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<document id="DCDD78CD64C564ECDE20B38D13BFD51A" ID-CLB-Dataset="30691" ID-DOI="10.11646/zootaxa.4382.2.3" ID-GBIF-Dataset="6dc55986-74fa-4ac0-bca6-b301f3831180" ID-ISSN="1175-5326" ID-Zenodo-Dep="1182167" ID-ZooBank="8E248E6F-FDA5-4829-9DD7-4CC0EFDB08B6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1519199964901" checkinUser="plazi" docAuthor="Okamoto, Makoto &amp; Gon, Ofer" docDate="2018" docId="03A2BF386E614064FF185E95147C9CDA" docLanguage="en" docName="zootaxa.4382.2.3.pdf" docOrigin="Zootaxa 4382 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Epigonus exodon Okamoto &amp; Motomura 2012" docType="treatment" docVersion="7" lastPageNumber="274" masterDocId="FF9BC7406E6D4069FF8F5C1A170D9F12" masterDocTitle="A review of the deepwater cardinalfish genus Epigonus (Perciformes: Epigonidae) of the Western Indian Ocean, with description of two new species" masterLastPageNumber="291" masterPageNumber="261" pageNumber="273" updateTime="1698523467852" updateUser="ExternalLinkService">
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<mods:title id="46C55DB6B41A5799F53F315A6F1AB332">A review of the deepwater cardinalfish genus Epigonus (Perciformes: Epigonidae) of the Western Indian Ocean, with description of two new species</mods:title>
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<mods:namePart id="10065ED60F998A3C0A57F17F376E7D4E">Okamoto, Makoto</mods:namePart>
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<mods:namePart id="B3ECFA6CEAD6A858D4C22D4081E82E86">Gon, Ofer</mods:namePart>
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<taxonomicName id="4C0B75AD6E614065FF185E9515D39DB8" ID-CoL="6FTJG" authority="Okamoto &amp; Motomura, 2012" authorityName="Okamoto &amp; Motomura" authorityYear="2012" box="[151,734,655,682]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="273" phylum="Chordata" rank="species" species="exodon">
<emphasis id="B97FD23C6E614065FF185E9515D39DB8" bold="true" box="[151,734,655,682]" pageId="12" pageNumber="273">
<emphasis id="B97FD23C6E614065FF185E9516679DBB" bold="true" box="[151,362,655,681]" italics="true" pageId="12" pageNumber="273">Epigonus exodon</emphasis>
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</emphasis>
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</paragraph>
<paragraph id="8BB40E2E6E614065FF185EA916939DFD" blockId="12.[151,823,655,751]" pageId="12" pageNumber="273">
(New English name: Buck-toothed Deepwater Cardinalfish) (
<figureCitation id="133012AB6E614065FF105ECC17E19DFD" box="[159,236,726,751]" captionStart="FIGURE 1" captionStartId="1.[151,250,1983,2005]" captionTargetBox="[376,1217,1316,1962]" captionTargetId="figure@1.[346,1242,1316,1975]" captionTargetPageId="1" captionText="FIGURE 1. Distribution of four species of the Epigonus oligolepis group and the E. telescopus group in the Western Indian Ocean." httpUri="https://zenodo.org/record/1182169/files/figure.png" pageId="12" pageNumber="273">Figs. 1</figureCitation>
,
<figureCitation id="133012AB6E614065FF755ECD16149DFD" box="[250,281,727,751]" captionStart="FIGURE 10" captionStartId="12.[151,250,1827,1849]" captionTargetBox="[188,1398,926,1805]" captionTargetId="figure@12.[188,1398,901,1805]" captionTargetPageId="12" captionText="FIGURE 10. Epigonus exodon, MNHN 1991-0155, holotype, 97.0 mm SL. Overall body (A); and dorsal (B) and ventral (C) views of tipof the lower jaw, arrows show the anteriorly projecting teeth." httpUri="https://zenodo.org/record/1182187/files/figure.png" pageId="12" pageNumber="273">10</figureCitation>
; Table 1, 2)
</paragraph>
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<paragraph id="8BB40E2E6E614065FF185F3A14CB9C41" blockId="12.[151,1436,798,851]" pageId="12" pageNumber="273">
<materialsCitation id="3B6304736E614065FF185F3A14CB9C41" ID-GBIF-Occurrence="1820370023" collectionCode="MNHN" country="Réunion" elevation="465" location="western South Indian Ocean" pageId="12" pageNumber="273" specimenCode="MNHN 1991-0155" specimenCount="2" typeStatus="holotype">
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<taxonomicName id="4C0B75AD6E614065FF185F3A15AB9C26" ID-CoL="6FTJG" authority="Okamoto &amp; Motomura, 2012: 85" authorityName="Okamoto &amp; Motomura" authorityPageNumber="85" authorityYear="2012" box="[151,678,798,821]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="273" phylum="Chordata" rank="species" species="exodon">
<emphasis id="B97FD23C6E614065FF185F3A164A9C26" box="[151,327,798,821]" italics="true" pageId="12" pageNumber="273">Epigonus exodon</emphasis>
<treatmentCitation id="0AAA283F6E614065FEC05F0415AB9C26" author="" box="[335,678,798,820]" page="85" pageId="12" pageNumber="273">
<bibRefCitation id="EF9A73DF6E614065FEC05F04158C9C26" box="[335,641,798,820]" pageId="12" pageNumber="273" refId="ref21276" refString="Okamoto, M., Bartsch, P. &amp; Motomura, H. (2012) Epigonus merleni, a junior synonym of Epigonus macrops (Actinopterygii: Perciformes: Epigonidae). Species Diversity, 17, 123 - 126. https: // doi. org / 10.12782 / sd. 17.2.123" type="journal article">Okamoto &amp; Motomura, 2012</bibRefCitation>
: 85
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, fig. 1 (original description;
<typeStatus id="54B0B08C6E614065FC5F5F0514F19C26" box="[976,1020,799,820]" pageId="12" pageNumber="273">type</typeStatus>
locality:
<collectingCountry id="F31C4EBE6E614065FBD15F0413BB9C26" box="[1118,1206,798,820]" name="Réunion" pageId="12" pageNumber="273">Réunion</collectingCountry>
,
<location id="8ED458F56E614065FB4D5F0516079C41" LSID="urn:lsid:plazi:treatment:03A2BF386E614064FF185E95147C9CDA:8ED458F56E614065FB4D5F0516079C41" country="Réunion" name="western South Indian Ocean" pageId="12" pageNumber="273">western South Indian Ocean</location>
, 21°03 S, 55°09 E,
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<quantity id="4CF3A3CB6E614065FE5E5F2715499C41" box="[465,580,829,851]" metricMagnitude="2" metricUnit="m" metricValue="4.65" metricValueMax="4.8" metricValueMin="4.5" pageId="12" pageNumber="273" unit="m" value="465.0" valueMax="480.0" valueMin="450.0">450480 m</quantity>
</elevation>
depth;
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:
<specimenCode id="DBADA6556E614065FD7A5F2414B69C41" box="[757,955,829,851]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="273">MNHN 1991-0155</specimenCode>
).
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</paragraph>
</subSubSection>
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<emphasis id="B97FD23C6E614065FF185B39162D982A" bold="true" box="[151,288,1827,1849]" pageId="12" pageNumber="273">FIGURE 10.</emphasis>
<taxonomicName id="4C0B75AD6E614065FEA85B3E16DA982B" authorityName="Okamoto &amp; Motomura" authorityYear="2012" box="[295,471,1827,1849]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="273" phylum="Chordata" rank="species" species="exodon">
<emphasis id="B97FD23C6E614065FEA85B3E16DA982B" box="[295,471,1827,1849]" italics="true" pageId="12" pageNumber="273">Epigonus exodon</emphasis>
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, MNHN 1991-0155, holotype, 97.0 mm SL. Overall body (A); and dorsal (B) and ventral (C) views of tip of the lower jaw, arrows show the anteriorly projecting teeth.
</paragraph>
</caption>
<subSubSection id="C3115DA56E614065FF185B9E169898F7" pageId="12" pageNumber="273" type="materials_examined">
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<materialsCitation id="3B6304736E614065FF185B9E15FC98D2" ID-GBIF-Occurrence="1820369962" collectingDate="1982-09-08" collectionCode="MNHN" country="Réunion" elevation="465" location="western South Indian Ocean" pageId="12" pageNumber="273" specimenCode="MNHN 1991-0155" specimenCount="1" typeStatus="holotype">
<emphasis id="B97FD23C6E614065FF185B9E168C988F" bold="true" box="[151,385,1924,1949]" pageId="12" pageNumber="273">Material examined.</emphasis>
<specimenCode id="DBADA6556E614065FE075B9F156D988F" box="[392,608,1924,1949]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="273">MNHN 1991-0155</specimenCode>
,
<typeStatus id="54B0B08C6E614065FDE35B9E15DC988F" box="[620,721,1924,1949]" pageId="12" pageNumber="273" type="holotype">holotype</typeStatus>
, 97.0 mm SL, male, 21°03 S, 55°09 E,
<collectingCountry id="F31C4EBE6E614065FB005B9E13FC988F" box="[1167,1265,1924,1949]" name="Réunion" pageId="12" pageNumber="273">Réunion</collectingCountry>
,
<location id="8ED458F56E614065FB735B9F163F98D2" LSID="urn:lsid:plazi:treatment:03A2BF386E614064FF185E95147C9CDA:8ED458F56E614065FB735B9F163F98D2" country="Réunion" name="western South Indian Ocean" pageId="12" pageNumber="273">western South Indian Ocean</location>
,
<elevation id="0026E91D6E614065FECE5BB216CE98D2" box="[321,451,1960,1985]" metricMagnitude="2" metricUnit="m" metricValue="4.65" metricValueMax="4.8" metricValueMin="4.5" pageId="12" pageNumber="273" unit="m" value="465.0" valueMax="480.0" valueMin="450.0">
<quantity id="4CF3A3CB6E614065FECE5BB216CE98D2" box="[321,451,1960,1985]" metricMagnitude="2" metricUnit="m" metricValue="4.65" metricValueMax="4.8" metricValueMin="4.5" pageId="12" pageNumber="273" unit="m" value="465.0" valueMax="480.0" valueMin="450.0">450480 m</quantity>
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depth,
<collectingDate id="EFF1D1066E614065FD935BB215FC98D2" box="[540,753,1960,1985]" pageId="12" pageNumber="273" value="1982-09-08">
<date id="FFB528EE6E614065FD935BB215FC98D2" box="[540,753,1960,1985]" pageId="12" pageNumber="273" value="1982-09-08">8 September 1982</date>
</collectingDate>
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;
<materialsCitation id="3B6304736E614065FC8F5BB3169C98F7" ID-GBIF-Occurrence="1820370037" collectingDate="1982-09-08" collectionCode="MNHN" country="Réunion" elevation="465" location="western South Indian Ocean" pageId="12" pageNumber="273" specimenCode="MNHN 2012-0134" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBADA6556E614065FC8F5BB314D698D2" box="[768,987,1960,1984]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="273">MNHN 2012-0134</specimenCode>
,
<typeStatus id="54B0B08C6E614065FC655BB3134098D3" box="[1002,1101,1961,1985]" pageId="12" pageNumber="273" type="paratype">paratype</typeStatus>
,
<quantity id="4CF3A3CB6E614065FBD45BB213CF98D2" box="[1115,1218,1960,1985]" metricMagnitude="-2" metricUnit="m" metricValue="6.02" pageId="12" pageNumber="273" unit="mm" value="60.2">60.2 mm</quantity>
SL, sex unknown, same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3115DA56E604064FF485C8D13399E0F" pageId="13" pageNumber="274" type="diagnosis">
<paragraph id="8BB40E2E6E604064FF485C8D13399E0F" blockId="13.[151,1437,151,968]" pageId="13" pageNumber="274">
<emphasis id="B97FD23C6E604064FF485C8D164C9FA2" bold="true" box="[199,321,151,176]" pageId="13" pageNumber="274">Diagnosis.</emphasis>
Dorsal-fin rays VII-I, 10; pectoral-fin rays 1617; total gill rakers 2628; vertebrae 10 + 15; pyloric caeca 7; pored lateral-line scales 3537 + 3; opercular spine absent; maxillary mustache-like processes absent; ribs on last abdominal vertebra absent; anteriorly projecting teeth present on each side of symphysis of lower jaw; tongue narrow; shallow V-shaped tooth patch present on rear two-thirds of tongue.
</paragraph>
</subSubSection>
<subSubSection id="C3115DA56E604064FF485D3D15729D96" pageId="13" pageNumber="274" type="description">
<paragraph id="8BB40E2E6E604064FF485D3D15729D96" blockId="13.[151,1437,151,968]" pageId="13" pageNumber="274">
<emphasis id="B97FD23C6E604064FF485D3D16D19E52" bold="true" box="[199,476,295,320]" pageId="13" pageNumber="274">Measurements (% SL)</emphasis>
; counts are given in Table 1. Head length 33.434.2; head width 14.016.9; head height 16.417.3; body depth 23.924.6; body width 13.114.3; caudal-peduncle depth 8.89.6; caudal-peduncle length 28.028.9; orbital diameter 14.515.5; interorbital width 9.09.6; postorbital length 10.413.0; upper-jaw length 14.114.8; lower-jaw length 14.515.8; snout length 7.17.8; pre-first dorsal-fin length 34.736.2; presecond dorsal-fin length 55.356.6; pre-pectoral-fin length 34.736.0; pre-pelvic-fin length 38.241.8; pre-anus length 55.356.3; pre-anal-fin length 60.963.8; first spine length on first dorsal fin 2.73.2; second spine length on first dorsal fin 14.915.4; third spine length on first dorsal fin 16.0; second dorsal-fin spine length 13.314.1; first anal-fin spine length 1.82.3; second anal-fin spine length 12.513.3; pelvic-fin spine length 13.614.2; first dorsal-fin base length 11.5; second dorsal-fin base length 11.412.5; anal-fin base length 10.510.6; pectoral-fin length 20.220.4; pelvic-fin length 8.99.0.
</paragraph>
</subSubSection>
<subSubSection id="C3115DA56E604064FF485E95147C9CDA" pageId="13" pageNumber="274" type="distribution">
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<emphasis id="B97FD23C6E604064FF485E95166D9DBA" bold="true" box="[199,352,655,680]" pageId="13" pageNumber="274">Distribution.</emphasis>
<collectingRegion id="49CFC0CC6E604064FEE85E9516C49DBA" box="[359,457,655,680]" country="France" name="Reunion" pageId="13" pageNumber="274">
<collectingCountry id="F31C4EBE6E604064FEE85E9516C49DBA" box="[359,457,655,680]" name="Réunion" pageId="13" pageNumber="274">Réunion</collectingCountry>
</collectingRegion>
, western
<collectingRegion id="49CFC0CC6E604064FDBB5E9515789DBA" box="[564,629,655,680]" country="Tanzania" name="Kusini Unguja" pageId="13" pageNumber="274">South</collectingRegion>
Indian Ocean, at
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depth (
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) (
<figureCitation id="133012AB6E604064FAE05E9517A99DDE" captionStart="FIGURE 1" captionStartId="1.[151,250,1983,2005]" captionTargetBox="[376,1217,1316,1962]" captionTargetId="figure@1.[346,1242,1316,1975]" captionTargetPageId="1" captionText="FIGURE 1. Distribution of four species of the Epigonus oligolepis group and the E. telescopus group in the Western Indian Ocean." httpUri="https://zenodo.org/record/1182169/files/figure.png" pageId="13" pageNumber="274">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8BB40E2E6E604064FF485ECD147C9CDA" blockId="13.[151,1437,151,968]" pageId="13" pageNumber="274">
<emphasis id="B97FD23C6E604064FF485ECD15199DE2" bold="true" box="[199,532,727,752]" pageId="13" pageNumber="274">Comparisons and Remarks.</emphasis>
<taxonomicName id="4C0B75AD6E604064FD945EC215D19DE2" authorityName="Okamoto &amp; Motomura" authorityYear="2012" box="[539,732,728,752]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="274" phylum="Chordata" rank="species" species="exodon">
<emphasis id="B97FD23C6E604064FD945EC215D19DE2" box="[539,732,728,752]" italics="true" pageId="13" pageNumber="274">Epigonus exodon</emphasis>
</taxonomicName>
belongs the
<taxonomicName id="4C0B75AD6E604064FCE65EC314FB9DE2" authorityName="Mayer" authorityYear="1974" box="[873,1014,728,752]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="274" phylum="Chordata" rank="species" species="oligolepis">
<emphasis id="B97FD23C6E604064FCE65EC314FB9DE2" box="[873,1014,728,752]" italics="true" pageId="13" pageNumber="274">E. oligolepis</emphasis>
</taxonomicName>
group, defined as having less than 40 pored lateral-line scales (
<figureCitation id="133012AB6E604064FE395EE615129C06" box="[438,543,764,789]" captionStart="FIGURE 10" captionStartId="12.[151,250,1827,1849]" captionTargetBox="[188,1398,926,1805]" captionTargetId="figure@12.[188,1398,901,1805]" captionTargetPageId="12" captionText="FIGURE 10. Epigonus exodon, MNHN 1991-0155, holotype, 97.0 mm SL. Overall body (A); and dorsal (B) and ventral (C) views of tipof the lower jaw, arrows show the anteriorly projecting teeth." httpUri="https://zenodo.org/record/1182187/files/figure.png" pageId="13" pageNumber="274">Fig. 10A</figureCitation>
), lacking an opercular spine and ribs on the last abdominal vertebra. It can be easily distinguished from other congeners of the
<collectingRegion id="49CFC0CC6E604064FD405F3A14279C25" box="[719,810,800,823]" country="Papua New Guinea" name="Western" pageId="13" pageNumber="274">Western</collectingRegion>
Indian Ocean by having 3537 lateral-line scales and anteriorly projecting teeth on the symphysis of lower jaw (vs. more than 45 lateral-line scales and lacking projecting teeth on the symphysis of lower jaw in other congeners;
<taxonomicName id="4C0B75AD6E604064FC005F7313299C92" authorityName="Okamoto &amp; Gon" authorityYear="2018" box="[911,1060,872,896]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="274" phylum="Chordata" rank="species" species="bispinosus">
<emphasis id="B97FD23C6E604064FC005F7313299C92" box="[911,1060,872,896]" italics="true" pageId="13" pageNumber="274">E. bispinosus</emphasis>
</taxonomicName>
has two spines on the symphysis of lower jaw).
<bibRefCitation id="EF9A73DF6E604064FECF5F9615949CB6" box="[320,665,908,933]" pageId="13" pageNumber="274" refId="ref21276" refString="Okamoto, M., Bartsch, P. &amp; Motomura, H. (2012) Epigonus merleni, a junior synonym of Epigonus macrops (Actinopterygii: Perciformes: Epigonidae). Species Diversity, 17, 123 - 126. https: // doi. org / 10.12782 / sd. 17.2.123" type="journal article">Okamoto &amp; Motomura (2012)</bibRefCitation>
reported an ontogenetic change in the anteriorly projecting teeth of the lower jaw (
<typeStatus id="54B0B08C6E604064FEB15FAA16AE9CDA" box="[318,419,944,968]" pageId="13" pageNumber="274" type="paratype">paratype</typeStatus>
,
<quantity id="4CF3A3CB6E604064FE215FB5151F9CD5" box="[430,530,943,968]" metricMagnitude="-2" metricUnit="m" metricValue="6.02" pageId="13" pageNumber="274" unit="mm" value="60.2">60.2 mm</quantity>
SL, lacks the projecting teeth).
</paragraph>
</subSubSection>
</treatment>
</document>