114 lines
19 KiB
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114 lines
19 KiB
XML
<document id="80E9E50706D3B2F1A7800BC4296D40A8" ID-DOI="10.1206/0003-0082(2000)287<0001:ROTNTT>2.0.CO;2" ID-ISSN="0003-0082" ID-Zenodo-Dep="10110744" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630265040447" checkinUser="carolina" docAuthor="SMETANA, ALEŠ & DAVIES, ANTHONY" docDate="2000" docId="0384878FD831FB3CFCA5694E0836FE27" docLanguage="en" docName="N3287.pdf" docOrigin="American Museum Novitates 3287" docSource="http://www.bioone.org/doi/abs/10.1206/0003-0082%282000%29287%3C0001%3AROTNTT%3E2.0.CO%3B2" docStyle="DocumentStyle:059C9B7006157ECE636E30DEF6793B3A.3:AmMusNovit.2000-2010.journal_article.type2" docStyleId="059C9B7006157ECE636E30DEF6793B3A" docStyleName="AmMusNovit.2000-2010.journal_article.type2" docStyleVersion="3" docTitle="Staphylinus Sensu Lato, Including Comments 1758" docType="treatment" docVersion="3" masterDocId="FFBDFFF7D829FB25FFD86B530951FFD4" masterDocTitle="Reclassification of the North Temperate Taxa Associated with Staphylinus Sensu Lato, Including Comments on Relevant Subtribes of Staphylinini (Coleoptera: Staphylinidae)" masterLastPageNumber="1" masterPageNumber="1" pageNumber="1" updateTime="1699682540511" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="05A48238381A54E5A9B14E60F62DD846">Reclassification of the North Temperate Taxa Associated with Staphylinus Sensu Lato, Including Comments on Relevant Subtribes of Staphylinini (Coleoptera: Staphylinidae)</mods:title>
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<mods:namePart id="FB57E0AFA3225AD6A48D5971446E7C19">SMETANA, ALEŠ</mods:namePart>
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<mods:namePart id="D7970D479E9FF70CD26DE98D7D0D9803">DAVIES, ANTHONY</mods:namePart>
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<mods:typeOfResource id="937AFF06A1E897BCF111D17E400119B1">text</mods:typeOfResource>
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<mods:title id="F69370F4BB7F2365B3CA05AB3E1F89E5">American Museum Novitates</mods:title>
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</mods:titleInfo>
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<mods:part id="F7AE06CA7B64868BBC61A8E3EDFC52A1">
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<mods:date id="3EBCC6815FF26AE94EC9EE1F94F67184">2000</mods:date>
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<mods:detail id="FD8180421E5159A9B7782F251471AED3" type="pubDate">
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<mods:number id="A66D3B8CD3D9F01DBB77EAFC6DABC46F">2000-02-02</mods:number>
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<mods:detail id="BF8104A57EB7185F1179106B97AC522C" type="volume">
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<mods:number id="9014A06E558BB69E71D9EDE20756D1E7">3287</mods:number>
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<mods:url id="599F729D80723429CDBCB5EF4CEFB283">http://www.bioone.org/doi/abs/10.1206/0003-0082%282000%29287%3C0001%3AROTNTT%3E2.0.CO%3B2</mods:url>
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<mods:classification id="B81AFD859EF484D0168031089CAB4B9C">journal article</mods:classification>
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<mods:identifier id="963882FC70E487A7817AAE35B461C5B1" type="DOI">10.1206/0003-0082(2000)287<0001:ROTNTT>2.0.CO;2</mods:identifier>
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<mods:identifier id="CE89366410665694658B139B5D76A0EA" type="ISSN">0003-0082</mods:identifier>
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<treatment id="0384878FD831FB3CFCA5694E0836FE27" LSID="urn:lsid:plazi:treatment:0384878FD831FB3CFCA5694E0836FE27" httpUri="http://treatment.plazi.org/id/0384878FD831FB3CFCA5694E0836FE27" lastPageId="25" pageId="24">
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<subSubSection id="C3376512D831FB3DFCA5694E0C65FDE1" box="[893,1332,541,565]" pageId="24" type="nomenclature">
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<paragraph id="8B923699D831FB3DFCA5694E0C65FDE1" blockId="24.[893,1332,541,565]" box="[893,1332,541,565]" pageId="24">
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<heading id="D0DA81F5D831FB3DFCA5694E0C65FDE1" box="[893,1332,541,565]" centered="true" fontSize="36" level="2" pageId="24" reason="8">
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<taxonomicName id="4C2D4D1AD831FB3DFCA5694E0D5AFDE1" ID-CoL="8VS43" authorityName="Sensu Lato, Including Comments" authorityYear="1758" box="[893,1035,541,565]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="24" phylum="Arthropoda" rank="genus">
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<emphasis id="B959EA8BD831FB3DFCA5694E0D5AFDE1" bold="true" box="[893,1035,541,565]" italics="true" pageId="24">Staphylinus</emphasis>
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</taxonomicName>
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Linne´, 1758 (
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<emphasis id="B959EA8BD831FB3DFB6F694E0C78FDE1" bold="true" box="[1207,1321,541,565]" pageId="24">stat. nov.</emphasis>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3376512D831FB3DFCB569020AB6FD51" pageId="24" type="materials_examined">
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<paragraph id="8B923699D831FB3DFCB569020AB6FD51" blockId="24.[848,1377,593,1731]" pageId="24">
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<materialsCitation id="3B453CC4D831FB3DFCB569020AB2FD51" collectionCode="SPECIES" pageId="24" specimenCount="1" typeStatus="holotype">
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<typeStatus id="5496883BD831FB3DFCB569020AF6FDB3" box="[877,935,593,616]" pageId="24">TYPE</typeStatus>
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<collectionCode id="ED3CAE5CD831FB3DFC6169020D41FDB3" box="[953,1040,593,616]" pageId="24">SPECIES</collectionCode>
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:
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<taxonomicName id="4C2D4D1AD831FB3DFBF269020C30FDBC" authorityName="Linnaeus" authorityYear="1758" box="[1066,1377,593,616]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="24" phylum="Arthropoda" rank="species" species="erythropterus">
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<emphasis id="B959EA8BD831FB3DFBF269020C30FDBC" box="[1066,1377,593,616]" italics="true" pageId="24">Staphylinus erythropterus</emphasis>
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</taxonomicName>
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Linne´, 1758
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3376512D831FB3CFCB569DF0836FE27" lastPageId="25" pageId="24" type="discussion">
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<paragraph id="8B923699D831FB3DFCB569DF0DA3FBF4" blockId="24.[848,1377,593,1731]" pageId="24">
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COMMENTS: All members of this genus share the synapomorphy of the middle coxal cavities not margined posteromedially (
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<figureCitation id="13162A1CD831FB3DFAE169950A20FD2F" captionStart="Figs" captionStartId="70.[298,346,1603,1624]" captionTargetBox="[285,1362,187,1582]" captionTargetId="figure-36@70.[285,1362,187,1582]" captionTargetPageId="70" captionText="Figs. 59–64. 59. Dinothenarus pubescens, series of long setae (some represented only by their sockets) on mesosternum. 60. Miobdelus sp., series of long setae on mesosternum. 61. Craspedomerus sp., series of long setae on mesosternum without transverse ridge. 62. Tasgius melanarius, mesocoxal acetabulum. 63. Ocypus weisei, mesocoxal acetabulum. 64. Staphylinus dimidiaticornis, mesocoxal cavities, not margined posteromedially." figureDoi="http://doi.org/10.5281/zenodo.10110770" httpUri="https://zenodo.org/record/10110770/files/figure.png" pageId="24">fig. 64</figureCitation>
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). In addition, they share (
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<bibRefCitation id="EFBC4B68D831FB3DFB7369B70DE6FD2F" author="Natural History Museum, London" box="[1195,1207,740,763]" pageId="24" pagination="204" refId="ref25242" refString="1. Ocypus almorensis (Cameron, 1932). Cameron (1932: 204) described the species (in Staphylinus) from specimens from '' West Almora: Kali Valley, alt. 9000 feet' ' and '' Punjab: Triun, alt. 9300 feet. '' We were able to study one of the male syntypes in the Natural History Museum, London, England, labelled as follows: '' Triun, 9,300 ft Dharm- sala. Punjab. H. G. C. / M. Cameron. Bequest. B. M. 1955 - 147. / Staphylinus (Ocypus) almorensis. '' The specimen is in very good shape. It was dissected, and the aedeagus and the genital segment were glued to the plate with the beetle. It is hereby designated as the lectotype of Staphylinus almorensis; the label '' LECTOTYPE Staphylinus almorensis Cameron A. Smetana des. 1999 '' was attached to it. The specimen cannot be distinguished from those of Ocypus helleni J. Muller that occur in northern Pakistan and in Kashmir. The name O. almorensis becomes a junior synonym of O. helleni (syn. nov.). The corresponding determination label was attached to the specimen." type="journal article" year="1955">1</bibRefCitation>
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) superior line of pronotal hypomeron markedly and relatively abruptly deflexed ventrad, before or at most at middle of pronotum, with its anterior portion situated markedly below basal portion in lateral view (
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<figureCitation id="13162A1CD831FB3DFB9368250DFCFC59" box="[1099,1197,886,909]" captionStart="Figs" captionStartId="79.[264,312,1523,1544]" captionTargetBox="[272,1307,179,1502]" captionTargetId="figure-34@79.[272,1307,179,1502]" captionTargetPageId="79" captionText="Figs. 111–116. 111. Dinothenarus pubescens, pronotum showing both marginal lines of pronotal hypomeron, and two lateral macrosetae, lateral view. 112. Staphylinus erythropterus, pronotum showing both marginal lines of pronotal hypomeron, and two lateral macrosetae, lateral view. 113. Ocypus ophthalmicus ophthalmicus, pronotum showing both marginal lines of pronotal hypomeron, and two lateral macrosetae, ventrolateral view. 114. Ocypus nitens semialatus, pronotum showing both marginal lines of pronotal hypomeron, and two lateral macrosetae, lateral view. 115. Physetops sp., dorsal face of neck, with dorsal portion of nuchal ridge missing. 116. Ontholestes cingulatus, dorsal face of neck, with nuchal ridge." figureDoi="http://doi.org/10.5281/zenodo.10110788" httpUri="https://zenodo.org/record/10110788/files/figure.png" pageId="24">fig. 112</figureCitation>
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); (
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<bibRefCitation id="EFBC4B68D831FB3DFB1768250D8EFC59" author="Ocypus" box="[1231,1247,886,909]" pageId="24" pagination="170" refId="ref25465" refString="2. Ocypus auricomus (Lindberg, 1953: 4), described in Staphylinus, is preoccupied by Staphylinus auricomus Brulle', 1842: pl. 5, fig. 6, now a synonym of Glenus chrysis (Gravenhorst, 1806), and by Staphylinus auricomus Cameron, 1929: 65. The name Ocypus mateui (Coiffait, 1954: 170), listed at present as a junior synonym of O. auricomus, becomes the valid name of this species (stat. nov.)." type="journal article" year="1842">2</bibRefCitation>
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) bicolored black body with red elytra and legs red excluding coxae; and (
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<bibRefCitation id="EFBC4B68D831FB3DFB8968E20D31FC1C" author="Lordithon" box="[1105,1120,945,968]" pageId="24" refId="ref25558" refString="3. Ocypus bimaculatus (Cameron, 1932: 207), described in Staphylinus, is preoccupied by several names in Staphylinus, the oldest one being Staphylinus bimaculatus Schrank, 1798: 644, now a synonym of Lordithon trinotatus (Erichson, 1839 b: 409). The replacement name Ocypus cameroni, nom. nov., is hereby proposed for Ocypus bimaculatus (Cameron, 1932)." type="journal volume" year="1839">3</bibRefCitation>
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) presence of yellow tomentose pubescence on head and usually also on pronotum, and paired yellow tomentose patches on abdominal tergites.
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</paragraph>
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<paragraph id="8B923699D831FB3CFCB56F750BD9FEB4" blockId="24.[848,1377,593,1731]" lastBlockId="25.[238,767,182,499]" lastPageId="25" pageId="24">
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Further character states include: right mandible with one stout, apically truncate, subemarginate or slightly emarginate tooth, left mandible with two dorsal teeth, and with one somewhat smaller tooth in more ventral plane (
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<figureCitation id="13162A1CD831FB3DFC7D6FEA0D58FB04" box="[933,1033,1209,1232]" captionStart="Figs" captionStartId="83.[264,312,1571,1592]" captionTargetBox="[242,1338,176,1549]" captionTargetId="figure-34@83.[242,1338,176,1549]" captionTargetPageId="83" captionText="Figs. 136–141. 136. Platydracus stercorarius, pronotal hypomeron, ventral view. 137. Platydracus stercorarius, right coxal cavity, ventral view. 138. Staphylinus dimidiaticornis, left mandible, ventral view. 139. Staphylinus erythropterus, last segment of labial palpus. 140. Dinothenarus pubescens, posteromedial portion of head and neck with pubescence, dorsal view. 141. Dinothenarus pubescens, pronotum with pubescence." figureDoi="http://doi.org/10.5281/zenodo.10110796" httpUri="https://zenodo.org/record/10110796/files/figure.png" pageId="24">fig. 138</figureCitation>
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); mandibular prostheca narrow, elongate, with ciliae present only at apex (
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<figureCitation id="13162A1CD831FB3DFC406FA70AB9FADF" box="[920,1000,1268,1291]" captionStart="Figs" captionStartId="62.[298,346,1373,1394]" captionTargetBox="[280,1368,176,1352]" captionTargetId="figure-36@62.[280,1368,176,1352]" captionTargetPageId="62" captionText="Figs. 11–16. 11. Platydracus stercorarius, right mandibular prostheca, ventral view. 12. Abemus chloropterus, left mandibular prostheca, ventral view. 13. Ocypus picipennis, left mandibular prostheca, ventral view. 14. Tasgius ater, left mandibular prostheca, ventral view. 15. Staphylinus dimidiaticornis, left mandibular prostheca, ventral view. 16. Tasgius melanarius, left mandibular prostheca, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10110754" httpUri="https://zenodo.org/record/10110754/files/figure.png" pageId="24">fig. 15</figureCitation>
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); last segment of maxillary palpus fusiform, narrower and markedly shorter than segment three, asetose (
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<figureCitation id="13162A1CD831FB3DFB696E7D0C54FA91" box="[1201,1285,1326,1349]" captionStart="Figs" captionStartId="64.[298,346,1603,1624]" captionTargetBox="[289,1358,181,1581]" captionTargetId="figure-36@64.[289,1358,181,1581]" captionTargetPageId="64" captionText="Figs. 22–27. 22. Dinothenarus pubescens, last segment of maxillary palpus. 23. Ocypus sylvaticus, last segment of maxillary palpus. 24. Staphylinus dimidiaticornis, last segment of maxillary palpus. 25. Ocypus tenebricosus, last segment of maxillary palpus. 26. Staphylinus erythropterus, last segment of maxillary palpus. 27. Staphylinus erythropterus, peglike setae in glandular openings on last segment of maxillary palpus." figureDoi="http://doi.org/10.5281/zenodo.10110758" httpUri="https://zenodo.org/record/10110758/files/figure.png" pageId="24">fig. 24</figureCitation>
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); ligula emarginate; last segment of labial palpus fusiform, narrower and markedly longer than segment two, asetose (
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<figureCitation id="13162A1CD831FB3DFB8E6ED50DE5FA49" box="[1110,1204,1414,1437]" captionStart="Figs" captionStartId="83.[264,312,1571,1592]" captionTargetBox="[242,1338,176,1549]" captionTargetId="figure-34@83.[242,1338,176,1549]" captionTargetPageId="83" captionText="Figs. 136–141. 136. Platydracus stercorarius, pronotal hypomeron, ventral view. 137. Platydracus stercorarius, right coxal cavity, ventral view. 138. Staphylinus dimidiaticornis, left mandible, ventral view. 139. Staphylinus erythropterus, last segment of labial palpus. 140. Dinothenarus pubescens, posteromedial portion of head and neck with pubescence, dorsal view. 141. Dinothenarus pubescens, pronotum with pubescence." figureDoi="http://doi.org/10.5281/zenodo.10110796" httpUri="https://zenodo.org/record/10110796/files/figure.png" pageId="24">fig. 139</figureCitation>
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); anterior margin of submentum inwardly arcuate, without bead; postmandibular ridge rudimentary to very short; base of infraorbital ridge on the neck distinctly recurved toward nuchal ridge dorsally, sometimes reaching it; superior and inferior lines of pronotal hypomeron connected well before anterior angles of pronotum (
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<figureCitation id="13162A1CD831FB3DFC496D220AB4F95C" box="[913,997,1649,1672]" captionStart="Figs" captionStartId="69.[264,312,1480,1501]" captionTargetBox="[251,1328,176,1459]" captionTargetId="figure-34@69.[251,1328,176,1459]" captionTargetPageId="69" captionText="Figs. 53–58. 53. Xanthopygus analis, disc of pronotum virtually invisible in ventral view. 54. Dinothenarus pubescens, joining of superior and inferior marginal lines of pronotal hypomeron and prosternum with strongly arched sternacostal ridge (arrow), ventral view. 55. Quedius laticollis, proepimeron, ventral view. 56. Abemus chloropterus, proepimeron, ventral view. 57. Staphylinus erythropterus, proepimeron absent. 58. Ocypus ophthalmicus, medial margin of sternacostal ridge forming a prominent finlike carina." figureDoi="http://doi.org/10.5281/zenodo.10110768" httpUri="https://zenodo.org/record/10110768/files/figure.png" pageId="24">fig. 57</figureCitation>
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); epimeron of prothorax absent (
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<figureCitation id="13162A1CD831FB3DFC806DDD0AFAF971" box="[856,939,1678,1701]" captionStart="Figs" captionStartId="68.[298,346,1509,1530]" captionTargetBox="[309,1338,176,1488]" captionTargetId="figure-35@68.[309,1338,176,1488]" captionTargetPageId="68" captionText="Figs. 45–52. 45. Philonthus caeruleipennis, superior and inferior marginal lines of pronotal hypomeron not joined, ventral view. 46. Agelosus carinatus, superior and inferior marginal lines of pronotal hypomeron distinctly not joined, ventral view. 47. Ocypus weisei, superior and inferior marginal lines of pronotal hypomeron not quite joined, ventral view. 48. Ocypus tenebricosus, superior and inferior marginal lines of pronotal hypomeron not quite joined, ventral view. 49. Creophilus maxillosus, superior marginal line of pronotal hypomeron becoming obsolete near anterior angles, ventral view. 50. Staphylinus dimidiaticornis, joining of superior and inferior marginal lines of pronotal hypomeron, ventral view. 51. Miobdelus sp., joining of superior and inferior marginal lines of pronotal hypomeron, ventral view. 52. Platydracus stercorarius, superior and inferior marginal lines of pronotal hypomeron not quite joined, appearing as a grooved carina anteriorly, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10110766" httpUri="https://zenodo.org/record/10110766/files/figure.png" pageId="24">fig. 50</figureCitation>
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); prosternum with middle portion of basisternite broadly rounded or gradually sloping anteriad; serial long setae on mesosternum originating in moderately impressed punctures in subtransverse to broadly arcuate arrangement; spines on dorsal apicolateral lobe of hind coxae small, almost indistinguishable in size from other setae.
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</paragraph>
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<paragraph id="8B923699D830FB3CFED36A350836FE27" blockId="25.[238,767,182,499]" pageId="25">
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This is a new concept of this genus, restricted to the species around
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<taxonomicName id="4C2D4D1AD830FB3CFD836AD70842FE6C" authorityName="Linnaeus" authorityYear="1758" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="25" phylum="Arthropoda" rank="species" species="erythropterus">
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<emphasis id="B959EA8BD830FB3CFD836AD70842FE6C" italics="true" pageId="25">S. erythropterus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C2D4D1AD830FB3CFE886AF208B5FE6C" authorityName="Cederhjelm" authorityYear="1798" box="[336,484,417,440]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="25" phylum="Arthropoda" rank="species" species="caesareus">
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<emphasis id="B959EA8BD830FB3CFE886AF208B5FE6C" box="[336,484,417,440]" italics="true" pageId="25">S. caesareus</emphasis>
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</taxonomicName>
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. Six Palearctic and one Nearctic species are at present included in the genus.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |