treatments-xml/data/6C/35/51/6C35515BFF832F62B991F901970CFDCC.xml
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<document id="C5541E8010455F3C3E924AC0E6B50C1A" ID-CLB-Dataset="29835" ID-DOI="10.11646/zootaxa.4438.2.11" ID-GBIF-Dataset="27cbe469-a6fd-43c1-a663-4367786bf528" ID-ISSN="1175-5326" ID-Zenodo-Dep="1294883" ID-ZooBank="7A2A9C66-5721-4D95-8BAD-57407773FDDD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1529569804037" checkinUser="plazi" docAuthor="Solís-Marín, Francisco Alonso, Arriaga-Ochoa, Julio Adrian, Villa, Cristian Moises Galvan &amp; Laguarda-Figueras, Alfredo" docDate="2018" docId="6C35515BFF832F62B991F901970CFDCC" docLanguage="en" docName="zootaxa.4438.2.11.pdf.imf" docOrigin="Zootaxa 4438 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lissothuria imbricata Solís-Marín &amp; Arriaga-Ochoa &amp; Villa &amp; Laguarda-Figueras 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="379" masterDocId="900C2923FF822F64B906FFAE933AFFD4" masterDocTitle="Lissothuria imbricata sp. nov., a new species of sea cucumber (Echinodermata: Holothuroidea) from the central-eastern Pacific Ocean" masterLastPageNumber="380" masterPageNumber="373" pageNumber="374" updateTime="1698550277557" updateUser="ExternalLinkService">
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<mods:title id="64CD2EAC131F12E0A53C5E59FB36026B">Lissothuria imbricata sp. nov., a new species of sea cucumber (Echinodermata: Holothuroidea) from the central-eastern Pacific Ocean</mods:title>
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<mods:namePart id="0D7819559FBB54C4EC09FDD781D4ABAE">Solís-Marín, Francisco Alonso</mods:namePart>
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<mods:namePart id="38A1E27EDC09B96FADE385BA2FDD2081">Arriaga-Ochoa, Julio Adrian</mods:namePart>
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<mods:namePart id="5189AD246035F1F231C5E44E0756A78B">Villa, Cristian Moises Galvan</mods:namePart>
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<mods:namePart id="70F4F4387FA5B1E45616BAC7BDD37777">Laguarda-Figueras, Alfredo</mods:namePart>
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<mods:date id="4EBA3F03C8845486D8F8385D42A9B039">2018</mods:date>
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<mods:number id="706025F1E6F088AE40E8BE4732B79120">2018-06-21</mods:number>
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<treatment id="6C35515BFF832F62B991F901970CFDCC" ID-DOI="http://doi.org/10.5281/zenodo.5976869" ID-GBIF-Taxon="146016156" ID-Zenodo-Dep="5976869" LSID="urn:lsid:plazi:treatment:6C35515BFF832F62B991F901970CFDCC" httpUri="http://treatment.plazi.org/id/6C35515BFF832F62B991F901970CFDCC" lastPageId="6" lastPageNumber="379" pageId="1" pageNumber="374">
<subSubSection id="AC86B3C6FF832F65B991F901922BF93F" pageId="1" pageNumber="374" type="nomenclature">
<paragraph id="E423E04DFF832F65B991F901913BF91E" blockId="1.[151,513,1711,1771]" box="[151,513,1711,1738]" pageId="1" pageNumber="374">
<heading id="BF6B5721FF832F65B991F901913BF91E" bold="true" box="[151,513,1711,1738]" fontSize="11" level="1" pageId="1" pageNumber="374" reason="1">
<emphasis id="D6E83C5FFF832F65B991F901913BF91E" bold="true" box="[151,513,1711,1738]" pageId="1" pageNumber="374">
<taxonomicName id="239C9BCEFF832F65B991F90192A6F91D" ID-CoL="3VGGD" authority="Solís-Marín &amp; Arriaga-Ochoa &amp; Villa &amp; Laguarda-Figueras, 2018" authorityName="Solís-Marín &amp; Arriaga-Ochoa &amp; Villa &amp; Laguarda-Figueras" authorityYear="2018" box="[151,412,1711,1737]" class="Holothuroidea" family="Psolidae" genus="Lissothuria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="1" pageNumber="374" phylum="Echinodermata" rank="species" species="imbricata" status="sp. nov.">
<emphasis id="D6E83C5FFF832F65B991F90192A6F91D" bold="true" box="[151,412,1711,1737]" italics="true" pageId="1" pageNumber="374">Lissothuria imbricata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CDDB8124FF832F65B8A5F91E913BF91E" box="[419,513,1712,1738]" pageId="1" pageNumber="374" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="E423E04DFF832F65B991F97C922BF93F" blockId="1.[151,513,1711,1771]" box="[151,273,1746,1771]" pageId="1" pageNumber="374">
(
<figureCitation id="7CA7FCC8FF832F65B999F97C9233F93F" box="[159,265,1746,1771]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1773,1795]" captionStartId-1="4.[151,250,1781,1803]" captionStartId-2="5.[151,250,1872,1894]" captionTargetBox-0="[151,1436,303,1748]" captionTargetBox-1="[284,1303,315,1755]" captionTargetBox-2="[167,1411,193,1833]" captionTargetId-0="figure@2.[151,1436,303,1748]" captionTargetId-1="figure@4.[284,1303,315,1755]" captionTargetId-2="figure@5.[167,1411,193,1833]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Lissothuria imbricata sp. nov., holotype ICML-UNAM 18000, A, Dorsal view; B, Ventral view. Paratype ICML- UNAM 12135; C, In situ picture showing habitat preferences; D, Live specimen, lateral view. All scale bars: 5 mm." captionText-1="FIGURE 2. Lissothuria imbricata sp. nov., holotype ICML-UNAM 18000, A, Dorsal perforated scale for passage of tube feet, scale bar: 100 µm; B, Detail of perforations in the dorsal scale; CE, Massive towers from the dorsal side, with slightly spiny apex, scale bar: 50 µm; FI, Baskets from the dorsal side, scale bar: 40 µm; JK, Dorsal tube feet vestigial end plates, scale bar: 50 µm." captionText-2="FIGURE 3. Lissothuria imbricata sp. nov., holotype ICML-UNAM 18000, AC, Ventral ossicles, knobbed buttons, scale bar: 50 µm; DF, Ventral ossicles, various shapes of perforated plates with smooth or knobbed margin, scale bar: 50 µm; GI, Ventral ossicles, small shallow baskets, with knobbed edges, scale bar: 30 µm; JK, Tentacles ossicles, small rosettes, scale bar: 10 µm; L, Tentacles ossicles, flat, curved and perforated rods, scale bar: 50 µm; and M, Tentacles ossicles, perforated plates, scale bar: 100 µm." httpUri-0="https://zenodo.org/record/1294885/files/figure.png" httpUri-1="https://zenodo.org/record/1294887/files/figure.png" httpUri-2="https://zenodo.org/record/1294889/files/figure.png" pageId="1" pageNumber="374">Figs. 13</figureCitation>
)
</paragraph>
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<subSubSection id="AC86B3C6FF832F65B991F8B49081F83B" pageId="1" pageNumber="374" type="description">
<paragraph id="E423E04DFF832F65B991F8B49630F88E" blockId="1.[151,1436,1818,2032]" pageId="1" pageNumber="374">
<materialsCitation id="54F4EA10FF832F65B991F8B49630F88E" ID-GBIF-Occurrence="1851020277" collectingDate="2014-11-14" collectorName="La Huerta" country="Mexico" elevation="5" latitude="19.545776" location="Isla Cocinas" longLatPrecision="1" longitude="-105.10953" municipality="Chamela Bay" pageId="1" pageNumber="374" specimenCount="2" stateProvince="Jalisco" typeStatus="holotype">
<emphasis id="D6E83C5FFF832F65B991F8B493D9F8E7" bold="true" box="[151,227,1818,1843]" pageId="1" pageNumber="374">
<typeStatus id="3B275EEFFF832F65B991F8B493E5F8E7" box="[151,223,1818,1843]" pageId="1" pageNumber="374">Types</typeStatus>
.
</emphasis>
<typeStatus id="3B275EEFFF832F65B9E9F8B4926DF8E7" box="[239,343,1818,1843]" pageId="1" pageNumber="374" type="holotype">Holotype</typeStatus>
ICML-UNAM 18000,
<quantity id="23644DA8FF832F65BB6DF8B491D8F8E6" box="[619,738,1818,1843]" metricMagnitude="-2" metricUnit="m" metricValue="1.646" pageId="1" pageNumber="374" unit="mm" value="16.46">16.46 mm</quantity>
Total Length (TL),
<location id="E143B696FF832F65BAD5F8B4975EF8E7" LSID="urn:lsid:plazi:treatment:6C35515BFF832F62B991F901970CFDCC:E143B696FF832F65BAD5F8B4975EF8E7" box="[979,1124,1818,1843]" country="Mexico" latitude="19.545776" longLatPrecision="1" longitude="-105.10953" municipality="Chamela Bay" name="Isla Cocinas" pageId="1" pageNumber="374" stateProvince="Jalisco">Isla Cocinas</location>
,
<collectingMunicipality id="04477A37FF832F65BD75F8B4962AF8E7" box="[1139,1296,1818,1843]" pageId="1" pageNumber="374">Chamela Bay</collectingMunicipality>
,
<collectorName id="4969859BFF832F65BC26F8B596A2F8E6" box="[1312,1432,1819,1842]" pageId="1" pageNumber="374">La Huerta</collectorName>
,
<collectingRegion id="26582EAFFF832F65B991F8EF93DCF88E" box="[151,230,1857,1882]" country="Mexico" name="Jalisco" pageId="1" pageNumber="374">Jalisco</collectingRegion>
,
<collectingCountry id="9C8BA0DDFF832F65B9F7F8EF9273F88E" box="[241,329,1857,1882]" name="Mexico" pageId="1" pageNumber="374">Mexico</collectingCountry>
, Pacific Ocean
<geoCoordinate id="81A8868AFF832F65B8FEF8EC9190F88D" box="[504,682,1857,1882]" direction="north" orientation="latitude" pageId="1" pageNumber="374" precision="1" value="19.545776">19° 32 44.8″ N</geoCoordinate>
,
<geoCoordinate id="81A8868AFF832F65BBB3F8EF9043F88D" box="[693,889,1857,1882]" direction="west" orientation="longitude" pageId="1" pageNumber="374" precision="1" value="-105.10953">105° 06 34.3″ W</geoCoordinate>
,
<quantity id="23644DA8FF832F65BA82F8EF90F1F88D" box="[900,971,1857,1882]" metricMagnitude="0" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="1" pageNumber="374" unit="m" value="4.5" valueMax="6.0" valueMin="3.0">
<elevation id="6FB1077EFF832F65BA82F8EF90F1F88D" box="[900,971,1857,1882]" metricMagnitude="0" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="1" pageNumber="374" unit="m" value="4.5" valueMax="6.0" valueMin="3.0">36 m</elevation>
</quantity>
depth,
<date id="9022C68DFF832F65BD18F8EF963CF88E" box="[1054,1286,1854,1882]" pageId="1" pageNumber="374" value="2014-11-14">
<collectingDate id="80663F65FF832F65BD18F8EF963CF88E" box="[1054,1286,1854,1882]" pageId="1" pageNumber="374" value="2014-11-14">November 14th 2014</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="E423E04DFF832F65B9C1F8C89081F83B" blockId="1.[151,1436,1818,2032]" pageId="1" pageNumber="374">
<typeStatus id="3B275EEFFF832F65B9C1F8C89203F8AA" box="[199,313,1894,1918]" pageId="1" pageNumber="374" type="paratype">Paratypes</typeStatus>
: ICML-UNAM 12135,
<specimenCount id="F29A2BC4FF832F65BB44F8C891E6F8AA" box="[578,732,1893,1918]" pageId="1" pageNumber="374" type="generic">13 specimens</specimenCount>
, same data as the
<typeStatus id="3B275EEFFF832F65BAA2F8CB9733F8AA" box="[932,1033,1893,1918]" pageId="1" pageNumber="374" type="holotype">holotype</typeStatus>
; LEMA-CEQ627,
<specimenCount id="F29A2BC4FF832F65BDDAF8C89653F8AA" box="[1244,1385,1893,1918]" pageId="1" pageNumber="374" type="generic">4 specimens</specimenCount>
,
<collectingCountry id="9C8BA0DDFF832F65BC72F8CB96A6F8AA" box="[1396,1436,1893,1918]" name="Iceland" pageId="1" pageNumber="374">Isla</collectingCountry>
Cocinas, Chamela Bay, La Huerta,
<collectingRegion id="26582EAFFF832F65BB18F8279157F876" box="[542,621,1929,1954]" country="Mexico" name="Jalisco" pageId="1" pageNumber="374">Jalisco</collectingRegion>
,
<collectingCountry id="9C8BA0DDFF832F65BB7EF82791EAF876" box="[632,720,1929,1954]" name="Mexico" pageId="1" pageNumber="374">
<collectingRegion id="26582EAFFF832F65BB7EF82791EAF876" box="[632,720,1929,1954]" country="Mexico" name="Mexico" pageId="1" pageNumber="374">Mexico</collectingRegion>
</collectingCountry>
, Pacific Ocean,
<geoCoordinate id="81A8868AFF832F65BA82F8249778F875" box="[900,1090,1929,1954]" direction="north" orientation="latitude" pageId="1" pageNumber="374" precision="1" value="19.54603">19° 32 45.71″ N</geoCoordinate>
,
<geoCoordinate id="81A8868AFF832F65BD4AF8279626F875" box="[1100,1308,1929,1954]" direction="west" orientation="longitude" pageId="1" pageNumber="374" precision="1" value="-105.107704">105° 06 27.73″ W</geoCoordinate>
,
<quantity id="23644DA8FF832F65BC20F824966BF875" box="[1318,1361,1930,1953]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="374" unit="m" value="7.0">7 m</quantity>
depth,
<date id="9022C68DFF832F65B991F81F9204F81C" box="[151,318,1965,1993]" pageId="1" pageNumber="374" value="2013-06-25">June 25th 2013</date>
; LEMA-CEQ632,
<specimenCount id="F29A2BC4FF832F65BB10F81F9199F81D" box="[534,675,1968,1993]" pageId="1" pageNumber="374" type="generic">7 specimens</specimenCount>
,
<collectingCountry id="9C8BA0DDFF832F65BBB6F81E91E2F81D" box="[688,728,1968,1993]" name="Iceland" pageId="1" pageNumber="374">Isla</collectingCountry>
<collectingCountry id="9C8BA0DDFF832F65BBE7F81E9062F81D" box="[737,856,1968,1993]" name="Saint Pierre and Miquelon" pageId="1" pageNumber="374">San Pedro</collectingCountry>
, Chamela Bay, La Huerta,
<collectingRegion id="26582EAFFF832F65BD88F81E97E7F81D" box="[1166,1245,1968,1993]" country="Mexico" name="Jalisco" pageId="1" pageNumber="374">Jalisco</collectingRegion>
,
<collectingCountry id="9C8BA0DDFF832F65BDEFF81E967BF81D" box="[1257,1345,1968,1993]" name="Mexico" pageId="1" pageNumber="374">
<collectingRegion id="26582EAFFF832F65BDEFF81E967BF81D" box="[1257,1345,1968,1993]" country="Mexico" name="Mexico" pageId="1" pageNumber="374">Mexico</collectingRegion>
</collectingCountry>
, Pacific Ocean,
<geoCoordinate id="81A8868AFF832F65B9EAF8799296F83B" box="[236,428,2007,2032]" direction="north" orientation="latitude" pageId="1" pageNumber="374" precision="1" value="19.533613">19° 32 01.01″ N</geoCoordinate>
,
<geoCoordinate id="81A8868AFF832F65B8B1F87991B0F83B" box="[439,650,2007,2032]" direction="west" orientation="longitude" pageId="1" pageNumber="374" precision="1" value="-105.088326">105° 05 17.97″ W</geoCoordinate>
,
<quantity id="23644DA8FF832F65BB92F8799185F83B" box="[660,703,2007,2032]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="374" unit="m" value="5.0">5 m</quantity>
depth,
<date id="9022C68DFF832F65BA14F8769082F83B" box="[786,952,2004,2032]" pageId="1" pageNumber="374" value="2013-06-26">June 26th 2013</date>
.
</paragraph>
</subSubSection>
<subSubSection id="AC86B3C6FF802F66B9C1FF399235FF2C" pageId="2" pageNumber="375" type="materials_examined">
<paragraph id="E423E04DFF802F66B9C1FF399235FF2C" blockId="2.[151,1436,151,248]" pageId="2" pageNumber="375">
<emphasis id="D6E83C5FFF802F66B9C1FF39925FFF64" bold="true" box="[199,357,151,176]" pageId="2" pageNumber="375">
<typeStatus id="3B275EEFFF802F66B9C1FF399238FF64" box="[199,258,151,176]" pageId="2" pageNumber="375">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="9C8BA0DDFF802F66B86AFF3992AFFF64" box="[364,405,151,176]" name="Iceland" pageId="2" pageNumber="375">Isla</collectingCountry>
Cocinas, Chamela Bay,
<collectingRegion id="26582EAFFF802F66BBA1FF3991CCFF64" box="[679,758,151,176]" country="Mexico" name="Jalisco" pageId="2" pageNumber="375">Jalisco</collectingRegion>
, Pacific Ocean,
<collectingCountry id="9C8BA0DDFF802F66BAAAFF39973BFF64" box="[940,1025,151,176]" name="Mexico" pageId="2" pageNumber="375">
<collectingRegion id="26582EAFFF802F66BAAAFF39973BFF64" box="[940,1025,151,176]" country="Mexico" name="Mexico" pageId="2" pageNumber="375">Mexico</collectingRegion>
</collectingCountry>
<geoCoordinate id="81A8868AFF802F66BD0FFF369781FF7B" box="[1033,1211,151,176]" direction="north" orientation="latitude" pageId="2" pageNumber="375" precision="1" value="19.545776">19° 32 44.8″ N</geoCoordinate>
, 105° 06 34.3″.
<emphasis id="D6E83C5FFF802F66B9C1FF129292FF01" bold="true" box="[199,424,188,213]" pageId="2" pageNumber="375">Non-type material.</emphasis>
<specimenCount id="F29A2BC4FF802F66B8A9FF12916FFF01" box="[431,597,188,213]" pageId="2" pageNumber="375" type="generic">153 specimens</specimenCount>
from Chamela Bay, La Huerta,
<collectingRegion id="26582EAFFF802F66BABCFF129733FF01" box="[954,1033,188,213]" country="Mexico" name="Jalisco" pageId="2" pageNumber="375">Jalisco</collectingRegion>
,
<collectingCountry id="9C8BA0DDFF802F66BD12FF129756FF01" box="[1044,1132,188,213]" name="Mexico" pageId="2" pageNumber="375">
<collectingRegion id="26582EAFFF802F66BD12FF129756FF01" box="[1044,1132,188,213]" country="Mexico" name="Mexico" pageId="2" pageNumber="375">Mexico</collectingRegion>
</collectingCountry>
, Pacific Ocean. See table 1 for details.
</paragraph>
</subSubSection>
<caption id="B0E3B0C5FF802F66B991F9439611F8F6" httpUri="https://zenodo.org/record/1294885/files/figure.png" pageId="2" pageNumber="375" startId="2.[151,250,1773,1795]" targetBox="[151,1436,303,1748]" targetPageId="2">
<paragraph id="E423E04DFF802F66B991F9439611F8F6" blockId="2.[151,1436,1773,1826]" pageId="2" pageNumber="375">
<emphasis id="D6E83C5FFF802F66B991F9439229F8D6" bold="true" box="[151,275,1773,1795]" pageId="2" pageNumber="375">FIGURE 1.</emphasis>
<taxonomicName id="239C9BCEFF802F66B81CF94392CDF8D7" authority="Solís-Marín &amp; Arriaga-Ochoa &amp; Villa &amp; Laguarda-Figueras, 2018" authorityName="Solís-Marín &amp; Arriaga-Ochoa &amp; Villa &amp; Laguarda-Figueras" authorityYear="2018" box="[282,503,1773,1795]" class="Holothuroidea" family="Psolidae" genus="Lissothuria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="2" pageNumber="375" phylum="Echinodermata" rank="species" species="imbricata" status="sp. nov.">
<emphasis id="D6E83C5FFF802F66B81CF94392CDF8D7" box="[282,503,1773,1795]" italics="true" pageId="2" pageNumber="375">Lissothuria imbricata</emphasis>
</taxonomicName>
<emphasis id="D6E83C5FFF802F66B8FBF9439174F8D6" bold="true" box="[509,590,1773,1794]" pageId="2" pageNumber="375">
<taxonomicNameLabel id="CDDB8124FF802F66B8FBF9439174F8D6" box="[509,590,1773,1794]" pageId="2" pageNumber="375" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype ICML-UNAM 18000, A, Dorsal view; B, Ventral view. Paratype ICML- UNAM 12135; C,
<emphasis id="D6E83C5FFF802F66B852F8A392AFF8F6" box="[340,405,1805,1826]" italics="true" pageId="2" pageNumber="375">In situ</emphasis>
picture showing habitat preferences; D, Live specimen, lateral view. All scale bars: 5 mm.
</paragraph>
</caption>
<subSubSection id="AC86B3C6FF802F67B9C1F8E797D3FEC8" lastPageId="3" lastPageNumber="376" pageId="2" pageNumber="375" type="diagnosis">
<paragraph id="E423E04DFF802F67B9C1F8E797D3FEC8" blockId="2.[151,1436,1865,2034]" lastBlockId="3.[151,1436,151,285]" lastPageId="3" lastPageNumber="376" pageId="2" pageNumber="375">
<emphasis id="D6E83C5FFF802F66B9C1F8E7927BF8B6" bold="true" box="[199,321,1865,1890]" pageId="2" pageNumber="375">Diagnosis.</emphasis>
Body wide almost the half of the body length. Dorsal scale covering complete, with 11 to 16 scales between the mouth and anus, 20 pointed narrow valves surrounding the mouth; the anus is surrounded by a varying number of rounded valves. Ten short tentacles, the two most ventral tentacles are smaller than the others. Sole elongated, fringed by an outer peripheral single series of smaller tube feet; inner peripheral series with three rows of closely crowded tube feet, which form a continuous margin, mid-ventral radius with a double row of tube feet, becoming crowded anterior and posteriorly. Sole leathery, translucent. Ossicles of dorsal surface are perforated plates, knobbed baskets; each dorsal scale covered by 720 short, wide, massive towers with a slightly spiny apex, giving the dorsal area a granular appearance. Ossicles of sole are perforated plates, which are smooth or knobbed; baskets with knobbed margin. Tentacles with irregular, perforated, flattened rods, and few rosettes.
</paragraph>
</subSubSection>
<caption id="B0E3B0C5FF812F67B991FEE8927BFEA9" pageId="3" pageNumber="376">
<paragraph id="E423E04DFF812F67B991FEE8927BFEA9" blockId="3.[151,1436,325,381]" pageId="3" pageNumber="376">
<emphasis id="D6E83C5FFF812F67B991FEE89234FE89" bold="true" box="[151,270,326,349]" pageId="3" pageNumber="376">TABLE 1.</emphasis>
<taxonomicName id="239C9BCEFF812F67B81FFEEB913CFE89" authority="Solís-Marín &amp; Arriaga-Ochoa &amp; Villa &amp; Laguarda-Figueras, 2018" authorityName="Solís-Marín &amp; Arriaga-Ochoa &amp; Villa &amp; Laguarda-Figueras" authorityYear="2018" box="[281,518,325,349]" class="Holothuroidea" family="Psolidae" genus="Lissothuria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="3" pageNumber="376" phylum="Echinodermata" rank="species" species="imbricata" status="sp. nov.">
<emphasis id="D6E83C5FFF812F67B81FFEEB913CFE89" box="[281,518,325,349]" italics="true" pageId="3" pageNumber="376">Lissothuria imbricata</emphasis>
</taxonomicName>
<emphasis id="D6E83C5FFF812F67BB16FEE89150FE89" bold="true" box="[528,618,326,349]" pageId="3" pageNumber="376">
<taxonomicNameLabel id="CDDB8124FF812F67BB16FEE89150FE89" box="[528,618,326,349]" pageId="3" pageNumber="376" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, non-type material collected at Chamela Bay, La Huerta, Jalisco, Pacific Ocean, Mexico.
</paragraph>
</caption>
<paragraph id="E423E04DFF812F67B999FE37966EFCBF" pageId="3" pageNumber="376">
<table id="969C12EDFF81D09BB999FE3796B0FCBF" box="[159,1418,409,875]" gridcols="5" gridrows="11" pageId="3" pageNumber="376">
<tr id="5AACE20FFF81D09BB999FE3796B0FE1A" box="[159,1418,409,462]" gridrow="0" pageId="3" pageNumber="376">
<th id="197D8B73FF81D09BB999FE379242FE1A" box="[159,376,409,462]" gridcol="0" gridrow="0" pageId="3" pageNumber="376">Catalog number</th>
<th id="197D8B73FF81D09BB89EFE37913CFE1A" box="[408,518,409,462]" gridcol="1" gridrow="0" pageId="3" pageNumber="376">Number of specimens</th>
<th id="197D8B73FF81D09BBB20FE37971FFE1A" box="[550,1061,409,462]" gridcol="2" gridrow="0" pageId="3" pageNumber="376">Locality</th>
<th id="197D8B73FF81D09BBD3FFE37979BFE1A" box="[1081,1185,409,462]" gridcol="3" gridrow="0" pageId="3" pageNumber="376">Depth (m)</th>
<th id="197D8B73FF81D09BBDC6FE3796B0FE1A" box="[1216,1418,409,462]" gridcol="4" gridrow="0" pageId="3" pageNumber="376">Date</th>
</tr>
<tr id="5AACE20FFF81D09BB999FE4F96B0FE23" box="[159,1418,481,503]" gridrow="1" pageId="3" pageNumber="376">
<th id="197D8B73FF81D09BB999FE4F9242FE23" box="[159,376,481,503]" gridcol="0" gridrow="1" pageId="3" pageNumber="376">ICML-UNAM 11804</th>
<td id="197D8B73FF81D09BB89EFE4F913CFE23" box="[408,518,481,503]" gridcol="1" gridrow="1" pageId="3" pageNumber="376">4</td>
<td id="197D8B73FF81D09BBB20FE4F971FFE23" box="[550,1061,481,503]" gridcol="2" gridrow="1" pageId="3" pageNumber="376">Isla Cocinas, 19° 32' 45.4&quot; N, 105° 06' 35.59&quot; W</td>
<td id="197D8B73FF81D09BBD3FFE4F979BFE23" box="[1081,1185,481,503]" gridcol="3" gridrow="1" pageId="3" pageNumber="376">2</td>
<td id="197D8B73FF81D09BBDC6FE4F96B0FE23" box="[1216,1418,481,503]" gridcol="4" gridrow="1" pageId="3" pageNumber="376">November 18, 2013</td>
</tr>
<tr id="5AACE20FFF81D09BB999FDA496B0FDF4" box="[159,1418,522,544]" gridrow="2" pageId="3" pageNumber="376">
<th id="197D8B73FF81D09BB999FDA49242FDF4" box="[159,376,522,544]" gridcol="0" gridrow="2" pageId="3" pageNumber="376">ICML-UNAM 11852</th>
<td id="197D8B73FF81D09BB89EFDA4913CFDF4" box="[408,518,522,544]" gridcol="1" gridrow="2" pageId="3" pageNumber="376">9</td>
<td id="197D8B73FF81D09BBB20FDA4971FFDF4" box="[550,1061,522,544]" gridcol="2" gridrow="2" pageId="3" pageNumber="376">Isla Cocinas, 19° 32 49.3″ N, 105° 06 32.2″ W</td>
<td id="197D8B73FF81D09BBD3FFDA4979BFDF4" box="[1081,1185,522,544]" gridcol="3" gridrow="2" pageId="3" pageNumber="376">6.5</td>
<td id="197D8B73FF81D09BBDC6FDA496B0FDF4" box="[1216,1418,522,544]" gridcol="4" gridrow="2" pageId="3" pageNumber="376">November 20, 2013</td>
</tr>
<tr id="5AACE20FFF81D09BB999FD9A96B0FD9E" box="[159,1418,564,586]" gridrow="3" pageId="3" pageNumber="376">
<th id="197D8B73FF81D09BB999FD9A9242FD9E" box="[159,376,564,586]" gridcol="0" gridrow="3" pageId="3" pageNumber="376">ICML-UNAM 11868</th>
<td id="197D8B73FF81D09BB89EFD9A913CFD9E" box="[408,518,564,586]" gridcol="1" gridrow="3" pageId="3" pageNumber="376">103</td>
<td id="197D8B73FF81D09BBB20FD9A971FFD9E" box="[550,1061,564,586]" gridcol="2" gridrow="3" pageId="3" pageNumber="376">Isla San Pedro, 19° 33 13.2″ N, 105° 06 37.7″W</td>
<td id="197D8B73FF81D09BBD3FFD9A979BFD9E" box="[1081,1185,564,586]" gridcol="3" gridrow="3" pageId="3" pageNumber="376">5</td>
<td id="197D8B73FF81D09BBDC6FD9A96B0FD9E" box="[1216,1418,564,586]" gridcol="4" gridrow="3" pageId="3" pageNumber="376">November 15, 2013</td>
</tr>
<tr id="5AACE20FFF81D09BB999FDF396B0FDA7" box="[159,1418,605,627]" gridrow="4" pageId="3" pageNumber="376">
<th id="197D8B73FF81D09BB999FDF39242FDA7" box="[159,376,605,627]" gridcol="0" gridrow="4" pageId="3" pageNumber="376">ICML-UNAM 11882</th>
<td id="197D8B73FF81D09BB89EFDF3913CFDA7" box="[408,518,605,627]" gridcol="1" gridrow="4" pageId="3" pageNumber="376">1</td>
<td id="197D8B73FF81D09BBB20FDF3971FFDA7" box="[550,1061,605,627]" gridcol="2" gridrow="4" pageId="3" pageNumber="376">Isla Colorada, 19° 32 23.9″ N, 105° 05 31.9″ W</td>
<td id="197D8B73FF81D09BBD3FFDF3979BFDA7" box="[1081,1185,605,627]" gridcol="3" gridrow="4" pageId="3" pageNumber="376">6-8</td>
<td id="197D8B73FF81D09BBDC6FDF396B0FDA7" box="[1216,1418,605,627]" gridcol="4" gridrow="4" pageId="3" pageNumber="376">November 12, 2013</td>
</tr>
<tr id="5AACE20FFF81D09BB999FD2896B0FD48" box="[159,1418,646,668]" gridrow="5" pageId="3" pageNumber="376">
<th id="197D8B73FF81D09BB999FD289242FD48" box="[159,376,646,668]" gridcol="0" gridrow="5" pageId="3" pageNumber="376">ICML-UNAM 11898</th>
<td id="197D8B73FF81D09BB89EFD28913CFD48" box="[408,518,646,668]" gridcol="1" gridrow="5" pageId="3" pageNumber="376">14</td>
<td id="197D8B73FF81D09BBB20FD28971FFD48" box="[550,1061,646,668]" gridcol="2" gridrow="5" pageId="3" pageNumber="376">Isla San Pedro, 19° 32 1″ N, 105° 05 17.9″ W</td>
<td id="197D8B73FF81D09BBD3FFD28979BFD48" box="[1081,1185,646,668]" gridcol="3" gridrow="5" pageId="3" pageNumber="376">5</td>
<td id="197D8B73FF81D09BBDC6FD2896B0FD48" box="[1216,1418,646,668]" gridcol="4" gridrow="5" pageId="3" pageNumber="376">November 12, 2013</td>
</tr>
<tr id="5AACE20FFF81D09BB999FD1E96B0FD12" box="[159,1418,688,710]" gridrow="6" pageId="3" pageNumber="376">
<th id="197D8B73FF81D09BB999FD1E9242FD12" box="[159,376,688,710]" gridcol="0" gridrow="6" pageId="3" pageNumber="376">ICML-UNAM 11910</th>
<td id="197D8B73FF81D09BB89EFD1E913CFD12" box="[408,518,688,710]" gridcol="1" gridrow="6" pageId="3" pageNumber="376">14</td>
<td id="197D8B73FF81D09BBB20FD1E971FFD12" box="[550,1061,688,710]" gridcol="2" gridrow="6" pageId="3" pageNumber="376">Isla Pajarera, 19° 33 27.7″ N, 105° 07 0.6″ W</td>
<td id="197D8B73FF81D09BBD3FFD1E979BFD12" box="[1081,1185,688,710]" gridcol="3" gridrow="6" pageId="3" pageNumber="376">16</td>
<td id="197D8B73FF81D09BBDC6FD1E96B0FD12" box="[1216,1418,688,710]" gridcol="4" gridrow="6" pageId="3" pageNumber="376">November 13, 2013</td>
</tr>
<tr id="5AACE20FFF81D09BB999FD7796B0FD3B" box="[159,1418,729,751]" gridrow="7" pageId="3" pageNumber="376">
<th id="197D8B73FF81D09BB999FD779242FD3B" box="[159,376,729,751]" gridcol="0" gridrow="7" pageId="3" pageNumber="376">LEMA-CEQ144</th>
<td id="197D8B73FF81D09BB89EFD77913CFD3B" box="[408,518,729,751]" gridcol="1" gridrow="7" pageId="3" pageNumber="376">1</td>
<td id="197D8B73FF81D09BBB20FD77971FFD3B" box="[550,1061,729,751]" gridcol="2" gridrow="7" pageId="3" pageNumber="376">Isla Cocinas, 19° 33 7.9″ N, 105° 06 31.85″ W</td>
<td id="197D8B73FF81D09BBD3FFD77979BFD3B" box="[1081,1185,729,751]" gridcol="3" gridrow="7" pageId="3" pageNumber="376">8</td>
<td id="197D8B73FF81D09BBDC6FD7796B0FD3B" box="[1216,1418,729,751]" gridcol="4" gridrow="7" pageId="3" pageNumber="376">April 24, 2009</td>
</tr>
<tr id="5AACE20FFF81D09BB999FCAC96B0FCCC" box="[159,1418,770,792]" gridrow="8" pageId="3" pageNumber="376">
<th id="197D8B73FF81D09BB999FCAC9242FCCC" box="[159,376,770,792]" gridcol="0" gridrow="8" pageId="3" pageNumber="376">LEMA-CEQ624</th>
<td id="197D8B73FF81D09BB89EFCAC913CFCCC" box="[408,518,770,792]" gridcol="1" gridrow="8" pageId="3" pageNumber="376">4</td>
<td id="197D8B73FF81D09BBB20FCAC971FFCCC" box="[550,1061,770,792]" gridcol="2" gridrow="8" pageId="3" pageNumber="376">El Novillo, 19° 33 22.67″ N, 105° 06 31.13″ W</td>
<td id="197D8B73FF81D09BBD3FFCAC979BFCCC" box="[1081,1185,770,792]" gridcol="3" gridrow="8" pageId="3" pageNumber="376">3</td>
<td id="197D8B73FF81D09BBDC6FCAC96B0FCCC" box="[1216,1418,770,792]" gridcol="4" gridrow="8" pageId="3" pageNumber="376">November 28, 2011</td>
</tr>
<tr id="5AACE20FFF81D09BB999FC8296B0FC96" box="[159,1418,812,834]" gridrow="9" pageId="3" pageNumber="376">
<th id="197D8B73FF81D09BB999FC829242FC96" box="[159,376,812,834]" gridcol="0" gridrow="9" pageId="3" pageNumber="376">LEMA-CEQ625</th>
<td id="197D8B73FF81D09BB89EFC82913CFC96" box="[408,518,812,834]" gridcol="1" gridrow="9" pageId="3" pageNumber="376">2</td>
<td id="197D8B73FF81D09BBB20FC82971FFC96" box="[550,1061,812,834]" gridcol="2" gridrow="9" pageId="3" pageNumber="376">El Novillo, 19° 33 22.67″ N, 105°0631.13″ W</td>
<td id="197D8B73FF81D09BBD3FFC82979BFC96" box="[1081,1185,812,834]" gridcol="3" gridrow="9" pageId="3" pageNumber="376">3</td>
<td id="197D8B73FF81D09BBDC6FC8296B0FC96" box="[1216,1418,812,834]" gridcol="4" gridrow="9" pageId="3" pageNumber="376">November 28, 2011</td>
</tr>
<tr id="5AACE20FFF81D09BB999FCFB96B0FCBF" box="[159,1418,853,875]" gridrow="10" pageId="3" pageNumber="376">
<th id="197D8B73FF81D09BB999FCFB9242FCBF" box="[159,376,853,875]" gridcol="0" gridrow="10" pageId="3" pageNumber="376">LEMA-CEQ626</th>
<td id="197D8B73FF81D09BB89EFCFB913CFCBF" box="[408,518,853,875]" gridcol="1" gridrow="10" pageId="3" pageNumber="376">1</td>
<td id="197D8B73FF81D09BBB20FCFB971FFCBF" box="[550,1061,853,875]" gridcol="2" gridrow="10" pageId="3" pageNumber="376">Isla Pajarera, 19° 33 44.34″ N, 105° 06 42.24″ W</td>
<td id="197D8B73FF81D09BBD3FFCFB979BFCBF" box="[1081,1185,853,875]" gridcol="3" gridrow="10" pageId="3" pageNumber="376">2</td>
<td id="197D8B73FF81D09BBDC6FCFB96B0FCBF" box="[1216,1418,853,875]" gridcol="4" gridrow="10" pageId="3" pageNumber="376">March 7, 2013</td>
</tr>
</table>
</paragraph>
<subSubSection id="AC86B3C6FF812F67B9C1FC019033FC39" pageId="3" pageNumber="376" type="etymology">
<paragraph id="E423E04DFF812F67B9C1FC019033FC39" blockId="3.[151,1437,943,2013]" pageId="3" pageNumber="376">
<emphasis id="D6E83C5FFF812F67B9C1FC019274FC1C" bold="true" box="[199,334,943,968]" pageId="3" pageNumber="376">Etymology.</emphasis>
The specific epithet
<taxonomicName id="239C9BCEFF812F67BB5AFC1E9003FC1C" authority="in Latin" authorityName="in Latin" box="[604,825,943,968]" class="Holothuroidea" family="Psolidae" genus="Lissothuria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="3" pageNumber="376" phylum="Echinodermata" rank="species" species="imbricata">
<emphasis id="D6E83C5FFF812F67BB5AFC1E91F3FC1C" box="[604,713,944,968]" italics="true" pageId="3" pageNumber="376">imbricata</emphasis>
in Latin
</taxonomicName>
means “imbricated, overlapping”; it refers to the conspicuous appearance of the imbricated dorsal scales.
</paragraph>
</subSubSection>
<subSubSection id="AC86B3C6FF812F60B9C1FC5990F9FF2C" lastPageId="4" lastPageNumber="377" pageId="3" pageNumber="376" type="materials_examined">
<paragraph id="E423E04DFF812F67B9C1FC599242F9F9" blockId="3.[151,1437,943,2013]" pageId="3" pageNumber="376">
<emphasis id="D6E83C5FFF812F67B9C1FC5992F3FBC4" bold="true" box="[199,457,1015,1040]" pageId="3" pageNumber="376">
<typeStatus id="3B275EEFFF812F67B9C1FC59920EFBC4" box="[199,308,1015,1040]" pageId="3" pageNumber="376" type="holotype">Holotype</typeStatus>
description.
</emphasis>
<quantity id="23644DA8FF812F67B8D4FC56910DFBDB" box="[466,567,1016,1040]" metricMagnitude="-2" metricUnit="m" metricValue="1.72" pageId="3" pageNumber="376" unit="mm" value="17.2">17.2 mm</quantity>
total length (TL) and
<quantity id="23644DA8FF812F67BA31FC5690B5FBDB" box="[823,911,1016,1040]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="3" pageNumber="376" unit="mm" value="4.4">4.4 mm</quantity>
high at the central part of the body (
<quantity id="23644DA8FF812F67BC30FC5996A7FBDB" box="[1334,1437,1015,1040]" metricMagnitude="-2" metricUnit="m" metricValue="1.68" pageId="3" pageNumber="376" unit="mm" value="16.8">16.8 mm</quantity>
sole length,
<quantity id="23644DA8FF812F67B826FBB2924CFBE0" box="[288,374,1052,1076]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="3" pageNumber="376" unit="mm" value="8.4">8.4 mm</quantity>
wide). Elongated body; well calcified test. Mouth and anus elevated (
<figureCitation id="7CA7FCC8FF812F67BD8FFBB29624FBE0" box="[1161,1310,1052,1077]" captionStart="FIGURE 1" captionStartId="2.[151,250,1773,1795]" captionTargetBox="[151,1436,303,1748]" captionTargetId="figure@2.[151,1436,303,1748]" captionTargetPageId="2" captionText="FIGURE 1. Lissothuria imbricata sp. nov., holotype ICML-UNAM 18000, A, Dorsal view; B, Ventral view. Paratype ICML- UNAM 12135; C, In situ picture showing habitat preferences; D, Live specimen, lateral view. All scale bars: 5 mm." httpUri="https://zenodo.org/record/1294885/files/figure.png" pageId="3" pageNumber="376">Figs. 1A, 1D</figureCitation>
). Tentacles contracted (
<figureCitation id="7CA7FCC8FF812F67B819FB919246FB8C" box="[287,380,1087,1112]" captionStart="FIGURE 1" captionStartId="2.[151,250,1773,1795]" captionTargetBox="[151,1436,303,1748]" captionTargetId="figure@2.[151,1436,303,1748]" captionTargetPageId="2" captionText="FIGURE 1. Lissothuria imbricata sp. nov., holotype ICML-UNAM 18000, A, Dorsal view; B, Ventral view. Paratype ICML- UNAM 12135; C, In situ picture showing habitat preferences; D, Live specimen, lateral view. All scale bars: 5 mm." httpUri="https://zenodo.org/record/1294885/files/figure.png" pageId="3" pageNumber="376">Fig. 1A</figureCitation>
). Dorsal surface with thick imbricating scales approximately 600900 µm in diameter; 11 scales between oral and anal openings, the marginal scales are smaller (
<figureCitation id="7CA7FCC8FF812F67BAC5FBCA9727FBA8" box="[963,1053,1124,1149]" captionStart="FIGURE 1" captionStartId="2.[151,250,1773,1795]" captionTargetBox="[151,1436,303,1748]" captionTargetId="figure@2.[151,1436,303,1748]" captionTargetPageId="2" captionText="FIGURE 1. Lissothuria imbricata sp. nov., holotype ICML-UNAM 18000, A, Dorsal view; B, Ventral view. Paratype ICML- UNAM 12135; C, In situ picture showing habitat preferences; D, Live specimen, lateral view. All scale bars: 5 mm." httpUri="https://zenodo.org/record/1294885/files/figure.png" pageId="3" pageNumber="376">Fig. 1D</figureCitation>
). Scales perforated for passage of inconspicuous tube feet (
<figureCitation id="7CA7FCC8FF812F67B8B6FB299133FB74" box="[432,521,1159,1184]" captionStart="FIGURE 1" captionStartId="2.[151,250,1773,1795]" captionTargetBox="[151,1436,303,1748]" captionTargetId="figure@2.[151,1436,303,1748]" captionTargetPageId="2" captionText="FIGURE 1. Lissothuria imbricata sp. nov., holotype ICML-UNAM 18000, A, Dorsal view; B, Ventral view. Paratype ICML- UNAM 12135; C, In situ picture showing habitat preferences; D, Live specimen, lateral view. All scale bars: 5 mm." httpUri="https://zenodo.org/record/1294885/files/figure.png" pageId="3" pageNumber="376">Fig. 1A</figureCitation>
;
<figureCitation id="7CA7FCC8FF812F67BB10FB2991A8FB74" box="[534,658,1159,1184]" captionStart="FIGURE 2" captionStartId="4.[151,250,1781,1803]" captionTargetBox="[284,1303,315,1755]" captionTargetId="figure@4.[284,1303,315,1755]" captionTargetPageId="4" captionText="FIGURE 2. Lissothuria imbricata sp. nov., holotype ICML-UNAM 18000, A, Dorsal perforated scale for passage of tube feet, scale bar: 100 µm; B, Detail of perforations in the dorsal scale; CE, Massive towers from the dorsal side, with slightly spiny apex, scale bar: 50 µm; FI, Baskets from the dorsal side, scale bar: 40 µm; JK, Dorsal tube feet vestigial end plates, scale bar: 50 µm." httpUri="https://zenodo.org/record/1294887/files/figure.png" pageId="3" pageNumber="376">Fig. 2AB</figureCitation>
); 20 pointed narrow valves surrounding the oral opening; anal valves are small and rounded varying in number (
<figureCitation id="7CA7FCC8FF812F67BB70FB0291EAFB10" box="[630,720,1196,1221]" captionStart="FIGURE 1" captionStartId="2.[151,250,1773,1795]" captionTargetBox="[151,1436,303,1748]" captionTargetId="figure@2.[151,1436,303,1748]" captionTargetPageId="2" captionText="FIGURE 1. Lissothuria imbricata sp. nov., holotype ICML-UNAM 18000, A, Dorsal view; B, Ventral view. Paratype ICML- UNAM 12135; C, In situ picture showing habitat preferences; D, Live specimen, lateral view. All scale bars: 5 mm." httpUri="https://zenodo.org/record/1294885/files/figure.png" pageId="3" pageNumber="376">Fig. 1D</figureCitation>
). Sole leathery, translucent, elongated, fringed by a single outer peripheral line of small tube feet; inner peripheral series with three rows of closely crowded tube feet, which form a continuous margin, the mid-ventral radius with a double row of tube feet scarce in the midventral area and becoming crowded anterior and posteriorly (
<figureCitation id="7CA7FCC8FF812F67BBA4FAB991C7FAE4" box="[674,765,1303,1328]" captionStart="FIGURE 1" captionStartId="2.[151,250,1773,1795]" captionTargetBox="[151,1436,303,1748]" captionTargetId="figure@2.[151,1436,303,1748]" captionTargetPageId="2" captionText="FIGURE 1. Lissothuria imbricata sp. nov., holotype ICML-UNAM 18000, A, Dorsal view; B, Ventral view. Paratype ICML- UNAM 12135; C, In situ picture showing habitat preferences; D, Live specimen, lateral view. All scale bars: 5 mm." httpUri="https://zenodo.org/record/1294885/files/figure.png" pageId="3" pageNumber="376">Fig. 1B</figureCitation>
). Dorsal tube feet distributed in all dorsal surface, those located in the ambulacra are numerous and conspicuous (
<figureCitation id="7CA7FCC8FF812F67BA11FA929055FA80" box="[791,879,1340,1365]" captionStart="FIGURE 1" captionStartId="2.[151,250,1773,1795]" captionTargetBox="[151,1436,303,1748]" captionTargetId="figure@2.[151,1436,303,1748]" captionTargetPageId="2" captionText="FIGURE 1. Lissothuria imbricata sp. nov., holotype ICML-UNAM 18000, A, Dorsal view; B, Ventral view. Paratype ICML- UNAM 12135; C, In situ picture showing habitat preferences; D, Live specimen, lateral view. All scale bars: 5 mm." httpUri="https://zenodo.org/record/1294885/files/figure.png" pageId="3" pageNumber="376">Fig. 1A</figureCitation>
). Ten dendritic tentacles, the two ventral most are smaller than the rest. Calcareous ring simple, slim, low, radial and interradial plates about the same size and well calcified. Radial plates are larger than the interradials, in the upper part they have a small and narrow slit for the insertion of the retractor muscle; ventral and dorsal radial plates are equal in size and shape. The interradial plates have a shallow groove in the lower part; dorsal and ventral interradial pieces are equal in shape and size. One Polian vesicle. Color in alcohol yellow to light purple, tentacles white. Well-developed gonads that occupy most of the coelomic cavity.
</paragraph>
<paragraph id="E423E04DFF812F60B9C1F99690F9FF2C" blockId="3.[151,1437,943,2013]" lastBlockId="4.[151,1436,151,248]" lastPageId="4" lastPageNumber="377" pageId="3" pageNumber="376">
<emphasis id="D6E83C5FFF812F67B9C1F9969295F984" box="[199,431,1592,1616]" italics="true" pageId="3" pageNumber="376">Ossicle assemblage.</emphasis>
Ossicles of dorsal surface include dorsal scales, with one to six perforations for the passage of tube feet (
<figureCitation id="7CA7FCC8FF812F67B88EF9F2913FF9A0" box="[392,517,1628,1653]" captionStart="FIGURE 2" captionStartId="4.[151,250,1781,1803]" captionTargetBox="[284,1303,315,1755]" captionTargetId="figure@4.[284,1303,315,1755]" captionTargetPageId="4" captionText="FIGURE 2. Lissothuria imbricata sp. nov., holotype ICML-UNAM 18000, A, Dorsal perforated scale for passage of tube feet, scale bar: 100 µm; B, Detail of perforations in the dorsal scale; CE, Massive towers from the dorsal side, with slightly spiny apex, scale bar: 50 µm; FI, Baskets from the dorsal side, scale bar: 40 µm; JK, Dorsal tube feet vestigial end plates, scale bar: 50 µm." httpUri="https://zenodo.org/record/1294887/files/figure.png" pageId="3" pageNumber="376">Fig. 2AB</figureCitation>
). Small dorsal scales (between 500 to 700 µm) are present at the base of the test, while those of the rest of the body can reach up to
<quantity id="23644DA8FF812F67BBC1F92E901AF94C" box="[711,800,1664,1688]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="376" unit="mm" value="1.8">1.8 mm</quantity>
. Dorsal scales are oriented towards the mid dorsal line of the body; those that are surrounding the mouth and the anus, are forming concentric lines (
<figureCitation id="7CA7FCC8FF812F67BDBFF90A9636F969" box="[1209,1292,1700,1725]" captionStart="FIGURE 1" captionStartId="2.[151,250,1773,1795]" captionTargetBox="[151,1436,303,1748]" captionTargetId="figure@2.[151,1436,303,1748]" captionTargetPageId="2" captionText="FIGURE 1. Lissothuria imbricata sp. nov., holotype ICML-UNAM 18000, A, Dorsal view; B, Ventral view. Paratype ICML- UNAM 12135; C, In situ picture showing habitat preferences; D, Live specimen, lateral view. All scale bars: 5 mm." httpUri="https://zenodo.org/record/1294885/files/figure.png" pageId="3" pageNumber="376">Fig.1D</figureCitation>
). The dorsal dermis posses massive towers with slightly spiny apex, the base may be smooth or irregular, the base measures between 170 and 200 µm in diameter, the height can be up to 190 µm (
<figureCitation id="7CA7FCC8FF812F67BACCF9429779F8D0" box="[970,1091,1772,1797]" captionStart="FIGURE 2" captionStartId="4.[151,250,1781,1803]" captionTargetBox="[284,1303,315,1755]" captionTargetId="figure@4.[284,1303,315,1755]" captionTargetPageId="4" captionText="FIGURE 2. Lissothuria imbricata sp. nov., holotype ICML-UNAM 18000, A, Dorsal perforated scale for passage of tube feet, scale bar: 100 µm; B, Detail of perforations in the dorsal scale; CE, Massive towers from the dorsal side, with slightly spiny apex, scale bar: 50 µm; FI, Baskets from the dorsal side, scale bar: 40 µm; JK, Dorsal tube feet vestigial end plates, scale bar: 50 µm." httpUri="https://zenodo.org/record/1294887/files/figure.png" pageId="3" pageNumber="376">Fig. 2CE</figureCitation>
), these massive towers reduce their size when approaching the lateral areas of the test, reaching 70 and 100 µm in diameter, the height can be up to 90 µm. Big baskets are 50 to 60 µm in diameter, the edge of which is usually spiny; small baskets are 40 to 45 µm in diameter, the base and the edge of which are usually knobbed (
<figureCitation id="7CA7FCC8FF812F67BAA7F8F99735F8A4" box="[929,1039,1879,1904]" captionStart="FIGURE 2" captionStartId="4.[151,250,1781,1803]" captionTargetBox="[284,1303,315,1755]" captionTargetId="figure@4.[284,1303,315,1755]" captionTargetPageId="4" captionText="FIGURE 2. Lissothuria imbricata sp. nov., holotype ICML-UNAM 18000, A, Dorsal perforated scale for passage of tube feet, scale bar: 100 µm; B, Detail of perforations in the dorsal scale; CE, Massive towers from the dorsal side, with slightly spiny apex, scale bar: 50 µm; FI, Baskets from the dorsal side, scale bar: 40 µm; JK, Dorsal tube feet vestigial end plates, scale bar: 50 µm." httpUri="https://zenodo.org/record/1294887/files/figure.png" pageId="3" pageNumber="376">Fig. 2FI</figureCitation>
). Curved supporting rods in ventral tube feet, here are also perforated plates; end plates appear to be vestigial in the dorsal tube feet (
<figureCitation id="7CA7FCC8FF812F67BDE4F8D2966DF840" box="[1250,1367,1916,1941]" captionStart="FIGURE 2" captionStartId="4.[151,250,1781,1803]" captionTargetBox="[284,1303,315,1755]" captionTargetId="figure@4.[284,1303,315,1755]" captionTargetPageId="4" captionText="FIGURE 2. Lissothuria imbricata sp. nov., holotype ICML-UNAM 18000, A, Dorsal perforated scale for passage of tube feet, scale bar: 100 µm; B, Detail of perforations in the dorsal scale; CE, Massive towers from the dorsal side, with slightly spiny apex, scale bar: 50 µm; FI, Baskets from the dorsal side, scale bar: 40 µm; JK, Dorsal tube feet vestigial end plates, scale bar: 50 µm." httpUri="https://zenodo.org/record/1294887/files/figure.png" pageId="3" pageNumber="376">Fig. 2JK</figureCitation>
). Sole ossicles are knobbed buttons with four holes (
<figureCitation id="7CA7FCC8FF812F67BBB3F831900EF86C" box="[693,820,1951,1976]" captionStart="FIGURE 3" captionStartId="5.[151,250,1872,1894]" captionTargetBox="[167,1411,193,1833]" captionTargetId="figure@5.[167,1411,193,1833]" captionTargetPageId="5" captionText="FIGURE 3. Lissothuria imbricata sp. nov., holotype ICML-UNAM 18000, AC, Ventral ossicles, knobbed buttons, scale bar: 50 µm; DF, Ventral ossicles, various shapes of perforated plates with smooth or knobbed margin, scale bar: 50 µm; GI, Ventral ossicles, small shallow baskets, with knobbed edges, scale bar: 30 µm; JK, Tentacles ossicles, small rosettes, scale bar: 10 µm; L, Tentacles ossicles, flat, curved and perforated rods, scale bar: 50 µm; and M, Tentacles ossicles, perforated plates, scale bar: 100 µm." httpUri="https://zenodo.org/record/1294889/files/figure.png" pageId="3" pageNumber="376">Fig. 3AC</figureCitation>
), perforated plates in various shapes with smooth or knobbed margin, they reach up to 150 µm in length (
<figureCitation id="7CA7FCC8FF812F67BBEFF86A905BF808" box="[745,865,1988,2013]" captionStart="FIGURE 3" captionStartId="5.[151,250,1872,1894]" captionTargetBox="[167,1411,193,1833]" captionTargetId="figure@5.[167,1411,193,1833]" captionTargetPageId="5" captionText="FIGURE 3. Lissothuria imbricata sp. nov., holotype ICML-UNAM 18000, AC, Ventral ossicles, knobbed buttons, scale bar: 50 µm; DF, Ventral ossicles, various shapes of perforated plates with smooth or knobbed margin, scale bar: 50 µm; GI, Ventral ossicles, small shallow baskets, with knobbed edges, scale bar: 30 µm; JK, Tentacles ossicles, small rosettes, scale bar: 10 µm; L, Tentacles ossicles, flat, curved and perforated rods, scale bar: 50 µm; and M, Tentacles ossicles, perforated plates, scale bar: 100 µm." httpUri="https://zenodo.org/record/1294889/files/figure.png" pageId="3" pageNumber="376">Fig. 3DF</figureCitation>
), and there are also small shallow baskets 50 to 65 µm in diameter, with knobbed edges (
<figureCitation id="7CA7FCC8FF862F60BB4CFF399185FF64" box="[586,703,151,176]" captionStart="FIGURE 3" captionStartId="5.[151,250,1872,1894]" captionTargetBox="[167,1411,193,1833]" captionTargetId="figure@5.[167,1411,193,1833]" captionTargetPageId="5" captionText="FIGURE 3. Lissothuria imbricata sp. nov., holotype ICML-UNAM 18000, AC, Ventral ossicles, knobbed buttons, scale bar: 50 µm; DF, Ventral ossicles, various shapes of perforated plates with smooth or knobbed margin, scale bar: 50 µm; GI, Ventral ossicles, small shallow baskets, with knobbed edges, scale bar: 30 µm; JK, Tentacles ossicles, small rosettes, scale bar: 10 µm; L, Tentacles ossicles, flat, curved and perforated rods, scale bar: 50 µm; and M, Tentacles ossicles, perforated plates, scale bar: 100 µm." httpUri="https://zenodo.org/record/1294889/files/figure.png" pageId="4" pageNumber="377">Fig. 3GI</figureCitation>
). Tentacles generally with elaborate small rosettes, 40 to 50 µm wide (
<figureCitation id="7CA7FCC8FF862F60B9D8FF129269FF00" box="[222,339,188,213]" captionStart="FIGURE 3" captionStartId="5.[151,250,1872,1894]" captionTargetBox="[167,1411,193,1833]" captionTargetId="figure@5.[167,1411,193,1833]" captionTargetPageId="5" captionText="FIGURE 3. Lissothuria imbricata sp. nov., holotype ICML-UNAM 18000, AC, Ventral ossicles, knobbed buttons, scale bar: 50 µm; DF, Ventral ossicles, various shapes of perforated plates with smooth or knobbed margin, scale bar: 50 µm; GI, Ventral ossicles, small shallow baskets, with knobbed edges, scale bar: 30 µm; JK, Tentacles ossicles, small rosettes, scale bar: 10 µm; L, Tentacles ossicles, flat, curved and perforated rods, scale bar: 50 µm; and M, Tentacles ossicles, perforated plates, scale bar: 100 µm." httpUri="https://zenodo.org/record/1294889/files/figure.png" pageId="4" pageNumber="377">Fig. 3JK</figureCitation>
), flat, curved and perforated rods, up to 200 µm, most times wider in the center than at the end portions (
<figureCitation id="7CA7FCC8FF862F60B804FF71926CFF2C" box="[258,342,223,248]" captionStart="FIGURE 3" captionStartId="5.[151,250,1872,1894]" captionTargetBox="[167,1411,193,1833]" captionTargetId="figure@5.[167,1411,193,1833]" captionTargetPageId="5" captionText="FIGURE 3. Lissothuria imbricata sp. nov., holotype ICML-UNAM 18000, AC, Ventral ossicles, knobbed buttons, scale bar: 50 µm; DF, Ventral ossicles, various shapes of perforated plates with smooth or knobbed margin, scale bar: 50 µm; GI, Ventral ossicles, small shallow baskets, with knobbed edges, scale bar: 30 µm; JK, Tentacles ossicles, small rosettes, scale bar: 10 µm; L, Tentacles ossicles, flat, curved and perforated rods, scale bar: 50 µm; and M, Tentacles ossicles, perforated plates, scale bar: 100 µm." httpUri="https://zenodo.org/record/1294889/files/figure.png" pageId="4" pageNumber="377">Fig. 3L</figureCitation>
) and perforated plates 100 to 300 µm length (
<figureCitation id="7CA7FCC8FF862F60BA51FF71908CFF2C" box="[855,950,223,248]" captionStart="FIGURE 3" captionStartId="5.[151,250,1872,1894]" captionTargetBox="[167,1411,193,1833]" captionTargetId="figure@5.[167,1411,193,1833]" captionTargetPageId="5" captionText="FIGURE 3. Lissothuria imbricata sp. nov., holotype ICML-UNAM 18000, AC, Ventral ossicles, knobbed buttons, scale bar: 50 µm; DF, Ventral ossicles, various shapes of perforated plates with smooth or knobbed margin, scale bar: 50 µm; GI, Ventral ossicles, small shallow baskets, with knobbed edges, scale bar: 30 µm; JK, Tentacles ossicles, small rosettes, scale bar: 10 µm; L, Tentacles ossicles, flat, curved and perforated rods, scale bar: 50 µm; and M, Tentacles ossicles, perforated plates, scale bar: 100 µm." httpUri="https://zenodo.org/record/1294889/files/figure.png" pageId="4" pageNumber="377">Fig. 3M</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="B0E3B0C5FF862F60B991F95B93E5F8B2" httpUri="https://zenodo.org/record/1294887/files/figure.png" pageId="4" pageNumber="377" startId="4.[151,250,1781,1803]" targetBox="[284,1303,315,1755]" targetPageId="4">
<paragraph id="E423E04DFF862F60B991F95B93E5F8B2" blockId="4.[151,1437,1780,1894]" pageId="4" pageNumber="377">
<emphasis id="D6E83C5FFF862F60B991F95B9228F8DE" bold="true" box="[151,274,1781,1803]" pageId="4" pageNumber="377">FIGURE 2.</emphasis>
<taxonomicName id="239C9BCEFF862F60B81EF95A92C9F8DE" authority="Solís-Marín &amp; Arriaga-Ochoa &amp; Villa &amp; Laguarda-Figueras, 2018" authorityName="Solís-Marín &amp; Arriaga-Ochoa &amp; Villa &amp; Laguarda-Figueras" authorityYear="2018" box="[280,499,1780,1802]" class="Holothuroidea" family="Psolidae" genus="Lissothuria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="377" phylum="Echinodermata" rank="species" species="imbricata" status="sp. nov.">
<emphasis id="D6E83C5FFF862F60B81EF95A92C9F8DE" box="[280,499,1780,1802]" italics="true" pageId="4" pageNumber="377">Lissothuria imbricata</emphasis>
</taxonomicName>
<emphasis id="D6E83C5FFF862F60B8FCF95B9170F8DE" bold="true" box="[506,586,1781,1802]" pageId="4" pageNumber="377">
<taxonomicNameLabel id="CDDB8124FF862F60B8FCF95B9170F8DE" box="[506,586,1781,1802]" pageId="4" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype ICML-UNAM 18000, A, Dorsal perforated scale for passage of tube feet, scale bar: 100 µm; B, Detail of perforations in the dorsal scale; CE, Massive towers from the dorsal side, with slightly spiny apex, scale bar: 50 µm; FI, Baskets from the dorsal side, scale bar: 40 µm; JK, Dorsal tube feet vestigial end plates, scale bar: 50 µm.
</paragraph>
</caption>
<caption id="B0E3B0C5FF872F61B991F8FE9268F838" httpUri="https://zenodo.org/record/1294889/files/figure.png" pageId="5" pageNumber="378" startId="5.[151,250,1872,1894]" targetBox="[167,1411,193,1833]" targetPageId="5">
<paragraph id="E423E04DFF872F61B991F8FE9268F838" blockId="5.[151,1437,1872,2028]" pageId="5" pageNumber="378">
<emphasis id="D6E83C5FFF872F61B991F8FE9228F8B1" bold="true" box="[151,274,1872,1894]" pageId="5" pageNumber="378">FIGURE 3.</emphasis>
<taxonomicName id="239C9BCEFF872F61B81FF8FE92CCF8B2" authority="Solís-Marín &amp; Arriaga-Ochoa &amp; Villa &amp; Laguarda-Figueras, 2018" authorityName="Solís-Marín &amp; Arriaga-Ochoa &amp; Villa &amp; Laguarda-Figueras" authorityYear="2018" box="[281,502,1872,1894]" class="Holothuroidea" family="Psolidae" genus="Lissothuria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="5" pageNumber="378" phylum="Echinodermata" rank="species" species="imbricata" status="sp. nov.">
<emphasis id="D6E83C5FFF872F61B81FF8FE92CCF8B2" box="[281,502,1872,1894]" italics="true" pageId="5" pageNumber="378">Lissothuria imbricata</emphasis>
</taxonomicName>
<emphasis id="D6E83C5FFF872F61B8FAF8FF9176F8B2" bold="true" box="[508,588,1873,1894]" pageId="5" pageNumber="378">
<taxonomicNameLabel id="CDDB8124FF872F61B8FAF8FF9176F8B2" box="[508,588,1873,1894]" pageId="5" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype ICML-UNAM 18000, AC, Ventral ossicles, knobbed buttons, scale bar: 50 µm; DF, Ventral ossicles, various shapes of perforated plates with smooth or knobbed margin, scale bar: 50 µm; GI, Ventral ossicles, small shallow baskets, with knobbed edges, scale bar: 30 µm; JK, Tentacles ossicles, small rosettes, scale bar: 10 µm; L, Tentacles ossicles, flat, curved and perforated rods, scale bar: 50 µm; and M, Tentacles ossicles, perforated plates, scale bar: 100 µm.
</paragraph>
</caption>
<subSubSection id="AC86B3C6FF842F62B9C1FF3797BDFE94" pageId="6" pageNumber="379" type="description">
<paragraph id="E423E04DFF842F62B9C1FF3797BDFE94" blockId="6.[151,1437,151,536]" pageId="6" pageNumber="379">
<emphasis id="D6E83C5FFF842F62B9C1FF379290FF7B" box="[199,426,152,176]" italics="true" pageId="6" pageNumber="379">
<typeStatus id="3B275EEFFF842F62B9C1FF379214FF64" box="[199,302,153,176]" pageId="6" pageNumber="379" type="paratype">Paratype</typeStatus>
variations
</emphasis>
. Specimens slightly smaller than the
<typeStatus id="3B275EEFFF842F62BA67FF3990FCFF64" box="[865,966,151,176]" pageId="6" pageNumber="379" type="holotype">holotype</typeStatus>
,
<quantity id="23644DA8FF842F62BAD2FF36976BFF7B" box="[980,1105,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.7" metricValueMin="1.0" pageId="6" pageNumber="379" unit="mm" value="13.5" valueMax="17.0" valueMin="10.0">1017 mm</quantity>
TL and
<quantity id="23644DA8FF842F62BDBCFF36962EFF7B" box="[1210,1300,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="6" pageNumber="379" unit="mm" value="3.3">3.3 mm</quantity>
high in the central part of the body (
<quantity id="23644DA8FF842F62B8C7FF12910BFF00" box="[449,561,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.5" metricValueMin="0.9" pageId="6" pageNumber="379" unit="mm" value="12.0" valueMax="15.0" valueMin="9.0">915 mm</quantity>
sole length,
<quantity id="23644DA8FF842F62BBCBFF129014FF00" box="[717,814,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="8.0" metricValueMin="3.0" pageId="6" pageNumber="379" unit="mm" value="5.5" valueMax="8.0" valueMin="3.0">38 mm</quantity>
wide). Elongated body; well calcified test. 11 scales between oral and anal openings. Mouth, anus and internal morphology same as in the
<typeStatus id="3B275EEFFF842F62BDBAFF719618FF2C" box="[1212,1314,223,248]" pageId="6" pageNumber="379" type="holotype">holotype</typeStatus>
. Tentacles contracted. Dorsal scales approximately 500900 µm in diameter. Shape and size of the ossicles are the same as in the
<typeStatus id="3B275EEFFF842F62B9C6FE89921FFE94" box="[192,293,295,320]" pageId="6" pageNumber="379" type="holotype">holotype</typeStatus>
, except that in the smaller
<typeStatus id="3B275EEFFF842F62BB56FE86918BFE94" box="[592,689,296,320]" pageId="6" pageNumber="379" type="paratype">paratype</typeStatus>
(
<quantity id="23644DA8FF842F62BBC6FE86902DFEEB" box="[704,791,296,320]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="6" pageNumber="379" unit="mm" value="9.8">9.8 mm</quantity>
TL) ossicles are more abundant.
</paragraph>
</subSubSection>
<subSubSection id="AC86B3C6FF842F62B9C1FEE29060FE21" pageId="6" pageNumber="379" type="biology_ecology">
<paragraph id="E423E04DFF842F62B9C1FEE297E5FE5C" blockId="6.[151,1437,151,536]" pageId="6" pageNumber="379">
<emphasis id="D6E83C5FFF842F62B9C1FEE29211FEB1" bold="true" box="[199,299,332,357]" pageId="6" pageNumber="379">Ecology.</emphasis>
<taxonomicName id="239C9BCEFF842F62B835FEE2911DFEB0" authority="Solís-Marín &amp; Arriaga-Ochoa &amp; Villa &amp; Laguarda-Figueras, 2018" authorityName="Solís-Marín &amp; Arriaga-Ochoa &amp; Villa &amp; Laguarda-Figueras" authorityYear="2018" box="[307,551,332,356]" class="Holothuroidea" family="Psolidae" genus="Lissothuria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="6" pageNumber="379" phylum="Echinodermata" rank="species" species="imbricata" status="sp. nov.">
<emphasis id="D6E83C5FFF842F62B835FEE2911DFEB0" box="[307,551,332,356]" italics="true" pageId="6" pageNumber="379">Lissothuria imbricata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CDDB8124FF842F62BB28FEE391BCFEB0" box="[558,646,333,356]" pageId="6" pageNumber="379" rank="species">sp. nov.</taxonomicNameLabel>
inhabits rocky reefs with stone blocks, it can be found attached under or on top of rocks; it uses its tube feet to partially cover its dorsal side with algae debris (
<figureCitation id="7CA7FCC8FF842F62BD7CFEC197EBFE5C" box="[1146,1233,367,392]" captionStart="FIGURE 1" captionStartId="2.[151,250,1773,1795]" captionTargetBox="[151,1436,303,1748]" captionTargetId="figure@2.[151,1436,303,1748]" captionTargetPageId="2" captionText="FIGURE 1. Lissothuria imbricata sp. nov., holotype ICML-UNAM 18000, A, Dorsal view; B, Ventral view. Paratype ICML- UNAM 12135; C, In situ picture showing habitat preferences; D, Live specimen, lateral view. All scale bars: 5 mm." httpUri="https://zenodo.org/record/1294885/files/figure.png" pageId="6" pageNumber="379">Fig. 1C</figureCitation>
).
</paragraph>
<paragraph id="E423E04DFF842F62B9C1FE3A9060FE21" blockId="6.[151,1437,151,536]" pageId="6" pageNumber="379">
<emphasis id="D6E83C5FFF842F62B9C1FE3A9249FE79" bold="true" box="[199,371,404,429]" pageId="6" pageNumber="379">Reproduction.</emphasis>
<taxonomicName id="239C9BCEFF842F62B879FE3A9143FE78" authority="Solís-Marín &amp; Arriaga-Ochoa &amp; Villa &amp; Laguarda-Figueras, 2018" authorityName="Solís-Marín &amp; Arriaga-Ochoa &amp; Villa &amp; Laguarda-Figueras" authorityYear="2018" box="[383,633,404,428]" class="Holothuroidea" family="Psolidae" genus="Lissothuria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="6" pageNumber="379" phylum="Echinodermata" rank="species" species="imbricata" status="sp. nov.">
<emphasis id="D6E83C5FFF842F62B879FE3A9143FE78" box="[383,633,404,428]" italics="true" pageId="6" pageNumber="379">Lissothuria imbricata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CDDB8124FF842F62BB80FE3B91D9FE78" box="[646,739,405,428]" pageId="6" pageNumber="379" rank="species">sp. nov.</taxonomicNameLabel>
is a gonochoric species; females have lecithotrophic eggs between 420 and 470 µm in diameter, in this case, ripe gonads occupy more than 80% of the celomic cavity. Neither brooding or external sexual dimorphism was observed.
</paragraph>
</subSubSection>
<subSubSection id="AC86B3C6FF842F62B9C1FE51970CFDCC" box="[199,1078,511,536]" pageId="6" pageNumber="379" type="description">
<paragraph id="E423E04DFF842F62B9C1FE51970CFDCC" blockId="6.[151,1437,151,536]" box="[199,1078,511,536]" pageId="6" pageNumber="379">
<materialsCitation id="54F4EA10FF842F62B9C1FE51970CFDCC" ID-GBIF-Occurrence="1851020344" box="[199,1078,511,536]" country="Mexico" location="Chamela Bay" pageId="6" pageNumber="379" specimenCount="1" stateProvince="Jalisco">
<emphasis id="D6E83C5FFF842F62B9C1FE51913EFDCC" bold="true" box="[199,516,511,536]" pageId="6" pageNumber="379">Geographical distribution.</emphasis>
Only known from
<location id="E143B696FF842F62BBDDFE519049FDCC" LSID="urn:lsid:plazi:treatment:6C35515BFF832F62B991F901970CFDCC:E143B696FF842F62BBDDFE519049FDCC" box="[731,883,511,536]" country="Mexico" name="Chamela Bay" pageId="6" pageNumber="379" stateProvince="Jalisco">Chamela Bay</location>
,
<collectingRegion id="26582EAFFF842F62BA78FE5190F7FDCC" box="[894,973,511,536]" country="Mexico" name="Jalisco" pageId="6" pageNumber="379">Jalisco</collectingRegion>
,
<collectingCountry id="9C8BA0DDFF842F62BADFFE519709FDCC" box="[985,1075,511,536]" name="Mexico" pageId="6" pageNumber="379">Mexico</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>