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<document id="461E3270B1EA46FCF84C5F64806384B3" ID-DOI="10.5281/zenodo.155299" ID-GBIF-Dataset="22750794-cbfc-4d09-a4ae-bcdb17a35113" ID-Zenodo-Dep="155299" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1477431652181" checkinUser="plazi" docAuthor="Volker Mahnert &amp; Yun-chun Li" docDate="2016" docId="039980242835273BFC3999DBFD2FCA54" docLanguage="en" docName="7_Mahnert_Li.pdf" docOrigin="Revue suisse de Zoologie 123" docStyle="DocumentStyle:225056F0A23C51DDC06AC27EAEA15D5B.2:RevSuisseZool.2015-2019.journal_article" docStyleId="225056F0A23C51DDC06AC27EAEA15D5B" docStyleName="RevSuisseZool.2015-2019.journal_article" docStyleVersion="2" docTitle="Bisetocreagris cavernarum Mahnert &amp; Li, 2016, n. sp." docType="treatment" docUuid="848A7BC9-14E8-4144-8C50-75EB0F772FEF" docUuidSource="ZooBank" docVersion="11" lastPageNumber="262" masterDocId="FFA0F85C2837273EFF819F3AFFF7CB6F" masterDocTitle="Cave-inhabiting Neobisiidae (Arachnida: Pseudoscorpiones) from China, with description of four new species of Bisetocreagris Ćurčić" masterLastPageNumber="268" masterPageNumber="259" pageNumber="261" updateTime="1698390683281" updateUser="plazi">
<mods:mods id="85C4F044AC45159C686300F6AC3EF3CF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0C83A97CCB81901D4D5142F6D8DA7671">
<mods:title id="760D9F569BAB95306E68E4D97A24F0E3">Cave-inhabiting Neobisiidae (Arachnida: Pseudoscorpiones) from China, with description of four new species of Bisetocreagris Ćurčić</mods:title>
</mods:titleInfo>
<mods:name id="4563E9120E88CE8B69F3BEE0FEC9BF23" type="personal">
<mods:role id="6046FB7138EB3B8A8F3FD0F0EB349423">
<mods:roleTerm id="18DA1598C80C28B262B93C00B6151E37">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A79901F33CB1EF2857FE8E0D46CB39C4">Volker Mahnert</mods:namePart>
</mods:name>
<mods:name id="4529FE11292B75572F023094D967BD9D" type="personal">
<mods:role id="C8C148B5867C90452A92F9EC173736A6">
<mods:roleTerm id="CD16C979B423136507DDDA0765EC3AAB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E31781CEDD6792D737DC5B83CADF5482">Yun-chun Li</mods:namePart>
</mods:name>
<mods:typeOfResource id="46E4FB304E189A685CA739FA4311EDE2">text</mods:typeOfResource>
<mods:relatedItem id="E0BAB2DAB82661C176229F0E1BD00587" type="host">
<mods:titleInfo id="18E0975A8A88D4A37A8AAF96AB31F298">
<mods:title id="2ABA08BF413C1C1450CEF2E5C2C6F055">Revue suisse de Zoologie</mods:title>
</mods:titleInfo>
<mods:part id="3083704F057DB3D243C3C6205A36AF8C">
<mods:date id="166E70FE0C5D7D880E887E0D3AAAB560">2016</mods:date>
<mods:detail id="0121E60AE16A318893C911D3E72DE155" type="volume">
<mods:number id="78C76183320B030B2A9E2998997B0EB4">123</mods:number>
</mods:detail>
<mods:extent id="D734A09BBDBE4C382D3F33FAEDCF67B9" unit="page">
<mods:start id="0CCB77FEC6639C7A1EC6B5C0C3B8AB1D">259</mods:start>
<mods:end id="CD360EA0BB3291DD4EBD5F53ED0256EA">268</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="9A13D9B5E0217C7E5A2E93CD7F3565B7">journal article</mods:classification>
<mods:identifier id="EF16B0C062BEE94A4B6DF5B3FF450DA6" type="DOI">10.5281/zenodo.155299</mods:identifier>
<mods:identifier id="08697CECA4D3D78D62E33DB52A64BADB" type="GBIF-Dataset">22750794-cbfc-4d09-a4ae-bcdb17a35113</mods:identifier>
<mods:identifier id="59327B2B844E8B1BC47B17EED21B5134" type="Zenodo-Dep">155299</mods:identifier>
<mods:identifier id="CD78C4BD0F48A30A5E8E04911D189B79" type="ZooBank">7D59A703-571B-4802-BB25-79D016A0FE18</mods:identifier>
</mods:mods>
<treatment id="039980242835273BFC3999DBFD2FCA54" ID-DOI="http://doi.org/10.5281/zenodo.6080950" ID-GBIF-Taxon="124558069" ID-Zenodo-Dep="6080950" LSID="urn:lsid:zoobank.org:act:848A7BC9-14E8-4144-8C50-75EB0F772FEF" httpUri="http://treatment.plazi.org/id/039980242835273BFC3999DBFD2FCA54" lastPageId="5" lastPageNumber="262" pageId="2" pageNumber="261">
<subSubSection id="C32A62B92835273CFC3999DBFAD3CD95" box="[952,1316,1761,1786]" pageId="2" pageNumber="261" type="nomenclature">
<paragraph id="8B8F31322835273CFC3999DBFAD3CD95" blockId="2.[952,1316,1761,1817]" box="[952,1316,1761,1786]" pageId="2" pageNumber="261">
<heading id="D0C7865E2835273CFC3999DBFAD3CD95" bold="true" box="[952,1316,1761,1786]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="261" reason="4">
<emphasis id="B944ED202835273CFC3999DBFAD3CD95" bold="true" box="[952,1316,1761,1786]" pageId="2" pageNumber="261">
<taxonomicName id="4C304AB12835273CFC3999DBFB16CD97" ID-CoL="LWS4" box="[952,1249,1761,1785]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="261" phylum="Arthropoda" rank="species" species="cavernarum" status="sp. nov.">
<emphasis id="B944ED202835273CFC3999DBFB16CD97" bold="true" box="[952,1249,1761,1785]" italics="true" pageId="2" pageNumber="261">Bisetocreagris cavernarum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A277505B2835273CFB6999D8FAD3CD95" box="[1256,1316,1762,1786]" pageId="2" pageNumber="261" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32A62B92835273CFBBA9838FB57CC76" box="[1083,1184,1794,1817]" pageId="2" pageNumber="261" type="description">
<paragraph id="8B8F31322835273CFBBA9838FB57CC76" blockId="2.[952,1316,1761,1817]" box="[1083,1184,1794,1817]" pageId="2" pageNumber="261">
<figureCitation id="130B2DB72835273CFBBA9838FB57CC76" box="[1083,1184,1794,1817]" captionStart="Figs 1 - 10" captionStartId="4.[151,192,1851,1872]" captionTargetBox="[157,1430,264,1849]" captionTargetId="figure@4.[148,1439,254,1849]" captionTargetPageId="4" captionText="Figs 1 - 10. (1 - 5) Bisetocreagris chuanensis n. sp., holotype ♂. (1) Left chelicera. (2) “ spinneret ” (broken?) of ♂ (a) and of ♀ (b). (3) Rallum. (4 - 5) Pedipalp (vestitural setae of chela fingers omitted), trochanter to patella (4) and hand (5). (6 - 10) Bisetocreagris cavernarum n. sp., holotype ♂. (6) Left chelicera. (7) Galea of ♂ (a) and of ♀ (b). (8) Rallum. (9 - 10) Pedipalp (vestitural setae of chelal fingers omitted), trochanter to patella (9) and hand (10). Scales: 0.5 mm (4 - 5, 9 - 10); 0.2 mm (2 - 3, 7 - 8; 1; 6)." httpUri="https://zenodo.org/record/163238/files/figure.png" pageId="2" pageNumber="261">Figs 6-10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32A62B92835273CFCBE9809FAA9C372" pageId="2" pageNumber="261" type="materials_examined">
<paragraph id="8B8F31322835273CFCBE9809FADCCC82" blockId="2.[831,1437,1842,2030]" pageId="2" pageNumber="261">
<materialsCitation id="3B583B6F2835273CFCBE9809FA6ACCE4" ID-GBIF-Occurrence="1324024072" collectingDate="2015-10-08" collectionCode="MSWU" collectorName="Yun-chun Li." country="China" elevation="776" latitude="30.05539" location="Jin Dao Xia Town" longitude="106.62385" municipality="Bei Bei" pageId="2" pageNumber="261" specimenCount="1" specimenCount-male="1" stateProvince="Chongqing" typeStatus="holotype">
<emphasis id="B944ED202835273CFCBE9809FC47CC24" bold="true" box="[831,944,1843,1867]" pageId="2" pageNumber="261">
<typeStatus id="548B8F902835273CFCBE9809FC5CCC24" box="[831,939,1843,1867]" pageId="2" pageNumber="261" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="ED21A9F72835273CFC3B9809FBE7CC24" box="[954,1040,1843,1867]" collectionName="MSWU" pageId="2" pageNumber="261">MSWU</collectionCode>
;
<specimenCount id="9D36FABB2835273CFB9F9809FBC5CC25" box="[1054,1074,1843,1866]" pageId="2" pageNumber="261" type="male">♂</specimenCount>
;
<collectingCountry id="F32771A22835273CFBC29808FB72CC26" box="[1091,1157,1842,1865]" name="China" pageId="2" pageNumber="261">China</collectingCountry>
,
<collectingRegion id="49F4FFD02835273CFB139808FAFCCC26" box="[1170,1291,1842,1865]" country="China" name="Chongqing" pageId="2" pageNumber="261">Chongqing</collectingRegion>
,
<collectingMunicipality id="6BEBAB482835273CFA989808FA87CC26" box="[1305,1392,1842,1865]" pageId="2" pageNumber="261">Bei Bei</collectingMunicipality>
,
<location id="8EEF67E92835273CFAFC9808FC16CC04" LSID="urn:lsid:plazi:treatment:039980242835273BFC3999DBFD2FCA54:8EEF67E92835273CFAFC9808FC16CC04" country="China" municipality="Bei Bei" name="Jin Dao Xia Town" pageId="2" pageNumber="261" stateProvince="Chongqing">Jin Dao Xia Town</location>
,
<location id="8EEF67E92835273CFC6C986EFB78CC03" LSID="urn:lsid:plazi:treatment:039980242835273BFC3999DBFD2FCA54:8EEF67E92835273CFC6C986EFB78CC03" box="[1005,1167,1876,1900]" country="China" municipality="Bei Bei" name="Ert Long Cave" pageId="2" pageNumber="261" stateProvince="Chongqing">Ert Long Cave</location>
, alt.
<elevation id="001DD6012835273CFB449869FAFBCC04" box="[1221,1292,1875,1899]" metricMagnitude="2" metricUnit="m" metricValue="7.76" pageId="2" pageNumber="261" unit="m" value="776.0">
<quantity id="4CC89CD72835273CFB449869FAFBCC04" box="[1221,1292,1875,1899]" metricMagnitude="2" metricUnit="m" metricValue="7.76" pageId="2" pageNumber="261" unit="m" value="776.0">776 m</quantity>
</elevation>
,
<geoCoordinate id="EE0457F52835273CFA999869FC23CCE4" pageId="2" pageNumber="261">30°05539’N 106°62385’E</geoCoordinate>
;
<date id="FF8E17F22835273CFC6B984EFB5BCCE4" box="[1002,1196,1908,1932]" pageId="2" pageNumber="261" value="2015-10-08">
<collectingDate id="EFCAEE1A2835273CFC6B984EFB5BCCE4" box="[1002,1196,1908,1932]" pageId="2" pageNumber="261" value="2015-10-08">8 October 2015</collectingDate>
</date>
; coll.
<collectorName id="26C554E42835273CFA83984EFA6ACCE4" box="[1282,1437,1908,1931]" pageId="2" pageNumber="261">Yun-chun Li.</collectorName>
</materialsCitation>
The cave is about
<quantity id="4CC89CD72835273CFB9C98AFFB82CCC3" box="[1053,1141,1941,1965]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="261" unit="m" value="1200.0">1200 m</quantity>
long,
<quantity id="4CC89CD72835273CFB4798AFFB02CCC3" box="[1222,1269,1941,1964]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="261" unit="m" value="7.0">7 m</quantity>
high, with an underground river. Pseudoscorpions were collected by hand under rocks, together with some spiders.
</paragraph>
<paragraph id="8B8F31322835273CFCBE973CFAA9C372" blockId="2.[831,1374,2054,2078]" box="[831,1374,2054,2078]" pageId="2" pageNumber="261">
<emphasis id="B944ED202835273CFCBE973CFC47C371" bold="true" box="[831,944,2054,2078]" pageId="2" pageNumber="261">
<typeStatus id="548B8F902835273CFCBE973CFC5CC371" box="[831,939,2054,2078]" pageId="2" pageNumber="261" type="paratype">Paratype</typeStatus>
:
</emphasis>
<collectionCode id="ED21A9F72835273CFC36973CFBFBC371" box="[951,1036,2054,2078]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="2" pageNumber="261">MHNG</collectionCode>
;
<specimenCount id="9D36FABB2835273CFB97973CFBCFC371" box="[1046,1080,2054,2078]" pageId="2" pageNumber="261" type="female">1♀</specimenCount>
; same data as for
<typeStatus id="548B8F902835273CFB78973CFAADC372" box="[1273,1370,2054,2077]" pageId="2" pageNumber="261" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C32A62B92834273DFF169E38FD30C9D5" pageId="3" pageNumber="262" type="diagnosis">
<paragraph id="8B8F31322834273DFF169E38FD30C9D5" blockId="3.[151,756,258,698]" pageId="3" pageNumber="262">
<emphasis id="B944ED202834273DFF169E38FEE7CA76" bold="true" box="[151,272,258,281]" pageId="3" pageNumber="262">Diagnosis:</emphasis>
Troglobiomorphic habitus; carapace with 2 tiny plus 2 indistinct eyes; epistome absent; carapace with 6 setae on posterior margin, tergite I with 8-9 setae; teeth on movable cheliceral finger on thickened lamella, galea with several branches; pedipalp slender, smooth, excepted for fine granulation in distal half of hand; femur 5.8 times (length 1.92-2.1), patella 4.3- 4.7 times (length 1.78-2.01) longer than broad, pedicel (length 0.95-1.00) about half of total length of patella and as long as club, hand with pedicel 2.3 times (length 1.37-1.48), chela with pedicel 5.3-5.4 (length 3.27-3.51) times longer than broad, finger 1.4 (length 1.93-2.05) times longer than hand with pedicel. Trichobothrium
<emphasis id="B944ED202834273DFD579DB9FD03C9F5" box="[726,756,643,666]" italics="true" pageId="3" pageNumber="262">est</emphasis>
nearer to
<emphasis id="B944ED202834273DFF7D9D99FEF8C9D5" box="[252,271,675,698]" italics="true" pageId="3" pageNumber="262">et</emphasis>
than to
<emphasis id="B944ED202834273DFEE79D98FE72C9D6" box="[358,389,674,697]" italics="true" pageId="3" pageNumber="262">isb</emphasis>
,
<emphasis id="B944ED202834273DFE139D98FE5DC9D6" box="[402,426,674,697]" italics="true" pageId="3" pageNumber="262">sb</emphasis>
halfway between
<emphasis id="B944ED202834273DFDEC9D98FD8DC9D6" box="[621,634,674,697]" italics="true" pageId="3" pageNumber="262">b</emphasis>
and
<emphasis id="B944ED202834273DFD2F9D99FD37C9D5" box="[686,704,675,698]" italics="true" pageId="3" pageNumber="262">st</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C32A62B92834273DFF169DE8FD76C865" pageId="3" pageNumber="262" type="etymology">
<paragraph id="8B8F31322834273DFF169DE8FD76C865" blockId="3.[151,756,721,778]" pageId="3" pageNumber="262">
<emphasis id="B944ED202834273DFF169DE8FEEAC985" bold="true" box="[151,285,722,746]" pageId="3" pageNumber="262">Etymology:</emphasis>
From the Latin word caverna (= cave), genitive plural (meaning coming from caves).
</paragraph>
</subSubSection>
<subSubSection id="C32A62B92834273DFF169C1AFA72CD27" pageId="3" pageNumber="262" type="description">
<paragraph id="8B8F31322834273DFF169C1AFE37CDF8" blockId="3.[151,757,800,2073]" pageId="3" pageNumber="262">
<emphasis id="B944ED202834273DFF169C1AFD03C858" bold="true" box="[151,756,800,825]" pageId="3" pageNumber="262">
Description of
<typeStatus id="548B8F902834273DFECD9C1BFE58C856" box="[332,431,801,825]" pageId="3" pageNumber="262" type="holotype">holotype</typeStatus>
(
<typeStatus id="548B8F902834273DFE439C1AFDDCC858" box="[450,555,800,823]" pageId="3" pageNumber="262" type="paratype">paratype</typeStatus>
in parentheses):
</emphasis>
Pedipalps, chelicerae and anterior half of carapace yellowish red, carapace in basal half and tergites yellowish. Carapace smooth, 1.1 times longer than broad, anterior margin with a small rounded median knob, with 4 small eyes, 2 anterior eyes with indistinct lens, 1.5 diameters from anterior carapace margin, 2 posterior eyes indistinct and difficult to observe; with 27 (7-8-6- 6) (
<typeStatus id="548B8F902834273DFF3F9B1AFEE8CF58" box="[190,287,1056,1079]" pageId="3" pageNumber="262" type="holotype">holotype</typeStatus>
) to 32 (
<date id="FF8E17F22834273DFEFB9B1AFE20CF58" box="[378,471,1056,1079]" pageId="3" pageNumber="262" value="2011-08-07" valueMax="2011-08-06" valueMin="2011-08-07">8-11-7-6</date>
) (
<typeStatus id="548B8F902834273DFE719B1BFDB8CF56" box="[496,591,1057,1081]" pageId="3" pageNumber="262" type="paratype">paratype</typeStatus>
) setae; tergites undivided, chaetotaxy: 8/9/
<date id="FF8E17F22834273DFE559B7BFDC3CF36" box="[468,564,1089,1113]" pageId="3" pageNumber="262" value="2011-10-10">10/10/11</date>
/
<date id="FF8E17F22834273DFDBC9B7BFD6BCF36" box="[573,668,1089,1113]" pageId="3" pageNumber="262" value="2013-12-13">13/13/12</date>
/
<date id="FF8E17F22834273DFD249B7BFD03CF36" box="[677,756,1089,1113]" pageId="3" pageNumber="262" value="2013-08-11">13/11/8</date>
(4 tactile setae) (9/8/9/9/
<date id="FF8E17F22834273DFE419B5BFDD7CF16" box="[448,544,1121,1145]" pageId="3" pageNumber="262" value="2012-12-11">11/12/12</date>
/
<date id="FF8E17F22834273DFDA99B5BFD7DCF16" box="[552,650,1121,1145]" pageId="3" pageNumber="262" value="2012-10-13">13/10/12</date>
/8). Manducatory process with 5 marginal setae, pedipalpal coxa with 7 (
<specimenCount id="9D36FABB2834273DFEB99B9BFEBBCFD7" box="[312,332,1185,1208]" pageId="3" pageNumber="262">♀</specimenCount>
9-11) setae and 2 round lyrifissures (anteromedially, laterobasally), coxa
<date id="FF8E17F22834273DFDC49BFBFD9BCFB7" box="[581,620,1217,1240]" pageId="3" pageNumber="262">I 7</date>
(11) setae, lateral corner short and rounded, medial corner rectangular,
<date id="FF8E17F22834273DFEA59A3BFEA5CE76" box="[292,338,1281,1305]" pageId="3" pageNumber="262">II 9</date>
,
<date id="FF8E17F22834273DFEE49A3BFE6ECE78" box="[357,409,1280,1304]" pageId="3" pageNumber="262">III 5</date>
/6 (6),
<date id="FF8E17F22834273DFE719A3BFDC3CE76" box="[496,564,1281,1305]" pageId="3" pageNumber="262" value="2011-04">IV 11</date>
(12/13). Genital operculum with about 75 long acute setae, genital opening with 4/4 short acute interior setae, median genital sac undivided and short (reaching anterior half of sternite IV), wrinkled and with enlarged apex, lateral sacs wrinkled, dilated apically and bent anteriorly; sternite III with a small anteromedian groove (with 1/2 short setae at its margins) and with 22 setae (5 medial discal setae, not counting the tiny groove setae) (
<specimenCount id="9D36FABB2834273DFD2C993BFD36CD77" box="[685,705,1537,1560]" pageId="3" pageNumber="262">♀</specimenCount>
: 14) + 7/8 suprastigmal setae,
<date id="FF8E17F22834273DFE2F991BFE1ACD56" box="[430,493,1569,1593]" pageId="3" pageNumber="262" value="2012-04">IV 12</date>
(11) +
<date id="FF8E17F22834273DFDC0991AFDA9CD58" box="[577,606,1568,1591]" pageId="3" pageNumber="262">2x</date>
5, 16 (18)-17- 18 (16)-19 (17)-15 (16)-17(15)-6 (2 submedial tactile setae), submedial setae submarginally placed on VII-X. Pleural membrane granular.
</paragraph>
<paragraph id="8B8F31322834273DFF16999AFEE8CC98" blockId="3.[151,757,800,2073]" pageId="3" pageNumber="262">
<emphasis id="B944ED202834273DFF16999AFEF7CDD8" box="[151,256,1696,1719]" italics="true" pageId="3" pageNumber="262">Chelicera</emphasis>
(
<figureCitation id="130B2DB72834273DFE90999AFEA3CDD8" box="[273,340,1696,1719]" captionStart="Figs 1 - 10" captionStartId="4.[151,192,1851,1872]" captionTargetBox="[157,1430,264,1849]" captionTargetId="figure@4.[148,1439,254,1849]" captionTargetPageId="4" captionText="Figs 1 - 10. (1 - 5) Bisetocreagris chuanensis n. sp., holotype ♂. (1) Left chelicera. (2) “ spinneret ” (broken?) of ♂ (a) and of ♀ (b). (3) Rallum. (4 - 5) Pedipalp (vestitural setae of chela fingers omitted), trochanter to patella (4) and hand (5). (6 - 10) Bisetocreagris cavernarum n. sp., holotype ♂. (6) Left chelicera. (7) Galea of ♂ (a) and of ♀ (b). (8) Rallum. (9 - 10) Pedipalp (vestitural setae of chelal fingers omitted), trochanter to patella (9) and hand (10). Scales: 0.5 mm (4 - 5, 9 - 10); 0.2 mm (2 - 3, 7 - 8; 1; 6)." httpUri="https://zenodo.org/record/163238/files/figure.png" pageId="3" pageNumber="262">Fig. 6</figureCitation>
): Hand with 7 long, acute setae; fixed finger with about 16 small, acute, uniform teeth; movable finger with about 14 pointed teeth, the median ones on enlarged lamella, subgaleal seta reaching tip of galea. Galea (
<figureCitation id="130B2DB72834273DFF69981AFEDCCC57" box="[232,299,1824,1848]" captionStart="Figs 1 - 10" captionStartId="4.[151,192,1851,1872]" captionTargetBox="[157,1430,264,1849]" captionTargetId="figure@4.[148,1439,254,1849]" captionTargetPageId="4" captionText="Figs 1 - 10. (1 - 5) Bisetocreagris chuanensis n. sp., holotype ♂. (1) Left chelicera. (2) “ spinneret ” (broken?) of ♂ (a) and of ♀ (b). (3) Rallum. (4 - 5) Pedipalp (vestitural setae of chela fingers omitted), trochanter to patella (4) and hand (5). (6 - 10) Bisetocreagris cavernarum n. sp., holotype ♂. (6) Left chelicera. (7) Galea of ♂ (a) and of ♀ (b). (8) Rallum. (9 - 10) Pedipalp (vestitural setae of chelal fingers omitted), trochanter to patella (9) and hand (10). Scales: 0.5 mm (4 - 5, 9 - 10); 0.2 mm (2 - 3, 7 - 8; 1; 6)." httpUri="https://zenodo.org/record/163238/files/figure.png" pageId="3" pageNumber="262">Fig. 7</figureCitation>
a, b) divided in 2 main branches (
<specimenCount id="9D36FABB2834273DFD2A981BFD48CC57" box="[683,703,1825,1848]" pageId="3" pageNumber="262">♂</specimenCount>
: left galea with one main branch broken), each one forked apically; serrula exterior with about 44-48 blades, serrula interior with about 34, distal blade not forked. Rallum (
<figureCitation id="130B2DB72834273DFF1E989AFF12CCD6" box="[159,229,1952,1977]" captionStart="Figs 1 - 10" captionStartId="4.[151,192,1851,1872]" captionTargetBox="[157,1430,264,1849]" captionTargetId="figure@4.[148,1439,254,1849]" captionTargetPageId="4" captionText="Figs 1 - 10. (1 - 5) Bisetocreagris chuanensis n. sp., holotype ♂. (1) Left chelicera. (2) “ spinneret ” (broken?) of ♂ (a) and of ♀ (b). (3) Rallum. (4 - 5) Pedipalp (vestitural setae of chela fingers omitted), trochanter to patella (4) and hand (5). (6 - 10) Bisetocreagris cavernarum n. sp., holotype ♂. (6) Left chelicera. (7) Galea of ♂ (a) and of ♀ (b). (8) Rallum. (9 - 10) Pedipalp (vestitural setae of chelal fingers omitted), trochanter to patella (9) and hand (10). Scales: 0.5 mm (4 - 5, 9 - 10); 0.2 mm (2 - 3, 7 - 8; 1; 6)." httpUri="https://zenodo.org/record/163238/files/figure.png" pageId="3" pageNumber="262">Fig. 8</figureCitation>
) with 9 (8) pinnate setae, the first one with expanded base and separated from second one, proximal one shortest.
</paragraph>
<paragraph id="8B8F31322834273DFF16973AFAABC9B5" blockId="3.[151,757,800,2073]" lastBlockId="3.[831,1437,258,1145]" pageId="3" pageNumber="262">
<emphasis id="B944ED202834273DFF16973AFF00C378" box="[151,247,2048,2071]" italics="true" pageId="3" pageNumber="262">Pedipalp</emphasis>
(
<figureCitation id="130B2DB72834273DFE89973AFE81C376" box="[264,374,2048,2073]" captionStart="Figs 1 - 10" captionStartId="4.[151,192,1851,1872]" captionTargetBox="[157,1430,264,1849]" captionTargetId="figure@4.[148,1439,254,1849]" captionTargetPageId="4" captionText="Figs 1 - 10. (1 - 5) Bisetocreagris chuanensis n. sp., holotype ♂. (1) Left chelicera. (2) “ spinneret ” (broken?) of ♂ (a) and of ♀ (b). (3) Rallum. (4 - 5) Pedipalp (vestitural setae of chela fingers omitted), trochanter to patella (4) and hand (5). (6 - 10) Bisetocreagris cavernarum n. sp., holotype ♂. (6) Left chelicera. (7) Galea of ♂ (a) and of ♀ (b). (8) Rallum. (9 - 10) Pedipalp (vestitural setae of chelal fingers omitted), trochanter to patella (9) and hand (10). Scales: 0.5 mm (4 - 5, 9 - 10); 0.2 mm (2 - 3, 7 - 8; 1; 6)." httpUri="https://zenodo.org/record/163238/files/figure.png" pageId="3" pageNumber="262">Figs 9-10</figureCitation>
): Femur and patella smooth, distal part of hand dorsally and medially finely granular; trochanter 2.5 (2.7) times, with a distinct small ventral hump, femur 5.8 (5.8) times, patella 4.3 (4.7) times, club 2.0 (2-4) times longer than broad, pedicel as long as or longer than club, hand with pedicel 2.3 (2.3) times, chela with pedicel 5.4 (5.3) times, without pedicel 5.0 (4.9) times longer than broad, finger 1.4 (1.4) times longer than hand with pedicel. Fixed finger with 121 (122) small conical uniform teeth, tooth row as long as on movable finger, movable finger with 119 (122) small teeth, in distal row part with small cusps, others rounded; a very short venom duct present in fixed finger. Trichobothria:
<emphasis id="B944ED202834273DFCC19DB8FC71C9F6" box="[832,902,642,665]" italics="true" pageId="3" pageNumber="262">eb-esb</emphasis>
on lateral side of hand,
<emphasis id="B944ED202834273DFB089DB8FB1FC9F6" box="[1161,1256,642,665]" italics="true" pageId="3" pageNumber="262">ib-isb-ist</emphasis>
grouped at level of
<emphasis id="B944ED202834273DFCDC9D98FC9DC9D6" box="[861,874,674,697]" italics="true" pageId="3" pageNumber="262">b</emphasis>
(movable finger),
<emphasis id="B944ED202834273DFBB59D99FBA5C9D5" box="[1076,1106,675,698]" italics="true" pageId="3" pageNumber="262">est</emphasis>
nearer to
<emphasis id="B944ED202834273DFB3E9D99FB1EC9D5" box="[1215,1257,675,698]" italics="true" pageId="3" pageNumber="262">et/it</emphasis>
and proximal of
<emphasis id="B944ED202834273DFCC19DF9FC80C9B6" box="[832,887,706,730]" italics="true" pageId="3" pageNumber="262">st, sb</emphasis>
on movable finger halfway between
<emphasis id="B944ED202834273DFA839DF8FAF8C9B6" box="[1282,1295,706,729]" italics="true" pageId="3" pageNumber="262">b</emphasis>
and
<emphasis id="B944ED202834273DFAC29DF9FAABC9B5" box="[1347,1372,707,730]" italics="true" pageId="3" pageNumber="262">st.</emphasis>
</paragraph>
<paragraph id="8B8F31322834273DFCBE9DD9FC47C816" blockId="3.[831,1437,258,1145]" pageId="3" pageNumber="262">
<emphasis id="B944ED202834273DFCBE9DD9FC81C995" box="[831,886,739,762]" italics="true" pageId="3" pageNumber="262">Leg I</emphasis>
: Femur 4.9 (5.5) times longer than deep and 1.5-1.6 times longer than patella; patella 4.1 (4.1) times, tibia 6.8 (6.3) times, basitarsus 3.9 (3.2) times, telotarsus 6.1 (4.7) times longer than deep and 1.6 (1.5) times longer than basitarsus.
</paragraph>
<paragraph id="8B8F31322834273DFCBE9CB9FB23CF16" blockId="3.[831,1437,258,1145]" pageId="3" pageNumber="262">
<emphasis id="B944ED202834273DFCBE9CB9FC7FC8F5" box="[831,904,899,922]" italics="true" pageId="3" pageNumber="262">Leg IV</emphasis>
: Femur+patella 6.5 (6.5) times longer than deep, femur distinctly shorter than patella, tibia 9.4 (9.2) times, with one or two longer setae in distal half, basitarsus with a basal tactile seta, 4.0 (3.9) times, telotarsus with a tactile seta near middle of segment, 6.8 (5.6) times longer than deep and 1.5 times longer than basitarsus; claws long and slender, smooth, subterminal seta forked and dentate, arolia shorter than claws.
</paragraph>
<paragraph id="8B8F31322834273DFCBE9BA8FA72CD27" blockId="3.[831,1437,1169,1610]" pageId="3" pageNumber="262">
<emphasis id="B944ED202834273DFCBE9BA8FC2CCFA7" bold="true" pageId="3" pageNumber="262">
Measurements of
<typeStatus id="548B8F902834273DFBAF9BA8FB65CFC5" box="[1070,1170,1170,1194]" pageId="3" pageNumber="262" type="holotype">holotype</typeStatus>
<specimenCount id="9D36FABB2834273DFB299BABFB4BCFC7" box="[1192,1212,1169,1192]" pageId="3" pageNumber="262">♂</specimenCount>
(
<typeStatus id="548B8F902834273DFB5B9BABFAB3CFC7" box="[1242,1348,1169,1192]" pageId="3" pageNumber="262" type="paratype">paratype</typeStatus>
<specimenCount id="9D36FABB2834273DFADA9BABFA98CFC7" box="[1371,1391,1169,1192]" pageId="3" pageNumber="262">♀</specimenCount>
in parentheses):
</emphasis>
Body length 4.40 (5.03); carapace 1.17/10.05 (1.31/1.22). Pedipalp: trochanter 0.88/0.35 (1.01/0.38), femur 1.92/0.33 (2.10/0.36), patella 1.78/1.41 (2.01/0.43), pedicel length 0.95 (1.00); hand with pedicel 1.37/0.60 (1.48/0.66), length of pedicel 0.24 (0.27), finger length 1.93 (2.05), chela length with pedicel 3.27 (5.32), without pedicel 3.03 (3.24). Leg I: femur 1.11/0.23 (1.23/0.22), patella 0.75/0.18 (0.78/0.19), tibia 0.98/0.14 (1.03/0.16), basitarsus 0.47/0.12 (0.46/0.15), telotarsus 0.75/0.12 (0.71/0.15). Leg IV: femur+patella 1.87/0.29 (1.98/0.30), femur length 0.82 (0.88), tibia 1.74/0.18 (1.85/0.20), basitarsus 0.60/0.15 (0.61/0.16), telotarsus 0.90/0.14 (0.89/0.16).
</paragraph>
</subSubSection>
<subSubSection id="C32A62B92834273BFCBE995BFD2FCA54" lastPageId="5" lastPageNumber="264" pageId="3" pageNumber="262" type="discussion">
<paragraph id="8B8F31322834273BFCBE995BFD2FCA54" blockId="3.[831,1436,1632,2073]" lastBlockId="5.[151,756,258,315]" lastPageId="5" lastPageNumber="264" pageId="3" pageNumber="262">
<emphasis id="B944ED202834273DFCBE995BFC46CD16" bold="true" box="[831,945,1633,1657]" pageId="3" pageNumber="262">Remarks:</emphasis>
The new species,
<emphasis id="B944ED202834273DFB0E9958FACBCD16" box="[1167,1340,1634,1657]" italics="true" pageId="3" pageNumber="262">
<taxonomicName id="4C304AB12834273DFB0E9958FACFCD16" box="[1167,1336,1634,1657]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="262" phylum="Arthropoda" rank="species" species="cavernarum">B. cavernarum</taxonomicName>
,
</emphasis>
is most similar to
<emphasis id="B944ED202834273DFC2F99B8FB7BCDF8" box="[942,1164,1664,1689]" italics="true" pageId="3" pageNumber="262">
<taxonomicName id="4C304AB12834273DFC2F99B8FB7FCDF8" box="[942,1160,1664,1689]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="262" phylum="Arthropoda" rank="species" species="chinacavernicola">B. chinacavernicola</taxonomicName>
,
</emphasis>
sharing with this species a similar size, elongate pedipalps, a long patellar pedicel (about half of the total length of patella), 6 or 7 setae on posterior margin of carapace, and 8 or 9 setae on anterior tergites.
<taxonomicName id="4C304AB12834273DFBF1983BFA6BCC76" box="[1136,1436,1793,1817]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="262" phylum="Arthropoda" rank="species" species="cavernarum" status="sp. nov.">
<emphasis id="B944ED202834273DFBF1983BFA6BCC76" box="[1136,1436,1793,1817]" italics="true" pageId="3" pageNumber="262">Bisetocreagris cavernarum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A277505B2834273DFCBE981BFC8ACC57" box="[831,893,1825,1848]" pageId="3" pageNumber="262" rank="species">n. sp.</taxonomicNameLabel>
differs from
<emphasis id="B944ED202834273DFB959818FB01CC58" box="[1044,1270,1824,1849]" italics="true" pageId="3" pageNumber="262">
<taxonomicName id="4C304AB12834273DFB959818FB05CC58" box="[1044,1266,1824,1849]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="262" phylum="Arthropoda" rank="species" species="chinacavernicola">B. chinacavernicola</taxonomicName>
,
</emphasis>
recorded from the caves Dei (=Xian Nin) and Chao-Tian (both near Huaying), by the presence of indistinct lenses on the 2 anterior eyes and the presence of 2 indistinct posterior eye-spots, by the fine granulation of the palpal hand, a more slender chela (without pedicel 4.9-5.0 vs 4.4-4.86 times longer than broad), the position of trichobothrium
<emphasis id="B944ED202834273DFCBE9738FCA6C376" box="[831,849,2050,2073]" italics="true" pageId="3" pageNumber="262">st</emphasis>
which is distinctly nearer to
<emphasis id="B944ED202834273DFB2B9738FB46C376" box="[1194,1201,2050,2073]" italics="true" pageId="3" pageNumber="262">t</emphasis>
than to
<emphasis id="B944ED202834273DFA98973AFAC6C378" box="[1305,1329,2048,2071]" italics="true" pageId="3" pageNumber="262">sb</emphasis>
(halfway between
<emphasis id="B944ED202832273BFF7A9E38FEE4CA76" box="[251,275,258,281]" italics="true" pageId="5" pageNumber="264">sb</emphasis>
and
<emphasis id="B944ED202832273BFECC9E39FEA3CA75" box="[333,340,259,282]" italics="true" pageId="5" pageNumber="264">t</emphasis>
in
<taxonomicName id="4C304AB12832273BFEFF9E38FDCFCA76" box="[382,568,258,281]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="264" phylum="Arthropoda" rank="species" species="chinacavernicola">
<emphasis id="B944ED202832273BFEFF9E38FDCFCA76" box="[382,568,258,281]" italics="true" pageId="5" pageNumber="264">chinacavernicola</emphasis>
</taxonomicName>
) and perhaps by shorter legs (e.g. femur+patella IV 1.87-1.98 vs 2.18).
</paragraph>
</subSubSection>
</treatment>
</document>
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