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<mods:title id="C90A4ABF6B6BBAE9F34F28C4D04B89AD">A reappraisal of some basal lineages of the family Macrochelidae, with the description of a new genus (Acarina: Mesostigmata)</mods:title>
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<mods:namePart id="5EAEBBBB871EB8F1170E0F00D00D288C">Emberson, Rowan M.</mods:namePart>
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<treatment id="03997670FFB0E66C8ECBFC7A72DF691A" ID-DOI="http://doi.org/10.5281/zenodo.6212254" ID-GBIF-Taxon="119387888" ID-Zenodo-Dep="6212254" LSID="urn:lsid:plazi:treatment:03997670FFB0E66C8ECBFC7A72DF691A" httpUri="http://treatment.plazi.org/id/03997670FFB0E66C8ECBFC7A72DF691A" lastPageId="12" lastPageNumber="49" pageId="11" pageNumber="48">
<subSubSection id="C32A94EDFFB0E66B8ECBFC7A71016CBA" box="[151,530,898,924]" pageId="11" pageNumber="48" type="nomenclature">
<paragraph id="8B8FC766FFB0E66B8ECBFC7A71016CBA" blockId="11.[151,530,898,924]" box="[151,530,898,924]" pageId="11" pageNumber="48">
<heading id="D0C7700AFFB0E66B8ECBFC7A71016CBA" bold="true" box="[151,530,898,924]" fontSize="11" level="1" pageId="11" pageNumber="48" reason="1">
<taxonomicName id="4C30BCE5FFB0E66B8ECBFC7A71016CBA" authority="Oudemans, 1931" authorityName="Oudemans" authorityYear="1931" box="[151,530,898,924]" class="Arachnida" family="Macrochelidae" genus="Macrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFB0E66B8ECBFC7A71016CBA" bold="true" box="[151,530,898,924]" pageId="11" pageNumber="48">
<emphasis id="B9441B74FFB0E66B8ECBFC7A72286CBA" bold="true" box="[151,315,898,924]" italics="true" pageId="11" pageNumber="48">Macrholaspis</emphasis>
<bibRefCitation id="EFA1BA97FFB0E66B8F1FFC7A71016CBA" author="Oudemans" box="[323,530,898,924]" pageId="11" pageNumber="53" refString="Oudemans, A. C. (1931) Acarologisches Aanteekeningen CIX. Entomologische Berichten, 8, 272 - 280." type="journal article" year="1931">Oudemans, 1931</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
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<subSubSection id="C32A94EDFFB0E66B8ECBFC3272B46A22" pageId="11" pageNumber="48" type="reference_group">
<paragraph id="8B8FC766FFB0E66B8ECBFC3270026B27" blockId="11.[151,1281,970,1025]" pageId="11" pageNumber="48">
<treatmentCitationGroup id="AB20E048FFB0E66B8ECBFC3270026B27" pageId="11" pageNumber="48">
<taxonomicName id="4C30BCE5FFB0E66B8ECBFC32710E6CC7" authority="Oudemans, 1931: 272" authorityName="Oudemans" authorityPageNumber="272" authorityYear="1931" box="[151,541,970,993]" class="Arachnida" family="Macrochelidae" genus="Macrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFB0E66B8ECBFC32723A6CC7" box="[151,297,970,993]" italics="true" pageId="11" pageNumber="48">Macrholaspis</emphasis>
<treatmentCitation id="0A91E177FFB0E66B8F6CFC32710E6CC7" author="Oudemans" box="[304,541,970,993]" page="272" pageId="11" pageNumber="48" year="1931">
<bibRefCitation id="EFA1BA97FFB0E66B8F6CFC3272FB6CC7" author="Oudemans" box="[304,488,970,993]" pageId="11" pageNumber="53" refString="Oudemans, A. C. (1931) Acarologisches Aanteekeningen CIX. Entomologische Berichten, 8, 272 - 280." type="journal article" year="1931">Oudemans, 1931</bibRefCitation>
: 272
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="548B79C4FFB0E66B8C74FC32714E6CC7" box="[552,605,970,993]" pageId="11" pageNumber="48">Type</typeStatus>
species:
<taxonomicName id="4C30BCE5FFB0E66B8CE2FC3270EA6CC7" authority="Koch, 1839" authorityName="Koch" authorityYear="1839" box="[702,1017,970,993]" class="Arachnida" family="Ascidae" genus="Gamasus" higherTaxonomySource="CoL" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="opacus">
<emphasis id="B9441B74FFB0E66B8CE2FC3270676CC7" box="[702,884,970,993]" italics="true" pageId="11" pageNumber="48">Gamasus opacus</emphasis>
Koch, 1839
</taxonomicName>
, by original designation.
<taxonomicName id="4C30BCE5FFB0E66B8ECBFC1272616B27" box="[151,370,1002,1025]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="opacus">
<emphasis id="B9441B74FFB0E66B8ECBFC1272616B27" box="[151,370,1002,1025]" italics="true" pageId="11" pageNumber="48">Macrocheles opacus</emphasis>
</taxonomicName>
species group.—
<treatmentCitation id="0A91E177FFB0E66B8C6EFC12701E6B27" author="Johnston" box="[562,781,1002,1025]" page="148" pageId="11" pageNumber="48" year="1970">
<bibRefCitation id="EFA1BA97FFB0E66B8C6EFC1271C46B27" author="Johnston" box="[562,727,1002,1025]" pageId="11" pageNumber="52" refString="Johnston, D. E. (1970) Notes on a collection of Austrian Macrochelidae with the description of Macrocheles beieri, n. sp. (Acari - Parasitiformes). Annalen des Naturhistorischen Museums in Wien, 74, 145 - 150." type="journal article" year="1970">Johnston, 1970</bibRefCitation>
: 148
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8FC766FFB0E66B8ECBFBC872B46A22" blockId="11.[151,1437,1070,2031]" pageId="11" pageNumber="48">Medium sized (female dorsal shield usually 600–1100 μm in length), litter-dwelling Macrochelini; dorsal shield with 28 pairs of setae, plus 1–4 unpaired or paired setae; setae long, tapered and usually entirely pilose; j1 laterally expanded; dorsal shield tapered posteriorly, lateral and posterior margins dentate, crenulate or smooth; ventri-anal shield rounded or subcircular, with 1–3 pairs of pre-anal setae; three pairs of postepigynal platelets free in membranous cuticle; lateral processes of epistome reduced; cheliceral dorsal seta pectinate laterodistally.</paragraph>
</subSubSection>
<subSubSection id="C32A94EDFFB0E66C8E98FAE873A96CC1" lastPageId="12" lastPageNumber="49" pageId="11" pageNumber="48" type="description">
<paragraph id="8B8FC766FFB0E66B8E98FAE8721F68EC" blockId="11.[151,1437,1070,2031]" pageId="11" pageNumber="48">
The dorsal shield has a basic complement of 28 pairs of setae plus 1–4 additional unpaired or paired setae; setae J3 are always present and a single seta, or a pair of setae, or 3–4 unpaired setae are present between setae j6 and J3. Setae j1 are short and laterally expanded; the other setae are long, tapered and pilose along their entire length (except for
<taxonomicName id="4C30BCE5FFB0E66B8F88FA7871A56ABC" authority="Balogh" authorityName="Balogh" box="[468,694,1408,1434]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="evansi">
<emphasis id="B9441B74FFB0E66B8F88FA7872FF6ABF" box="[468,492,1408,1433]" italics="true" pageId="11" pageNumber="48">M</emphasis>
.
<emphasis id="B9441B74FFB0E66B8FA2FA78715D6ABF" box="[510,590,1408,1433]" italics="true" pageId="11" pageNumber="48">evansi</emphasis>
Balogh
</taxonomicName>
, in which setae j6, z6, J3 and a pair between j6 and J3 are aciculate). Setae z1 and J5 are often shorter than the other setae. The dorsal shield is tapered from the shoulders and does not cover the entire dorsum. It may be ornamented with microspicules or have reticulated surface sculpture. The lateral and posterior margins of the dorsal shield are often finely dentate or crenulate. The ventral sclerotisation is reduced and the setae are aciculate or pilose. The sternal shield is punctatereticulate without strong lineal patterns. The epigynal shield is reduced and rounded posteriorly, with the pores free in the membrane. The ventri-anal shield is more or less reduced, usually rounded or subcircular, with 1–3 pairs of pre-anal setae and a cribum with strong para-anal extensions. There are three pairs of sclerotised post-epigynal platelets free in membranous cuticle. The lateral processes of the epistome are usually reduced to narrow spikes, either attached to the median process or free (
<figureCitation id="130BDBE3FFB0E66B8A6BF928779D69CC" box="[1079,1166,1744,1770]" captionStart="FIGURES 12 – 17. 12 – 15" captionStartId="7.[151,270,1948,1972]" captionTargetBox="[202,1362,402,1947]" captionTargetId="figure@7.[202,1385,402,1988]" captionTargetPageId="7" captionText="FIGURES 12 – 17. 12 – 15. Reductholaspis analis (Hyatt &amp; Emberson) comb. nov., female. 12. Chelicera; 13. Epistome; 14. Anal shield; 15. gnathosoma venter, showing position of hypostomal setae. 16. Nothrholaspis carinatus (Koch), epistome. 17. Macrholaspis opacus (Koch), epistome." httpUri="https://zenodo.org/record/195838/files/figure.png" pageId="11" pageNumber="48">Fig. 17</figureCitation>
), but they are absent in
<taxonomicName id="4C30BCE5FFB0E66B8ECBF90D72CF6829" authority="Masan. The" authorityName="Masan. The" box="[151,476,1781,1807]" class="Arachnida" family="Macrochelidae" genus="Macrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="carpathicus">
<emphasis id="B9441B74FFB0E66B8ECBF90D73BC6828" box="[151,175,1781,1806]" italics="true" pageId="11" pageNumber="48">M</emphasis>
.
<emphasis id="B9441B74FFB0E66B8EE2F90D725A6828" box="[190,329,1781,1806]" italics="true" pageId="11" pageNumber="48">carpathicus</emphasis>
Mašán. The
</taxonomicName>
median process is bifurcate and usually dissected distally. The fixed chela has an offset subterminal tooth and a median tooth, and the female movable chela has two central teeth. The cheliceral dorsal seta is pectinate laterodistally. The legs are strongly rugose, with setation normal for the family. The setae are mainly pilose or occasionally blade-shaped, except on coxa and tarsus I where they are aciculate. The visible spermathecal structures include elongate tubuli and well developed rami; the sacculus is indistinct.
</paragraph>
<paragraph id="8B8FC766FFB0E66C8E98F82D70126FF1" blockId="11.[151,1437,1070,2031]" lastBlockId="12.[151,1437,152,1596]" lastPageId="12" lastPageNumber="49" pageId="11" pageNumber="48">
Males, where known (
<taxonomicName id="4C30BCE5FFB0E66B8F88F82D71DE68C9" authority="Johnston" authorityName="Johnston" box="[468,717,2005,2031]" class="Arachnida" family="Macrochelidae" genus="Macrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="beieri">
<emphasis id="B9441B74FFB0E66B8F88F82D72FF68C8" box="[468,492,2005,2030]" italics="true" pageId="11" pageNumber="48">M</emphasis>
.
<emphasis id="B9441B74FFB0E66B8FA1F82D715768C8" box="[509,580,2005,2030]" italics="true" pageId="11" pageNumber="48">beieri</emphasis>
(Johnston)
</taxonomicName>
and
<taxonomicName id="4C30BCE5FFB0E66B8D55F82D773368C9" authority="Petrova" authorityName="Petrova" box="[777,1056,2005,2031]" class="Arachnida" family="Macrochelidae" genus="Macrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="reductus">
<emphasis id="B9441B74FFB0E66B8D55F82D703268C8" box="[777,801,2005,2030]" italics="true" pageId="11" pageNumber="48">M</emphasis>
.
<emphasis id="B9441B74FFB0E66B8D6FF82D708968C8" box="[819,922,2005,2030]" italics="true" pageId="11" pageNumber="48">reductus</emphasis>
(Petrova))
</taxonomicName>
, have separate sternogynal and ventri-anal shields. The male movable chela is unidentate with a short, strongly tapered, posteriodorsally directed spermatodactyl. Femur II has a ventral spur.
</paragraph>
<paragraph id="8B8FC766FFB7E66C8E9AFF1A771E6EB4" blockId="12.[151,1437,152,1596]" pageId="12" pageNumber="49">
<emphasis id="B9441B74FFB7E66C8E9AFF1A72076FDA" bold="true" box="[198,276,226,252]" pageId="12" pageNumber="49">Notes.</emphasis>
<taxonomicName id="4C30BCE5FFB7E66C8F40FF1A72AE6FDD" box="[284,445,226,251]" class="Arachnida" family="Macrochelidae" genus="Macrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFB7E66C8F40FF1A72AE6FDD" box="[284,445,226,251]" italics="true" pageId="12" pageNumber="49">Macrholaspis</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4C30BCE5FFB7E66C8C95FF1A704C6FDD" box="[713,863,226,251]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFB7E66C8C95FF1A704C6FDD" box="[713,863,226,251]" italics="true" pageId="12" pageNumber="49">Macrocheles</emphasis>
</taxonomicName>
by the reduced ventral sclerotisation and by the presence of three pairs of sclerotised post-epigynal platelets in the membrane between the epigynal and ventri-anal shields, a feature shared with
<taxonomicName id="4C30BCE5FFB7E66C8CF0FED570736E60" box="[684,864,301,326]" class="Arachnida" family="Macrochelidae" genus="Nothrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFB7E66C8CF0FED570736E60" box="[684,864,301,326]" italics="true" pageId="12" pageNumber="49">Nothrholaspis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C30BCE5FFB7E66C8DC3FED5774D6E60" box="[927,1118,301,326]" class="Arachnida" family="Macrochelidae" genus="Geotrupacarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFB7E66C8DC3FED5774D6E60" box="[927,1118,301,326]" italics="true" pageId="12" pageNumber="49">Geotrupacarus</emphasis>
</taxonomicName>
.
<taxonomicName id="4C30BCE5FFB7E66C8A2CFED5760E6E60" box="[1136,1309,301,326]" class="Arachnida" family="Macrochelidae" genus="Macrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFB7E66C8A2CFED5760E6E60" box="[1136,1309,301,326]" italics="true" pageId="12" pageNumber="49">Macrholaspis</emphasis>
</taxonomicName>
is further distinguished from other genera by the distinctive shape of the epistome, the fully pilose dorsal setae, the laterally expanded setae j1, the litter dwelling habit, and the rarity of males.
</paragraph>
<paragraph id="8B8FC766FFB7E66C8E98FE65722D6D54" blockId="12.[151,1437,152,1596]" pageId="12" pageNumber="49">
<bibRefCitation id="EFA1BA97FFB7E66C8E98FE6571136E91" author="Hyatt" box="[196,512,413,439]" pageId="12" pageNumber="52" refString="Hyatt, K. H. &amp; Emberson, R. M. (1988) A review of the Macrochelidae (Acari: Mesostigmata) of the British Isles. Bulletin of the British Museum (Natural History), Zoology, 54, 63 - 125 + 6 plates." type="book chapter" year="1988">Hyatt &amp; Emberson (1988)</bibRefCitation>
saw no males among
<specimenCount id="9D360CEFFFB7E66C8D51FE65709A6E91" box="[781,905,413,439]" pageId="12" pageNumber="49" type="adult">116 adults</specimenCount>
of the three species occurring in the British Isles, and
<bibRefCitation id="EFA1BA97FFB7E66C8F53FE3A72A06EFA" author="Masan" box="[271,435,450,476]" pageId="12" pageNumber="53" refString="Masan, P. (2003) Macrochelid mites of Slovakia (Acari: Mesostigmata: Macrochelidae). Institute of Zoology, Slovak Academy of Sciences, Bratislava. 149 pp." type="book" year="2003">Mašán (2003)</bibRefCitation>
did not report any males among nearly
<specimenCount id="9D360CEFFFB7E66C8DD2FE3A772E6EFA" box="[910,1085,450,476]" pageId="12" pageNumber="49" type="generic">200 specimens</specimenCount>
of six species from
<collectingCountry id="F32787F6FFB7E66C8B72FE3A768A6EFA" box="[1326,1433,450,476]" name="Slovakia" pageId="12" pageNumber="49">Slovakia</collectingCountry>
.
<bibRefCitation id="EFA1BA97FFB7E66C8ECBFE10724A6D24" author="Johnston" box="[151,345,488,514]" pageId="12" pageNumber="52" refString="Johnston, D. E. (1970) Notes on a collection of Austrian Macrochelidae with the description of Macrocheles beieri, n. sp. (Acari - Parasitiformes). Annalen des Naturhistorischen Museums in Wien, 74, 145 - 150." type="journal article" year="1970">Johnston (1970)</bibRefCitation>
stated that
<taxonomicName id="4C30BCE5FFB7E66C8FB8FE1071D66D24" authority="Johnston" authorityName="Johnston" box="[484,709,488,514]" class="Arachnida" family="Macrochelidae" genus="Macrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="beieri">
<emphasis id="B9441B74FFB7E66C8FB8FE1072EF6D27" box="[484,508,488,513]" italics="true" pageId="12" pageNumber="49">M</emphasis>
.
<emphasis id="B9441B74FFB7E66C8C50FE1071416D27" box="[524,594,488,513]" italics="true" pageId="12" pageNumber="49">beieri</emphasis>
Johnston
</taxonomicName>
was the only species of the genus (species group) for which males were known. This, however, is not strictly correct;
<bibRefCitation id="EFA1BA97FFB7E66C8D14FDF570EE6D01" author="Petrova" box="[840,1021,525,551]" pageId="12" pageNumber="53" refString="Petrova, A. D. (1966) Macrocheles reductus a new species of gamasoid mite from the southern coast (Gamasoidea, Macrochelidae). Nauchnye Doklady Vysshei Shkoly: Biologicheski Nauki, 9, 18 - 23. [in Russian]." type="journal article" year="1966">Petrova (1966)</bibRefCitation>
, in describing
<taxonomicName id="4C30BCE5FFB7E66C8AF2FDF576286D00" box="[1198,1339,525,551]" class="Arachnida" family="Macrochelidae" genus="Macrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="reductus">
<emphasis id="B9441B74FFB7E66C8AF2FDF577D56D00" box="[1198,1222,525,550]" italics="true" pageId="12" pageNumber="49">M</emphasis>
.
<emphasis id="B9441B74FFB7E66C8A8AFDF576286D00" box="[1238,1339,525,550]" italics="true" pageId="12" pageNumber="49">reductus</emphasis>
</taxonomicName>
, clearly indicated and illustrated the male of the species, but compared it to species of
<taxonomicName id="4C30BCE5FFB7E66C8A3EFDCA761C6D6D" box="[1122,1295,562,587]" class="Arachnida" family="Macrochelidae" genus="Nothrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFB7E66C8A3EFDCA761C6D6D" box="[1122,1295,562,587]" italics="true" pageId="12" pageNumber="49">Nothrholaspis</emphasis>
</taxonomicName>
rather than
<taxonomicName id="4C30BCE5FFB7E66C8ECBFDA0722B6D57" box="[151,312,600,625]" class="Arachnida" family="Macrochelidae" genus="Macrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFB7E66C8ECBFDA0722B6D57" box="[151,312,600,625]" italics="true" pageId="12" pageNumber="49">Macrholaspis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B8FC766FFB7E66C8E98FD8573A96CC1" blockId="12.[151,1437,152,1596]" pageId="12" pageNumber="49">
<taxonomicName id="4C30BCE5FFB7E66C8E98FD8572766DB0" box="[196,357,637,662]" class="Arachnida" family="Macrochelidae" genus="Macrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFB7E66C8E98FD8572766DB0" box="[196,357,637,662]" italics="true" pageId="12" pageNumber="49">Macrholaspis</emphasis>
</taxonomicName>
is closely related to
<taxonomicName id="4C30BCE5FFB7E66C8C04FD8571ED6DB0" box="[600,766,637,662]" class="Arachnida" family="Macrochelidae" genus="Nothrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFB7E66C8C04FD8571ED6DB0" box="[600,766,637,662]" italics="true" pageId="12" pageNumber="49">Nothrholaspis</emphasis>
</taxonomicName>
, and a case could be made for regarding it as a subgenus of the latter. However, the epistomal structure of
<taxonomicName id="4C30BCE5FFB7E66C8C8DFD5A70616D9D" box="[721,882,674,699]" class="Arachnida" family="Macrochelidae" genus="Macrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFB7E66C8C8DFD5A70616D9D" box="[721,882,674,699]" italics="true" pageId="12" pageNumber="49">Macrholaspis</emphasis>
</taxonomicName>
is quite different from that of
<taxonomicName id="4C30BCE5FFB7E66C8A8CFD5A76666D9D" box="[1232,1397,674,699]" class="Arachnida" family="Macrochelidae" genus="Nothrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFB7E66C8A8CFD5A76666D9D" box="[1232,1397,674,699]" italics="true" pageId="12" pageNumber="49">Nothrholaspis</emphasis>
</taxonomicName>
, as are the fully pilose dorsal setae and the laterally expanded setae j1. The two genera share the presence of three pairs of sclerotised post-epigynal platelets with
<taxonomicName id="4C30BCE5FFB7E66C8C86FD15709C6C20" box="[730,911,749,774]" class="Arachnida" family="Macrochelidae" genus="Geotrupacarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFB7E66C8C86FD15709C6C20" box="[730,911,749,774]" italics="true" pageId="12" pageNumber="49">Geotrupacarus</emphasis>
</taxonomicName>
, a feature otherwise seen only rarely in the family. These three genera also have para-anal extensions of the cribrum, an apparently pleisiomorphic character that occurs widely in free-living, early derivative lines of the
<taxonomicName id="4C30BCE5FFB7E66C8A10FCC0761E6C74" box="[1100,1293,824,850]" class="Arachnida" family="Macrochelidae" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="family">Macrochelidae</taxonomicName>
(including Geholaspini), but is absent in most species of
<taxonomicName id="4C30BCE5FFB7E66C8CF0FCA570516C50" box="[684,834,861,886]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFB7E66C8CF0FCA570516C50" box="[684,834,861,886]" italics="true" pageId="12" pageNumber="49">Macrocheles</emphasis>
</taxonomicName>
. The males of these three genera also have separate sternogynal and ventri-anal shields and major spurs on the legs are limited to femora II. Males of more derived species of
<taxonomicName id="4C30BCE5FFB7E66C8F2AFC50711E6CE7" box="[374,525,936,961]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFB7E66C8F2AFC50711E6CE7" box="[374,525,936,961]" italics="true" pageId="12" pageNumber="49">Macrocheles</emphasis>
</taxonomicName>
often have holoventral shields and may have additional spurs on legs II and IV.
</paragraph>
</subSubSection>
<subSubSection id="C32A94EDFFB7E66C8E98FC0A76706A11" pageId="12" pageNumber="49" type="reference_group">
<paragraph id="8B8FC766FFB7E66C8E98FC0A72916B71" blockId="12.[151,1437,152,1596]" pageId="12" pageNumber="49">
<taxonomicName id="4C30BCE5FFB7E66C8E98FC0A72746B2D" box="[196,359,1010,1035]" class="Arachnida" family="Macrochelidae" genus="Macrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFB7E66C8E98FC0A72746B2D" box="[196,359,1010,1035]" italics="true" pageId="12" pageNumber="49">Macrholaspis</emphasis>
</taxonomicName>
is a Palaearctic genus with species distributed from
<collectingCountry id="F32787F6FFB7E66C8D80FC0A773D6B2A" box="[988,1070,1010,1036]" name="United Kingdom" pageId="12" pageNumber="49">Britain</collectingCountry>
and
<collectingCountry id="F32787F6FFB7E66C8A35FC0A77AE6B2A" box="[1129,1213,1010,1036]" name="Ireland" pageId="12" pageNumber="49">Ireland</collectingCountry>
to
<collectingCountry id="F32787F6FFB7E66C8AB9FC0A763A6B2A" box="[1253,1321,1010,1036]" name="Japan" pageId="12" pageNumber="49">Japan</collectingCountry>
(
<bibRefCitation id="EFA1BA97FFB7E66C8B65FC0A72466B14" author="Hyatt" pageId="12" pageNumber="52" refString="Hyatt, K. H. &amp; Emberson, R. M. (1988) A review of the Macrochelidae (Acari: Mesostigmata) of the British Isles. Bulletin of the British Museum (Natural History), Zoology, 54, 63 - 125 + 6 plates." type="book chapter" year="1988">Hyatt &amp; Emberson 1988</bibRefCitation>
,
<bibRefCitation id="EFA1BA97FFB7E66C8F3EFBE071086B14" author="Ishikawa" box="[354,539,1048,1074]" pageId="12" pageNumber="52" refString="Ishikawa, K. (1969) Taxonomic investigations on free-living mites in the subalpine forest on Shiga Heights IBP area. I. Mesostigmata (Part 1). Bulletin of the National Science Museum, Tokyo, 12, 39 - 64." type="journal article" year="1969">Ishikawa, 1969</bibRefCitation>
), but appears to be most diverse in Central Europe (
<bibRefCitation id="EFA1BA97FFB7E66C8AD4FBE076356B14" author="Balogh" box="[1160,1318,1048,1074]" pageId="12" pageNumber="51" refString="Balogh, J. (1958) Macrocheliden aus Bulgarien (Acarina, Mesostigmata). Acta entomologica Musei Nationalis Prague, 32, 247 - 256." type="journal article" year="1958">Balogh 1958</bibRefCitation>
,
<bibRefCitation id="EFA1BA97FFB7E66C8B6FFBE073C76B71" author="Johnston" pageId="12" pageNumber="52" refString="Johnston, D. E. (1970) Notes on a collection of Austrian Macrochelidae with the description of Macrocheles beieri, n. sp. (Acari - Parasitiformes). Annalen des Naturhistorischen Museums in Wien, 74, 145 - 150." type="journal article" year="1970">Johnston 1970</bibRefCitation>
,
<bibRefCitation id="EFA1BA97FFB7E66C8EBCFBC572656B71" author="Masan" box="[224,374,1085,1111]" pageId="12" pageNumber="53" refString="Masan, P. (2003) Macrochelid mites of Slovakia (Acari: Mesostigmata: Macrochelidae). Institute of Zoology, Slovak Academy of Sciences, Bratislava. 149 pp." type="book" year="2003">Mašán 2003</bibRefCitation>
).
</paragraph>
<paragraph id="8B8FC766FFB7E66C8E98FB9A76706A11" blockId="12.[151,1437,152,1596]" pageId="12" pageNumber="49">
<emphasis id="B9441B74FFB7E66C8E98FB9A72B16B5A" bold="true" box="[196,418,1122,1148]" pageId="12" pageNumber="49">Included species:</emphasis>
<taxonomicName id="4C30BCE5FFB7E66C8FF0FB9B71E66B5A" authority="Johnston, 1970" authorityName="Johnston" authorityYear="1970" box="[428,757,1122,1148]" class="Arachnida" family="Macrochelidae" genus="Macrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="beieri" status="comb. nov.">
<emphasis id="B9441B74FFB7E66C8FF0FB9B72D76B5A" box="[428,452,1123,1148]" italics="true" pageId="12" pageNumber="49">M</emphasis>
.
<emphasis id="B9441B74FFB7E66C8F88FB9B71086B5A" box="[468,539,1123,1148]" italics="true" pageId="12" pageNumber="49">beieri</emphasis>
(
<bibRefCitation id="EFA1BA97FFB7E66C8C71FB9A71FF6B5A" author="Johnston" box="[557,748,1122,1148]" pageId="12" pageNumber="52" refString="Johnston, D. E. (1970) Notes on a collection of Austrian Macrochelidae with the description of Macrocheles beieri, n. sp. (Acari - Parasitiformes). Annalen des Naturhistorischen Museums in Wien, 74, 145 - 150." type="journal article" year="1970">Johnston, 1970</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A277A60FFFB7E66C8D5EFB9A70CA6B5A" box="[770,985,1122,1148]" pageId="12" pageNumber="49">new combination</taxonomicNameLabel>
;
<taxonomicName id="4C30BCE5FFB7E66C8DBAFB9B764E6B5A" authority="Masan, 2003" authorityName="Masan" authorityYear="2003" box="[998,1373,1122,1148]" class="Arachnida" family="Macrochelidae" genus="Macrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="carpathicus" status="comb. nov.">
<emphasis id="B9441B74FFB7E66C8DBAFB9B70ED6B5A" box="[998,1022,1123,1148]" italics="true" pageId="12" pageNumber="49">M</emphasis>
.
<emphasis id="B9441B74FFB7E66C8A53FB9B778C6B5A" box="[1039,1183,1123,1148]" italics="true" pageId="12" pageNumber="49">carpathicus</emphasis>
(
<bibRefCitation id="EFA1BA97FFB7E66C8AECFB9A76476B5A" author="Masan" box="[1200,1364,1122,1148]" pageId="12" pageNumber="53" refString="Masan, P. (2003) Macrochelid mites of Slovakia (Acari: Mesostigmata: Macrochelidae). Institute of Zoology, Slovak Academy of Sciences, Bratislava. 149 pp." type="book" year="2003">Mašán, 2003</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A277A60FFFB7E66C8B36FB9A72266B84" pageId="12" pageNumber="49">new combination</taxonomicNameLabel>
;
<taxonomicName id="4C30BCE5FFB7E66C8F1EFB70700F6B84" authority="Evans &amp; Browning, 1956" authorityName="Evans &amp; Browning" authorityYear="1956" box="[322,796,1160,1186]" class="Arachnida" family="Macrochelidae" genus="Macrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="B9441B74FFB7E66C8F1EFB7072496B87" box="[322,346,1160,1185]" italics="true" pageId="12" pageNumber="49">M</emphasis>
.
<emphasis id="B9441B74FFB7E66C8F37FB7072C06B87" box="[363,467,1160,1185]" italics="true" pageId="12" pageNumber="49">dentatus</emphasis>
<bibRefCitation id="EFA1BA97FFB7E66C8F80FB70700F6B84" author="Evans" box="[476,796,1160,1186]" pageId="12" pageNumber="52" refString="Evans, G. O. &amp; Browning, E. (1956) British mites of the subfamily Macrochelinae Tragardh (Gamasina - Macrochelidae). Bulletin of the British Museum (Natural History), Zoology, 4, 1 - 56 + 4 plates." type="book chapter" year="1956">Evans &amp; Browning, 1956</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C30BCE5FFB7E66C8D75FB7077476B84" authority="Balogh, 1958" authorityName="Balogh" authorityYear="1958" box="[809,1108,1160,1186]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="evansi">
<emphasis id="B9441B74FFB7E66C8D75FB7070526B87" box="[809,833,1160,1185]" italics="true" pageId="12" pageNumber="49">M</emphasis>
.
<emphasis id="B9441B74FFB7E66C8D0FFB7070B26B87" box="[851,929,1160,1185]" italics="true" pageId="12" pageNumber="49">evansi</emphasis>
<bibRefCitation id="EFA1BA97FFB7E66C8DF6FB7077476B84" author="Balogh" box="[938,1108,1160,1186]" pageId="12" pageNumber="51" refString="Balogh, J. (1958) Macrocheliden aus Bulgarien (Acarina, Mesostigmata). Acta entomologica Musei Nationalis Prague, 32, 247 - 256." type="journal article" year="1958">Balogh, 1958</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C30BCE5FFB7E66C8A3EFB7073F36BE1" authority="Bregetova, 1977" authorityName="Bregetova" authorityYear="1977" class="Arachnida" family="Macrochelidae" genus="Macrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="georgicus" status="comb. nov.">
<emphasis id="B9441B74FFB7E66C8A3EFB7077696B87" box="[1122,1146,1160,1185]" italics="true" pageId="12" pageNumber="49">M</emphasis>
.
<emphasis id="B9441B74FFB7E66C8AD7FB7076106B87" box="[1163,1283,1160,1185]" italics="true" pageId="12" pageNumber="49">georgicus</emphasis>
(
<bibRefCitation id="EFA1BA97FFB7E66C8B49FB7073C46BE1" author="Bregetova" pageId="12" pageNumber="51" refString="Bregetova, N. G. (1977) Sem. Macrochelidae. In: Bregetova, N. G., Vainshtein, B. A., Kadite, B. A., Koroleva, E. V., Petrova, A. D., Tikhomirov, S. I. &amp; Shcherbak, G. I. (Eds.), Opredelitel obitayushchikh v pochve kleshchei, Mesostigmata. Izdatelstvo Nauka, Leningrad, pp. 621 - 632. [in Russian]." type="book chapter" year="1977">Bregetova, 1977</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A277A60FFFB7E66C8EB1FB5572D06BE1" box="[237,451,1197,1223]" pageId="12" pageNumber="49">new combination</taxonomicNameLabel>
;
<taxonomicName id="4C30BCE5FFB7E66C8F8CFB5570406BE1" authority="(Ishikawa, 1969)" baseAuthorityName="Ishikawa" baseAuthorityYear="1969" box="[464,851,1197,1223]" class="Arachnida" family="Macrochelidae" genus="Macrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="morikawai">
<emphasis id="B9441B74FFB7E66C8F8CFB5572FB6BE0" box="[464,488,1197,1222]" italics="true" pageId="12" pageNumber="49">M</emphasis>
.
<emphasis id="B9441B74FFB7E66C8FA5FB5571696BE0" box="[505,634,1197,1222]" italics="true" pageId="12" pageNumber="49">morikawai</emphasis>
(
<bibRefCitation id="EFA1BA97FFB7E66C8CD6FB55705A6BE1" author="Ishikawa" box="[650,841,1197,1223]" pageId="12" pageNumber="52" refString="Ishikawa, K. (1969) Taxonomic investigations on free-living mites in the subalpine forest on Shiga Heights IBP area. I. Mesostigmata (Part 1). Bulletin of the National Science Museum, Tokyo, 12, 39 - 64." type="journal article" year="1969">Ishikawa, 1969</bibRefCitation>
)
</taxonomicName>
, new combination;
<taxonomicName id="4C30BCE5FFB7E66C8A1EFB55767D6BE1" authority="Koch, 1839" authorityName="Koch" authorityYear="1839" box="[1090,1390,1197,1223]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="opacus">
<emphasis id="B9441B74FFB7E66C8A1EFB5577496BE0" box="[1090,1114,1197,1222]" italics="true" pageId="12" pageNumber="49">M</emphasis>
.
<emphasis id="B9441B74FFB7E66C8A37FB5577D36BE0" box="[1131,1216,1197,1222]" italics="true" pageId="12" pageNumber="49">opacus</emphasis>
(Koch, 1839)
</taxonomicName>
;
<taxonomicName id="4C30BCE5FFB7E66C8B20FB55715E6BCA" authority="Bregetova &amp; Koroleva, 1960" authorityName="Bregetova &amp; Koroleva" authorityYear="1960" class="Arachnida" family="Macrochelidae" genus="Macrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="recki" status="comb. nov.">
<emphasis id="B9441B74FFB7E66C8B20FB5576876BE0" box="[1404,1428,1197,1222]" italics="true" pageId="12" pageNumber="49">M</emphasis>
.
<emphasis id="B9441B74FFB7E66C8ECBFB2B73C16BCA" box="[151,210,1235,1260]" italics="true" pageId="12" pageNumber="49">recki</emphasis>
(
<bibRefCitation id="EFA1BA97FFB7E66C8EBEFB2A71566BCA" author="Bregetova" box="[226,581,1234,1260]" pageId="12" pageNumber="51" refString="Bregetova, N. G. &amp; Koroleva, E. V. (1960) The macrochelid mites (Gamasoidea, Macrochelidae) in the USSR. Akademiia Nauk SSSR Zoologicheskii institut Parazitologicheskii Sbornik, 29, 32 - 154. [in Russian]." type="journal article" year="1960">Bregetova &amp; Koroleva, 1960</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A277A60FFFB7E66C8C0AFB2A70396BCA" box="[598,810,1234,1260]" pageId="12" pageNumber="49">new combination</taxonomicNameLabel>
;
<taxonomicName id="4C30BCE5FFB7E66C8D6BFB2B779D6BCA" authority="Petrova, 1966" authorityName="Petrova" authorityYear="1966" box="[823,1166,1234,1260]" class="Arachnida" family="Macrochelidae" genus="Macrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="reductus" status="comb. nov.">
<emphasis id="B9441B74FFB7E66C8D6BFB2B705C6BCA" box="[823,847,1235,1260]" italics="true" pageId="12" pageNumber="49">M</emphasis>
.
<emphasis id="B9441B74FFB7E66C8D3CFB2B70D46BCA" box="[864,967,1235,1260]" italics="true" pageId="12" pageNumber="49">reductus</emphasis>
(
<bibRefCitation id="EFA1BA97FFB7E66C8D8BFB2A77976BCA" author="Petrova" box="[983,1156,1234,1260]" pageId="12" pageNumber="53" refString="Petrova, A. D. (1966) Macrocheles reductus a new species of gamasoid mite from the southern coast (Gamasoidea, Macrochelidae). Nauchnye Doklady Vysshei Shkoly: Biologicheski Nauki, 9, 18 - 23. [in Russian]." type="journal article" year="1966">Petrova, 1966</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A277A60FFFB7E66C8AC7FB2A767C6BCA" box="[1179,1391,1234,1260]" pageId="12" pageNumber="49">new combination</taxonomicNameLabel>
;
<taxonomicName id="4C30BCE5FFB7E66C8B20FB2B72F86A34" authority="Masan, 2003" authorityName="Masan" authorityYear="2003" class="Arachnida" family="Macrochelidae" genus="Macrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="similiopacus" status="comb. nov.">
<emphasis id="B9441B74FFB7E66C8B20FB2B76876BCA" box="[1404,1428,1235,1260]" italics="true" pageId="12" pageNumber="49">M</emphasis>
.
<emphasis id="B9441B74FFB7E66C8ECBFB00723C6A37" box="[151,303,1272,1297]" italics="true" pageId="12" pageNumber="49">similiopacus</emphasis>
(
<bibRefCitation id="EFA1BA97FFB7E66C8F1CFB0072F26A34" author="Masan" box="[320,481,1272,1298]" pageId="12" pageNumber="53" refString="Masan, P. (2003) Macrochelid mites of Slovakia (Acari: Mesostigmata: Macrochelidae). Institute of Zoology, Slovak Academy of Sciences, Bratislava. 149 pp." type="book" year="2003">Mašán, 2003</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A277A60FFFB7E66C8FA4FB0071DF6A34" box="[504,716,1272,1298]" pageId="12" pageNumber="49">new combination</taxonomicNameLabel>
;
<taxonomicName id="4C30BCE5FFB7E66C8C85FB0077226A34" authority="Krauss, 1970" authorityName="Krauss" authorityYear="1970" box="[729,1073,1272,1298]" class="Arachnida" family="Macrochelidae" genus="Macrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="stammeri" status="comb. nov.">
<emphasis id="B9441B74FFB7E66C8C85FB0071E26A37" box="[729,753,1272,1297]" italics="true" pageId="12" pageNumber="49">M</emphasis>
.
<emphasis id="B9441B74FFB7E66C8D5EFB0070626A37" box="[770,881,1272,1297]" italics="true" pageId="12" pageNumber="49">stammeri</emphasis>
(
<bibRefCitation id="EFA1BA97FFB7E66C8DDEFB00773B6A34" author="Krauss" box="[898,1064,1272,1298]" pageId="12" pageNumber="52" refString="Krauss, W. (1970) Die europaischen Arten der Gattungen Macrocheles Latreille 1829 und Geholaspis Berlese 1918. (Eine systematische Studie aus dem Jahr 1960). Acarologie, Schriftenreihe fur Vergleichende Milbenkunde, 14, 1 - 43 + 20 plates." type="book chapter" year="1970">Krauss, 1970</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A277A60FFFB7E66C8A62FB0076016A34" box="[1086,1298,1272,1298]" pageId="12" pageNumber="49">new combination</taxonomicNameLabel>
;
<taxonomicName id="4C30BCE5FFB7E66C8B43FB0072EB6A11" authority="Canestrini &amp; Fanzago, 1877" authorityName="Canestrini &amp; Fanzago" authorityYear="1877" class="Arachnida" family="Macrochelidae" genus="Macrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="terreus" status="comb. nov.">
<emphasis id="B9441B74FFB7E66C8B43FB0076246A37" box="[1311,1335,1272,1297]" italics="true" pageId="12" pageNumber="49">M</emphasis>
.
<emphasis id="B9441B74FFB7E66C8B14FB00768F6A37" box="[1352,1436,1272,1297]" italics="true" pageId="12" pageNumber="49">terreus</emphasis>
(
<bibRefCitation id="EFA1BA97FFB7E66C8EC3FAE572FD6A11" author="Canestrini" box="[159,494,1309,1335]" pageId="12" pageNumber="51" refString="Canestrini, G., &amp; Fanzago, F. (1877) Intorno agli Acari italiani. Atti del Reale Istituto Veneto di Scienze, Lettere ed Arti (series 5) 4, 69 - 208." type="book chapter" year="1877">Canestrini &amp; Fanzago, 1877</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A277A60FFFB7E66C8C5FFAE571C26A11" box="[515,721,1309,1335]" pageId="12" pageNumber="49">new combination</taxonomicNameLabel>
;
<taxonomicName id="4C30BCE5FFB7E66C8C81FAE577966A11" authority="Bregetova, 1977" authorityName="Bregetova" authorityYear="1977" box="[733,1157,1309,1335]" class="Arachnida" family="Macrochelidae" genus="Macrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="tianschanicus" status="comb. nov.">
<emphasis id="B9441B74FFB7E66C8C81FAE571E66A10" box="[733,757,1309,1334]" italics="true" pageId="12" pageNumber="49">M</emphasis>
.
<emphasis id="B9441B74FFB7E66C8D58FAE570B66A10" box="[772,933,1309,1334]" italics="true" pageId="12" pageNumber="49">tianschanicus</emphasis>
(
<bibRefCitation id="EFA1BA97FFB7E66C8DE9FAE577686A11" author="Bregetova" box="[949,1147,1309,1335]" pageId="12" pageNumber="51" refString="Bregetova, N. G. (1977) Sem. Macrochelidae. In: Bregetova, N. G., Vainshtein, B. A., Kadite, B. A., Koroleva, E. V., Petrova, A. D., Tikhomirov, S. I. &amp; Shcherbak, G. I. (Eds.), Opredelitel obitayushchikh v pochve kleshchei, Mesostigmata. Izdatelstvo Nauka, Leningrad, pp. 621 - 632. [in Russian]." type="book chapter" year="1977">Bregetova, 1977</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A277A60FFFB7E66C8ACCFAE5764C6A11" box="[1168,1375,1309,1335]" pageId="12" pageNumber="49">new combination</taxonomicNameLabel>
.
</paragraph>
</subSubSection>
<subSubSection id="C32A94EDFFB7E66C8E9AFABA72DF691A" pageId="12" pageNumber="49" type="description">
<paragraph id="8B8FC766FFB7E66C8E9AFABA72DF691A" blockId="12.[151,1437,152,1596]" pageId="12" pageNumber="49">
<collectingCountry id="F32787F6FFB7E66C8E9AFABA73F46A7A" box="[198,231,1346,1372]" name="American Samoa" pageId="12" pageNumber="49">As</collectingCountry>
pointed out by
<bibRefCitation id="EFA1BA97FFB7E66C8FFAFABA715D6A7A" author="Masan" box="[422,590,1346,1372]" pageId="12" pageNumber="53" refString="Masan, P. (2003) Macrochelid mites of Slovakia (Acari: Mesostigmata: Macrochelidae). Institute of Zoology, Slovak Academy of Sciences, Bratislava. 149 pp." type="book" year="2003">Mašán (2003)</bibRefCitation>
,
<taxonomicName id="4C30BCE5FFB7E66C8C00FABB77196A7A" authority="Berlese" authorityName="Berlese" box="[604,1034,1346,1372]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="punctatissimus">
<emphasis id="B9441B74FFB7E66C8C00FABB70BA6A7A" box="[604,937,1347,1372]" italics="true" pageId="12" pageNumber="49">Macrocheles punctatissimus</emphasis>
Berlese
</taxonomicName>
is not a species of
<taxonomicName id="4C30BCE5FFB7E66C8AA8FABB76866A7A" box="[1268,1429,1347,1372]" class="Arachnida" family="Macrochelidae" genus="Macrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFB7E66C8AA8FABB76866A7A" box="[1268,1429,1347,1372]" italics="true" pageId="12" pageNumber="49">Macrholaspis</emphasis>
</taxonomicName>
, although it was tentatively included in the
<taxonomicName id="4C30BCE5FFB7E66C8CC7FA9071E36AA7" box="[667,752,1384,1409]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="opacus">
<emphasis id="B9441B74FFB7E66C8CC7FA9071E36AA7" box="[667,752,1384,1409]" italics="true" pageId="12" pageNumber="49">opacus</emphasis>
</taxonomicName>
species group by
<bibRefCitation id="EFA1BA97FFB7E66C8D90FA9076186AA4" author="Hyatt" box="[972,1291,1384,1410]" pageId="12" pageNumber="52" refString="Hyatt, K. H. &amp; Emberson, R. M. (1988) A review of the Macrochelidae (Acari: Mesostigmata) of the British Isles. Bulletin of the British Museum (Natural History), Zoology, 54, 63 - 125 + 6 plates." type="book chapter" year="1988">Hyatt &amp; Emberson (1988)</bibRefCitation>
. It lacks the definitive characters of the genus, particularly the reduced ventral sclerotisation, the free post-epigynal platelets, the reduced lateral processes of the epistome, and the pectinate cheliceral dorsal seta.
<taxonomicName id="4C30BCE5FFB7E66C8B59FA4B72DC6AD4" authority="Hyatt &amp; Emberson" authorityName="Hyatt &amp; Emberson" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="analis">
<emphasis id="B9441B74FFB7E66C8B59FA4B73F26AD7" italics="true" pageId="12" pageNumber="49">Macrocheles analis</emphasis>
Hyatt &amp; Emberson
</taxonomicName>
, also placed in the
<taxonomicName id="4C30BCE5FFB7E66C8CE4FA20701E6AD7" box="[696,781,1496,1521]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="opacus">
<emphasis id="B9441B74FFB7E66C8CE4FA20701E6AD7" box="[696,781,1496,1521]" italics="true" pageId="12" pageNumber="49">opacus</emphasis>
</taxonomicName>
species group by
<bibRefCitation id="EFA1BA97FFB7E66C8DB4FA2076356AD4" author="Hyatt" box="[1000,1318,1496,1522]" pageId="12" pageNumber="52" refString="Hyatt, K. H. &amp; Emberson, R. M. (1988) A review of the Macrochelidae (Acari: Mesostigmata) of the British Isles. Bulletin of the British Museum (Natural History), Zoology, 54, 63 - 125 + 6 plates." type="book chapter" year="1988">Hyatt &amp; Emberson (1988)</bibRefCitation>
, has been transferred to
<taxonomicName id="4C30BCE5FFB7E66C8F60FA0572FC6930" box="[316,495,1533,1558]" class="Arachnida" family="Macrochelidae" genus="Reductholaspis" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="45" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9441B74FFB7E66C8F60FA0572FC6930" box="[316,495,1533,1558]" italics="true" pageId="12" pageNumber="49">Reductholaspis</emphasis>
</taxonomicName>
, a distinctive new genus apparently more closely related to
<taxonomicName id="4C30BCE5FFB7E66C8AE9FA0576506930" box="[1205,1347,1533,1558]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFB7E66C8AE9FA0576506930" box="[1205,1347,1533,1558]" italics="true" pageId="12" pageNumber="49">Longicheles</emphasis>
</taxonomicName>
than to
<taxonomicName id="4C30BCE5FFB7E66C8ECBF9DB722B691A" box="[151,312,1571,1596]" class="Arachnida" family="Macrochelidae" genus="Macrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFB7E66C8ECBF9DB722B691A" box="[151,312,1571,1596]" italics="true" pageId="12" pageNumber="49">Macrholaspis</emphasis>
</taxonomicName>
(see above).
</paragraph>
</subSubSection>
</treatment>
</document>
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