234 lines
35 KiB
XML
234 lines
35 KiB
XML
<document id="250CE18489A28A36B44B30DE952684A0" ID-CLB-Dataset="32021" ID-DOI="10.11646/zootaxa.4323.3.1" ID-GBIF-Dataset="f1be4566-adeb-4995-83c7-2aebab3c7458" ID-ISSN="1175-5326" ID-Zenodo-Dep="919606" ID-ZooBank="3E58F99B-2A30-437A-95Ae-D68A1D3Ecc45" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1506057765384" checkinUser="plazi" docAuthor="Schmidt, Joachim" docDate="2017" docId="03956A03A3467F50FF67F8AB8865DF16" docLanguage="en" docName="zootaxa.4323.3.1.pdf" docOrigin="Zootaxa 4323 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trichotichnus (Bottchrus) martensi Schmidt 2017, sp. n." docType="treatment" docVersion="7" lastPageNumber="346" masterDocId="FFAC127BA36D7F7DFFF0FFAE8B5CD930" masterDocTitle="Brachypterous ground beetles of the Trichotichnus subgenus Bottchrus Jedlička (Coleoptera, Carabidae) from the Himalaya, with description of fifteen new species" masterLastPageNumber="358" masterPageNumber="301" pageNumber="344" updateTime="1698468866275" updateUser="ExternalLinkService">
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<mods:title id="CB6E21C40BFE94BEE9A16071F9AA0C5A">Brachypterous ground beetles of the Trichotichnus subgenus Bottchrus Jedlička (Coleoptera, Carabidae) from the Himalaya, with description of fifteen new species</mods:title>
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<mods:namePart id="E82FD3DA25E334F9EBF95C538F8D3128">Schmidt, Joachim</mods:namePart>
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<mods:title id="7D59DC0CB89E143E9B4620DAA21CDC72">Zootaxa</mods:title>
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<mods:date id="E6D4EF5205B50A6D6D653EE6BA002960">2017</mods:date>
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<mods:number id="FAD46E479A07F2FCF11FAFE8052AD165">2017-09-22</mods:number>
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<treatment id="03956A03A3467F50FF67F8AB8865DF16" ID-DOI="http://doi.org/10.5281/zenodo.6015771" ID-GBIF-Taxon="134839429" ID-Zenodo-Dep="6015771" LSID="urn:lsid:plazi:treatment:03956A03A3467F50FF67F8AB8865DF16" httpUri="http://treatment.plazi.org/id/03956A03A3467F50FF67F8AB8865DF16" lastPageId="45" lastPageNumber="346" pageId="43" pageNumber="344">
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<subSubSection id="C326889EA3467F56FF67F8AB8A80DE70" pageId="43" pageNumber="344" type="nomenclature">
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<paragraph id="8B83DB15A3467F56FF67F8AB89D1DE2F" blockId="43.[151,653,1797,1856]" box="[151,653,1797,1823]" pageId="43" pageNumber="344">
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<heading id="D0CB6C79A3467F56FF67F8AB89D1DE2F" bold="true" box="[151,653,1797,1823]" fontSize="11" level="1" pageId="43" pageNumber="344" reason="1">
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<emphasis id="B9480707A3467F56FF67F8AB89D1DE2F" bold="true" box="[151,653,1797,1823]" pageId="43" pageNumber="344">
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<taxonomicName id="4C3CA096A3467F56FF67F8AB8918DE2F" ID-CoL="58LH9" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[151,580,1797,1823]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="344" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov." subGenus="Bottchrus">
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<emphasis id="B9480707A3467F56FF67F8AB8A1DDE2F" bold="true" box="[151,321,1797,1823]" italics="true" pageId="43" pageNumber="344">Trichotichnus</emphasis>
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(
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<emphasis id="B9480707A3467F56FEA2F8AB8A95DE2F" bold="true" box="[338,457,1797,1823]" italics="true" pageId="43" pageNumber="344">Bottchrus</emphasis>
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)
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<emphasis id="B9480707A3467F56FE2AF8AB8918DE2F" bold="true" box="[474,580,1797,1823]" italics="true" pageId="43" pageNumber="344">martensi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27BBA7CA3467F56FDBBF8AB89D1DE2F" box="[587,653,1797,1823]" pageId="43" pageNumber="344" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B83DB15A3467F56FF67F8898A80DE70" blockId="43.[151,653,1797,1856]" box="[151,476,1831,1856]" pageId="43" pageNumber="344">
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(
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<figureCitation id="1307C790A3467F56FF6FF8898A5FDE70" box="[159,259,1831,1856]" captionStart="FIGURES 120 – 123" captionStartId="41.[151,264,1942,1965]" captionTargetBox="[229,1356,202,1919]" captionTargetId="figure@41.[227,1360,193,1921]" captionTargetPageId="41" captionText="FIGURES 120 – 123. Himalayan species of Trichotichnus subgenus Bottchrus, photographs of habitus. 120, T. sikkimensis sp. n., holotype; 121, T. martensi sp. n., holotype; 122, T. hingstoni Andrewes, 1930, male from Kechu Pheri Lake; 123, T. birmanicus Bates, 1892, male from the Kali Gandaki, Tatopani." httpUri="https://zenodo.org/record/919672/files/figure.png" pageId="43" pageNumber="344">Figs 121</figureCitation>
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,
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<figureCitation id="1307C790A3467F56FEFFF8868A28DE70" box="[271,372,1832,1856]" captionStart="FIGURES 130 – 134" captionStartId="45.[151,264,1138,1161]" captionTargetBox="[222,1362,204,1113]" captionTargetId="figure@45.[208,1378,193,1118]" captionTargetPageId="45" captionText="FIGURES 130 – 134. Trichotichnus (Bottchrus) martensi sp. n. (holotype). 130, left metepisternum; 131, apex of last (VII) abdominal sternite, male; 132, 134, median lobe of aedeagus; 133, apical capitulum. 130 – 131, ventral view; 132, lateral view; 133, caudal view; 134, dorsal view. Scale bars = 0.25 mm (A: Figs 130 – 132, 134; B: Fig. 133)." httpUri="https://zenodo.org/record/919680/files/figure.png" pageId="43" pageNumber="344">130–134</figureCitation>
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,
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<figureCitation id="1307C790A3467F56FE8FF8898AF7DE70" box="[383,427,1831,1856]" captionStart="FIGURE 151" captionStartId="53.[151,250,1080,1102]" captionTargetBox="[151,1436,339,1059]" captionTargetId="figure@53.[151,1436,339,1059]" captionTargetPageId="53" captionText="FIGURE 151. Distribution of the Himalayan species of Trichotichnus subgenus Bottchrus. 1, T. holzschuhi Kirschenhofer, 1992; 2, T. baglungensis sp. n.; 3, T. parvulus sp. n.; 4, T. bubsaensis sp. n.; 5, T. schawalleri sp. n.; 6, T. obliquebasalis sp. n.; 7, T. panchhaseensis sp. n.; 8, T. siklesensis sp. n.; 9, T. pusillus sp. n.; 10, T. ganeshensis sp. n.; 11, T. minutus sp. n.; 12, T. cyanescens Ito, 1998; 13, T. martensi sp. n.; 14, T. opacus Ito, 1998." httpUri="https://zenodo.org/record/919694/files/figure.png" pageId="43" pageNumber="344">151</figureCitation>
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,
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<figureCitation id="1307C790A3467F56FE47F8868A8FDE70" box="[439,467,1832,1856]" captionStart="FIGURES 5 – 13" captionStartId="6.[151,264,1196,1219]" captionTargetBox="[152,1425,180,1152]" captionTargetId="figure@6.[151,1436,180,1193]" captionTargetPageId="6" captionText="FIGURES 5 – 13. Trichotichnus (Bottchrus) holzschuhi Kirschenhofer, 1992 (5 – 7, paratype; 8 – 13, specimens from Sete). 5, terminal lamella of aedeagus; 6 – 8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5 – 8, 10 – 13; B: Fig. 9)." httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="43" pageNumber="344">
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<emphasis id="B9480707A3467F56FE47F8868A8FDE70" box="[439,467,1832,1856]" italics="true" pageId="43" pageNumber="344">13</emphasis>
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</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C326889EA3467F56FF67F8C18E11DE9C" pageId="43" pageNumber="344" type="materials_examined">
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<paragraph id="8B83DB15A3467F56FF67F8C18E11DE9C" blockId="43.[151,1437,1903,2037]" pageId="43" pageNumber="344">
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<materialsCitation id="3B54D148A3467F56FF67F8C18E11DE9C" ID-GBIF-Occurrence="1655986723" collectingDate="1988-04-21" collectionCode="SMNS" county="Nepal-Expeditionen Jochen Martens" elevation="1900" location="Taplejung Distr." municipality="Worebung Pass" pageId="43" pageNumber="344" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<emphasis id="B9480707A3467F56FF67F8C18A1FDEB8" bold="true" box="[151,323,1903,1928]" pageId="43" pageNumber="344">
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<typeStatus id="548765B7A3467F56FF67F8C18B8EDEB8" box="[151,210,1903,1928]" pageId="43" pageNumber="344">Type</typeStatus>
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material
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</emphasis>
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.
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<typeStatus id="548765B7A3467F56FEA6F8C18A9FDEB8" box="[342,451,1903,1928]" pageId="43" pageNumber="344" type="holotype">Holotype</typeStatus>
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:
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<specimenCount id="9D3A109CA3467F56FE22F8DE8AB5DEB7" box="[466,489,1904,1927]" pageId="43" pageNumber="344" type="male">♂</specimenCount>
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, “333
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<location id="8EE38DCEA3467F56FDCDF8C189A6DEB8" LSID="urn:lsid:plazi:treatment:03956A03A3467F50FF67F8AB8865DF16:8EE38DCEA3467F56FDCDF8C189A6DEB8" box="[573,762,1903,1928]" county="Nepal-Expeditionen Jochen Martens" municipality="Worebung Pass" name="Taplejung Distr." pageId="43" pageNumber="344">Taplejung Distr.</location>
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, from
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<location id="8EE38DCEA3467F56FCBCF8C188CADEB8" LSID="urn:lsid:plazi:treatment:03956A03A3467F50FF67F8AB8865DF16:8EE38DCEA3467F56FCBCF8C188CADEB8" box="[844,918,1903,1928]" county="Nepal-Expeditionen Jochen Martens" municipality="Worebung Pass" name="Yektin" pageId="43" pageNumber="344">Yektin</location>
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to
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<collectingMunicipality id="6BE7416FA3467F56FC34F8C18F25DEB7" box="[964,1145,1903,1928]" pageId="43" pageNumber="344">Worebung Pass</collectingMunicipality>
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,
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<elevation id="00113C26A3467F56FB79F8DE8E70DEB7" box="[1161,1324,1904,1928]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="43" pageNumber="344" unit="m" value="1900.0" valueMax="2000.0" valueMin="1800.0">
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<quantity id="4CC476F0A3467F56FB79F8DE8E70DEB7" box="[1161,1324,1904,1928]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="43" pageNumber="344" unit="m" value="1900.0" valueMax="2000.0" valueMin="1800.0">1800–2000 m</quantity>
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</elevation>
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, tree-rich cultural land,
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<collectingDate id="EFC6043DA3467F56FEC2F83A8AF9DE9C" box="[306,421,1940,1964]" pageId="43" pageNumber="344" value="1988-04-21">
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||
<date id="FF82FDD5A3467F56FEC2F83A8AF9DE9C" box="[306,421,1940,1964]" pageId="43" pageNumber="344" value="1988-04-21">21 Apr 88</date>
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</collectingDate>
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, MARTENS & SCHAWALLER”, “
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<collectingCounty id="62E2A399A3467F56FCC7F83A8F89DE9C" box="[823,1237,1940,1965]" pageId="43" pageNumber="344">Nepal-Expeditionen Jochen Martens</collectingCounty>
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” (
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<collectionCode id="ED2D43D0A3467F56FB1EF83A8E1CDE9C" box="[1262,1344,1940,1964]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="43" pageNumber="344">SMNS</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C326889EA3467F51FF37F816889ADE7D" lastPageId="44" lastPageNumber="345" pageId="43" pageNumber="344" type="description">
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<paragraph id="8B83DB15A3467F51FF37F8168910D9E5" blockId="43.[151,1437,1903,2037]" lastBlockId="44.[151,1437,151,1977]" lastPageId="44" lastPageNumber="345" pageId="43" pageNumber="344">
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<emphasis id="B9480707A3467F56FF37F8168A11DEE1" bold="true" box="[199,333,1976,2001]" pageId="43" pageNumber="344">Description</emphasis>
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(male). Habitus as in
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<figureCitation id="1307C790A3467F56FDC7F81689C8DEE0" box="[567,660,1976,2001]" captionStart="FIGURES 120 – 123" captionStartId="41.[151,264,1942,1965]" captionTargetBox="[229,1356,202,1919]" captionTargetId="figure@41.[227,1360,193,1921]" captionTargetPageId="41" captionText="FIGURES 120 – 123. Himalayan species of Trichotichnus subgenus Bottchrus, photographs of habitus. 120, T. sikkimensis sp. n., holotype; 121, T. martensi sp. n., holotype; 122, T. hingstoni Andrewes, 1930, male from Kechu Pheri Lake; 123, T. birmanicus Bates, 1892, male from the Kali Gandaki, Tatopani." httpUri="https://zenodo.org/record/919672/files/figure.png" pageId="43" pageNumber="344">Fig. 121</figureCitation>
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. Comparatively robust and convex. Body length
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<quantity id="4CC476F0A3467F56FB50F8168FA9DEE0" box="[1184,1269,1976,2001]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="43" pageNumber="344" unit="mm" value="6.6">6.6 mm</quantity>
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, width 3.0 mm. Colour: Body black piceous, shiny on dorsum, with light green tinge on pronotum and elytra; labrum, very narrow margins of pronotum and apical portion of elytral margins paler, reddish brown; palpi, antennae and legs brownish yellow, femora not infuscate.
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</paragraph>
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<paragraph id="8B83DB15A3417F51FF37FF718E7FDB50" blockId="44.[151,1437,151,1977]" pageId="44" pageNumber="345">
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Head: Large, with wide neck (HWmax/PWmax = 0.68; HWmin/PWmax = 0.55). Eyes markedly convex (HWmax/HWmin = 1.22), somewhat wide oval in lateral view. Tempora about half as long as eye, slightly convex, somewhat abrupt to neck. Genae wide, about 1.7 as wide as width of antennomere 1, with few short setae. Lateral margin of head between antenna and eye slightly protruded laterally, forming an obtuse angle near eyes, distinctly bordered. Area between supraorbital furrow and upper margin of eye slightly narrower than width of antennomere 2 basally. Supraorbital setigerous pore located just behind level of posterior margin of eye, separated from supraorbital furrow approximately by width of antennomere 3 apically. Labrum moderately concave anteriorly. Clypeus slightly emarginate along anterior margin, slightly convex, with one setigerous pore at each apical angle. Frons and vertex convex.
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<collectingRegion id="49F815F7A3417F51FE4BFDAE8946DB27" box="[443,538,512,535]" country="Myanmar" name="Chin" pageId="44" pageNumber="345">Mentum</collectingRegion>
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and submentum completely fused, submentum on each side with one long seta. Left mandible not truncate, slightly blunted at apex. Dorsal microsculpture not evident. Antennae comparatively slender, slightly surpassing basal edge of pronotum, with antennomeres 4–8 markedly longer than wide.
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</paragraph>
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<paragraph id="8B83DB15A3417F51FF37FDC38F84DD20" blockId="44.[151,1437,151,1977]" pageId="44" pageNumber="345">Pronotum: Comparatively wide (PWmax/PL = 1.42), widest slightly before middle, moderately narrowed posteriad (PWmax/PWmin = 1.17), with one lateral seta inserted slightly before widest point. Sides rounded anteriorly, almost straightly converging in posterior third. Apical margin slightly emarginate, almost straight in middle portion, bordered throughout, with very fine border at middle. Apical angles rounded, very slightly protruded anteriad. Basal margin slightly sinuate bilaterally, broadly rounded at middle and at basal angles, vaguely bordered almost throughout, noticeably longer than apical margin and only slightly shorter than base of elytra between humeral angles. Basal angles obtuse, not denticulate, slightly blunted at tip. Pronotal disc convex, markedly sloped to apical angles and moderately so to basal ones, not depressed basally; small area near basal angles slightly flattened. Lateral gutter very narrow basally, very slightly widened in apical half. Median line very fine, superficial, crenulate, ended markedly before apical and basal margins. Basal foveae absent. Surface very finely and densely punctate along base and in latero-basal area, also with very fine scattered punctures at apical margin medially. Dorsal microsculpture present throughout, consisting of fine transverse meshes.</paragraph>
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<paragraph id="8B83DB15A3417F51FF37FBB28962DC48" blockId="44.[151,1437,151,1977]" pageId="44" pageNumber="345">Elytra: In lateral and caudal view convex, in dorsal view oval, moderately wide (EL/EW = 1.35; EL/PL = 2.38; EW/PWmax = 1.24), widest at middle, with somewhat acute apex. Humeri prominent, angularly rounded at tip, without denticle. Subapical sinuation weak but distinct. Sutural angle acutangular, not blunted at tip. Basal edge slightly sinuate laterally, forming a very obtuse angle with lateral margin. Striae smooth, striae 1–5 somewhat superficial, striae 6–8 slightly impressed along entire length. Parascutellar setigerous pore present, large. Parascutellar striole on right elytron as long as distance from parascutellar pore to suture, with apex free, on left elytron slightly longer, with apex connected with stria 1. Intervals nearly flat, external ones subconvex apically. Interval 3 with one discal setigerous pore at stria 2 behind middle. Marginal umbilicate series widely interrupted at middle, with 8–9 pores in anterior group and 9–10 pores in posterior group. Microsculpture strongly obliterate, consisting of poorly visible thin lines.</paragraph>
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<paragraph id="8B83DB15A3417F51FF37FA2A890FDCAD" blockId="44.[151,1437,151,1977]" box="[199,595,1412,1437]" pageId="44" pageNumber="345">Hindwings: Reduced to tiny scales.</paragraph>
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<paragraph id="8B83DB15A3417F51FF37FA098E30DCD4" blockId="44.[151,1437,151,1977]" pageId="44" pageNumber="345">
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Ventral surface: Prosternum and metasternum finely pubescent. Metepisternum (
|
||
<figureCitation id="1307C790A3417F51FBA7FA098FEBDCF0" box="[1111,1207,1447,1472]" captionStart="FIGURES 130 – 134" captionStartId="45.[151,264,1138,1161]" captionTargetBox="[222,1362,204,1113]" captionTargetId="figure@45.[208,1378,193,1118]" captionTargetPageId="45" captionText="FIGURES 130 – 134. Trichotichnus (Bottchrus) martensi sp. n. (holotype). 130, left metepisternum; 131, apex of last (VII) abdominal sternite, male; 132, 134, median lobe of aedeagus; 133, apical capitulum. 130 – 131, ventral view; 132, lateral view; 133, caudal view; 134, dorsal view. Scale bars = 0.25 mm (A: Figs 130 – 132, 134; B: Fig. 133)." httpUri="https://zenodo.org/record/919680/files/figure.png" pageId="44" pageNumber="345">Fig. 130</figureCitation>
|
||
) slightly wider than long, markedly narrowed posteriad. Apex of last visible (VII) abdominal sternite (in male) rounded (
|
||
<figureCitation id="1307C790A3417F51FB0DFA628E03DCD4" box="[1277,1375,1484,1509]" captionStart="FIGURES 130 – 134" captionStartId="45.[151,264,1138,1161]" captionTargetBox="[222,1362,204,1113]" captionTargetId="figure@45.[208,1378,193,1118]" captionTargetPageId="45" captionText="FIGURES 130 – 134. Trichotichnus (Bottchrus) martensi sp. n. (holotype). 130, left metepisternum; 131, apex of last (VII) abdominal sternite, male; 132, 134, median lobe of aedeagus; 133, apical capitulum. 130 – 131, ventral view; 132, lateral view; 133, caudal view; 134, dorsal view. Scale bars = 0.25 mm (A: Figs 130 – 132, 134; B: Fig. 133)." httpUri="https://zenodo.org/record/919680/files/figure.png" pageId="44" pageNumber="345">Fig. 131</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B83DB15A3417F51FF37FA5E888BDF45" blockId="44.[151,1437,151,1977]" pageId="44" pageNumber="345">Legs: Protibia not sulcate on upper surface. Profemur with four and six setae on anterio-ventral margin. Tarsi short, metatarsus noticeably shorter than HWmin, with metatarsomeres (particularly 2–4) moderately widened posteriad, metatarsomere 1 approximately as long as metatarsomeres 2+3. Tarsomere 5 with two pairs of ventrolateral setae. Pro- and mesotarsi (of male) comparatively weakly widened.</paragraph>
|
||
<paragraph id="8B83DB15A3417F51FF37F9D1889ADE7D" blockId="44.[151,1437,151,1977]" pageId="44" pageNumber="345">
|
||
Male genitalia: Median lobe of aedeagus (
|
||
<figureCitation id="1307C790A3417F51FD43F9D1880DDFA8" box="[691,849,1663,1688]" captionStart="FIGURES 130 – 134" captionStartId="45.[151,264,1138,1161]" captionTargetBox="[222,1362,204,1113]" captionTargetId="figure@45.[208,1378,193,1118]" captionTargetPageId="45" captionText="FIGURES 130 – 134. Trichotichnus (Bottchrus) martensi sp. n. (holotype). 130, left metepisternum; 131, apex of last (VII) abdominal sternite, male; 132, 134, median lobe of aedeagus; 133, apical capitulum. 130 – 131, ventral view; 132, lateral view; 133, caudal view; 134, dorsal view. Scale bars = 0.25 mm (A: Figs 130 – 132, 134; B: Fig. 133)." httpUri="https://zenodo.org/record/919680/files/figure.png" pageId="44" pageNumber="345">Figs 132–134</figureCitation>
|
||
) relatively large, in lateral view strongly bent just behind basal bulb, with ventral margin very widely sinuate in medial portion and slightly convex in apical third; in dorsal view median lobe in distal half evenly narrowed to apex and slightly curved to the right side. Terminal lamella in dorsal view short and narrow, with sides converging apicad. Apical capitulum in lateral view (
|
||
<figureCitation id="1307C790A3417F51FAC4F9428EC8DE34" box="[1332,1428,1772,1797]" captionStart="FIGURES 130 – 134" captionStartId="45.[151,264,1138,1161]" captionTargetBox="[222,1362,204,1113]" captionTargetId="figure@45.[208,1378,193,1118]" captionTargetPageId="45" captionText="FIGURES 130 – 134. Trichotichnus (Bottchrus) martensi sp. n. (holotype). 130, left metepisternum; 131, apex of last (VII) abdominal sternite, male; 132, 134, median lobe of aedeagus; 133, apical capitulum. 130 – 131, ventral view; 132, lateral view; 133, caudal view; 134, dorsal view. Scale bars = 0.25 mm (A: Figs 130 – 132, 134; B: Fig. 133)." httpUri="https://zenodo.org/record/919680/files/figure.png" pageId="44" pageNumber="345">Fig. 132</figureCitation>
|
||
) large, slightly oblique, approximately equally prominent ventrally and dorsally, in caudal view (
|
||
<figureCitation id="1307C790A3417F51FB27F8A18E6BDE18" box="[1239,1335,1807,1832]" captionStart="FIGURES 130 – 134" captionStartId="45.[151,264,1138,1161]" captionTargetBox="[222,1362,204,1113]" captionTargetId="figure@45.[208,1378,193,1118]" captionTargetPageId="45" captionText="FIGURES 130 – 134. Trichotichnus (Bottchrus) martensi sp. n. (holotype). 130, left metepisternum; 131, apex of last (VII) abdominal sternite, male; 132, 134, median lobe of aedeagus; 133, apical capitulum. 130 – 131, ventral view; 132, lateral view; 133, caudal view; 134, dorsal view. Scale bars = 0.25 mm (A: Figs 130 – 132, 134; B: Fig. 133)." httpUri="https://zenodo.org/record/919680/files/figure.png" pageId="44" pageNumber="345">Fig. 133</figureCitation>
|
||
) narrow, elongate oval, with acute apex. Internal sac with a spiny patch at middle.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C326889EA3417F50FF37F8F98E54DCEE" lastPageId="45" lastPageNumber="346" pageId="44" pageNumber="345" type="etymology">
|
||
<paragraph id="8B83DB15A3417F51FF37F8F98AFCDE89" blockId="44.[151,1437,151,1977]" pageId="44" pageNumber="345">
|
||
<emphasis id="B9480707A3417F51FF37F8F98A1ADE40" bold="true" box="[199,326,1879,1904]" pageId="44" pageNumber="345">Etymology</emphasis>
|
||
. This new species is named in honour of Prof. Dr. Jochen Martens (Mainz), renowned biogeographer of High Asia and participant of the 1988 expedition to the
|
||
<collectingRegion id="49F815F7A3417F51FC38F8D38F40DEA4" box="[968,1052,1917,1940]" country="Sri Lanka" name="Eastern" pageId="44" pageNumber="345">Eastern</collectingRegion>
|
||
<collectingCountry id="F32B9B85A3417F51FBD3F8D28F3ADEA5" box="[1059,1126,1916,1941]" name="Nepal" pageId="44" pageNumber="345">Nepal</collectingCountry>
|
||
Himalaya during which the new species was found.
|
||
</paragraph>
|
||
<caption id="DF438B9DA3407F50FF67FBDC8F06DDF6" httpUri="https://zenodo.org/record/919680/files/figure.png" pageId="45" pageNumber="346" targetBox="[222,1362,204,1113]" targetPageId="45">
|
||
<paragraph id="8B83DB15A3407F50FF67FBDC8F06DDF6" blockId="45.[151,1436,1138,1222]" pageId="45" pageNumber="346">
|
||
<emphasis id="B9480707A3407F50FF67FBDC8A2CDDB8" bold="true" box="[151,368,1138,1161]" pageId="45" pageNumber="346">FIGURES 130–134.</emphasis>
|
||
<taxonomicName id="4C3CA096A3407F50FE88FBDD89BEDDB9" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[376,738,1139,1161]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="346" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A3407F50FE88FBDD8958DDB9" box="[376,516,1139,1161]" italics="true" pageId="45" pageNumber="346">Trichotichnus</emphasis>
|
||
(
|
||
<emphasis id="B9480707A3407F50FDE5FBDD892BDDB9" box="[533,631,1139,1161]" italics="true" pageId="45" pageNumber="346">Bottchrus</emphasis>
|
||
)
|
||
<emphasis id="B9480707A3407F50FD79FBDA89BEDDB9" box="[649,738,1140,1161]" italics="true" pageId="45" pageNumber="346">martensi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA3407F50FD1BFBDD887ADDB8" box="[747,806,1139,1160]" pageId="45" pageNumber="346" rank="species">
|
||
<emphasis id="B9480707A3407F50FD1BFBDD887ADDB8" bold="true" box="[747,806,1139,1160]" pageId="45" pageNumber="346">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(holotype). 130, left metepisternum; 131, apex of last (VII) abdominal sternite, male; 132, 134, median lobe of aedeagus; 133, apical capitulum. 130–131, ventral view; 132, lateral view; 133, caudal view; 134, dorsal view. Scale bars = 0.25 mm (A: Figs 130–132, 134; B: Fig. 133).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B83DB15A3407F50FF37FB438E54DCEE" blockId="45.[151,1436,1261,1574]" pageId="45" pageNumber="346">
|
||
<emphasis id="B9480707A3407F50FF37FB438A06DC36" bold="true" box="[199,346,1261,1286]" pageId="45" pageNumber="346">Comparison</emphasis>
|
||
. This new species is most similar to
|
||
<taxonomicName id="4C3CA096A3407F50FD03FB4188E0DC36" authorityName="Ito" authorityYear="1998" box="[755,956,1262,1286]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="346" phylum="Arthropoda" rank="species" species="cyanescens" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A3407F50FD03FB418854DC36" box="[755,776,1263,1286]" italics="true" pageId="45" pageNumber="346">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A3407F50FCE7FB418874DC36" box="[791,808,1263,1286]" italics="true" pageId="45" pageNumber="346">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A3407F50FCCEFB4188E0DC36" box="[830,956,1263,1286]" italics="true" pageId="45" pageNumber="346">cyanescens</emphasis>
|
||
</taxonomicName>
|
||
and particularly to
|
||
<taxonomicName id="4C3CA096A3407F50FB65FB418E7DDC36" authorityName="Ito" authorityYear="1998" box="[1173,1313,1261,1286]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="346" phylum="Arthropoda" rank="species" species="loebli" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A3407F50FB65FB418FF6DC36" box="[1173,1194,1263,1286]" italics="true" pageId="45" pageNumber="346">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A3407F50FB49FB418F96DC36" box="[1209,1226,1263,1286]" italics="true" pageId="45" pageNumber="346">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A3407F50FB11FB438E7DDC36" box="[1249,1313,1261,1286]" italics="true" pageId="45" pageNumber="346">loebli</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3CA096A3407F50FAA6FB418A4BDC1B" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="346" phylum="Arthropoda" rank="species" species="sikkimensis" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A3407F50FAA6FB418E37DC36" box="[1366,1387,1263,1286]" italics="true" pageId="45" pageNumber="346">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A3407F50FA8AFB418ED7DC36" box="[1402,1419,1263,1286]" italics="true" pageId="45" pageNumber="346">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A3407F50FF67FABC8A4BDC1B" box="[151,279,1298,1323]" italics="true" pageId="45" pageNumber="346">sikkimensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA3407F50FED3FABC8A3BDC1B" box="[291,359,1298,1323]" pageId="45" pageNumber="346" rank="species">
|
||
<emphasis id="B9480707A3407F50FED3FABC8A3BDC1B" bold="true" box="[291,359,1298,1323]" pageId="45" pageNumber="346">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in comparatively large size and habitus, also in having short metepisternum, not infuscate femora, pronotal lateral gutter slightly widened anteriad, pronotal and elytral discs with fine microsculpture, and humerus without denticle at tip. In addition, the pronotal basal angles are slightly blunted at apex as in
|
||
<taxonomicName id="4C3CA096A3407F50FAA2FAF58B8BDCA6" authorityName="Ito" authorityYear="1998" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="346" phylum="Arthropoda" rank="species" species="loebli" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A3407F50FAA2FAF58E3BDC42" box="[1362,1383,1371,1394]" italics="true" pageId="45" pageNumber="346">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A3407F50FA8AFAF58ED7DC42" box="[1402,1419,1371,1394]" italics="true" pageId="45" pageNumber="346">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A3407F50FF67FAD38B8BDCA6" box="[151,215,1405,1430]" italics="true" pageId="45" pageNumber="346">loebli</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C3CA096A3407F50FF14FAD38944DCA6" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[228,536,1405,1430]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="346" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A3407F50FF14FAD38A22DCA6" box="[228,382,1405,1430]" italics="true" pageId="45" pageNumber="346">Trichotichnus</emphasis>
|
||
(
|
||
<emphasis id="B9480707A3407F50FE7EFAD18AC3DCA6" box="[398,415,1407,1430]" italics="true" pageId="45" pageNumber="346">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A3407F50FE46FAD18944DCA6" box="[438,536,1407,1430]" italics="true" pageId="45" pageNumber="346">martensi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA3407F50FDEEFAD38901DCA6" box="[542,605,1405,1430]" pageId="45" pageNumber="346" rank="species">
|
||
<emphasis id="B9480707A3407F50FDEEFAD38901DCA6" bold="true" box="[542,605,1405,1430]" pageId="45" pageNumber="346">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
markedly differs from all these species in having mentum and submentum completely fused, protibia not sulcate on dorsal side, and median lobe of the aedeagus in lateral view straighter, with almost straight ventral margin and with a very large slightly oblique apical capitulum (
|
||
<figureCitation id="1307C790A3407F50FB69FA6B8FA7DCEE" box="[1177,1275,1477,1502]" captionStart="FIGURES 130 – 134" captionStartId="45.[151,264,1138,1161]" captionTargetBox="[222,1362,204,1113]" captionTargetId="figure@45.[208,1378,193,1118]" captionTargetPageId="45" captionText="FIGURES 130 – 134. Trichotichnus (Bottchrus) martensi sp. n. (holotype). 130, left metepisternum; 131, apex of last (VII) abdominal sternite, male; 132, 134, median lobe of aedeagus; 133, apical capitulum. 130 – 131, ventral view; 132, lateral view; 133, caudal view; 134, dorsal view. Scale bars = 0.25 mm (A: Figs 130 – 132, 134; B: Fig. 133)." httpUri="https://zenodo.org/record/919680/files/figure.png" pageId="45" pageNumber="346">Fig. 132</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C326889EA3407F50FF37FA448865DF16" pageId="45" pageNumber="346" type="distribution">
|
||
<paragraph id="8B83DB15A3407F50FF37FA448865DF16" blockId="45.[151,1436,1261,1574]" pageId="45" pageNumber="346">
|
||
<emphasis id="B9480707A3407F50FF37FA448A05DF33" bold="true" box="[199,345,1514,1539]" pageId="45" pageNumber="346">Distribution</emphasis>
|
||
(
|
||
<figureCitation id="1307C790A3407F50FE9BFA448A93DF33" box="[363,463,1514,1539]" captionStart="FIGURE 151" captionStartId="53.[151,250,1080,1102]" captionTargetBox="[151,1436,339,1059]" captionTargetId="figure@53.[151,1436,339,1059]" captionTargetPageId="53" captionText="FIGURE 151. Distribution of the Himalayan species of Trichotichnus subgenus Bottchrus. 1, T. holzschuhi Kirschenhofer, 1992; 2, T. baglungensis sp. n.; 3, T. parvulus sp. n.; 4, T. bubsaensis sp. n.; 5, T. schawalleri sp. n.; 6, T. obliquebasalis sp. n.; 7, T. panchhaseensis sp. n.; 8, T. siklesensis sp. n.; 9, T. pusillus sp. n.; 10, T. ganeshensis sp. n.; 11, T. minutus sp. n.; 12, T. cyanescens Ito, 1998; 13, T. martensi sp. n.; 14, T. opacus Ito, 1998." httpUri="https://zenodo.org/record/919694/files/figure.png" pageId="45" pageNumber="346">Fig. 151</figureCitation>
|
||
,
|
||
<figureCitation id="1307C790A3407F50FE2EFA448AA6DF32" box="[478,506,1514,1538]" captionStart="FIGURES 5 – 13" captionStartId="6.[151,264,1196,1219]" captionTargetBox="[152,1425,180,1152]" captionTargetId="figure@6.[151,1436,180,1193]" captionTargetPageId="6" captionText="FIGURES 5 – 13. Trichotichnus (Bottchrus) holzschuhi Kirschenhofer, 1992 (5 – 7, paratype; 8 – 13, specimens from Sete). 5, terminal lamella of aedeagus; 6 – 8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5 – 8, 10 – 13; B: Fig. 9)." httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="45" pageNumber="346">
|
||
<emphasis id="B9480707A3407F50FE2EFA448AA6DF32" box="[478,506,1514,1538]" italics="true" pageId="45" pageNumber="346">13</emphasis>
|
||
</figureCitation>
|
||
). Up to today only found on the western slope of the Singalila mountain range, Taplejung District, East
|
||
<collectingCountry id="F32B9B85A3407F50FE56F9A38AB0DF16" box="[422,492,1549,1574]" name="Nepal" pageId="45" pageNumber="346">Nepal</collectingCountry>
|
||
, at altitudes of
|
||
<quantity id="4CC476F0A3407F50FD67F9A08869DF16" box="[663,821,1550,1574]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="45" pageNumber="346" unit="m" value="1900.0" valueMax="2000.0" valueMin="1800.0">1800–2000 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |