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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1081.75927" ID-GBIF-Dataset="6ebea377-f492-43a0-b87f-eea36d326201" ID-PMC="PMC8782822" ID-Pensoft-Pub="1313-2970-1081-127" ID-Pensoft-UUID="A101072FE2F95283985CF69DC922CED0" ID-PubMed="35087299" ID-ZooBank="AD2D54F556D14DB2B7F8F3B3E0528394" ModsDocID="1313-2970-1081-127" checkinTime="1642207361831" checkinUser="pensoft" docAuthor="Yoshitomi, Hiroyuki &amp; Pham, Thai Hong" docDate="2022" docId="079B2E1C8B25532C8E8654CF99A0156F" docLanguage="en" docName="ZooKeys 1081: 127-136" docOrigin="ZooKeys 1081" docPubDate="2022-01-14" docSource="http://dx.doi.org/10.3897/zookeys.1081.75927" docTitle="Cacodaemon proavus Strohecker 1964" docType="treatment" docVersion="4" id="A101072FE2F95283985CF69DC922CED0" lastPageNumber="127" masterDocId="A101072FE2F95283985CF69DC922CED0" masterDocTitle="The genus Cacodaemon (Coleoptera, Endomychidae) of Vietnam" masterLastPageNumber="136" masterPageNumber="127" pageNumber="127" updateTime="1668151305564" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The genus Cacodaemon (Coleoptera, Endomychidae) of Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yoshitomi, Hiroyuki</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8416-9249</mods:nameIdentifier>
<mods:affiliation>Entomological Laboratory, Faculty of Agriculture, Ehime University, Tarumi 3 - 5 - 7, Matsuyama, 790 - 8566 Japan</mods:affiliation>
<mods:nameIdentifier type="email">hymushi@agr.ehime-u.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pham, Thai Hong</mods:namePart>
<mods:affiliation>Mientrung Institute for Scientific Research, Vietnam National Museum of Nature, VAST, 321 Huynh Thuc Khang, Hue, Vietnam &amp; Graduate School of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-01-14</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1081</mods:number>
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<mods:start>127</mods:start>
<mods:end>136</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1081.75927</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1081.75927</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1081-127</mods:identifier>
<mods:identifier type="ZooBank">AD2D54F556D14DB2B7F8F3B3E0528394</mods:identifier>
<mods:identifier type="Pensoft-UUID">A101072FE2F95283985CF69DC922CED0</mods:identifier>
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<subSection pageId="0" pageNumber="127" type="taxonomy">
<treatment ID-GBIF-Taxon="192143027" LSID="urn:lsid:plazi:treatment:079B2E1C8B25532C8E8654CF99A0156F" httpUri="http://treatment.plazi.org/id/079B2E1C8B25532C8E8654CF99A0156F" lastPageNumber="127" pageId="0" pageNumber="127">
<subSubSection pageId="0" pageNumber="127" type="nomenclature">
<paragraph pageId="0" pageNumber="127">
<taxonomicName LSID="079B2E1C-8B25-532C-8E86-54CF99A0156F" authority="Strohecker, 1964" authorityName="Strohecker" authorityYear="1964" class="Insecta" family="Endomychidae" genus="Cacodaemon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cacodaemon proavus" order="Coleoptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="proavus">Cacodaemon proavus Strohecker, 1964</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="description">
<paragraph pageId="0" pageNumber="127">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Cacodaemon spp. A-D C. laotinus laotinus, Vietnamese specimens (A, B) and Laotian specimens (C, D) E C. vietnamensis sp. nov., holotype F C. proavus A, B male C, D-F female. Scales: 1.0 mm." figureDoi="10.3897/zookeys.1081.75927.figure1" httpUri="https://binary.pensoft.net/fig/634063" pageId="0" pageNumber="127">Figs 1F</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Fore tibiae of Cacodaemon spp. A-D C. laotinus laotinus, Vietnamese specimens (A, B) and Laotian specimens (C, D) E C. vietnamensis sp. nov., holotype F C. proavus A, C male B, D-F female. Scale: 1.0 mm." figureDoi="10.3897/zookeys.1081.75927.figure2" httpUri="https://binary.pensoft.net/fig/634064" pageId="0" pageNumber="127">, 2F</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Elytral appendages of Cacodaemon spp. in lateral view A-D C. laotinus laotinus, Vietnamese specimens (A, B) and Laotian specimens (C, D) E C. vietnamensis sp. nov., holotype F C. proavus A, C male B, D-F female. Scale: 1.0 mm." figureDoi="10.3897/zookeys.1081.75927.figure3" httpUri="https://binary.pensoft.net/fig/634065" pageId="0" pageNumber="127">, 3F</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Female genitalia of Cacodaemon spp. A, B C. laotinus laotinus C, D C. vietnamensis sp. nov. E, F C. proavus A, C, E sternite VIII B, D, F ovipositor. Scale: 0.5 mm." figureDoi="10.3897/zookeys.1081.75927.figure5" httpUri="https://binary.pensoft.net/fig/634067" pageId="0" pageNumber="127">, 5E</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Female genitalia of Cacodaemon spp. A, B C. laotinus laotinus C, D C. vietnamensis sp. nov. E, F C. proavus A, C, E sternite VIII B, D, F ovipositor. Scale: 0.5 mm." figureDoi="10.3897/zookeys.1081.75927.figure5" httpUri="https://binary.pensoft.net/fig/634067" pageId="0" pageNumber="127">, 5F</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="reference_group">
<paragraph pageId="0" pageNumber="127">
<taxonomicName authorityName="Strohecker" authorityYear="1964" class="Insecta" family="Endomychidae" genus="Cacodaemon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cacodaemon proavus" order="Coleoptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="proavus">Cacodaemon proavus</taxonomicName>
Strohecker, 1964: 346;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.1999.1.1" author="Shockley, FW" journalOrPublisher="Zootaxa" pageId="0" pageNumber="127" pagination="1 - 113" refId="B5" refString="Shockley, FW, Tomaszewska, KW, McHugh, JV, 2009. An annotated checklist of the handsome fungus beetles of the world (Coleoptera: Cucujoidea: Endomychidae). Zootaxa 1999: 1 - 113, DOI: https://doi.org/10.11646/zootaxa.1999.1.1" title="An annotated checklist of the handsome fungus beetles of the world (Coleoptera: Cucujoidea: Endomychidae)." url="https://doi.org/10.11646/zootaxa.1999.1.1" volume="1999" year="2009">Shockley et al. 2009</bibRefCitation>
: 36.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="materials_examined">
<paragraph pageId="0" pageNumber="127">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="127">
<materialsCitation collectingDate="1999-05-22" collectorName="S. Nomura" elevation="1200" location="Mt. Pia Oac" municipality="Cao Bang Prov." specimenCount="1" specimenCount-female="1">
<specimenCount type="female">1 female</specimenCount>
(NSMT), &quot;
<location LSID="urn:lsid:plazi:treatment:079B2E1C8B25532C8E8654CF99A0156F:90EA5C5FB7AF0C4737D0EB0BA69639B6" municipality="Cao Bang Prov." name="Mt. Pia Oac">Mt. Pia Oac</location>
(LT:
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">1,200m</elevation>
</quantity>
)
<collectingMunicipality>Cao Bang Prov.</collectingMunicipality>
[N-Vietnam]
<collectingDate value="1999-05-22">22. V. 1999</collectingDate>
,
<collectorName>S. Nomura</collectorName>
leg.&quot;
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="diagnosis">
<paragraph pageId="0" pageNumber="127">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="127">
This is a distinct species in the genus by having the following characteristics: BA, DA, PA in the form of tubercles; HA carinate; apex of elytra rounded. In general appearance, this species is similar to
<taxonomicName class="Insecta" family="Endomychidae" genus="Amphisternus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Amphisternus sordidus" order="Coleoptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="sordidus">
<emphasis italics="true" pageId="0" pageNumber="127">Amphisternus sordidus</emphasis>
</taxonomicName>
Arrow, 1928 known from Vietnam and Laos, but differs from it by the mat and smooth dorsal surface (shiny and rugose in
<taxonomicName lsidName="A. sordidus" pageId="0" pageNumber="127" rank="species" species="sordidus">
<emphasis italics="true" pageId="0" pageNumber="127">A. sordidus</emphasis>
</taxonomicName>
), widely carinate HA (narrowly carinate in
<taxonomicName lsidName="A. sordidus" pageId="0" pageNumber="127" rank="species" species="sordidus">
<emphasis italics="true" pageId="0" pageNumber="127">A. sordidus</emphasis>
</taxonomicName>
), and BA in the form of a tubercle (carinate in
<taxonomicName lsidName="A. sordidus" pageId="0" pageNumber="127" rank="species" species="sordidus">
<emphasis italics="true" pageId="0" pageNumber="127">A. sordidus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="redescription">
<paragraph pageId="0" pageNumber="127">Redescription.</paragraph>
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" pageId="0" pageNumber="127">Female.</emphasis>
Body (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Cacodaemon spp. A-D C. laotinus laotinus, Vietnamese specimens (A, B) and Laotian specimens (C, D) E C. vietnamensis sp. nov., holotype F C. proavus A, B male C, D-F female. Scales: 1.0 mm." figureDoi="10.3897/zookeys.1081.75927.figure1" httpUri="https://binary.pensoft.net/fig/634063" pageId="0" pageNumber="127">1F</figureCitation>
) oval, slightly convex dorsally, weakly shiny. Coloration of body black, but DA and PA faintly dull orange.
</paragraph>
<paragraph pageId="0" pageNumber="127">
Head moderate in size, finely punctate. Antennae long, relatively stout; antennomere 3 longest, shorter than antennomeres 4 and 5 combined; club (antennomeres 9-11) weakly widened. Pronotum indistinctly and finely punctate, microreticulate, widest at middle, widely upturned in lateral parts; front corners triangular, minutely pointed at apices; lateral margins arcuate; posterior corners right-angled; PLM/PLS 0.75; PWM/PWA 1.52; PWM/PLM 1.80; PWM/PLS 1.35. Fore tibia (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Fore tibiae of Cacodaemon spp. A-D C. laotinus laotinus, Vietnamese specimens (A, B) and Laotian specimens (C, D) E C. vietnamensis sp. nov., holotype F C. proavus A, C male B, D-F female. Scale: 1.0 mm." figureDoi="10.3897/zookeys.1081.75927.figure2" httpUri="https://binary.pensoft.net/fig/634064" pageId="0" pageNumber="127">2F</figureCitation>
) straight, relatively stout. Elytra (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Elytral appendages of Cacodaemon spp. in lateral view A-D C. laotinus laotinus, Vietnamese specimens (A, B) and Laotian specimens (C, D) E C. vietnamensis sp. nov., holotype F C. proavus A, C male B, D-F female. Scale: 1.0 mm." figureDoi="10.3897/zookeys.1081.75927.figure3" httpUri="https://binary.pensoft.net/fig/634065" pageId="0" pageNumber="127">3F</figureCitation>
) minutely and sparsely punctate, microreticulate; basal and lateral grooves shallow; BA in the form of a tubercle; HA carinate; DA tubercle, small and low; PA tubercle, small and low; apex of elytra rounded; EL/EWH 0.99; EL/PLM 2.66; EWH/PWM 1.49; EWH/EWM 1.21; TL/EWH 1.37.
</paragraph>
<paragraph pageId="0" pageNumber="127">
Sternite VIII (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Female genitalia of Cacodaemon spp. A, B C. laotinus laotinus C, D C. vietnamensis sp. nov. E, F C. proavus A, C, E sternite VIII B, D, F ovipositor. Scale: 0.5 mm." figureDoi="10.3897/zookeys.1081.75927.figure5" httpUri="https://binary.pensoft.net/fig/634067" pageId="0" pageNumber="127">5E</figureCitation>
) shallowly concave at posterior margin. Ovipositor (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Female genitalia of Cacodaemon spp. A, B C. laotinus laotinus C, D C. vietnamensis sp. nov. E, F C. proavus A, C, E sternite VIII B, D, F ovipositor. Scale: 0.5 mm." figureDoi="10.3897/zookeys.1081.75927.figure5" httpUri="https://binary.pensoft.net/fig/634067" pageId="0" pageNumber="127">5F</figureCitation>
) bearing short setae along lateral margins of fused coxites; posterior margin of fused coxites weakly protruding; styli bearing short setae.
</paragraph>
<caption doi="10.3897/zookeys.1081.75927.figure5" httpUri="https://binary.pensoft.net/fig/634067" pageId="0" pageNumber="127" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" pageId="0" pageNumber="127">Figure 5.</emphasis>
Female genitalia of
<taxonomicName authorityName="Thomson" authorityYear="1857" class="Insecta" family="Endomychidae" genus="Cacodaemon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cacodaemon" order="Coleoptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="127">Cacodaemon</emphasis>
</taxonomicName>
spp.
<emphasis bold="true" pageId="0" pageNumber="127">A, B</emphasis>
<taxonomicName authority="laotinus" lsidName="C. laotinus subsp. laotinus" pageId="0" pageNumber="127" rank="subSpecies" species="laotinus" subSpecies="laotinus">
<emphasis italics="true" pageId="0" pageNumber="127">C. laotinus laotinus</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="127">C, D</emphasis>
<taxonomicName lsidName="C. vietnamensis" pageId="0" pageNumber="127" rank="species" species="vietnamensis">
<emphasis italics="true" pageId="0" pageNumber="127">C. vietnamensis</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="127">E, F</emphasis>
<taxonomicName lsidName="C. proavus" pageId="0" pageNumber="127" rank="species" species="proavus">
<emphasis italics="true" pageId="0" pageNumber="127">C. proavus</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="127">A, C, E</emphasis>
sternite VIII
<emphasis bold="true" pageId="0" pageNumber="127">B, D, F</emphasis>
ovipositor. Scale: 0.5 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" pageId="0" pageNumber="127">Male.</emphasis>
Not examined. Male genitalia figured by
<bibRefCitation author="Strohecker, HF" journalOrPublisher="Pacific Insects" pageId="0" pageNumber="127" pagination="319 - 357" refId="B7" refString="Strohecker, HF, 1964. A synopsis of the Amphisternini (Coleoptera: Endomychidae). Pacific Insects 6: 319 - 357" title="A synopsis of the Amphisternini (Coleoptera: Endomychidae)." volume="6" year="1964">Strohecker (1964)</bibRefCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" pageId="0" pageNumber="127">Measurement.</emphasis>
Female (
<emphasis italics="true" pageId="0" pageNumber="127">N</emphasis>
= 1). TL 7.43 mm; PWM 3.65 mm; PWA 2.40 mm; PLM 2.03 mm; PLS 2.70 mm; EL 5.40 mm; EWH 5.43 mm; EWM 4.50 mm.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="distribution">
<paragraph pageId="0" pageNumber="127">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="127">Vietnam.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="remarks">
<paragraph pageId="0" pageNumber="127">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="127">
<bibRefCitation author="Strohecker, HF" journalOrPublisher="Pacific Insects" pageId="0" pageNumber="127" pagination="319 - 357" refId="B7" refString="Strohecker, HF, 1964. A synopsis of the Amphisternini (Coleoptera: Endomychidae). Pacific Insects 6: 319 - 357" title="A synopsis of the Amphisternini (Coleoptera: Endomychidae)." volume="6" year="1964">Strohecker (1964)</bibRefCitation>
described this species based on three specimens collected from &quot;Mauson, Tonkin&quot; (= Mt. Mauson, Loc Binh District, Lang Son Province, Northeastern Vietnam).
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>