306 lines
43 KiB
XML
306 lines
43 KiB
XML
<document ID-DOI="10.11646/zootaxa.5277.1.3" ID-GBIF-Dataset="195c7011-1a00-4c5f-a339-dc529aa9e104" ID-ISSN="1175-5326" ID-Zenodo-Dep="7891898" ID-ZooBank="2EBD6E19-89E9-4918-8EE3-3324DF3BBA13" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1682930661848" checkinUser="plazi" docAuthor="Jałoszyński, Paweł, Hlaváč, Peter & Anlaş, Sinan" docDate="2023" docId="03AB722FA107FFED25C9FE2696C8FD75" docLanguage="en" docName="zootaxa.5277.1.3.pdf" docOrigin="Zootaxa 5277 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5277.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thaumastocephalus Poggi, Nonveiller, Colla, Pavicevic & Rada 2001" docType="treatment" docVersion="3" lastPageNumber="86" masterDocId="FF920A57A10BFFE2255EFFB59768FFDC" masterDocTitle="Percussiopalpus gen. n. from Turkey, a free-living relative of cave-dwelling Thaumastocephalus, and revised placement of Thaumastocephalini (Coleoptera: Staphylinidae: Pselaphinae)" masterLastPageNumber="90" masterPageNumber="71" pageNumber="83" updateTime="1683121264666" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo>
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<mods:title>Percussiopalpus gen. n. from Turkey, a free-living relative of cave-dwelling Thaumastocephalus, and revised placement of Thaumastocephalini (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Jałoszyński, Paweł</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0003-2973-1803</mods:nameIdentifier>
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<mods:affiliation>Museum ofNaturalHistory, UniversityofWrocław, Sienkiewicza 21, 50 - 335 Wrocław, Poland. https: // orcid. org / 0000 - 0003 - 2973 - 1803 Department of Entomology, National Museum, Natural History Museum, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Počernice, The Czech Republic. peterhlavac @ gmail. com; https: // orcid. org / 0000 - 0001 - 5060 - 0811</mods:affiliation>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Hlaváč, Peter</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0001-5060-0811</mods:nameIdentifier>
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<mods:nameIdentifier type="email">peterhlavac@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Anlaş, Sinan</mods:namePart>
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<mods:affiliation>Manisa Celal Bayar University, Alaşehir Vocational School, TR- 45600, Alaşehir, Manisa, Turkey.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-05-01</mods:number>
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<mods:detail type="volume">
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<mods:number>5277</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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<mods:start>71</mods:start>
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<mods:end>90</mods:end>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5277.1.3</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5277.1.3</mods:identifier>
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<mods:identifier type="GBIF-Dataset">195c7011-1a00-4c5f-a339-dc529aa9e104</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7891898</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7891956" ID-Zenodo-Dep="7891956" LSID="urn:lsid:plazi:treatment:03AB722FA107FFED25C9FE2696C8FD75" httpUri="http://treatment.plazi.org/id/03AB722FA107FFED25C9FE2696C8FD75" lastPageId="15" lastPageNumber="86" pageId="12" pageNumber="83">
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<subSubSection box="[151,1008,402,430]" pageId="12" pageNumber="83" type="nomenclature">
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<paragraph blockId="12.[151,1008,402,466]" box="[151,1008,402,430]" pageId="12" pageNumber="83">
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<heading bold="true" box="[151,1008,402,430]" fontSize="11" level="1" pageId="12" pageNumber="83" reason="1">
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<taxonomicName ID-CoL="8XQBS" authority="Poggi, Nonveiller, Colla, Pavicevic & Rada, 2001" authorityName="Poggi, Nonveiller, Colla, Pavicevic & Rada" authorityYear="2001" box="[151,1008,402,430]" class="Insecta" family="Staphylinidae" genus="Thaumastocephalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="83" phylum="Arthropoda" rank="genus">
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<emphasis bold="true" box="[151,1008,402,430]" pageId="12" pageNumber="83">
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<emphasis bold="true" box="[151,395,403,430]" italics="true" pageId="12" pageNumber="83">Thaumastocephalus</emphasis>
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<bibRefCitation author="Poggi, R. & Nonveiller, G. & Colla, A. & Pavicevic, D. & Rada, T." box="[403,1008,402,429]" pageId="12" pageNumber="83" pagination="1 - 20" refId="ref10503" refString="Poggi, R., Nonveiller, G., Colla, A., Pavicevic, D. & Rada, T. (2001) Thaumastocephalini, a new tribe of Pselaphinae for Thaumastocephalus folliculipalpus n. gen., n. sp., an interesting new troglobious species from central Dalmatia (Croatia) (Coleoptera, Staphylinidae, Pselaphinae). Annali del Museo Civico di Storia Naturale " Giacomo Doria ", 54, 1 - 20." type="journal article" year="2001">Poggi, Nonveiller, Colla, Pavićević & Rada, 2001</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,295,439,466]" pageId="12" pageNumber="83" type="description">
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<paragraph blockId="12.[151,1008,402,466]" box="[151,295,439,466]" pageId="12" pageNumber="83">
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(
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<figureCitation box="[159,287,439,466]" captionStart-0="FIGURES 41–44" captionStart-1="FIGURES 45–50" captionStart-2="FIGURES 51–55" captionStartId-0="13.[151,264,1540,1564]" captionStartId-1="14.[151,264,1679,1703]" captionStartId-2="15.[151,264,1925,1949]" captionTargetBox-0="[161,1436,328,1505]" captionTargetBox-1="[151,1436,634,1644]" captionTargetBox-2="[160,1436,713,1902]" captionTargetId-0="figure-77@13.[151,1436,309,1517]" captionTargetId-1="figure-191@14.[151,1436,633,1656]" captionTargetId-2="figure-238@15.[151,1436,705,1902]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="15" captionText-0="FIGURES 41–44. Thaumastocephalus sp., male from Croatia. Dorsal habitus (41), head in dorsal view (42), prothorax and elytral base in dorsal view (43), anterior body region in lateral view (44)." captionText-1="FIGURES 45–50. Thaumastocephalus sp., male from Croatia. Head in ventral (45) and lateral view (46), antenna in lateral view (47), prothorax and proximal region of fore legs in ventral view (48), pterothorax in ventral view (49), mesoventrite and anterior region of metaventrite in ventral view (50) (arrowheads indicate foveae)." captionText-2="FIGURES 51–55. Thaumastocephalus sp., male from Croatia. elytra and mesonotum in dorsal view (51), anterior region of elytra and mesonotum in dorsal view (52), abdomen in ventral view (53), abdominal tergites III and IV in dorsal view (54), abdominal sternites III and IV in ventral view (55) (arrowheads indicate foveae)." figureDoi-0="http://doi.org/10.5281/zenodo.7891934" figureDoi-1="http://doi.org/10.5281/zenodo.7891936" figureDoi-2="http://doi.org/10.5281/zenodo.7891938" httpUri-0="https://zenodo.org/record/7891934/files/figure.png" httpUri-1="https://zenodo.org/record/7891936/files/figure.png" httpUri-2="https://zenodo.org/record/7891938/files/figure.png" pageId="12" pageNumber="83">Figs 41–55</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,572,508,532]" pageId="12" pageNumber="83" type="reference_group">
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<paragraph blockId="12.[151,572,508,532]" box="[151,572,508,532]" pageId="12" pageNumber="83">
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<treatmentCitationGroup box="[151,572,508,532]" pageId="12" pageNumber="83">
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<treatmentCitation author="Poggi, R. & Nonveiller, G. & Colla, A. & Pavicevic, D. & Rada, T." box="[151,568,508,532]" page="3" pageId="12" pageNumber="83" year="2001">
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<taxonomicName ID-CoL="8XQBS" authority="Poggi et al., 2001: 3" authorityName="Poggi" authorityPageNumber="3" authorityYear="2001" box="[151,568,508,532]" class="Insecta" family="Staphylinidae" genus="Thaumastocephalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="83" phylum="Arthropoda" rank="genus">
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<emphasis box="[151,354,508,532]" italics="true" pageId="12" pageNumber="83">Thaumastocephalus</emphasis>
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<bibRefCitation author="Poggi, R. & Nonveiller, G. & Colla, A. & Pavicevic, D. & Rada, T." box="[360,568,508,532]" pageId="12" pageNumber="83" pagination="1 - 20" refId="ref10503" refString="Poggi, R., Nonveiller, G., Colla, A., Pavicevic, D. & Rada, T. (2001) Thaumastocephalini, a new tribe of Pselaphinae for Thaumastocephalus folliculipalpus n. gen., n. sp., an interesting new troglobious species from central Dalmatia (Croatia) (Coleoptera, Staphylinidae, Pselaphinae). Annali del Museo Civico di Storia Naturale " Giacomo Doria ", 54, 1 - 20." type="journal article" year="2001">
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Poggi
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<emphasis box="[426,482,508,532]" italics="true" pageId="12" pageNumber="83">et al.,</emphasis>
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2001: 3
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</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="12" pageNumber="83" type="diagnosis">
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<paragraph blockId="12.[151,1437,577,2008]" pageId="12" pageNumber="83">
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<emphasis bold="true" box="[151,374,577,603]" pageId="12" pageNumber="83">Revised diagnosis.</emphasis>
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<taxonomicName authorityName="Jałoszyński & Hlaváč & Anlaş" authorityYear="2023" box="[386,613,578,604]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="71" phylum="Arthropoda" rank="tribe" tribe="Thaumastocephalini">Thaumastocephalini</taxonomicName>
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with frontal sulcus with duffuse margin; head behind frontal rostrum abruptly broadening posteriorly in step-wise manner; tempora gradually confluent with neck region; posterior tentorial pits situated closer to submental bulge than to neck region; maxillary palpomere 4 with concave distal margin; pronotum barrel-shaped, lacking lateral foveae and lateral longitudinal sulci; median longitudinal sulcus lacking or faint; mesoscutellar shield strongly elongate; metaventrite with two pairs of foveae (lateral to mesocoxal insertions and just posterior to mesocoxal rests); elytra lacking basal foveae and striae; tergite IV lacking discal carinae, with two pairs of minute lateral foveae; sternite IV lacking pair of submedian foveae.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="15" lastPageNumber="86" pageId="12" pageNumber="83" type="description">
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<paragraph blockId="12.[151,1437,577,2008]" pageId="12" pageNumber="83">
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<emphasis bold="true" box="[199,373,829,855]" pageId="12" pageNumber="83">Redescription.</emphasis>
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Body (
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<figureCitation box="[456,538,830,856]" captionStart="FIGURES 41–44" captionStartId="13.[151,264,1540,1564]" captionTargetBox="[161,1436,328,1505]" captionTargetId="figure-77@13.[151,1436,309,1517]" captionTargetPageId="13" captionText="FIGURES 41–44. Thaumastocephalus sp., male from Croatia. Dorsal habitus (41), head in dorsal view (42), prothorax and elytral base in dorsal view (43), anterior body region in lateral view (44)." figureDoi="http://doi.org/10.5281/zenodo.7891934" httpUri="https://zenodo.org/record/7891934/files/figure.png" pageId="12" pageNumber="83">Fig. 41</figureCitation>
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) elongate, slender, moderately convex, with distinct constrictions between head and prothorax and between prothorax and elytra.
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</paragraph>
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<paragraph blockId="12.[151,1437,577,2008]" pageId="12" pageNumber="83">
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Head (
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<figureCitation box="[274,490,902,928]" captionStart-0="FIGURES 41–44" captionStart-1="FIGURES 45–50" captionStartId-0="13.[151,264,1540,1564]" captionStartId-1="14.[151,264,1679,1703]" captionTargetBox-0="[161,1436,328,1505]" captionTargetBox-1="[151,1436,634,1644]" captionTargetId-0="figure-77@13.[151,1436,309,1517]" captionTargetId-1="figure-191@14.[151,1436,633,1656]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="FIGURES 41–44. Thaumastocephalus sp., male from Croatia. Dorsal habitus (41), head in dorsal view (42), prothorax and elytral base in dorsal view (43), anterior body region in lateral view (44)." captionText-1="FIGURES 45–50. Thaumastocephalus sp., male from Croatia. Head in ventral (45) and lateral view (46), antenna in lateral view (47), prothorax and proximal region of fore legs in ventral view (48), pterothorax in ventral view (49), mesoventrite and anterior region of metaventrite in ventral view (50) (arrowheads indicate foveae)." figureDoi-0="http://doi.org/10.5281/zenodo.7891934" figureDoi-1="http://doi.org/10.5281/zenodo.7891936" httpUri-0="https://zenodo.org/record/7891934/files/figure.png" httpUri-1="https://zenodo.org/record/7891936/files/figure.png" pageId="12" pageNumber="83">Figs 41, 42, 44‒46</figureCitation>
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) distinctly elongate, brodest near middle or slightly anteriorly, strongly flattened in posterior half. Temples rounded; vertex and frons confluent, lacking frontal fovea; dorsal tentorial pits very shallow but accentuated by dark spots marking invagination sites of dorsal tentorial arms, head with broad and shallow U-shaped vertexal-frontal sulcus with diffuse margins; compound eyes lacking. Gular sutures lacking; posterior tentorial pits large, circular, widely separated, situated far anterior to transverse impression demarcating neck region ventrally; submental region strongly transverse, with prominent oval submental bulge strongly projecting ventrally and slightly constricted at base, situated in posterior submental region, close to posterior tentorial pits; hypostomal ridge vestigial.
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</paragraph>
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<paragraph blockId="12.[151,1437,577,2008]" pageId="12" pageNumber="83">
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Antennae (
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<figureCitation box="[324,408,1190,1216]" captionStart="FIGURES 45–50" captionStartId="14.[151,264,1679,1703]" captionTargetBox="[151,1436,634,1644]" captionTargetId="figure-191@14.[151,1436,633,1656]" captionTargetPageId="14" captionText="FIGURES 45–50. Thaumastocephalus sp., male from Croatia. Head in ventral (45) and lateral view (46), antenna in lateral view (47), prothorax and proximal region of fore legs in ventral view (48), pterothorax in ventral view (49), mesoventrite and anterior region of metaventrite in ventral view (50) (arrowheads indicate foveae)." figureDoi="http://doi.org/10.5281/zenodo.7891936" httpUri="https://zenodo.org/record/7891936/files/figure.png" pageId="12" pageNumber="83">Fig. 47</figureCitation>
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) composed of eleven antennomeres, slender, antennal insertions broadly separated. Scape cylindrical and elongate, longer than pedicel, antennal club trimerous and very indistinctly demarcated, antennomere 11 elongate, symmetric, rapidly narrowing in distal region to pointed apex, lacking cavities and clusters of basiconic sensilla.
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</paragraph>
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<paragraph blockId="12.[151,1437,577,2008]" pageId="12" pageNumber="83">Labium with elongate mentum which is inversely subtrapezoidal, broadest near anterior third, with sinuate sides; prementum short, transverse, lacking ligula; labial palpi trimerous, palpomere 1 short, nearly annular, palpomere 2 strongly elongate and weakly clavate, palpomere 3 setiform, barely discernible, obscured by long outer apical seta of palpomere 2.</paragraph>
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<paragraph blockId="12.[151,1437,577,2008]" pageId="12" pageNumber="83">
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Maxilla with large, nearly semicircular cardo; basistipes triangular, elongate, mediostipes subtriangular, about as long as basistipes; galea elongate and with fringe of dense, thick setae along mesal margin; lacinia similar to galea but shorter; palpifer elongate. Maxillary palpi (
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<figureCitation box="[748,830,1550,1577]" captionStart="FIGURES 45–50" captionStartId="14.[151,264,1679,1703]" captionTargetBox="[151,1436,634,1644]" captionTargetId="figure-191@14.[151,1436,633,1656]" captionTargetPageId="14" captionText="FIGURES 45–50. Thaumastocephalus sp., male from Croatia. Head in ventral (45) and lateral view (46), antenna in lateral view (47), prothorax and proximal region of fore legs in ventral view (48), pterothorax in ventral view (49), mesoventrite and anterior region of metaventrite in ventral view (50) (arrowheads indicate foveae)." figureDoi="http://doi.org/10.5281/zenodo.7891936" httpUri="https://zenodo.org/record/7891936/files/figure.png" pageId="12" pageNumber="83">Fig. 45</figureCitation>
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) only slightly shorter than head, palpomere 1 minute, elongate, about twice as long as wide; palpomere 2 pedunculate, rapidly and strongly brodening in distal half, distal region strongly transverse and projecting outwards, with lateral sensory process inserted on conspicuously long and slender petiole broadened distally to form short collar bearing large sphaerical structure sparsely covered with flattened setae; palpomere 3 non-pedunculate, strongly transverse and projecting outwards, with similar sensory process as palpomere 2; palpomere 4 strongly transverse and projecting inwards, conspicuosly curved, with concave anterior and convex posterior margin, wider than each of palpomeres 2 and 3, with apical sensory appendage rod-like, short and slender, pointed at apex, inserted in socket surrounded by distinctly brodened and raised margin.
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</paragraph>
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<paragraph blockId="12.[151,1437,577,2008]" pageId="12" pageNumber="83">Mandibles subtriangular, slightly convex dorsally and concave ventrally, large, slightly asymmetric, differing in shapes and numbers of mesal row of several teeth gradually reducing in length towards mandibular base, outer mandibular margin with conspicuously long seta; prostheca and mola lacking.</paragraph>
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<paragraph blockId="12.[151,1437,577,2008]" pageId="12" pageNumber="83">Labrum transverse, subtrapezoidal, broadest anteriorly, with nearly straight anterior and rounded lateral margins; anteroventral margin with one pair of short peg-like sensilla inserted on anterior epipharyngeal region.</paragraph>
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<paragraph blockId="13.[151,1437,151,285]" pageId="13" pageNumber="84">
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Pronotum (
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<figureCitation box="[329,415,151,177]" captionStart="FIGURES 41–44" captionStartId="13.[151,264,1540,1564]" captionTargetBox="[161,1436,328,1505]" captionTargetId="figure-77@13.[151,1436,309,1517]" captionTargetPageId="13" captionText="FIGURES 41–44. Thaumastocephalus sp., male from Croatia. Dorsal habitus (41), head in dorsal view (42), prothorax and elytral base in dorsal view (43), anterior body region in lateral view (44)." figureDoi="http://doi.org/10.5281/zenodo.7891934" httpUri="https://zenodo.org/record/7891934/files/figure.png" pageId="13" pageNumber="84">Fig. 43</figureCitation>
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) widest near middle, barrel-shaped, strongly narrowing both anteriorly and posteriorly; lateral margins strongly rounded, anterior pronotal margin subequal in width to posterior one, anterior and posterior pronotal corners strongly obtuse-angled; pronotal base weakly arcuate. Pronotal disc lacking foveae and sulci or with shallow vestiges of asetose median antebasal fovea and faint trace of median longitudinal sulcus.
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7891934" ID-Zenodo-Dep="7891934" httpUri="https://zenodo.org/record/7891934/files/figure.png" pageId="13" pageNumber="84" startId="13.[151,264,1540,1564]" targetBox="[161,1436,328,1505]" targetPageId="13" targetType="figure">
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<paragraph blockId="13.[151,1437,1540,1601]" pageId="13" pageNumber="84">
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<emphasis bold="true" box="[151,342,1540,1564]" pageId="13" pageNumber="84">FIGURES 41–44.</emphasis>
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<taxonomicName box="[350,592,1540,1564]" class="Insecta" family="Staphylinidae" genus="Thaumastocephalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="84" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis box="[350,553,1540,1564]" italics="true" pageId="13" pageNumber="84">Thaumastocephalus</emphasis>
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sp.
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</taxonomicName>
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, male from Croatia. Dorsal habitus (
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<emphasis bold="true" box="[975,1000,1540,1564]" pageId="13" pageNumber="84">41</emphasis>
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), head in dorsal view (
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<emphasis bold="true" box="[1244,1269,1540,1564]" pageId="13" pageNumber="84">42</emphasis>
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), prothorax and elytral base in dorsal view (
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<emphasis bold="true" box="[432,457,1576,1600]" pageId="13" pageNumber="84">43</emphasis>
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), anterior body region in lateral view (
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<emphasis bold="true" box="[851,876,1576,1600]" pageId="13" pageNumber="84">44</emphasis>
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).
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</paragraph>
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</caption>
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<paragraph blockId="13.[151,1437,1647,2033]" pageId="13" pageNumber="84">
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Prosternum (
|
||
<figureCitation box="[346,429,1647,1673]" captionStart="FIGURES 45–50" captionStartId="14.[151,264,1679,1703]" captionTargetBox="[151,1436,634,1644]" captionTargetId="figure-191@14.[151,1436,633,1656]" captionTargetPageId="14" captionText="FIGURES 45–50. Thaumastocephalus sp., male from Croatia. Head in ventral (45) and lateral view (46), antenna in lateral view (47), prothorax and proximal region of fore legs in ventral view (48), pterothorax in ventral view (49), mesoventrite and anterior region of metaventrite in ventral view (50) (arrowheads indicate foveae)." figureDoi="http://doi.org/10.5281/zenodo.7891936" httpUri="https://zenodo.org/record/7891936/files/figure.png" pageId="13" pageNumber="84">Fig. 48</figureCitation>
|
||
) fused with hypomera, but with short grooves on anterior margin representing probable rudiments of notosternal sutures; hypomera broad and posteriorly convex or indistinctly flattened, each with only trace of barely discernible hypomeral groove; basisternal region much longer than very short coxal region (= furcasternum); prosternum with one pair of submedian posterior foveae near middle of posterior basisternal margin, foveal pockets directed dorsally and slightly outwards from body midline (
|
||
<figureCitation box="[1105,1191,1791,1817]" captionStart="FIGURES 45–50" captionStartId="14.[151,264,1679,1703]" captionTargetBox="[151,1436,634,1644]" captionTargetId="figure-191@14.[151,1436,633,1656]" captionTargetPageId="14" captionText="FIGURES 45–50. Thaumastocephalus sp., male from Croatia. Head in ventral (45) and lateral view (46), antenna in lateral view (47), prothorax and proximal region of fore legs in ventral view (48), pterothorax in ventral view (49), mesoventrite and anterior region of metaventrite in ventral view (50) (arrowheads indicate foveae)." figureDoi="http://doi.org/10.5281/zenodo.7891936" httpUri="https://zenodo.org/record/7891936/files/figure.png" pageId="13" pageNumber="84">Fig. 48</figureCitation>
|
||
; arrowhead); anterior prosternal margin smooth; prosternal process indistinct, not separating procoxae. Hypomera with hypomeral ridges demarcating conspicuously narrow, parallel-sided inner (adcoxal) regions.
|
||
</paragraph>
|
||
<paragraph blockId="13.[151,1437,1647,2033]" pageId="13" pageNumber="84">
|
||
Mesonotum (
|
||
<figureCitation box="[345,466,1898,1925]" captionStart="FIGURES 51–55" captionStartId="15.[151,264,1925,1949]" captionTargetBox="[160,1436,713,1902]" captionTargetId="figure-238@15.[151,1436,705,1902]" captionTargetPageId="15" captionText="FIGURES 51–55. Thaumastocephalus sp., male from Croatia. elytra and mesonotum in dorsal view (51), anterior region of elytra and mesonotum in dorsal view (52), abdomen in ventral view (53), abdominal tergites III and IV in dorsal view (54), abdominal sternites III and IV in ventral view (55) (arrowheads indicate foveae)." figureDoi="http://doi.org/10.5281/zenodo.7891938" httpUri="https://zenodo.org/record/7891938/files/figure.png" pageId="13" pageNumber="84">Figs51, 52</figureCitation>
|
||
) with mesoscutum and mesoscutellum fused and forming strongly elongate subtriangular scutellar shield, only its minute capitate tip projects posteriorly beyond basal articulating lobe of elytra, so that scutellar shield is barely visible between elytral bases in intact specimens.
|
||
</paragraph>
|
||
<paragraph blockId="13.[151,1437,1647,2033]" box="[199,1154,2007,2033]" pageId="13" pageNumber="84">Metanotum vestigial, strongly shortened and weakly sclerotized due to loss of wings.</paragraph>
|
||
<paragraph blockId="14.[151,1437,150,609]" pageId="14" pageNumber="85">
|
||
Mesoventrite (
|
||
<figureCitation box="[364,497,150,177]" captionStart="FIGURES 45–50" captionStartId="14.[151,264,1679,1703]" captionTargetBox="[151,1436,634,1644]" captionTargetId="figure-191@14.[151,1436,633,1656]" captionTargetPageId="14" captionText="FIGURES 45–50. Thaumastocephalus sp., male from Croatia. Head in ventral (45) and lateral view (46), antenna in lateral view (47), prothorax and proximal region of fore legs in ventral view (48), pterothorax in ventral view (49), mesoventrite and anterior region of metaventrite in ventral view (50) (arrowheads indicate foveae)." figureDoi="http://doi.org/10.5281/zenodo.7891936" httpUri="https://zenodo.org/record/7891936/files/figure.png" pageId="14" pageNumber="85">Figs 49, 50</figureCitation>
|
||
) broadening posteriorly, with massive prepectus; mesoventrite with three pairs of foveae in its anterior half (
|
||
<figureCitation box="[456,540,186,213]" captionStart="FIGURES 45–50" captionStartId="14.[151,264,1679,1703]" captionTargetBox="[151,1436,634,1644]" captionTargetId="figure-191@14.[151,1436,633,1656]" captionTargetPageId="14" captionText="FIGURES 45–50. Thaumastocephalus sp., male from Croatia. Head in ventral (45) and lateral view (46), antenna in lateral view (47), prothorax and proximal region of fore legs in ventral view (48), pterothorax in ventral view (49), mesoventrite and anterior region of metaventrite in ventral view (50) (arrowheads indicate foveae)." figureDoi="http://doi.org/10.5281/zenodo.7891936" httpUri="https://zenodo.org/record/7891936/files/figure.png" pageId="14" pageNumber="85">Fig. 50</figureCitation>
|
||
, arrowheads): one sublateral fovea directed laterally with ventral opening, one lateral fovea directed anteromesally, and one lateral fovea directed mesally, lateral foveae share one large laterally visible setose opening (
|
||
<figureCitation box="[411,494,259,285]" captionStart="FIGURES 41–44" captionStartId="13.[151,264,1540,1564]" captionTargetBox="[161,1436,328,1505]" captionTargetId="figure-77@13.[151,1436,309,1517]" captionTargetPageId="13" captionText="FIGURES 41–44. Thaumastocephalus sp., male from Croatia. Dorsal habitus (41), head in dorsal view (42), prothorax and elytral base in dorsal view (43), anterior body region in lateral view (44)." figureDoi="http://doi.org/10.5281/zenodo.7891934" httpUri="https://zenodo.org/record/7891934/files/figure.png" pageId="14" pageNumber="85">Fig. 44</figureCitation>
|
||
, upper arrowhead). Median subtrapezoidal area of mesoventrite demarcated laterally by complete ridges. Mesocoxae narrowly separated by elongate, subtriangular mesoventral process with rounded apex reaching middle of mesocoxae, meeting, but not fused with anterior tip of anterior metaventral process, both processes flat. Division between meso- and metaventrite marked with narrow sutures lateral to mesocoxal cavities, but division obliterated within mesocoxal rests.
|
||
</paragraph>
|
||
<paragraph blockId="14.[151,1437,150,609]" pageId="14" pageNumber="85">
|
||
Metaventrite (
|
||
<figureCitation box="[360,445,439,465]" captionStart="FIGURES 45–50" captionStartId="14.[151,264,1679,1703]" captionTargetBox="[151,1436,634,1644]" captionTargetId="figure-191@14.[151,1436,633,1656]" captionTargetPageId="14" captionText="FIGURES 45–50. Thaumastocephalus sp., male from Croatia. Head in ventral (45) and lateral view (46), antenna in lateral view (47), prothorax and proximal region of fore legs in ventral view (48), pterothorax in ventral view (49), mesoventrite and anterior region of metaventrite in ventral view (50) (arrowheads indicate foveae)." figureDoi="http://doi.org/10.5281/zenodo.7891936" httpUri="https://zenodo.org/record/7891936/files/figure.png" pageId="14" pageNumber="85">Fig. 49</figureCitation>
|
||
) strongly transverse and broadening posteriorly; posterior margins of mesocoxal rests carinate; metaventrite with pair of lateral setose foveae directed anteromesally just lateral to mesocoxal insertion (
|
||
<figureCitation captionStart="FIGURES 45–50" captionStartId="14.[151,264,1679,1703]" captionTargetBox="[151,1436,634,1644]" captionTargetId="figure-191@14.[151,1436,633,1656]" captionTargetPageId="14" captionText="FIGURES 45–50. Thaumastocephalus sp., male from Croatia. Head in ventral (45) and lateral view (46), antenna in lateral view (47), prothorax and proximal region of fore legs in ventral view (48), pterothorax in ventral view (49), mesoventrite and anterior region of metaventrite in ventral view (50) (arrowheads indicate foveae)." figureDoi="http://doi.org/10.5281/zenodo.7891936" httpUri="https://zenodo.org/record/7891936/files/figure.png" pageId="14" pageNumber="85">Fig. 50</figureCitation>
|
||
; upper arrowhead), and pair of setose foveae directed anteromesally just behind posterior margins of mesocoxal rests (
|
||
<figureCitation box="[218,301,546,573]" captionStart="FIGURES 45–50" captionStartId="14.[151,264,1679,1703]" captionTargetBox="[151,1436,634,1644]" captionTargetId="figure-191@14.[151,1436,633,1656]" captionTargetPageId="14" captionText="FIGURES 45–50. Thaumastocephalus sp., male from Croatia. Head in ventral (45) and lateral view (46), antenna in lateral view (47), prothorax and proximal region of fore legs in ventral view (48), pterothorax in ventral view (49), mesoventrite and anterior region of metaventrite in ventral view (50) (arrowheads indicate foveae)." figureDoi="http://doi.org/10.5281/zenodo.7891936" httpUri="https://zenodo.org/record/7891936/files/figure.png" pageId="14" pageNumber="85">Fig. 50</figureCitation>
|
||
; lower arrowhead); metaventral intermetacoxal process broad and clearly separating metacoxae, with posteromedian emargination.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7891936" ID-Zenodo-Dep="7891936" httpUri="https://zenodo.org/record/7891936/files/figure.png" pageId="14" pageNumber="85" startId="14.[151,264,1679,1703]" targetBox="[151,1436,634,1644]" targetPageId="14" targetType="figure">
|
||
<paragraph blockId="14.[151,1437,1679,1776]" pageId="14" pageNumber="85">
|
||
<emphasis bold="true" box="[151,342,1679,1704]" pageId="14" pageNumber="85">FIGURES 45–50.</emphasis>
|
||
<taxonomicName box="[350,592,1680,1704]" class="Insecta" family="Staphylinidae" genus="Thaumastocephalus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="85" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis box="[350,553,1680,1704]" italics="true" pageId="14" pageNumber="85">Thaumastocephalus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
, male from Croatia. Head in ventral (
|
||
<emphasis bold="true" box="[985,1010,1680,1704]" pageId="14" pageNumber="85">45</emphasis>
|
||
) and lateral view (
|
||
<emphasis bold="true" box="[1210,1235,1680,1704]" pageId="14" pageNumber="85">46</emphasis>
|
||
), antenna in lateral view (
|
||
<emphasis bold="true" box="[216,241,1716,1740]" pageId="14" pageNumber="85">47</emphasis>
|
||
), prothorax and proximal region of fore legs in ventral view (
|
||
<emphasis bold="true" box="[875,900,1716,1740]" pageId="14" pageNumber="85">48</emphasis>
|
||
), pterothorax in ventral view (
|
||
<emphasis bold="true" box="[1215,1240,1716,1740]" pageId="14" pageNumber="85">49</emphasis>
|
||
), mesoventrite and anterior region of metaventrite in ventral view (
|
||
<emphasis bold="true" box="[634,659,1752,1776]" pageId="14" pageNumber="85">50</emphasis>
|
||
) (arrowheads indicate foveae).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="14.[151,1437,1822,2029]" box="[199,1425,1822,1849]" pageId="14" pageNumber="85">Metendosternite with broad and subquadrate lamellate stem and short lateral furcal arms diverging anteriorly.</paragraph>
|
||
<paragraph blockId="14.[151,1437,1822,2029]" pageId="14" pageNumber="85">
|
||
Elytra (
|
||
<figureCitation box="[285,419,1858,1885]" captionStart="FIGURES 51–55" captionStartId="15.[151,264,1925,1949]" captionTargetBox="[160,1436,713,1902]" captionTargetId="figure-238@15.[151,1436,705,1902]" captionTargetPageId="15" captionText="FIGURES 51–55. Thaumastocephalus sp., male from Croatia. elytra and mesonotum in dorsal view (51), anterior region of elytra and mesonotum in dorsal view (52), abdomen in ventral view (53), abdominal tergites III and IV in dorsal view (54), abdominal sternites III and IV in ventral view (55) (arrowheads indicate foveae)." figureDoi="http://doi.org/10.5281/zenodo.7891938" httpUri="https://zenodo.org/record/7891938/files/figure.png" pageId="14" pageNumber="85">Figs 51, 52</figureCitation>
|
||
) together subtrapezoidal, dorsally flattened, lacking humeral denticle, foveae and striae; posterior elytral margin slightly concave and lacking fringe of modified setae.
|
||
</paragraph>
|
||
<paragraph blockId="14.[151,1437,1822,2029]" lastBlockId="15.[151,1437,150,681]" lastPageId="15" lastPageNumber="86" pageId="14" pageNumber="85">
|
||
Abdomen (
|
||
<figureCitation box="[328,460,1930,1957]" captionStart="FIGURES 51–55" captionStartId="15.[151,264,1925,1949]" captionTargetBox="[160,1436,713,1902]" captionTargetId="figure-238@15.[151,1436,705,1902]" captionTargetPageId="15" captionText="FIGURES 51–55. Thaumastocephalus sp., male from Croatia. elytra and mesonotum in dorsal view (51), anterior region of elytra and mesonotum in dorsal view (52), abdomen in ventral view (53), abdominal tergites III and IV in dorsal view (54), abdominal sternites III and IV in ventral view (55) (arrowheads indicate foveae)." figureDoi="http://doi.org/10.5281/zenodo.7891938" httpUri="https://zenodo.org/record/7891938/files/figure.png" pageId="14" pageNumber="85">Figs 53–55</figureCitation>
|
||
) broadest near middle, weakly convex dorsally and strongly convex ventrally, with well-developed broad paratergites on segments IV–VI. Tergite IV distinctly longer than remaining dorsally visible tergites (i.e., V–VII) combined, with poorly marked transverse mediobasal impression and lacking discal carinae, with two pairs of minute basolateral foveae directed laterally (
|
||
<figureCitation box="[868,954,150,177]" captionStart="FIGURES 51–55" captionStartId="15.[151,264,1925,1949]" captionTargetBox="[160,1436,713,1902]" captionTargetId="figure-238@15.[151,1436,705,1902]" captionTargetPageId="15" captionText="FIGURES 51–55. Thaumastocephalus sp., male from Croatia. elytra and mesonotum in dorsal view (51), anterior region of elytra and mesonotum in dorsal view (52), abdomen in ventral view (53), abdominal tergites III and IV in dorsal view (54), abdominal sternites III and IV in ventral view (55) (arrowheads indicate foveae)." figureDoi="http://doi.org/10.5281/zenodo.7891938" httpUri="https://zenodo.org/record/7891938/files/figure.png" pageId="15" pageNumber="86">Fig. 54</figureCitation>
|
||
; arrowheads). Sternite III short and partly exposed only between metacoxae, with dense fringe of long setae along posterior margin; sternite IV about as long as remaining visible sternites (i.e., V–IX in male) combined, with one pair of lateral foveae directed mesally (
|
||
<figureCitation captionStart="FIGURES 51–55" captionStartId="15.[151,264,1925,1949]" captionTargetBox="[160,1436,713,1902]" captionTargetId="figure-238@15.[151,1436,705,1902]" captionTargetPageId="15" captionText="FIGURES 51–55. Thaumastocephalus sp., male from Croatia. elytra and mesonotum in dorsal view (51), anterior region of elytra and mesonotum in dorsal view (52), abdomen in ventral view (53), abdominal tergites III and IV in dorsal view (54), abdominal sternites III and IV in ventral view (55) (arrowheads indicate foveae)." figureDoi="http://doi.org/10.5281/zenodo.7891938" httpUri="https://zenodo.org/record/7891938/files/figure.png" pageId="15" pageNumber="86">Fig. 55</figureCitation>
|
||
; arrowhead). Male sternite IX with large and elongate median penial plate, lateral portions subtriangular and shorter than penial plate, weakly visible in intact specimens.
|
||
</paragraph>
|
||
<paragraph blockId="15.[151,1437,150,681]" pageId="15" pageNumber="86">Legs long and slender, all femora slightly clavate, tibiae and tarsi slender; tarsomere 1 minuscule, tarsomere 2 longer than tarsomere 3; tarsal claw only slightly shorter than tarsomere 3; all tarsi shorter than half length of tibiae.</paragraph>
|
||
<paragraph blockId="15.[151,1437,150,681]" pageId="15" pageNumber="86">
|
||
Aedeagus (illustrated in
|
||
<bibRefCitation author="Poggi, R. & Nonveiller, G. & Colla, A. & Pavicevic, D. & Rada, T." box="[475,667,439,465]" pageId="15" pageNumber="86" pagination="1 - 20" refId="ref10503" refString="Poggi, R., Nonveiller, G., Colla, A., Pavicevic, D. & Rada, T. (2001) Thaumastocephalini, a new tribe of Pselaphinae for Thaumastocephalus folliculipalpus n. gen., n. sp., an interesting new troglobious species from central Dalmatia (Croatia) (Coleoptera, Staphylinidae, Pselaphinae). Annali del Museo Civico di Storia Naturale " Giacomo Doria ", 54, 1 - 20." type="journal article" year="2001">
|
||
Poggi
|
||
<emphasis box="[548,604,439,465]" italics="true" pageId="15" pageNumber="86">et al.</emphasis>
|
||
2001
|
||
</bibRefCitation>
|
||
and
|
||
<bibRefCitation author="Hlavac, P. & Bregovic, P. & Jalzic, B." box="[723,931,439,465]" pageId="15" pageNumber="86" pagination="90 - 110" refId="ref9606" refString="Hlavac, P., Bregovic, P. & Jalzic, B. (2019) Endogean and Cavernicolous Coleoptera of the Balkans. XVIII. Strong radiation in caves of central Dinarides: seven new species of Thaumastocephalus Poggi et al., 2001 (Staphylinidae: Pselaphinae). Zootaxa, 4559 (1), 90 - 110. https: // doi. org / 10.11646 / zootaxa. 4559.1.3" type="journal article" year="2019">
|
||
Hlaváč
|
||
<emphasis box="[809,859,439,465]" italics="true" pageId="15" pageNumber="86">et al</emphasis>
|
||
. 2019
|
||
</bibRefCitation>
|
||
) elongate, with weakly sclerotized walls and asymmetrical median lobe bearing circular or oval dorsal diaphragm; endophallic sclerites elongate and asymmetric, parameres asymmetric, long and broad, with subapical and/or apical setae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="15" pageNumber="86" type="distribution">
|
||
<paragraph blockId="15.[151,1437,150,681]" box="[199,1236,546,573]" pageId="15" pageNumber="86">
|
||
<emphasis bold="true" box="[199,353,546,572]" pageId="15" pageNumber="86">Distribution.</emphasis>
|
||
Balkan Region: caves in central Dinarides (
|
||
<collectingCountry box="[846,933,547,573]" name="Croatia" pageId="15" pageNumber="86">Croatia</collectingCountry>
|
||
,
|
||
<collectingCountry box="[944,1224,547,573]" name="Bosnia and Herzegovina" pageId="15" pageNumber="86">Bosnia and Herzegovina</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="15.[151,1437,150,681]" pageId="15" pageNumber="86">
|
||
<emphasis bold="true" box="[199,407,582,608]" pageId="15" pageNumber="86">Species included.</emphasis>
|
||
<taxonomicName authority="Hlavac et al." authorityName="Hlavac" box="[417,934,583,609]" class="Insecta" family="Staphylinidae" genus="Thaumastocephalus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="species" species="bilandzijae">
|
||
<emphasis box="[417,775,583,609]" italics="true" pageId="15" pageNumber="86">Thaumastocephalus bilandzijae</emphasis>
|
||
Hlaváč
|
||
<emphasis box="[876,929,583,609]" italics="true" pageId="15" pageNumber="86">et al</emphasis>
|
||
.
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Hlavac et al." authorityName="Hlavac" box="[953,1225,583,609]" class="Insecta" family="Staphylinidae" genus="Thaumastocephalus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="species" species="dahnae">
|
||
<emphasis box="[953,1066,583,609]" italics="true" pageId="15" pageNumber="86">T. dahnae</emphasis>
|
||
Hlaváč
|
||
<emphasis box="[1167,1219,583,609]" italics="true" pageId="15" pageNumber="86">et al</emphasis>
|
||
.
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Poggi et al." authorityName="Poggi" class="Insecta" family="Staphylinidae" genus="Thaumastocephalus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="species" species="folliculipalpus">
|
||
<emphasis box="[1243,1436,583,609]" italics="true" pageId="15" pageNumber="86">T. folliculipalpus</emphasis>
|
||
Poggi
|
||
<emphasis box="[223,279,619,645]" italics="true" pageId="15" pageNumber="86">et al.</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Hlavac et al." authorityName="Hlavac" box="[293,528,619,645]" class="Insecta" family="Staphylinidae" genus="Thaumastocephalus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="species" species="kirini">
|
||
<emphasis box="[293,381,619,645]" italics="true" pageId="15" pageNumber="86">T. kirini</emphasis>
|
||
Hlaváč
|
||
<emphasis box="[473,522,619,645]" italics="true" pageId="15" pageNumber="86">et al</emphasis>
|
||
.
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Hlavac et al." authorityName="Hlavac" box="[542,801,619,645]" class="Insecta" family="Staphylinidae" genus="Thaumastocephalus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="species" species="marsici">
|
||
<emphasis box="[542,653,619,644]" italics="true" pageId="15" pageNumber="86">T. marsici</emphasis>
|
||
Hlaváč
|
||
<emphasis box="[746,795,619,645]" italics="true" pageId="15" pageNumber="86">et al</emphasis>
|
||
.
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Hlavac et al." authorityName="Hlavac" box="[815,1113,619,645]" class="Insecta" family="Staphylinidae" genus="Thaumastocephalus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="species" species="rujnicensis">
|
||
<emphasis box="[815,966,619,644]" italics="true" pageId="15" pageNumber="86">T. rujnicensis</emphasis>
|
||
Hlaváč
|
||
<emphasis box="[1058,1107,619,645]" italics="true" pageId="15" pageNumber="86">et al</emphasis>
|
||
.
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Hlavac et al." authorityName="Hlavac" box="[1127,1381,619,645]" class="Insecta" family="Staphylinidae" genus="Thaumastocephalus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="species" species="slavkoi">
|
||
<emphasis box="[1127,1234,619,645]" italics="true" pageId="15" pageNumber="86">T. slavkoi</emphasis>
|
||
Hlaváč
|
||
<emphasis box="[1326,1375,619,645]" italics="true" pageId="15" pageNumber="86">et al</emphasis>
|
||
.
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName authority="Hlavac et al." authorityName="Hlavac" box="[151,416,655,681]" class="Insecta" family="Staphylinidae" genus="Thaumastocephalus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="86" phylum="Arthropoda" rank="species" species="troglavi">
|
||
<emphasis box="[151,267,655,681]" italics="true" pageId="15" pageNumber="86">T. troglavi</emphasis>
|
||
Hlaváč
|
||
<emphasis box="[360,409,655,681]" italics="true" pageId="15" pageNumber="86">et al</emphasis>
|
||
.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |