435 lines
54 KiB
XML
435 lines
54 KiB
XML
<document id="18F9CD692448796416199E253F089D10" ID="10.11646/zootaxa.4204.1.1" ID-DOI="10.11646/zootaxa.4204.1.1" ID-GBIF-Dataset="3f8866d2-529e-43ad-b971-29fc52a13858" ID-ISSN="1175-5326" ID-Zenodo-Dep="192302" ID-ZooBank="C050058A-774D-49C0-93F9-7A055B51C2A0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1480667017343" checkinUser="plazi" docAuthor="Williams, Paul H., Huang, Jiaxing, Rasmont, Pierre & An, Jiandong" docDate="2016" docId="03AE67547C4D332DB090A088A7ADFB04" docLanguage="en" docName="zootaxa.4204.1.1.pdf" docOrigin="Zootaxa 4204 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Bombus convexus Wang" docType="treatment" docVersion="9" lastPageNumber="34" masterDocId="FF971F2C7C6D330CB007A424A336FFA4" masterDocTitle="Early-diverging bumblebees from across the roof of the world: the high-mountain subgenus Mendacibombus revised from species’ gene coalescents and morphology (Hymenoptera, Apidae)" masterLastPageNumber="72" masterPageNumber="1" pageNumber="33" updateTime="1698395689147" updateUser="plazi">
|
||
<mods:mods id="54972E8203F6F2BB903C3E377BEC8976" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="09D6A94652393EF97E31967EC502E7F5">
|
||
<mods:title id="A5A11A8CAB36460A3B1F175EF6FADDC2">Early-diverging bumblebees from across the roof of the world: the high-mountain subgenus Mendacibombus revised from species’ gene coalescents and morphology (Hymenoptera, Apidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="A137D9C5550F28D56311B88E58571386" type="personal">
|
||
<mods:role id="FD8DACC21ACC957D9952F2B0AF0C10F3">
|
||
<mods:roleTerm id="07DD7336E58EE5295C5934873A5ADCA9">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="A299DDA9A2DF4B695C4370890269668B">Williams, Paul H.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="BB3D87CB56179D78C9D467DCF23C24E8" type="personal">
|
||
<mods:role id="6AC41A5C2A39FDCEFD6F28275BED370F">
|
||
<mods:roleTerm id="7BA5AD1BF8BC06D5A692C9ACDA5A5243">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B5AA7D700116E5C26DBFDB98FEB2B820">Huang, Jiaxing</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="5D0232B47C1034BD93D41480DB7E77DD" type="personal">
|
||
<mods:role id="C76855674D2C4A04A347CD764A2A4603">
|
||
<mods:roleTerm id="C73052A353A71F9AB8CD863ABB15488D">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="F19E619EC3E5EDB0424C315A7BB24F7C">Rasmont, Pierre</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="72E1E50F81490FEA54364F58A1551457" type="personal">
|
||
<mods:role id="377E6EC4A7CCD415DEA2F02232914522">
|
||
<mods:roleTerm id="B01D5FC953D95F081CBA8A74BC6051C8">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="59F53BD53E32EAB2926338F03B722D03">An, Jiandong</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="834F2AF1F0F53E9BCD2E550425517431">text</mods:typeOfResource>
|
||
<mods:relatedItem id="78E0D09332016A56E6F297EE684CBBB9" type="host">
|
||
<mods:titleInfo id="65E8938BBE4477A879A7E2686C430FD1">
|
||
<mods:title id="B65EAE8C98C800654D588BDF79C813E1">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="B799E52CFF7A52478A58A23370488CD2">
|
||
<mods:date id="8932859F8EFC113EA2F1D3685F0C8CF0">2016</mods:date>
|
||
<mods:detail id="2D1D73993E1116FE7953D2BE685AEA40" type="volume">
|
||
<mods:number id="AB335502A064A4CA7F71EB2551082273">4204</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="C99C9D7A4FB6DAC0B00F18FB423B51CE" type="issue">
|
||
<mods:number id="D0E021D571C5CE29493875818E306CAF">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="2F9811061068868B21CEF68DBC487D8E" unit="page">
|
||
<mods:start id="96F1AC2E60BA9F874D00854BBBAA43A4">1</mods:start>
|
||
<mods:end id="567EDBB6254E4BB57FCD3725DF453088">72</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="5316CBB40EC3FB1655DA041C30F46D7B">journal article</mods:classification>
|
||
<mods:identifier id="F1BA0DFE5D3CF5CD2450AE27BD194333" type="DOI">10.11646/zootaxa.4204.1.1</mods:identifier>
|
||
<mods:identifier id="B4B35CB30C85D2288280AD73B2F6F94F" type="GBIF-Dataset">3f8866d2-529e-43ad-b971-29fc52a13858</mods:identifier>
|
||
<mods:identifier id="166CFC0983F3CB34117EDD85998104D6" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="62BEE6BE1A44A36F54FE57A06396275F" type="Zenodo-Dep">192302</mods:identifier>
|
||
<mods:identifier id="ECA76476BD10ED44A1135EF3E1B3F2CA" type="ZooBank">C050058A-774D-49C0-93F9-7A055B51C2A0</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03AE67547C4D332DB090A088A7ADFB04" ID-DOI="http://doi.org/10.5281/zenodo.5625289" ID-GBIF-Taxon="124940395" ID-Zenodo-Dep="5625289" LSID="urn:lsid:plazi:treatment:03AE67547C4D332DB090A088A7ADFB04" httpUri="http://treatment.plazi.org/id/03AE67547C4D332DB090A088A7ADFB04" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
|
||
<subSubSection id="C31D85C97C4D332CB090A088A2E8FB62" box="[151,478,1196,1222]" pageId="32" pageNumber="33" type="nomenclature">
|
||
<paragraph id="8BB8D6427C4D332CB090A088A2E8FB62" blockId="32.[151,478,1196,1255]" box="[151,478,1196,1222]" pageId="32" pageNumber="33">
|
||
<heading id="D0F0612E7C4D332CB090A088A2E8FB62" bold="true" box="[151,478,1196,1222]" fontSize="11" level="1" pageId="32" pageNumber="33" reason="1">
|
||
<emphasis id="B9730A507C4D332CB090A088A2E8FB62" bold="true" box="[151,478,1196,1222]" pageId="32" pageNumber="33">
|
||
3.
|
||
<taxonomicName id="4C07ADC17C4D332CB0B3A088A2E8FB62" ID-CoL="MFGK" authority="Wang" authorityName="Wang" box="[180,478,1196,1222]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="convexus">
|
||
<emphasis id="B9730A507C4D332CB0B3A088A2B9FB62" bold="true" box="[180,399,1196,1222]" italics="true" pageId="32" pageNumber="33">Bombus convexus</emphasis>
|
||
Wang
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D85C97C4D332CB090A0EAA2A1FB43" box="[151,407,1230,1255]" pageId="32" pageNumber="33" type="description">
|
||
<paragraph id="8BB8D6427C4D332CB090A0EAA2A1FB43" blockId="32.[151,478,1196,1255]" box="[151,407,1230,1255]" pageId="32" pageNumber="33">
|
||
(
|
||
<figureCitation id="133CCAC77C4D332CB098A0EAA3D1FB43" box="[159,231,1230,1255]" captionStart="FIGURES 1 ‒ 6" captionStartId="5.[151,264,1819,1841]" captionTargetBox="[168,1389,193,1768]" captionTargetId="figure@5.[168,1412,193,1784]" captionTargetPageId="5" captionText="FIGURES 1 ‒ 6. Views of perching males of species of the subgenus Mendacibombus, showing the anteriorly extended antennae and laterally extended wings that are characteristic of their form of male mate-searching behaviour: 1, B. waltoni at Diebu ca 3910 m, Minshan mountains, Gansu, China, 29. viii. 2009; 2, B. convexus at Diebu ca 3340 m, Minshan mountains, Gansu, China, 29. viii. 2009; 3, B. avinoviellus at Gulmarg ca 2700 m, Pir Panjal mountains, Kashmir, India, 16. viii. 1985; 4, B. himalayanus at Apharwat ca 4100 m, Pir Panjal mountains, Kashmir, India, 30. viii. 1986; 5 ‒ 6, B. margreiteri near Dulan ca 3520 m, Burhan Budai mountains, Qinghai, China, 31. viii. 2010. Photos by PW (Figs 3 – 4 are the best available for these species)." httpUri="https://zenodo.org/record/192303/files/figure.png" pageId="32" pageNumber="33">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="133CCAC77C4D332CB0F4A0EAA224FB43" box="[243,274,1230,1255]" captionStart="FIGURES 24 ‒ 35" captionStartId="26.[151,264,1867,1889]" captionTargetBox="[233,1387,193,1789]" captionTargetId="figure@26.[233,1390,193,1894]" captionTargetPageId="26" captionText="FIGURES 24 ‒ 35. Morphology of the male genitalia for species of the subgenus Mendacibombus from the dorsal aspect, anterior to the left of the image, posterior to the right: 24, B. superbus, China MD # 339; 25, B. waltoni, China MD # 216; 26, B. convexus, China MD # 215; 27, B. makarjini, Kazakhstan MD # 1240; 28, B. marussinus, Pakistan MD # 408; 29, B. avinoviellus, India MD # 404; 30, B. himalayanus, India MD # 426; 31, B. turkestanicus, China MD # 351; 32, B. defector, Kyrgyzstan MD # 367; 33, B. margreiteri, China MD # 294; 34, B. handlirschianus, Russia MD # 835; 35, B. mendax, Spain MD # 689." httpUri="https://zenodo.org/record/192312/files/figure.png" pageId="32" pageNumber="33">26</figureCitation>
|
||
,
|
||
<figureCitation id="133CCAC77C4D332CB11AA0EBA253FB43" box="[285,357,1231,1255]" captionStart="FIGURES 36 ‒ 55" captionStartId="29.[151,264,1875,1897]" captionTargetBox="[151,1415,299,1829]" captionTargetId="figure@29.[151,1436,193,1853]" captionTargetPageId="29" captionText="FIGURES 36 ‒ 55. Morphology of parts of the male genitalia from the dorsal aspect of the left side, anterior to the left of the image, posterior to the right, white bars indicating morphological characters referred to in the key to males: 36, B. waltoni; 37, B. mendax; 38, B. superbus; 39, B. convexus; 40, B. waltoni; 41, B. convexus; 42, B. defector; 43, B. mendax; 44, B. mendax; 45, B. turkestanicus; 46, B. avinoviellus; 47, B. mendax; 48, B. mendax; 49, B. makarjini; 50, B. handlirschianus; 51, B. himalayanus; 52, B. turkestanicus; 53, B. himalayanus; 54, B. himalayanus; 55, B. marussinus." httpUri="https://zenodo.org/record/192314/files/figure.png" pageId="32" pageNumber="33">39, 41</figureCitation>
|
||
,
|
||
<figureCitation id="133CCAC77C4D332CB176A0EAA2B8FB43" box="[369,398,1230,1255]" captionStart="FIGURES 56 ‒ 67" captionStartId="35.[151,264,1891,1913]" captionTargetBox="[320,1245,193,1835]" captionTargetId="figure@35.[320,1245,193,1843]" captionTargetPageId="35" captionText="FIGURES 56 ‒ 67. Maps of the global distributions of species of the subgenus Mendacibombus from the material examined, with grey spots for specimen records and black spots for the samples with COI sequences (each species is unknown from regions outside the map boundaries shown); grey crosses for records of all species of Mendacibombus combined. Relief map with hill shading, Cartesian orthonormal projection, grey lines show borders to national administration according to UN maps, north at the top of the map. Image created in ArcGIS using World _ Shaded _ Relief basemap © 2014 Esri." httpUri="https://zenodo.org/record/192315/files/figure.png" pageId="32" pageNumber="33">58</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D85C97C4D332CB090A133A275F943" pageId="32" pageNumber="33" type="reference_group">
|
||
<paragraph id="8BB8D6427C4D332CB090A133A094FACE" blockId="32.[151,1437,1302,1631]" pageId="32" pageNumber="33">
|
||
<materialsCitation id="3B6FDC1F7C4D332CB090A133A094FACE" ID-GBIF-Occurrence="1413110428" collectionCode="ZISP" country="China" county="Note" location="Cyrillic" municipality="Williams" pageId="32" pageNumber="33" specimenCount="1" stateProvince="Gansu" typeStatus="holotype">
|
||
<
|
||
<taxonomicName id="4C07ADC17C4D332CB0A2A132A3C0FA88" ID-CoL="62H8K" box="[165,246,1302,1324]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9730A507C4D332CB0A2A132A3C0FA88" box="[165,246,1302,1324]" italics="true" pageId="32" pageNumber="33">Bombus</emphasis>
|
||
</taxonomicName>
|
||
>
|
||
<taxonomicName id="4C07ADC17C4D332CB10BA132A104FA88" ID-CoL="4PK2K" authority="Morawitz 1880:339" authorityName="Morawitz" authorityPageNumber="339" authorityYear="1880" box="[268,562,1302,1324]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="lugubris">
|
||
<emphasis id="B9730A507C4D332CB10BA132A257FA88" box="[268,353,1302,1324]" italics="true" pageId="32" pageNumber="33">lugubris</emphasis>
|
||
<bibRefCitation id="EF96ABB37C4D332CB16FA132A131FA88" author="Morawitz" box="[360,519,1302,1324]" pageId="32" pageNumber="33" refString="Morawitz, F. F. (1880) Ein Beitrag zur Bienen-Fauna mittel-Asiens. Izvestiya Imperatorskoi akademii nauk, 26, 337 - 379." type="journal article" year="1880">Morawitz 1880</bibRefCitation>
|
||
:339
|
||
</taxonomicName>
|
||
(not of
|
||
<bibRefCitation id="EF96ABB37C4D332CB281A132A063FA88" author="Kriechbaumer" box="[646,853,1302,1324]" pageId="32" pageNumber="33" refString="Kriechbaumer, J. (1870) Vier neue Hummelarten. Verhandlungen der Zoologisch - botanischen Gesellschaft in Wien, 20, 157 - 160." type="journal article" year="1870">Kriechbaumer 1870</bibRefCitation>
|
||
:159, =
|
||
<taxonomicName id="4C07ADC17C4D332CB3A5A13CA7B3FA88" ID-CoL="68NQ6" authority="Kirby" authorityName="Kirby" box="[930,1157,1302,1325]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" sensu="lato" species="barbutellus">
|
||
<emphasis id="B9730A507C4D332CB3A5A13CA704FA88" box="[930,1074,1302,1325]" italics="true" pageId="32" pageNumber="33">B. barbutellus</emphasis>
|
||
(Kirby)
|
||
</taxonomicName>
|
||
<emphasis id="B9730A507C4D332CB48AA13CA784FA88" box="[1165,1202,1302,1325]" italics="true" pageId="32" pageNumber="33">
|
||
<taxonomicNameLabel id="A240B72B7C4D332CB48AA13CA784FA88" box="[1165,1202,1302,1325]" pageId="32" pageNumber="33" sensu="lato">s. l.</taxonomicNameLabel>
|
||
</emphasis>
|
||
), type-locality citation ‘Gan-su’.
|
||
<typeStatus id="54BC68E07C4D332CB136A111A2B9FAEF" box="[305,399,1333,1355]" pageId="32" pageNumber="33" type="holotype">Holotype</typeStatus>
|
||
worker by monotypy
|
||
<collectionCode id="ED164E877C4D332CB27BA111A186FAEF" box="[636,688,1333,1355]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="32" pageNumber="33">ZISP</collectionCode>
|
||
examined, (
|
||
<location id="8ED880997C4D332CB334A111A0B5FAEF" LSID="urn:lsid:plazi:treatment:03AE67547C4D332DB090A088A7ADFB04:8ED880997C4D332CB334A111A0B5FAEF" box="[819,899,1333,1355]" country="China" county="Note" municipality="Williams" name="Cyrillic" pageId="32" pageNumber="33" stateProvince="Gansu">Cyrillic</location>
|
||
) ‘[
|
||
<collectingRegion id="49C318A07C4D332CB398A111A0DAFAEF" box="[927,1004,1333,1355]" country="China" name="Gansu" pageId="32" pageNumber="33">Gansu</collectingRegion>
|
||
]’ (but probably actually
|
||
<collectingRegion id="49C318A07C4D332CB4E8A111A675FAEF" box="[1263,1347,1333,1355]" country="China" name="Qinghai" pageId="32" pageNumber="33">Qinghai</collectingRegion>
|
||
,
|
||
<collectingCountry id="F31096D27C4D332CB556A111A6A4FAEF" box="[1361,1426,1333,1355]" name="China" pageId="32" pageNumber="33">China</collectingCountry>
|
||
).
|
||
<location id="8ED880997C4D332CB0C0A170A267FACE" LSID="urn:lsid:plazi:treatment:03AE67547C4D332DB090A088A7ADFB04:8ED880997C4D332CB0C0A170A267FACE" box="[199,337,1364,1386]" country="China" county="Note" municipality="Williams" name="Synonymised" pageId="32" pageNumber="33" stateProvince="Gansu">Synonymised</location>
|
||
with
|
||
<taxonomicName id="4C07ADC17C4D332CB18DA170A1B6FACE" ID-CoL="MFGK" authority="Wang" authorityName="Wang" box="[394,640,1364,1386]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="convexus">
|
||
<emphasis id="B9730A507C4D332CB18DA170A109FACE" box="[394,575,1364,1386]" italics="true" pageId="32" pageNumber="33">Bombus convexus</emphasis>
|
||
Wang
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EF96ABB37C4D332CB2A1A170A078FACE" author="Williams" box="[678,846,1364,1386]" pageId="32" pageNumber="33" refString="Williams, P. H. (1991) The bumble bees of the Kashmir Himalaya (Hymenoptera: Apidae, Bombini). Bulletin of the British Museum (Natural History) (Entomology), 60, 1 - 204." type="journal article" year="1991">
|
||
<collectingMunicipality id="6BDC4C387C4D332CB2A1A170A034FACE" box="[678,770,1364,1386]" pageId="32" pageNumber="33">Williams</collectingMunicipality>
|
||
(1991)
|
||
</bibRefCitation>
|
||
.
|
||
<collectingCounty id="62D9AECE7C4D332CB35FA171A0BFFACE" box="[856,905,1365,1386]" pageId="32" pageNumber="33">Note</collectingCounty>
|
||
1.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BB8D6427C4D332CB090A156A029FA2C" blockId="32.[151,1437,1302,1631]" box="[151,799,1394,1416]" pageId="32" pageNumber="33">
|
||
<treatmentCitationGroup id="AB17F16C7C4D332CB090A156A029FA2C" box="[151,799,1394,1416]" pageId="32" pageNumber="33">
|
||
<treatmentCitation id="0AA6F0537C4D332CB090A156A1B2FA2C" author="Morawitz" box="[151,644,1394,1416]" page="243" pageId="32" pageNumber="33" year="1881">
|
||
<taxonomicName id="4C07ADC17C4D332CB090A156A161FA2C" ID-CoL="4PK2K" authority="Morawitz" authorityName="Morawitz" authorityYear="1881" box="[151,599,1394,1416]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="lugubris">
|
||
<emphasis id="B9730A507C4D332CB090A156A272FA2C" box="[151,324,1394,1416]" italics="true" pageId="32" pageNumber="33">Bombus lugubris</emphasis>
|
||
<bibRefCitation id="EF96ABB37C4D332CB14EA156A161FA2C" author="Morawitz" box="[329,599,1394,1416]" pageId="32" pageNumber="33" refString="Morawitz, F. F. (1881) Die russischen Bombus - Arten in der Sammlung der Kaiserlichen Akademie der Wissenschaften. Izvestiya Imperatorskoi akademii nauk, 27, 213 - 265." type="journal article" year="1881">Morawitz; Morawitz 1881</bibRefCitation>
|
||
</taxonomicName>
|
||
:243
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA6F0537C4D332CB289A157A02DFA2C" author="Wu" box="[654,795,1394,1416]" page="281" pageId="32" pageNumber="33" year="1941">
|
||
<bibRefCitation id="EF96ABB37C4D332CB289A157A1D8FA2C" author="Wu" box="[654,750,1394,1416]" pageId="32" pageNumber="33" refString="Wu, C. - F. (1941) Catalogus insectorum sinensium (Vol. 6). Peiping, Department of Biology, Yenching University." type="journal volume" year="1941">Wu 1941</bibRefCitation>
|
||
:281
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BB8D6427C4D332CB090A1B5A088FA03" blockId="32.[151,1437,1302,1631]" box="[151,958,1425,1447]" pageId="32" pageNumber="33">
|
||
<treatmentCitationGroup id="AB17F16C7C4D332CB090A1B5A088FA03" box="[151,958,1425,1447]" pageId="32" pageNumber="33">
|
||
<taxonomicName id="4C07ADC17C4D332CB090A1B5A123FA03" authority="Morawitz" authorityName="Morawitz" box="[151,533,1425,1447]" class="Insecta" family="Apidae" genus="Mendacibombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="lugubris">
|
||
<emphasis id="B9730A507C4D332CB090A1B5A2ACFA03" box="[151,410,1425,1447]" italics="true" pageId="32" pageNumber="33">Mendacibombus lugubris</emphasis>
|
||
(Morawitz)
|
||
</taxonomicName>
|
||
;
|
||
<treatmentCitation id="0AA6F0537C4D332CB218A1B5A1D0FA03" author="Skorikov" box="[543,742,1425,1447]" page="149" pageId="32" pageNumber="33" year="1923">
|
||
<bibRefCitation id="EF96ABB37C4D332CB218A1B5A18CFA03" author="Skorikov" box="[543,698,1425,1447]" pageId="32" pageNumber="33" refString="Skorikov, A. S. (1923) [Palaearctic bumblebees. Part I. General biology (including zoogeography)]. Izvestiya Severnoi oblastnoi stantsii zashchity rastenii ot vreditelei, 4 (1922), 1 - 160." type="journal article" year="1923">Skorikov 1923</bibRefCitation>
|
||
:149
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA6F0537C4D332CB2F7A1B5A08CFA03" author="Skorikov" box="[752,954,1425,1447]" page="213" pageId="32" pageNumber="33" year="1931">
|
||
<bibRefCitation id="EF96ABB37C4D332CB2F7A1B5A0BAFA03" author="Skorikov" box="[752,908,1425,1447]" pageId="32" pageNumber="33" refString="Skorikov, A. S. (1931) Die Hummelfauna Turkestans und ihre Beziehungen zur zentralasiatischen Fauna (Hymenoptera, Bombidae). In: V. A. Lindholm (Ed.), Abhandlungen der Pamir - Expedition 1928. Academy of Sciences of the USSR, Leningrad, pp. 175 - 247." type="journal article" year="1931">Skorikov, 1931</bibRefCitation>
|
||
:213
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BB8D6427C4D332CB090A194A787FA62" blockId="32.[151,1437,1302,1631]" box="[151,1201,1456,1478]" pageId="32" pageNumber="33">
|
||
<treatmentCitationGroup id="AB17F16C7C4D332CB090A194A787FA62" box="[151,1201,1456,1478]" pageId="32" pageNumber="33">
|
||
<taxonomicName id="4C07ADC17C4D332CB090A194A158FA62" ID-CoL="4PK2K" authority="Morawitz" authorityName="Morawitz" box="[151,622,1456,1478]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="lugubris" subGenus="Mendacibombus">
|
||
<emphasis id="B9730A507C4D332CB090A194A134FA62" box="[151,514,1456,1478]" italics="true" pageId="32" pageNumber="33">Bombus (Mendacibombus) lugubris</emphasis>
|
||
Morawitz
|
||
</taxonomicName>
|
||
;
|
||
<treatmentCitation id="0AA6F0537C4D332CB270A194A01DFA62" author="Bischoff" box="[631,811,1456,1478]" page="17" pageId="32" pageNumber="33" year="1936">
|
||
<bibRefCitation id="EF96ABB37C4D332CB270A194A03BFA62" author="Bischoff" box="[631,781,1456,1478]" pageId="32" pageNumber="33" refString="Bischoff, H. (1936) Schwedisch - chinesische wissenschaftliche Expedition nach den nordwestlichen Provinzen Chinas, unter Leitung von Dr. Sven Hedin und Prof. Su Ping - chang. Insekten gesammelt vom schwedischen Arzt der Expedition Dr. David Hummel 1927 - 1930. 56. Hymenoptera. 10. Bombinae. Arkiv for zoologi, 27 A, 1 - 27." type="journal article" year="1936">Bischoff 1936</bibRefCitation>
|
||
:17
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA6F0537C4D332CB332A194A0C5FA62" author="Panfilov" box="[821,1011,1456,1478]" page="235" pageId="32" pageNumber="33" year="1957">
|
||
<bibRefCitation id="EF96ABB37C4D332CB332A194A0F1FA62" author="Panfilov" box="[821,967,1456,1478]" pageId="32" pageNumber="33" refString="Panfilov, D. V. (1957) [On the geographical distribution of bumblebees (Bombus) in China]. Acta geographica sinica, 23, 221 - 239." type="journal article" year="1957">Panfilov 1957</bibRefCitation>
|
||
:235
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA6F0537C4D332CB3F9A194A79BFA62" author="Tkalcu" box="[1022,1197,1456,1478]" page="369" pageId="32" pageNumber="33" year="1961">
|
||
<bibRefCitation id="EF96ABB37C4D332CB3F9A194A749FA62" author="Tkalcu" box="[1022,1151,1456,1478]" pageId="32" pageNumber="33" refString="Tkalcu, B. (1961) Zur Hummelfauna der Umgebung Kuku - Nors (Hymenoptera, Bombinae). Casopis Ceskoslovenske spolecnosti entomologicke, 58, 344 - 379." type="journal article" year="1961">Tkalců 1961</bibRefCitation>
|
||
:369
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BB8D6427C4D332CB090A1EAA0F0F9A7" blockId="32.[151,1437,1302,1631]" pageId="32" pageNumber="33">
|
||
<materialsCitation id="3B6FDC1F7C4D332CB090A1EAA0F0F9A7" ID-GBIF-Occurrence="1413110426" collectionCode="IZB" country="China" location="Chinese" municipality="Markam" pageId="32" pageNumber="33" specimenCount="1" stateProvince="Xizang" typeStatus="holotype">
|
||
<taxonomicName id="4C07ADC17C4D332CB090A1EAA18EFA40" ID-CoL="MFGK" authority="Wang, 1979:190" authorityName="Wang" authorityPageNumber="190" authorityYear="1979" box="[151,696,1486,1509]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="convexus" subGenus="Mendacibombus">
|
||
<emphasis id="B9730A507C4D332CB090A1EAA13FFA41" box="[151,521,1486,1509]" italics="true" pageId="32" pageNumber="33">Bombus (Mendacibombus) convexus</emphasis>
|
||
Wang, 1979:190
|
||
</taxonomicName>
|
||
, type-locality citation (Chinese) ‘[
|
||
<collectingRegion id="49C318A07C4D332CB40BA1EAA769FA40" box="[1036,1119,1486,1508]" country="China" name="Xizang" pageId="32" pageNumber="33">Xizang</collectingRegion>
|
||
: Markam]’.
|
||
<typeStatus id="54BC68E07C4D332CB4DBA1EAA60CFA40" box="[1244,1338,1486,1508]" pageId="32" pageNumber="33" type="holotype">Holotype</typeStatus>
|
||
queen by original designation
|
||
<collectionCode id="ED164E877C4D332CB190A1CAA2F6F9A7" box="[407,448,1518,1539]" pageId="32" pageNumber="33">IZB</collectionCode>
|
||
examined, (
|
||
<location id="8ED880997C4D332CB23BA1C9A1A6F9A7" LSID="urn:lsid:plazi:treatment:03AE67547C4D332DB090A088A7ADFB04:8ED880997C4D332CB23BA1C9A1A6F9A7" box="[572,656,1517,1539]" country="China" municipality="Markam" name="Chinese" pageId="32" pageNumber="33" stateProvince="Xizang">Chinese</location>
|
||
) ‘[
|
||
<collectingMunicipality id="6BDC4C387C4D332CB2AFA1C9A03DF9A7" box="[680,779,1517,1539]" pageId="32" pageNumber="33">Markam</collectingMunicipality>
|
||
]’ (
|
||
<collectingRegion id="49C318A07C4D332CB325A1C9A059F9A7" box="[802,879,1517,1539]" country="China" name="Xizang" pageId="32" pageNumber="33">Xizang</collectingRegion>
|
||
,
|
||
<collectingCountry id="F31096D27C4D332CB37EA1C9A08CF9A7" box="[889,954,1517,1539]" name="China" pageId="32" pageNumber="33">China</collectingCountry>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BB8D6427C4D332CB090A228A126F9FB" blockId="32.[151,1437,1302,1631]" pageId="32" pageNumber="33">
|
||
<treatmentCitationGroup id="AB17F16C7C4D332CB090A228A126F9FB" pageId="32" pageNumber="33">
|
||
<treatmentCitation id="0AA6F0537C4D332CB090A228A002F986" author="Wang" box="[151,820,1548,1570]" page="429" pageId="32" pageNumber="33" year="1982">
|
||
<taxonomicName id="4C07ADC17C4D332CB090A228A03EF986" ID-CoL="MFGK" authority="Wang" authorityName="Wang" authorityYear="1982" box="[151,776,1548,1570]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="convexus" subGenus="Mendacibombus">
|
||
<emphasis id="B9730A507C4D332CB090A228A13DF986" box="[151,523,1548,1570]" italics="true" pageId="32" pageNumber="33">Bombus (Mendacibombus) convexus</emphasis>
|
||
<bibRefCitation id="EF96ABB37C4D332CB215A229A03EF986" author="Wang" box="[530,776,1548,1570]" pageId="32" pageNumber="33" refString="Wang, S. - F. (1982) Hymenoptera: Apidae - Bombus. In: Q. - T. P. C. S. I. T. o. t. C. A. o. Sciences (Ed), Insects of Xizang. Science Press, Beijing, pp. 427 - 447." type="book chapter" year="1982">Wang; S.-F. Wang 1982</bibRefCitation>
|
||
</taxonomicName>
|
||
:429
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA6F0537C4D332CB338A229A71AF986" author="Yao" box="[831,1068,1548,1570]" page="890" pageId="32" pageNumber="33" year="2005">
|
||
<bibRefCitation id="EF96ABB37C4D332CB338A229A736F986" author="Yao" box="[831,1024,1548,1570]" pageId="32" pageNumber="33" refString="Yao, J. & Wang, S. - F. (2005) Hymenoptera, Apidae, Bombini. In: X. Yang (Ed), Insect fauna of middle - west Qinling range and south mountains of Gansu province. Science Press, Beijing, pp. 890 - 896." type="book chapter" year="2005">Yao & Wang 2005</bibRefCitation>
|
||
:890
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA6F0537C4D332CB430A229A616F986" author="Williams" box="[1079,1312,1548,1570]" page="42" pageId="32" pageNumber="33" year="1991">
|
||
<bibRefCitation id="EF96ABB37C4D332CB430A229A636F986" author="Williams" box="[1079,1280,1548,1570]" pageId="32" pageNumber="33" refString="Williams, P. H. (1991) The bumble bees of the Kashmir Himalaya (Hymenoptera: Apidae, Bombini). Bulletin of the British Museum (Natural History) (Entomology), 60, 1 - 204." type="journal article" year="1991">P.H. Williams 1991</bibRefCitation>
|
||
:42
|
||
</treatmentCitation>
|
||
; S.-
|
||
<treatmentCitation id="0AA6F0537C4D332CB541A228A200F9E4" author="Wang" page="1424" pageId="32" pageNumber="33" year="1992">
|
||
<bibRefCitation id="EF96ABB37C4D332CB541A228A3CBF9E4" author="Wang" pageId="32" pageNumber="33" refString="Wang, S. - F. (1992) Hymenoptera: Apoidae (II) - Bombus. In: The series of the scientific expedition to the Hengduan Mountains region of Qinghai - Xizang Plateau. Chinese Academy of Sciences, pp. 1422 - 1430." type="book chapter" year="1992">F. Wang 1992</bibRefCitation>
|
||
:1424
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA6F0537C4D332CB146A20FA101F9E4" author="Williams" box="[321,567,1578,1600]" page="100" pageId="32" pageNumber="33" year="1998">
|
||
<bibRefCitation id="EF96ABB37C4D332CB146A20FA13DF9E4" author="Williams" box="[321,523,1578,1600]" pageId="32" pageNumber="33" refString="Williams, P. H. (1998) An annotated checklist of bumble bees with an analysis of patterns of description (Hymenoptera: Apidae, Bombini). Bulletin of The Natural History Museum (Entomology), 67, 79 - 152. [updated at www. nhm. ac. uk / bombus / accessed 2015]." type="journal article" year="1998">P.H. Williams 1998</bibRefCitation>
|
||
:100
|
||
</treatmentCitation>
|
||
;
|
||
<bibRefCitation id="EF96ABB37C4D332CB245A20FA03BF9E4" author="Williams" box="[578,781,1578,1600]" pageId="32" pageNumber="33" refString="Williams, P. H. (2004) Genus Bombus Latreille. In: R. K. Gupta (Ed), An annotated catalogue of the bee species of the Indian Region. Jai Narain Vyas University, Jodhpur, p. Web page providing a list of bumblebees of India and adjacent countries with common synonyms and a summary of their distributions within the region." type="book" year="2004">P.H. Williams 2004</bibRefCitation>
|
||
:no. 26;
|
||
<treatmentCitation id="0AA6F0537C4D332CB35AA20EA76BF9E4" author="Cameron" box="[861,1117,1578,1600]" page="165" pageId="32" pageNumber="33" year="2007">
|
||
<bibRefCitation id="EF96ABB37C4D332CB35AA20EA704F9E4" author="Cameron" box="[861,1074,1578,1600]" pageId="32" pageNumber="33" refString="Cameron, S. A., Hines, H. M. & Williams, P. H. (2007) A comprehensive phylogeny of the bumble bees (Bombus). Biological Journal of the Linnean Society, 91, 161 - 188. http: // dx. doi. org / 10.1111 / j. 1095 - 8312.2007.00784. x" type="journal article" year="2007">
|
||
Cameron
|
||
<emphasis id="B9730A507C4D332CB3C7A20FA0C2F9E4" box="[960,1012,1578,1600]" italics="true" pageId="32" pageNumber="33">et al.</emphasis>
|
||
2007
|
||
</bibRefCitation>
|
||
:165
|
||
</treatmentCitation>
|
||
; P.H.
|
||
<treatmentCitation id="0AA6F0537C4D332CB49EA20EA6AEF9E4" author="Williams" box="[1177,1432,1578,1600]" page="130" pageId="32" pageNumber="33" year="2009">
|
||
<bibRefCitation id="EF96ABB37C4D332CB49EA20EA65BF9E4" author="Williams" box="[1177,1389,1578,1600]" pageId="32" pageNumber="33" refString="Williams, P. H., Tang, Y., Yao, J. & Cameron, S. (2009) The bumblebees of Sichuan (Hymenoptera: Apidae, Bombini). Systematics and Biodiversity, 7, 101 - 190." type="journal article" year="2009">
|
||
Williams
|
||
<emphasis id="B9730A507C4D332CB4FCA20FA619F9E4" box="[1275,1327,1578,1600]" italics="true" pageId="32" pageNumber="33">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
:130
|
||
</treatmentCitation>
|
||
; An
|
||
<emphasis id="B9730A507C4D332CB0EBA26EA229F9FB" box="[236,287,1609,1631]" italics="true" pageId="32" pageNumber="33">et al.</emphasis>
|
||
2011:5;
|
||
<bibRefCitation id="EF96ABB37C4D332CB17FA26EA13AF9FB" author="An" box="[376,524,1609,1631]" pageId="32" pageNumber="33" refString="An, J. - D., Huang, J. - X., Shao, Y. - Q., Zhang, S. - W., Wang, B., Liu, X. - Y., Wu, J. & Williams, P. H. (2014) The bumblebees of North China (Apidae, Bombus Latreille). Zootaxa, 3830 (1), 1 - 89. http: // dx. doi. org / 10.11646 / zootaxa. 3830.1.1" type="journal article" year="2014">
|
||
An
|
||
<emphasis id="B9730A507C4D332CB19AA26EA2E6F9FB" box="[413,464,1609,1631]" italics="true" pageId="32" pageNumber="33">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BB8D6427C4D332CB090A2A3A275F943" blockId="32.[151,1437,1670,2019]" pageId="32" pageNumber="33">
|
||
Note 1 (
|
||
<taxonomicName id="4C07ADC17C4D332CB0F2A2A2A264F93B" ID-CoL="4PK2K" box="[245,338,1670,1695]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="lugubris">
|
||
<emphasis id="B9730A507C4D332CB0F2A2A2A264F93B" box="[245,338,1670,1695]" italics="true" pageId="32" pageNumber="33">lugubris</emphasis>
|
||
</taxonomicName>
|
||
). The original publication specifies that there was only one type (worker) specimen of the taxon
|
||
<taxonomicName id="4C07ADC17C4D332CB090A28EA25BF967" ID-CoL="4PK2K" authority="Morawitz" authorityName="Morawitz" box="[151,365,1706,1731]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="lugubris">
|
||
<emphasis id="B9730A507C4D332CB090A28EA3C2F967" box="[151,244,1706,1731]" italics="true" pageId="32" pageNumber="33">lugubris</emphasis>
|
||
Morawitz
|
||
</taxonomicName>
|
||
, so the worker in the ZISP collection is regarded as the holotype by monotypy (
|
||
<bibRefCitation id="EF96ABB37C4D332CB50EA28FA201F943" author="ICZN" pageId="32" pageNumber="33" refString="ICZN (1999) International code of zoological nomenclature (4 ed.). London, International Commission on Zoological Nomenclature." type="book chapter" year="1999">ICZN, 1999: Article 73.1.2</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D85C97C4D332CB0C0A2D6A0B9F81B" pageId="32" pageNumber="33" type="etymology">
|
||
<paragraph id="8BB8D6427C4D332CB0C0A2D6A6ABF88B" blockId="32.[151,1437,1670,2019]" pageId="32" pageNumber="33">
|
||
<emphasis id="B9730A507C4D332CB0C0A2D6A278F8AF" bold="true" box="[199,334,1778,1803]" pageId="32" pageNumber="33">Etymology.</emphasis>
|
||
The species is named from the Latin
|
||
<taxonomicName id="4C07ADC17C4D332CB321A2D0A0BAF8AF" box="[806,908,1780,1803]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="convexus">
|
||
<emphasis id="B9730A507C4D332CB321A2D0A0BAF8AF" box="[806,908,1780,1803]" italics="true" pageId="32" pageNumber="33">convexus</emphasis>
|
||
</taxonomicName>
|
||
for ‘rounded’, a reference to a pronounced convexity or rounded posterior projection of the median posterior edge of metasomal T
|
||
<quantity id="4CFF7BA77C4D332CB465A333A7BAF88B" box="[1122,1164,1814,1839]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="32" pageNumber="33" unit="in" value="2.0">2 in</quantity>
|
||
the original description.
|
||
</paragraph>
|
||
<paragraph id="8BB8D6427C4D332CB0C0A31EA0B9F81B" blockId="32.[151,1437,1670,2019]" pageId="32" pageNumber="33">
|
||
<emphasis id="B9730A507C4D332CB0C0A31EA136F8F7" bold="true" box="[199,512,1850,1875]" pageId="32" pageNumber="33">Taxonomy and variation.</emphasis>
|
||
This species shows effectively a single colour pattern of the hair with little variation. This is the pattern in the original description, which is unique within the subgenus. All specimens have a white-banded and none has a yellow-banded colour pattern. The form of the female median posterior edge of T2 (although slightly variable) and of the male genitalia are diagnostic.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D85C97C4D332DB0C0A3EEA03BFD54" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="description">
|
||
<paragraph id="8BB8D6427C4D332DB0C0A3EEA03BFD54" blockId="32.[151,1437,1670,2019]" lastBlockId="33.[151,1437,151,1184]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
|
||
<emphasis id="B9730A507C4D332CB0C0A3EEA2E9F847" bold="true" box="[199,479,1994,2019]" pageId="32" pageNumber="33">Diagnostic description.</emphasis>
|
||
Wings nearly clear. Hair long, uneven and sparse.
|
||
<emphasis id="B9730A507C4D332CB448A3EEA6AAF847" box="[1103,1436,1994,2019]" italics="true" pageId="32" pageNumber="33">Female hair colour pattern:</emphasis>
|
||
generally black, but with white hair intermixed especially as short hairs on the face, intermixed in a transverse band anteriorly on the thoracic dorsum and extending laterally and ventrally without black hairs to the midleg base, in a lateral patch posteriorly on the thoracic dorsum (lateral patch on the scutellum, metanotum, and propodeum, so that the thoracic dorsum between the wing bases has the hair entirely black), on T1 and anteriorly on T2, and intermixed on T4‒6 (
|
||
<emphasis id="B9730A507C4C332DB18CA50CA2A9FEE4" box="[395,415,296,320]" italics="true" pageId="33" pageNumber="34">cf</emphasis>
|
||
. all other
|
||
<taxonomicName id="4C07ADC17C4C332DB211A503A1F9FEE4" box="[534,719,295,320]" class="Insecta" family="Apidae" genus="Mendacibombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9730A507C4C332DB211A503A1F9FEE4" box="[534,719,295,320]" italics="true" pageId="33" pageNumber="34">Mendacibombus</emphasis>
|
||
</taxonomicName>
|
||
species). Hindleg tibia with the corbicular fringes black, often with a few hairs with orange or white tips.
|
||
<emphasis id="B9730A507C4C332DB290A568A0B1FEC1" box="[663,903,332,357]" italics="true" pageId="33" pageNumber="34">Female morphology:</emphasis>
|
||
labrum with the basal depression narrow, the transverse ridge moderately broad and high, medially not subsiding or interrupted and in the median third with scattered large punctures, lateral tubercles with a few scattered small punctures. T2 usually with a posteriorlydirected convexity of its median posterior edge, affecting a quarter of its breadth, especially pronounced in queens, for which there may be edge concavities and even slight pre-marginal depressions lateral to the convexity.
|
||
<emphasis id="B9730A507C4C332DB565A5F8A228FDBC" italics="true" pageId="33" pageNumber="34">Male morphology</emphasis>
|
||
: beard of the mandible long, dense and black, but the short hairs and a few of the long hairs brown; T2 and sometimes T3 with posteriorly-directed convexities of their median posterior edge, affecting a quarter or less of their breadth. Genitalia (
|
||
<figureCitation id="133CCAC77C4C332DB1D1A663A11CFDC4" box="[470,554,583,608]" captionStart="FIGURES 24 ‒ 35" captionStartId="26.[151,264,1867,1889]" captionTargetBox="[233,1387,193,1789]" captionTargetId="figure@26.[233,1390,193,1894]" captionTargetPageId="26" captionText="FIGURES 24 ‒ 35. Morphology of the male genitalia for species of the subgenus Mendacibombus from the dorsal aspect, anterior to the left of the image, posterior to the right: 24, B. superbus, China MD # 339; 25, B. waltoni, China MD # 216; 26, B. convexus, China MD # 215; 27, B. makarjini, Kazakhstan MD # 1240; 28, B. marussinus, Pakistan MD # 408; 29, B. avinoviellus, India MD # 404; 30, B. himalayanus, India MD # 426; 31, B. turkestanicus, China MD # 351; 32, B. defector, Kyrgyzstan MD # 367; 33, B. margreiteri, China MD # 294; 34, B. handlirschianus, Russia MD # 835; 35, B. mendax, Spain MD # 689." httpUri="https://zenodo.org/record/192312/files/figure.png" pageId="33" pageNumber="34">Fig. 26</figureCitation>
|
||
) with the volsella at its broadest near the midpoint of its length, the dorsal surface just distal to this point with a raised curved ridge, often with small teeth, just inside the inner margin, running for 0.3× the remaining distal length of the volsella; volsella distally sharply acute (pointed) and curled back dorsally and anteriorly. Gonostylus length 2× its greatest breadth. Penis-valve head length 0.25× the length of the penis valve distal to the broadest point of the spatha.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D85C97C4C332DB0C0A6D8A6AEFB90" pageId="33" pageNumber="34" type="materials_examined">
|
||
<paragraph id="8BB8D6427C4C332DB0C0A6D8A7CEFC9C" blockId="33.[151,1437,151,1184]" pageId="33" pageNumber="34">
|
||
<emphasis id="B9730A507C4C332DB0C0A6D8A29FFCB1" bold="true" box="[199,425,764,789]" pageId="33" pageNumber="34">Material examined</emphasis>
|
||
. 6 queens
|
||
<specimenCount id="9D011DCB7C4C332DB227A6D8A19DFCB1" box="[544,683,764,789]" pageId="33" pageNumber="34" type="worker">296 workers</specimenCount>
|
||
<specimenCount id="9D011DCB7C4C332DB2B5A6D8A02FFCB1" box="[690,793,764,789]" pageId="33" pageNumber="34" type="male">51 males</specimenCount>
|
||
, from
|
||
<collectingCountry id="F31096D27C4C332DB366A6D8A092FCB1" box="[865,932,764,789]" name="China" pageId="33" pageNumber="34">China</collectingCountry>
|
||
(
|
||
<figureCitation id="133CCAC77C4C332DB3B4A6D8A731FCB1" box="[947,1031,764,789]" captionStart="FIGURES 56 ‒ 67" captionStartId="35.[151,264,1891,1913]" captionTargetBox="[320,1245,193,1835]" captionTargetId="figure@35.[320,1245,193,1843]" captionTargetPageId="35" captionText="FIGURES 56 ‒ 67. Maps of the global distributions of species of the subgenus Mendacibombus from the material examined, with grey spots for specimen records and black spots for the samples with COI sequences (each species is unknown from regions outside the map boundaries shown); grey crosses for records of all species of Mendacibombus combined. Relief map with hill shading, Cartesian orthonormal projection, grey lines show borders to national administration according to UN maps, north at the top of the map. Image created in ArcGIS using World _ Shaded _ Relief basemap © 2014 Esri." httpUri="https://zenodo.org/record/192315/files/figure.png" pageId="33" pageNumber="34">Fig. 58</figureCitation>
|
||
:
|
||
<collectionCode id="ED164E877C4C332DB415A6D8A75CFCB0" box="[1042,1130,764,788]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="33" pageNumber="34">AMNH</collectionCode>
|
||
,
|
||
<collectionCode id="ED164E877C4C332DB472A6D8A79EFCB0" box="[1141,1192,764,788]" pageId="33" pageNumber="34">IAR</collectionCode>
|
||
,
|
||
<collectionCode id="ED164E877C4C332DB4B4A6D9A7D5FCB0" box="[1203,1251,765,788]" pageId="33" pageNumber="34">IZB</collectionCode>
|
||
,
|
||
<collectionCode id="ED164E877C4C332DB4E9A6D9A604FCB0" box="[1262,1330,765,788]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="33" pageNumber="34">NHM</collectionCode>
|
||
, PW, SC,
|
||
<collectionCode id="ED164E877C4C332DB090A704A3DDFC9C" box="[151,235,800,824]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="33" pageNumber="34">USNM</collectionCode>
|
||
, YT,
|
||
<collectionCode id="ED164E877C4C332DB120A704A256FC9C" box="[295,352,800,824]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="33" pageNumber="34">ZISP</collectionCode>
|
||
, ZX), with
|
||
<specimenCount id="9D011DCB7C4C332DB1E6A73BA15DFC9C" box="[481,619,799,824]" pageId="33" pageNumber="34" type="generic">6 specimens</specimenCount>
|
||
sequenced (interpretable sequences listed in
|
||
<figureCitation id="133CCAC77C4C332DB463A73BA7DCFC9C" box="[1124,1258,799,824]" captionStart-0="FIGURES 11 ‒ 12" captionStart-1="FIGURE 13" captionStartId-0="15.[151,264,1630,1652]" captionStartId-1="16.[151,250,1733,1755]" captionTargetBox-0="[300,1121,202,1610]" captionTargetBox-1="[314,1260,194,1721]" captionTargetId-0="figure@15.[292,1266,193,1625]" captionTargetId-1="figure@16.[314,1261,193,1722]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="FIGURES 11 ‒ 12. Recognising species from an estimate of phylogeny for 30 unique COI haplotypes of the subgenus Mendacibombus by BEAST, using GMYC models and the PTP procedure. From the GMYC models (11), the single threshold T is shown by the vertical grey bar so that the intersecting lineages are interpreted as indicating 12 separate species (black spots at coalescent nodes; outgroups not shown). Values below the nodes of the maximum clade-credibility tree are Bayesian posterior probabilities showing branch support (groups with values <0.8 are considered unreliable) from the BEAST analysis with two billion MCMC generations and a 1 % burn-in. The x - scale axis is calibrated in substitutions per nucleotide site. From the PTP procedure (12), the Bayesian PTP solution with highest support also indicates 12 species. Values above the nodes are PTP Bayesian support values that all daughter haplotypes are parts of a single species; values below the nodes are Bayesian posterior probabilities showing branch support. The scale bar is calibrated in substitutions per nucleotide site. Each unique haplotype is represented by one of the longest available sample sequences, labelled with: the sequence length in number of base pairs, a taxon name or name for an unpublished colour pattern (light / dark / yellow / white / pale), and then a code that consists of a specimen identifier (MD #) from the project database and (after the hyphen) a sample identifier from BOLD or from GenBank, followed with its geographic origin." captionText-1="FIGURE 13. Assigning names to species from an estimate of phylogeny for 130 COI sequences of the subgenus Mendacibombus. Based on a BEAST analysis of the 30 unique haplotypes (Fig. 11), with the remaining sequences interpolated back into the tree with 0 branch lengths from their matching samples (where a sequence matches more than one longer sequence within a species in the Collapse results, it is added after the first matching sequence, with the addition of the two short-sequence yellow unbanded specimens from Pakistan, MD # 4058, 4059). The sequence labels follow the format used in Figs 11, 12. Lineages accepted as species from the GMYC models (Fig. 11) are shown as thick branches, the most recent common ancestors of each species are shown with a black spot, and branches within species are shown as thin branches. Asterisks mark samples used as informal proxies for the type specimens of each of the taxon names (Table 1). The proxy sample for the type specimen for the oldest available name (the valid name) for each species (from Fig. 11) is marked in bold. The interpretations of the species with their valid names are shown in grey to the right." httpUri-0="https://zenodo.org/record/192308/files/figure.png" httpUri-1="https://zenodo.org/record/192309/files/figure.png" pageId="33" pageNumber="34">Figs. 11–13</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB8D6427C4C332DB0C0A760A6AEFB90" blockId="33.[151,1437,151,1184]" pageId="33" pageNumber="34">
|
||
<emphasis id="B9730A507C4C332DB0C0A760A2C5FCF9" bold="true" box="[199,499,836,861]" pageId="33" pageNumber="34">Habitat and distribution</emphasis>
|
||
. Flower-rich alpine and subalpine grassland, at elevations 2196‒(3449)‒
|
||
<quantity id="4CFF7BA77C4C332DB545A760A6AAFCF8" box="[1346,1436,836,861]" metricMagnitude="3" metricUnit="m" metricValue="4.5" pageId="33" pageNumber="34" unit="m" value="4500.0">4500 m</quantity>
|
||
a.s.l.. A species of the east Qinghai-Tibetan plateau, in the east Himalayan, Hengduan, and Qinghai-Gansu mountains, including the Min Shan and Qilian Shan. Compared to
|
||
<taxonomicName id="4C07ADC17C4C332DB3AFA7A9A729FC01" box="[936,1055,908,933]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="waltoni">
|
||
<emphasis id="B9730A507C4C332DB3AFA7A9A729FC01" box="[936,1055,908,933]" italics="true" pageId="33" pageNumber="34">B. waltoni</emphasis>
|
||
</taxonomicName>
|
||
, distributions of the two species overlap broadly, but
|
||
<taxonomicName id="4C07ADC17C4C332DB185A795A13EFC6C" box="[386,520,945,968]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="convexus">
|
||
<emphasis id="B9730A507C4C332DB185A795A13EFC6C" box="[386,520,945,968]" italics="true" pageId="33" pageNumber="34">B. convexus</emphasis>
|
||
</taxonomicName>
|
||
extends less far to the north and west and tends to occur at lower elevation (and the two species rarely occur together at a site).
|
||
<taxonomicName id="4C07ADC17C4C332DB2ACA7F0A045FC48" box="[683,883,980,1005]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="convexus">
|
||
<emphasis id="B9730A507C4C332DB2ACA7F0A045FC48" box="[683,883,980,1005]" italics="true" pageId="33" pageNumber="34">Bombus convexus</emphasis>
|
||
</taxonomicName>
|
||
replaces the western
|
||
<taxonomicName id="4C07ADC17C4C332DB461A7F1A63BFC49" box="[1126,1293,980,1005]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="avinoviellus">
|
||
<emphasis id="B9730A507C4C332DB461A7F1A63BFC49" box="[1126,1293,980,1005]" italics="true" pageId="33" pageNumber="34">B. avinoviellus</emphasis>
|
||
</taxonomicName>
|
||
in the lower alpine zone and upper forest wet meadows of the eastern Himalaya, where it appears to be rare. Regional distribution maps are available for
|
||
<collectingRegion id="49C318A07C4C332DB219A038A14EFB91" box="[542,632,1052,1077]" country="China" name="Sichuan" pageId="33" pageNumber="34">Sichuan</collectingRegion>
|
||
(P.H.
|
||
<bibRefCitation id="EF96ABB37C4C332DB2B9A038A092FB90" author="Williams" box="[702,932,1052,1077]" pageId="33" pageNumber="34" refString="Williams, P. H., Tang, Y., Yao, J. & Cameron, S. (2009) The bumblebees of Sichuan (Hymenoptera: Apidae, Bombini). Systematics and Biodiversity, 7, 101 - 190." type="journal article" year="2009">
|
||
Williams
|
||
<emphasis id="B9730A507C4C332DB32CA039A055FB91" box="[811,867,1052,1077]" italics="true" pageId="33" pageNumber="34">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
) and
|
||
<collectingRegion id="49C318A07C4C332DB3E6A038A71FFB90" box="[993,1065,1052,1076]" country="China" name="Gansu" pageId="33" pageNumber="34">Gansu</collectingRegion>
|
||
(An
|
||
<emphasis id="B9730A507C4C332DB464A039A7ADFB91" box="[1123,1179,1052,1077]" italics="true" pageId="33" pageNumber="34">et al.</emphasis>
|
||
2011;
|
||
<bibRefCitation id="EF96ABB37C4C332DB4E0A038A6BAFB90" author="An" box="[1255,1420,1052,1077]" pageId="33" pageNumber="34" refString="An, J. - D., Huang, J. - X., Shao, Y. - Q., Zhang, S. - W., Wang, B., Liu, X. - Y., Wu, J. & Williams, P. H. (2014) The bumblebees of North China (Apidae, Bombus Latreille). Zootaxa, 3830 (1), 1 - 89. http: // dx. doi. org / 10.11646 / zootaxa. 3830.1.1" type="journal article" year="2014">
|
||
An
|
||
<emphasis id="B9730A507C4C332DB517A039A67FFB91" box="[1296,1353,1052,1077]" italics="true" pageId="33" pageNumber="34">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D85C97C4C332DB0C0A01BA7ADFB04" pageId="33" pageNumber="34" type="description">
|
||
<paragraph id="8BB8D6427C4C332DB0C0A01BA050FBFC" blockId="33.[151,1437,151,1184]" box="[199,870,1087,1112]" pageId="33" pageNumber="34">
|
||
<emphasis id="B9730A507C4C332DB0C0A01BA262FBFC" bold="true" box="[199,340,1087,1112]" pageId="33" pageNumber="34">Food plants</emphasis>
|
||
.
|
||
<bibRefCitation id="EF96ABB37C4C332DB165A01BA16CFBFC" box="[354,602,1087,1112]" pageId="33" pageNumber="34" refString="Williams, P. H., Tang, Y., Yao, J. & Cameron, S. (2009) The bumblebees of Sichuan (Hymenoptera: Apidae, Bombini). Systematics and Biodiversity, 7, 101 - 190." type="journal article">
|
||
Williams
|
||
<emphasis id="B9730A507C4C332DB1C9A065A2C9FBFC" box="[462,511,1087,1112]" italics="true" pageId="33" pageNumber="34">et al</emphasis>
|
||
. (2009)
|
||
</bibRefCitation>
|
||
, An
|
||
<emphasis id="B9730A507C4C332DB288A065A1F6FBFC" box="[655,704,1087,1112]" italics="true" pageId="33" pageNumber="34">et al</emphasis>
|
||
. (2011; 2014).
|
||
</paragraph>
|
||
<paragraph id="8BB8D6427C4C332DB0C0A040A7ADFB04" blockId="33.[151,1437,151,1184]" pageId="33" pageNumber="34">
|
||
<materialsCitation id="3B6FDC1F7C4C332DB0C0A040A7ADFB04" ID-GBIF-Occurrence="1413110427" collectingDate="2009-08-29" country="China" location="Diebu" pageId="33" pageNumber="34" specimenCount="1" stateProvince="Gansu">
|
||
<emphasis id="B9730A507C4C332DB0C0A040A276FBD9" bold="true" box="[199,320,1124,1149]" pageId="33" pageNumber="34">Behaviour</emphasis>
|
||
. Mate-searching males perch on bare patches of ground and pursue other bees that fly past before the males return to the same perch (PW:
|
||
<figureCitation id="133CCAC77C4C332DB258A0A3A194FB04" box="[607,674,1159,1184]" captionStart="FIGURES 1 ‒ 6" captionStartId="5.[151,264,1819,1841]" captionTargetBox="[168,1389,193,1768]" captionTargetId="figure@5.[168,1412,193,1784]" captionTargetPageId="5" captionText="FIGURES 1 ‒ 6. Views of perching males of species of the subgenus Mendacibombus, showing the anteriorly extended antennae and laterally extended wings that are characteristic of their form of male mate-searching behaviour: 1, B. waltoni at Diebu ca 3910 m, Minshan mountains, Gansu, China, 29. viii. 2009; 2, B. convexus at Diebu ca 3340 m, Minshan mountains, Gansu, China, 29. viii. 2009; 3, B. avinoviellus at Gulmarg ca 2700 m, Pir Panjal mountains, Kashmir, India, 16. viii. 1985; 4, B. himalayanus at Apharwat ca 4100 m, Pir Panjal mountains, Kashmir, India, 30. viii. 1986; 5 ‒ 6, B. margreiteri near Dulan ca 3520 m, Burhan Budai mountains, Qinghai, China, 31. viii. 2010. Photos by PW (Figs 3 – 4 are the best available for these species)." httpUri="https://zenodo.org/record/192303/files/figure.png" pageId="33" pageNumber="34">Fig. 2</figureCitation>
|
||
,
|
||
<date id="FFB9F0827C4C332DB2A8A0A3A00EFB04" box="[687,824,1159,1184]" pageId="33" pageNumber="34" value="2009-08-29">
|
||
<collectingDate id="EFFD096A7C4C332DB2A8A0A3A00EFB04" box="[687,824,1159,1184]" pageId="33" pageNumber="34" value="2009-08-29">
|
||
<figureCitation id="133CCAC77C4C332DB2A8A0A3A1FAFB04" box="[687,716,1159,1184]" captionStart="FIGURES 24 ‒ 35" captionStartId="26.[151,264,1867,1889]" captionTargetBox="[233,1387,193,1789]" captionTargetId="figure@26.[233,1390,193,1894]" captionTargetPageId="26" captionText="FIGURES 24 ‒ 35. Morphology of the male genitalia for species of the subgenus Mendacibombus from the dorsal aspect, anterior to the left of the image, posterior to the right: 24, B. superbus, China MD # 339; 25, B. waltoni, China MD # 216; 26, B. convexus, China MD # 215; 27, B. makarjini, Kazakhstan MD # 1240; 28, B. marussinus, Pakistan MD # 408; 29, B. avinoviellus, India MD # 404; 30, B. himalayanus, India MD # 426; 31, B. turkestanicus, China MD # 351; 32, B. defector, Kyrgyzstan MD # 367; 33, B. margreiteri, China MD # 294; 34, B. handlirschianus, Russia MD # 835; 35, B. mendax, Spain MD # 689." httpUri="https://zenodo.org/record/192312/files/figure.png" pageId="33" pageNumber="34">29</figureCitation>
|
||
.viii.
|
||
<quantity id="4CFF7BA77C4C332DB2FCA0A3A00EFB04" box="[763,824,1159,1184]" metricMagnitude="3" metricUnit="m" metricValue="2.0093339" pageId="33" pageNumber="34" unit="m" value="2009.3339">2009</quantity>
|
||
</collectingDate>
|
||
</date>
|
||
, 3339 m
|
||
<location id="8ED880997C4C332DB3A7A0A3A0D1FB04" LSID="urn:lsid:plazi:treatment:03AE67547C4D332DB090A088A7ADFB04:8ED880997C4C332DB3A7A0A3A0D1FB04" box="[928,999,1159,1184]" country="China" name="Diebu" pageId="33" pageNumber="34" stateProvince="Gansu">Diebu</location>
|
||
,
|
||
<collectingRegion id="49C318A07C4C332DB3F5A0ACA70AFB04" box="[1010,1084,1160,1184]" country="China" name="Gansu" pageId="33" pageNumber="34">Gansu</collectingRegion>
|
||
,
|
||
<collectingCountry id="F31096D27C4C332DB44FA0A3A7A6FB04" box="[1096,1168,1159,1184]" name="China" pageId="33" pageNumber="34">China</collectingCountry>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |