,1937,1958]" italics="true" pageId="29" pageNumber="30">B. convexus</emphasis>
|
||
</taxonomicName>
|
||
; 40,
|
||
<taxonomicName id="4C07ADC17C703311B292A3B5A1C8F802" box="[661,766,1936,1958]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="waltoni">
|
||
<emphasis id="B9730A507C703311B292A3B5A1C8F802" box="[661,766,1936,1958]" italics="true" pageId="29" pageNumber="30">B. waltoni</emphasis>
|
||
</taxonomicName>
|
||
; 41,
|
||
<taxonomicName id="4C07ADC17C703311B334A3B5A09BF802" box="[819,941,1937,1958]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="convexus">
|
||
<emphasis id="B9730A507C703311B334A3B5A09BF802" box="[819,941,1937,1958]" italics="true" pageId="29" pageNumber="30">B. convexus</emphasis>
|
||
</taxonomicName>
|
||
; 42,
|
||
<taxonomicName id="4C07ADC17C703311B3E4A3B5A765F802" box="[995,1107,1936,1958]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="defector">
|
||
<emphasis id="B9730A507C703311B3E4A3B5A765F802" box="[995,1107,1936,1958]" italics="true" pageId="29" pageNumber="30">B. defector</emphasis>
|
||
</taxonomicName>
|
||
; 43,
|
||
<taxonomicName id="4C07ADC17C703311B48FA3B5A7C5F802" box="[1160,1267,1936,1958]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="mendax">
|
||
<emphasis id="B9730A507C703311B48FA3B5A7C5F802" box="[1160,1267,1936,1958]" italics="true" pageId="29" pageNumber="30">B. mendax</emphasis>
|
||
</taxonomicName>
|
||
; 44,
|
||
<taxonomicName id="4C07ADC17C703311B52EA3B5A6A3F802" box="[1321,1429,1936,1958]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="mendax">
|
||
<emphasis id="B9730A507C703311B52EA3B5A6A3F802" box="[1321,1429,1936,1958]" italics="true" pageId="29" pageNumber="30">B. mendax</emphasis>
|
||
</taxonomicName>
|
||
; 45,
|
||
<taxonomicName id="4C07ADC17C703311B0C6A394A25EF860" box="[193,360,1966,1989]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="turkestanicus">
|
||
<emphasis id="B9730A507C703311B0C6A394A25EF860" box="[193,360,1966,1989]" italics="true" pageId="29" pageNumber="30">B. turkestanicus</emphasis>
|
||
</taxonomicName>
|
||
; 46,
|
||
<taxonomicName id="4C07ADC17C703311B1A5A394A10BF860" box="[418,573,1966,1989]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="avinoviellus">
|
||
<emphasis id="B9730A507C703311B1A5A394A10BF860" box="[418,573,1966,1989]" italics="true" pageId="29" pageNumber="30">B. avinoviellus</emphasis>
|
||
</taxonomicName>
|
||
; 47,
|
||
<taxonomicName id="4C07ADC17C703311B27FA394A1D0F860" box="[632,742,1966,1989]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="mendax">
|
||
<emphasis id="B9730A507C703311B27FA394A1D0F860" box="[632,742,1966,1989]" italics="true" pageId="29" pageNumber="30">B. mendax</emphasis>
|
||
</taxonomicName>
|
||
; 48,
|
||
<taxonomicName id="4C07ADC17C703311B326A394A0B9F860" box="[801,911,1966,1989]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="mendax">
|
||
<emphasis id="B9730A507C703311B326A394A0B9F860" box="[801,911,1966,1989]" italics="true" pageId="29" pageNumber="30">B. mendax</emphasis>
|
||
</taxonomicName>
|
||
; 49,
|
||
<taxonomicName id="4C07ADC17C703311B3CCA394A778F860" box="[971,1102,1966,1989]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="makarjini">
|
||
<emphasis id="B9730A507C703311B3CCA394A778F860" box="[971,1102,1966,1989]" italics="true" pageId="29" pageNumber="30">B. makarjini</emphasis>
|
||
</taxonomicName>
|
||
; 50,
|
||
<taxonomicName id="4C07ADC17C703311B48FA394A67DF860" box="[1160,1355,1966,1989]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="handlirschianus">
|
||
<emphasis id="B9730A507C703311B48FA394A67DF860" box="[1160,1355,1966,1989]" italics="true" pageId="29" pageNumber="30">B. handlirschianus</emphasis>
|
||
</taxonomicName>
|
||
; 51,
|
||
<taxonomicName id="4C07ADC17C703311B581A394A22EF847" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="himalayanus">
|
||
<emphasis id="B9730A507C703311B581A394A22EF847" italics="true" pageId="29" pageNumber="30">B. himalayanus</emphasis>
|
||
</taxonomicName>
|
||
; 52,
|
||
<taxonomicName id="4C07ADC17C703311B14BA3EAA2D9F847" box="[332,495,1997,2019]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="turkestanicus">
|
||
<emphasis id="B9730A507C703311B14BA3EAA2D9F847" box="[332,495,1997,2019]" italics="true" pageId="29" pageNumber="30">B. turkestanicus</emphasis>
|
||
</taxonomicName>
|
||
; 53,
|
||
<taxonomicName id="4C07ADC17C703311B225A3EAA189F847" box="[546,703,1997,2019]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="himalayanus">
|
||
<emphasis id="B9730A507C703311B225A3EAA189F847" box="[546,703,1997,2019]" italics="true" pageId="29" pageNumber="30">B. himalayanus</emphasis>
|
||
</taxonomicName>
|
||
; 54,
|
||
<taxonomicName id="4C07ADC17C703311B2F4A3EAA0A6F847" box="[755,912,1997,2019]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="himalayanus">
|
||
<emphasis id="B9730A507C703311B2F4A3EAA0A6F847" box="[755,912,1997,2019]" italics="true" pageId="29" pageNumber="30">B. himalayanus</emphasis>
|
||
</taxonomicName>
|
||
; 55,
|
||
<taxonomicName id="4C07ADC17C703311B3C3A3EAA765F847" box="[964,1107,1998,2019]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="marussinus">
|
||
<emphasis id="B9730A507C703311B3C3A3EAA765F847" box="[964,1107,1998,2019]" italics="true" pageId="29" pageNumber="30">B. marussinus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB8D6427C73332CB090A38FA258FE09" blockId="30.[151,468,1963,1985]" lastBlockId="32.[151,1436,151,1112]" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31">
|
||
<figureCitation id="133CCAC77C733312B090A38FA27BF865" box="[151,333,1963,1985]" captionStart="FIGURES 36 ‒ 55" captionStartId="29.[151,264,1875,1897]" captionTargetBox="[151,1415,299,1829]" captionTargetId="figure@29.[151,1436,193,1853]" captionTargetPageId="29" captionText="FIGURES 36 ‒ 55. Morphology of parts of the male genitalia from the dorsal aspect of the left side, anterior to the left of the image, posterior to the right, white bars indicating morphological characters referred to in the key to males: 36, B. waltoni; 37, B. mendax; 38, B. superbus; 39, B. convexus; 40, B. waltoni; 41, B. convexus; 42, B. defector; 43, B. mendax; 44, B. mendax; 45, B. turkestanicus; 46, B. avinoviellus; 47, B. mendax; 48, B. mendax; 49, B. makarjini; 50, B. handlirschianus; 51, B. himalayanus; 52, B. turkestanicus; 53, B. himalayanus; 54, B. himalayanus; 55, B. marussinus." httpUri="https://zenodo.org/record/192314/files/figure.png" pageId="30" pageNumber="31">
|
||
<emphasis id="B9730A507C733312B090A38FA27BF865" bold="true" box="[151,333,1963,1985]" pageId="30" pageNumber="31">FIGURES 36‒55</emphasis>
|
||
</figureCitation>
|
||
. (Continued)
|
||
<figureCitation id="133CCAC77C723313B090A28AA27BF960" box="[151,333,1710,1732]" captionStart="FIGURES 36 ‒ 55" captionStartId="29.[151,264,1875,1897]" captionTargetBox="[151,1415,299,1829]" captionTargetId="figure@29.[151,1436,193,1853]" captionTargetPageId="29" captionText="FIGURES 36 ‒ 55. Morphology of parts of the male genitalia from the dorsal aspect of the left side, anterior to the left of the image, posterior to the right, white bars indicating morphological characters referred to in the key to males: 36, B. waltoni; 37, B. mendax; 38, B. superbus; 39, B. convexus; 40, B. waltoni; 41, B. convexus; 42, B. defector; 43, B. mendax; 44, B. mendax; 45, B. turkestanicus; 46, B. avinoviellus; 47, B. mendax; 48, B. mendax; 49, B. makarjini; 50, B. handlirschianus; 51, B. himalayanus; 52, B. turkestanicus; 53, B. himalayanus; 54, B. himalayanus; 55, B. marussinus." httpUri="https://zenodo.org/record/192314/files/figure.png" pageId="31" pageNumber="32">
|
||
<emphasis id="B9730A507C723313B090A28AA27BF960" bold="true" box="[151,333,1710,1732]" pageId="31" pageNumber="32">FIGURES 36‒55</emphasis>
|
||
</figureCitation>
|
||
. (Continued)
|
||
<emphasis id="B9730A507C723313B0C0A330A2D6F889" bold="true" box="[199,480,1812,1837]" pageId="31" pageNumber="32">Diagnostic description.</emphasis>
|
||
Wings nearly clear. Hair long, uneven and slightly sparse.
|
||
<emphasis id="B9730A507C723313B4B2A330A3C7F8F5" italics="true" pageId="31" pageNumber="32">Female hair colour pattern:</emphasis>
|
||
generally black, but the thoracic dorsum often with many grey-white hairs intermixed with black, on the side of the thorax and on T1 and on T2 anteriorly the black hairs often have white tips (
|
||
<emphasis id="B9730A507C723313B458A379A745F8D1" box="[1119,1139,1885,1909]" italics="true" pageId="31" pageNumber="32">cf</emphasis>
|
||
. all other
|
||
<taxonomicName id="4C07ADC17C723313B4E4A379A6AAF8D2" box="[1251,1436,1885,1910]" class="Insecta" family="Apidae" genus="Mendacibombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9730A507C723313B4E4A379A6AAF8D2" box="[1251,1436,1885,1910]" italics="true" pageId="31" pageNumber="32">Mendacibombus</emphasis>
|
||
</taxonomicName>
|
||
species), on T2 posteriorly and on T3‒6 the predominantly orange hairs often have white tips, T6 with few black hairs. Hindleg tibia with the corbicular fringes with black and orange hairs, many with white tips.
|
||
<emphasis id="B9730A507C723313B54FA381A211F846" italics="true" pageId="31" pageNumber="32">Female morphology:</emphasis>
|
||
labrum with the basal depression narrow, the transverse ridge very broad and high, medially not subsiding or interrupted and in the median third shining with very few scattered large punctures, lateral tubercles almost without punctures. T2 at most (in queens) with only a very subtle posteriorly-directed convexity of its median posterior edge (
|
||
<emphasis id="B9730A507C4D332CB1A6A4C4A17DFF5C" box="[417,587,224,248]" italics="true" pageId="32" pageNumber="33">
|
||
cf.
|
||
<taxonomicName id="4C07ADC17C4D332CB1C3A4C5A17DFF5C" box="[452,587,225,248]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="convexus">B. convexus</taxonomicName>
|
||
</emphasis>
|
||
).
|
||
<emphasis id="B9730A507C4D332CB265A4FBA01CFF5C" box="[610,810,223,248]" italics="true" pageId="32" pageNumber="33">Male morphology</emphasis>
|
||
: beard of the mandible long, dense and black; genitalia (
|
||
<figureCitation id="133CCAC77C4D332CB098A520A3C7FEB9" box="[159,241,260,285]" captionStart="FIGURES 24 ‒ 35" captionStartId="26.[151,264,1867,1889]" captionTargetBox="[233,1387,193,1789]" captionTargetId="figure@26.[233,1390,193,1894]" captionTargetPageId="26" captionText="FIGURES 24 ‒ 35. Morphology of the male genitalia for species of the subgenus Mendacibombus from the dorsal aspect, anterior to the left of the image, posterior to the right: 24, B. superbus, China MD # 339; 25, B. waltoni, China MD # 216; 26, B. convexus, China MD # 215; 27, B. makarjini, Kazakhstan MD # 1240; 28, B. marussinus, Pakistan MD # 408; 29, B. avinoviellus, India MD # 404; 30, B. himalayanus, India MD # 426; 31, B. turkestanicus, China MD # 351; 32, B. defector, Kyrgyzstan MD # 367; 33, B. margreiteri, China MD # 294; 34, B. handlirschianus, Russia MD # 835; 35, B. mendax, Spain MD # 689." httpUri="https://zenodo.org/record/192312/files/figure.png" pageId="32" pageNumber="33">Fig. 25</figureCitation>
|
||
) with the volsella at its broadest near the midpoint of its length, the dorsal surface just distal to this point with a raised curved ridge, often with small teeth, just inside the inner margin, running for 0.5× the remaining distal length of the volsella; volsella distally sharply acute (pointed) and curled back dorsally and anteriorly. Gonostylus length 1.25× its greatest breadth. Penis-valve head length 0.33× the length of the penis valve distal to the broadest point of the spatha.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D85C97C4D332CB0C0A593A766FC6C" pageId="32" pageNumber="33" type="materials_examined">
|
||
<paragraph id="8BB8D6427C4D332CB0C0A593A135FDBC" blockId="32.[151,1436,151,1112]" pageId="32" pageNumber="33">
|
||
<emphasis id="B9730A507C4D332CB0C0A593A285FE74" bold="true" box="[199,435,439,464]" pageId="32" pageNumber="33">Material examined.</emphasis>
|
||
29 queens
|
||
<specimenCount id="9D011DCB7C4D332CB23EA593A1F1FE74" box="[569,711,439,464]" pageId="32" pageNumber="33" type="worker">296 workers</specimenCount>
|
||
<specimenCount id="9D011DCB7C4D332CB2D6A593A00DFE74" box="[721,827,439,464]" pageId="32" pageNumber="33" type="male">56 males</specimenCount>
|
||
, from
|
||
<collectingCountry id="F31096D27C4D332CB38EA593A0F9FE74" box="[905,975,439,464]" name="China" pageId="32" pageNumber="33">China</collectingCountry>
|
||
,
|
||
<collectingCountry id="F31096D27C4D332CB3DAA593A72FFE74" box="[989,1049,439,464]" name="India" pageId="32" pageNumber="33">India</collectingCountry>
|
||
, and
|
||
<collectingCountry id="F31096D27C4D332CB45EA593A7AAFE74" box="[1113,1180,439,464]" name="Nepal" pageId="32" pageNumber="33">Nepal</collectingCountry>
|
||
(
|
||
<figureCitation id="133CCAC77C4D332CB4A9A593A633FE74" box="[1198,1285,439,464]" captionStart="FIGURES 56 ‒ 67" captionStartId="35.[151,264,1891,1913]" captionTargetBox="[320,1245,193,1835]" captionTargetId="figure@35.[320,1245,193,1843]" captionTargetPageId="35" captionText="FIGURES 56 ‒ 67. Maps of the global distributions of species of the subgenus Mendacibombus from the material examined, with grey spots for specimen records and black spots for the samples with COI sequences (each species is unknown from regions outside the map boundaries shown); grey crosses for records of all species of Mendacibombus combined. Relief map with hill shading, Cartesian orthonormal projection, grey lines show borders to national administration according to UN maps, north at the top of the map. Image created in ArcGIS using World _ Shaded _ Relief basemap © 2014 Esri." httpUri="https://zenodo.org/record/192315/files/figure.png" pageId="32" pageNumber="33">Fig. 57</figureCitation>
|
||
:
|
||
<collectionCode id="ED164E877C4D332CB515A59CA673FE74" box="[1298,1349,440,464]" pageId="32" pageNumber="33">IAR</collectionCode>
|
||
,
|
||
<collectionCode id="ED164E877C4D332CB554A59CA6A1FE74" box="[1363,1431,440,464]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34797" name="Illinois Natural History Survey" pageId="32" pageNumber="33">INHS</collectionCode>
|
||
,
|
||
<collectionCode id="ED164E877C4D332CB090A5F9A3F1FE50" box="[151,199,477,500]" pageId="32" pageNumber="33">IZB</collectionCode>
|
||
,
|
||
<collectionCode id="ED164E877C4D332CB0D4A5F9A21DFE50" box="[211,299,477,500]" pageId="32" pageNumber="33">MNHU</collectionCode>
|
||
,
|
||
<collectionCode id="ED164E877C4D332CB13FA5F8A25AFE50" box="[312,364,476,500]" country="Philippines" httpUri="http://grbio.org/cool/g413-dnr0" name="Marine Science Institute, University of the Philippines" pageId="32" pageNumber="33">MSI</collectionCode>
|
||
,
|
||
<collectionCode id="ED164E877C4D332CB17EA5F9A28BFE50" box="[377,445,477,500]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="32" pageNumber="33">NHM</collectionCode>
|
||
,
|
||
<collectionCode id="ED164E877C4D332CB1CDA5F9A13DFE50" box="[458,523,477,500]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="32" pageNumber="33">NME</collectionCode>
|
||
,
|
||
<collectionCode id="ED164E877C4D332CB210A5F8A166FE50" box="[535,592,476,500]" pageId="32" pageNumber="33">OLL</collectionCode>
|
||
, PW,
|
||
<collectionCode id="ED164E877C4D332CB293A5F9A1DDFE50" box="[660,747,477,500]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="32" pageNumber="33">RMNH</collectionCode>
|
||
, SC, YT,
|
||
<collectionCode id="ED164E877C4D332CB35CA5F8A0AFFE50" box="[859,921,476,500]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="32" pageNumber="33">ZISP</collectionCode>
|
||
), with
|
||
<specimenCount id="9D011DCB7C4D332CB3E0A5F8A7B6FE51" box="[999,1152,476,501]" pageId="32" pageNumber="33" type="generic">14 specimens</specimenCount>
|
||
sequenced (interpretable sequences listed in
|
||
<figureCitation id="133CCAC77C4D332CB177A5DBA2C0FDBC" box="[368,502,511,536]" captionStart-0="FIGURES 11 ‒ 12" captionStart-1="FIGURE 13" captionStartId-0="15.[151,264,1630,1652]" captionStartId-1="16.[151,250,1733,1755]" captionTargetBox-0="[300,1121,202,1610]" captionTargetBox-1="[314,1260,194,1721]" captionTargetId-0="figure@15.[292,1266,193,1625]" captionTargetId-1="figure@16.[314,1261,193,1722]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="FIGURES 11 ‒ 12. Recognising species from an estimate of phylogeny for 30 unique COI haplotypes of the subgenus Mendacibombus by BEAST, using GMYC models and the PTP procedure. From the GMYC models (11), the single threshold T is shown by the vertical grey bar so that the intersecting lineages are interpreted as indicating 12 separate species (black spots at coalescent nodes; outgroups not shown). Values below the nodes of the maximum clade-credibility tree are Bayesian posterior probabilities showing branch support (groups with values <0.8 are considered unreliable) from the BEAST analysis with two billion MCMC generations and a 1 % burn-in. The x - scale axis is calibrated in substitutions per nucleotide site. From the PTP procedure (12), the Bayesian PTP solution with highest support also indicates 12 species. Values above the nodes are PTP Bayesian support values that all daughter haplotypes are parts of a single species; values below the nodes are Bayesian posterior probabilities showing branch support. The scale bar is calibrated in substitutions per nucleotide site. Each unique haplotype is represented by one of the longest available sample sequences, labelled with: the sequence length in number of base pairs, a taxon name or name for an unpublished colour pattern (light / dark / yellow / white / pale), and then a code that consists of a specimen identifier (MD #) from the project database and (after the hyphen) a sample identifier from BOLD or from GenBank, followed with its geographic origin." captionText-1="FIGURE 13. Assigning names to species from an estimate of phylogeny for 130 COI sequences of the subgenus Mendacibombus. Based on a BEAST analysis of the 30 unique haplotypes (Fig. 11), with the remaining sequences interpolated back into the tree with 0 branch lengths from their matching samples (where a sequence matches more than one longer sequence within a species in the Collapse results, it is added after the first matching sequence, with the addition of the two short-sequence yellow unbanded specimens from Pakistan, MD # 4058, 4059). The sequence labels follow the format used in Figs 11, 12. Lineages accepted as species from the GMYC models (Fig. 11) are shown as thick branches, the most recent common ancestors of each species are shown with a black spot, and branches within species are shown as thin branches. Asterisks mark samples used as informal proxies for the type specimens of each of the taxon names (Table 1). The proxy sample for the type specimen for the oldest available name (the valid name) for each species (from Fig. 11) is marked in bold. The interpretations of the species with their valid names are shown in grey to the right." httpUri-0="https://zenodo.org/record/192308/files/figure.png" httpUri-1="https://zenodo.org/record/192309/files/figure.png" pageId="32" pageNumber="33">Figs. 11–13</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB8D6427C4D332CB0C0A600A766FC6C" blockId="32.[151,1436,151,1112]" pageId="32" pageNumber="33">
|
||
<emphasis id="B9730A507C4D332CB0C0A600A2C4FD99" bold="true" box="[199,498,548,573]" pageId="32" pageNumber="33">Habitat and distribution.</emphasis>
|
||
Flower-rich alpine grassland, at elevations 2604‒(3927)‒
|
||
<quantity id="4CFF7BA77C4D332CB472A600A7FDFD98" box="[1141,1227,548,573]" metricMagnitude="3" metricUnit="m" metricValue="5.22" pageId="32" pageNumber="33" unit="m" value="5220.0">5220 m</quantity>
|
||
a.s.l.. A species of the east Qinghai-Tibetan plateau, in the east Himalayan, Hengduan, and Qinghai-Gansu mountains, including the Tanggula Shan, Kunlun Shan, Burhan Budai Shan,
|
||
<collectingRegion id="49C318A07C4D332CB2D4A648A01BFD21" box="[723,813,620,645]" country="China" name="Qinghai" pageId="32" pageNumber="33">Qinghai</collectingRegion>
|
||
Nan Shan, Qiliang Shan, and Min Shan, but much rarer in the western Himalaya. The lack of records in the northwest Tibetan plateau (
|
||
<figureCitation id="133CCAC77C4D332CB40FA6ABA76FFD0C" box="[1032,1113,655,680]" captionStart="FIGURES 56 ‒ 67" captionStartId="35.[151,264,1891,1913]" captionTargetBox="[320,1245,193,1835]" captionTargetId="figure@35.[320,1245,193,1843]" captionTargetPageId="35" captionText="FIGURES 56 ‒ 67. Maps of the global distributions of species of the subgenus Mendacibombus from the material examined, with grey spots for specimen records and black spots for the samples with COI sequences (each species is unknown from regions outside the map boundaries shown); grey crosses for records of all species of Mendacibombus combined. Relief map with hill shading, Cartesian orthonormal projection, grey lines show borders to national administration according to UN maps, north at the top of the map. Image created in ArcGIS using World _ Shaded _ Relief basemap © 2014 Esri." httpUri="https://zenodo.org/record/192315/files/figure.png" pageId="32" pageNumber="33">Fig. 57</figureCitation>
|
||
) may at least in part reflect a lack of sampling (see the comments on
|
||
<taxonomicName id="4C07ADC17C4D332CB25EA691A1E8FD69" box="[601,734,692,717]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="superbus">
|
||
<emphasis id="B9730A507C4D332CB25EA691A1E8FD69" box="[601,734,692,717]" italics="true" pageId="32" pageNumber="33">B. superbus</emphasis>
|
||
</taxonomicName>
|
||
) or may reflect a true absence (P.H.
|
||
<bibRefCitation id="EF96ABB37C4D332CB47AA690A3C0FD54" author="Williams" pageId="32" pageNumber="33" refString="Williams, P. H., Bystriakova, N., Huang, J. - X., Miao, Z. - Y. & An, J. - D. (2015) Bumblebees, climate and glaciers across the Tibetan plateau (Apidae: Bombus Latreille). Systematics and Biodiversity, 13, 164 - 181." type="journal article" year="2015">
|
||
Williams, Bystriakova,
|
||
<emphasis id="B9730A507C4D332CB580A691A382FD54" italics="true" pageId="32" pageNumber="33">et al.</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
) from a semi-arid region (the Qiangtang plateau, although modelling climate suitability shows that areas near the centre near Siling lake might be suitable). Compared to
|
||
<taxonomicName id="4C07ADC17C4D332CB393A6D9A728FCB0" box="[916,1054,765,788]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="convexus">
|
||
<emphasis id="B9730A507C4D332CB393A6D9A728FCB0" box="[916,1054,765,788]" italics="true" pageId="32" pageNumber="33">B. convexus</emphasis>
|
||
</taxonomicName>
|
||
, distributions of the two species overlap broadly, but
|
||
<taxonomicName id="4C07ADC17C4D332CB17BA705A2D9FC9C" box="[380,495,799,824]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="waltoni">
|
||
<emphasis id="B9730A507C4D332CB17BA705A2D9FC9C" box="[380,495,799,824]" italics="true" pageId="32" pageNumber="33">B. waltoni</emphasis>
|
||
</taxonomicName>
|
||
extends further to the north and west and tends to occur at higher elevation (and it is uncommon for the two species to occur together at a site).
|
||
<taxonomicName id="4C07ADC17C4D332CB32EA760A0E9FCF9" box="[809,991,836,861]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="waltoni">
|
||
<emphasis id="B9730A507C4D332CB32EA760A0E9FCF9" box="[809,991,836,861]" italics="true" pageId="32" pageNumber="33">Bombus waltoni</emphasis>
|
||
</taxonomicName>
|
||
replaces the western
|
||
<taxonomicName id="4C07ADC17C4D332CB4D7A761A648FCF9" box="[1232,1406,836,861]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="himalayanus">
|
||
<emphasis id="B9730A507C4D332CB4D7A761A648FCF9" box="[1232,1406,836,861]" italics="true" pageId="32" pageNumber="33">B. himalayanus</emphasis>
|
||
</taxonomicName>
|
||
in the wetter meadows of the higher alpine zone of the eastern Himalaya, where it overlaps with
|
||
<taxonomicName id="4C07ADC17C4D332CB4CAA74DA665FC24" box="[1229,1363,871,896]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="superbus">
|
||
<emphasis id="B9730A507C4D332CB4CAA74DA665FC24" box="[1229,1363,871,896]" italics="true" pageId="32" pageNumber="33">B. superbus</emphasis>
|
||
</taxonomicName>
|
||
in the Hohxil and the Tanggula mountains. Regional distribution maps are available for
|
||
<collectingRegion id="49C318A07C4D332CB44CA7A8A793FC01" box="[1099,1189,908,933]" country="China" name="Sichuan" pageId="32" pageNumber="33">Sichuan</collectingRegion>
|
||
(P.H.
|
||
<bibRefCitation id="EF96ABB37C4D332CB4F6A7A8A3E5FC6C" author="Williams" pageId="32" pageNumber="33" refString="Williams, P. H., Tang, Y., Yao, J. & Cameron, S. (2009) The bumblebees of Sichuan (Hymenoptera: Apidae, Bombini). Systematics and Biodiversity, 7, 101 - 190." type="journal article" year="2009">
|
||
Williams
|
||
<emphasis id="B9730A507C4D332CB566A7A9A6AAFC01" box="[1377,1436,908,933]" italics="true" pageId="32" pageNumber="33">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
),
|
||
<collectingRegion id="49C318A07C4D332CB0E1A794A218FC6C" box="[230,302,944,968]" country="China" name="Gansu" pageId="32" pageNumber="33">Gansu</collectingRegion>
|
||
(An
|
||
<emphasis id="B9730A507C4D332CB160A795A2A9FC6C" box="[359,415,943,968]" italics="true" pageId="32" pageNumber="33">et al.</emphasis>
|
||
2011;
|
||
<bibRefCitation id="EF96ABB37C4D332CB1EBA794A1A6FC6C" author="An" box="[492,656,943,968]" pageId="32" pageNumber="33" refString="An, J. - D., Huang, J. - X., Shao, Y. - Q., Zhang, S. - W., Wang, B., Liu, X. - Y., Wu, J. & Williams, P. H. (2014) The bumblebees of North China (Apidae, Bombus Latreille). Zootaxa, 3830 (1), 1 - 89. http: // dx. doi. org / 10.11646 / zootaxa. 3830.1.1" type="journal article" year="2014">
|
||
An
|
||
<emphasis id="B9730A507C4D332CB212A795A17BFC6C" box="[533,589,943,968]" italics="true" pageId="32" pageNumber="33">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
), and
|
||
<collectingCountry id="F31096D27C4D332CB2D4A78BA020FC6C" box="[723,790,943,968]" name="Nepal" pageId="32" pageNumber="33">Nepal</collectingCountry>
|
||
(P.H. Williams
|
||
<emphasis id="B9730A507C4D332CB3CFA795A737FC6C" box="[968,1025,943,968]" italics="true" pageId="32" pageNumber="33">et al.</emphasis>
|
||
2010).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D85C97C4D332CB0C0A7F0A1E0FBFC" pageId="32" pageNumber="33" type="description">
|
||
<paragraph id="8BB8D6427C4D332CB0C0A7F0A050FC48" blockId="32.[151,1436,151,1112]" box="[199,870,980,1005]" pageId="32" pageNumber="33">
|
||
<emphasis id="B9730A507C4D332CB0C0A7F0A26DFC49" bold="true" box="[199,347,980,1005]" pageId="32" pageNumber="33">Food plants.</emphasis>
|
||
<bibRefCitation id="EF96ABB37C4D332CB165A7F0A16CFC48" box="[354,602,980,1005]" pageId="32" pageNumber="33" refString="Williams, P. H., Tang, Y., Yao, J. & Cameron, S. (2009) The bumblebees of Sichuan (Hymenoptera: Apidae, Bombini). Systematics and Biodiversity, 7, 101 - 190." type="journal article">
|
||
Williams
|
||
<emphasis id="B9730A507C4D332CB1C9A7F1A2C9FC49" box="[462,511,980,1005]" italics="true" pageId="32" pageNumber="33">et al</emphasis>
|
||
. (2009)
|
||
</bibRefCitation>
|
||
, An
|
||
<emphasis id="B9730A507C4D332CB288A7F1A1F6FC49" box="[655,704,980,1005]" italics="true" pageId="32" pageNumber="33">et al</emphasis>
|
||
. (2011; 2014).
|
||
</paragraph>
|
||
<paragraph id="8BB8D6427C4D332CB0C0A7D3A1E0FBFC" blockId="32.[151,1436,151,1112]" pageId="32" pageNumber="33">
|
||
<materialsCitation id="3B6FDC1F7C4D332CB0C0A7D3A1E0FBFC" ID-GBIF-Occurrence="1413110437" collectingDate="2002-08-05" collectingDateMax="2009-08-29" collectingDateMin="2002-08-05" country="China" location="Diebu" pageId="32" pageNumber="33" specimenCount="1" stateProvince="Sichuan">
|
||
<emphasis id="B9730A507C4D332CB0C0A7D3A27EFBB4" bold="true" box="[199,328,1015,1040]" pageId="32" pageNumber="33">Behaviour.</emphasis>
|
||
Mate-searching males perch on bare patches of ground or low bushes and pursue other bees that fly past before the males return to the same perch (PW:
|
||
<figureCitation id="133CCAC77C4D332CB317A038A058FB91" box="[784,878,1052,1077]" captionStart="FIGURES 1 ‒ 6" captionStartId="5.[151,264,1819,1841]" captionTargetBox="[168,1389,193,1768]" captionTargetId="figure@5.[168,1412,193,1784]" captionTargetPageId="5" captionText="FIGURES 1 ‒ 6. Views of perching males of species of the subgenus Mendacibombus, showing the anteriorly extended antennae and laterally extended wings that are characteristic of their form of male mate-searching behaviour: 1, B. waltoni at Diebu ca 3910 m, Minshan mountains, Gansu, China, 29. viii. 2009; 2, B. convexus at Diebu ca 3340 m, Minshan mountains, Gansu, China, 29. viii. 2009; 3, B. avinoviellus at Gulmarg ca 2700 m, Pir Panjal mountains, Kashmir, India, 16. viii. 1985; 4, B. himalayanus at Apharwat ca 4100 m, Pir Panjal mountains, Kashmir, India, 30. viii. 1986; 5 ‒ 6, B. margreiteri near Dulan ca 3520 m, Burhan Budai mountains, Qinghai, China, 31. viii. 2010. Photos by PW (Figs 3 – 4 are the best available for these species)." httpUri="https://zenodo.org/record/192303/files/figure.png" pageId="32" pageNumber="33">
|
||
Fig. 1,
|
||
<collectingDate id="EFFD096A7C4D332CB366A038A058FB91" box="[865,878,1052,1077]" pageId="32" pageNumber="33" value="2002-08-05">
|
||
<date id="FFB9F0827C4D332CB366A038A058FB91" box="[865,878,1052,1077]" pageId="32" pageNumber="33" value="2002-08-05">5</date>
|
||
</collectingDate>
|
||
</figureCitation>
|
||
.viii.
|
||
<quantity id="4CFF7BA77C4D332CB399A038A708FB90" box="[926,1086,1052,1077]" metricMagnitude="3" metricUnit="m" metricValue="2.0023764" pageId="32" pageNumber="33" unit="m" value="2002.3764">2002, 3764 m</quantity>
|
||
Aba-Hongyuan road,
|
||
<collectingRegion id="49C318A07C4D332CB53CA038A6AEFB91" box="[1339,1432,1052,1077]" country="China" name="Sichuan" pageId="32" pageNumber="33">Sichuan</collectingRegion>
|
||
,
|
||
<collectingCountry id="F31096D27C4D332CB090A01BA3E8FBFC" box="[151,222,1087,1112]" name="China" pageId="32" pageNumber="33">China</collectingCountry>
|
||
;
|
||
<date id="FFB9F0827C4D332CB0EFA01BA244FBFC" box="[232,370,1087,1112]" pageId="32" pageNumber="33" value="2009-08-29">
|
||
<collectingDate id="EFFD096A7C4D332CB0EFA01BA244FBFC" box="[232,370,1087,1112]" pageId="32" pageNumber="33" value="2009-08-29">
|
||
29.viii.
|
||
<quantity id="4CFF7BA77C4D332CB132A01BA244FBFC" box="[309,370,1087,1112]" metricMagnitude="3" metricUnit="m" metricValue="2.0093914" pageId="32" pageNumber="33" unit="m" value="2009.3914">2009</quantity>
|
||
</collectingDate>
|
||
</date>
|
||
, 3914 m
|
||
<location id="8ED880997C4D332CB1DDA01BA117FBFC" LSID="urn:lsid:plazi:treatment:03AE67547C76332CB090A079A1E0FBFC:8ED880997C4D332CB1DDA01BA117FBFC" box="[474,545,1087,1112]" country="China" name="Diebu" pageId="32" pageNumber="33" stateProvince="Sichuan">Diebu</location>
|
||
,
|
||
<collectingRegion id="49C318A07C4D332CB22BA064A141FBFC" box="[556,631,1088,1112]" country="China" name="Gansu" pageId="32" pageNumber="33">Gansu</collectingRegion>
|
||
,
|
||
<collectingCountry id="F31096D27C4D332CB285A01BA1FCFBFC" box="[642,714,1087,1112]" name="China" pageId="32" pageNumber="33">China</collectingCountry>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |