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<document id="8B4E1EF8E14ED7ADA9381A990FA20869" ID-DOI="10.11646/zootaxa.4013.3.4" ID-GBIF-Dataset="d8878b8c-0bd2-446f-a5eb-52c83620e3ad" ID-ISSN="1175-5326" ID-Zenodo-Dep="241262" ID-ZooBank="1F9B736F-25DF-47A9-9D16-CC9ADBDFD8E6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461248028428" checkinUser="plazi" docAuthor="Komai, Tomoyuki, Okuno, Junji &amp; Minemizu, Ryo" docDate="2015" docId="03959A028A646B0BFF2EFC2CFD88F98A" docLanguage="en" docName="zt04013p412.pdf" docOrigin="Zootaxa 4013 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thor leptochelus Xu &amp; Li 2015, n. comb." docType="treatment" docVersion="8" lastPageNumber="406" masterDocId="FFACE27A8A656B0CFFB9FFB3FFCBFFFE" masterDocTitle="New records of two species of the coral reef shrimp genus Thor Kingsley, 1878 (Crustacea: Decapoda: Thoridae) from the Ryukyu Islands, Japan" masterLastPageNumber="412" masterPageNumber="399" pageNumber="400" updateTime="1698606911097" updateUser="plazi">
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<mods:title id="EE644F39003CB66B90D66F9E9774BFBC">New records of two species of the coral reef shrimp genus Thor Kingsley, 1878 (Crustacea: Decapoda: Thoridae) from the Ryukyu Islands, Japan</mods:title>
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<mods:namePart id="C58E70B9961AF4D83FB0888B9834B9AD">Komai, Tomoyuki</mods:namePart>
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<mods:namePart id="2A8027A9EDDD442E64D803CAECB2467B">Okuno, Junji</mods:namePart>
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<mods:namePart id="9ACC763C2DB0FEA3707BAC3D46CD0252">Minemizu, Ryo</mods:namePart>
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<mods:title id="8081899A0DF70BF4992B02AE52FBB463">Zootaxa</mods:title>
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<mods:date id="D697EB7F98FB43E0AC3A8E4DAAFAD786">2015</mods:date>
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<mods:number id="391CE77DD34BECD42771639017DE5F75">4013</mods:number>
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<mods:identifier id="1AAD2B2261B7911C45FD0FB8D5C7148D" type="DOI">10.11646/zootaxa.4013.3.4</mods:identifier>
<mods:identifier id="2E8D3567DB65548510D4A25439CD9012" type="GBIF-Dataset">d8878b8c-0bd2-446f-a5eb-52c83620e3ad</mods:identifier>
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<treatment id="03959A028A646B0BFF2EFC2CFD88F98A" ID-DOI="http://doi.org/10.5281/zenodo.6112410" ID-GBIF-Taxon="119635244" ID-Zenodo-Dep="6112410" LSID="urn:lsid:plazi:treatment:03959A028A646B0BFF2EFC2CFD88F98A" httpUri="http://treatment.plazi.org/id/03959A028A646B0BFF2EFC2CFD88F98A" lastPageId="7" lastPageNumber="406" pageId="1" pageNumber="400">
<subSubSection id="C326789F8A646B0DFF2EFC2CFEE4FC00" pageId="1" pageNumber="400" type="nomenclature">
<paragraph id="8B832B148A646B0DFF2EFC2CFD6AFC47" blockId="1.[151,721,927,1022]" box="[151,673,927,953]" pageId="1" pageNumber="400">
<heading id="D0CB9C788A646B0DFF2EFC2CFD6AFC47" bold="true" box="[151,673,927,953]" fontSize="11" level="1" pageId="1" pageNumber="400" reason="1">
<emphasis id="B948F7068A646B0DFF2EFC2CFD6AFC47" bold="true" box="[151,673,927,953]" pageId="1" pageNumber="400">
<taxonomicName id="4C3C50978A646B0DFF2EFC2CFDE4FC47" authority="Xu &amp; Li, 2015" authorityName="Xu &amp; Li" authorityYear="2015" box="[151,559,927,953]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="400" phylum="Arthropoda" rank="species" species="leptochelus" status="comb. nov.">
<emphasis id="B948F7068A646B0DFF2EFC2CFEA8FC47" bold="true" box="[151,355,927,953]" italics="true" pageId="1" pageNumber="400">Thor leptochelus</emphasis>
(
<bibRefCitation id="EFAD56E58A646B0DFECBFC2CFDECFC47" author="Xu" box="[370,551,927,953]" pageId="1" pageNumber="400" refString="Xu, P. &amp; Li, X. (2015) Report on the Hippolytidae Bate (sensu lato) from China seas. Zoological Systematics, 40, 107 - 165." type="journal article" year="2015">Xu &amp; Li, 2015</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A27B4A7D8A646B0DFD8FFC2CFD6AFC47" box="[566,673,927,953]" pageId="1" pageNumber="400">n. comb.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B832B148A646B0DFF2EFC71FEE4FC00" blockId="1.[151,721,927,1022]" pageId="1" pageNumber="400">
[New Japanese name: Zunguri-hime-sango-moebi]
<figureCitation id="130737918A646B0DFF2EFC56FECAFC00" box="[151,257,997,1022]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,250,1312,1334]" captionStartId-1="4.[151,250,1822,1844]" captionStartId-2="5.[151,250,1935,1957]" captionStartId-3="6.[151,250,1083,1105]" captionTargetBox-0="[157,1409,336,1311]" captionTargetBox-1="[238,1348,213,1793]" captionTargetBox-2="[197,1388,220,1891]" captionTargetBox-3="[167,1388,276,956]" captionTargetId-0="figure@2.[157,1477,304,1317]" captionTargetId-1="figure@4.[212,1372,188,1805]" captionTargetId-2="figure@5.[172,1414,198,1920]" captionTargetId-3="figure@6.[165,1430,244,1003]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 1. Thor leptochelus (Xu &amp; Li, 2015) n. comb., ovigerous female (cl 2.3 mm), CBM-ZC 13091, habitus, lateral view. Scale bar: 2 mm." captionText-1="FIGURE 2. Thor leptochelus (Xu &amp; Li, 2015) n. comb. A, B, D – N, ovigerous female (cl 2.3 mm), CBM-ZC 13091; C, paratype, female (cl 1.5 mm), CBM-ZC 13090. Mouthparts dissected from right side. A, carapace and eyes, dorsal view; B, rostrum and anterior part of carapace, lateral view; C, rostrum, dorsolateral view; D, telson and right uropod, dorsal view (setae omitted); E, posterior part of telson, dorsal view; F, right antennular peduncle and antennal basicerite, lateral view; G, right antennular peduncle, dorsolateral view; H, right antenna, ventral view (flagellum omitted); I, mandible, external view; J, same, internal view; K, maxillule, outer view (proximal endite missing); L, maxilla, external view; M, first maxilliped, external view; N, second maxilliped, external view. Scale bars: 1 mm for A, B, F; 0.5 mm for C – E, G – N." captionText-2="FIGURE 3. Thor leptochelus (Xu &amp; Li, 2015) n. comb., ovigerous female (cl 2.3 mm), CBM-ZC 13091. Thoracic appendages in lateral view. A, right third maxilliped; B, right first pereopod; C, right second pereopod; D, left third pereopod; E, right third pereopod, dactylus and propodus; F, right fourth pereopod; G, right fifth pereopod. Scale bar: 1 mm." captionText-3="FIGURE 4. Thor leptochelus (Xu &amp; Li, 2015) n. comb., ovigerous female (cl 2.3 mm), CBM-ZC 13091. A, right third maxilliped, distal part of ultimate segment, showing spination; B, same, ultimate segment (= carpus) and distal part of antepenultimate segment, mesial view, showing setation and armature; C, same, detail of distolateral armature of antepenultimate segment; D, right first pereopod, chela and distal part of carpus, flexor view; E, same, merus and ischium, mesial view; F, left third pereopod, dactylus and distal part of propodus, lateral view; G, same, distal part of propodus, showing armature on flexor surface; H, right fourth pereopod, dactylus and distal part of propodus, lateral view; G, right fifth pereopod, dactylus and distal part of propodus, lateral view. Scale bars: 0.5 mm." httpUri-0="https://zenodo.org/record/241263/files/figure.png" httpUri-1="https://zenodo.org/record/241264/files/figure.png" httpUri-2="https://zenodo.org/record/241265/files/figure.png" httpUri-3="https://zenodo.org/record/241266/files/figure.png" pageId="1" pageNumber="400">Figs. 1–4</figureCitation>
,
<figureCitation id="130737918A646B0DFEB4FC56FEE4FC00" box="[269,303,997,1022]" captionStart="FIGURE 5. A" captionStartId="9.[151,250,1917,1939]" captionTargetBox="[172,1409,195,1897]" captionTargetId="figure@9.[172,1410,195,1897]" captionTargetPageId="9" captionText="FIGURE 5. A, Thor leptochelus (Xu &amp; Li, 2015) n. comb., ovigerous female (cl 2.3 mm), CBM-ZC 13091, in situ; B, Thor marguitae Bruce, 1978, ovigerous female (cl 2.7 mm), CBM-ZC 11889, in situ, attaching to solitary fungiid coral Fungia repanda. Photography by R. Minemizu." httpUri="https://zenodo.org/record/241267/files/figure.png" pageId="1" pageNumber="400">5A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C326789F8A646B0DFF2EFB9EFA9DFB4C" pageId="1" pageNumber="400" type="materials_examined">
<paragraph id="8B832B148A646B0DFF2EFB9EFA9DFB4C" blockId="1.[151,1437,1069,2025]" pageId="1" pageNumber="400">
<emphasis id="B948F7068A646B0DFF2EFB9EFEB2FBB8" bold="true" box="[151,377,1069,1094]" pageId="1" pageNumber="400">Material examined</emphasis>
. Cape Maeda, Onna, Okinawa Island,
<quantity id="4CC486F18A646B0DFC94FB9DFC91FBBB" box="[813,858,1070,1093]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="400" unit="m" value="7.0">7 m</quantity>
,
<date id="FF820DD48A646B0DFCDFFB9DFC34FBB8" box="[870,1023,1070,1094]" pageId="1" pageNumber="400" value="2012-06-23">23 June 2012</date>
,
<collectionCode id="ED2DB3D18A646B0DFBB3FB9DFBACFBB8" box="[1034,1127,1070,1094]" pageId="1" pageNumber="400">SCUBA</collectionCode>
diving, coll. N. Shirakawa,
<specimenCount id="9D3AE09D8A646B0DFF2EFBE1FF33FB94" box="[151,248,1105,1130]" pageId="1" pageNumber="400" type="female">1 female</specimenCount>
(cl
<quantity id="4CC486F18A646B0DFE9DFBE2FEB5FB94" box="[292,382,1105,1130]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="400" unit="mm" value="1.5">1.5 mm</quantity>
), 1 ovigerous female (cl
<quantity id="4CC486F18A646B0DFD2CFBE2FD24FB94" box="[661,751,1105,1130]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="400" unit="mm" value="1.6">1.6 mm</quantity>
), CBM-ZC 13090; Sunabe, Chatan, Okinawa Island,
<quantity id="4CC486F18A646B0DFAFEFBE1FABFFB97" box="[1351,1396,1106,1130]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="400" unit="m" value="8.0">8 m</quantity>
,
<date id="FF820DD48A646B0DFA39FBE2FE99FB70" pageId="1" pageNumber="400" value="2013-11-15">15 November 2013</date>
,
<collectionCode id="ED2DB3D18A646B0DFED8FBC5FE75FB70" box="[353,446,1142,1166]" pageId="1" pageNumber="400">SCUBA</collectionCode>
diving, coll. N. Shirakawa, ovigerous female (cl
<quantity id="4CC486F18A646B0DFBBAFBC5FBAAFB70" box="[1027,1121,1142,1166]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="400" unit="mm" value="2.3">2.3 mm</quantity>
), CBM-ZC 13091; Shichugama, Kume Island,
<quantity id="4CC486F18A646B0DFEC6FB29FE67FB4C" box="[383,428,1178,1202]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="400" unit="m" value="8.0">8 m</quantity>
,
<date id="FF820DD48A646B0DFE01FB29FD89FB4C" box="[440,578,1178,1202]" pageId="1" pageNumber="400" value="2002-06-08">8 June 2002</date>
, coll. T. Kawamoto, 1 ovigerous female (cl
<quantity id="4CC486F18A646B0DFB89FB29FB41FB4C" box="[1072,1162,1178,1202]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="400" unit="mm" value="1.8">1.8 mm</quantity>
), CMNH-ZC 997.
</paragraph>
</subSubSection>
<subSubSection id="C326789F8A646B0DFF7EFB0BFC07FA0F" pageId="1" pageNumber="400" type="diagnosis">
<paragraph id="8B832B148A646B0DFF7EFB0BFC07FA0F" blockId="1.[151,1437,1069,2025]" pageId="1" pageNumber="400">
<emphasis id="B948F7068A646B0DFF7EFB0BFEF1FB2F" bold="true" box="[199,314,1208,1233]" pageId="1" pageNumber="400">Diagnosis</emphasis>
. Body very robust in adult female. Rostrum short, about half-length of carapace; distal part compressed laterally, proximal part abruptly widened in dorsal view; dorsal margin armed with 1–3 teeth, none postrostral; tip acute; ventral margin unarmed or armed with 1 tooth subterminally. Carapace with pair of supraorbital teeth; suborbital angle well defined, subacute; pterygostomial angle rounded, unarmed. Pleura of fourth and fifth pleomeres each with posteroventral tooth. Telson with 3 pairs of dorsolateral spines; posterior margin convex, with 2 pairs of spines and 1 mesial pair of plumose spines. Antennular stylocerite unarmed on dorsolateral margin proximally. No strap-like epipods on third maxilliped and pereopods. First pereopod with slender chela tapered distally and longer than carpus; merus unarmed in distal half of ventral margin. Meri of third and fourth pereopods each with 1 spine subdistally, that of fifth unarmed.
</paragraph>
</subSubSection>
<subSubSection id="C326789F8A646B0FFF7EFA4FFC52F820" lastPageId="3" lastPageNumber="402" pageId="1" pageNumber="400" type="description">
<paragraph id="8B832B148A646B0DFF7EFA4FFA59F9EB" blockId="1.[151,1437,1069,2025]" box="[199,1426,1532,1557]" pageId="1" pageNumber="400">
<emphasis id="B948F7068A646B0DFF7EFA4FFE9AF9EB" bold="true" box="[199,337,1532,1557]" pageId="1" pageNumber="400">Description</emphasis>
. Body (
<figureCitation id="130737918A646B0DFE12FA4FFE27F9EB" box="[427,492,1532,1557]" captionStart="FIGURE 1" captionStartId="2.[151,250,1312,1334]" captionTargetBox="[157,1409,336,1311]" captionTargetId="figure@2.[157,1477,304,1317]" captionTargetPageId="2" captionText="FIGURE 1. Thor leptochelus (Xu &amp; Li, 2015) n. comb., ovigerous female (cl 2.3 mm), CBM-ZC 13091, habitus, lateral view. Scale bar: 2 mm." httpUri="https://zenodo.org/record/241263/files/figure.png" pageId="1" pageNumber="400">Fig. 1</figureCitation>
) very robust particularly in ovigerous females, carapace height greater than length.
</paragraph>
<paragraph id="8B832B148A646B0DFF7EF992FF31F8A7" blockId="1.[151,1437,1069,2025]" pageId="1" pageNumber="400">
Rostrum (
<figureCitation id="130737918A646B0DFE84F993FE4EF9C6" box="[317,389,1568,1593]" captionStart="FIGURE 2" captionStartId="4.[151,250,1822,1844]" captionTargetBox="[238,1348,213,1793]" captionTargetId="figure@4.[212,1372,188,1805]" captionTargetPageId="4" captionText="FIGURE 2. Thor leptochelus (Xu &amp; Li, 2015) n. comb. A, B, D – N, ovigerous female (cl 2.3 mm), CBM-ZC 13091; C, paratype, female (cl 1.5 mm), CBM-ZC 13090. Mouthparts dissected from right side. A, carapace and eyes, dorsal view; B, rostrum and anterior part of carapace, lateral view; C, rostrum, dorsolateral view; D, telson and right uropod, dorsal view (setae omitted); E, posterior part of telson, dorsal view; F, right antennular peduncle and antennal basicerite, lateral view; G, right antennular peduncle, dorsolateral view; H, right antenna, ventral view (flagellum omitted); I, mandible, external view; J, same, internal view; K, maxillule, outer view (proximal endite missing); L, maxilla, external view; M, first maxilliped, external view; N, second maxilliped, external view. Scale bars: 1 mm for A, B, F; 0.5 mm for C – E, G – N." httpUri="https://zenodo.org/record/241264/files/figure.png" pageId="1" pageNumber="400">Fig. 2</figureCitation>
A, B) short, about half-length of carapace, directed forward, not reaching distal end of antennular peduncle, strongly widened both transversely and vertically at base; dorsal margin sloping anteriorly, armed with 1–3 teeth, rostral carina following last tooth extending slightly beyond level of posterior margin of orbit; tip acute; ventral margin with 1 conspicuous subterminal tooth or unarmed (
<figureCitation id="130737918A646B0DFBF7F93FFB5FF95B" box="[1102,1172,1676,1701]" captionStart="FIGURE 2" captionStartId="4.[151,250,1822,1844]" captionTargetBox="[238,1348,213,1793]" captionTargetId="figure@4.[212,1372,188,1805]" captionTargetPageId="4" captionText="FIGURE 2. Thor leptochelus (Xu &amp; Li, 2015) n. comb. A, B, D – N, ovigerous female (cl 2.3 mm), CBM-ZC 13091; C, paratype, female (cl 1.5 mm), CBM-ZC 13090. Mouthparts dissected from right side. A, carapace and eyes, dorsal view; B, rostrum and anterior part of carapace, lateral view; C, rostrum, dorsolateral view; D, telson and right uropod, dorsal view (setae omitted); E, posterior part of telson, dorsal view; F, right antennular peduncle and antennal basicerite, lateral view; G, right antennular peduncle, dorsolateral view; H, right antenna, ventral view (flagellum omitted); I, mandible, external view; J, same, internal view; K, maxillule, outer view (proximal endite missing); L, maxilla, external view; M, first maxilliped, external view; N, second maxilliped, external view. Scale bars: 1 mm for A, B, F; 0.5 mm for C – E, G – N." httpUri="https://zenodo.org/record/241264/files/figure.png" pageId="1" pageNumber="400">Fig. 2</figureCitation>
C); ventrolateral carina distinct in proximal two-thirds, merging into orbital margin. Carapace (
<figureCitation id="130737918A646B0DFC0FF903FC35F937" box="[950,1022,1712,1737]" captionStart="FIGURE 1" captionStartId="2.[151,250,1312,1334]" captionTargetBox="[157,1409,336,1311]" captionTargetId="figure@2.[157,1477,304,1317]" captionTargetPageId="2" captionText="FIGURE 1. Thor leptochelus (Xu &amp; Li, 2015) n. comb., ovigerous female (cl 2.3 mm), CBM-ZC 13091, habitus, lateral view. Scale bar: 2 mm." httpUri="https://zenodo.org/record/241263/files/figure.png" pageId="1" pageNumber="400">Figs.1</figureCitation>
,
<figureCitation id="130737918A646B0DFBB3F903FBD3F936" box="[1034,1048,1712,1736]" captionStart="FIGURE 2" captionStartId="4.[151,250,1822,1844]" captionTargetBox="[238,1348,213,1793]" captionTargetId="figure@4.[212,1372,188,1805]" captionTargetPageId="4" captionText="FIGURE 2. Thor leptochelus (Xu &amp; Li, 2015) n. comb. A, B, D – N, ovigerous female (cl 2.3 mm), CBM-ZC 13091; C, paratype, female (cl 1.5 mm), CBM-ZC 13090. Mouthparts dissected from right side. A, carapace and eyes, dorsal view; B, rostrum and anterior part of carapace, lateral view; C, rostrum, dorsolateral view; D, telson and right uropod, dorsal view (setae omitted); E, posterior part of telson, dorsal view; F, right antennular peduncle and antennal basicerite, lateral view; G, right antennular peduncle, dorsolateral view; H, right antenna, ventral view (flagellum omitted); I, mandible, external view; J, same, internal view; K, maxillule, outer view (proximal endite missing); L, maxilla, external view; M, first maxilliped, external view; N, second maxilliped, external view. Scale bars: 1 mm for A, B, F; 0.5 mm for C – E, G – N." httpUri="https://zenodo.org/record/241264/files/figure.png" pageId="1" pageNumber="400">2</figureCitation>
A) with height greater than length; dorsal midline without postrostral tooth, but with minute tubercle; pair of acute supraorbital teeth present; dorsum slightly inflated posteriorly; postorbital region somewhat depressed; antennal tooth small, slightly ascending; suborbital angle produced into subacute triangular process somewhat ascending; pterygostomial angle broadly rounded.
</paragraph>
<paragraph id="8B832B148A646B0EFF7EF8D7FF03FF2B" blockId="1.[151,1437,1069,2025]" lastBlockId="2.[151,1436,151,285]" lastPageId="2" lastPageNumber="401" pageId="1" pageNumber="400">
Pleon (
<figureCitation id="130737918A646B0DFEA4F8D7FEAFF883" box="[285,356,1892,1917]" captionStart="FIGURE 1" captionStartId="2.[151,250,1312,1334]" captionTargetBox="[157,1409,336,1311]" captionTargetId="figure@2.[157,1477,304,1317]" captionTargetPageId="2" captionText="FIGURE 1. Thor leptochelus (Xu &amp; Li, 2015) n. comb., ovigerous female (cl 2.3 mm), CBM-ZC 13091, habitus, lateral view. Scale bar: 2 mm." httpUri="https://zenodo.org/record/241263/files/figure.png" pageId="1" pageNumber="400">Fig. 1</figureCitation>
) dorsally rounded. First to third pleura rounded, fourth and fifth pleura each with acute posteroventral tooth. Sixth pleomere 1.9 times as long as fifth pleomere and 1.4 times as long as proximal height. Telson (
<figureCitation id="130737918A646B0DFF4AF81FFEF3F83B" box="[243,312,1964,1989]" captionStart="FIGURE 2" captionStartId="4.[151,250,1822,1844]" captionTargetBox="[238,1348,213,1793]" captionTargetId="figure@4.[212,1372,188,1805]" captionTargetPageId="4" captionText="FIGURE 2. Thor leptochelus (Xu &amp; Li, 2015) n. comb. A, B, D – N, ovigerous female (cl 2.3 mm), CBM-ZC 13091; C, paratype, female (cl 1.5 mm), CBM-ZC 13090. Mouthparts dissected from right side. A, carapace and eyes, dorsal view; B, rostrum and anterior part of carapace, lateral view; C, rostrum, dorsolateral view; D, telson and right uropod, dorsal view (setae omitted); E, posterior part of telson, dorsal view; F, right antennular peduncle and antennal basicerite, lateral view; G, right antennular peduncle, dorsolateral view; H, right antenna, ventral view (flagellum omitted); I, mandible, external view; J, same, internal view; K, maxillule, outer view (proximal endite missing); L, maxilla, external view; M, first maxilliped, external view; N, second maxilliped, external view. Scale bars: 1 mm for A, B, F; 0.5 mm for C – E, G – N." httpUri="https://zenodo.org/record/241264/files/figure.png" pageId="1" pageNumber="400">Fig. 2</figureCitation>
D) 1.5 times as long as sixth pleomere, tapering to gently convex posterior margin, armed with 3 pairs of dorsolateral spines; posterior margin (
<figureCitation id="130737918A646B0DFD22F863FD16F816" box="[667,733,2000,2025]" captionStart="FIGURE 2" captionStartId="4.[151,250,1822,1844]" captionTargetBox="[238,1348,213,1793]" captionTargetId="figure@4.[212,1372,188,1805]" captionTargetPageId="4" captionText="FIGURE 2. Thor leptochelus (Xu &amp; Li, 2015) n. comb. A, B, D – N, ovigerous female (cl 2.3 mm), CBM-ZC 13091; C, paratype, female (cl 1.5 mm), CBM-ZC 13090. Mouthparts dissected from right side. A, carapace and eyes, dorsal view; B, rostrum and anterior part of carapace, lateral view; C, rostrum, dorsolateral view; D, telson and right uropod, dorsal view (setae omitted); E, posterior part of telson, dorsal view; F, right antennular peduncle and antennal basicerite, lateral view; G, right antennular peduncle, dorsolateral view; H, right antenna, ventral view (flagellum omitted); I, mandible, external view; J, same, internal view; K, maxillule, outer view (proximal endite missing); L, maxilla, external view; M, first maxilliped, external view; N, second maxilliped, external view. Scale bars: 1 mm for A, B, F; 0.5 mm for C – E, G – N." httpUri="https://zenodo.org/record/241264/files/figure.png" pageId="1" pageNumber="400">Fig. 2</figureCitation>
E) without median tooth, but with 3 pairs of spines, lateral pair at posterolateral angle of telson short, intermediate pair longest, mesial pair plumose, shorter than intermediate pair.
</paragraph>
<paragraph id="8B832B148A676B0EFF7EFF53FC29FEE3" blockId="2.[151,1436,151,285]" pageId="2" pageNumber="401">
Eye (
<figureCitation id="130737918A676B0EFEBBFF6CFE81FF06" box="[258,330,223,248]" captionStart="FIGURE 1" captionStartId="2.[151,250,1312,1334]" captionTargetBox="[157,1409,336,1311]" captionTargetId="figure@2.[157,1477,304,1317]" captionTargetPageId="2" captionText="FIGURE 1. Thor leptochelus (Xu &amp; Li, 2015) n. comb., ovigerous female (cl 2.3 mm), CBM-ZC 13091, habitus, lateral view. Scale bar: 2 mm." httpUri="https://zenodo.org/record/241263/files/figure.png" pageId="2" pageNumber="401">Figs 1</figureCitation>
,
<figureCitation id="130737918A676B0EFEE3FF53FEA3FF06" box="[346,360,224,248]" captionStart="FIGURE 2" captionStartId="4.[151,250,1822,1844]" captionTargetBox="[238,1348,213,1793]" captionTargetId="figure@4.[212,1372,188,1805]" captionTargetPageId="4" captionText="FIGURE 2. Thor leptochelus (Xu &amp; Li, 2015) n. comb. A, B, D – N, ovigerous female (cl 2.3 mm), CBM-ZC 13091; C, paratype, female (cl 1.5 mm), CBM-ZC 13090. Mouthparts dissected from right side. A, carapace and eyes, dorsal view; B, rostrum and anterior part of carapace, lateral view; C, rostrum, dorsolateral view; D, telson and right uropod, dorsal view (setae omitted); E, posterior part of telson, dorsal view; F, right antennular peduncle and antennal basicerite, lateral view; G, right antennular peduncle, dorsolateral view; H, right antenna, ventral view (flagellum omitted); I, mandible, external view; J, same, internal view; K, maxillule, outer view (proximal endite missing); L, maxilla, external view; M, first maxilliped, external view; N, second maxilliped, external view. Scale bars: 1 mm for A, B, F; 0.5 mm for C – E, G – N." httpUri="https://zenodo.org/record/241264/files/figure.png" pageId="2" pageNumber="401">2</figureCitation>
A) large, stout, nearly subcylindrical; cornea slightly wider and distinctly longer than eyestalk, semispherical, with ocellar spot; inner surface of eyestalk strongly inflated.
</paragraph>
<caption id="DF437B9C8A676B0EFF2EFA93FE88FAAB" httpUri="https://zenodo.org/record/241263/files/figure.png" pageId="2" pageNumber="401" targetBox="[157,1409,336,1311]" targetPageId="2">
<paragraph id="8B832B148A676B0EFF2EFA93FE88FAAB" blockId="2.[151,1436,1312,1365]" pageId="2" pageNumber="401">
<emphasis id="B948F7068A676B0EFF2EFA93FED8FACB" bold="true" box="[151,275,1312,1334]" pageId="2" pageNumber="401">FIGURE 1.</emphasis>
<taxonomicName id="4C3C50978A676B0EFEA0FA93FDBAFAC8" authority="Xu &amp; Li, 2015" authorityName="Xu &amp; Li" authorityYear="2015" box="[281,625,1312,1334]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="401" phylum="Arthropoda" rank="species" species="leptochelus" status="comb. nov.">
<emphasis id="B948F7068A676B0EFEA0FA93FE0EFAC8" box="[281,453,1312,1334]" italics="true" pageId="2" pageNumber="401">Thor leptochelus</emphasis>
(Xu &amp; Li, 2015)
</taxonomicName>
<taxonomicNameLabel id="A27B4A7D8A676B0EFDC1FA92FD18FAC8" box="[632,723,1312,1334]" pageId="2" pageNumber="401">n. comb.</taxonomicNameLabel>
, ovigerous female (cl 2.3 mm), CBM-ZC 13091, habitus, lateral view. Scale bar: 2 mm.
</paragraph>
</caption>
<paragraph id="8B832B148A676B0EFF7EFA31FA83F968" blockId="2.[151,1437,1410,2011]" pageId="2" pageNumber="401">
Antennular peduncle (
<figureCitation id="130737918A676B0EFE7FFA31FDC7FA64" box="[454,524,1410,1435]" captionStart="FIGURE 2" captionStartId="4.[151,250,1822,1844]" captionTargetBox="[238,1348,213,1793]" captionTargetId="figure@4.[212,1372,188,1805]" captionTargetPageId="4" captionText="FIGURE 2. Thor leptochelus (Xu &amp; Li, 2015) n. comb. A, B, D – N, ovigerous female (cl 2.3 mm), CBM-ZC 13091; C, paratype, female (cl 1.5 mm), CBM-ZC 13090. Mouthparts dissected from right side. A, carapace and eyes, dorsal view; B, rostrum and anterior part of carapace, lateral view; C, rostrum, dorsolateral view; D, telson and right uropod, dorsal view (setae omitted); E, posterior part of telson, dorsal view; F, right antennular peduncle and antennal basicerite, lateral view; G, right antennular peduncle, dorsolateral view; H, right antenna, ventral view (flagellum omitted); I, mandible, external view; J, same, internal view; K, maxillule, outer view (proximal endite missing); L, maxilla, external view; M, first maxilliped, external view; N, second maxilliped, external view. Scale bars: 1 mm for A, B, F; 0.5 mm for C – E, G – N." httpUri="https://zenodo.org/record/241264/files/figure.png" pageId="2" pageNumber="401">Fig. 2</figureCitation>
E, G) somewhat flattened dorsoventrally, perpendicularly erect in habitus. First segment with lateral margin expanded in rounded lobe; stylocerite acuminate, overreaching distolateral margin of third segment, slightly sinuous in lateral view, unarmed on dorsolateral margin proximally; ventral surface with low swelling proximally, mesial margin terminating in small tooth. Second segment very short, with sharp distolateral tooth. Third segment bearing triangular plate articulated at dorsodistal margin. Upper flagellum very stout and fairly long, thickened aesthetasc-bearing portion slightly longer than carapace; aesthetascs extremely thick and fairly long; slender terminal portion less than half-length of thickened portion. Lower flagellum slightly shorter than total length of upper flagellum, each article with very short seta on lower side of distal margin.
</paragraph>
<paragraph id="8B832B148A676B0EFF7EF911FBF7F8D9" blockId="2.[151,1437,1410,2011]" pageId="2" pageNumber="401">
Antennal peduncle (
<figureCitation id="130737918A676B0EFE14F911FE39F944" box="[429,498,1698,1723]" captionStart="FIGURE 2" captionStartId="4.[151,250,1822,1844]" captionTargetBox="[238,1348,213,1793]" captionTargetId="figure@4.[212,1372,188,1805]" captionTargetPageId="4" captionText="FIGURE 2. Thor leptochelus (Xu &amp; Li, 2015) n. comb. A, B, D – N, ovigerous female (cl 2.3 mm), CBM-ZC 13091; C, paratype, female (cl 1.5 mm), CBM-ZC 13090. Mouthparts dissected from right side. A, carapace and eyes, dorsal view; B, rostrum and anterior part of carapace, lateral view; C, rostrum, dorsolateral view; D, telson and right uropod, dorsal view (setae omitted); E, posterior part of telson, dorsal view; F, right antennular peduncle and antennal basicerite, lateral view; G, right antennular peduncle, dorsolateral view; H, right antenna, ventral view (flagellum omitted); I, mandible, external view; J, same, internal view; K, maxillule, outer view (proximal endite missing); L, maxilla, external view; M, first maxilliped, external view; N, second maxilliped, external view. Scale bars: 1 mm for A, B, F; 0.5 mm for C – E, G – N." httpUri="https://zenodo.org/record/241264/files/figure.png" pageId="2" pageNumber="401">Fig. 2</figureCitation>
F, H) with stout basicerite bearing moderate ventrolateral distal tooth. Carpocerite (fifth segment) short and stout, reaching midlength of antennal scale. Antennal scale (
<figureCitation id="130737918A676B0EFBE1F975FB50F921" box="[1112,1179,1734,1759]" captionStart="FIGURE 2" captionStartId="4.[151,250,1822,1844]" captionTargetBox="[238,1348,213,1793]" captionTargetId="figure@4.[212,1372,188,1805]" captionTargetPageId="4" captionText="FIGURE 2. Thor leptochelus (Xu &amp; Li, 2015) n. comb. A, B, D – N, ovigerous female (cl 2.3 mm), CBM-ZC 13091; C, paratype, female (cl 1.5 mm), CBM-ZC 13090. Mouthparts dissected from right side. A, carapace and eyes, dorsal view; B, rostrum and anterior part of carapace, lateral view; C, rostrum, dorsolateral view; D, telson and right uropod, dorsal view (setae omitted); E, posterior part of telson, dorsal view; F, right antennular peduncle and antennal basicerite, lateral view; G, right antennular peduncle, dorsolateral view; H, right antenna, ventral view (flagellum omitted); I, mandible, external view; J, same, internal view; K, maxillule, outer view (proximal endite missing); L, maxilla, external view; M, first maxilliped, external view; N, second maxilliped, external view. Scale bars: 1 mm for A, B, F; 0.5 mm for C – E, G – N." httpUri="https://zenodo.org/record/241264/files/figure.png" pageId="2" pageNumber="401">Fig. 2</figureCitation>
H) 0.9 times as long as carapace, 1.6 times as long as wide, obliquely ascending (angle against horizontal plane of carapace about 45°); lateral margin straight, distolateral tooth reaching rounded distal margin of lamella.
</paragraph>
<paragraph id="8B832B148A676B0FFF7EF881FE0FFE2E" blockId="2.[151,1437,1410,2011]" lastBlockId="3.[151,1437,151,2014]" lastPageId="3" lastPageNumber="402" pageId="2" pageNumber="401">
Mandible (
<figureCitation id="130737918A676B0EFEFBF881FE4DF8B4" box="[322,390,1842,1867]" captionStart="FIGURE 2" captionStartId="4.[151,250,1822,1844]" captionTargetBox="[238,1348,213,1793]" captionTargetId="figure@4.[212,1372,188,1805]" captionTargetPageId="4" captionText="FIGURE 2. Thor leptochelus (Xu &amp; Li, 2015) n. comb. A, B, D – N, ovigerous female (cl 2.3 mm), CBM-ZC 13091; C, paratype, female (cl 1.5 mm), CBM-ZC 13090. Mouthparts dissected from right side. A, carapace and eyes, dorsal view; B, rostrum and anterior part of carapace, lateral view; C, rostrum, dorsolateral view; D, telson and right uropod, dorsal view (setae omitted); E, posterior part of telson, dorsal view; F, right antennular peduncle and antennal basicerite, lateral view; G, right antennular peduncle, dorsolateral view; H, right antenna, ventral view (flagellum omitted); I, mandible, external view; J, same, internal view; K, maxillule, outer view (proximal endite missing); L, maxilla, external view; M, first maxilliped, external view; N, second maxilliped, external view. Scale bars: 1 mm for A, B, F; 0.5 mm for C – E, G – N." httpUri="https://zenodo.org/record/241264/files/figure.png" pageId="2" pageNumber="401">
Fig.
<date id="FF820DD48A676B0EFECEF881FE4DF8B4" box="[375,390,1842,1866]" pageId="2" pageNumber="401">2</date>
</figureCitation>
I, J) without palp; molar process moderately stout, with complex masticatory surface; incisor process slender, distally with fine, closely spaced acute teeth. Maxillule (
<figureCitation id="130737918A676B0EFC7DF8E5FBCDF891" box="[964,1030,1878,1903]" captionStart="FIGURE 2" captionStartId="4.[151,250,1822,1844]" captionTargetBox="[238,1348,213,1793]" captionTargetId="figure@4.[212,1372,188,1805]" captionTargetPageId="4" captionText="FIGURE 2. Thor leptochelus (Xu &amp; Li, 2015) n. comb. A, B, D – N, ovigerous female (cl 2.3 mm), CBM-ZC 13091; C, paratype, female (cl 1.5 mm), CBM-ZC 13090. Mouthparts dissected from right side. A, carapace and eyes, dorsal view; B, rostrum and anterior part of carapace, lateral view; C, rostrum, dorsolateral view; D, telson and right uropod, dorsal view (setae omitted); E, posterior part of telson, dorsal view; F, right antennular peduncle and antennal basicerite, lateral view; G, right antennular peduncle, dorsolateral view; H, right antenna, ventral view (flagellum omitted); I, mandible, external view; J, same, internal view; K, maxillule, outer view (proximal endite missing); L, maxilla, external view; M, first maxilliped, external view; N, second maxilliped, external view. Scale bars: 1 mm for A, B, F; 0.5 mm for C – E, G – N." httpUri="https://zenodo.org/record/241264/files/figure.png" pageId="2" pageNumber="401">Fig. 2</figureCitation>
K) with bilobed endopod, outer lobe much smaller than inner lobe, inner lobe with apparently simple, apical seta; proximal endite damaged during dissection; distal endite broad, truncate mesial margin with numerous spiniform setae. Maxilla (
<figureCitation id="130737918A676B0EFB71F82DFAC1F849" box="[1224,1290,1950,1975]" captionStart="FIGURE 2" captionStartId="4.[151,250,1822,1844]" captionTargetBox="[238,1348,213,1793]" captionTargetId="figure@4.[212,1372,188,1805]" captionTargetPageId="4" captionText="FIGURE 2. Thor leptochelus (Xu &amp; Li, 2015) n. comb. A, B, D – N, ovigerous female (cl 2.3 mm), CBM-ZC 13091; C, paratype, female (cl 1.5 mm), CBM-ZC 13090. Mouthparts dissected from right side. A, carapace and eyes, dorsal view; B, rostrum and anterior part of carapace, lateral view; C, rostrum, dorsolateral view; D, telson and right uropod, dorsal view (setae omitted); E, posterior part of telson, dorsal view; F, right antennular peduncle and antennal basicerite, lateral view; G, right antennular peduncle, dorsolateral view; H, right antenna, ventral view (flagellum omitted); I, mandible, external view; J, same, internal view; K, maxillule, outer view (proximal endite missing); L, maxilla, external view; M, first maxilliped, external view; N, second maxilliped, external view. Scale bars: 1 mm for A, B, F; 0.5 mm for C – E, G – N." httpUri="https://zenodo.org/record/241264/files/figure.png" pageId="2" pageNumber="401">Fig. 2</figureCitation>
L) with short endopod (= palp) far falling short of distal margin of basial endite; distal endite deeply bilobed, finely setose on mesial margin; proximal endite reduced, setose on mesial margin; scaphognathite with anterior lobe large and broadly rounded, posterior lobe small. First maxilliped (
<figureCitation id="130737918A666B0FFCB5FF0FFC85FF2A" box="[780,846,188,213]" captionStart="FIGURE 2" captionStartId="4.[151,250,1822,1844]" captionTargetBox="[238,1348,213,1793]" captionTargetId="figure@4.[212,1372,188,1805]" captionTargetPageId="4" captionText="FIGURE 2. Thor leptochelus (Xu &amp; Li, 2015) n. comb. A, B, D – N, ovigerous female (cl 2.3 mm), CBM-ZC 13091; C, paratype, female (cl 1.5 mm), CBM-ZC 13090. Mouthparts dissected from right side. A, carapace and eyes, dorsal view; B, rostrum and anterior part of carapace, lateral view; C, rostrum, dorsolateral view; D, telson and right uropod, dorsal view (setae omitted); E, posterior part of telson, dorsal view; F, right antennular peduncle and antennal basicerite, lateral view; G, right antennular peduncle, dorsolateral view; H, right antenna, ventral view (flagellum omitted); I, mandible, external view; J, same, internal view; K, maxillule, outer view (proximal endite missing); L, maxilla, external view; M, first maxilliped, external view; N, second maxilliped, external view. Scale bars: 1 mm for A, B, F; 0.5 mm for C – E, G – N." httpUri="https://zenodo.org/record/241264/files/figure.png" pageId="3" pageNumber="402">Fig. 2</figureCitation>
M) with endopod (= palp) very broad, uni-articulate, bearing numerous short setae along mesial margin; distal endite broad, mesially setose; proximal endite also broad, also setose on mesial margin; exopod with flagellum well developed with numerous plumose distal setae; caridean lobe small with several marginal plumose setae. epipod large, bilobed. Second maxilliped (
<figureCitation id="130737918A666B0FFB6AFE94FAD0FEBE" box="[1235,1307,295,320]" captionStart="FIGURE 2" captionStartId="4.[151,250,1822,1844]" captionTargetBox="[238,1348,213,1793]" captionTargetId="figure@4.[212,1372,188,1805]" captionTargetPageId="4" captionText="FIGURE 2. Thor leptochelus (Xu &amp; Li, 2015) n. comb. A, B, D – N, ovigerous female (cl 2.3 mm), CBM-ZC 13091; C, paratype, female (cl 1.5 mm), CBM-ZC 13090. Mouthparts dissected from right side. A, carapace and eyes, dorsal view; B, rostrum and anterior part of carapace, lateral view; C, rostrum, dorsolateral view; D, telson and right uropod, dorsal view (setae omitted); E, posterior part of telson, dorsal view; F, right antennular peduncle and antennal basicerite, lateral view; G, right antennular peduncle, dorsolateral view; H, right antenna, ventral view (flagellum omitted); I, mandible, external view; J, same, internal view; K, maxillule, outer view (proximal endite missing); L, maxilla, external view; M, first maxilliped, external view; N, second maxilliped, external view. Scale bars: 1 mm for A, B, F; 0.5 mm for C – E, G – N." httpUri="https://zenodo.org/record/241264/files/figure.png" pageId="3" pageNumber="402">Fig. 2</figureCitation>
N) normal; dactylus small and narrow, with dense spiniform setae on mesial margin; propodus broad, with slender spiniform setae on mesial margin; carpus with weak tooth on outer distal margin; merus stout; ischium and basis fused, mesial face forming shallow excavation; exopodal flagellum well developed; epipod roundly triangular, with small, non-lamellate podobranch.
</paragraph>
<paragraph id="8B832B148A666B0FFF7EFE6FFB4EFCF0" blockId="3.[151,1437,151,2014]" pageId="3" pageNumber="402">
Third maxilliped (
<figureCitation id="130737918A666B0FFE27FE6FFE2EFE0A" box="[414,485,476,501]" captionStart="FIGURE 3" captionStartId="5.[151,250,1935,1957]" captionTargetBox="[197,1388,220,1891]" captionTargetId="figure@5.[172,1414,198,1920]" captionTargetPageId="5" captionText="FIGURE 3. Thor leptochelus (Xu &amp; Li, 2015) n. comb., ovigerous female (cl 2.3 mm), CBM-ZC 13091. Thoracic appendages in lateral view. A, right third maxilliped; B, right first pereopod; C, right second pereopod; D, left third pereopod; E, right third pereopod, dactylus and propodus; F, right fourth pereopod; G, right fifth pereopod. Scale bar: 1 mm." httpUri="https://zenodo.org/record/241265/files/figure.png" pageId="3" pageNumber="402">Fig. 3</figureCitation>
A) stout, overreaching distal margin of antennal scale by 0.3 length of ultimate segment. Ultimate segment about 4.0 times as long as penultimate segment (= carpus), tapering distally, tip encircled by 6 corneous spinules (
<figureCitation id="130737918A666B0FFDA6FD91FDAFFDC5" box="[543,612,546,571]" captionStart="FIGURE 4" captionStartId="6.[151,250,1083,1105]" captionTargetBox="[167,1388,276,956]" captionTargetId="figure@6.[165,1430,244,1003]" captionTargetPageId="6" captionText="FIGURE 4. Thor leptochelus (Xu &amp; Li, 2015) n. comb., ovigerous female (cl 2.3 mm), CBM-ZC 13091. A, right third maxilliped, distal part of ultimate segment, showing spination; B, same, ultimate segment (= carpus) and distal part of antepenultimate segment, mesial view, showing setation and armature; C, same, detail of distolateral armature of antepenultimate segment; D, right first pereopod, chela and distal part of carpus, flexor view; E, same, merus and ischium, mesial view; F, left third pereopod, dactylus and distal part of propodus, lateral view; G, same, distal part of propodus, showing armature on flexor surface; H, right fourth pereopod, dactylus and distal part of propodus, lateral view; G, right fifth pereopod, dactylus and distal part of propodus, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/241266/files/figure.png" pageId="3" pageNumber="402">Fig. 4</figureCitation>
A); dorsal and ventral margins with row of tufts of stiff setae becoming shorter distally; mesial face with numerous transverse tracts of stiff setae, forming grooming apparatus. Penultimate segment (= carpus) with 2 transverse tracts of long, stiff setae on dorsomesial surface (
<figureCitation id="130737918A666B0FFA8BFDDBFAB0FD7F" box="[1330,1403,616,641]" captionStart="FIGURE 4" captionStartId="6.[151,250,1083,1105]" captionTargetBox="[167,1388,276,956]" captionTargetId="figure@6.[165,1430,244,1003]" captionTargetPageId="6" captionText="FIGURE 4. Thor leptochelus (Xu &amp; Li, 2015) n. comb., ovigerous female (cl 2.3 mm), CBM-ZC 13091. A, right third maxilliped, distal part of ultimate segment, showing spination; B, same, ultimate segment (= carpus) and distal part of antepenultimate segment, mesial view, showing setation and armature; C, same, detail of distolateral armature of antepenultimate segment; D, right first pereopod, chela and distal part of carpus, flexor view; E, same, merus and ischium, mesial view; F, left third pereopod, dactylus and distal part of propodus, lateral view; G, same, distal part of propodus, showing armature on flexor surface; H, right fourth pereopod, dactylus and distal part of propodus, lateral view; G, right fifth pereopod, dactylus and distal part of propodus, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/241266/files/figure.png" pageId="3" pageNumber="402">Fig. 4</figureCitation>
B). Antepenultimate segment depressed proximally, with row of long stiff setae on dorsodistal margin; distolateral margin with 1 small tooth obscured by single long spiniform seta (
<figureCitation id="130737918A666B0FFC21FD1CFC14FD39" box="[920,991,687,712]" captionStart="FIGURE 4" captionStartId="6.[151,250,1083,1105]" captionTargetBox="[167,1388,276,956]" captionTargetId="figure@6.[165,1430,244,1003]" captionTargetPageId="6" captionText="FIGURE 4. Thor leptochelus (Xu &amp; Li, 2015) n. comb., ovigerous female (cl 2.3 mm), CBM-ZC 13091. A, right third maxilliped, distal part of ultimate segment, showing spination; B, same, ultimate segment (= carpus) and distal part of antepenultimate segment, mesial view, showing setation and armature; C, same, detail of distolateral armature of antepenultimate segment; D, right first pereopod, chela and distal part of carpus, flexor view; E, same, merus and ischium, mesial view; F, left third pereopod, dactylus and distal part of propodus, lateral view; G, same, distal part of propodus, showing armature on flexor surface; H, right fourth pereopod, dactylus and distal part of propodus, lateral view; G, right fifth pereopod, dactylus and distal part of propodus, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/241266/files/figure.png" pageId="3" pageNumber="402">Fig. 4</figureCitation>
C); ventrolateral margin with 1 spinule distally, ventromesial margin also with 1 spiniform seta subdistally (
<figureCitation id="130737918A666B0FFC29FD61FC1FFD15" box="[912,980,722,747]" captionStart="FIGURE 4" captionStartId="6.[151,250,1083,1105]" captionTargetBox="[167,1388,276,956]" captionTargetId="figure@6.[165,1430,244,1003]" captionTargetPageId="6" captionText="FIGURE 4. Thor leptochelus (Xu &amp; Li, 2015) n. comb., ovigerous female (cl 2.3 mm), CBM-ZC 13091. A, right third maxilliped, distal part of ultimate segment, showing spination; B, same, ultimate segment (= carpus) and distal part of antepenultimate segment, mesial view, showing setation and armature; C, same, detail of distolateral armature of antepenultimate segment; D, right first pereopod, chela and distal part of carpus, flexor view; E, same, merus and ischium, mesial view; F, left third pereopod, dactylus and distal part of propodus, lateral view; G, same, distal part of propodus, showing armature on flexor surface; H, right fourth pereopod, dactylus and distal part of propodus, lateral view; G, right fifth pereopod, dactylus and distal part of propodus, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/241266/files/figure.png" pageId="3" pageNumber="402">Fig. 4</figureCitation>
B). Coxa somewhat inflated laterally, but without strap-like epipod. Exopod not reaching distal margin of antepenultimate segment.
</paragraph>
<paragraph id="8B832B148A666B0FFF7EFCAAFDBDFCCF" blockId="3.[151,1437,151,2014]" box="[199,630,792,817]" pageId="3" pageNumber="402">No strap-like epipod on any pereopod.</paragraph>
<paragraph id="8B832B148A666B0FFF7EFC8FFB2EFBD6" blockId="3.[151,1437,151,2014]" pageId="3" pageNumber="402">
First pereopod (
<figureCitation id="130737918A666B0FFEC3FC8FFE70FCAA" box="[378,443,828,853]" captionStart="FIGURE 3" captionStartId="5.[151,250,1935,1957]" captionTargetBox="[197,1388,220,1891]" captionTargetId="figure@5.[172,1414,198,1920]" captionTargetPageId="5" captionText="FIGURE 3. Thor leptochelus (Xu &amp; Li, 2015) n. comb., ovigerous female (cl 2.3 mm), CBM-ZC 13091. Thoracic appendages in lateral view. A, right third maxilliped; B, right first pereopod; C, right second pereopod; D, left third pereopod; E, right third pereopod, dactylus and propodus; F, right fourth pereopod; G, right fifth pereopod. Scale bar: 1 mm." httpUri="https://zenodo.org/record/241265/files/figure.png" pageId="3" pageNumber="402">Fig. 3</figureCitation>
B) shortest among pereopods, not reaching midlength of antennal scale. Chela (
<figureCitation id="130737918A666B0FFA85FC8FFAB4FCAA" box="[1340,1407,828,853]" captionStart="FIGURE 4" captionStartId="6.[151,250,1083,1105]" captionTargetBox="[167,1388,276,956]" captionTargetId="figure@6.[165,1430,244,1003]" captionTargetPageId="6" captionText="FIGURE 4. Thor leptochelus (Xu &amp; Li, 2015) n. comb., ovigerous female (cl 2.3 mm), CBM-ZC 13091. A, right third maxilliped, distal part of ultimate segment, showing spination; B, same, ultimate segment (= carpus) and distal part of antepenultimate segment, mesial view, showing setation and armature; C, same, detail of distolateral armature of antepenultimate segment; D, right first pereopod, chela and distal part of carpus, flexor view; E, same, merus and ischium, mesial view; F, left third pereopod, dactylus and distal part of propodus, lateral view; G, same, distal part of propodus, showing armature on flexor surface; H, right fourth pereopod, dactylus and distal part of propodus, lateral view; G, right fifth pereopod, dactylus and distal part of propodus, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/241266/files/figure.png" pageId="3" pageNumber="402">Fig. 4</figureCitation>
D) slender, tapering distally, 1.2 times as long as carpus; dactylus short, about half length of palm, terminating in 2 curved ungues; fixed finger terminating in slender unguis about twice length of those on dactylus; palm with short oblique row of stiff setae proximally on flexor surface. Carpus about twice as long as distal width, slightly shorter than palm, with grooming apparatus on mesial face. Merus slightly inflated ventrally; ventromesial margin with cluster of minute spinules medially (
<figureCitation id="130737918A666B0FFD88FC5FFDBFFBFA" box="[561,628,1004,1029]" captionStart="FIGURE 4" captionStartId="6.[151,250,1083,1105]" captionTargetBox="[167,1388,276,956]" captionTargetId="figure@6.[165,1430,244,1003]" captionTargetPageId="6" captionText="FIGURE 4. Thor leptochelus (Xu &amp; Li, 2015) n. comb., ovigerous female (cl 2.3 mm), CBM-ZC 13091. A, right third maxilliped, distal part of ultimate segment, showing spination; B, same, ultimate segment (= carpus) and distal part of antepenultimate segment, mesial view, showing setation and armature; C, same, detail of distolateral armature of antepenultimate segment; D, right first pereopod, chela and distal part of carpus, flexor view; E, same, merus and ischium, mesial view; F, left third pereopod, dactylus and distal part of propodus, lateral view; G, same, distal part of propodus, showing armature on flexor surface; H, right fourth pereopod, dactylus and distal part of propodus, lateral view; G, right fifth pereopod, dactylus and distal part of propodus, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/241266/files/figure.png" pageId="3" pageNumber="402">Fig. 4</figureCitation>
E). Ischium shorter than merus, with set of 3 spiniform setae subdistally on ventromesial face (
<figureCitation id="130737918A666B0FFE36FBBCFE18FBD9" box="[399,467,1039,1064]" captionStart="FIGURE 4" captionStartId="6.[151,250,1083,1105]" captionTargetBox="[167,1388,276,956]" captionTargetId="figure@6.[165,1430,244,1003]" captionTargetPageId="6" captionText="FIGURE 4. Thor leptochelus (Xu &amp; Li, 2015) n. comb., ovigerous female (cl 2.3 mm), CBM-ZC 13091. A, right third maxilliped, distal part of ultimate segment, showing spination; B, same, ultimate segment (= carpus) and distal part of antepenultimate segment, mesial view, showing setation and armature; C, same, detail of distolateral armature of antepenultimate segment; D, right first pereopod, chela and distal part of carpus, flexor view; E, same, merus and ischium, mesial view; F, left third pereopod, dactylus and distal part of propodus, lateral view; G, same, distal part of propodus, showing armature on flexor surface; H, right fourth pereopod, dactylus and distal part of propodus, lateral view; G, right fifth pereopod, dactylus and distal part of propodus, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/241266/files/figure.png" pageId="3" pageNumber="402">Fig. 4</figureCitation>
E). Ventral margin of merus-ischium distinctly notched at articulation.
</paragraph>
<paragraph id="8B832B148A666B0FFF7EFB81FA51FB6F" blockId="3.[151,1437,151,2014]" pageId="3" pageNumber="402">
Second pereopod (
<figureCitation id="130737918A666B0FFE2EFB81FE12FBB5" box="[407,473,1074,1099]" captionStart="FIGURE 3" captionStartId="5.[151,250,1935,1957]" captionTargetBox="[197,1388,220,1891]" captionTargetId="figure@5.[172,1414,198,1920]" captionTargetPageId="5" captionText="FIGURE 3. Thor leptochelus (Xu &amp; Li, 2015) n. comb., ovigerous female (cl 2.3 mm), CBM-ZC 13091. Thoracic appendages in lateral view. A, right third maxilliped; B, right first pereopod; C, right second pereopod; D, left third pereopod; E, right third pereopod, dactylus and propodus; F, right fourth pereopod; G, right fifth pereopod. Scale bar: 1 mm." httpUri="https://zenodo.org/record/241265/files/figure.png" pageId="3" pageNumber="402">Fig. 3</figureCitation>
C) overreaching antennal scale by length of chela. Dactylus about 0.7 times as long as palm. Carpus 3.2 times as long as chela, divided into 6 articles of unequal length, third article longest. Merus 1.3 times as long as ischium. Ischium wider than merus, with set of few spiniform setae proximally on ventral margin.
</paragraph>
<paragraph id="8B832B148A666B0FFF7EFB2FFB11FA55" blockId="3.[151,1437,151,2014]" pageId="3" pageNumber="402">
Third to fifth pereopods generally similar in structure, but decreasing in length and stoutness toward posterior. Third pereopod (
<figureCitation id="130737918A666B0FFEECFB0CFE5CFB29" box="[341,407,1215,1240]" captionStart="FIGURE 3" captionStartId="5.[151,250,1935,1957]" captionTargetBox="[197,1388,220,1891]" captionTargetId="figure@5.[172,1414,198,1920]" captionTargetPageId="5" captionText="FIGURE 3. Thor leptochelus (Xu &amp; Li, 2015) n. comb., ovigerous female (cl 2.3 mm), CBM-ZC 13091. Thoracic appendages in lateral view. A, right third maxilliped; B, right first pereopod; C, right second pereopod; D, left third pereopod; E, right third pereopod, dactylus and propodus; F, right fourth pereopod; G, right fifth pereopod. Scale bar: 1 mm." httpUri="https://zenodo.org/record/241265/files/figure.png" pageId="3" pageNumber="402">Fig. 3</figureCitation>
D) overreaching antennal scale by length of dactylus. Dactylus (
<figureCitation id="130737918A666B0FFBD5FB0CFB64FB29" box="[1132,1199,1215,1240]" captionStart="FIGURE 4" captionStartId="6.[151,250,1083,1105]" captionTargetBox="[167,1388,276,956]" captionTargetId="figure@6.[165,1430,244,1003]" captionTargetPageId="6" captionText="FIGURE 4. Thor leptochelus (Xu &amp; Li, 2015) n. comb., ovigerous female (cl 2.3 mm), CBM-ZC 13091. A, right third maxilliped, distal part of ultimate segment, showing spination; B, same, ultimate segment (= carpus) and distal part of antepenultimate segment, mesial view, showing setation and armature; C, same, detail of distolateral armature of antepenultimate segment; D, right first pereopod, chela and distal part of carpus, flexor view; E, same, merus and ischium, mesial view; F, left third pereopod, dactylus and distal part of propodus, lateral view; G, same, distal part of propodus, showing armature on flexor surface; H, right fourth pereopod, dactylus and distal part of propodus, lateral view; G, right fifth pereopod, dactylus and distal part of propodus, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/241266/files/figure.png" pageId="3" pageNumber="402">Fig. 4</figureCitation>
F) about 0.3 times as long as propodus, 3.0 times longer than high (including unguis), distally biunguiculate (penultimate unguis curved, shorter but almost as wide as ultimate unguis); flexor margin with 4 or 5 slender accessory spinules over entire length. Propodus with field of minute spinules in distal one-third, followed by 2 rows of spinules, on flexor surface in largest specimen (
<figureCitation id="130737918A666B0FFE3CFAFFFE00FA9A" box="[389,459,1356,1381]" captionStart="FIGURE 4" captionStartId="6.[151,250,1083,1105]" captionTargetBox="[167,1388,276,956]" captionTargetId="figure@6.[165,1430,244,1003]" captionTargetPageId="6" captionText="FIGURE 4. Thor leptochelus (Xu &amp; Li, 2015) n. comb., ovigerous female (cl 2.3 mm), CBM-ZC 13091. A, right third maxilliped, distal part of ultimate segment, showing spination; B, same, ultimate segment (= carpus) and distal part of antepenultimate segment, mesial view, showing setation and armature; C, same, detail of distolateral armature of antepenultimate segment; D, right first pereopod, chela and distal part of carpus, flexor view; E, same, merus and ischium, mesial view; F, left third pereopod, dactylus and distal part of propodus, lateral view; G, same, distal part of propodus, showing armature on flexor surface; H, right fourth pereopod, dactylus and distal part of propodus, lateral view; G, right fifth pereopod, dactylus and distal part of propodus, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/241266/files/figure.png" pageId="3" pageNumber="402">Fig. 4</figureCitation>
G), and row of several tufts of long setae on extensor surface; flexor surface of right propodus only with 2 rows of spinules in other three specimens (
<figureCitation id="130737918A666B0FFCC4FADCFC0AFA79" box="[893,961,1391,1416]" captionStart="FIGURE 3" captionStartId="5.[151,250,1935,1957]" captionTargetBox="[197,1388,220,1891]" captionTargetId="figure@5.[172,1414,198,1920]" captionTargetPageId="5" captionText="FIGURE 3. Thor leptochelus (Xu &amp; Li, 2015) n. comb., ovigerous female (cl 2.3 mm), CBM-ZC 13091. Thoracic appendages in lateral view. A, right third maxilliped; B, right first pereopod; C, right second pereopod; D, left third pereopod; E, right third pereopod, dactylus and propodus; F, right fourth pereopod; G, right fifth pereopod. Scale bar: 1 mm." httpUri="https://zenodo.org/record/241265/files/figure.png" pageId="3" pageNumber="402">Fig. 3</figureCitation>
E). Carpus about half length of propodus. Merus about 4.0 times as long as wide, armed with 1 spine located near ventrolateral distal angle.
</paragraph>
<paragraph id="8B832B148A666B0FFF7EFA06FCC3F980" blockId="3.[151,1437,151,2014]" pageId="3" pageNumber="402">
Fourth pereopod (
<figureCitation id="130737918A666B0FFE29FA06FE1AFA30" box="[400,465,1461,1486]" captionStart="FIGURE 3" captionStartId="5.[151,250,1935,1957]" captionTargetBox="[197,1388,220,1891]" captionTargetId="figure@5.[172,1414,198,1920]" captionTargetPageId="5" captionText="FIGURE 3. Thor leptochelus (Xu &amp; Li, 2015) n. comb., ovigerous female (cl 2.3 mm), CBM-ZC 13091. Thoracic appendages in lateral view. A, right third maxilliped; B, right first pereopod; C, right second pereopod; D, left third pereopod; E, right third pereopod, dactylus and propodus; F, right fourth pereopod; G, right fifth pereopod. Scale bar: 1 mm." httpUri="https://zenodo.org/record/241265/files/figure.png" pageId="3" pageNumber="402">Fig. 3</figureCitation>
F) overreaching midlength of antennal scale by tip of propodus, but not reaching distal margin of antennal scale; flexor margin of dactylus with 4 or 5 accessory spinules in addition to penultimate unguis; flexor surface of propodus with short field of minute spinules in distal one-fourth in either side; merus with 1 spine. Fifth pereopod (
<figureCitation id="130737918A666B0FFE09F9ACFE38F9C9" box="[432,499,1567,1592]" captionStart="FIGURE 3" captionStartId="5.[151,250,1935,1957]" captionTargetBox="[197,1388,220,1891]" captionTargetId="figure@5.[172,1414,198,1920]" captionTargetPageId="5" captionText="FIGURE 3. Thor leptochelus (Xu &amp; Li, 2015) n. comb., ovigerous female (cl 2.3 mm), CBM-ZC 13091. Thoracic appendages in lateral view. A, right third maxilliped; B, right first pereopod; C, right second pereopod; D, left third pereopod; E, right third pereopod, dactylus and propodus; F, right fourth pereopod; G, right fifth pereopod. Scale bar: 1 mm." httpUri="https://zenodo.org/record/241265/files/figure.png" pageId="3" pageNumber="402">Fig. 3</figureCitation>
G) reaching or overreaching midlength of antennal scale by tip of dactylus; flexor margin of dactylus armed with 4 or 5 accessory spinules in addition to penultimate unguis; flexor surface of propodus only with 2 rows of spinules; merus unarmed.
</paragraph>
<paragraph id="8B832B148A666B0FFF7EF93BFA46F916" blockId="3.[151,1437,151,2014]" pageId="3" pageNumber="402">
Protopods of first to fourth pleopods (
<figureCitation id="130737918A666B0FFDCFF93BFD73F95F" box="[630,696,1672,1697]" captionStart="FIGURE 1" captionStartId="2.[151,250,1312,1334]" captionTargetBox="[157,1409,336,1311]" captionTargetId="figure@2.[157,1477,304,1317]" captionTargetPageId="2" captionText="FIGURE 1. Thor leptochelus (Xu &amp; Li, 2015) n. comb., ovigerous female (cl 2.3 mm), CBM-ZC 13091, habitus, lateral view. Scale bar: 2 mm." httpUri="https://zenodo.org/record/241263/files/figure.png" pageId="3" pageNumber="402">Fig. 1</figureCitation>
) each with strongly broadened ventrolateral lamella in spawning molt, while moderately broad in non-ovigerous female. Uropod (
<figureCitation id="130737918A666B0FFCCBF91FFC7FF93A" box="[882,948,1708,1733]" captionStart="FIGURE 2" captionStartId="4.[151,250,1822,1844]" captionTargetBox="[238,1348,213,1793]" captionTargetId="figure@4.[212,1372,188,1805]" captionTargetPageId="4" captionText="FIGURE 2. Thor leptochelus (Xu &amp; Li, 2015) n. comb. A, B, D – N, ovigerous female (cl 2.3 mm), CBM-ZC 13091; C, paratype, female (cl 1.5 mm), CBM-ZC 13090. Mouthparts dissected from right side. A, carapace and eyes, dorsal view; B, rostrum and anterior part of carapace, lateral view; C, rostrum, dorsolateral view; D, telson and right uropod, dorsal view (setae omitted); E, posterior part of telson, dorsal view; F, right antennular peduncle and antennal basicerite, lateral view; G, right antennular peduncle, dorsolateral view; H, right antenna, ventral view (flagellum omitted); I, mandible, external view; J, same, internal view; K, maxillule, outer view (proximal endite missing); L, maxilla, external view; M, first maxilliped, external view; N, second maxilliped, external view. Scale bars: 1 mm for A, B, F; 0.5 mm for C – E, G – N." httpUri="https://zenodo.org/record/241264/files/figure.png" pageId="3" pageNumber="402">Fig. 2</figureCitation>
D) with endopod reaching posterior margin of telson; exopod slightly longer than endopod, armed with small spine just mesial to minute posterolateral tooth.
</paragraph>
<paragraph id="8B832B148A666B0FFF7EF941FD5BF8F4" blockId="3.[151,1437,151,2014]" box="[199,656,1778,1803]" pageId="3" pageNumber="402">
<emphasis id="B948F7068A666B0FFF7EF941FF3CF8F5" bold="true" box="[199,247,1778,1803]" pageId="3" pageNumber="402">Size</emphasis>
. Ovigerous females: cl
<quantity id="4CC486F18A666B0FFDBBF941FD47F8F4" box="[514,652,1778,1803]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.3" metricValueMin="1.6" pageId="3" pageNumber="402" unit="mm" value="1.95" valueMax="2.3" valueMin="1.6">1.6–2.3 mm</quantity>
.
</paragraph>
<paragraph id="8B832B148A666B0FFF7EF8A6FC52F820" blockId="3.[151,1437,151,2014]" pageId="3" pageNumber="402">
<emphasis id="B948F7068A666B0FFF7EF8A6FE60F8D0" bold="true" box="[199,427,1813,1838]" pageId="3" pageNumber="402">Colouration in life</emphasis>
. Carapace and anterior three pleomeres mottled milky white, with scattered brownish blotches or spots; fourth to sixth pleomeres and tail fan translucent. Cornea grayish. Antennular peduncle milky white; outer flagellum reddish brown, mottled with white. Antennal peduncle and scale translucent in general, carpocerite with reddish ring distally. Third maxilliped to second pereopods generally brownish. Third to fifth pereopods with dactyli to carpi translucent with brown tint, meri darker brown. Protopods of anterior three pleopods milky white, those of fourth and fifth pleopods translucent.
</paragraph>
</subSubSection>
<caption id="DF437B9C8A616B08FF2EF8ADFBE1F812" httpUri="https://zenodo.org/record/241264/files/figure.png" pageId="4" pageNumber="403" targetBox="[238,1348,213,1793]" targetPageId="4">
<paragraph id="8B832B148A616B08FF2EF8ADFBE1F812" blockId="4.[151,1436,1822,2028]" pageId="4" pageNumber="403">
<emphasis id="B948F7068A616B08FF2EF8ADFEDBF8CD" bold="true" box="[151,272,1822,1844]" pageId="4" pageNumber="403">FIGURE 2</emphasis>
.
<taxonomicName id="4C3C50978A616B08FE99F8ADFD41F8CA" authority="Xu &amp; Li, 2015" authorityName="Xu &amp; Li" authorityYear="2015" box="[288,650,1822,1844]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="403" phylum="Arthropoda" rank="species" species="leptochelus" status="comb. nov.">
<emphasis id="B948F7068A616B08FE99F8ADFE05F8CA" box="[288,462,1822,1844]" italics="true" pageId="4" pageNumber="403">Thor leptochelus</emphasis>
(Xu &amp; Li, 2015)
</taxonomicName>
<taxonomicNameLabel id="A27B4A7D8A616B08FD2AF8ACFD3BF8CA" box="[659,752,1822,1844]" pageId="4" pageNumber="403">n. comb.</taxonomicNameLabel>
A, B, D–N, ovigerous female (cl 2.3 mm), CBM-ZC 13091; C, paratype, female (cl 1.5 mm), CBM-ZC 13090. Mouthparts dissected from right side. A, carapace and eyes, dorsal view; B, rostrum and anterior part of carapace, lateral view; C, rostrum, dorsolateral view; D, telson and right uropod, dorsal view (setae omitted); E, posterior part of telson, dorsal view; F, right antennular peduncle and antennal basicerite, lateral view; G, right antennular peduncle, dorsolateral view; H, right antenna, ventral view (flagellum omitted); I, mandible, external view; J, same, internal view; K, maxillule, outer view (proximal endite missing); L, maxilla, external view; M, first maxilliped, external view; N, second maxilliped, external view. Scale bars: 1 mm for A, B, F; 0.5 mm for C–E, G–N.
</paragraph>
</caption>
<caption id="DF437B9C8A606B09FF2EF83CFB46F814" httpUri="https://zenodo.org/record/241265/files/figure.png" pageId="5" pageNumber="404" targetBox="[197,1388,220,1891]" targetPageId="5">
<paragraph id="8B832B148A606B09FF2EF83CFB46F814" blockId="5.[151,1436,1935,2026]" pageId="5" pageNumber="404">
<emphasis id="B948F7068A606B09FF2EF83CFED9F85A" bold="true" box="[151,274,1935,1957]" pageId="5" pageNumber="404">FIGURE 3.</emphasis>
<taxonomicName id="4C3C50978A606B09FEA0F83CFDA4F85B" authority="Xu &amp; Li, 2015" authorityName="Xu &amp; Li" authorityYear="2015" box="[281,623,1935,1957]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="404" phylum="Arthropoda" rank="species" species="leptochelus" status="comb. nov.">
<emphasis id="B948F7068A606B09FEA0F83CFE08F85B" box="[281,451,1935,1957]" italics="true" pageId="5" pageNumber="404">Thor leptochelus</emphasis>
(Xu &amp; Li, 2015)
</taxonomicName>
<taxonomicNameLabel id="A27B4A7D8A606B09FDCFF823FD1AF85B" box="[630,721,1935,1957]" pageId="5" pageNumber="404">n. comb.</taxonomicNameLabel>
, ovigerous female (cl 2.3 mm), CBM-ZC 13091. Thoracic appendages in lateral view. A, right third maxilliped; B, right first pereopod; C, right second pereopod; D, left third pereopod; E, right third pereopod, dactylus and propodus; F, right fourth pereopod; G, right fifth pereopod. Scale bar: 1 mm.
</paragraph>
</caption>
<caption id="DF437B9C8A636B0AFF2EFB88FC93FAF4" httpUri="https://zenodo.org/record/241266/files/figure.png" pageId="6" pageNumber="405" targetBox="[167,1388,276,956]" targetPageId="6">
<paragraph id="8B832B148A636B0AFF2EFB88FC93FAF4" blockId="6.[151,1436,1083,1290]" pageId="6" pageNumber="405">
<emphasis id="B948F7068A636B0AFF2EFB88FEDCFBAE" bold="true" box="[151,279,1083,1105]" pageId="6" pageNumber="405">FIGURE 4.</emphasis>
<taxonomicName id="4C3C50978A636B0AFE98FB88FD46FBAF" authority="Xu &amp; Li, 2015" authorityName="Xu &amp; Li" authorityYear="2015" box="[289,653,1083,1105]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="405" phylum="Arthropoda" rank="species" species="leptochelus" status="comb. nov.">
<emphasis id="B948F7068A636B0AFE98FB88FE1BFBAF" box="[289,464,1083,1105]" italics="true" pageId="6" pageNumber="405">Thor leptochelus</emphasis>
(Xu &amp; Li, 2015)
</taxonomicName>
<taxonomicNameLabel id="A27B4A7D8A636B0AFD21FB8FFD3CFBAF" box="[664,759,1083,1105]" pageId="6" pageNumber="405">n. comb.</taxonomicNameLabel>
, ovigerous female (cl 2.3 mm), CBM-ZC 13091. A, right third maxilliped, distal part of ultimate segment, showing spination; B, same, ultimate segment (= carpus) and distal part of antepenultimate segment, mesial view, showing setation and armature; C, same, detail of distolateral armature of antepenultimate segment; D, right first pereopod, chela and distal part of carpus, flexor view; E, same, merus and ischium, mesial view; F, left third pereopod, dactylus and distal part of propodus, lateral view; G, same, distal part of propodus, showing armature on flexor surface; H, right fourth pereopod, dactylus and distal part of propodus, lateral view; G, right fifth pereopod, dactylus and distal part of propodus, lateral view. Scale bars: 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C326789F8A636B0AFF7EFA85FB25FA45" pageId="6" pageNumber="405" type="distribution">
<paragraph id="8B832B148A636B0AFF7EFA85FC9EFA8C" blockId="6.[151,1437,1334,2007]" pageId="6" pageNumber="405">
<emphasis id="B948F7068A636B0AFF7EFA85FE92FAB1" bold="true" box="[199,345,1334,1359]" pageId="6" pageNumber="405">Distribution</emphasis>
. Presently known only from Xisha Islands in the South
<collectingCountry id="F32B6B848A636B0AFC50FA85FBE7FAB1" box="[1001,1068,1334,1359]" name="China" pageId="6" pageNumber="405">China</collectingCountry>
Sea (
<typeStatus id="548795B68A636B0AFBC9FA84FB69FAB1" box="[1136,1186,1335,1359]" pageId="6" pageNumber="405">type</typeStatus>
locality) and Ryukyu Archipelago (Okinawa and Kume islands), at depths of
<quantity id="4CC486F18A636B0AFCBEFAE8FC9AFA8C" box="[775,849,1371,1395]" metricMagnitude="0" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="6" pageNumber="405" unit="m" value="7.5" valueMax="8.0" valueMin="7.0">
7–
<specimenCount id="9D3AE09D8A636B0AFC9CFAE8FC9AFA8C" box="[805,849,1371,1395]" pageId="6" pageNumber="405" type="generic">8 m</specimenCount>
</quantity>
.
</paragraph>
<paragraph id="8B832B148A636B0AFF7EFACDFB25FA45" blockId="6.[151,1437,1334,2007]" pageId="6" pageNumber="405">
<emphasis id="B948F7068A636B0AFF7EFACDFEE1FA69" bold="true" box="[199,298,1406,1431]" pageId="6" pageNumber="405">Habitat.</emphasis>
No association with other invertebrates was observed during sampling. Individuals were seen in crevices on reefs, but only during the night. They tried to escape immediately when light was used.
</paragraph>
</subSubSection>
<subSubSection id="C326789F8A636B0AFF7EFA75FE61F8B9" pageId="6" pageNumber="405" type="description">
<paragraph id="8B832B148A636B0AFF7EFA75FCBFF9FD" blockId="6.[151,1437,1334,2007]" pageId="6" pageNumber="405">
<emphasis id="B948F7068A636B0AFF7EFA75FEFEFA21" bold="true" box="[199,309,1478,1503]" pageId="6" pageNumber="405">Variation</emphasis>
. The body is less robust in the non-ovigerous female
<typeStatus id="548795B68A636B0AFC36FA74FC24FA21" box="[911,1007,1479,1503]" pageId="6" pageNumber="405" type="paratype">paratype</typeStatus>
than in the three ovigerous specimens (
<typeStatus id="548795B68A636B0AFF27FA59FEC9F9FD" box="[158,258,1514,1539]" pageId="6" pageNumber="405" type="holotype">holotype</typeStatus>
and
<typeStatus id="548795B68A636B0AFE80FA58FE62F9FD" box="[313,425,1515,1539]" pageId="6" pageNumber="405" type="paratype">paratypes</typeStatus>
), and this variation could be size-related.
</paragraph>
<paragraph id="8B832B148A636B0AFF7EF9BDFE52F96C" blockId="6.[151,1437,1334,2007]" pageId="6" pageNumber="405">
Notable intraspecific variation is seen in the dentition of the rostrum: there are three dorsal and one ventral teeth in the
<typeStatus id="548795B68A636B0AFE98F981FE4DF9B5" box="[289,390,1586,1611]" pageId="6" pageNumber="405" type="holotype">holotype</typeStatus>
, one dorsal and one ventral teeth in the ovigerous female
<typeStatus id="548795B68A636B0AFB9BF980FB5BF9B5" box="[1058,1168,1587,1611]" pageId="6" pageNumber="405" type="paratype">paratypes</typeStatus>
, and one dorsal and no ventral tooth in the non-ovigerous female
<typeStatus id="548795B68A636B0AFDC6F9E4FD28F991" box="[639,739,1623,1647]" pageId="6" pageNumber="405" type="paratype">paratype</typeStatus>
.
<bibRefCitation id="EFAD56E58A636B0AFD49F9E4FC43F991" author="Bruce" box="[752,904,1622,1647]" pageId="6" pageNumber="405" refString="Bruce, A. J. (1976) A report on a small collection of shrimps from the Kenya National Marine Parks at Malindi, with notes on selected species. Zoologische Verhandelingen, 145, 1 - 72." type="journal article" year="1976">Bruce (1976)</bibRefCitation>
reported on the occasional absence of ventral tooth in
<taxonomicName id="4C3C50978A636B0AFF4DF9C9FE59F96C" box="[244,402,1658,1683]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="405" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B948F7068A636B0AFF4DF9C9FE59F96C" box="[244,402,1658,1683]" italics="true" pageId="6" pageNumber="405">Thor spinosus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B832B148A636B0AFF7EF92DFE61F8B9" blockId="6.[151,1437,1334,2007]" pageId="6" pageNumber="405">The armature of the flexor surface of the propodi of the third and fourth pereopods is also variable among the specimens examined. In the largest specimen, there is a field of numerous minute spinules on the distal part of the flexor surface of the left third pereopod propodus and the both fourth pereopod propodi, whereas such a spinulose field is absent in the right third pereopod propodus. In the other three specimens, such spinulose field is not seen on every pereopod propodi.</paragraph>
</subSubSection>
<subSubSection id="C326789F8A636B0BFF7EF8E1FBD7FD11" lastPageId="7" lastPageNumber="406" pageId="6" pageNumber="405" type="discussion">
<paragraph id="8B832B148A636B0BFF7EF8E1FB18FEBE" blockId="6.[151,1437,1334,2007]" lastBlockId="7.[151,1437,151,1653]" lastPageId="7" lastPageNumber="406" pageId="6" pageNumber="405">
<emphasis id="B948F7068A636B0AFF7EF8E1FEFFF895" bold="true" box="[199,308,1874,1899]" pageId="6" pageNumber="405">Remarks</emphasis>
. The present specimens agree well with the original description of
<taxonomicName id="4C3C50978A636B0AFB97F8E1FAC4F895" box="[1070,1295,1874,1899]" class="Malacostraca" family="Hippolytidae" genus="Thinora" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="405" phylum="Arthropoda" rank="species" species="leptochelus">
<emphasis id="B948F7068A636B0AFB97F8E1FAC4F895" box="[1070,1295,1874,1899]" italics="true" pageId="6" pageNumber="405">Thinora leptochelus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFAD56E58A636B0AFA82F8E0FF2FF871" author="Xu" pageId="6" pageNumber="405" refString="Xu, P. &amp; Li, X. (2015) Report on the Hippolytidae Bate (sensu lato) from China seas. Zoological Systematics, 40, 107 - 165." type="journal article" year="2015">Xu &amp; Li (2015)</bibRefCitation>
, particularly in the very robust body, the ascending suborbital angle of the carapace, similar armature of the telson, absence of a conspicuous tooth on the antennular stylocerite and the tapered chela of the first pereopod. Minor discrepancies are as follows.
<bibRefCitation id="EFAD56E58A636B0AFD87F80CFD32F829" author="Xu" box="[574,761,1982,2007]" pageId="6" pageNumber="405" refString="Xu, P. &amp; Li, X. (2015) Report on the Hippolytidae Bate (sensu lato) from China seas. Zoological Systematics, 40, 107 - 165." type="journal article" year="2015">Xu &amp; Li (2015)</bibRefCitation>
stated that the antennular stylocerite is “armed with little indistinct tooth near proximal end of lateral margin”, but in our specimens, there is no trace of any tooth-like structure. According to the description by
<bibRefCitation id="EFAD56E58A626B0BFDC9FF0EFCE8FF2B" author="Xu" box="[624,803,188,213]" pageId="7" pageNumber="406" refString="Xu, P. &amp; Li, X. (2015) Report on the Hippolytidae Bate (sensu lato) from China seas. Zoological Systematics, 40, 107 - 165." type="journal article" year="2015">Xu &amp; Li (2015)</bibRefCitation>
, there are three accessory spinules on the dactylus of the third pereopod, whereas in our specimens the dactylus of the third pereopod is armed with four or five accessory spinules. The rostral armature shows variation in our specimens as mentioned above, but two of the four available specimens match the
<typeStatus id="548795B68A626B0BFE3EFE94FE22FEBE" box="[391,489,295,320]" pageId="7" pageNumber="406" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C3C50978A626B0BFDB7FE94FD24FEBE" box="[526,751,295,320]" class="Malacostraca" family="Hippolytidae" genus="Thinora" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="species" species="leptochelus">
<emphasis id="B948F7068A626B0BFDB7FE94FD24FEBE" box="[526,751,295,320]" italics="true" pageId="7" pageNumber="406">Thinora leptochelus</emphasis>
</taxonomicName>
in having one dorsal and one ventral teeth.
</paragraph>
<paragraph id="8B832B148A626B0BFF7EFEFFFBD7FD11" blockId="7.[151,1437,151,1653]" pageId="7" pageNumber="406">
This species was originally assigned to
<taxonomicName id="4C3C50978A626B0BFD21FEFFFD39FE9B" box="[664,754,332,357]" class="Malacostraca" family="Hippolytidae" genus="Thinora" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="genus">
<emphasis id="B948F7068A626B0BFD21FEFFFD39FE9B" box="[664,754,332,357]" italics="true" pageId="7" pageNumber="406">Thinora</emphasis>
</taxonomicName>
in reference only to the rostral armature and the unarmed pterygostomial angle of the carapace (
<bibRefCitation id="EFAD56E58A626B0BFDE9FEC3FD37FE76" author="Xu" box="[592,764,367,392]" pageId="7" pageNumber="406" refString="Xu, P. &amp; Li, X. (2015) Report on the Hippolytidae Bate (sensu lato) from China seas. Zoological Systematics, 40, 107 - 165." type="journal article" year="2015">Xu &amp; Li 2015</bibRefCitation>
).
<taxonomicName id="4C3C50978A626B0BFCA8FEDCFCA0FE76" box="[785,875,367,392]" class="Malacostraca" family="Hippolytidae" genus="Thinora" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="genus">
<emphasis id="B948F7068A626B0BFCA8FEDCFCA0FE76" box="[785,875,367,392]" italics="true" pageId="7" pageNumber="406">Thinora</emphasis>
</taxonomicName>
was diagnosed as having a single dorsal and no ventral teeth on the rostrum (
<bibRefCitation id="EFAD56E58A626B0BFE50FE27FDB8FE52" author="Bruce" box="[489,627,404,428]" pageId="7" pageNumber="406" refString="Bruce, A. J. (1997) A new genus of hippolytid shrimp (Crustacea: Decapoda: Hippolytidae) for Thor maldivensis Borradaile. Memoirs of the Queensland Museum, 42, 13 - 23." type="journal article" year="1997">Bruce 1997</bibRefCitation>
), but substantial variation in the rostral armature has been reported in species of
<taxonomicName id="4C3C50978A626B0BFEB6FE04FE8DFE2E" box="[271,326,439,464]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="genus">
<emphasis id="B948F7068A626B0BFEB6FE04FE8DFE2E" box="[271,326,439,464]" italics="true" pageId="7" pageNumber="406">Thor</emphasis>
</taxonomicName>
(e.g.,
<bibRefCitation id="EFAD56E58A626B0BFE37FE0BFDD8FE2E" author="Bruce" box="[398,531,439,464]" pageId="7" pageNumber="406" refString="Bruce, A. J. (1976) A report on a small collection of shrimps from the Kenya National Marine Parks at Malindi, with notes on selected species. Zoologische Verhandelingen, 145, 1 - 72." type="journal article" year="1976">Bruce 1976</bibRefCitation>
,
<bibRefCitation id="EFAD56E58A626B0BFD99FE0BFD97FE2E" author="Bruce" box="[544,604,440,464]" pageId="7" pageNumber="406" refString="Bruce, A. J. (1978) Thor marguitae sp. nov., a new hippolytid shrimp from Heron Island, Australia. Crustaceana, 35, 160 - 169. http: // dx. doi. org / 10.1163 / 156854078 X 00060" type="journal article" year="1978">1978</bibRefCitation>
). The present specimens show that the rostral armature is also variable in
<taxonomicName id="4C3C50978A626B0BFF2EFE6FFEB1FE0B" box="[151,378,476,501]" class="Malacostraca" family="Hippolytidae" genus="Thinora" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="species" species="leptochelus">
<emphasis id="B948F7068A626B0BFF2EFE6FFEB1FE0B" box="[151,378,476,501]" italics="true" pageId="7" pageNumber="406">Thinora leptochelus</emphasis>
</taxonomicName>
, as described above, and the observation led us to reevaluate the generic assignment of the species.
<taxonomicName id="4C3C50978A626B0BFF4CFE4CFE1DFDE6" box="[245,470,511,536]" class="Malacostraca" family="Hippolytidae" genus="Thinora" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="species" species="leptochelus">
<emphasis id="B948F7068A626B0BFF4CFE4CFE1DFDE6" box="[245,470,511,536]" italics="true" pageId="7" pageNumber="406">Thinora leptochelus</emphasis>
</taxonomicName>
further differs from
<taxonomicName id="4C3C50978A626B0BFD05FE4CFC69FDE6" box="[700,930,511,536]" class="Malacostraca" family="Hippolytidae" genus="Thinora" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="species" species="maldivensis">
<emphasis id="B948F7068A626B0BFD05FE4CFC69FDE6" box="[700,930,511,536]" italics="true" pageId="7" pageNumber="406">Thinora maldivensis</emphasis>
</taxonomicName>
and is rather similar to species of
<taxonomicName id="4C3C50978A626B0BFA99FE4CFA9DFDE6" box="[1312,1366,511,536]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="genus">
<emphasis id="B948F7068A626B0BFA99FE4CFA9DFDE6" box="[1312,1366,511,536]" italics="true" pageId="7" pageNumber="406">Thor</emphasis>
</taxonomicName>
in the non-dorsoventrally flattened rostrum and the dactyli of the third to fifth pereopods each bearing three to five accessory spinules in addition to penultimate unguis, rather than having only two accessory spinules in
<taxonomicName id="4C3C50978A626B0BFAFBFDF4FED0FD7B" class="Malacostraca" family="Hippolytidae" genus="Thinora" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="species" species="maldivensis">
<emphasis id="B948F7068A626B0BFAFBFDF4FED0FD7B" italics="true" pageId="7" pageNumber="406">Thinora maldivensis</emphasis>
</taxonomicName>
. The shape of the pterygostomial angle of the carapace is not reliable in differentiating
<taxonomicName id="4C3C50978A626B0BFAA9FDDFFAA0FD7B" box="[1296,1387,620,645]" class="Malacostraca" family="Hippolytidae" genus="Thinora" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="genus">
<emphasis id="B948F7068A626B0BFAA9FDDFFAA0FD7B" box="[1296,1387,620,645]" italics="true" pageId="7" pageNumber="406">Thinora</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3C50978A626B0BFF2EFD3CFF07FD56" box="[151,204,655,680]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="genus">
<emphasis id="B948F7068A626B0BFF2EFD3CFF07FD56" box="[151,204,655,680]" italics="true" pageId="7" pageNumber="406">Thor</emphasis>
</taxonomicName>
, and none of previous authors did mention this character to be of possible generic significance (e.g.,
<bibRefCitation id="EFAD56E58A626B0BFAEFFD3CFF19FD32" author="Chace" pageId="7" pageNumber="406" refString="Chace, F. A. Jr. (1997) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition 1907 - 1910, Part 7: Families Atyidae, Eugonatonotidae, Rhynchocinetidae, Bathypalaemonellidae, Processidae, and Hippolytidae. Smithsonian Contributions to Zoology, 587, 1 - 106." type="journal article" year="1997">Chace 1997</bibRefCitation>
;
<bibRefCitation id="EFAD56E58A626B0BFF59FD06FEA2FD32" author="Bruce" box="[224,361,692,716]" pageId="7" pageNumber="406" refString="Bruce, A. J. (1997) A new genus of hippolytid shrimp (Crustacea: Decapoda: Hippolytidae) for Thor maldivensis Borradaile. Memoirs of the Queensland Museum, 42, 13 - 23." type="journal article" year="1997">Bruce 1997</bibRefCitation>
). At present, male characteristics need to be verified for
<taxonomicName id="4C3C50978A626B0BFC47FD07FB75FD33" box="[1022,1214,692,717]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="species" species="leptochelus">
<emphasis id="B948F7068A626B0BFC47FD07FB75FD33" box="[1022,1214,692,717]" italics="true" pageId="7" pageNumber="406">Thor leptochelus</emphasis>
</taxonomicName>
, but the diagnostic characters noted above clearly shows that the species should be referred to
<taxonomicName id="4C3C50978A626B0BFC59FD64FBDEFD0E" box="[992,1045,727,752]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="genus">
<emphasis id="B948F7068A626B0BFC59FD64FBDEFD0E" box="[992,1045,727,752]" italics="true" pageId="7" pageNumber="406">Thor</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C326789F8A626B0BFF7EFD4FFD88F98A" pageId="7" pageNumber="406" type="reference_group">
<paragraph id="8B832B148A626B0BFF7EFD4FFBF8FACE" blockId="7.[151,1437,151,1653]" pageId="7" pageNumber="406">
<taxonomicName id="4C3C50978A626B0BFF7EFD4FFE40FCEB" box="[199,395,764,789]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="species" species="leptochelus" status="comb. nov.">
<emphasis id="B948F7068A626B0BFF7EFD4FFE40FCEB" box="[199,395,764,789]" italics="true" pageId="7" pageNumber="406">Thor leptochelus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B4A7D8A626B0BFE23FD4EFDCFFCEB" box="[410,516,764,789]" pageId="7" pageNumber="406">n. comb.</taxonomicNameLabel>
is the fourth species of
<taxonomicName id="4C3C50978A626B0BFC84FD4FFCBFFCEB" box="[829,884,764,789]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="genus">
<emphasis id="B948F7068A626B0BFC84FD4FFCBFFCEB" box="[829,884,764,789]" italics="true" pageId="7" pageNumber="406">Thor</emphasis>
</taxonomicName>
characterized by the possession of a pair of conspicuous supraorbital teeth on the carapace. The other three species are
<taxonomicName id="4C3C50978A626B0BFC51FC92FB91FCC6" box="[1000,1114,799,824]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="species" species="cordelli">
<emphasis id="B948F7068A626B0BFC51FC92FB91FCC6" box="[1000,1114,799,824]" italics="true" pageId="7" pageNumber="406">T. cordelli</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3C50978A626B0BFBD0FC92FB2AFCC6" box="[1129,1249,801,824]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="B948F7068A626B0BFBD0FC92FB2AFCC6" box="[1129,1249,801,824]" italics="true" pageId="7" pageNumber="406">T. spinipes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3C50978A626B0BFAA1FC92FA59FCC6" box="[1304,1426,801,824]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B948F7068A626B0BFAA1FC92FA59FCC6" box="[1304,1426,801,824]" italics="true" pageId="7" pageNumber="406">T. spinosus</emphasis>
</taxonomicName>
.
<taxonomicName id="4C3C50978A626B0BFF2EFCF7FE93FCA3" box="[151,344,836,861]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="species" species="leptochelus">
<emphasis id="B948F7068A626B0BFF2EFCF7FE93FCA3" box="[151,344,836,861]" italics="true" pageId="7" pageNumber="406">Thor leptochelus</emphasis>
</taxonomicName>
differs from the latter three species in the lack of a small tooth near the proximal end of the antennular stylocerite and the slender, tapered chela of the first pereopod. In
<taxonomicName id="4C3C50978A626B0BFC54FCDAFB94FC7E" box="[1005,1119,871,896]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="species" species="cordelli">
<emphasis id="B948F7068A626B0BFC54FCDAFB94FC7E" box="[1005,1119,871,896]" italics="true" pageId="7" pageNumber="406">T. cordelli</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3C50978A626B0BFBD5FCDAFB2FFC7E" box="[1132,1252,873,896]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="B948F7068A626B0BFBD5FCDAFB2FFC7E" box="[1132,1252,873,896]" italics="true" pageId="7" pageNumber="406">T. spinipes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3C50978A626B0BFAA0FCDAFA59FC7E" box="[1305,1426,873,896]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B948F7068A626B0BFAA0FCDAFA59FC7E" box="[1305,1426,873,896]" italics="true" pageId="7" pageNumber="406">T. spinosus</emphasis>
</taxonomicName>
, the antennular stylocerite bears a small tooth on the proximal end of the stylocerite; the chela of the first pereopod is robust and not markedly tapered (
<bibRefCitation id="EFAD56E58A626B0BFD8DFC03FD77FC36" author="Bruce" box="[564,700,943,968]" pageId="7" pageNumber="406" refString="Bruce, A. J. (1976) A report on a small collection of shrimps from the Kenya National Marine Parks at Malindi, with notes on selected species. Zoologische Verhandelingen, 145, 1 - 72." type="journal article" year="1976">Bruce 1976</bibRefCitation>
,
<bibRefCitation id="EFAD56E58A626B0BFD71FC03FCC8FC36" author="Bruce" box="[712,771,944,968]" pageId="7" pageNumber="406" refString="Bruce, A. J. (1983) Thor spinipes sp. nov., a new hippolytid shrimp from the Cobourg Peninsula, Northern Australia. The Beagle, Records of the Museums and Art Galleries of the Northern Territory Museum of Arts and Sciences, 1, 1 - 10." type="journal article" year="1983">1983</bibRefCitation>
;
<bibRefCitation id="EFAD56E58A626B0BFCA8FC1CFC75FC36" author="Wicksten" box="[785,958,943,968]" pageId="7" pageNumber="406" refString="Wicksten, M. K. (1996) A new species of hippolytid shrimp from Rocas Alijos. In: Schmieder, R. W. (Ed.), Rocas Alijos. Kluwer Academic Publishers, Dordrecht, pp. 295 - 298." type="book chapter" year="1996">Wicksten 1996</bibRefCitation>
). The extremely robust body in ovigerous females could be also characteristic to the present new species, although this character might be size- or sexrelated. In the structure of the chela of the first pereopod,
<taxonomicName id="4C3C50978A626B0BFCF9FC4AFC15FBEE" box="[832,990,1015,1040]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="species" species="leptochelus">
<emphasis id="B948F7068A626B0BFCF9FC4AFC15FBEE" box="[832,990,1015,1040]" italics="true" pageId="7" pageNumber="406">T. leptochelus</emphasis>
</taxonomicName>
is rather similar to
<taxonomicName id="4C3C50978A626B0BFB70FC4AFAA0FBEE" box="[1225,1387,1015,1040]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="B948F7068A626B0BFB70FC4AFAA0FBEE" box="[1225,1387,1015,1040]" italics="true" pageId="7" pageNumber="406">T. intermedius</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="EFAD56E58A626B0BFF2EFBAFFEF4FBCA" author="Holthuis" box="[151,319,1052,1077]" pageId="7" pageNumber="406" refString="Holthuis, L. B. (1947) The Decapoda of the Siboga Expedition. Part IX. The Hippolytidae and Rhynchocinetidae collected by the Siboga and Snellius expeditions with remarks on other species. Siboga Expeditie, 39 a 8, 1 - 100." type="journal article" year="1947">Holthuis 1947</bibRefCitation>
) and
<taxonomicName id="4C3C50978A626B0BFE37FBAEFDE8FBCA" box="[398,547,1053,1076]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="species" species="marguitae">
<emphasis id="B948F7068A626B0BFE37FBAEFDE8FBCA" box="[398,547,1053,1076]" italics="true" pageId="7" pageNumber="406">T. marguitae</emphasis>
</taxonomicName>
(cf. Bruce 1973; this study). From
<taxonomicName id="4C3C50978A626B0BFC5EFBAEFBADFBCA" box="[999,1126,1053,1076]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="B948F7068A626B0BFC5EFBAEFBADFBCA" box="[999,1126,1053,1076]" italics="true" pageId="7" pageNumber="406">T. spinipes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3C50978A626B0BFBC5FBAEFAD4FBCB" box="[1148,1311,1052,1077]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="species" species="leptochelus">
<emphasis id="B948F7068A626B0BFBC5FBAEFAD4FBCB" box="[1148,1311,1052,1077]" italics="true" pageId="7" pageNumber="406">T. leptochelus</emphasis>
</taxonomicName>
is further distinguished by the comparatively wider antennal scale (1.6 times as long as wide versus about 2.5 times as long) and the much more robust ultimate segment of the third maxilliped. From
<taxonomicName id="4C3C50978A626B0BFBBBFBD6FB4AFB82" box="[1026,1153,1125,1148]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B948F7068A626B0BFBBBFBD6FB4AFB82" box="[1026,1153,1125,1148]" italics="true" pageId="7" pageNumber="406">T. spinosus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3C50978A626B0BFB2DFBD6FAF8FB83" box="[1172,1331,1124,1149]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="species" species="leptochelus">
<emphasis id="B948F7068A626B0BFB2DFBD6FAF8FB83" box="[1172,1331,1124,1149]" italics="true" pageId="7" pageNumber="406">T. leptochelus</emphasis>
</taxonomicName>
is easily separated by the different armature of the posterior margin of the telson and the less stout dactyli of the third to fifth pereopods (about 3.0 times as long as wide versus about 2.0 times as long). In
<taxonomicName id="4C3C50978A626B0BFBB7FB1EFB61FB3B" box="[1038,1194,1196,1221]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="species" species="leptochelus">
<emphasis id="B948F7068A626B0BFBB7FB1EFB61FB3B" box="[1038,1194,1196,1221]" italics="true" pageId="7" pageNumber="406">T. leptochelus</emphasis>
</taxonomicName>
, the posterior margin of the telson bears one mesial pair of plumose setae on either side of the unarmed midpoint, whereas in
<taxonomicName id="4C3C50978A626B0BFAA0FB62FA59FB16" box="[1305,1426,1233,1256]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B948F7068A626B0BFAA0FB62FA59FB16" box="[1305,1426,1233,1256]" italics="true" pageId="7" pageNumber="406">T. spinosus</emphasis>
</taxonomicName>
, there are two mesial pairs of plumose setae on either side of the tiny median tooth. Finally, the new species differs from
<taxonomicName id="4C3C50978A626B0BFF6DFAAAFE8DFACE" box="[212,326,1303,1328]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="species" species="cordelli">
<emphasis id="B948F7068A626B0BFF6DFAAAFE8DFACE" box="[212,326,1303,1328]" italics="true" pageId="7" pageNumber="406">T. cordelli</emphasis>
</taxonomicName>
in the lack of a median tooth on the posterior margin of the telson.
</paragraph>
<paragraph id="8B832B148A626B0BFF7EFA8FFD88F98A" blockId="7.[151,1437,151,1653]" pageId="7" pageNumber="406">
The living coloration of the new species seems to be also characteristic, in particular in the milky white cephalothorax and anterior three pleomeres and the translucent posterior three pleomeres and tail fan (telson + uropods), giving a curious impression that the body of the shrimp can only half be seen. According to
<bibRefCitation id="EFAD56E58A626B0BFAE0FA36FF28FA3E" author="Bruce" pageId="7" pageNumber="406" refString="Bruce, A. J. (1983) Thor spinipes sp. nov., a new hippolytid shrimp from the Cobourg Peninsula, Northern Australia. The Beagle, Records of the Museums and Art Galleries of the Northern Territory Museum of Arts and Sciences, 1, 1 - 10." type="journal article" year="1983">Bruce (1983)</bibRefCitation>
, in
<taxonomicName id="4C3C50978A626B0BFEA8FA1AFE40FA3E" box="[273,395,1449,1472]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="B948F7068A626B0BFEA8FA1AFE40FA3E" box="[273,395,1449,1472]" italics="true" pageId="7" pageNumber="406">T. spinipes</emphasis>
</taxonomicName>
, the body is semitransparent but heavily mottled with superficial and deep patches of dull yellow-brown.
<taxonomicName id="4C3C50978A626B0BFEFAFA7FFE29FA1A" box="[323,482,1484,1509]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B948F7068A626B0BFEFAFA7FFE29FA1A" box="[323,482,1484,1509]" italics="true" pageId="7" pageNumber="406">Thor spinosus</emphasis>
</taxonomicName>
also shows a quite characteristic colour pattern on the body: the ground color of the body is dark olive brown with a superimposed pattern of fine striae; a pair of small dorsolateral, dark bluish-red eyespots is present on the third pleomere and a pair of similar spots is present on the endopods of the uropod; a single large median ventral spot is also present on the sixth pleomere (
<bibRefCitation id="EFAD56E58A626B0BFC01F98BFB8AF9AE" author="Bruce" box="[952,1089,1591,1616]" pageId="7" pageNumber="406" refString="Bruce, A. J. (1976) A report on a small collection of shrimps from the Kenya National Marine Parks at Malindi, with notes on selected species. Zoologische Verhandelingen, 145, 1 - 72." type="journal article" year="1976">Bruce 1976</bibRefCitation>
). No information on the living colouration is available for
<taxonomicName id="4C3C50978A626B0BFE72F9EEFDF7F98B" box="[459,572,1628,1653]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="406" phylum="Arthropoda" rank="species" species="cordelli">
<emphasis id="B948F7068A626B0BFE72F9EEFDF7F98B" box="[459,572,1628,1653]" italics="true" pageId="7" pageNumber="406">T. cordelli</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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