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<document id="8D82D28C45D45C8A75DD9F7553F7716D" ID-CLB-Dataset="7654" ID-DOI="10.5852/ejt.2021.739.1275" ID-GBIF-Dataset="a4c1ff18-1aa7-4dde-a8d0-2b0739cd4621" ID-ISSN="2118-9773" ID-Zenodo-Dep="4620965" ID-ZooBank="27DF46F5-C89A-45E9-87C9-9EF5C90CFAC1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616145751554" checkinUser="plazi" docAuthor="Matsumoto, Keita" docDate="2021" docId="F035D85E990FFFF9816CFA1DFBF054A6" docLanguage="en" docName="ejt-739_168-184.pdf" docOrigin="European Journal of Taxonomy 739" docSource="http://dx.doi.org/10.5852/ejt.2021.739.1275" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Caccothryptus tardarsauceae Matsumoto 2021, sp. nov." docType="treatment" docUuid="3DAF222E-1DF9-4254-BF38-627576985EA4" docUuidSource="ZooBank" docVersion="7" lastPageNumber="180" masterDocId="0C0CA0269905FFF58355FFA4FFC1525C" masterDocTitle="Six new species of the genus Caccothryptus from the Himalayas (Coleoptera: Limnichidae)" masterLastPageNumber="184" masterPageNumber="168" pageNumber="178" updateTime="1698934364920" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="2ABC402C1ED86BE3AF0F8F3DBE1F6388">Six new species of the genus Caccothryptus from the Himalayas (Coleoptera: Limnichidae)</mods:title>
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<mods:namePart id="81D0F08B61731A739EF3A110B67A80F9">Matsumoto, Keita</mods:namePart>
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<mods:affiliation id="706F03ED33E18090EF5E520D79DDD38F">Department of Life Sciences, Natural History Museum, London, SW 7 5 BD, UK.</mods:affiliation>
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<mods:title id="CE52F5C79A84FCB49DB90688729FBACF">European Journal of Taxonomy</mods:title>
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<mods:date id="CB701E8465167E886D4C11B8AFD91F76">2021</mods:date>
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<paragraph id="78236948990FFFFF816CFA1DFC3A5789" blockId="10.[372,1216,1465,1563]" box="[569,1019,1465,1493]" pageId="10" pageNumber="178">
<heading id="236BDE24990FFFFF816CFA1DFC3A5789" box="[569,1019,1465,1493]" centered="true" fontSize="11" level="2" pageId="10" pageNumber="178" reason="2">
<taxonomicName id="BF9C12CB990FFFFF816CFA1DFC585788" authority="Matsumoto, 2021" authorityName="Matsumoto" authorityYear="2021" box="[569,921,1465,1492]" class="Insecta" family="Limnichidae" genus="Caccothryptus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="178" phylum="Arthropoda" rank="species" species="tardarsauceae" status="sp. nov.">
<emphasis id="4AE8B55A990FFFFF816CFA1DFC585788" bold="true" box="[569,921,1465,1492]" italics="true" pageId="10" pageNumber="178">Caccothryptus tardarsauceae</emphasis>
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<taxonomicNameLabel id="51DB0821990FFFFF80F4FA1FFC3A5789" box="[929,1019,1467,1493]" pageId="10" pageNumber="178" rank="species">sp. nov.</taxonomicNameLabel>
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<figureCitation id="E0A775CD990FFFFF818BF9A4FCE95447" box="[734,808,1536,1563]" captionStart="Fig" captionStartId="11.[190,233,1963,1989]" captionTargetBox="[259,1333,685,1911]" captionTargetId="figure-67@11.[235,1353,658,1928]" captionTargetPageId="11" captionText="Fig. 5. Caccothryptus tardarsauceae sp. nov., ♂, holotype (NHMUK011225274). A. Dorsal habitus. B. Genitalia, dorsal. C. Genitalia, lateral. D. Genitalia, ventral." figureDoi="http://doi.org/10.5281/zenodo.4620981" httpUri="https://zenodo.org/record/4620981/files/figure.png" pageId="10" pageNumber="178">
Figs
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,
<figureCitation id="E0A775CD990FFFFF8060F9A5FC975447" box="[821,854,1537,1563]" captionStart="Fig" captionStartId="15.[189,232,1892,1918]" captionTargetBox="[356,1263,697,1831]" captionTargetId="figure-59@15.[292,1296,669,1860]" captionTargetPageId="15" captionText="Fig. 7. Close-up of Caccothryptus spp. genitalia ventral apex, ♂, holotype. A. C. brendelli sp. nov. (NHMUK011225276). B. C. arakawae sp. nov. (NHMUK011225268). C. C. championi sp. nov. (NHMUK011225382). D. C. larryi sp. nov. (NHMUK011225324). E. C. tardarsauceae sp. nov. (NHMUK011225274). F. C. abboti sp. nov. (NHMUK011225275)." figureDoi="http://doi.org/10.5281/zenodo.4620985" httpUri="https://zenodo.org/record/4620985/files/figure.png" pageId="10" pageNumber="178">7E</figureCitation>
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<heading id="236BDE24990FFFFF83E8F9E6FEF75400" bold="true" box="[189,310,1602,1628]" fontSize="11" level="3" pageId="10" pageNumber="178" reason="3">
<emphasis id="4AE8B55A990FFFFF83E8F9E6FEF75400" bold="true" box="[189,310,1602,1628]" pageId="10" pageNumber="178">Diagnosis</emphasis>
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<taxonomicName id="BF9C12CB990FFFFF83E8F9D4FDD654D6" authority="Matsumoto, 2021" authorityName="Matsumoto" authorityYear="2021" box="[189,535,1648,1674]" class="Insecta" family="Limnichidae" genus="Caccothryptus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="178" phylum="Arthropoda" rank="species" species="tardarsauceae" status="sp. nov.">
<emphasis id="4AE8B55A990FFFFF83E8F9D4FDD654D6" box="[189,535,1648,1674]" italics="true" pageId="10" pageNumber="178">Caccothryptus tardarsauceae</emphasis>
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<taxonomicNameLabel id="51DB0821990FFFFF8148F9D5FDB654D7" box="[541,631,1649,1675]" pageId="10" pageNumber="178" rank="species">sp. nov.</taxonomicNameLabel>
is within the
<taxonomicName id="BF9C12CB990FFFFF8047F9D5FC4954D6" box="[786,904,1648,1675]" class="Insecta" family="Limnichidae" genus="Caccothryptus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="178" phylum="Arthropoda" rank="species" species="testudo">
<emphasis id="4AE8B55A990FFFFF8047F9D5FC4954D6" box="[786,904,1648,1675]" italics="true" pageId="10" pageNumber="178">C. testudo</emphasis>
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group and can be distinguished by several genital characters: (1) apex of median lobe narrow with tip slightly pointed in ventral view; (2) concave structure between apical emargination narrowly V-shaped with gently pointed end; (3) dorsal side of parameres bisinuate in lateral view; (4) apical emargination of parameres V-shaped in ventral view.
</paragraph>
</subSubSection>
<subSubSection id="30863AC3990FFFFF83E8F8B8FF3755D4" pageId="10" pageNumber="178" type="etymology">
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<emphasis id="4AE8B55A990FFFFF83E8F8B8FE85556A" bold="true" box="[189,324,1820,1846]" pageId="10" pageNumber="178">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="78236948990FFFFF83E8F8EEFF3755D4" blockId="10.[189,1399,1866,1928]" pageId="10" pageNumber="178">This species is named after the American internet celebrity cat ʻTardar Sauceʼ, nicknamed ʻGrumpy Catʼ.</paragraph>
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<emphasis id="4AE8B55A990EFFFE83E8FEACFE6B537E" bold="true" box="[189,426,264,290]" pageId="11" pageNumber="179">Material examined</emphasis>
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<materialsCitation id="C8F46315990EFFFE83BFFE93FDEA53C4" ID-GBIF-Occurrence="3058402311" collectorName="H. G. Champion" country="India" elevation="1829" location="Jhatingri" pageId="11" pageNumber="179" specimenCount="1" stateProvince="Punjab" typeStatus="holotype">
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<typeStatus id="A727D7EA990EFFFE83BFFE93FE9D530D" box="[234,348,311,337]" pageId="11" pageNumber="179" type="holotype">Holotype</typeStatus>
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; “
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, Mandi.
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.
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H.G.C. //
<collectorName id="D5690C9E990EFFFE80EAFEFEFB455328" box="[959,1156,346,372]" pageId="11" pageNumber="179">H.G. Champion</collectorName>
B.M. 1953-156 // NHMUK011225274”;
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.
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<emphasis id="4AE8B55A990EFFFE83E8FE1BFE8F5385" bold="true" box="[189,334,447,473]" pageId="11" pageNumber="179">Description</emphasis>
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<emphasis id="4AE8B55A990EFFFE83BFFE4AFEEB5054" bold="true" box="[234,298,494,520]" pageId="11" pageNumber="179">Male</emphasis>
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BODY. Elongate, oval, dorsum convex, EL:
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, EW:
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, PL:
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, PW:
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. Colour: dorsal side brown; venter brown, tibia reddish brown, tarsal claws light brown. Dorsal surface
</paragraph>
<caption id="2CE339C0990EFFFE83EBF80FFC5C55B4" ID-DOI="http://doi.org/10.5281/zenodo.4620981" ID-Zenodo-Dep="4620981" httpUri="https://zenodo.org/record/4620981/files/figure.png" pageId="11" pageNumber="179" startId="11.[190,233,1963,1989]" targetBox="[259,1333,685,1911]" targetPageId="11">
<paragraph id="78236948990EFFFE83EBF80FFC5C55B4" blockId="11.[189,1399,1963,2025]" pageId="11" pageNumber="179">
<emphasis id="4AE8B55A990EFFFE83EBF80FFECF5599" bold="true" box="[190,270,1963,1989]" pageId="11" pageNumber="179">Fig. 5.</emphasis>
<taxonomicName id="BF9C12CB990EFFFE8242F80FFDB55599" authority="Matsumoto, 2021" authorityName="Matsumoto" authorityYear="2021" box="[279,628,1963,1989]" class="Insecta" family="Limnichidae" genus="Caccothryptus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="179" phylum="Arthropoda" rank="species" species="tardarsauceae" status="sp. nov.">
<emphasis id="4AE8B55A990EFFFE8242F80FFDB55599" box="[279,628,1963,1989]" italics="true" pageId="11" pageNumber="179">Caccothryptus tardarsauceae</emphasis>
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<taxonomicNameLabel id="51DB0821990EFFFE812AF808FD1F559A" box="[639,734,1964,1990]" pageId="11" pageNumber="179" rank="species">sp. nov.</taxonomicNameLabel>
, ♂, holotype (NHMUK011225274).
<emphasis id="4AE8B55A990EFFFE87C9F80FFB705599" bold="true" box="[1180,1201,1963,1989]" pageId="11" pageNumber="179">A</emphasis>
. Dorsal habitus.
<emphasis id="4AE8B55A990EFFFE83E8F86AFF1055B4" bold="true" box="[189,209,1998,2024]" pageId="11" pageNumber="179">B</emphasis>
. Genitalia, dorsal.
<emphasis id="4AE8B55A990EFFFE82E5F86AFE0455B4" bold="true" box="[432,453,1998,2024]" pageId="11" pageNumber="179">C</emphasis>
. Genitalia, lateral.
<emphasis id="4AE8B55A990EFFFE81F3F86AFD7A55B4" bold="true" box="[678,699,1998,2024]" pageId="11" pageNumber="179">D</emphasis>
. Genitalia, ventral.
</paragraph>
</caption>
<paragraph id="782369489909FFF983E8FEADFE71531B" blockId="12.[188,1398,265,327]" pageId="12" pageNumber="180">densely covered with long setae with patches of white setae forming a wavy pattern, ventral surface with relatively long setae.</paragraph>
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DORSUM. Head: partially retracted into pronotum; overall surface smooth, densely punctured across surface. Antennae: 11 segmented, filiform, fully pubescent, antennomeres long and slender, 1
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widest, 2
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narrow and long, 3
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<collectingCountry id="008B29D89909FFF98123FE10FD5A5392" box="[630,667,436,462]" name="American Samoa" pageId="12" pageNumber="180">AS</collectingCountry>
smallest, 4
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11
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long, thin and obtusely angled on one side. Pronotum: transverse, widest at posterior end, narrowest at anterior end, narrower than elytra; anterior margin convex, posterior margin trisinuate, lateral margin convex; surface covered with dense punctation, surface between punctures smooth and shiny. Scutellum: subtriangular, large, lateral sides gently convex. Elytra: longer than wide; anterior margin slightly convex, sinuate; anterior lateral angle nearly right angled, pointed; lateral margins strongly bordered; surface with mixtures of small fine sized punctuations and medium sized punctuations forming irregular near the elytral suture; elytral striae more clearly visible towards lateral end; surface between punctures smooth and shiny.
</paragraph>
<paragraph id="782369489909FFF983E8FD48FEC051A1" blockId="12.[187,1401,748,1021]" pageId="12" pageNumber="180">
VENTER. Prosternum: concave on anterior margin, outer lateral margin concave; prosternal process as long as wide, lateral margin sinuated, posterior margin rounded. Mesoventrite: wider than long, narrowed towards posterior; anterior end concave where prosternal process fits, discrimen visible; laterally subparallel and gently curved, posteriorly flat, anteriorly trisinuate. Metaventrite: long and wide, flat on the middle; discrimen thin visible across each end. Abdomen: five clearly visible ventrites. Intercoxal process with pointed apex. Posterior margin of 1
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and 2
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ventrites convex, 3
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4
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ventrites slightly concave around the middle, 5
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ventrite semioval. Surface covered with long, light yellow, recumbent setae.
</paragraph>
<paragraph id="782369489909FFF983E8FB80FE9756FB" blockId="12.[189,1399,1060,1191]" pageId="12" pageNumber="180">
LEGS. Tibia: protibia with outer and inner lateral margin curved. Meso- and metatibia with outer lateral margin almost straight, inner lateral margin straight. Tarsi: simple, segments increase in length towards the apex, 5
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tarsal segment as long as 1
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4
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tarsal segment combined. Tarsal claws: long, narrow, symmetrical.
</paragraph>
<paragraph id="782369489909FFF983E8FB6BFC725479" blockId="12.[188,1400,1231,1574]" pageId="12" pageNumber="180">GENITALIA. Aedeagus: elongate, strongly sclerotised. Base: asymmetrical, widest around basal 0.2 from both lateral and ventral view. Median lobe: approximately ¼ length of genitalia; longer than parameres; gently curved on dorsal side, slightly bisinuate on ventral side, widest at middle, narrowing towards the apex, apex with round point in lateral view; apex pointed, split into two on anterior half and edge pointed, lower half of lateral end relatively straight in dorsal view; apical end smooth and narrows inwards in ventral view. Parameres: wider than median lobe in ventral view; apical emargination wide V-shaped, concave structure between parameres elongated and wide V-shaped, with gently pointed end; bisinuate, lower half of sinuation long and shallow, upper half short and slightly deep on dorsal side in lateral view; gently curved on ventral side in lateral view; apex pointed and overlaps with median lobe, majority of parameres overlaps with median lobe in lateral view.</paragraph>
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<emphasis id="4AE8B55A9909FFF983BFF9E9FE84543B" bold="true" box="[234,325,1613,1639]" pageId="12" pageNumber="180">Female</emphasis>
</paragraph>
<paragraph id="782369489909FFF983E8F9D4FEF854D6" blockId="12.[189,325,1613,1674]" box="[189,313,1648,1674]" pageId="12" pageNumber="180">Unknown.</paragraph>
</subSubSection>
<subSubSection id="30863AC39909FFF983E8F915FBF054A6" box="[189,1073,1713,1787]" pageId="12" pageNumber="180" type="distribution">
<paragraph id="782369489909FFF983E8F915FE975497" blockId="12.[189,342,1713,1739]" box="[189,342,1713,1739]" pageId="12" pageNumber="180">
<heading id="236BDE249909FFF983E8F915FE975497" bold="true" box="[189,342,1713,1739]" fontSize="11" level="3" pageId="12" pageNumber="180" reason="3">
<emphasis id="4AE8B55A9909FFF983E8F915FE975497" bold="true" box="[189,342,1713,1739]" pageId="12" pageNumber="180">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="782369489909FFF983E8F944FBF054A6" blockId="12.[189,1073,1760,1787]" box="[189,1073,1760,1787]" pageId="12" pageNumber="180">
<taxonomicName id="BF9C12CB9909FFF983E8F944FDD654A6" authority="Matsumoto, 2021" authorityName="Matsumoto" authorityYear="2021" box="[189,535,1760,1786]" class="Insecta" family="Limnichidae" genus="Caccothryptus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="180" phylum="Arthropoda" rank="species" species="tardarsauceae" status="sp. nov.">
<emphasis id="4AE8B55A9909FFF983E8F944FDD654A6" box="[189,535,1760,1786]" italics="true" pageId="12" pageNumber="180">Caccothryptus tardarsauceae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="51DB08219909FFF9814AF945FDBB54A7" box="[543,634,1761,1787]" pageId="12" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
is known only from the
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locality.
</paragraph>
</subSubSection>
</treatment>
</document>