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<mods:title id="A0A43B08574C7A62BB3CD333EAC0A707">Johnwellsia, a new intertidal genus of Parastenheliidae (Copepoda, Harpacticoida) from the Taiwan Strait, China, including a review of the family and key to genera</mods:title>
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<paragraph id="1D0E36FCFFECFFB1FF51D276E5AEFE53" blockId="47.[151,494,366,393]" box="[151,494,366,393]" pageId="47" pageNumber="283">
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<taxonomicName id="DAB14D7FFFECFFB1FF51D276E5AEFE53" authority="Monard, 1935 a" authorityName="Monard" authorityYear="1935" box="[151,494,366,393]" class="Hexanauplia" family="Parastenheliidae" genus="Thalestrella" kingdom="Animalia" order="Harpacticoida" pageId="47" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="2FC5EAEEFFECFFB1FF51D276E5AEFE53" bold="true" box="[151,494,366,393]" pageId="47" pageNumber="283">
<emphasis id="2FC5EAEEFFECFFB1FF51D276E568FE52" bold="true" box="[151,296,366,393]" italics="true" pageId="47" pageNumber="283">Thalestrella</emphasis>
<bibRefCitation id="79204B0DFFECFFB1FEF6D277E5AEFE53" author="Monard, A." box="[304,494,366,393]" pageId="47" pageNumber="283" pagination="5 - 88" refId="ref76147" refString="Monard, A. (1935 a) Etude sur la faune des Harpacticoides marins de Roscoff. Travaux de la Station biologique de Roscoff, 13, 5 - 88, figs. 1 - 235." type="journal article" year="1935">Monard, 1935a</bibRefCitation>
</emphasis>
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<taxonomicName id="DAB14D7FFFECFFB1FF51D2ABE5CCFE10" authority="Noodt, 1964" authorityName="Noodt" authorityYear="1964" box="[151,396,435,459]" class="Hexanauplia" family="Parastenheliidae" genus="Karllangia" kingdom="Animalia" order="Harpacticoida" pageId="47" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="2FC5EAEEFFECFFB1FF51D2ABE546FE10" box="[151,262,435,459]" italics="true" pageId="47" pageNumber="283">Karllangia</emphasis>
<bibRefCitation id="79204B0DFFECFFB1FECBD2ABE5CCFE10" author="Noodt, W." box="[269,396,435,459]" pageId="47" pageNumber="283" pagination="128 - 154" refId="ref76696" refString="Noodt, W. (1964) Copepoda Harpacticoidea aus dem Litoral des Roten Meeres. Kieler Meeresforschungen, 20, Sonderheft, 128 - 154." type="journal article" year="1964">Noodt, 1964</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="1D0E36FCFFECFFB1FF51D2E1E0C5FD7F" blockId="47.[151,1437,505,2008]" pageId="47" pageNumber="283">
<bibRefCitation id="79204B0DFFECFFB1FF51D2E1E539FDCF" author="Monard, A." box="[151,377,505,532]" pageId="47" pageNumber="283" pagination="5 - 88" refId="ref76147" refString="Monard, A. (1935 a) Etude sur la faune des Harpacticoides marins de Roscoff. Travaux de la Station biologique de Roscoff, 13, 5 - 88, figs. 1 - 235." type="journal article" year="1935">Monard (1935a: 45)</bibRefCitation>
described
<taxonomicName id="DAB14D7FFFECFFB1FE34D2E1E743FDC8" authorityName="Monard." authorityYear="1935" baseAuthorityName="sensu Por" baseAuthorityYear="1935" box="[498,771,505,531]" class="Hexanauplia" family="Parastenheliidae" genus="Thalestrella" kingdom="Animalia" order="Harpacticoida" pageId="47" pageNumber="283" phylum="Arthropoda" rank="species" species="ornatissima">
<emphasis id="2FC5EAEEFFECFFB1FE34D2E1E743FDC8" box="[498,771,505,531]" italics="true" pageId="47" pageNumber="283">Thalestrella ornatissima</emphasis>
</taxonomicName>
as the
<typeStatus id="C20A885EFFECFFB1FC94D2E2E7C2FDCF" box="[850,898,506,532]" pageId="47" pageNumber="283">type</typeStatus>
species of his new genus
<taxonomicName id="DAB14D7FFFECFFB1FB63D2E1E16BFDC8" authorityName="Monard" authorityYear="1935" box="[1189,1323,505,531]" class="Hexanauplia" family="Parastenheliidae" genus="Thalestrella" kingdom="Animalia" order="Harpacticoida" pageId="47" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="2FC5EAEEFFECFFB1FB63D2E1E16BFDC8" box="[1189,1323,505,531]" italics="true" pageId="47" pageNumber="283">Thalestrella</emphasis>
</taxonomicName>
, based on females dredged from sand near Roch Iliévec in the Roscoff area (
<collectingRegion id="DF75F81EFFECFFB1FCBFD105E79BFDE3" box="[889,987,541,568]" country="France" name="Bretagne" pageId="47" pageNumber="283">Brittany</collectingRegion>
).
<bibRefCitation id="79204B0DFFECFFB1FC2AD106E02FFDEC" author="Lang, K." box="[1004,1135,541,568]" pageId="47" pageNumber="283" refId="ref75245" refString="Lang, K. (1944) Monographie der Harpacticiden (Vorlaufige Mitteilung). Almqvist &amp; Wiksells Boktryckeri Ab, Uppsala, 39 pp." type="book" year="1944">Lang (1944</bibRefCitation>
,
<bibRefCitation id="79204B0DFFECFFB1FBBFD105E0F2FDEC" author="Lang, K." box="[1145,1202,541,567]" pageId="47" pageNumber="283" refId="ref75271" refString="Lang, K. (1948) Monographie der Harpacticiden. Vols. 1 - 2. Hakan Ohlsson, Lund, 1682 pp." type="book" year="1948">1948</bibRefCitation>
) rejected the validity of
<taxonomicName id="DAB14D7FFFECFFB1FF73D159E57BFD80" authorityName="Monard" authorityYear="1935" box="[181,315,577,603]" class="Hexanauplia" family="Parastenheliidae" genus="Thalestrella" kingdom="Animalia" order="Harpacticoida" pageId="47" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="2FC5EAEEFFECFFB1FF73D159E57BFD80" box="[181,315,577,603]" italics="true" pageId="47" pageNumber="283">Thalestrella</emphasis>
</taxonomicName>
and transferred
<taxonomicName id="DAB14D7FFFECFFB1FE34D15AE6D2FD80" authorityName="Monard." authorityYear="1935" baseAuthorityName="sensu Por" baseAuthorityYear="1935" box="[498,658,578,603]" class="Hexanauplia" family="Parastenheliidae" genus="Thalestrella" kingdom="Animalia" order="Harpacticoida" pageId="47" pageNumber="283" phylum="Arthropoda" rank="species" species="ornatissima">
<emphasis id="2FC5EAEEFFECFFB1FE34D15AE6D2FD80" box="[498,658,578,603]" italics="true" pageId="47" pageNumber="283">T. ornatissima</emphasis>
</taxonomicName>
to
<taxonomicName id="DAB14D7FFFECFFB1FD70D159E713FD80" authorityName="Sewell" authorityYear="1940" box="[694,851,577,603]" class="Hexanauplia" family="Parastenheliidae" genus="Parastenhelia" kingdom="Animalia" order="Harpacticoida" pageId="47" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="2FC5EAEEFFECFFB1FD70D159E713FD80" box="[694,851,577,603]" italics="true" pageId="47" pageNumber="283">Parastenhelia</emphasis>
</taxonomicName>
, claiming a close relationship with
<taxonomicName id="DAB14D7FFFECFFB1FB24D15AE111FD80" authorityName="sensu Wells" authorityYear="1961" baseAuthorityName="Norman &amp; Scott" baseAuthorityYear="1905" box="[1250,1361,577,603]" class="Hexanauplia" family="Parastenheliidae" genus="Parastenhelia" kingdom="Animalia" order="Harpacticoida" pageId="47" pageNumber="283" phylum="Arthropoda" rank="species" species="anglica">
<emphasis id="2FC5EAEEFFECFFB1FB24D15AE111FD80" box="[1250,1361,577,603]" italics="true" pageId="47" pageNumber="283">P. anglica</emphasis>
</taxonomicName>
and
<taxonomicName id="DAB14D7FFFECFFB1FA4ED15AE4B0FDA4" authorityName="Thompson &amp; Scott" authorityYear="1903" class="Hexanauplia" family="Parastenheliidae" genus="Parastenhelia" kingdom="Animalia" order="Harpacticoida" pageId="47" pageNumber="283" phylum="Arthropoda" rank="species" species="hornelli">
<emphasis id="2FC5EAEEFFECFFB1FA4ED15AE4B0FDA4" italics="true" pageId="47" pageNumber="283">P. hornelli</emphasis>
</taxonomicName>
.
<bibRefCitation id="79204B0DFFECFFB1FF3BD17DE53BFD5B" author="Gee, J. M." box="[253,379,613,640]" pageId="47" pageNumber="283" pagination="2611 - 2651" refId="ref71824" refString="Gee, J. M. (2006) Parastenheliidae (Copepoda: Harpacticoida) from the Isles of Scilly. Journal of natural History, 40, 2611 - 2651. https: // doi. org / 10.1080 / 00222930601108194" type="journal article" year="2006">Gee (2006)</bibRefCitation>
recently assigned the species to
<taxonomicName id="DAB14D7FFFECFFB1FD22D17DE71FFDA4" authorityName="Noodt" authorityYear="1964" box="[740,863,613,639]" class="Hexanauplia" family="Parastenheliidae" genus="Karllangia" kingdom="Animalia" order="Harpacticoida" pageId="47" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="2FC5EAEEFFECFFB1FD22D17DE71FFDA4" box="[740,863,613,639]" italics="true" pageId="47" pageNumber="283">Karllangia</emphasis>
</taxonomicName>
, repeatedly stating that the original description was based on only a single female but there is no evidence for that in
<bibRefCitation id="79204B0DFFECFFB1FCB7D191E07AFD7F" author="Monard, A." box="[881,1082,649,676]" pageId="47" pageNumber="283" pagination="5 - 88" refId="ref76147" refString="Monard, A. (1935 a) Etude sur la faune des Harpacticoides marins de Roscoff. Travaux de la Station biologique de Roscoff, 13, 5 - 88, figs. 1 - 235." type="journal article" year="1935">Monards (1935a)</bibRefCitation>
paper.
</paragraph>
</subSubSection>
<subSubSection id="55AB6577FFECFFB1FF01D1B5E576FA67" pageId="47" pageNumber="283" type="description">
<paragraph id="1D0E36FCFFECFFB1FF01D1B5E576FA67" blockId="47.[151,1437,505,2008]" pageId="47" pageNumber="283">
<bibRefCitation id="79204B0DFFECFFB1FF01D1B5E5FAFD13" author="Mielke, W." box="[199,442,685,712]" pageId="47" pageNumber="283" pagination="141 - 153" refId="ref75832" refString="Mielke, W. (1994 b) Two co-occuring new Karllangia species (Copepoda: Ameiridae) from the Caribbean coast of Costa Rica. Revista de Biologia tropical, 42, 141 - 153." type="journal article" year="1994">Mielke (1994b: 152)</bibRefCitation>
recognized two evolutionary lineages within
<taxonomicName id="DAB14D7FFFECFFB1FC1CD1B5E015FD1C" authorityName="Noodt" authorityYear="1964" box="[986,1109,685,711]" class="Hexanauplia" family="Parastenheliidae" genus="Karllangia" kingdom="Animalia" order="Harpacticoida" pageId="47" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="2FC5EAEEFFECFFB1FC1CD1B5E015FD1C" box="[986,1109,685,711]" italics="true" pageId="47" pageNumber="283">Karllangia</emphasis>
</taxonomicName>
, the more primitive branch comprising
<taxonomicName id="DAB14D7FFFECFFB1FED8D1CAE65EFD30" authority="arenicola" box="[286,542,721,747]" class="Hexanauplia" family="Parastenheliidae" genus="Thalestrella" kingdom="Animalia" order="Harpacticoida" pageId="47" pageNumber="283" phylum="Arthropoda" rank="subSpecies" species="arenicola" subSpecies="arenicola">
<emphasis id="2FC5EAEEFFECFFB1FED8D1CAE65EFD30" box="[286,542,721,747]" italics="true" pageId="47" pageNumber="283">K. arenicola arenicola</emphasis>
</taxonomicName>
and
<taxonomicName id="DAB14D7FFFECFFB1FD9ED1CAE6F8FD30" baseAuthorityName="Kunz" baseAuthorityYear="1975" box="[600,696,722,747]" class="Hexanauplia" family="Parastenheliidae" genus="Foweya" kingdom="Animalia" order="Harpacticoida" pageId="47" pageNumber="283" phylum="Arthropoda" rank="species" species="tertia">
<emphasis id="2FC5EAEEFFECFFB1FD9ED1CAE6F8FD30" box="[600,696,722,747]" italics="true" pageId="47" pageNumber="283">K. tertia</emphasis>
</taxonomicName>
, and the more advanced one represented by
<taxonomicName id="DAB14D7FFFECFFB1FB7FD1CAE133FD30" baseAuthorityName="Wells" baseAuthorityYear="1967" box="[1209,1395,721,747]" class="Hexanauplia" family="Parastenheliidae" genus="Thalestrella" kingdom="Animalia" order="Harpacticoida" pageId="47" pageNumber="283" phylum="Arthropoda" rank="species" species="psammophila">
<emphasis id="2FC5EAEEFFECFFB1FB7FD1CAE133FD30" box="[1209,1395,721,747]" italics="true" pageId="47" pageNumber="283">K. psammophila</emphasis>
</taxonomicName>
,
<taxonomicName id="DAB14D7FFFECFFB1FA45D1CAE5CEFCD4" authorityName="Wells &amp; Rao" authorityYear="1987" class="Hexanauplia" family="Parastenheliidae" genus="Thalestrella" kingdom="Animalia" order="Harpacticoida" pageId="47" pageNumber="283" phylum="Arthropoda" rank="subSpecies" species="arenicola" subSpecies="bengalensis">
<emphasis id="2FC5EAEEFFECFFB1FA45D1CAE5CEFCD4" italics="true" pageId="47" pageNumber="283">K. arenicola bengalensis</emphasis>
</taxonomicName>
and both
<collectingRegion id="DF75F81EFFECFFB1FDC2D1EDE604FCD4" box="[516,580,757,783]" country="Tanzania" name="Pwani" pageId="47" pageNumber="283">Costa</collectingRegion>
Rican species (
<taxonomicName id="DAB14D7FFFECFFB1FD3CD1EEE737FCD4" baseAuthorityName="Mielke" baseAuthorityYear="1994" box="[762,887,757,783]" class="Hexanauplia" family="Parastenheliidae" genus="Thalestrella" kingdom="Animalia" order="Harpacticoida" pageId="47" pageNumber="283" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="2FC5EAEEFFECFFB1FD3CD1EEE737FCD4" box="[762,887,757,783]" italics="true" pageId="47" pageNumber="283">K. obscura</emphasis>
</taxonomicName>
,
<taxonomicName id="DAB14D7FFFECFFB1FC41D1EEE041FCD4" baseAuthorityName="Mielke" baseAuthorityYear="1994" box="[903,1025,757,783]" class="Hexanauplia" family="Parastenheliidae" genus="Thalestrella" kingdom="Animalia" order="Harpacticoida" pageId="47" pageNumber="283" phylum="Arthropoda" rank="species" species="pulchra">
<emphasis id="2FC5EAEEFFECFFB1FC41D1EEE041FCD4" box="[903,1025,757,783]" italics="true" pageId="47" pageNumber="283">K. pulchra</emphasis>
</taxonomicName>
). As pointed out by
<bibRefCitation id="79204B0DFFECFFB1FB36D1EDE131FCCB" author="Gee, J. M." box="[1264,1393,757,784]" pageId="47" pageNumber="283" pagination="2611 - 2651" refId="ref71824" refString="Gee, J. M. (2006) Parastenheliidae (Copepoda: Harpacticoida) from the Isles of Scilly. Journal of natural History, 40, 2611 - 2651. https: // doi. org / 10.1080 / 00222930601108194" type="journal article" year="2006">Gee (2006)</bibRefCitation>
the outstanding problem within the genus
<taxonomicName id="DAB14D7FFFECFFB1FD81D001E682FCE8" authorityName="Noodt" authorityYear="1964" box="[583,706,793,819]" class="Hexanauplia" family="Parastenheliidae" genus="Karllangia" kingdom="Animalia" order="Harpacticoida" pageId="47" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="2FC5EAEEFFECFFB1FD81D001E682FCE8" box="[583,706,793,819]" italics="true" pageId="47" pageNumber="283">Karllangia</emphasis>
</taxonomicName>
is the fact that its
<typeStatus id="C20A885EFFECFFB1FC56D002E780FCEF" box="[912,960,794,820]" pageId="47" pageNumber="283">type</typeStatus>
species,
<taxonomicName id="DAB14D7FFFECFFB1FBE1D002E0F2FCE8" baseAuthorityName="Noodt" baseAuthorityYear="1964" box="[1063,1202,793,819]" class="Hexanauplia" family="Parastenheliidae" genus="Thalestrella" kingdom="Animalia" order="Harpacticoida" pageId="47" pageNumber="283" phylum="Arthropoda" rank="species" species="arenicola">
<emphasis id="2FC5EAEEFFECFFB1FBE1D002E0F2FCE8" box="[1063,1202,793,819]" italics="true" pageId="47" pageNumber="283">K. arenicola</emphasis>
</taxonomicName>
, apparently does not display the most important synapomorphy that supports the common ancestry of the remaining species (including the
<typeStatus id="C20A885EFFECFFB1FF04D07AE4B2FCA7" box="[194,242,866,892]" pageId="47" pageNumber="283">type</typeStatus>
species of
<taxonomicName id="DAB14D7FFFECFFB1FEB0D079E5BCFCA0" authorityName="Monard" authorityYear="1935" box="[374,508,865,891]" class="Hexanauplia" family="Parastenheliidae" genus="Thalestrella" kingdom="Animalia" order="Harpacticoida" pageId="47" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="2FC5EAEEFFECFFB1FEB0D079E5BCFCA0" box="[374,508,865,891]" italics="true" pageId="47" pageNumber="283">Thalestrella</emphasis>
</taxonomicName>
). This character state which distinguishes
<taxonomicName id="DAB14D7FFFECFFB1FC22D079E01FFCA0" authorityName="Noodt" authorityYear="1964" box="[996,1119,865,891]" class="Hexanauplia" family="Parastenheliidae" genus="Karllangia" kingdom="Animalia" order="Harpacticoida" pageId="47" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="2FC5EAEEFFECFFB1FC22D079E01FFCA0" box="[996,1119,865,891]" italics="true" pageId="47" pageNumber="283">Karllangia</emphasis>
</taxonomicName>
from other members of the
<taxonomicName id="DAB14D7FFFECFFB1FF51D09DE50FFC44" baseAuthorityName="Willen" baseAuthorityYear="2000" box="[151,335,901,927]" class="Hexanauplia" family="Parastenheliidae" kingdom="Animalia" order="Harpacticoida" pageId="47" pageNumber="283" phylum="Arthropoda" rank="family">Parastenheliidae</taxonomicName>
is the modification of the male antenna (allobasal seta much more plumose, exp-1 smaller and with only one strong, plumose seta, exp-2 broader with more strongly plumose setae than in
<specimenCount id="0BB7FD75FFECFFB1FBA3D0B2E039FC1F" box="[1125,1145,938,964]" pageId="47" pageNumber="283" type="female"></specimenCount>
). The original description by
<bibRefCitation id="79204B0DFFECFFB1FF7DD0D5E514FC33" author="Noodt, W." box="[187,340,973,1000]" pageId="47" pageNumber="283" pagination="128 - 154" refId="ref76696" refString="Noodt, W. (1964) Copepoda Harpacticoidea aus dem Litoral des Roten Meeres. Kieler Meeresforschungen, 20, Sonderheft, 128 - 154." type="journal article" year="1964">Noodt (1964)</bibRefCitation>
of
<taxonomicName id="DAB14D7FFFECFFB1FEBDD0D6E647FC3C" baseAuthorityName="Noodt" baseAuthorityYear="1964" box="[379,519,973,999]" class="Hexanauplia" family="Parastenheliidae" genus="Thalestrella" kingdom="Animalia" order="Harpacticoida" pageId="47" pageNumber="283" phylum="Arthropoda" rank="species" species="arenicola">
<emphasis id="2FC5EAEEFFECFFB1FEBDD0D6E647FC3C" box="[379,519,973,999]" italics="true" pageId="47" pageNumber="283">K. arenicola</emphasis>
</taxonomicName>
from coralline sands on the Egyptian coast of the
<collectingRegion id="DF75F81EFFECFFB1FB82D0D5E0E1FC3C" box="[1092,1185,973,999]" country="Egypt" name="Al Bahr al Ahmar" pageId="47" pageNumber="283">Red Sea</collectingRegion>
does not mention any sexual dimorphism in the male antenna which could be attributed to either an observational error or to genuine absence. This ambiguity surrounding the
<typeStatus id="C20A885EFFECFFB1FDB4D70EE6E2FBEB" box="[626,674,1046,1072]" pageId="47" pageNumber="283">type</typeStatus>
species has now been removed by synonymizing
<taxonomicName id="DAB14D7FFFECFFB1FB20D70DE121FBF4" authorityName="Noodt" authorityYear="1964" box="[1254,1377,1045,1071]" class="Hexanauplia" family="Parastenheliidae" genus="Karllangia" kingdom="Animalia" order="Harpacticoida" pageId="47" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="2FC5EAEEFFECFFB1FB20D70DE121FBF4" box="[1254,1377,1045,1071]" italics="true" pageId="47" pageNumber="283">Karllangia</emphasis>
</taxonomicName>
with its senior subjective synonym
<taxonomicName id="DAB14D7FFFECFFB1FE33D721E63BFB88" authorityName="Monard" authorityYear="1935" box="[501,635,1081,1107]" class="Hexanauplia" family="Parastenheliidae" genus="Thalestrella" kingdom="Animalia" order="Harpacticoida" pageId="47" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="2FC5EAEEFFECFFB1FE33D721E63BFB88" box="[501,635,1081,1107]" italics="true" pageId="47" pageNumber="283">Thalestrella</emphasis>
</taxonomicName>
. The presence of antennary sexual dimorphism in its
<typeStatus id="C20A885EFFECFFB1FB2AD722E15CFB8F" box="[1260,1308,1082,1108]" pageId="47" pageNumber="283">type</typeStatus>
species,
<taxonomicName id="DAB14D7FFFECFFB1FA4ED722E55BFBAC" authorityName="Monard." authorityYear="1935" baseAuthorityName="sensu Por" baseAuthorityYear="1935" class="Hexanauplia" family="Parastenheliidae" genus="Thalestrella" kingdom="Animalia" order="Harpacticoida" pageId="47" pageNumber="283" phylum="Arthropoda" rank="species" species="ornatissima">
<emphasis id="2FC5EAEEFFECFFB1FA4ED722E55BFBAC" italics="true" pageId="47" pageNumber="283">T. ornatissima</emphasis>
</taxonomicName>
, has been demonstrated by
<bibRefCitation id="79204B0DFFECFFB1FD88D745E68DFBA3" author="Gee, J. M." box="[590,717,1117,1144]" pageId="47" pageNumber="283" pagination="2611 - 2651" refId="ref71824" refString="Gee, J. M. (2006) Parastenheliidae (Copepoda: Harpacticoida) from the Isles of Scilly. Journal of natural History, 40, 2611 - 2651. https: // doi. org / 10.1080 / 00222930601108194" type="journal article" year="2006">Gee (2006)</bibRefCitation>
who confirmed a number of additional genus-level apomorphies in the species, including the compressed appearance of the distal antennulary segments in the female, the presence of only three setae on exp-2 of the antenna, two setae on the mandibular exopod, and one seta on the maxillipedal syncoxa; the absence of the inner seta on P2P4 exp-1; and, a male P5 exopod with only four setae. Additional apomorphies which were not observed by
<bibRefCitation id="79204B0DFFECFFB1FDB0D7F5E6B6FAD3" author="Gee, J. M." box="[630,758,1261,1288]" pageId="47" pageNumber="283" pagination="2611 - 2651" refId="ref71824" refString="Gee, J. M. (2006) Parastenheliidae (Copepoda: Harpacticoida) from the Isles of Scilly. Journal of natural History, 40, 2611 - 2651. https: // doi. org / 10.1080 / 00222930601108194" type="journal article" year="2006">Gee (2006)</bibRefCitation>
include (a) male antennule subchirocer, compound segment 5 (XIVXX) swollen; (b) the male antennule with only three segments distal to the geniculation (XXIXXIII, XXIVXXV, XXVIXXVIII); (c) ventral surface of segment 2 of male antennule with characteristic, backwardly directed, spiniform element, either displaying a serrate or flagellate tip (
<bibRefCitation id="79204B0DFFECFFB1FC19D641E0FEFAA8" author="Wells, J. B. J. &amp; Rao, G. C." box="[991,1214,1369,1396]" pageId="47" pageNumber="283" pagination="1 - 385" refId="ref82468" refString="Wells, J. B. J. &amp; Rao, G. C. (1987) Littoral Harpacticoida (Crustacea: Copepoda) from Andaman and Nicobar Islands. Memoirs of the zoological Survey of India, 16 (4), 1 - 385." type="journal article" year="1987">Wells &amp; Rao 1987</bibRefCitation>
: Fig. 110b;
<bibRefCitation id="79204B0DFFECFFB1FA8BD641E4A2FA4C" author="Mielke, W." pageId="47" pageNumber="283" pagination="141 - 153" refId="ref75832" refString="Mielke, W. (1994 b) Two co-occuring new Karllangia species (Copepoda: Ameiridae) from the Caribbean coast of Costa Rica. Revista de Biologia tropical, 42, 141 - 153." type="journal article" year="1994">Mielke 1994b</bibRefCitation>
:
<figureCitation id="858A2A79FFECFFB1FF2AD665E505FA43" box="[236,325,1405,1432]" captionStart="FIGURE 6" captionStartId="14.[151,250,1812,1836]" captionTargetBox="[249,1337,188,1787]" captionTargetPageId="14" captionText="FIGURE 6. Sexual dimorphism of P2 (AE) and P4 (FG) endopodal segmentation and armature within the Parastenheliidae. A, Foweya tertia (Kunz, 1975); B, Microthalestris costata (Pallares, 1982) comb. nov., M. variabilis sp. nov., Parastenhelia similis Thompson &amp; Scott, 1903, P. willemvervoorti sp. nov., Porirualia pyriformis (Song, Kim &amp; Chang, 2003) comb. nov.; C, Johnwellsia bipartita sp. nov.; D, Parastenhelia hornelli Thompson &amp; Scott, 1903 sensu Apostolov (1973); E, Parastenhelia oligochaeta Wells &amp; Rao, 1987; F, Microthalestris bulbosa (Gee, 2006) comb. nov., Parastenhelia willemvervoorti sp. nov.; G, Microthalestris minuta (Pallares, 1982) comb. nov., Penicillicaris penicillata sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5572433" httpUri="https://zenodo.org/record/5572433/files/figure.png" pageId="47" pageNumber="283">Fig. 6A</figureCitation>
); and (d) outer spine on P3 enp-3 reduced in size in
<specimenCount id="0BB7FD75FFECFFB1FC57D666E7E6FA43" box="[913,934,1406,1432]" pageId="47" pageNumber="283" type="male"></specimenCount>
(
<bibRefCitation id="79204B0DFFECFFB1FC72D665E017FA4C" author="Mielke, W." box="[948,1111,1405,1431]" pageId="47" pageNumber="283" pagination="141 - 153" refId="ref75832" refString="Mielke, W. (1994 b) Two co-occuring new Karllangia species (Copepoda: Ameiridae) from the Caribbean coast of Costa Rica. Revista de Biologia tropical, 42, 141 - 153." type="journal article" year="1994">Mielke 1994b</bibRefCitation>
:
<figureCitation id="858A2A79FFECFFB1FBA4D665E0F9FA4C" box="[1122,1209,1405,1431]" captionStart="FIGURE 6" captionStartId="14.[151,250,1812,1836]" captionTargetBox="[249,1337,188,1787]" captionTargetPageId="14" captionText="FIGURE 6. Sexual dimorphism of P2 (AE) and P4 (FG) endopodal segmentation and armature within the Parastenheliidae. A, Foweya tertia (Kunz, 1975); B, Microthalestris costata (Pallares, 1982) comb. nov., M. variabilis sp. nov., Parastenhelia similis Thompson &amp; Scott, 1903, P. willemvervoorti sp. nov., Porirualia pyriformis (Song, Kim &amp; Chang, 2003) comb. nov.; C, Johnwellsia bipartita sp. nov.; D, Parastenhelia hornelli Thompson &amp; Scott, 1903 sensu Apostolov (1973); E, Parastenhelia oligochaeta Wells &amp; Rao, 1987; F, Microthalestris bulbosa (Gee, 2006) comb. nov., Parastenhelia willemvervoorti sp. nov.; G, Microthalestris minuta (Pallares, 1982) comb. nov., Penicillicaris penicillata sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5572433" httpUri="https://zenodo.org/record/5572433/files/figure.png" pageId="47" pageNumber="283">Fig. 6C</figureCitation>
;
<bibRefCitation id="79204B0DFFECFFB1FB03D665E175FA4C" author="Gee, J. M." box="[1221,1333,1405,1431]" pageId="47" pageNumber="283" pagination="2611 - 2651" refId="ref71824" refString="Gee, J. M. (2006) Parastenheliidae (Copepoda: Harpacticoida) from the Isles of Scilly. Journal of natural History, 40, 2611 - 2651. https: // doi. org / 10.1080 / 00222930601108194" type="journal article" year="2006">Gee 2006</bibRefCitation>
:
<figureCitation id="858A2A79FFECFFB1FA86D665E1D7FA43" box="[1344,1431,1405,1432]" captionStart="FIGURE 5" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[185,1407,194,1907]" captionTargetId="figure-17@11.[172,1416,181,1919]" captionTargetPageId="11" captionText="FIGURE 5. Johnwellsia bipartita gen. et sp. nov. (♂): A, antennule, ventral (armature omitted); B, antennule, ventral (disarticulated, showing armature); C, P2 endopod, anterior; D, P3 endopod, anterior; E, P4 endopod, anterior; F, P5, anterior; G, P6; H, maxilliped." figureDoi="http://doi.org/10.5281/zenodo.5572431" httpUri="https://zenodo.org/record/5572431/files/figure.png" pageId="47" pageNumber="283">Fig. 5C</figureCitation>
; Fig. 7 herein).
</paragraph>
</subSubSection>
<subSubSection id="55AB6577FFECFFAEFF01D6DDE019FC06" lastPageId="48" lastPageNumber="284" pageId="47" pageNumber="283" type="diagnosis">
<paragraph id="1D0E36FCFFECFFB1FF01D6DDE5EEF963" blockId="47.[151,1437,505,2008]" pageId="47" pageNumber="283">
<emphasis id="2FC5EAEEFFECFFB1FF01D6DDE502FA04" bold="true" box="[199,322,1477,1503]" pageId="47" pageNumber="283">Diagnosis.</emphasis>
<taxonomicName id="DAB14D7FFFECFFB1FE8BD6DDE648FA04" baseAuthorityName="Willen" baseAuthorityYear="2000" box="[333,520,1477,1503]" class="Hexanauplia" family="Parastenheliidae" kingdom="Animalia" order="Harpacticoida" pageId="47" pageNumber="283" phylum="Arthropoda" rank="family">Parastenheliidae</taxonomicName>
. Sexual dimorphism in antennule, P3 enp-3 outer spine, P5P6 and urosomal segmentation. Body subcylindrical; posterior margin of cephalothorax (typically) and all somites (except anal somite) typically with deeply divided, denticulodigitate or lobate, hyaline frills (plain in
<taxonomicName id="DAB14D7FFFECFFB1FBB7D516E0A6F9FC" baseAuthorityName="Mielke" baseAuthorityYear="1994" box="[1137,1254,1549,1575]" class="Hexanauplia" family="Parastenheliidae" genus="Thalestrella" kingdom="Animalia" order="Harpacticoida" pageId="47" pageNumber="283" phylum="Arthropoda" rank="species" species="obscura" status="comb. nov.">
<emphasis id="2FC5EAEEFFECFFB1FBB7D516E0A6F9FC" box="[1137,1254,1549,1575]" italics="true" pageId="47" pageNumber="283">T. obscura</emphasis>
</taxonomicName>
<emphasis id="2FC5EAEEFFECFFB1FB2BD515E12DF9FC" bold="true" box="[1261,1389,1549,1575]" pageId="47" pageNumber="283">
<taxonomicNameLabel id="34F65795FFECFFB1FB2BD515E12DF9FC" box="[1261,1389,1549,1575]" pageId="47" pageNumber="283" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
and possibly
<taxonomicName id="DAB14D7FFFECFFB1FF3BD52AE5C5F990" baseAuthorityName="Noodt" baseAuthorityYear="1964" box="[253,389,1585,1611]" class="Hexanauplia" family="Parastenheliidae" genus="Thalestrella" kingdom="Animalia" order="Harpacticoida" pageId="47" pageNumber="283" phylum="Arthropoda" rank="species" species="arenicola" status="comb. nov.">
<emphasis id="2FC5EAEEFFECFFB1FF3BD52AE5C5F990" box="[253,389,1585,1611]" italics="true" pageId="47" pageNumber="283">T. arenicola</emphasis>
</taxonomicName>
<emphasis id="2FC5EAEEFFECFFB1FE48D529E650F990" bold="true" box="[398,528,1585,1611]" pageId="47" pageNumber="283">
<taxonomicNameLabel id="34F65795FFECFFB1FE48D529E650F990" box="[398,528,1585,1611]" pageId="47" pageNumber="283" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
). Rostrum defined at base, of moderate size, reaching at most to end of second segment of antennule; linguiform. Anal operculum semicircular, variously ornamented; usually bordered with fine spinules or denticles. Caudal ramus at most as wide as long, with dorsal spinular row; with seven setae, setae IVV not basally inflated in
<specimenCount id="0BB7FD75FFECFFB1FE54D586E5E8F963" box="[402,424,1694,1720]" pageId="47" pageNumber="283" type="female"></specimenCount>
.
</paragraph>
<paragraph id="1D0E36FCFFECFFAEFF01D5D9E6A9FE76" blockId="47.[151,1437,505,2008]" lastBlockId="48.[151,1437,151,645]" lastPageId="48" lastPageNumber="284" pageId="47" pageNumber="283">
<materialsCitation id="ADD93CA1FFECFFAEFF01D5D9E6A5FE76" ID-GBIF-Occurrence="3387337313" collectionCode="XIV, XX, VIII, IX, XII, XIII, XXI, XXIII, XXIV, XXV, XXVI, XXVIII" county="Antenna" lastPageId="48" lastPageNumber="284" location="Mandible" municipality="Female" pageId="47" pageNumber="283" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Maale">
Antennule
<specimenCount id="0BB7FD75FFECFFB1FE80D5DAE51BF907" box="[326,347,1730,1756]" pageId="47" pageNumber="283" type="female"></specimenCount>
compact, 8- or indistinctly 9-segmented; segment 1 not elongate, distal four or five segments small with combined length only slightly longer than segment 4; with aesthetascs on segments 4 and 8 (or 9). Antennule
<specimenCount id="0BB7FD75FFECFFB1FED1D412E56CF8FF" box="[279,300,1802,1828]" pageId="47" pageNumber="283" type="male"></specimenCount>
subchirocer, 7-, 8- or indistinctly 9-segmented; ventral surface of segment 2 with characteristic, backwardly directed, spiniform element; with geniculation between segments 5 and 6 (or 6 and 7 when indistinctly 9-segmented) and 23 segments distal to geniculation; compound segment 5 (
<collectionCode id="7BA0AE39FFECFFB1FBEED449E01BF8B0" box="[1064,1115,1873,1899]" pageId="47" pageNumber="283">XIV</collectionCode>
<collectionCode id="7BA0AE39FFECFFB1FBACD449E0D4F8B0" box="[1130,1172,1873,1899]" pageId="47" pageNumber="283">XX</collectionCode>
) swollen, occasionally subdivided dorsally by transverse surface suture; with aesthetasc on segments 5 and 7 or 8; segmental homologies of 8-segmented antennule: I, II
<collectionCode id="7BA0AE39FFECFFB1FDC4D482E674F86F" box="[514,564,1946,1972]" pageId="47" pageNumber="283">VIII</collectionCode>
,
<collectionCode id="7BA0AE39FFECFFB1FD86D482E61EF86F" box="[576,606,1946,1972]" pageId="47" pageNumber="283">IX</collectionCode>
<collectionCode id="7BA0AE39FFECFFB1FDABD482E6D5F86F" box="[621,661,1946,1972]" pageId="47" pageNumber="283">XII</collectionCode>
,
<collectionCode id="7BA0AE39FFECFFB1FD66D482E693F86F" box="[672,723,1946,1972]" pageId="47" pageNumber="283">XIII</collectionCode>
,
<collectionCode id="7BA0AE39FFECFFB1FD19D482E752F86F" box="[735,786,1946,1972]" pageId="47" pageNumber="283">XIV</collectionCode>
<collectionCode id="7BA0AE39FFECFFB1FCE7D482E70BF86F" box="[801,843,1946,1972]" pageId="47" pageNumber="283">XX</collectionCode>
,
<collectionCode id="7BA0AE39FFECFFB1FC90D482E7C9F86F" box="[854,905,1946,1972]" pageId="47" pageNumber="283">XXI</collectionCode>
<collectionCode id="7BA0AE39FFECFFB1FC51D482E79DF86F" box="[919,989,1946,1972]" pageId="47" pageNumber="283">XXIII</collectionCode>
,
<collectionCode id="7BA0AE39FFECFFB1FC2ED482E06EF86F" box="[1000,1070,1946,1972]" pageId="47" pageNumber="283">XXIV</collectionCode>
<collectionCode id="7BA0AE39FFECFFB1FBFAD482E038F86F" box="[1084,1144,1946,1972]" pageId="47" pageNumber="283">XXV</collectionCode>
,
<collectionCode id="7BA0AE39FFECFFB1FB42D482E08BF86F" box="[1156,1227,1946,1972]" pageId="47" pageNumber="283">XXVI</collectionCode>
<collectionCode id="7BA0AE39FFECFFB1FB1CD482E174F86F" box="[1242,1332,1946,1972]" pageId="47" pageNumber="283">XXVIII</collectionCode>
.
<collectingCounty id="F46F4E70FFECFFB1FAFBD482E1DCF86F" box="[1341,1436,1946,1972]" pageId="47" pageNumber="283">Antenna</collectingCounty>
sexually dimorphic.
<collectingMunicipality id="FD6AAC86FFECFFB1FEBCD4A5E58CF80C" box="[378,460,1981,2007]" pageId="47" pageNumber="283">Female</collectingMunicipality>
antenna with proximal endopodal segment largely fused to basis forming allobasis, with short pinnate seta on abexopodal margin; exopod 2-segmented (or segments partially fused), proximal segment with 12 setae, distal segment with one lateral and two apical elements (surrounded at base by serrate hyaline frill); distal endopodal segment without penicillate elements.
<collectingRegion id="DF75F81EFFF3FFAEFD08D3C7E748FF22" box="[718,776,223,249]" country="Maldives" name="Maale" pageId="48" pageNumber="284">Male</collectingRegion>
antenna with enlarged, highly plumose seta on allobasis; exopodal segments of different size, with only one seta on proximal segment and all lateral setae enlarged and highly plumose.
<location id="186E6027FFF3FFAEFE92D23FE5FFFE9A" LSID="urn:lsid:plazi:treatment:951887EAFFECFFAEFF51D276E0FFFB01:186E6027FFF3FFAEFE92D23FE5FFFE9A" box="[340,447,295,321]" county="Antenna" municipality="Female" name="Mandible" pageId="48" pageNumber="284" stateProvince="Maale">Mandible</location>
with three elements on basis; endopod with eight setae; exopod 1-segmented, with 24 setae.
<location id="186E6027FFF3FFAEFF1AD253E520FEBE" LSID="urn:lsid:plazi:treatment:951887EAFFECFFAEFF51D276E0FFFB01:186E6027FFF3FFAEFF1AD253E520FEBE" box="[220,352,331,357]" county="Antenna" municipality="Female" name="Maxillulary" pageId="48" pageNumber="284" stateProvince="Maale">Maxillulary</location>
coxal epipodite represented by one seta.
<location id="186E6027FFF3FFAEFCE3D253E73CFEBE" LSID="urn:lsid:plazi:treatment:951887EAFFECFFAEFF51D276E0FFFB01:186E6027FFF3FFAEFCE3D253E73CFEBE" box="[805,892,331,357]" county="Antenna" municipality="Female" name="Maxilla" pageId="48" pageNumber="284" stateProvince="Maale">Maxilla</location>
with three endites on syncoxa; endopod discrete, with 45 setae.
<location id="186E6027FFF3FFAEFE8ED277E583FE52" LSID="urn:lsid:plazi:treatment:951887EAFFECFFAEFF51D276E0FFFB01:186E6027FFF3FFAEFE8ED277E583FE52" box="[328,451,367,393]" county="Antenna" municipality="Female" name="Maxilliped" pageId="48" pageNumber="284" stateProvince="Maale">Maxilliped</location>
with one seta on syncoxa; basis with 02 setae on palmar margin; endopod represented by pinnate claw, accompanied by one accessory seta
</materialsCitation>
.
</paragraph>
<paragraph id="1D0E36FCFFF3FFAEFF01D2AFE552FD5E" blockId="48.[151,1437,151,645]" pageId="48" pageNumber="284">
P1 exopod 3-segmented; segments subequal; exp-2 with or without inner seta; exp-3 with two pinnate spines and one geniculate spine and one (non-)geniculate seta. P1 endopod 2-segmented; enp-1 elongate, about twice as long as exopod, with well developed, pinnate inner seta originating from proximal half in a region of reduced chitinization of segment wall; enp-2 very small, with one naked minute seta and two unipinnate claws, longest of which geniculate. P2P4 rami 3-segmented; outer spine of P3 enp-
<specimenCount id="0BB7FD75FFF3FFAEFC44D15FE7EAFDBA" box="[898,938,583,609]" pageId="48" pageNumber="284" type="male">3 ♂</specimenCount>
reduced in size. Armature formula of P2P4 as follows:
</paragraph>
<paragraph id="1D0E36FCFFF3FFAEFD8BD1A0E03BFC97" pageId="48" pageNumber="284">
<table id="6FB1C45CFFF30061FEA6D1A0E03BFC97" box="[352,1147,696,844]" gridcols="3" gridrows="4" pageId="48" pageNumber="284" rowsContinueFrom="46.[352,1113,162,310]">
<tr id="A38134BEFFF30061FEA6D1A0E03BFD09" box="[352,1147,696,722]" gridrow="0" pageId="48" pageNumber="284" rowspan-0="1">
<th id="E0505DC2FFF30061FD8BD1A0E690FD09" box="[589,720,696,722]" gridcol="1" gridrow="0" pageId="48" pageNumber="284">exopod</th>
<th id="E0505DC2FFF30061FC3FD1A0E03BFD09" box="[1017,1147,696,722]" gridcol="2" gridrow="0" pageId="48" pageNumber="284">endopod</th>
</tr>
<tr id="A38134BEFFF30061FEA6D1F9E03BFD20" box="[352,1147,737,763]" gridrow="1" pageId="48" pageNumber="284">
<th id="E0505DC2FFF30061FEA6D1F9E53AFD20" box="[352,378,737,763]" gridcol="0" gridrow="1" pageId="48" pageNumber="284">P2</th>
<td id="E0505DC2FFF30061FD8BD1F9E690FD20" box="[589,720,737,763]" gridcol="1" gridrow="1" pageId="48" pageNumber="284">0.1.223</td>
<td id="E0505DC2FFF30061FC3FD1F9E03BFD20" box="[1017,1147,737,763]" gridcol="2" gridrow="1" pageId="48" pageNumber="284">1.1.121</td>
</tr>
<tr id="A38134BEFFF30061FEA6D012E03BFCFF" box="[352,1147,778,804]" gridrow="2" pageId="48" pageNumber="284">
<th id="E0505DC2FFF30061FEA6D012E53AFCFF" box="[352,378,778,804]" gridcol="0" gridrow="2" pageId="48" pageNumber="284">P3</th>
<td id="E0505DC2FFF30061FD8BD012E690FCFF" box="[589,720,778,804]" gridcol="1" gridrow="2" pageId="48" pageNumber="284">0.1.(23)23</td>
<td id="E0505DC2FFF30061FC3FD012E03BFCFF" box="[1017,1147,778,804]" gridcol="2" gridrow="2" pageId="48" pageNumber="284">1.1.(12)21</td>
</tr>
<tr id="A38134BEFFF30061FEA6D02AE03BFC97" box="[352,1147,818,844]" gridrow="3" pageId="48" pageNumber="284">
<th id="E0505DC2FFF30061FEA6D02AE53AFC97" box="[352,378,818,844]" gridcol="0" gridrow="3" pageId="48" pageNumber="284">P4</th>
<td id="E0505DC2FFF30061FD8BD02AE690FC97" box="[589,720,818,844]" gridcol="1" gridrow="3" pageId="48" pageNumber="284">0.1.(23)23</td>
<td id="E0505DC2FFF30061FC3FD02AE03BFC97" box="[1017,1147,818,844]" gridcol="2" gridrow="3" pageId="48" pageNumber="284">1.1.(12)21</td>
</tr>
</table>
</paragraph>
<paragraph id="1D0E36FCFFF3FFAEFF01D063E019FC06" blockId="48.[151,1437,891,1242]" pageId="48" pageNumber="284">
P
<specimenCount id="0BB7FD75FFF3FFAEFF1ED063E542FC4E" box="[216,258,891,918]" pageId="48" pageNumber="284" type="female">5 ♀</specimenCount>
endopodal lobe triangular, with five setae; inner margin without transverse striae; exopod elongate (25 times longer than wide), with 56 elements. P
<specimenCount id="0BB7FD75FFF3FFAEFD57D087E6F9FC62" box="[657,697,927,954]" pageId="48" pageNumber="284" type="male">5 ♂</specimenCount>
endopodal lobe with two elements; exopod 1-segmented, with 45 elements. Vestigial P
<specimenCount id="0BB7FD75FFF3FFAEFE47D0DBE5EBFC06" box="[385,427,963,989]" pageId="48" pageNumber="284" type="female">6 ♀</specimenCount>
represented by 23 short setae. P
<specimenCount id="0BB7FD75FFF3FFAEFCE5D0DBE70CFC06" box="[803,844,963,989]" pageId="48" pageNumber="284" type="male">6 ♂</specimenCount>
with three setae/spines.
</paragraph>
</subSubSection>
<subSubSection id="55AB6577FFF3FFAEFF01D0FFE0FFFB01" pageId="48" pageNumber="284" type="type_taxon">
<paragraph id="1D0E36FCFFF3FFAEFF01D0FFE7A3FBDA" blockId="48.[151,1437,891,1242]" box="[199,995,999,1026]" pageId="48" pageNumber="284">
<emphasis id="2FC5EAEEFFF3FFAEFF01D0FFE522FBDA" bold="true" box="[199,354,999,1025]" pageId="48" pageNumber="284">
<typeStatus id="C20A885EFFF3FFAEFF01D0FFE542FBDA" box="[199,258,999,1025]" pageId="48" pageNumber="284">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="DAB14D7FFFF3FFAEFEAFD0FFE76CFBD9" authority="Monard, 1935 a" authorityName="Monard." authorityYear="1935" baseAuthorityName="sensu Por" baseAuthorityYear="1935" box="[361,812,999,1026]" class="Hexanauplia" family="Parastenheliidae" genus="Thalestrella" kingdom="Animalia" order="Harpacticoida" pageId="48" pageNumber="284" phylum="Arthropoda" rank="species" species="ornatissima">
<emphasis id="2FC5EAEEFFF3FFAEFEAFD0FFE63BFBDA" box="[361,635,999,1025]" italics="true" pageId="48" pageNumber="284">Thalestrella ornatissima</emphasis>
<bibRefCitation id="79204B0DFFF3FFAEFD44D0FFE76CFBD9" author="Monard, A." box="[642,812,999,1026]" pageId="48" pageNumber="284" pagination="5 - 88" refId="ref76147" refString="Monard, A. (1935 a) Etude sur la faune des Harpacticoides marins de Roscoff. Travaux de la Station biologique de Roscoff, 13, 5 - 88, figs. 1 - 235." type="journal article" year="1935">Monard, 1935a</bibRefCitation>
</taxonomicName>
(by monotypy).
</paragraph>
<paragraph id="1D0E36FCFFF3FFAEFF01D713E673FB6E" blockId="48.[151,1437,891,1242]" pageId="48" pageNumber="284">
<emphasis id="2FC5EAEEFFF3FFAEFF01D713E530FBFE" bold="true" box="[199,368,1035,1061]" pageId="48" pageNumber="284">Other species.</emphasis>
<taxonomicName id="DAB14D7FFFF3FFAEFEBED713E6BBFBFE" authority="Noodt, 1964" authorityName="Noodt" authorityYear="1964" box="[376,763,1035,1061]" class="Hexanauplia" family="Parastenheliidae" genus="Karllangia" kingdom="Animalia" order="Harpacticoida" pageId="48" pageNumber="284" phylum="Arthropoda" rank="species" species="arenicola">
<emphasis id="2FC5EAEEFFF3FFAEFEBED713E625FBFE" box="[376,613,1035,1061]" italics="true" pageId="48" pageNumber="284">Karllangia arenicola</emphasis>
<bibRefCitation id="79204B0DFFF3FFAEFDA8D713E6BBFBFE" author="Noodt, W." box="[622,763,1035,1061]" pageId="48" pageNumber="284" pagination="128 - 154" refId="ref76696" refString="Noodt, W. (1964) Copepoda Harpacticoidea aus dem Litoral des Roten Meeres. Kieler Meeresforschungen, 20, Sonderheft, 128 - 154." type="journal article" year="1964">Noodt, 1964</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="DAB14D7FFFF3FFAEFCDDD714E00AFBFD" authority="(Noodt, 1964)" baseAuthorityName="Noodt" baseAuthorityYear="1964" box="[795,1098,1035,1062]" class="Hexanauplia" family="Parastenheliidae" genus="Karllangia" kingdom="Animalia" order="Harpacticoida" pageId="48" pageNumber="284" phylum="Arthropoda" rank="species" species="arenicola" status="comb. nov.">
<emphasis id="2FC5EAEEFFF3FFAEFCDDD714E7E2FBFE" box="[795,930,1035,1061]" italics="true" pageId="48" pageNumber="284">T. arenicola</emphasis>
(
<bibRefCitation id="79204B0DFFF3FFAEFC74D713E001FBFD" author="Noodt, W." box="[946,1089,1035,1062]" pageId="48" pageNumber="284" pagination="128 - 154" refId="ref76696" refString="Noodt, W. (1964) Copepoda Harpacticoidea aus dem Litoral des Roten Meeres. Kieler Meeresforschungen, 20, Sonderheft, 128 - 154." type="journal article" year="1964">Noodt, 1964</bibRefCitation>
)
</taxonomicName>
<emphasis id="2FC5EAEEFFF3FFAEFB94D713E094FBFE" bold="true" box="[1106,1236,1035,1061]" pageId="48" pageNumber="284">
<taxonomicNameLabel id="34F65795FFF3FFAEFB94D713E094FBFE" box="[1106,1236,1035,1061]" pageId="48" pageNumber="284" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
;
<taxonomicName id="DAB14D7FFFF3FFAEFB25D714E560FB92" authority="Wells, 1967" authorityName="Wells" authorityYear="1967" class="Hexanauplia" family="Parastenheliidae" genus="Thalestrella" kingdom="Animalia" order="Harpacticoida" pageId="48" pageNumber="284" phylum="Arthropoda" rank="species" species="psammophila">
<emphasis id="2FC5EAEEFFF3FFAEFB25D714E1DCFBFE" box="[1251,1436,1035,1061]" italics="true" pageId="48" pageNumber="284">K. psammophila</emphasis>
<bibRefCitation id="79204B0DFFF3FFAEFF51D737E560FB92" author="Wells, J. B. J." box="[151,288,1071,1097]" pageId="48" pageNumber="284" pagination="189 - 358" refId="ref82201" refString="Wells, J. B. J. (1967) The littoral Copepoda (Crustacea) of Inhaca Island, Mozambique. Transactions of the Royal Society of Edinburgh, 67 (7), 189 - 358. https: // doi. org / 10.1017 / s 0080456800024017" type="journal article" year="1967">Wells, 1967</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="DAB14D7FFFF3FFAEFE83D728E6E1FB91" authority="(Wells, 1967)" baseAuthorityName="Wells" baseAuthorityYear="1967" box="[325,673,1071,1098]" class="Hexanauplia" family="Parastenheliidae" genus="Thalestrella" kingdom="Animalia" order="Harpacticoida" pageId="48" pageNumber="284" phylum="Arthropoda" rank="species" species="psammophila" status="comb. nov.">
<emphasis id="2FC5EAEEFFF3FFAEFE83D728E5BBFB92" box="[325,507,1071,1097]" italics="true" pageId="48" pageNumber="284">T. psammophila</emphasis>
(
<bibRefCitation id="79204B0DFFF3FFAEFDCBD737E6D9FB91" author="Wells, J. B. J." box="[525,665,1071,1098]" pageId="48" pageNumber="284" pagination="189 - 358" refId="ref82201" refString="Wells, J. B. J. (1967) The littoral Copepoda (Crustacea) of Inhaca Island, Mozambique. Transactions of the Royal Society of Edinburgh, 67 (7), 189 - 358. https: // doi. org / 10.1017 / s 0080456800024017" type="journal article" year="1967">Wells, 1967</bibRefCitation>
)
</taxonomicName>
<emphasis id="2FC5EAEEFFF3FFAEFD6DD737E76FFB92" bold="true" box="[683,815,1071,1097]" pageId="48" pageNumber="284">
<taxonomicNameLabel id="34F65795FFF3FFAEFD6DD737E76FFB92" box="[683,815,1071,1097]" pageId="48" pageNumber="284" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
;
<taxonomicName id="DAB14D7FFFF3FFAEFC84D737E144FB91" authority="Apostolov, 1975" authorityName="Apostolov" authorityYear="1975" baseAuthorityName="Apostolov" baseAuthorityYear="1975" box="[834,1284,1071,1098]" class="Hexanauplia" family="Parastenheliidae" genus="Parastenhelia" kingdom="Animalia" order="Harpacticoida" pageId="48" pageNumber="284" phylum="Arthropoda" rank="species" species="reducta">
<emphasis id="2FC5EAEEFFF3FFAEFC84D737E07FFB92" box="[834,1087,1071,1097]" italics="true" pageId="48" pageNumber="284">Parastenhelia reducta</emphasis>
<bibRefCitation id="79204B0DFFF3FFAEFB81D737E144FB91" author="Apostolov, A." box="[1095,1284,1071,1098]" pageId="48" pageNumber="284" pagination="165 - 178" refId="ref67876" refString="Apostolov, A. (1975) Les Harpacticoides marins de la mer Noire. Description de quelques formes nouvelles. Vie et Milieu, (A), 25, 165 - 178." type="journal article" year="1975">Apostolov, 1975</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="DAB14D7FFFF3FFAEFAEFD728E523FBB5" authority="(Apostolov, 1975)" baseAuthorityName="Apostolov" baseAuthorityYear="1975" class="Hexanauplia" family="Parastenheliidae" genus="Parastenhelia" kingdom="Animalia" order="Harpacticoida" pageId="48" pageNumber="284" phylum="Arthropoda" rank="species" species="reducta" status="comb. nov.">
<emphasis id="2FC5EAEEFFF3FFAEFAEFD728E1DDFB92" box="[1321,1437,1071,1097]" italics="true" pageId="48" pageNumber="284">T. reducta</emphasis>
(
<bibRefCitation id="79204B0DFFF3FFAEFF59D74BE51BFBB5" author="Apostolov, A." box="[159,347,1107,1134]" pageId="48" pageNumber="284" pagination="165 - 178" refId="ref67876" refString="Apostolov, A. (1975) Les Harpacticoides marins de la mer Noire. Description de quelques formes nouvelles. Vie et Milieu, (A), 25, 165 - 178." type="journal article" year="1975">Apostolov, 1975</bibRefCitation>
)
</taxonomicName>
<emphasis id="2FC5EAEEFFF3FFAEFEACD74BE5ABFBB6" bold="true" box="[362,491,1107,1133]" pageId="48" pageNumber="284">
<taxonomicNameLabel id="34F65795FFF3FFAEFEACD74BE5ABFBB6" box="[362,491,1107,1133]" pageId="48" pageNumber="284" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
;
<taxonomicName id="DAB14D7FFFF3FFAEFE3CD74CE7AEFBB6" authority="Wells &amp; Rao, 1987" authorityName="Wells &amp; Rao" authorityYear="1987" box="[506,1006,1107,1133]" class="Hexanauplia" family="Parastenheliidae" genus="Karllangia" kingdom="Animalia" order="Harpacticoida" pageId="48" pageNumber="284" phylum="Arthropoda" rank="subSpecies" species="arenicola" subSpecies="bengalensis">
<emphasis id="2FC5EAEEFFF3FFAEFE3CD74CE750FBB6" box="[506,784,1107,1133]" italics="true" pageId="48" pageNumber="284">K. arenicola bengalensis</emphasis>
<bibRefCitation id="79204B0DFFF3FFAEFCD1D74BE7AEFBB6" author="Wells, J. B. J. &amp; Rao, G. C." box="[791,1006,1107,1133]" pageId="48" pageNumber="284" pagination="1 - 385" refId="ref82468" refString="Wells, J. B. J. &amp; Rao, G. C. (1987) Littoral Harpacticoida (Crustacea: Copepoda) from Andaman and Nicobar Islands. Memoirs of the zoological Survey of India, 16 (4), 1 - 385." type="journal article" year="1987">Wells &amp; Rao, 1987</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="DAB14D7FFFF3FFAEFBCAD74CE1DCFBB6" authority="(Wells &amp; Rao, 1987)" baseAuthorityName="Wells &amp; Rao" baseAuthorityYear="1987" box="[1036,1436,1107,1133]" class="Hexanauplia" family="Parastenheliidae" genus="Thalestrella" kingdom="Animalia" order="Harpacticoida" pageId="48" pageNumber="284" phylum="Arthropoda" rank="species" species="bengalensis" status="comb. nov.">
<emphasis id="2FC5EAEEFFF3FFAEFBCAD74CE0ECFBB6" box="[1036,1196,1107,1133]" italics="true" pageId="48" pageNumber="284">T. bengalensis</emphasis>
(
<bibRefCitation id="79204B0DFFF3FFAEFB7CD74BE1D4FBB6" author="Wells, J. B. J. &amp; Rao, G. C." box="[1210,1428,1107,1133]" pageId="48" pageNumber="284" pagination="1 - 385" refId="ref82468" refString="Wells, J. B. J. &amp; Rao, G. C. (1987) Littoral Harpacticoida (Crustacea: Copepoda) from Andaman and Nicobar Islands. Memoirs of the zoological Survey of India, 16 (4), 1 - 385." type="journal article" year="1987">Wells &amp; Rao, 1987</bibRefCitation>
)
</taxonomicName>
<emphasis id="2FC5EAEEFFF3FFAEFF51D76FE559FB4A" bold="true" box="[151,281,1143,1169]" pageId="48" pageNumber="284">
<taxonomicNameLabel id="34F65795FFF3FFAEFF51D76FE559FB4A" box="[151,281,1143,1169]" pageId="48" pageNumber="284" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
;
<taxonomicName id="DAB14D7FFFF3FFAEFEEFD760E612FB4A" authority="Mielke, 1994 b" authorityName="Mielke" authorityYear="1994" box="[297,594,1143,1169]" class="Hexanauplia" family="Parastenheliidae" genus="Thalestrella" kingdom="Animalia" order="Harpacticoida" pageId="48" pageNumber="284" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="2FC5EAEEFFF3FFAEFEEFD760E5E5FB4A" box="[297,421,1143,1169]" italics="true" pageId="48" pageNumber="284">K. obscura</emphasis>
<bibRefCitation id="79204B0DFFF3FFAEFE6BD76FE612FB4A" author="Mielke, W." box="[429,594,1143,1169]" pageId="48" pageNumber="284" pagination="141 - 153" refId="ref75832" refString="Mielke, W. (1994 b) Two co-occuring new Karllangia species (Copepoda: Ameiridae) from the Caribbean coast of Costa Rica. Revista de Biologia tropical, 42, 141 - 153." type="journal article" year="1994">Mielke, 1994b</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="DAB14D7FFFF3FFAEFDB4D760E7E8FB49" authority="(Mielke, 1994 b)" baseAuthorityName="Mielke" baseAuthorityYear="1994" box="[626,936,1143,1170]" class="Hexanauplia" family="Parastenheliidae" genus="Thalestrella" kingdom="Animalia" order="Harpacticoida" pageId="48" pageNumber="284" phylum="Arthropoda" rank="species" species="obscura" status="comb. nov.">
<emphasis id="2FC5EAEEFFF3FFAEFDB4D760E6A8FB4A" box="[626,744,1143,1169]" italics="true" pageId="48" pageNumber="284">T. obscura</emphasis>
(
<bibRefCitation id="79204B0DFFF3FFAEFD3FD76FE7E0FB49" author="Mielke, W." box="[761,928,1143,1170]" pageId="48" pageNumber="284" pagination="141 - 153" refId="ref75832" refString="Mielke, W. (1994 b) Two co-occuring new Karllangia species (Copepoda: Ameiridae) from the Caribbean coast of Costa Rica. Revista de Biologia tropical, 42, 141 - 153." type="journal article" year="1994">Mielke, 1994b</bibRefCitation>
)
</taxonomicName>
<emphasis id="2FC5EAEEFFF3FFAEFC76D76FE072FB4A" bold="true" box="[944,1074,1143,1169]" pageId="48" pageNumber="284">
<taxonomicNameLabel id="34F65795FFF3FFAEFC76D76FE072FB4A" box="[944,1074,1143,1169]" pageId="48" pageNumber="284" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
;
<taxonomicName id="DAB14D7FFFF3FFAEFB84D760E128FB4A" authority="Mielke, 1994 b" authorityName="Mielke" authorityYear="1994" box="[1090,1384,1143,1169]" class="Hexanauplia" family="Parastenheliidae" genus="Thalestrella" kingdom="Animalia" order="Harpacticoida" pageId="48" pageNumber="284" phylum="Arthropoda" rank="species" species="pulchra">
<emphasis id="2FC5EAEEFFF3FFAEFB84D760E0FBFB4A" box="[1090,1211,1143,1169]" italics="true" pageId="48" pageNumber="284">K. pulchra</emphasis>
<bibRefCitation id="79204B0DFFF3FFAEFB05D76FE128FB4A" author="Mielke, W." box="[1219,1384,1143,1169]" pageId="48" pageNumber="284" pagination="141 - 153" refId="ref75832" refString="Mielke, W. (1994 b) Two co-occuring new Karllangia species (Copepoda: Ameiridae) from the Caribbean coast of Costa Rica. Revista de Biologia tropical, 42, 141 - 153." type="journal article" year="1994">Mielke, 1994b</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="DAB14D7FFFF3FFAEFA4ED760E5EBFB6D" authority="(Mielke, 1994 b)" baseAuthorityName="Mielke" baseAuthorityYear="1994" class="Hexanauplia" family="Parastenheliidae" genus="Thalestrella" kingdom="Animalia" order="Harpacticoida" pageId="48" pageNumber="284" phylum="Arthropoda" rank="species" species="pulchra" status="comb. nov.">
<emphasis id="2FC5EAEEFFF3FFAEFA4ED760E4AEFB6E" italics="true" pageId="48" pageNumber="284">T. pulchra</emphasis>
(
<bibRefCitation id="79204B0DFFF3FFAEFF3BD783E5E3FB6D" author="Mielke, W." box="[253,419,1179,1206]" pageId="48" pageNumber="284" pagination="141 - 153" refId="ref75832" refString="Mielke, W. (1994 b) Two co-occuring new Karllangia species (Copepoda: Ameiridae) from the Caribbean coast of Costa Rica. Revista de Biologia tropical, 42, 141 - 153." type="journal article" year="1994">Mielke, 1994b</bibRefCitation>
)
</taxonomicName>
<emphasis id="2FC5EAEEFFF3FFAEFE74D783E673FB6E" bold="true" box="[434,563,1179,1205]" pageId="48" pageNumber="284">
<taxonomicNameLabel id="34F65795FFF3FFAEFE74D783E673FB6E" box="[434,563,1179,1205]" pageId="48" pageNumber="284" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="1D0E36FCFFF3FFAEFF01D7A7E0FFFB01" blockId="48.[151,1437,891,1242]" box="[199,1215,1214,1242]" pageId="48" pageNumber="284">
Species inquirenda.
<taxonomicName id="DAB14D7FFFF3FFAEFE77D7A7E0FAFB01" authority="(Norman &amp; Scott, 1905) sensu Por (1964)" authorityName="sensu Por" authorityYear="1964" baseAuthorityName="Norman &amp; Scott" baseAuthorityYear="1905" box="[433,1210,1215,1242]" class="Hexanauplia" family="Parastenheliidae" genus="Parastenhelia" kingdom="Animalia" order="Harpacticoida" pageId="48" pageNumber="284" phylum="Arthropoda" rank="species" species="ornatissima">
<emphasis id="2FC5EAEEFFF3FFAEFE77D7A7E699FB02" box="[433,729,1215,1241]" italics="true" pageId="48" pageNumber="284">Parastenhelia ornatissima</emphasis>
(
<bibRefCitation id="79204B0DFFF3FFAEFD2ED7A7E7AAFB01" author="Norman, A. M. &amp; Scott, T." box="[744,1002,1215,1242]" pageId="48" pageNumber="284" pagination="284 - 300" refId="ref76779" refString="Norman, A. M. &amp; Scott, T. (1905) Crustacea Copepoda new to Science from Devon and Cornwall. Annals and Magazine of natural History, Series 7, 15, 284 - 300. https: // doi. org / 10.1080 / 03745480509443044" type="journal article" year="1905">Norman &amp; Scott, 1905</bibRefCitation>
) sensu
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</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>