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<document id="E32D7C3DEC22F980BF8E3004C7711639" ID-DOI="10.11646/zootaxa.3478.1.32" ID-GBIF-Dataset="f3e4ca1e-a40b-465a-a900-7a898e472cde" ID-ISSN="1175-5326" ID-Zenodo-Dep="282268" ID-ZooBank="FC92E481-4FF7-4DD8-B7C9-9F192F373D2E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460524755954" checkinUser="plazi" docAuthor="Aarvik, Leif, Karisch, Timm &amp; Marthinsen, Gunnhild" docDate="2012" docId="03C13418FFEEFF961AE7F981388AFE7A" docLanguage="en" docName="zt03478p372.pdf" docOrigin="Zootaxa 3478" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Dracontogena metamorphica Meyrick 1928" docType="treatment" docVersion="4" lastPageNumber="357" masterDocId="FFF84C60FFE5FF9A1A70FFA73C2AFFB5" masterDocTitle="Review of the Afrotropical genus Dracontogena Diakonoff, 1970 (Lepidoptera, Tortricidae) with descriptions of eight new species" masterLastPageNumber="372" masterPageNumber="345" pageNumber="356" updateTime="1698315014682" updateUser="plazi">
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<mods:title id="13A5AEA9A437A65830E740C2F0B92F9C">Review of the Afrotropical genus Dracontogena Diakonoff, 1970 (Lepidoptera, Tortricidae) with descriptions of eight new species</mods:title>
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<mods:namePart id="F66919C892D75811B2674AA8426E1DD4">Aarvik, Leif</mods:namePart>
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<mods:namePart id="1C5B4F0396C89E1E8599EB3E59BE5B42">Karisch, Timm</mods:namePart>
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<mods:namePart id="DF87ADC2909B55EC314349C52514EB69">Marthinsen, Gunnhild</mods:namePart>
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<mods:date id="427C85FCA6A561536D6873FA8F8F8DB6">2012</mods:date>
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<treatment id="03C13418FFEEFF961AE7F981388AFE7A" ID-DOI="http://doi.org/10.5281/zenodo.5697414" ID-GBIF-Taxon="194203686" ID-Zenodo-Dep="5697414" LSID="urn:lsid:plazi:treatment:03C13418FFEEFF961AE7F981388AFE7A" httpUri="http://treatment.plazi.org/id/03C13418FFEEFF961AE7F981388AFE7A" lastPageId="12" lastPageNumber="357" pageId="11" pageNumber="356">
<subSubSection id="C372D685FFEEFF911AE7F9813E8DF9F5" box="[151,679,1574,1600]" pageId="11" pageNumber="356" type="nomenclature">
<paragraph id="8BD7850EFFEEFF911AE7F9813E8DF9F5" blockId="11.[151,679,1574,1600]" box="[151,679,1574,1600]" pageId="11" pageNumber="356">
<heading id="D09F3262FFEEFF911AE7F9813E8DF9F5" bold="true" box="[151,679,1574,1600]" fontSize="11" level="1" pageId="11" pageNumber="356" reason="1">
<emphasis id="B91C591CFFEEFF911AE7F9813E8DF9F5" bold="true" box="[151,679,1574,1600]" pageId="11" pageNumber="356">
<taxonomicName id="4C68FE8DFFEEFF911AE7F9813DD3F9F5" ID-CoL="94S3N" authority="Meyrick, 1928" authorityName="Meyrick" authorityYear="1928" box="[151,505,1574,1600]" class="Insecta" family="Tortricidae" genus="Dracontogena" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="356" phylum="Arthropoda" rank="species" species="metamorphica">
<emphasis id="B91C591CFFEEFF911AE7F9813DD3F9F5" bold="true" box="[151,505,1574,1600]" italics="true" pageId="11" pageNumber="356">Dracontogena metamorphica</emphasis>
</taxonomicName>
species group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C372D685FFEEFF911AE7F9CA3D7DF8E9" pageId="11" pageNumber="356" type="description">
<paragraph id="8BD7850EFFEEFF911AE7F9CA3F59F978" blockId="11.[151,1437,1645,1742]" pageId="11" pageNumber="356">Male hindwing without modifications, valva lacking internal circular scale patch, male abdomen without coremata. The spatulate scales at the hindwing anal corner have a serrate distal edge. The species are variable, and in general, dissection of the genitalia is necessary for accurate identification.</paragraph>
<paragraph id="8BD7850EFFEEFF911AE7F8863EECF88F" blockId="11.[151,710,1824,1884]" box="[151,710,1824,1851]" pageId="11" pageNumber="356">
<heading id="D09F3262FFEEFF911AE7F8863EECF88F" bold="true" box="[151,710,1824,1851]" fontSize="11" level="1" pageId="11" pageNumber="356" reason="1">
<taxonomicName id="4C68FE8DFFEEFF911AE7F8863EECF88F" authority="Meyrick, 1928" authorityName="Meyrick" authorityYear="1928" box="[151,710,1824,1851]" class="Insecta" family="Tortricidae" genus="Dracontogena" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="356" phylum="Arthropoda" rank="species" species="metamorphica">
<emphasis id="B91C591CFFEEFF911AE7F8863EECF88F" bold="true" box="[151,710,1824,1851]" pageId="11" pageNumber="356">
<emphasis id="B91C591CFFEEFF911AE7F8863DD3F88E" bold="true" box="[151,505,1825,1851]" italics="true" pageId="11" pageNumber="356">Dracontogena metamorphica</emphasis>
(
<bibRefCitation id="EFF9F8FFFFEEFF911878F8873E94F88F" author="Meyrick, E." box="[520,702,1824,1850]" firstAuthor="Meyrick" pageId="11" pageNumber="356" pagination="417 - 448" part="3" refId="ref14767" refString="Meyrick, E. (1928) Exotic Microlepidoptera, 3 (14), 417 - 448." title="Exotic Microlepidoptera" type="journal article" year="1928">Meyrick, 1928</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BD7850EFFEEFF911AE7F8E33D7DF8E9" blockId="11.[151,710,1824,1884]" box="[151,343,1860,1884]" pageId="11" pageNumber="356">
(
<figureCitation id="1353998BFFEEFF911AEFF8E33CD7F8E9" box="[159,253,1860,1884]" captionStart="FIGURES 41 46" captionStartId="15.[151,264,1745,1768]" captionTargetBox="[168,1435,469,1734]" captionTargetId="figure@15.[151,1436,469,1757]" captionTargetPageId="15" captionText="FIGURES 41 46. Male genitalia in Dracontogena. 41. D. bernardi Karisch. (slide L. Aarvik 2808). 42. D. bernardi Karisch, phallus (slide L. Aarvik 2808). 43. D. solii n. sp. (slide NHMO 2010). 44. D. solii n. sp. phallus (slide NHMO 2010). 45. D. tonitrualis (Meyrick). 46. D. metamorphica (Meyrick). Scale figs. 41, 43: 1 mm., figs. 42, 44: 0.5 mm. Figs. 45 46 from Karisch (2005)." httpUri="https://zenodo.org/record/282275/files/figure.png" pageId="11" pageNumber="356">Figs. 46</figureCitation>
,
<figureCitation id="1353998BFFEEFF911B78F8E33D65F8E9" box="[264,335,1860,1884]" captionStart="FIGURES 60 63" captionStartId="19.[151,264,1924,1947]" captionTargetBox="[177,1407,216,1880]" captionTargetId="figure@19.[161,1425,193,1903]" captionTargetPageId="19" captionText="FIGURES 60 63. Female genitalia of Dracontogena. 60. D. agassizi n. sp. (slide USNM 137,547). 61. D. agassizi n. sp. sterigma and sternite 7 (slide T. Karisch 1923). 62 63. D. metamorphica (Meyrick) (slide L. Aarvik 2011.025). Scale figs. 60, 62: 1 mm., figs. 61, 63: 0.5 mm." httpUri="https://zenodo.org/record/282278/files/figure.png" pageId="11" pageNumber="356">62, 63</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C372D685FFEEFF911AE7F82A3F85F875" pageId="11" pageNumber="356" type="reference_group">
<paragraph id="8BD7850EFFEEFF911AE7F82A3FC8F814" blockId="11.[151,994,1931,1984]" box="[151,994,1931,1954]" pageId="11" pageNumber="356">
<treatmentCitationGroup id="AB78A220FFEEFF911AE7F82A3FC8F814" box="[151,994,1931,1954]" pageId="11" pageNumber="356">
<treatmentCitation id="0AC9A31FFFEEFF911AE7F82A3E74F814" author="Meyrick" box="[151,606,1931,1954]" page="439" pageId="11" pageNumber="356" year="1928">
<taxonomicName id="4C68FE8DFFEEFF911AE7F82A3E74F814" ID-CoL="3BY3C" authority="Meyrick, 1928: 439" authorityName="Meyrick" authorityPageNumber="439" authorityYear="1928" box="[151,606,1931,1954]" class="Insecta" family="Tortricidae" genus="Eucosma" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="356" phylum="Arthropoda" rank="species" species="metamorphica">
<emphasis id="B91C591CFFEEFF911AE7F82A3DA6F817" box="[151,396,1932,1954]" italics="true" pageId="11" pageNumber="356">Eucosma metamorphica</emphasis>
<bibRefCitation id="EFF9F8FFFFEEFF911BE3F82C3E74F814" author="Meyrick, E." box="[403,606,1931,1953]" firstAuthor="Meyrick" page="439" pageId="11" pageNumber="356" pagination="417 - 448" part="3" refId="ref14767" refString="Meyrick, E. (1928) Exotic Microlepidoptera, 3 (14), 417 - 448." title="Exotic Microlepidoptera" type="journal article" year="1928">Meyrick, 1928: 439</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0AC9A31FFFEEFF911818F82C3F30F814" author="Clarke" box="[616,794,1931,1953]" page="375" pageId="11" pageNumber="356" year="1958">
<bibRefCitation id="EFF9F8FFFFEEFF911818F82C3F30F814" author="Clarke, J. F. G." box="[616,794,1931,1953]" firstAuthor="Clarke" journalOrPublisher="Tortricidae, Olethreutidae, Noctuidae" page="375" pageId="11" pageNumber="356" pagination="1 - 600" refId="ref14308" refString="Clarke, J. F. G. (1958) Catalogue of the Type Specimens of Microlepidoptera in the British Museum (Natural History) described by Edward Meyrick. Vol. III. Tortricidae, Olethreutidae, Noctuidae, 1 - 600." title="Catalogue of the Type Specimens of Microlepidoptera in the British Museum (Natural History) described by Edward Meyrick. Vol. III" type="book chapter" year="1958">Clarke 1958: 375</bibRefCitation>
</treatmentCitation>
, pl. 186, figs. 1, 1a.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD7850EFFEEFF911AE7F80C3F85F875" blockId="11.[151,994,1931,1984]" box="[151,943,1962,1984]" pageId="11" pageNumber="356">
<treatmentCitationGroup id="AB78A220FFEEFF911AE7F80C3F85F875" box="[151,943,1962,1984]" pageId="11" pageNumber="356">
<treatmentCitation id="0AC9A31FFFEEFF911AE7F80C3EACF875" author="Karisch" box="[151,646,1962,1984]" page="462" pageId="11" pageNumber="356" year="2005">
<taxonomicName id="4C68FE8DFFEEFF911AE7F80C3EACF875" ID-CoL="94S3N" authority="Karisch 2005: 462" authorityName="Karisch" authorityPageNumber="462" authorityYear="2005" box="[151,646,1962,1984]" class="Insecta" family="Tortricidae" genus="Dracontogena" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="356" phylum="Arthropoda" rank="species" species="metamorphica">
<emphasis id="B91C591CFFEEFF911AE7F80C3DEBF875" box="[151,449,1962,1984]" italics="true" pageId="11" pageNumber="356">Dracontogena metamorphica</emphasis>
<bibRefCitation id="EFF9F8FFFFEEFF911BB8F80D3EACF875" author="Karisch, T." box="[456,646,1962,1984]" firstAuthor="Karisch" journalOrPublisher="Linzer biologische Beitrage" page="462" pageId="11" pageNumber="356" pagination="457 - 476" part="37" refId="ref14694" refString="Karisch, T. (2005) Ubersicht uber die Gattung Dracontogena Diakonoff 1970 (Lepidoptera: Tortricidae: Olethreutinae: Grapholitini). Linzer biologische Beitrage, 37, 457 - 476." title="Ubersicht uber die Gattung Dracontogena Diakonoff 1970 (Lepidoptera: Tortricidae: Olethreutinae: Grapholitini)" type="journal article" year="2005">Karisch 2005: 462</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, text figs. 13, 15, pl. 1, fig. 8.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C372D685FFE9FF961AE7FF303EA2FF42" pageId="12" pageNumber="357" type="materials_examined">
<paragraph id="8BD7850EFFE9FF961AE7FF303EA2FF42" blockId="12.[151,1437,151,463]" pageId="12" pageNumber="357">
Material examined.
<materialsCitation id="3B008F53FFE9FF961BF1FF3F3D89FF61" ID-GBIF-Occurrence="3712346311" collectionCode="BMNH" collectorName="T. A. Barns" country="Sao Tome and Principe" location="February" municipality="February" pageId="12" pageNumber="357" specimenCode="BMNH 9396" specimenCount="1">
Lectotype of
<taxonomicName id="4C68FE8DFFE9FF96186FFF3E3F19FF05" box="[543,819,152,176]" class="Insecta" family="Tortricidae" genus="Eucosma" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="357" phylum="Arthropoda" rank="species" species="metamorphica">
<emphasis id="B91C591CFFE9FF96186FFF3E3F19FF05" box="[543,819,152,176]" italics="true" pageId="12" pageNumber="357">
<originalDetermination id="A8763DF2FFE9FF96186FFF3E3F19FF05" box="[543,819,152,176]" pageId="12" pageNumber="357">Eucosma metamorphica</originalDetermination>
</emphasis>
</taxonomicName>
, 3,
<collectingCountry id="F37FC59EFFE9FF96191DFF303FCEFF1A" box="[877,996,151,175]" name="Sao Tome and Principe" pageId="12" pageNumber="357">Sao Tomé</collectingCountry>
,
<collectingMunicipality id="6BB31F74FFE9FF961984FF303870FF1A" box="[1012,1114,151,175]" country="Sao Tome and Principe" name="February" pageId="12" pageNumber="357">February</collectingMunicipality>
,
<collectorName id="269DE0D8FFE9FF961E1AFF3F38C1FF1A" box="[1130,1259,152,176]" pageId="12" pageNumber="357">T.A. Barns</collectorName>
, genitalia slide
<specimenCode id="DBCE2D75FFE9FF961AE7FF1A3D06FF61" box="[151,300,188,212]" pageId="12" pageNumber="357">BMNH 9396</specimenCode>
(
<collectionCode id="ED791DCBFFE9FF961B4DFF1B3DBDFF61" box="[317,407,188,212]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="12" pageNumber="357">BMNH</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B008F53FFE9FF961BDDFF1B387DFF61" ID-GBIF-Occurrence="3712346308" box="[429,1111,188,212]" collectionCode="BMNH" pageId="12" pageNumber="357" specimenCount="1">
1Ƥ, same data as lectotype, genitalia slide 30576 (
<collectionCode id="ED791DCBFFE9FF961981FF1B3861FF61" box="[1009,1099,188,212]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="12" pageNumber="357">BMNH</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B008F53FFE9FF961E10FF1B3EA2FF42" ID-GBIF-Occurrence="3712346304" collectionCode="USNM" collectorName="L. Aarvik" pageId="12" pageNumber="357" specimenCount="1" specimenCount-pupa="1">
<specimenCount id="9D6E4E87FFE9FF961E10FF1B38A9FF66" box="[1120,1155,188,212]" pageId="12" pageNumber="357" type="pupa"></specimenCount>
, same data as lectotype, genitalia slide
<collectorName id="269DE0D8FFE9FF961B4AFF473D8DFF42" box="[314,423,223,247]" pageId="12" pageNumber="357">L. Aarvik</collectorName>
2011.025 (
<collectionCode id="ED791DCBFFE9FF961855FF783E56FF42" LSID="urn:lsid:biocol.org:col:34871" box="[549,636,223,247]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="357">USNM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C372D685FFE9FF961AB7FEA3380CFE19" pageId="12" pageNumber="357" type="diagnosis">
<paragraph id="8BD7850EFFE9FF961AB7FEA3380CFE19" blockId="12.[151,1437,151,463]" pageId="12" pageNumber="357">
Diagnosis. Wingspan 20.025.0 mm. Externally,
<taxonomicName id="4C68FE8DFFE9FF96197EFEA23FF2FEA9" box="[782,984,260,284]" class="Insecta" family="Tortricidae" genus="Eucosma" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="357" phylum="Arthropoda" rank="species" species="metamorphica">
<emphasis id="B91C591CFFE9FF96197EFEA23F08FEA9" box="[782,802,261,284]" italics="true" pageId="12" pageNumber="357">D</emphasis>
.
<emphasis id="B91C591CFFE9FF961946FEA33FF2FEA9" box="[822,984,260,284]" italics="true" pageId="12" pageNumber="357">metamorphica</emphasis>
</taxonomicName>
resembles the bicoloured form of
<taxonomicName id="4C68FE8DFFE9FF961FF1FEA23CD4FEF5" class="Insecta" family="Tortricidae" genus="Dracontogena" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="357" phylum="Arthropoda" rank="species" species="kingstoni" status="sp. nov.">
<emphasis id="B91C591CFFE9FF961FF1FEA23CD4FEF5" italics="true" pageId="12" pageNumber="357">D. kingstoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22FE467FFE9FF961B75FE8F3D6BFE8A" box="[261,321,296,319]" pageId="12" pageNumber="357" rank="species">n. sp.</taxonomicNameLabel>
(see below). The male genitalia (
<figureCitation id="1353998BFFE9FF9618CBFE803F27FE8A" box="[699,781,295,319]" captionStart="FIGURES 41 46" captionStartId="15.[151,264,1745,1768]" captionTargetBox="[168,1435,469,1734]" captionTargetId="figure@15.[151,1436,469,1757]" captionTargetPageId="15" captionText="FIGURES 41 46. Male genitalia in Dracontogena. 41. D. bernardi Karisch. (slide L. Aarvik 2808). 42. D. bernardi Karisch, phallus (slide L. Aarvik 2808). 43. D. solii n. sp. (slide NHMO 2010). 44. D. solii n. sp. phallus (slide NHMO 2010). 45. D. tonitrualis (Meyrick). 46. D. metamorphica (Meyrick). Scale figs. 41, 43: 1 mm., figs. 42, 44: 0.5 mm. Figs. 45 46 from Karisch (2005)." httpUri="https://zenodo.org/record/282275/files/figure.png" pageId="12" pageNumber="357">Fig. 46</figureCitation>
) are distinct by the deeply excavate ventral edge of valva. The female genitalia (
<figureCitation id="1353998BFFE9FF961BE6FEEB3E32FED1" box="[406,536,332,356]" captionStart="FIGURES 60 63" captionStartId="19.[151,264,1924,1947]" captionTargetBox="[177,1407,216,1880]" captionTargetId="figure@19.[161,1425,193,1903]" captionTargetPageId="19" captionText="FIGURES 60 63. Female genitalia of Dracontogena. 60. D. agassizi n. sp. (slide USNM 137,547). 61. D. agassizi n. sp. sterigma and sternite 7 (slide T. Karisch 1923). 62 63. D. metamorphica (Meyrick) (slide L. Aarvik 2011.025). Scale figs. 60, 62: 1 mm., figs. 61, 63: 0.5 mm." httpUri="https://zenodo.org/record/282278/files/figure.png" pageId="12" pageNumber="357">Fig. 62, 63</figureCitation>
) have sternite 7 deeply excavate with two membranous bands anterior of the excavation; segment 7 with conspicuous scale sockets anterolaterally; the sterigma large, becoming wider posteriorly; and the paired oval sclerites posterior of the sterigma small, separate.
</paragraph>
</subSubSection>
<subSubSection id="C372D685FFE9FF961AB7FE10388AFE7A" box="[199,1184,439,463]" pageId="12" pageNumber="357" type="distribution">
<paragraph id="8BD7850EFFE9FF961AB7FE10388AFE7A" blockId="12.[151,1437,151,463]" box="[199,1184,439,463]" pageId="12" pageNumber="357">
Distribution. Known only from
<collectingCountry id="F37FC59EFFE9FF96185CFE103EB5FE7A" box="[556,671,439,463]" name="Sao Tome and Principe" pageId="12" pageNumber="357">Sao Tomé</collectingCountry>
, an island in the Gulf of
<collectingCountry id="F37FC59EFFE9FF9619C4FE103822FE7A" box="[948,1032,439,463]" name="Guinea-Bissau" pageId="12" pageNumber="357">Guinea</collectingCountry>
, West Africa.
</paragraph>
</subSubSection>
</treatment>
</document>