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<document ID-DOI="http://doi.org/10.5281/zenodo.3793008" ID-GBIF-Dataset="6a4658b7-ce15-45fb-8985-b925e1490704" ID-GBIF-Taxon="163747379" ID-ISSN="1313–2970" ID-Zenodo-Dep="3793008" approvalRequired="27" approvalRequired_for_taxonomicNames="1" approvalRequired_for_treatments="26" checkinTime="1587952747731" checkinUser="plazi" docAuthor="Majka, Christopher, Klimaszewski, Jan &amp; Lauff, Randolph" docDate="2008" docId="03848788466E662F4BD99E21FD31D3F3" docLanguage="en" docName="ZK_article_1941.pdf.imd" docOrigin="ZooKeys 2 (2)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@2e16c743" docStyleName="zookeys.2008.journal_article" docTitle="Creophilus maxillosus subsp. villosus" docType="treatment" docVersion="9" lastPageNumber="134" masterDocId="FFBDFFF0467D663C4B559C4EFF95D647" masterDocTitle="The coastal rove beetles (Coleoptera, Staphylinidae) of Atlantic Canada: a survey and new records" masterLastPageNumber="150" masterPageNumber="115" pageId="19" pageNumber="134" updateTime="1643570856658" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The coastal rove beetles (Coleoptera, Staphylinidae) of Atlantic Canada: a survey and new records</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Majka, Christopher</mods:namePart>
<mods:affiliation>Nova Scotia Museum, Halifax, NS, Canada</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Klimaszewski, Jan</mods:namePart>
<mods:affiliation>St. Francis Xavier University, Department of Biology, Antigonish, Nova Scotia, Canada</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lauff, Randolph</mods:namePart>
<mods:affiliation>St. Francis Xavier University, Antigonish ,, Canada</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2008</mods:date>
<mods:detail type="pubDate">
<mods:number>2008-09-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>2</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>115</mods:start>
<mods:end>150</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.2.2</mods:identifier>
<mods:identifier type="GBIF-Dataset">6a4658b7-ce15-45fb-8985-b925e1490704</mods:identifier>
<mods:identifier type="ISSN">1313–2970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576408</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3793008" ID-GBIF-Taxon="163747379" ID-Zenodo-Dep="3793008" LSID="urn:lsid:plazi:treatment:03848788466E662F4BD99E21FD31D3F3" httpUri="http://treatment.plazi.org/id/03848788466E662F4BD99E21FD31D3F3" lastPageNumber="134" pageId="19" pageNumber="134">
<subSubSection pageId="19" pageNumber="134" type="nomenclature">
<paragraph blockId="19.[140,733,623,650]" box="[140,733,623,650]" pageId="19" pageNumber="134">
<heading box="[140,733,623,650]" fontSize="11" level="2" pageId="19" pageNumber="134" reason="3">
<taxonomicName authority="(Gravenhorst, 1802)" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" box="[140,733,623,650]" class="Insecta" family="Staphylinidae" genus="Creophilus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="134" phylum="Arthropoda" rank="subSpecies" species="maxillosus" subSpecies="villosus">
<emphasis bold="true" box="[140,493,623,650]" italics="true" pageId="19" pageNumber="134">Creophilus maxillosus villosus</emphasis>
(Gravenhorst, 1802)
</taxonomicName>
</heading>
</paragraph>
<paragraph blockId="19.[140,1108,694,1460]" pageId="19" pageNumber="134">
<materialsCitation ID-GBIF-Occurrence="2608811587" collectingDate="2000-08-29" collectingDateMax="2006-06-07" collectingDateMin="2000-08-29" collectionCode="CGMC, STFX" collectorName="D. S. Christie &amp; H. J. Watts" county="Albert Co." location="Antigonish Co." municipality="Mary's Point" pageId="19" pageNumber="134" specimenCount="1" stateProvince="NEW BRUNSWICK">
<emphasis bold="true" box="[140,519,694,721]" pageId="19" pageNumber="134">
<collectingRegion box="[140,385,694,721]" country="Canada" name="New Brunswick" pageId="19" pageNumber="134">NEW BRUNSWICK</collectingRegion>
:
<collectingCounty box="[393,513,694,720]" pageId="19" pageNumber="134">Albert Co.</collectingCounty>
:
</emphasis>
<collectingMunicipality box="[522,660,694,721]" pageId="19" pageNumber="134">Mary’s Point</collectingMunicipality>
,
<date box="[668,785,694,720]" pageId="19" pageNumber="134" value="2006-06-07">
<collectingDate box="[668,785,694,720]" pageId="19" pageNumber="134" value="2006-06-07">7.VI.2006</collectingDate>
</date>
,
<collectorName box="[793,938,694,721]" pageId="19" pageNumber="134">D.S.Christie</collectorName>
, on dead Minke Whale, (2,
<collectionCode box="[263,356,730,756]" collectionName="Christopher C. Majka" pageId="19" pageNumber="134">CGMC</collectionCode>
).
<emphasis bold="true" box="[373,760,729,755]" pageId="19" pageNumber="134">
<collectingRegion box="[373,563,729,755]" country="Canada" name="Nova Scotia" pageId="19" pageNumber="134">NOVA SCOTIA</collectingRegion>
:
<location LSID="urn:lsid:plazi:treatment:03848788466E662F4BD99E21FD31D3F3:8EF26045466E662F496B9E97FD67D4B4" box="[574,754,729,755]" county="Albert Co." municipality="Mary's Point" name="Antigonish Co." pageId="19" pageNumber="134" stateProvince="NEW BRUNSWICK">Antigonish Co.</location>
:
</emphasis>
Pomquet Beach,
<date box="[951,1103,730,756]" pageId="19" pageNumber="134" value="2000-08-29">
<collectingDate box="[951,1103,730,756]" pageId="19" pageNumber="134" value="2000-08-29">29.VIII.2000</collectingDate>
</date>
,
<collectorName box="[140,260,765,791]" pageId="19" pageNumber="134">H.J. Watts</collectorName>
, coastal beach, (1,
<collectionCode box="[467,538,765,791]" collectionName="Canada, Nova Scotia, Antigonish, St. Francis Xavier University" pageId="19" pageNumber="134">STFX</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608811588" collectingDate="2004-05-06" collectionCode="STFX" collectorName="M. A. Rossong" location="Pomquet Beach" pageId="19" pageNumber="134" specimenCount="1" stateProvince="New Brunswick">
<location LSID="urn:lsid:plazi:treatment:03848788466E662F4BD99E21FD31D3F3:8EF26045466E662F497B9EB3FD4BD550" box="[558,734,765,791]" name="Pomquet Beach" pageId="19" pageNumber="134" stateProvince="New Brunswick">Pomquet Beach</location>
,
<date box="[745,850,765,791]" pageId="19" pageNumber="134" value="2004-05-06">
<collectingDate box="[745,850,765,791]" pageId="19" pageNumber="134" value="2004-05-06">6.V.2004</collectingDate>
</date>
,
<collectorName box="[861,1022,765,791]" pageId="19" pageNumber="134">M.A. Rossong</collectorName>
, coastal beach, (1,
<collectionCode box="[260,331,800,826]" collectionName="Canada, Nova Scotia, Antigonish, St. Francis Xavier University" pageId="19" pageNumber="134">STFX</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608811553" box="[352,1103,800,827]" collectingDate="2001-05-16" collectionCode="STFX" collectorName="B. F. Leahey" location="Pomquet Beach" pageId="19" pageNumber="134" specimenCount="1" stateProvince="New Brunswick">
<location LSID="urn:lsid:plazi:treatment:03848788466E662F4BD99E21FD31D3F3:8EF26045466E662F4A359F6FFD86D57D" box="[352,531,800,827]" name="Pomquet Beach" pageId="19" pageNumber="134" stateProvince="New Brunswick">Pomquet Beach</location>
,
<date box="[545,663,800,826]" pageId="19" pageNumber="134" value="2001-05-16">
<collectingDate box="[545,663,800,826]" pageId="19" pageNumber="134" value="2001-05-16">16.V.2001</collectingDate>
</date>
,
<collectorName box="[677,808,800,827]" pageId="19" pageNumber="134">B.F. Leahey</collectorName>
, coastal beach, (1,
<collectionCode box="[1024,1096,800,826]" collectionName="Canada, Nova Scotia, Antigonish, St. Francis Xavier University" pageId="19" pageNumber="134">STFX</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608811582" collectingDate="2005-08-10" collectionCode="CBU" collectorName="K. R. Aikens" county="Cape Breton Co." location="Savage Cove" municipality="Scatarie Island" pageId="19" pageNumber="134" specimenCount="1" stateProvince="New Brunswick">
<emphasis bold="true" box="[140,347,835,861]" pageId="19" pageNumber="134">
<collectingCounty box="[140,341,835,861]" pageId="19" pageNumber="134">
<emphasis box="[140,200,835,861]" inLineHeading="true" pageId="19" pageNumber="134" reason="1">Cape</emphasis>
Breton Co.
</collectingCounty>
:
</emphasis>
<collectingMunicipality box="[355,518,835,861]" pageId="19" pageNumber="134">Scatarie Island</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03848788466E662F4BD99E21FD31D3F3:8EF26045466E662F49419F0DFD37D51A" box="[532,674,835,861]" county="Cape Breton Co." municipality="Scatarie Island" name="Savage Cove" pageId="19" pageNumber="134" stateProvince="New Brunswick">Savage Cove</location>
,
<date box="[687,838,835,862]" pageId="19" pageNumber="134" value="2005-08-10">
<collectingDate box="[687,838,835,862]" pageId="19" pageNumber="134" value="2005-08-10">10.VIII.2005</collectingDate>
</date>
,
<collectorName box="[851,990,835,862]" pageId="19" pageNumber="134">K.R. Aikens</collectorName>
, sea beach under wrack, (1,
<collectionCode box="[328,390,871,897]" collectionName="Canada, Nova Scotia, Sydney, Cape Breton University" country="Canada" httpUri="http://grbio.org/cool/yp8i-58ys" name="Cape Breton University" pageId="19" pageNumber="134">CBU</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608811565" box="[409,906,870,897]" collectingDate="1995-08-03" collectionCode="CBU" collectorName="A. Brown" location="South Bar" pageId="19" pageNumber="134" specimenCount="1" stateProvince="New Brunswick">
<location LSID="urn:lsid:plazi:treatment:03848788466E662F4BD99E21FD31D3F3:8EF26045466E662F4ACC9F28FD9CD5C6" box="[409,521,870,897]" name="South Bar" pageId="19" pageNumber="134" stateProvince="New Brunswick">South Bar</location>
,
<date box="[531,668,870,897]" pageId="19" pageNumber="134" value="1995-08-03">
<collectingDate box="[531,668,870,897]" pageId="19" pageNumber="134" value="1995-08-03">3.VIII.1995</collectingDate>
</date>
,
<collectorName box="[679,788,871,897]" pageId="19" pageNumber="134">A. Brown</collectorName>
, (1,
<collectionCode box="[836,898,871,897]" collectionName="Canada, Nova Scotia, Sydney, Cape Breton University" country="Canada" httpUri="http://grbio.org/cool/yp8i-58ys" name="Cape Breton University" pageId="19" pageNumber="134">CBU</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608811589" collectingDate="28. VII. 2003, 15. IX. 2004" collectingMethod="pitfall trap" collectionCode="JOC" collectorName="J. Ogden &amp; K. Goodwin" county="Digby Co." location="Pond Cove" municipality="Brier Island" pageId="19" pageNumber="134" specimenCount="1" stateProvince="New Brunswick">
<emphasis bold="true" box="[917,1046,870,896]" pageId="19" pageNumber="134">
<collectingCounty box="[917,1041,870,896]" pageId="19" pageNumber="134">Digby Co.</collectingCounty>
:
</emphasis>
<collectingMunicipality pageId="19" pageNumber="134">Brier Island</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03848788466E662F4BD99E21FD31D3F3:8EF26045466E662F4B8A9FC4FECAD5E3" box="[223,351,906,932]" county="Digby Co." municipality="Brier Island" name="Pond Cove" pageId="19" pageNumber="134" stateProvince="New Brunswick">Pond Cove</location>
,
<collectingDate box="[365,651,905,932]" isEnumeration="true" pageId="19" pageNumber="134">
<date box="[365,507,906,932]" pageId="19" pageNumber="134" value="2003-07-28">28.VII.2003</date>
,
<date box="[521,651,905,932]" pageId="19" pageNumber="134" value="2004-09-15">15.IX.2004</date>
</collectingDate>
,
<collectorName box="[666,769,906,932]" pageId="19" pageNumber="134">J. Ogden</collectorName>
&amp;
<collectorName box="[812,957,906,932]" pageId="19" pageNumber="134">K. Goodwin</collectorName>
, rocky shore,
<collectingMethod box="[140,259,941,967]" pageId="19" pageNumber="134">pitfall trap</collectingMethod>
, (3,
<collectionCode box="[310,366,941,967]" collectionName="Jeffrey Ogden" pageId="19" pageNumber="134">JOC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608811575" collectingDate="1993-07-20" collectionCode="JCC" collectorName="J. &amp; T. Cook" location="Mavilette Beach" pageId="19" pageNumber="134" specimenCount="1" stateProvince="New Brunswick">
<location LSID="urn:lsid:plazi:treatment:03848788466E662F4BD99E21FD31D3F3:8EF26045466E662F4AD69FE3FDACD580" box="[387,569,941,967]" name="Mavilette Beach" pageId="19" pageNumber="134" stateProvince="New Brunswick">Mavilette Beach</location>
,
<date box="[581,723,941,967]" pageId="19" pageNumber="134" value="1993-07-20">
<collectingDate box="[581,723,941,967]" pageId="19" pageNumber="134" value="1993-07-20">20.VII.1993</collectingDate>
</date>
,
<collectorName box="[736,887,941,967]" pageId="19" pageNumber="134">J. &amp; T. Cook</collectorName>
, beach dune, under driftwood, (1,
<collectionCode box="[308,362,976,1002]" collectionName="JCC" pageId="19" pageNumber="134">JCC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608811579" collectingDate="20. VI. 1976, 9. VI. 1977, 10. VI. 1977, 12. VI. 1977, 12. IX. 1977" collectionCode="NSMC" collectorName="B. Wright" county="Halifax Co." location="Sable Island" pageId="19" pageNumber="134" specimenCount="1" stateProvince="New Brunswick">
<emphasis bold="true" box="[384,530,976,1002]" pageId="19" pageNumber="134">
<collectingCounty box="[384,524,976,1002]" pageId="19" pageNumber="134">Halifax Co.</collectingCounty>
:
</emphasis>
<location LSID="urn:lsid:plazi:treatment:03848788466E662F4BD99E21FD31D3F3:8EF26045466E662F49499F9EFD30D5AD" box="[540,677,976,1002]" county="Halifax Co." name="Sable Island" pageId="19" pageNumber="134" stateProvince="New Brunswick">Sable Island</location>
,
<collectingDate isEnumeration="true" pageId="19" pageNumber="134">
<date box="[692,824,976,1002]" pageId="19" pageNumber="134" value="1976-06-20">20.VI.1976</date>
,
<date box="[838,956,976,1002]" pageId="19" pageNumber="134" value="1977-06-09">9.VI.1977</date>
,
<date box="[971,1103,976,1002]" pageId="19" pageNumber="134" value="1977-06-10">10.VI.1977</date>
,
<date box="[140,272,1011,1037]" pageId="19" pageNumber="134" value="1977-06-12">12.VI.1977</date>
,
<date box="[285,416,1011,1037]" pageId="19" pageNumber="134" value="1977-09-12">12.IX.1977</date>
</collectingDate>
,
<collectorName box="[429,542,1011,1038]" pageId="19" pageNumber="134">B. Wright</collectorName>
, (17,
<collectionCode box="[609,696,1011,1037]" collectionName="USA, Nevada, Carson City, Nevada State Museum" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34980" lsid="urn:lsid:biocol.org:col:34980" name="Nova Scotia Museum" pageId="19" pageNumber="134">NSMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608811564" collectingDate="2007-08-18" collectionCode="NSMC" collectorName="R. Gorham" county="Shelburne Co." location="Daniel's Head Beach" municipality="Cape Sable Island" pageId="19" pageNumber="134" specimenCount="1" stateProvince="New Brunswick">
<emphasis bold="true" box="[716,893,1011,1037]" pageId="19" pageNumber="134">
<collectingCounty box="[716,887,1011,1037]" pageId="19" pageNumber="134">Shelburne Co.</collectingCounty>
:
</emphasis>
<collectingMunicipality box="[901,1104,1011,1037]" pageId="19" pageNumber="134">Cape Sable Island</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03848788466E662F4BD99E21FD31D3F3:8EF26045466E662F4BD99858FEE6D277" box="[140,371,1046,1072]" county="Shelburne Co." municipality="Cape Sable Island" name="Daniel's Head Beach" pageId="19" pageNumber="134" stateProvince="New Brunswick">Daniel’s Head Beach</location>
,
<date box="[382,534,1047,1073]" pageId="19" pageNumber="134" value="2007-08-18">
<collectingDate box="[382,534,1047,1073]" pageId="19" pageNumber="134" value="2007-08-18">18.VIII.2007</collectingDate>
</date>
,
<collectorName box="[545,675,1046,1073]" pageId="19" pageNumber="134">R. Gorham</collectorName>
, on seal carcass, (1,
<collectionCode box="[893,981,1047,1073]" collectionName="USA, Nevada, Carson City, Nevada State Museum" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34980" lsid="urn:lsid:biocol.org:col:34980" name="Nova Scotia Museum" pageId="19" pageNumber="134">NSMC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="134" type="description">
<paragraph blockId="19.[140,1108,694,1460]" pageId="19" pageNumber="134">
This cosmopolitan species has been known in the
<collectingRegion box="[748,855,1082,1108]" country="Russia" name="Primorskiy" pageId="19" pageNumber="134">Maritime</collectingRegion>
Provinces since
<date pageId="19" pageNumber="134" value="1827" valueMax="1828">1827- 28</date>
from collections made in
<collectingRegion box="[484,624,1117,1143]" country="Canada" name="Nova Scotia" pageId="19" pageNumber="134">Nova Scotia</collectingRegion>
by Captain Basil Hall (
<bibRefCitation author="Kirby" box="[909,1046,1117,1143]" firstAuthor="Kirby" pageId="19" pageNumber="134" refId="ref16926" refString="Kirby WF (1837) The insects. In: Richardson J (Ed) Fauna boreali-Americana or the Zoology of the northern parts of British America, containing descriptions of the objects of natural history collected on the late northern land expeditions, under the command of Captain Sir John Franklin, RN. Fletcher, Norwich, xxxix + 325 pp." type="book" year="1837">Kirby 1837</bibRefCitation>
) and
<bibRefCitation author="Prevost and Bain" box="[140,416,1152,1178]" firstAuthor="Prevost" pageId="19" pageNumber="134" pagination="205 - 216" refId="ref17882" refString="Prevost M-A, Bain A (2007) L'implantation d'une colonie terre-neuvienne au XVIIe siecle: l'apport des analyses archeobotanique et archeoentomologique. In: Bain A, Chabot J, Moussette M (Eds) La mesure du passe: contributions a la recherche en archeometrie (2000 - 2006), British Archaeological Reports, International Series 1700. Oxford Archaeopress, Oxford, 205 - 216." type="book chapter" year="2007">Prévost and Bain (2007)</bibRefCitation>
found it in deposits dated
<quantity box="[722,811,1152,1178]" metricMagnitude="1" metricUnit="m" metricValue="4.1148" pageId="19" pageNumber="134" unit="in" value="1620.0">1620 in</quantity>
Newfoundland. Its distribution in the
<collectingRegion box="[295,402,1187,1213]" country="Russia" name="Primorskiy" pageId="19" pageNumber="134">Maritime</collectingRegion>
Provinces is shown in
<figureCitation box="[657,739,1187,1214]" captionStart="Fig" captionStartId="18.[139,172,1590,1612]" captionTargetBox="[139,1108,841,1571]" captionTargetId="figure@18.[139,1108,841,1569]" captionTargetPageId="18" captionText="Fig. 15. The distribution of Gyrohypnus angustatus, Creophilus maxillosus villosus, Cafius bistriatus, and Gabrius astutoides in Atlantic Canada. Note: Two sites from central Labrador (Goose Bay and Cartwright) are not shown on the map." figureDoi="http://doi.org/10.5281/zenodo.3767195" httpUri="https://zenodo.org/record/3767195/files/figure.png" pageId="19" pageNumber="134">Fig. 15</figureCitation>
. As the above records indicate, it is frequently encountered in coastal situations (where adults and larvae feed on maggots found on various kinds of carrion and decaying matter), however, it also occurs in many kinds of natural habitats and in synanthropic situations (
<bibRefCitation author="Newton" box="[865,1095,1293,1319]" etAl="et al." firstAuthor="Newton" pageId="19" pageNumber="134" pagination="272 - 418" refId="ref17697" refString="Newton AF, Thayer MK, Ashe JS, Chandler DS (2000) Staphylinidae Latreille, 1802. In: Arnett RH, Jr, Thomas, MC (Eds) American Beetles, Volume 1: Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Raton, 272 - 418." type="book chapter" year="2000">Newton et al. 2000</bibRefCitation>
).
<bibRefCitation author="Newton" box="[140,381,1328,1355]" etAl="et al." firstAuthor="Newton" pageId="19" pageNumber="134" pagination="272 - 418" refId="ref17697" refString="Newton AF, Thayer MK, Ashe JS, Chandler DS (2000) Staphylinidae Latreille, 1802. In: Arnett RH, Jr, Thomas, MC (Eds) American Beetles, Volume 1: Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Raton, 272 - 418." type="book chapter" year="2000">Newton et al. (2000)</bibRefCitation>
pointed out that the North American specimens of the subspecies
<taxonomicName baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" box="[191,427,1363,1390]" class="Insecta" family="Staphylinidae" genus="Creophilus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="134" phylum="Arthropoda" rank="subSpecies" species="maxillosus" subSpecies="villosus">
<emphasis box="[191,427,1363,1390]" italics="true" pageId="19" pageNumber="134">C. maxillosus villosus</emphasis>
</taxonomicName>
, are distinguishable from the Palaearctic
<emphasis box="[912,1100,1363,1390]" italics="true" pageId="19" pageNumber="134">
C. m.
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[989,1100,1363,1389]" class="Insecta" family="Staphylinidae" genus="Creophilus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="134" phylum="Arthropoda" rank="species" species="maxillosus">maxillosus</taxonomicName>
</emphasis>
.
<emphasis box="[140,375,1398,1424]" italics="true" pageId="19" pageNumber="134">
<taxonomicName authorityName="Leach" authorityYear="1819" box="[140,255,1398,1424]" class="Insecta" family="Staphylinidae" genus="Creophilus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="134" phylum="Arthropoda" rank="genus">Creophilus</taxonomicName>
m. villosus
</emphasis>
has a broad pre-human North American distribution, and hence can be considered a native, Nearctic subspecies.
</paragraph>
</subSubSection>
</treatment>
</document>
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