treatments-xml/data/EC/0B/70/EC0B7049FF904123FF55FF71CC4A1B68.xml

<document id="56B44B4A28CFBD11D7EB43E043DA0963" ID-CLB-Dataset="46650" ID-DOI="10.5281/zenodo.277908" ID-GBIF-Dataset="5351b061-b2f0-435e-87f4-73e0db3bfa9b" ID-ISSN="1175-5326" ID-Zenodo-Dep="277908" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460370290096" checkinUser="plazi" docAuthor="Rojas-Runjaic, Fernando J. M., Infante-Rivero, Edwin E. &amp; Barrio-Amorós, César L." docDate="2011" docId="EC0B7049FF904123FF55FF71CC4A1B68" docLanguage="en" docName="zt02919p050.pdf" docOrigin="Zootaxa 2919" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Aromobates tokuko Rojas-Runjaic, Infante-Rivero &amp; Barrio-Amorós, 2011, sp. nov." docType="treatment" docVersion="10" lastPageNumber="48" masterDocId="10320831FF924128FFC2FFE6CD50187F" masterDocTitle="A new frog of the genus Aromobates (Anura, Dendrobatidae) from Sierra de Perijá, Venezuela" masterLastPageNumber="50" masterPageNumber="37" pageNumber="39" updateTime="1698295642514" updateUser="ExternalLinkService">
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<mods:title id="2C9110B69A3640BE28D85771030E824A">A new frog of the genus Aromobates (Anura, Dendrobatidae) from Sierra de Perijá, Venezuela</mods:title>
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<mods:namePart id="8E553DB4030464B093D9E57D8E42D0D3">Rojas-Runjaic, Fernando J. M.</mods:namePart>
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<mods:namePart id="3C565F8CFCABB5E2333B2AD533ACC915">Infante-Rivero, Edwin E.</mods:namePart>
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<mods:namePart id="028F0E1854691B8443E65155D2FBDE93">Barrio-Amorós, César L.</mods:namePart>
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<mods:date id="D9F7DBB1FA12724DEF382C3E6FB16925">2011</mods:date>
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<treatment id="EC0B7049FF904123FF55FF71CC4A1B68" ID-DOI="http://doi.org/10.5281/zenodo.5658016" ID-GBIF-Taxon="119411637" ID-Zenodo-Dep="5658016" LSID="urn:lsid:plazi:treatment:EC0B7049FF904123FF55FF71CC4A1B68" httpUri="http://treatment.plazi.org/id/EC0B7049FF904123FF55FF71CC4A1B68" lastPageId="11" lastPageNumber="48" pageId="2" pageNumber="39">
<subSubSection id="2CB892D4FF90412AFF55FF71CCDF1963" pageId="2" pageNumber="39" type="nomenclature">
<paragraph id="641DC15FFF90412AFF55FF71CC8818CF" blockId="2.[151,582,151,284]" box="[151,472,151,176]" pageId="2" pageNumber="39">
<heading id="3F557633FF90412AFF55FF71CC8818CF" bold="true" box="[151,472,151,176]" fontSize="10" level="2" pageId="2" pageNumber="39" reason="2">
<emphasis id="56D61D4DFF90412AFF55FF71CC8818CF" bold="true" box="[151,472,151,176]" pageId="2" pageNumber="39">
<taxonomicName id="A3A2BADCFF90412AFF55FF71CC2718CF" ID-CoL="GTRY" box="[151,375,151,176]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="2" pageNumber="39" phylum="Chordata" rank="species" species="tokuko" status="sp. nov.">
<emphasis id="56D61D4DFF90412AFF55FF71CC2718CF" bold="true" box="[151,375,151,176]" italics="true" pageId="2" pageNumber="39">Aromobates tokuko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4DE5A036FF90412AFEBCFF7ECC8818CF" box="[382,472,152,176]" pageId="2" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="641DC15FFF90412AFF55FF5ACCDF1963" blockId="2.[151,582,151,284]" pageId="2" pageNumber="39">
English name: Perija’s Nurse Frog Spanish name: Sapito Niñera de Perijá (
<figureCitation id="FC99DDDAFF90412AFF5CFEE2CC761963" box="[158,294,260,284]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1985,2008]" captionStartId-1="3.[151,250,1710,1733]" captionStartId-2="8.[151,250,1860,1883]" captionTargetBox-0="[151,1436,483,1961]" captionTargetBox-1="[151,1436,193,1670]" captionTargetBox-2="[151,1436,193,1838]" captionTargetId-0="figure@2.[151,1436,483,1961]" captionTargetId-1="figure@3.[151,1436,193,1671]" captionTargetId-2="figure@8.[151,1436,193,1839]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="8" captionText-0="FIGURE 1. Aromobates tokuko sp. nov. Dorsal (a) and ventral (b) views of the male holotype (MHNLS 18479). Dorsal (c) and ventral (d) views of the female paratype (MHNLS 18568). Scale bars represent 10 mm." captionText-1="FIGURE 2. Aromobates tokuko sp. nov. (male holotype, MHNLS 18479). Ventral (a), dorsal (b), and lateral (c) views of the head. Details of left hand (d), and left foot (e). Scale bars represent 2 mm." captionText-2="FIGURE 3. Color pattern variation in Aromobates tokuko sp. nov. (a) male paratype (MHNLS 18499) with well-defined dorsolateral stripes and dorsal pattern; (b) female paratype (MHNLS 18490) with uniform dorsal and lateral color, and incomplete dorsolateral stripe." httpUri-0="https://zenodo.org/record/277909/files/figure.png" httpUri-1="https://zenodo.org/record/277910/files/figure.png" httpUri-2="https://zenodo.org/record/277911/files/figure.png" pageId="2" pageNumber="39">Figures 1–3</figureCitation>
, 
<tableCitation id="2920F4E4FF90412AFEF1FEE2CCD71963" box="[307,391,260,284]" captionStart="TABLE 2" captionStartId="7.[151,235,766,787]" captionTargetBox="[151,1435,851,1630]" captionTargetPageId="7" captionText="TABLE 2. Morphometric variation (in mm) in the type series of Aromobates tokuko sp. nov. ‾ X ± SD: mean ± standard deviation; Min – Max: minimum – maximum." httpUri="http://table.plazi.org/id/30DD91D7FF95412FFF55FD18CF4B1B48" pageId="2" pageNumber="39" tableUuid="30DD91D7FF95412FFF55FD18CF4B1B48">Table 2</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="2CB892D4FF90412CFF55FEAACEB519B0" lastPageId="4" lastPageNumber="41" pageId="2" pageNumber="39" type="materials_examined">
<paragraph id="641DC15FFF90412AFF55FEAACEA519D3" blockId="2.[151,1436,332,428]" pageId="2" pageNumber="39">
<emphasis id="56D61D4DFF90412AFF55FEAACC5B191B" bold="true" box="[151,267,332,356]" pageId="2" pageNumber="39">
<typeStatus id="BB197FFDFF90412AFF55FEAACC56191B" box="[151,262,332,356]" pageId="2" pageNumber="39" type="holotype">Holotype</typeStatus>
.
</emphasis>
Adult male, 
<collectionCode id="02B3599AFF90412AFE5EFEAACCAE191B" box="[412,510,332,356]" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="2" pageNumber="39">MHNLS</collectionCode>
18479, from the surroundings of Ipika, Yukpa indigenous community, Río 
<taxonomicName id="A3A2BADCFF90412AFA85FEAAC8CC191B" box="[1351,1436,332,356]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="2" pageNumber="48" phylum="Chordata" rank="species" species="tokuko">Tokuko</taxonomicName>
basin, Municipio Machiques de Perijá, Sierra de Perijá, Estado Zulia, 
<collectingCountry id="1CB581CFFF90412AFC75FE89C97A19F8" box="[951,1066,367,391]" name="Venezuela" pageId="2" pageNumber="39">Venezuela</collectingCountry>
(
<geoCoordinate id="0196A798FF90412AFBFEFE89C9F919F8" box="[1084,1193,367,391]" direction="north" orientation="latitude" pageId="2" pageNumber="39" precision="925" value="9.866667">09°52’ N</geoCoordinate>
, 
<geoCoordinate id="0196A798FF90412AFB74FE89C87919F8" box="[1206,1321,367,391]" direction="west" orientation="longitude" pageId="2" pageNumber="39" precision="925" value="-72.85">72°51’ W</geoCoordinate>
; elevation 
<quantity id="A35A6CBAFF90412AFF55FE72CDB119D3" box="[151,225,404,428]" metricMagnitude="2" metricUnit="m" metricValue="5.95" pageId="2" pageNumber="39" unit="m" value="595.0">595 m</quantity>
), collected on 
<date id="101CE79FFF90412AFE46FE72CF0F19D3" box="[388,607,404,428]" pageId="2" pageNumber="39" value="2006-11-26">26 November 2006</date>
, by F. Rojas-Runjaic and E. Infante.
</paragraph>
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<paragraph id="641DC15FFF90412AFF55F827C95D1F89" blockId="2.[151,1437,1985,2038]" pageId="2" pageNumber="39">
<emphasis id="56D61D4DFF90412AFF55F827CC421FA8" bold="true" box="[151,274,1985,2008]" pageId="2" pageNumber="39">FIGURE 1.</emphasis>
<taxonomicName id="A3A2BADCFF90412AFEDAF827CC8A1FA8" box="[280,474,1985,2007]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="2" pageNumber="39" phylum="Chordata" rank="species" species="tokuko" status="sp. nov.">
<emphasis id="56D61D4DFF90412AFEDAF827CC8A1FA8" box="[280,474,1985,2007]" italics="true" pageId="2" pageNumber="39">Aromobates tokuko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4DE5A036FF90412AFE22F824CF601FA8" box="[480,560,1986,2007]" pageId="2" pageNumber="39" rank="species">
<emphasis id="56D61D4DFF90412AFE22F824CF601FA8" bold="true" box="[480,560,1986,2007]" pageId="2" pageNumber="39">sp. nov.</emphasis>
</taxonomicNameLabel>
Dorsal (a) and ventral (b) views of the male holotype (MHNLS 18479). Dorsal (c) and ventral (d) views of the female paratype (MHNLS 18568). Scale bars represent 10 mm.
</paragraph>
</caption>
<caption id="30DD91D7FF91412BFF55F948CED41E9C" httpUri="https://zenodo.org/record/277910/files/figure.png" pageId="3" pageNumber="40" targetBox="[151,1436,193,1670]" targetPageId="3">
<paragraph id="641DC15FFF91412BFF55F948CED41E9C" blockId="3.[151,1436,1710,1763]" pageId="3" pageNumber="40">
<emphasis id="56D61D4DFF91412BFF55F948CC431EBC" bold="true" box="[151,275,1710,1733]" pageId="3" pageNumber="40">FIGURE 2.</emphasis>
<taxonomicName id="A3A2BADCFF91412BFED9F948CC8F1EBB" box="[283,479,1710,1732]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="3" pageNumber="40" phylum="Chordata" rank="species" species="tokuko" status="sp. nov.">
<emphasis id="56D61D4DFF91412BFED9F948CC8F1EBB" box="[283,479,1710,1732]" italics="true" pageId="3" pageNumber="40">Aromobates tokuko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4DE5A036FF91412BFE25F949CF681EBB" box="[487,568,1711,1732]" pageId="3" pageNumber="40" rank="species">
<emphasis id="56D61D4DFF91412BFE25F949CF681EBB" bold="true" box="[487,568,1711,1732]" pageId="3" pageNumber="40">sp. nov.</emphasis>
</taxonomicNameLabel>
(male holotype, MHNLS 18479). Ventral (a), dorsal (b), and lateral (c) views of the head. Details of left hand (d), and left foot (e). Scale bars represent 2 mm.
</paragraph>
</caption>
<paragraph id="641DC15FFF91412CFF07F8F6C806191B" blockId="3.[151,1436,1807,2011]" lastBlockId="4.[151,1437,151,2012]" lastPageId="4" lastPageNumber="41" pageId="3" pageNumber="40">
<emphasis id="56D61D4DFF91412BFF07F8F6CC141F57" bold="true" box="[197,324,1808,1832]" pageId="3" pageNumber="40">
<typeStatus id="BB197FFDFF91412BFF07F8F6CC6F1F57" box="[197,319,1808,1832]" pageId="3" pageNumber="40" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Twenty-one specimens: four adult females (
<collectionCode id="02B3599AFF91412BFC86F8E9CEC81F58" box="[836,920,1807,1831]" httpUri="http://grbio.org/cool/4jmm-iazx" name="Musee Guimet d'Histoire Naturelle de Lyon" pageId="3" pageNumber="40">MHNL</collectionCode>
18471, 18474, 18478, 18480) and two adult males (
<collectionCode id="02B3599AFF91412BFF25F8D5CC1B1F34" box="[231,331,1843,1867]" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="3" pageNumber="40">MHNLS</collectionCode>
18473, 18476) with the same collection data as the 
<typeStatus id="BB197FFDFF91412BFC50F8D5CEA81F34" box="[914,1016,1843,1867]" pageId="3" pageNumber="40" type="holotype">holotype</typeStatus>
; four adult females (
<collectionCode id="02B3599AFF91412BFB1FF8D5C8111F34" box="[1245,1345,1843,1867]" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="3" pageNumber="40">MHNLS</collectionCode>
18490– 91, 18493, 18495) collected on 
<date id="101CE79FFF91412BFE3FF8B1CF871F10" box="[509,727,1879,1903]" pageId="3" pageNumber="40" value="2006-12-27">27 December 2006</date>
, at the 
<typeStatus id="BB197FFDFF91412BFCEBF8B1CEDB1F10" box="[809,907,1879,1903]" pageId="3" pageNumber="40" type="holotype">holotype</typeStatus>
locality; two adult males (
<collectionCode id="02B3599AFF91412BFB75F8B1C84B1F10" box="[1207,1307,1879,1903]" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="3" pageNumber="40">MHNLS</collectionCode>
18498–99) from Kusare waterfall, near Ipika, Río Tukuko basin, Municipio Machiques de Perijá, Sierra de Perijá, Estado Zulia, 
<collectingCountry id="1CB581CFFF91412BFF22F879CC031FC8" box="[224,339,1951,1975]" name="Venezuela" pageId="3" pageNumber="40">Venezuela</collectingCountry>
(
<geoCoordinate id="0196A798FF91412BFEA1F879CC9E1FC7" box="[355,462,1951,1976]" direction="north" orientation="latitude" pageId="3" pageNumber="40" precision="925" value="9.866667">09°52’ N</geoCoordinate>
, 
<geoCoordinate id="0196A798FF91412BFE18F879CF1A1FC7" box="[474,586,1951,1976]" direction="west" orientation="longitude" pageId="3" pageNumber="40" precision="925" value="-72.833336">72°50’ W</geoCoordinate>
; elevation 
<quantity id="A35A6CBAFF91412BFD01F879CE5B1FC8" box="[707,779,1951,1975]" metricMagnitude="2" metricUnit="m" metricValue="5.79" pageId="3" pageNumber="40" unit="m" value="579.0">579 m</quantity>
), collected on 
<date id="101CE79FFF91412BFC73F879C9D61FC8" box="[945,1158,1951,1975]" pageId="3" pageNumber="40" value="2006-12-28">28 December 2006</date>
by F. Rojas-Runjaic and E. Infante; three adult females (
<collectionCode id="02B3599AFF91412BFDC6F825CF381FA4" box="[516,616,1987,2011]" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="3" pageNumber="40">MHNLS</collectionCode>
18521–23) and one adult male (
<collectionCode id="02B3599AFF91412BFC20F825C9161FA4" box="[994,1094,1987,2011]" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="3" pageNumber="40">MHNLS</collectionCode>
18520) collected on 
<date id="101CE79FFF91412CFAFEF825CD8218D0" lastPageId="4" lastPageNumber="41" pageId="3" pageNumber="40" value="2007-04-11">11 April 2007</date>
, with the same collection data as the 
<typeStatus id="BB197FFDFF96412CFD52FF71CFA618D0" box="[656,758,151,175]" pageId="4" pageNumber="41" type="holotype">holotype</typeStatus>
; one adult female (
<collectionCode id="02B3599AFF96412CFC1BFF71C96D18D0" box="[985,1085,151,175]" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="4" pageNumber="41">MHNLS</collectionCode>
18568) and three adult males (
<collectionCode id="02B3599AFF96412CFF5CFF5ACC5218AB" box="[158,258,188,212]" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="4" pageNumber="41">MHNLS</collectionCode>
18566–67, 18569), from Kiriponsa, Yukpa indigenous community, Río 
<taxonomicName id="A3A2BADCFF96412CFBB0FF5AC99718AB" box="[1138,1223,188,212]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="4" pageNumber="48" phylum="Chordata" rank="species" species="tokuko">Tokuko</taxonomicName>
basin, Municipio Machiques de Perijá, Sierra de Perijá, Estado Zulia, 
<collectingCountry id="1CB581CFFF96412CFD24FF39CE091888" box="[742,857,223,247]" name="Venezuela" pageId="4" pageNumber="41">Venezuela</collectingCountry>
(
<geoCoordinate id="0196A798FF96412CFCA8FF39CE841888" box="[874,980,223,247]" direction="north" orientation="latitude" pageId="4" pageNumber="41" precision="925" value="9.866667">09º52’ N</geoCoordinate>
, 
<geoCoordinate id="0196A798FF96412CFC22FF39C91F1888" box="[992,1103,223,247]" direction="west" orientation="longitude" pageId="4" pageNumber="41" precision="925" value="-72.86667">72º52’ W</geoCoordinate>
; elevation 
<quantity id="A35A6CBAFF96412CFB0BFF39C8721888" box="[1225,1314,223,247]" metricMagnitude="3" metricUnit="m" metricValue="1.005" pageId="4" pageNumber="41" unit="m" value="1005.0">1005 m</quantity>
), collected on 
<date id="101CE79FFF96412CFF7BFEE2CC051963" box="[185,341,260,284]" pageId="4" pageNumber="41" value="2007-04-12">12 April 2007</date>
by F. Rojas-Runjaic and E. Infante; one adult female (
<collectionCode id="02B3599AFF96412CFC73FEE2C9451963" box="[945,1045,260,284]" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="4" pageNumber="41">MHNLS</collectionCode>
19748) from near Terakibu, Yukpa indigenous community, Río 
<taxonomicName id="A3A2BADCFF96412CFE1CFEC1CF661940" box="[478,566,295,319]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="4" pageNumber="48" phylum="Chordata" rank="species" species="tokuko">Tokuko</taxonomicName>
, Municipio Machiques de Perijá, Sierra de Perijá, Estado Zulia, 
<collectingCountry id="1CB581CFFF96412CFAEBFEC1C8CC1940" box="[1321,1436,295,319]" name="Venezuela" pageId="4" pageNumber="41">Venezuela</collectingCountry>
(
<geoCoordinate id="0196A798FF96412CFF5DFEAACC58191B" box="[159,264,332,356]" direction="north" orientation="latitude" pageId="4" pageNumber="41" precision="925" value="9.866667">09º52’ N</geoCoordinate>
, 
<geoCoordinate id="0196A798FF96412CFED1FEAACCD1191B" box="[275,385,332,356]" direction="west" orientation="longitude" pageId="4" pageNumber="41" precision="925" value="-72.816666">72º49’ W</geoCoordinate>
; elevation 
<quantity id="A35A6CBAFF96412CFE38FEAACF14191B" box="[506,580,332,356]" metricMagnitude="2" metricUnit="m" metricValue="4.1899999999999995" pageId="4" pageNumber="41" unit="m" value="419.0">419 m</quantity>
), collected on 
<date id="101CE79FFF96412CFD25FEAACE91191B" box="[743,961,332,356]" pageId="4" pageNumber="41" value="2008-09-16">16 September 2008</date>
by F. Rojas-Runjaic and P. Cabello.
</paragraph>
<paragraph id="641DC15FFF96412CFF07FE89CEB519B0" blockId="4.[151,1437,151,2012]" pageId="4" pageNumber="41">
<emphasis id="56D61D4DFF96412CFF07FE89CCE919F7" bold="true" box="[197,441,367,392]" pageId="4" pageNumber="41">Referred specimens.</emphasis>
Eight specimens: four juveniles (
<collectionCode id="02B3599AFF96412CFCF8FE89CECE19F8" box="[826,926,367,391]" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="4" pageNumber="41">MHNLS</collectionCode>
18477, 18481–83), with the same collection data as the 
<typeStatus id="BB197FFDFF96412CFED4FE72CC2B19D3" box="[278,379,404,428]" pageId="4" pageNumber="41" type="holotype">holotype</typeStatus>
; three juveniles (
<collectionCode id="02B3599AFF96412CFDF8FE72CFCE19D3" box="[570,670,404,428]" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="4" pageNumber="41">MHNLS</collectionCode>
18492, 18494, 18496) and one premetamorph (
<collectionCode id="02B3599AFF96412CFB75FE72C85B19D3" box="[1207,1291,404,428]" httpUri="http://grbio.org/cool/4jmm-iazx" name="Musee Guimet d'Histoire Naturelle de Lyon" pageId="4" pageNumber="41">MHNL</collectionCode>
18497), collected on 
<date id="101CE79FFF96412CFEC1FE51CC8C19B0" box="[259,476,439,463]" pageId="4" pageNumber="41" value="2006-12-27">27 December 2006</date>
, with the same collection data as the 
<typeStatus id="BB197FFDFF96412CFCBEFE51CEB219B0" box="[892,994,439,463]" pageId="4" pageNumber="41" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="2CB892D4FF96412DFF07FE3DCEB41AB3" lastPageId="5" lastPageNumber="42" pageId="4" pageNumber="41" type="diagnosis">
<paragraph id="641DC15FFF96412CFF07FE3DCF7A1D07" blockId="4.[151,1437,151,2012]" pageId="4" pageNumber="41">
<emphasis id="56D61D4DFF96412CFF07FE3DCC6F198B" bold="true" box="[197,319,475,500]" pageId="4" pageNumber="41">Diagnosis.</emphasis>
(1) Skin of posterior third of dorsum, flanks, prehumeral area, dorsum of thighs and shanks granular; (2) Paired dorsal digital scutes conspicuous and protuberant on all toes; (3) Distal subarticular tubercle under FIV circular and conspicuous; (4) FIV extending past distal subarticular tubercle of FIII; (5) FI shorter than FII; (6) Digital discs present; (7) Finger discs moderately expanded; (8) Lateral fringes absent on FI and FIV, preaxials on FII and FIII, present and conspicuous on all toes; (9) Metacarpal ridge absent or, if present, very weak and almost indistinct; (10) FIII of adult males not swollen; (11) Carpal pad absent; (12) Male excrescences on thumb absent; (13) Thenar tubercle small (approximately 1/3 of the palmar tubercle and equal in size and shape to the proximal subarticular tubercle of FII) elliptical and conspicuous; (14) Black arm gland in adult males absent; (15) Tarsal keel present, well-defined and slightly curved from proximal edge of inner metatarsal tubercle, transverse across tarsus, and reaching almost the mid-tarsus where it becomes slightly wider; (16) Toe discs slightly expanded, disc of TI barely wider than distal end of adjacent phalanx (including fringes), discs of TII–TV about 1.5 times wider than distal end of adjacent phalanx; (17) Toe webbing basal between TI–TIV; (18) Metatarsal fold flat and very weak, almost indistinct; (19) dorsum uniformly dark brown or with a pale brown background and irregularly-spaced dark brown spots; (20) Dorsolateral stripe present, pale golden to cream, from tip of snout to midbody or to upper level of the groin, eventually continual with paracloacal horizontal stripes; (21) Oblique lateral stripe present but diffuse, formed by small whitish spots coincident with small non-keratinized tubercles, extending from groin to midbody; (22) Paracloacal horizontal stripe present, diffuse to very well defined; (23) Ventrolateral stripe absent; (24) Throat grayish ivory (with profusion of evenly spaced melanophores when seen under magnification) on almost all males and some females, immaculate ivory on almost all females and some males; (25) Throat collar absent; (26) Chest and belly white to ivory, rarely with pale gray or pale brown spots on chest and anterior third of belly; (27) Ventral surface of arms, thighs, shanks and tarsi pale orange; (28) Palms and soles pale gray; (29) Axillae, groins and hidden parts of hind limbs weakly yellowish; (30) Iris golden and finely dotted with black, with a transversal wide blackish band and a bright golden pupil ring; (31) Large intestine unpigmented, creamish white; (32) Adult testis whitish; (33) Median lingual process absent; (34) Tympanum ill-defined, about 1/4 concealed posterodorsally; (35) Vocal sac structure not distinct; (36) Teeth present on the maxillary arch; (37) Small size, with males up to 
<quantity id="A35A6CBAFF96412CFAFBFADAC8CC1D2B" box="[1337,1436,1340,1364]" metricMagnitude="-2" metricUnit="m" metricValue="2.32" pageId="4" pageNumber="41" unit="mm" value="23.2">23.2 mm</quantity>
of SVL and females up to 
<quantity id="A35A6CBAFF96412CFE7CFA86CF751D07" box="[446,549,1376,1400]" metricMagnitude="-2" metricUnit="m" metricValue="2.92" pageId="4" pageNumber="41" unit="mm" value="29.2">29.2 mm</quantity>
.
</paragraph>
<paragraph id="641DC15FFF96412CFF07FA65CED71FA3" blockId="4.[151,1437,151,2012]" pageId="4" pageNumber="41">
<emphasis id="56D61D4DFF96412CFF07FA65CF6E1DE3" bold="true" box="[197,574,1411,1436]" pageId="4" pageNumber="41">Comparison with other species.</emphasis>
<taxonomicName id="A3A2BADCFF96412CFD87FA62CE4E1DE3" box="[581,798,1412,1436]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="4" pageNumber="41" phylum="Chordata" rank="species" species="tokuko">
<emphasis id="56D61D4DFF96412CFD87FA62CE4E1DE3" box="[581,798,1412,1436]" italics="true" pageId="4" pageNumber="41">Aromobates tokuko</emphasis>
</taxonomicName>
(characters in parentheses) differs by its small size from large species of the genus, such as 
<taxonomicName id="A3A2BADCFF96412CFDDCFA4ECECF1DBF" authority="Boulenger, 1903" authorityName="Boulenger" authorityYear="1903" box="[542,927,1448,1472]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="4" pageNumber="41" phylum="Chordata" rank="species" species="alboguttatus">
<emphasis id="56D61D4DFF96412CFDDCFA4ECF991DBF" box="[542,713,1448,1472]" italics="true" pageId="4" pageNumber="41">A. alboguttatus</emphasis>
(Boulenger, 1903)
</taxonomicName>
, 
<taxonomicName id="A3A2BADCFF96412CFC69FA4EC9A71DBF" authority="Rivero, 1978" authorityName="Rivero" authorityYear="1978" box="[939,1271,1448,1472]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="4" pageNumber="41" phylum="Chordata" rank="species" species="leopardalis">
<emphasis id="56D61D4DFF96412CFC69FA4EC9191DBF" box="[939,1097,1448,1472]" italics="true" pageId="4" pageNumber="41">A. leopardalis</emphasis>
(
<bibRefCitation id="0033BCAEFF96412CFB95FA4EC9A01DBF" author="Rivero" box="[1111,1264,1448,1472]" pageId="4" pageNumber="41" refString="Rivero, J. A. (1978) Notas sobre los anfibios de Venezuela II. Sobre los Colostethus de los Andes Venezolanos. Memoria de la Sociedad de Ciencias Naturales La Salle, 35, 327 - 344." type="journal article" year="1978">Rivero, 1978</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="A3A2BADCFF96412CFAC1FA4ECCCC1D9B" authority="Dole &amp; Durant, 1973" authorityName="Dole &amp; Durant" authorityYear="1973" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="4" pageNumber="41" phylum="Chordata" rank="species" species="meridensis">
<emphasis id="56D61D4DFF96412CFAC1FA4EC8CB1DBF" box="[1283,1435,1448,1472]" italics="true" pageId="4" pageNumber="41">A. meridensis</emphasis>
(Dole &amp; Durant, 1973)
</taxonomicName>
, 
<taxonomicName id="A3A2BADCFF96412CFE6AFA2ACEF31D9B" authority="Myers, Paolillo &amp; Daly, 1991" authorityName="Myers, Paolillo &amp; Daly" authorityYear="1991" box="[424,931,1484,1508]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="4" pageNumber="41" phylum="Chordata" rank="species" species="nocturnus">
<emphasis id="56D61D4DFF96412CFE6AFA2ACF681D9B" box="[424,568,1484,1508]" italics="true" pageId="4" pageNumber="41">A. nocturnus</emphasis>
(
<bibRefCitation id="0033BCAEFF96412CFD8AFA2ACECA1D9B" author="Myers" box="[584,922,1484,1508]" pageId="4" pageNumber="41" refString="Myers, C. W., Paolillo, A. &amp; Daly, J. W. (1991) Discovery of a defensively malodorous and nocturnal frog in the family Dendrobatidae: phylogenetic significances of a new genus and species from the Venezuelan Andes. American Museum Novitates, 3002, 1 - 33." type="journal article" year="1991">Myers, Paolillo &amp; Daly, 1991</bibRefCitation>
)
</taxonomicName>
, and 
<taxonomicName id="A3A2BADCFF96412CFC1DFA2AC86A1D9B" authority="Pefaur, 1993" authorityName="Pefaur" authorityYear="1993" box="[991,1338,1484,1508]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="4" pageNumber="41" phylum="Chordata" rank="species" species="capurinensis">
<emphasis id="56D61D4DFF96412CFC1DFA2AC9DE1D9B" box="[991,1166,1484,1508]" italics="true" pageId="4" pageNumber="41">A. capurinensis</emphasis>
(
<bibRefCitation id="0033BCAEFF96412CFB5CFA2AC8611D9B" author="Pefaur" box="[1182,1329,1484,1508]" pageId="4" pageNumber="41" refString="Pefaur, J. E. (1993) Description of a new Colostethus (Dendrobatidae) with some natural history on the genus in Venezuela. Alytes, 11, 88 - 96." type="journal article" year="1993">Péfaur, 1993</bibRefCitation>
)
</taxonomicName>
, all with maximum SVL&gt; 
<quantity id="A35A6CBAFF96412CFEA0FA16CCE11E77" box="[354,433,1520,1544]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="4" pageNumber="41" unit="mm" value="31.0">31 mm</quantity>
(up to 
<quantity id="A35A6CBAFF96412CFDC3FA16CF381E77" box="[513,616,1520,1544]" metricMagnitude="-2" metricUnit="m" metricValue="2.92" pageId="4" pageNumber="41" unit="mm" value="29.2">29.2 mm</quantity>
). 
<taxonomicName id="A3A2BADCFF96412CFDB9FA16CEC51E77" box="[635,917,1520,1544]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="4" pageNumber="41" phylum="Chordata" rank="species" species="alboguttatus">
<emphasis id="56D61D4DFF96412CFDB9FA16CEC51E77" box="[635,917,1520,1544]" italics="true" pageId="4" pageNumber="41">Aromobates alboguttatus</emphasis>
</taxonomicName>
has smooth dorsal skin (granular) and its venter is brown spotted with white (ivory white). 
<taxonomicName id="A3A2BADCFF96412CFD06F9F2CE851E53" box="[708,981,1556,1580]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="4" pageNumber="41" phylum="Chordata" rank="species" species="leopardalis">
<emphasis id="56D61D4DFF96412CFD06F9F2CE851E53" box="[708,981,1556,1580]" italics="true" pageId="4" pageNumber="41">Aromobates leopardalis</emphasis>
</taxonomicName>
has toes 1/3 webbed (basally webbed between TI–TIV), dorsal skin smooth with a few tubercles posteriorly (granular), a strongly spotted dorsum (without consistent pattern). 
<taxonomicName id="A3A2BADCFF96412CFE5EF9BACFF21E0B" box="[412,674,1628,1652]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="4" pageNumber="41" phylum="Chordata" rank="species" species="meridensis">
<emphasis id="56D61D4DFF96412CFE5EF9BACFF21E0B" box="[412,674,1628,1652]" italics="true" pageId="4" pageNumber="41">Aromobates meridensis</emphasis>
</taxonomicName>
has FI slightly longer than FII (FI shorter than FII), subarticular distal tubercle of FIV indistinct (conspicuous), fringes absent on fingers (preaxial fringes on FII and FIII), tarsal keel straight and indistinct (slightly curved and well defined), dorsolateral stripes absent or, if present, very ill-defined (dorsolateral stripes always present, generally well-defined), and belly bright yellow with diffuse dark reticulation (belly immaculate ivory, occasionally with diffuse pale brown or grayish spots on anterior half, never reticulated). 
<taxonomicName id="A3A2BADCFF96412CFF55F8F6CCC81F57" box="[151,408,1808,1832]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="4" pageNumber="41" phylum="Chordata" rank="species" species="nocturnus">
<emphasis id="56D61D4DFF96412CFF55F8F6CCC81F57" box="[151,408,1808,1832]" italics="true" pageId="4" pageNumber="41">Aromobates nocturnus</emphasis>
</taxonomicName>
is the most peculiar and largest congener, with SVL reaching 
<quantity id="A35A6CBAFF96412CFBBFF8F6C9801F57" box="[1149,1232,1808,1832]" metricMagnitude="-2" metricUnit="m" metricValue="6.2" pageId="4" pageNumber="41" unit="mm" value="62.0">62 mm</quantity>
(
<quantity id="A35A6CBAFF96412CFB20F8F6C8001F57" box="[1250,1360,1808,1832]" metricMagnitude="-2" metricUnit="m" metricValue="2.92" pageId="4" pageNumber="41" unit="mm" value="29.2">29.2 mm</quantity>
), fully webbed toes (basally webbed), belly coloration mottled with grayish white in life (immaculate ivory, occasionally with diffuse pale brown or grayish spots on anterior third), and unique nocturnal (diurnal), aquatic (riparian), and releasing a mercaptanlike odor when molested (absent). 
<taxonomicName id="A3A2BADCFF96412CFCDCF89AC96D1FEB" box="[798,1085,1916,1940]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="4" pageNumber="41" phylum="Chordata" rank="species" species="capurinensis">
<emphasis id="56D61D4DFF96412CFCDCF89AC96D1FEB" box="[798,1085,1916,1940]" italics="true" pageId="4" pageNumber="41">Aromobates capurinensis</emphasis>
</taxonomicName>
is a larger frog, mean SVL of females 
<quantity id="A35A6CBAFF96412CFF35F846CC0E1FC7" box="[247,350,1952,1976]" metricMagnitude="-2" metricUnit="m" metricValue="3.34" pageId="4" pageNumber="41" unit="mm" value="33.4">33.4 mm</quantity>
(
<quantity id="A35A6CBAFF96412CFEADF846CC8B1FC7" box="[367,475,1952,1976]" metricMagnitude="-2" metricUnit="m" metricValue="2.55" pageId="4" pageNumber="41" unit="mm" value="25.5">25.5 mm</quantity>
), with dorsal skin smooth (granular), dorsolateral stripes present but indistinct (distinct), oblique lateral stripe absent (formed by small whitish spots).
</paragraph>
<paragraph id="641DC15FFF97412DFF07FF71CEB41AB3" blockId="5.[151,1437,151,2012]" pageId="5" pageNumber="42">
The seven other species of 
<taxonomicName id="A3A2BADCFF97412DFE3CFF7ECFD418CF" box="[510,644,152,176]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="genus">
<emphasis id="56D61D4DFF97412DFE3CFF7ECFD418CF" box="[510,644,152,176]" italics="true" pageId="5" pageNumber="42">Aromobates</emphasis>
</taxonomicName>
(
<taxonomicName id="A3A2BADCFF97412DFD55FF7ECE5B18CF" box="[663,779,152,176]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="duranti">
<emphasis id="56D61D4DFF97412DFD55FF7ECE5B18CF" box="[663,779,152,176]" italics="true" pageId="5" pageNumber="42">A. duranti</emphasis>
</taxonomicName>
[
<bibRefCitation id="0033BCAEFF97412DFCD8FF71CEE218D0" author="Pefaur" box="[794,946,151,175]" pageId="5" pageNumber="42" refString="Pefaur, J. E. (1985) New species of Venezuelan Colostethus (Dendrobatidae) .. Journal of Herpetology, 19, 321 - 327." type="journal article" year="1985">Péfaur, 1985</bibRefCitation>
], 
<taxonomicName id="A3A2BADCFF97412DFC04FF7EC90118CF" box="[966,1105,152,176]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="haydeeae">
<emphasis id="56D61D4DFF97412DFC04FF7EC90118CF" box="[966,1105,152,176]" italics="true" pageId="5" pageNumber="42">A. haydeeae</emphasis>
</taxonomicName>
[
<bibRefCitation id="0033BCAEFF97412DFBA2FF71C9AC18D0" author="Rivero" box="[1120,1276,151,175]" pageId="5" pageNumber="42" refString="Rivero, J. A. (1978) Notas sobre los anfibios de Venezuela II. Sobre los Colostethus de los Andes Venezolanos. Memoria de la Sociedad de Ciencias Naturales La Salle, 35, 327 - 344." type="journal article" year="1978">Rivero, 1978</bibRefCitation>
], 
<taxonomicName id="A3A2BADCFF97412DFAD3FF7EC8CC18CF" box="[1297,1436,152,176]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="mayorgai">
<emphasis id="56D61D4DFF97412DFAD3FF7EC8CC18CF" box="[1297,1436,152,176]" italics="true" pageId="5" pageNumber="42">A. mayorgai</emphasis>
</taxonomicName>
[
<bibRefCitation id="0033BCAEFF97412DFF5FFF5ACC6918AB" author="Rivero" box="[157,313,188,212]" pageId="5" pageNumber="42" refString="Rivero, J. A. (1980) Notas sobre los anfibios de Venezuela III. Nuevos Colostethus de los Andes Venezolanos. Memoria de la Sociedad de Ciencias Naturales La Salle, 38, 95 - 111." type="journal article" year="1980">Rivero, 1980</bibRefCitation>
], 
<taxonomicName id="A3A2BADCFF97412DFE8CFF5ACC8718AB" box="[334,471,188,212]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="molinarii">
<emphasis id="56D61D4DFF97412DFE8CFF5ACC8718AB" box="[334,471,188,212]" italics="true" pageId="5" pageNumber="42">A. molinarii</emphasis>
</taxonomicName>
[
<bibRefCitation id="0033BCAEFF97412DFE25FF5BCFF518AB" author="La" box="[487,677,188,212]" pageId="5" pageNumber="42" refString="La Marca, E. (1985) A new species of Colostethus (Anura: Dendrobatidae) from the Cordillera de Merida, Northern Andes, South America. Occasional Papers of the Museum of Zoology, University of Michigan, 710, 1 - 10." type="journal article" year="1985">La Marca, 1985</bibRefCitation>
], 
<taxonomicName id="A3A2BADCFF97412DFD7BFF5ACE1318AB" box="[697,835,188,212]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="orostoma">
<emphasis id="56D61D4DFF97412DFD7BFF5ACE1318AB" box="[697,835,188,212]" italics="true" pageId="5" pageNumber="42">A. orostoma</emphasis>
</taxonomicName>
[
<bibRefCitation id="0033BCAEFF97412DFC90FF5ACEBE18AB" author="Rivero" box="[850,1006,188,212]" pageId="5" pageNumber="42" refString="Rivero, J. A. (1978) Notas sobre los anfibios de Venezuela II. Sobre los Colostethus de los Andes Venezolanos. Memoria de la Sociedad de Ciencias Naturales La Salle, 35, 327 - 344." type="journal article" year="1978">Rivero, 1978</bibRefCitation>
], 
<taxonomicName id="A3A2BADCFF97412DFBC1FF5AC9C118AB" box="[1027,1169,188,212]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="saltuensis">
<emphasis id="56D61D4DFF97412DFBC1FF5AC9C118AB" box="[1027,1169,188,212]" italics="true" pageId="5" pageNumber="42">A. saltuensis</emphasis>
</taxonomicName>
[
<bibRefCitation id="0033BCAEFF97412DFB60FF5AC86E18AB" author="Rivero" box="[1186,1342,188,212]" pageId="5" pageNumber="42" refString="Rivero, J. A. (1980) Notas sobre los anfibios de Venezuela III. Nuevos Colostethus de los Andes Venezolanos. Memoria de la Sociedad de Ciencias Naturales La Salle, 38, 95 - 111." type="journal article" year="1980">Rivero, 1980</bibRefCitation>
], and 
<taxonomicName id="A3A2BADCFF97412DFA46FF5ACDA91887" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="serranus">
<emphasis id="56D61D4DFF97412DFA46FF5ACDA91887" italics="true" pageId="5" pageNumber="42">A. serranus</emphasis>
</taxonomicName>
[
<bibRefCitation id="0033BCAEFF97412DFECBFF39CCF21888" author="Pefaur" box="[265,418,223,247]" pageId="5" pageNumber="42" refString="Pefaur, J. E. (1985) New species of Venezuelan Colostethus (Dendrobatidae) .. Journal of Herpetology, 19, 321 - 327." type="journal article" year="1985">Péfaur, 1985</bibRefCitation>
]), are similar in size to 
<taxonomicName id="A3A2BADCFF97412DFD78FF06CEC51887" box="[698,917,224,248]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="tokuko">
<emphasis id="56D61D4DFF97412DFD78FF06CEC51887" box="[698,917,224,248]" italics="true" pageId="5" pageNumber="42">Aromobates tokuko</emphasis>
</taxonomicName>
. 
<taxonomicName id="A3A2BADCFF97412DFC64FF06C9D81887" box="[934,1160,224,248]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="duranti">
<emphasis id="56D61D4DFF97412DFC64FF06C9D81887" box="[934,1160,224,248]" italics="true" pageId="5" pageNumber="42">Aromobates duranti</emphasis>
</taxonomicName>
has smooth dorsal skin (granular), FI longer than FII (FI shorter than FII), finger discs not expanded (expanded), and a gray venter with white spots (unicolor ivory without spots). 
<taxonomicName id="A3A2BADCFF97412DFD4CFECECED8193F" box="[654,904,296,320]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="haydeeae">
<emphasis id="56D61D4DFF97412DFD4CFECECED8193F" box="[654,904,296,320]" italics="true" pageId="5" pageNumber="42">Aromobates haydeeae</emphasis>
</taxonomicName>
has no fringes on fingers (fringes on FII and FIII), FI longer than FII (FI shorter than FII), tarsal fold ill-defined (well defined), dorsal color reddish copper (dark to light brown), and ventral parts orange (ivory white). 
<taxonomicName id="A3A2BADCFF97412DFC8EFE96C91419F7" box="[844,1092,368,392]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="mayorgai">
<emphasis id="56D61D4DFF97412DFC8EFE96C91419F7" box="[844,1092,368,392]" italics="true" pageId="5" pageNumber="42">Aromobates mayorgai</emphasis>
</taxonomicName>
has smooth dorsal skin with a few tubercles posteriorly (granular), ventral parts yellow (ivory white), and the oblique lateral stripe absent (formed by small whitish spots). 
<taxonomicName id="A3A2BADCFF97412DFDCDFE5ECE5619AF" box="[527,774,440,464]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="molinarii">
<emphasis id="56D61D4DFF97412DFDCDFE5ECE5619AF" box="[527,774,440,464]" italics="true" pageId="5" pageNumber="42">Aromobates molinarii</emphasis>
</taxonomicName>
has FI and II equal in length (FI shorter than FII), fringes absent on fingers (preaxial fringes on FII and FIII), tarsal fold slight (well defined), toes moderately webbed (basally webbed), and the oblique lateral stripe absent (formed by small whitish spots). 
<taxonomicName id="A3A2BADCFF97412DFBBAFDE6C83F1A67" box="[1144,1391,512,536]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="orostoma">
<emphasis id="56D61D4DFF97412DFBBAFDE6C83F1A67" box="[1144,1391,512,536]" italics="true" pageId="5" pageNumber="42">Aromobates orostoma</emphasis>
</taxonomicName>
has no fringes on fingers (fringes on FII and FIII), dorsal skin smooth (granular), and venter yellow (ivory white). 
<taxonomicName id="A3A2BADCFF97412DFAA8FDC2CC3A1A1F" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="saltuensis">
<emphasis id="56D61D4DFF97412DFAA8FDC2CC3A1A1F" italics="true" pageId="5" pageNumber="42">Aromobates saltuensis</emphasis>
</taxonomicName>
has FI and FII equal in length (FI shorter than FII), dorsal skin smooth with a few tubercles posteriorly (granular), supratympanic bulge absent (present), and no tympanum (present). 
<taxonomicName id="A3A2BADCFF97412DFBA8FD8AC80B1AFB" box="[1130,1371,620,644]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="5" pageNumber="42" phylum="Chordata" rank="species" species="serranus">
<emphasis id="56D61D4DFF97412DFBA8FD8AC80B1AFB" box="[1130,1371,620,644]" italics="true" pageId="5" pageNumber="42">Aromobates serranus</emphasis>
</taxonomicName>
has a distinct tympanum (ill-defined), dorsum brown with blotches or reticulum (usually uniform, but not reticulated), and ventral parts creamy white with brown reticulation (immaculate ivory).
</paragraph>
</subSubSection>
<subSubSection id="2CB892D4FF97412FFF07FD31CCE31AB3" lastPageId="7" lastPageNumber="44" pageId="5" pageNumber="42" type="description">
<paragraph id="641DC15FFF97412DFF07FD31CE481C97" blockId="5.[151,1437,151,2012]" pageId="5" pageNumber="42">
<emphasis id="56D61D4DFF97412DFF07FD31CF411A8F" bold="true" box="[197,529,727,752]" pageId="5" pageNumber="42">
Description of the 
<typeStatus id="BB197FFDFF97412DFE61FD3ECF5C1A8F" box="[419,524,728,752]" pageId="5" pageNumber="42" type="holotype">holotype</typeStatus>
.
</emphasis>
The 
<typeStatus id="BB197FFDFF97412DFD8EFD31CFFE1A90" box="[588,686,727,751]" pageId="5" pageNumber="42" type="holotype">holotype</typeStatus>
is an adult male of 
<quantity id="A35A6CBAFF97412DFC4DFD31CEA41A90" box="[911,1012,727,751]" metricMagnitude="-2" metricUnit="m" metricValue="2.26" pageId="5" pageNumber="42" unit="mm" value="22.6">22.6 mm</quantity>
SVL. Body robust, short and slightly wider than head (
<figureCitation id="FC99DDDAFF97412DFEA0FD1ACCF51B6B" box="[354,421,764,788]" captionStart="FIGURE 1" captionStartId="2.[151,250,1985,2008]" captionTargetBox="[151,1436,483,1961]" captionTargetId="figure@2.[151,1436,483,1961]" captionTargetPageId="2" captionText="FIGURE 1. Aromobates tokuko sp. nov. Dorsal (a) and ventral (b) views of the male holotype (MHNLS 18479). Dorsal (c) and ventral (d) views of the female paratype (MHNLS 18568). Scale bars represent 10 mm." httpUri="https://zenodo.org/record/277909/files/figure.png" pageId="5" pageNumber="42">Fig. 1</figureCitation>
a). Head almost as long as wide; head length 38.1% SVL; greatest head width between angles of jaws 38.7% SVL. Snout short and subacuminate in profile (
<figureCitation id="FC99DDDAFF97412DFC62FCF9CEB41B48" box="[928,996,799,823]" captionStart="FIGURE 2" captionStartId="3.[151,250,1710,1733]" captionTargetBox="[151,1436,193,1670]" captionTargetId="figure@3.[151,1436,193,1671]" captionTargetPageId="3" captionText="FIGURE 2. Aromobates tokuko sp. nov. (male holotype, MHNLS 18479). Ventral (a), dorsal (b), and lateral (c) views of the head. Details of left hand (d), and left foot (e). Scale bars represent 2 mm." httpUri="https://zenodo.org/record/277910/files/figure.png" pageId="5" pageNumber="42">Fig. 2</figureCitation>
c), nearly rounded in dorsal and ventral view (
<figureCitation id="FC99DDDAFF97412DFF1EFCA2CC791B23" box="[220,297,836,860]" captionStart="FIGURE 2" captionStartId="3.[151,250,1710,1733]" captionTargetBox="[151,1436,193,1670]" captionTargetId="figure@3.[151,1436,193,1671]" captionTargetPageId="3" captionText="FIGURE 2. Aromobates tokuko sp. nov. (male holotype, MHNLS 18479). Ventral (a), dorsal (b), and lateral (c) views of the head. Details of left hand (d), and left foot (e). Scale bars represent 2 mm." httpUri="https://zenodo.org/record/277910/files/figure.png" pageId="5" pageNumber="42">Figs. 2</figureCitation>
a–b). Distance eye-naris slightly shorter than eye diameter (EN/ED = 0.8). Nares are situated laterally to the tip of the snout; nares visible from the front, not visible dorsally; barely visible ventrally. 
<emphasis id="56D61D4DFF97412DFB1BFC8EC8CC1BFF" box="[1241,1436,872,896]" italics="true" pageId="5" pageNumber="42">Canthus rostralis</emphasis>
faintly angular; loreal region flat. Interorbital region wider than upper eyelid; snout longer than eye length (ETS/ ED = 1.8). Tympanum ill-defined, about 1/4 of the tympanum concealed posterodorsally by low supratympanic fold formed by superficial slip of 
<emphasis id="56D61D4DFF97412DFDD8FC33CE671B93" box="[538,823,980,1004]" italics="true" pageId="5" pageNumber="42">m. depressor mandibulae</emphasis>
; tympanum is positioned closely behind eye and low, nearly touching angle of jaws. Teeth are present on maxillary arch. Vocal slits large and long, from mid-level of tongue to near angles of jaws. Tongue is slightly cordiform, 2/3 free posteriorly, without median lingual process. Dorsal skin, including dorsal surfaces of hind limbs, granular, moderately granular on upper eyelids, and smooth on forelimbs. Skin of the flanks moderately granular; tuberculate on the prehumeral area (inferoposteriorly to angle of mouth), on ventral portion of the flanks, external sides of shanks and tarsi. Ventral skin smooth on throat, chest and lower portion of belly, moderately granular on low chest, and upper belly; ventral skin smooth on ventral surface of the arms, and slightly granular on undersurface of thighs.
</paragraph>
<paragraph id="641DC15FFF97412DFF07FB12CC651E53" blockId="5.[151,1437,151,2012]" pageId="5" pageNumber="42">
Hand (
<figureCitation id="FC99DDDAFF97412DFED2FB12CC021D73" box="[272,338,1268,1292]" captionStart="FIGURE 2" captionStartId="3.[151,250,1710,1733]" captionTargetBox="[151,1436,193,1670]" captionTargetId="figure@3.[151,1436,193,1671]" captionTargetPageId="3" captionText="FIGURE 2. Aromobates tokuko sp. nov. (male holotype, MHNLS 18479). Ventral (a), dorsal (b), and lateral (c) views of the head. Details of left hand (d), and left foot (e). Scale bars represent 2 mm." httpUri="https://zenodo.org/record/277910/files/figure.png" pageId="5" pageNumber="42">Fig. 2</figureCitation>
d) is moderate in size (26.8% SVL). Relative lengths of adpressed fingers are III&gt; IV&gt; II&gt; I; FIV reaches past the distal subarticular tubercle of FIII; discs of all fingers moderately expanded, round; disc on FIII 1.5 times wider than distal end of adjacent phalanx. FIII not swollen. Base of palm has a large, rounded palmar tubercle; small (approximately 1/3 of the palmar tubercle and equal in size and form to the proximal subarticular tubercle of FII), well-defined, and elliptical thenar tubercle on base of FI; one elliptical subarticular tubercle each on FI and FII, and two circular each on FIII and FIV; all tubercles very conspicuous and swollen; without supernumerary tubercles. Fringes on fingers absent on FI and FIV; preaxial fringes present on FII and FIII. Metacarpal ridge and carpal pad absent. Paired dorsal digital scutes conspicuous and protuberant in all fingers. Nuptial excrescences on thumb absent.
</paragraph>
<paragraph id="641DC15FFF97412DFF07F9DECFB11FEB" blockId="5.[151,1437,151,2012]" pageId="5" pageNumber="42">
<collectingCountry id="1CB581CFFF97412DFF07F9DECDAD1E2F" box="[197,253,1592,1616]" name="India" pageId="5" pageNumber="42">Hind</collectingCountry>
limbs of moderate length, shank 49.8% SVL. Relative lengths of adpressed toes are IV&gt; III&gt; V&gt; II&gt; I; TI short, the tip reaching the anterior third of subarticular tubercle of TII. Toe discs slightly expanded, about 1.5 times wider than distal end of adjacent phalanx. Paired dorsal digital scutes conspicuous and protuberant in all toes. Feet (
<figureCitation id="FC99DDDAFF97412DFF15F942CC4A1EC3" box="[215,282,1700,1724]" captionStart="FIGURE 2" captionStartId="3.[151,250,1710,1733]" captionTargetBox="[151,1436,193,1670]" captionTargetId="figure@3.[151,1436,193,1671]" captionTargetPageId="3" captionText="FIGURE 2. Aromobates tokuko sp. nov. (male holotype, MHNLS 18479). Ventral (a), dorsal (b), and lateral (c) views of the head. Details of left hand (d), and left foot (e). Scale bars represent 2 mm." httpUri="https://zenodo.org/record/277910/files/figure.png" pageId="5" pageNumber="42">Fig. 2</figureCitation>
e) with basal web from TI–TIV, absent between TIV and TV. One to three non-protuberant, small subarticular tubercles present (one on TI and TII, two on TIII and TV, three on TIV, the proximal one almost indistinct). Conspicuous fringes on all toes. Two metatarsal tubercles are present, including a small round outer, and an oval and protuberant inner tarsal tubercle, the latter twice the size of the former. Metatarsal fold flat and very weak, almost indistinct. With a well-defined and slightly curved tarsal keel from proximal edge of inner metatarsal tubercle, transverse across tarsus, and almost to the mid-tarsus, where it becomes slightly wider. Anal opening at upper quarter level of thighs, with a slight flap above vent.
</paragraph>
<paragraph id="641DC15FFF97412EFF07F879CCFC1A68" blockId="5.[151,1437,151,2012]" lastBlockId="6.[151,1437,151,2012]" lastPageId="6" lastPageNumber="43" pageId="5" pageNumber="42">
<emphasis id="56D61D4DFF97412DFF07F879CC051FC7" bold="true" box="[197,341,1951,1976]" pageId="5" pageNumber="42">Color in life</emphasis>
(based on field notes of FR, 
<date id="101CE79FFF97412DFD5CF846CE721FC7" box="[670,802,1952,1976]" pageId="5" pageNumber="42" value="2006-11-26">11/26/2006</date>
). The dorsal background color is pale brown, with some irregular dark brown spots extending from the interorbital region to the posterior third of the dorsum; two conspicuous paravertebral dark brown stripes present, beginning in the sacral area and joining distally above the vent. A dorsolateral pale golden stripe present on each side of dorsum, starting from tip of snout and continuing through the 
<emphasis id="56D61D4DFF94412EFF55FF06CC051887" box="[151,341,224,248]" italics="true" pageId="6" pageNumber="43">canthus rostralis</emphasis>
, external edge of the upper eyelid and dorsolaterally to reach the upper level of the groin; this stripe is solid and conspicuous, iridescent when seen under magnification. Sides of body with a well-defined blackish brown lateral band from tip of snout to upper level of groin, covering the naris, the two superior thirds of the eye, and the superior third of the tympanum; the upper border of this band is almost straight and in touch with the dorsolateral pale golden stripe, while the lower border becomes broader and diffuse posterior to the tympanum. A diffuse and inconspicuous oblique lateral stripe is present from groin to midbody; it is formed by small whitish spots, coincident with small non-keratinized tubercles. Ventrolateral stripe absent. Tubercles on prehumeral area and ventral portion of the flanks whitish. Paracloacal marks whitish and diffused, forming horizontal stripes from base to middle of thighs.
</paragraph>
<paragraph id="641DC15FFF94412EFF07FDC2CC091B47" blockId="6.[151,1437,151,2012]" pageId="6" pageNumber="43">Dorsal background of forelimbs pale brown. Shoulder and axilla slightly yellowish; anterior side of upper arm with a diffuse dark brown longitudinal stripe, posteriorly without marks. Forearm dorsally dotted with small and diffuse brown spots; with a ventrolateral row of irregular blackish brown spots extending from elbow to wrist. Fingers with alternating whitish and blackish brown bands; discs blackish brown with bluish digital scutes. Dorsal surfaces of hind limbs pale brown in background color; cross-barred with four to five well-defined blackish brown bars on thighs, shanks and tarsi; a longitudinal blackish brown stripe on anterior side of thigh, from groin to knee. Groin and hidden parts of hind limbs slightly yellowish. Toes banded as the fingers; discs blackish brown with bluish digital scutes.</paragraph>
<paragraph id="641DC15FFF94412EFF07FCA2C8CC1BDB" blockId="6.[151,1437,151,2012]" pageId="6" pageNumber="43">Ventrally the throat is grayish ivory; the chest and belly ivory (with profusion of evenly spaced melanophores when seen under magnification); ventral surfaces of arms, thighs, shanks and tarsi are pale orange. Palms and soles gray. Iris is golden and finely dotted of black, with a transversal wide blackish band and a bright golden pupil ring.</paragraph>
<paragraph id="641DC15FFF94412EFF07FC49C93D1C03" blockId="6.[151,1437,151,2012]" pageId="6" pageNumber="43">
<emphasis id="56D61D4DFF94412EFF07FC49CC951BB7" bold="true" box="[197,453,943,968]" pageId="6" pageNumber="43">Color in preservative</emphasis>
(after three years, 
<date id="101CE79FFF94412EFD62FC57CE6D1BB8" box="[672,829,943,969]" pageId="6" pageNumber="43" value="2010-01">January 2010</date>
). Dorsal background pale brown faded to pale beige; dorsal dark brown spots and stripes turned pale brown. The golden color of dorsolateral stripe faded to pale cream (
<figureCitation id="FC99DDDAFF94412EFF5DFC11CDBA1C70" box="[159,234,1015,1039]" captionStart="FIGURE 1" captionStartId="2.[151,250,1985,2008]" captionTargetBox="[151,1436,483,1961]" captionTargetId="figure@2.[151,1436,483,1961]" captionTargetPageId="2" captionText="FIGURE 1. Aromobates tokuko sp. nov. Dorsal (a) and ventral (b) views of the male holotype (MHNLS 18479). Dorsal (c) and ventral (d) views of the female paratype (MHNLS 18568). Scale bars represent 10 mm." httpUri="https://zenodo.org/record/277909/files/figure.png" pageId="6" pageNumber="43">Figs. 1</figureCitation>
a, 2b). Blackish brown lateral band, spots of the flanks, inner longitudinal stripe of the upper arm, spots of the forearms and cross-bands of the thighs, shanks, fingers and toes turned dark brown (
<figureCitation id="FC99DDDAFF94412EFBAAFBFAC9E21C4B" box="[1128,1202,1052,1076]" captionStart="FIGURE 1" captionStartId="2.[151,250,1985,2008]" captionTargetBox="[151,1436,483,1961]" captionTargetId="figure@2.[151,1436,483,1961]" captionTargetPageId="2" captionText="FIGURE 1. Aromobates tokuko sp. nov. Dorsal (a) and ventral (b) views of the male holotype (MHNLS 18479). Dorsal (c) and ventral (d) views of the female paratype (MHNLS 18568). Scale bars represent 10 mm." httpUri="https://zenodo.org/record/277909/files/figure.png" pageId="6" pageNumber="43">Figs. 1</figureCitation>
a, 2c–d). The ventral side pale ivory with a profusion of melanophores on the throat conferring a dirty aspect to this region (
<figureCitation id="FC99DDDAFF94412EFAD1FBA6C8021C27" box="[1299,1362,1088,1112]" captionStart="FIGURE 1" captionStartId="2.[151,250,1985,2008]" captionTargetBox="[151,1436,483,1961]" captionTargetId="figure@2.[151,1436,483,1961]" captionTargetPageId="2" captionText="FIGURE 1. Aromobates tokuko sp. nov. Dorsal (a) and ventral (b) views of the male holotype (MHNLS 18479). Dorsal (c) and ventral (d) views of the female paratype (MHNLS 18568). Scale bars represent 10 mm." httpUri="https://zenodo.org/record/277909/files/figure.png" pageId="6" pageNumber="43">Fig. 1</figureCitation>
b, 2a). Palms and soles are dirty ivory, fingers and toes spotted with pale gray (
<figureCitation id="FC99DDDAFF94412EFC7FFB82C9571C03" box="[957,1031,1124,1148]" captionStart="FIGURE 1" captionStartId="2.[151,250,1985,2008]" captionTargetBox="[151,1436,483,1961]" captionTargetId="figure@2.[151,1436,483,1961]" captionTargetPageId="2" captionText="FIGURE 1. Aromobates tokuko sp. nov. Dorsal (a) and ventral (b) views of the male holotype (MHNLS 18479). Dorsal (c) and ventral (d) views of the female paratype (MHNLS 18568). Scale bars represent 10 mm." httpUri="https://zenodo.org/record/277909/files/figure.png" pageId="6" pageNumber="43">Figs. 1</figureCitation>
b, 2d–e).
</paragraph>
<paragraph id="641DC15FFF94412EFF07FB6ECCF71C97" blockId="6.[151,1437,151,2012]" pageId="6" pageNumber="43">
<emphasis id="56D61D4DFF94412EFF07FB6ECF3D1CDF" bold="true" box="[197,621,1159,1184]" pageId="6" pageNumber="43">
Measurements of 
<typeStatus id="BB197FFDFF94412EFE5BFB6ECF501CDF" box="[409,512,1160,1184]" pageId="6" pageNumber="43" type="holotype">holotype</typeStatus>
(in mm).
</emphasis>
SVL: 22.6; SL: 11.3; ThL: 11.3; FL: 10.7; HaL: 6.1; HeL: 8.6; HW: 8.8; InD: 2.8; EN: 2.1; ED: 2.6; TD: 1.1; ETS: 4.5; F3D: 0.8; T4D: 1.0; 1FiL: 3.3: 2FiL: 3.9. See 
<tableCitation id="2920F4E4FF94412EFB71FB4AC8561CBB" box="[1203,1286,1196,1220]" captionStart="TABLE 2" captionStartId="7.[151,235,766,787]" captionTargetBox="[151,1435,851,1630]" captionTargetPageId="7" captionText="TABLE 2. Morphometric variation (in mm) in the type series of Aromobates tokuko sp. nov. ‾ X ± SD: mean ± standard deviation; Min – Max: minimum – maximum." httpUri="http://table.plazi.org/id/30DD91D7FF95412FFF55FD18CF4B1B48" pageId="6" pageNumber="43" tableUuid="30DD91D7FF95412FFF55FD18CF4B1B48">Table 2</tableCitation>
for measurements of the 
<typeStatus id="BB197FFDFF94412EFEEAFB37CC081C96" box="[296,344,1233,1257]" pageId="6" pageNumber="43">type</typeStatus>
series.
</paragraph>
<paragraph id="641DC15FFF94412EFF07FB15CDB41F0F" blockId="6.[151,1437,151,2012]" pageId="6" pageNumber="43">
<emphasis id="56D61D4DFF94412EFF07FB15CC691D73" bold="true" box="[197,313,1267,1292]" pageId="6" pageNumber="43">Variation.</emphasis>
<taxonomicName id="A3A2BADCFF94412EFE86FB12CF4D1D73" box="[324,541,1268,1292]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="6" pageNumber="43" phylum="Chordata" rank="species" species="tokuko">
<emphasis id="56D61D4DFF94412EFE86FB12CF4D1D73" box="[324,541,1268,1292]" italics="true" pageId="6" pageNumber="43">Aromobates tokuko</emphasis>
</taxonomicName>
is quite variable in color pattern in both sexes and it is difficult to differentiate males and females at first glance. Except by size no other sexually dimorphic characters are noticeable (
<figureCitation id="FC99DDDAFF94412EFAF1FAFEC82E1D4F" box="[1331,1406,1304,1328]" captionStart="FIGURE 1" captionStartId="2.[151,250,1985,2008]" captionTargetBox="[151,1436,483,1961]" captionTargetId="figure@2.[151,1436,483,1961]" captionTargetPageId="2" captionText="FIGURE 1. Aromobates tokuko sp. nov. Dorsal (a) and ventral (b) views of the male holotype (MHNLS 18479). Dorsal (c) and ventral (d) views of the female paratype (MHNLS 18568). Scale bars represent 10 mm." httpUri="https://zenodo.org/record/277909/files/figure.png" pageId="6" pageNumber="43">Figs. 1</figureCitation>
a– d). Males and females of similar size could only be identified by dissection or inspection of presence/absence of vocal slits. Adult females reach larger sizes than adult males; the nine adult males of the 
<typeStatus id="BB197FFDFF94412EFB46FA87C9E41D06" box="[1156,1204,1377,1401]" pageId="6" pageNumber="43">type</typeStatus>
series range in SVL from 
<quantity id="A35A6CBAFF94412EFF16FA62CC271DE3" box="[212,375,1412,1436]" metricMagnitude="-2" metricUnit="m" metricValue="2.245" metricValueMax="2.32" metricValueMin="2.17" pageId="6" pageNumber="43" unit="mm" value="22.45" valueMax="23.2" valueMin="21.7">21.7–23.2 mm</quantity>
(22.6 ± 0.5), while the thirteen adult females range from 
<quantity id="A35A6CBAFF94412EFC38FA62C9CD1DE3" box="[1018,1181,1412,1436]" metricMagnitude="-2" metricUnit="m" metricValue="2.5549999999999997" metricValueMax="2.92" metricValueMin="2.19" pageId="6" pageNumber="43" unit="mm" value="25.549999999999997" valueMax="29.2" valueMin="21.9">21.9–29.2 mm</quantity>
(25.5 ± 1.8). Variation of general morphometric characters of the 
<typeStatus id="BB197FFDFF94412EFD42FA4FCFE01DBE" box="[640,688,1449,1473]" pageId="6" pageNumber="43">type</typeStatus>
series is shown in 
<tableCitation id="2920F4E4FF94412EFC4CFA4ECEB61DBF" box="[910,998,1448,1472]" captionStart="TABLE 2" captionStartId="7.[151,235,766,787]" captionTargetBox="[151,1435,851,1630]" captionTargetPageId="7" captionText="TABLE 2. Morphometric variation (in mm) in the type series of Aromobates tokuko sp. nov. ‾ X ± SD: mean ± standard deviation; Min – Max: minimum – maximum." httpUri="http://table.plazi.org/id/30DD91D7FF95412FFF55FD18CF4B1B48" pageId="6" pageNumber="43" tableUuid="30DD91D7FF95412FFF55FD18CF4B1B48">Table 2</tableCitation>
. The head length ranges from 35.0%– 39.8% SVL (37.3 ± 1.3; 
<emphasis id="56D61D4DFF94412EFE68FA2BCCE81D9B" box="[426,440,1485,1508]" italics="true" pageId="6" pageNumber="43">n</emphasis>
= 23), and maximum head width from 35.1%–39.1% SVL (37.0 ± 1.1; 
<emphasis id="56D61D4DFF94412EFB1AFA2BC9B61D9B" box="[1240,1254,1485,1508]" italics="true" pageId="6" pageNumber="43">n</emphasis>
= 23). The ratio of width/length of head is somewhat variable, with the head sligthly longer than wide in 57% of the 
<typeStatus id="BB197FFDFF94412EFADEFA17C81C1E76" box="[1308,1356,1521,1545]" pageId="6" pageNumber="43">type</typeStatus>
series, slightly wider than long in 30%, and as wide as long in the remaining 13%. The ratio EN/ED is also variable but every distance eye-naris is slightly shorter than eye diameter (0.7–0.9 [0.8 ± 1.1; 
<emphasis id="56D61D4DFF94412EFBF8F9DFC9181E2F" box="[1082,1096,1593,1616]" italics="true" pageId="6" pageNumber="43">n</emphasis>
= 23]). The snout in males is slightly longer than the eye (ratio ETS/ED) and more variable than in females (males: 1.3–1.8 [1.4 ± 0.2; 
<emphasis id="56D61D4DFF94412EFA80F9BBC8001E0B" box="[1346,1360,1629,1652]" italics="true" pageId="6" pageNumber="43">n</emphasis>
= 10], 
<emphasis id="56D61D4DFF94412EFF55F967CDFE1EE7" box="[151,174,1665,1688]" italics="true" pageId="6" pageNumber="43">vs</emphasis>
. females: 1.2–1.5 [1.4 ± 0.1; 
<emphasis id="56D61D4DFF94412EFDC6F967CF421EE7" box="[516,530,1665,1688]" italics="true" pageId="6" pageNumber="43">n</emphasis>
= 13]). Hand length ranges from 23.3%–29.4% SVL (26.4 ± 1.3; 
<emphasis id="56D61D4DFF94412EFACEF967C84A1EE7" box="[1292,1306,1665,1688]" italics="true" pageId="6" pageNumber="43">n</emphasis>
= 23). The male 
<typeStatus id="BB197FFDFF94412EFF1AF943CC691EC2" box="[216,313,1701,1725]" pageId="6" pageNumber="43" type="paratype">paratype</typeStatus>
MHNLS 18473 has FI slightly longer than FII, while the norm is that FI is shorter than FII. A metacarpal ridge is present but very inconspicuous in MHNLS 18498 (male) and MHNLS 18522 (female), and it is absent in all other specimens of the 
<typeStatus id="BB197FFDFF94412EFDEEF90BCF0C1F7A" box="[556,604,1773,1797]" pageId="6" pageNumber="43">type</typeStatus>
series. A metatarsal fold is present but very weak and almost indistinct in MHNLS 18471, 18480, and 18490 (all females). Protuberant tubercles on the posterior third of dorsum, flanks, thighs and shanks are more evident in females (
<figureCitation id="FC99DDDAFF94412EFD79F8D2CE581F33" box="[699,776,1844,1868]" captionStart="FIGURE 1" captionStartId="2.[151,250,1985,2008]" captionTargetBox="[151,1436,483,1961]" captionTargetId="figure@2.[151,1436,483,1961]" captionTargetPageId="2" captionText="FIGURE 1. Aromobates tokuko sp. nov. Dorsal (a) and ventral (b) views of the male holotype (MHNLS 18479). Dorsal (c) and ventral (d) views of the female paratype (MHNLS 18568). Scale bars represent 10 mm." httpUri="https://zenodo.org/record/277909/files/figure.png" pageId="6" pageNumber="43">Figs. 1</figureCitation>
a, c), very conspicuous in MHNLS 18478, 18490–91 and 18495.
</paragraph>
<paragraph id="641DC15FFF94412FFF07F89ACCE31AB3" blockId="6.[151,1437,151,2012]" lastBlockId="7.[151,1436,151,716]" lastPageId="7" lastPageNumber="44" pageId="6" pageNumber="43">
Dorsal color patterns vary from uniform dark brown (males MHNLS 18520, 18498–99; females MHNLS 18480, 18490–91, 18523) (
<figureCitation id="FC99DDDAFF94412EFE0DF846CF441FC7" box="[463,532,1952,1976]" captionStart="FIGURE 3" captionStartId="8.[151,250,1860,1883]" captionTargetBox="[151,1436,193,1838]" captionTargetId="figure@8.[151,1436,193,1839]" captionTargetPageId="8" captionText="FIGURE 3. Color pattern variation in Aromobates tokuko sp. nov. (a) male paratype (MHNLS 18499) with well-defined dorsolateral stripes and dorsal pattern; (b) female paratype (MHNLS 18490) with uniform dorsal and lateral color, and incomplete dorsolateral stripe." httpUri="https://zenodo.org/record/277911/files/figure.png" pageId="6" pageNumber="43">Fig. 3</figureCitation>
b), to pale brown with irregularly-spaced dark brown spots (
<figureCitation id="FC99DDDAFF94412EFB0CF846C8431FC7" box="[1230,1299,1952,1976]" captionStart="FIGURE 3" captionStartId="8.[151,250,1860,1883]" captionTargetBox="[151,1436,193,1838]" captionTargetId="figure@8.[151,1436,193,1839]" captionTargetPageId="8" captionText="FIGURE 3. Color pattern variation in Aromobates tokuko sp. nov. (a) male paratype (MHNLS 18499) with well-defined dorsolateral stripes and dorsal pattern; (b) female paratype (MHNLS 18490) with uniform dorsal and lateral color, and incomplete dorsolateral stripe." httpUri="https://zenodo.org/record/277911/files/figure.png" pageId="6" pageNumber="43">Fig. 3</figureCitation>
a), as in the 
<typeStatus id="BB197FFDFF94412EFF55F822CDA91FA3" box="[151,249,1988,2012]" pageId="6" pageNumber="43" type="holotype">holotype</typeStatus>
(
<figureCitation id="FC99DDDAFF94412EFEC8F822CC1C1FA3" box="[266,332,1988,2012]" captionStart="FIGURE 1" captionStartId="2.[151,250,1985,2008]" captionTargetBox="[151,1436,483,1961]" captionTargetId="figure@2.[151,1436,483,1961]" captionTargetPageId="2" captionText="FIGURE 1. Aromobates tokuko sp. nov. Dorsal (a) and ventral (b) views of the male holotype (MHNLS 18479). Dorsal (c) and ventral (d) views of the female paratype (MHNLS 18568). Scale bars represent 10 mm." httpUri="https://zenodo.org/record/277909/files/figure.png" pageId="6" pageNumber="43">Fig. 1</figureCitation>
a). Dorsolateral stripes yellowish cream to pale golden, generally well-defined and extending to upper level of the groin (
<figureCitation id="FC99DDDAFF95412FFE7FFF71CF5218D0" box="[445,514,151,175]" captionStart="FIGURE 3" captionStartId="8.[151,250,1860,1883]" captionTargetBox="[151,1436,193,1838]" captionTargetId="figure@8.[151,1436,193,1839]" captionTargetPageId="8" captionText="FIGURE 3. Color pattern variation in Aromobates tokuko sp. nov. (a) male paratype (MHNLS 18499) with well-defined dorsolateral stripes and dorsal pattern; (b) female paratype (MHNLS 18490) with uniform dorsal and lateral color, and incomplete dorsolateral stripe." httpUri="https://zenodo.org/record/277911/files/figure.png" pageId="7" pageNumber="44">Fig. 3</figureCitation>
a), eventually comming into contact or continuing with paracloacal light stripes (males MHNLS 18473, 18498, 18567, 18569; females MHNLS 18478, 18480); or ill-defined and ending at level of mid-dorsum (males MHNLS 18476, 18520, 18566; females MHNLS 18471, 18490–91, 18495) (
<figureCitation id="FC99DDDAFF95412FFAC7FF39C8021888" box="[1285,1362,223,247]" captionStart="FIGURE 1" captionStartId="2.[151,250,1985,2008]" captionTargetBox="[151,1436,483,1961]" captionTargetId="figure@2.[151,1436,483,1961]" captionTargetPageId="2" captionText="FIGURE 1. Aromobates tokuko sp. nov. Dorsal (a) and ventral (b) views of the male holotype (MHNLS 18479). Dorsal (c) and ventral (d) views of the female paratype (MHNLS 18568). Scale bars represent 10 mm." httpUri="https://zenodo.org/record/277909/files/figure.png" pageId="7" pageNumber="44">Figs. 1</figureCitation>
c, 3b). Cross-bands on hind limbs range from four to seven, generally blackish brown and well-defined (
<figureCitation id="FC99DDDAFF95412FFB24FEE2C8791963" box="[1254,1321,260,284]" captionStart="FIGURE 3" captionStartId="8.[151,250,1860,1883]" captionTargetBox="[151,1436,193,1838]" captionTargetId="figure@8.[151,1436,193,1839]" captionTargetPageId="8" captionText="FIGURE 3. Color pattern variation in Aromobates tokuko sp. nov. (a) male paratype (MHNLS 18499) with well-defined dorsolateral stripes and dorsal pattern; (b) female paratype (MHNLS 18490) with uniform dorsal and lateral color, and incomplete dorsolateral stripe." httpUri="https://zenodo.org/record/277911/files/figure.png" pageId="7" pageNumber="44">Fig. 3</figureCitation>
a); diffuse pale brown in MHNLS 18491, 18523 (females), and 18476 (male); hind limbs irregularly spotted with blackish brown (not cross-banded) in MHNLS 18566. Light paracloacal stripes ill-defined or absent in specimens whose dorslateral stripe is also ill-defined (males MHNLS 18476, 18566; females MHNLS 18490–91), and very welldefined in specimens with conspicuous dorsolateral stripe (males MHNLS 18498–99, 18569; females MHNLS 18478, 18480). Oblique lateral stripe almost indistinct in MHNLS 18476 (lateral background uniformly pale brown), and MHNLS 18490 (lateral background uniformly dark brown); well-defined on males MHNLS 18473, 18498–99, and 18569. Ventrally the throat, chest and belly generally are immaculate ivory in females (
<figureCitation id="FC99DDDAFF95412FFACFFE19C81C1A68" box="[1293,1356,511,535]" captionStart="FIGURE 1" captionStartId="2.[151,250,1985,2008]" captionTargetBox="[151,1436,483,1961]" captionTargetId="figure@2.[151,1436,483,1961]" captionTargetPageId="2" captionText="FIGURE 1. Aromobates tokuko sp. nov. Dorsal (a) and ventral (b) views of the male holotype (MHNLS 18479). Dorsal (c) and ventral (d) views of the female paratype (MHNLS 18568). Scale bars represent 10 mm." httpUri="https://zenodo.org/record/277909/files/figure.png" pageId="7" pageNumber="44">Fig. 1</figureCitation>
d), and the throat is grayish ivory in males (
<figureCitation id="FC99DDDAFF95412FFDFEFDC2CF2E1A43" box="[572,638,548,572]" captionStart="FIGURE 1" captionStartId="2.[151,250,1985,2008]" captionTargetBox="[151,1436,483,1961]" captionTargetId="figure@2.[151,1436,483,1961]" captionTargetPageId="2" captionText="FIGURE 1. Aromobates tokuko sp. nov. Dorsal (a) and ventral (b) views of the male holotype (MHNLS 18479). Dorsal (c) and ventral (d) views of the female paratype (MHNLS 18568). Scale bars represent 10 mm." httpUri="https://zenodo.org/record/277909/files/figure.png" pageId="7" pageNumber="44">Fig. 1</figureCitation>
b), with a profusion of melanophores when seen under magnification (
<figureCitation id="FC99DDDAFF95412FFF5DFDA1CDB61A20" box="[159,230,583,607]" captionStart="FIGURE 2" captionStartId="3.[151,250,1710,1733]" captionTargetBox="[151,1436,193,1670]" captionTargetId="figure@3.[151,1436,193,1671]" captionTargetPageId="3" captionText="FIGURE 2. Aromobates tokuko sp. nov. (male holotype, MHNLS 18479). Ventral (a), dorsal (b), and lateral (c) views of the head. Details of left hand (d), and left foot (e). Scale bars represent 2 mm." httpUri="https://zenodo.org/record/277910/files/figure.png" pageId="7" pageNumber="44">Fig. 2</figureCitation>
a); even grayish ivory on chest in MHNLS 18499. In the males MHNLS 18473 and 18476 the throat is barely pigmented, and immaculate ivory in the male MHNLS 18499. Two females have the throat dirty ivory (MHNLS 18491 and 18495), and one female (MHNLS 18490) have weakly spotted of pale brown the throat, chest and anterior half of belly.
</paragraph>
</subSubSection>
<caption id="30DD91D7FF95412FFF55FD18CF4B1B48" ID-Table-UUID="30DD91D7FF95412FFF55FD18CF4B1B48" httpUri="http://table.plazi.org/id/30DD91D7FF95412FFF55FD18CF4B1B48" pageId="7" pageNumber="44" targetBox="[151,1435,851,1630]" targetIsTable="true" targetPageId="7">
<paragraph id="641DC15FFF95412FFF55FD18CF4B1B48" blockId="7.[151,1435,764,823]" pageId="7" pageNumber="44">
<emphasis id="56D61D4DFF95412FFF55FD18CC541B6C" bold="true" box="[151,260,766,787]" pageId="7" pageNumber="44">TABLE 2.</emphasis>
Morphometric variation (in mm) in the type series of 
<taxonomicName id="A3A2BADCFF95412FFCEFFD18CEA11B6B" box="[813,1009,766,788]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="7" pageNumber="44" phylum="Chordata" rank="species" species="tokuko" status="sp. nov.">
<emphasis id="56D61D4DFF95412FFCEFFD18CEA11B6B" box="[813,1009,766,788]" italics="true" pageId="7" pageNumber="44">Aromobates tokuko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4DE5A036FF95412FFC35FD18C9181B6C" box="[1015,1096,766,787]" pageId="7" pageNumber="44" rank="species">
<emphasis id="56D61D4DFF95412FFC35FD18C9181B6C" bold="true" box="[1015,1096,766,787]" pageId="7" pageNumber="44">sp. nov.</emphasis>
</taxonomicNameLabel>
‾ 
<emphasis id="56D61D4DFF95412FFB97FD19C9341B6B" box="[1109,1124,767,788]" italics="true" pageId="7" pageNumber="44">X</emphasis>
± SD: mean ± standard deviation; Min–Max: minimum–maximum.
</paragraph>
</caption>
<paragraph id="641DC15FFF95412FFF5DFCB5C9A11E21" pageId="7" pageNumber="44">
<table id="16A233FFFF95BED7FF55FCB5C8CB1E21" box="[151,1435,851,1630]" gridcols="5" gridrows="15" pageId="7" pageNumber="44">
<tr id="DA92C31DFF95BED7FF55FCB5C8CB1BEB" box="[151,1435,851,916]" gridrow="0" pageId="7" pageNumber="44">
<th id="9943AA61FF95BED7FF55FCB5CC611BEB" box="[151,305,851,916]" gridcol="0" gridrow="0" pageId="7" pageNumber="44">Morphometric characters</th>
<th id="9943AA61FF95BED7FE5EFCB5CF761BEB" box="[412,550,851,916]" gridcol="1" gridrow="0" pageId="7" pageNumber="44">
Males (
<emphasis id="56D61D4DFF95412FFE2BFCB3CCA61B15" box="[489,502,853,874]" italics="true" pageId="7" pageNumber="44">n</emphasis>
= 9) ‾ 
<emphasis id="56D61D4DFF95412FFE6BFC99CCA41BEB" bold="true" box="[425,500,893,916]" italics="true" pageId="7" pageNumber="44">X ± SD</emphasis>
</th>
<th id="9943AA61FF95BED7FD62FCB5CE5B1BEB" box="[672,779,851,916]" gridcol="2" gridrow="0" pageId="7" pageNumber="44">
<emphasis id="56D61D4DFF95412FFD62FC9BCE5B1BEC" bold="true" box="[672,779,893,916]" pageId="7" pageNumber="44">Min–Max</emphasis>
</th>
<th id="9943AA61FF95BED7FC59FCB5C9171BEB" box="[923,1095,851,916]" gridcol="3" gridrow="0" pageId="7" pageNumber="44">
Females (
<emphasis id="56D61D4DFF95412FFC3CFCB3C95B1B15" box="[1022,1035,853,874]" italics="true" pageId="7" pageNumber="44">n</emphasis>
= 13) ‾ 
<emphasis id="56D61D4DFF95412FFC6AFC99CEA31BEB" bold="true" box="[936,1011,893,916]" italics="true" pageId="7" pageNumber="44">X ± SD</emphasis>
</th>
<th id="9943AA61FF95BED7FB67FCB5C8CB1BEB" box="[1189,1435,851,916]" gridcol="4" gridrow="0" pageId="7" pageNumber="44">
<emphasis id="56D61D4DFF95412FFB67FC9BC85F1BEC" bold="true" box="[1189,1295,893,916]" pageId="7" pageNumber="44">Min–Max</emphasis>
</th>
</tr>
<tr id="DA92C31DFF95BED7FF55FC4EC8CB1BC1" box="[151,1435,936,958]" gridrow="1" pageId="7" pageNumber="44">
<th id="9943AA61FF95BED7FF55FC4ECC611BC1" box="[151,305,936,958]" gridcol="0" gridrow="1" pageId="7" pageNumber="44">SVL</th>
<td id="9943AA61FF95BED7FE5EFC4ECF761BC1" box="[412,550,936,958]" gridcol="1" gridrow="1" pageId="7" pageNumber="44">22.6 ± 0.5</td>
<td id="9943AA61FF95BED7FD62FC4ECE5B1BC1" box="[672,779,936,958]" gridcol="2" gridrow="1" pageId="7" pageNumber="44">21.7–23.2</td>
<td id="9943AA61FF95BED7FC59FC4EC9171BC1" box="[923,1095,936,958]" gridcol="3" gridrow="1" pageId="7" pageNumber="44">25.5 ± 1.8</td>
<td id="9943AA61FF95BED7FB67FC4EC8CB1BC1" box="[1189,1435,936,958]" gridcol="4" gridrow="1" pageId="7" pageNumber="44">21.9–29.2</td>
</tr>
<tr id="DA92C31DFF95BED7FF55FC34C8CB1C6D" box="[151,1435,978,1042]" gridrow="2" pageId="7" pageNumber="44">
<th id="9943AA61FF95BED7FF55FC34CC611C6D" box="[151,305,978,1042]" gridcol="0" gridrow="2" pageId="7" pageNumber="44">SL Thl</th>
<td id="9943AA61FF95BED7FE5EFC34CF761C6D" box="[412,550,978,1042]" gridcol="1" gridrow="2" pageId="7" pageNumber="44">11.6 ± 0.3 11.3 ± 0.5</td>
<td id="9943AA61FF95BED7FD62FC34CE5B1C6D" box="[672,779,978,1042]" gridcol="2" gridrow="2" pageId="7" pageNumber="44">11.3–12.3 10.4–11.9</td>
<td id="9943AA61FF95BED7FC59FC34C9171C6D" box="[923,1095,978,1042]" gridcol="3" gridrow="2" pageId="7" pageNumber="44">12.5 ± 0.7 12.4 ± 0.8</td>
<td id="9943AA61FF95BED7FB67FC34C8CB1C6D" box="[1189,1435,978,1042]" gridcol="4" gridrow="2" pageId="7" pageNumber="44">10.6–13.4 10.0–13.3</td>
</tr>
<tr id="DA92C31DFF95BED7FF55FBC0C8CB1C43" box="[151,1435,1062,1084]" gridrow="3" pageId="7" pageNumber="44">
<th id="9943AA61FF95BED7FF55FBC0CC611C43" box="[151,305,1062,1084]" gridcol="0" gridrow="3" pageId="7" pageNumber="44">FL</th>
<td id="9943AA61FF95BED7FE5EFBC0CF761C43" box="[412,550,1062,1084]" gridcol="1" gridrow="3" pageId="7" pageNumber="44">10.7 ± 0.4</td>
<td id="9943AA61FF95BED7FD62FBC0CE5B1C43" box="[672,779,1062,1084]" gridcol="2" gridrow="3" pageId="7" pageNumber="44">9.8–11.3</td>
<td id="9943AA61FF95BED7FC59FBC0C9171C43" box="[923,1095,1062,1084]" gridcol="3" gridrow="3" pageId="7" pageNumber="44">11.6 ± 0.8</td>
<td id="9943AA61FF95BED7FB67FBC0C8CB1C43" box="[1189,1435,1062,1084]" gridcol="4" gridrow="3" pageId="7" pageNumber="44">9.9–12.6</td>
</tr>
<tr id="DA92C31DFF95BED7FF55FBB6C8CB1C19" box="[151,1435,1104,1126]" gridrow="4" pageId="7" pageNumber="44">
<th id="9943AA61FF95BED7FF55FBB6CC611C19" box="[151,305,1104,1126]" gridcol="0" gridrow="4" pageId="7" pageNumber="44">HaL</th>
<td id="9943AA61FF95BED7FE5EFBB6CF761C19" box="[412,550,1104,1126]" gridcol="1" gridrow="4" pageId="7" pageNumber="44">6.0 ± 0.4</td>
<td id="9943AA61FF95BED7FD62FBB6CE5B1C19" box="[672,779,1104,1126]" gridcol="2" gridrow="4" pageId="7" pageNumber="44">5.1–6.8</td>
<td id="9943AA61FF95BED7FC59FBB6C9171C19" box="[923,1095,1104,1126]" gridcol="3" gridrow="4" pageId="7" pageNumber="44">6.7 ± 0.4</td>
<td id="9943AA61FF95BED7FB67FBB6C8CB1C19" box="[1189,1435,1104,1126]" gridcol="4" gridrow="4" pageId="7" pageNumber="44">5.6–7.4</td>
</tr>
<tr id="DA92C31DFF95BED7FF55FB9CC8CB1CEF" box="[151,1435,1146,1168]" gridrow="5" pageId="7" pageNumber="44">
<th id="9943AA61FF95BED7FF55FB9CCC611CEF" box="[151,305,1146,1168]" gridcol="0" gridrow="5" pageId="7" pageNumber="44">HeL</th>
<td id="9943AA61FF95BED7FE5EFB9CCF761CEF" box="[412,550,1146,1168]" gridcol="1" gridrow="5" pageId="7" pageNumber="44">8.5 ± 0.3</td>
<td id="9943AA61FF95BED7FD62FB9CCE5B1CEF" box="[672,779,1146,1168]" gridcol="2" gridrow="5" pageId="7" pageNumber="44">7.9–9.0</td>
<td id="9943AA61FF95BED7FC59FB9CC9171CEF" box="[923,1095,1146,1168]" gridcol="3" gridrow="5" pageId="7" pageNumber="44">9.4 ± 0.5</td>
<td id="9943AA61FF95BED7FB67FB9CC8CB1CEF" box="[1189,1435,1146,1168]" gridcol="4" gridrow="5" pageId="7" pageNumber="44">8.3–10.3</td>
</tr>
<tr id="DA92C31DFF95BED7FF55FB42C8CB1CC5" box="[151,1435,1188,1210]" gridrow="6" pageId="7" pageNumber="44">
<th id="9943AA61FF95BED7FF55FB42CC611CC5" box="[151,305,1188,1210]" gridcol="0" gridrow="6" pageId="7" pageNumber="44">HW</th>
<td id="9943AA61FF95BED7FE5EFB42CF761CC5" box="[412,550,1188,1210]" gridcol="1" gridrow="6" pageId="7" pageNumber="44">8.4 ± 0.3</td>
<td id="9943AA61FF95BED7FD62FB42CE5B1CC5" box="[672,779,1188,1210]" gridcol="2" gridrow="6" pageId="7" pageNumber="44">7.8–8.9</td>
<td id="9943AA61FF95BED7FC59FB42C9171CC5" box="[923,1095,1188,1210]" gridcol="3" gridrow="6" pageId="7" pageNumber="44">9.4 ± 0.7</td>
<td id="9943AA61FF95BED7FB67FB42C8CB1CC5" box="[1189,1435,1188,1210]" gridcol="4" gridrow="6" pageId="7" pageNumber="44">8.4–10.8</td>
</tr>
<tr id="DA92C31DFF95BED7FF55FB28C8CB1C9B" box="[151,1435,1230,1252]" gridrow="7" pageId="7" pageNumber="44">
<th id="9943AA61FF95BED7FF55FB28CC611C9B" box="[151,305,1230,1252]" gridcol="0" gridrow="7" pageId="7" pageNumber="44">IO</th>
<td id="9943AA61FF95BED7FE5EFB28CF761C9B" box="[412,550,1230,1252]" gridcol="1" gridrow="7" pageId="7" pageNumber="44">2.8 ± 0.2</td>
<td id="9943AA61FF95BED7FD62FB28CE5B1C9B" box="[672,779,1230,1252]" gridcol="2" gridrow="7" pageId="7" pageNumber="44">2.6–3.1</td>
<td id="9943AA61FF95BED7FC59FB28C9171C9B" box="[923,1095,1230,1252]" gridcol="3" gridrow="7" pageId="7" pageNumber="44">3.0 ± 0.2</td>
<td id="9943AA61FF95BED7FB67FB28C8CB1C9B" box="[1189,1435,1230,1252]" gridcol="4" gridrow="7" pageId="7" pageNumber="44">2.7–3.3</td>
</tr>
<tr id="DA92C31DFF95BED7FF55FB1EC8CB1D71" box="[151,1435,1272,1294]" gridrow="8" pageId="7" pageNumber="44">
<th id="9943AA61FF95BED7FF55FB1ECC611D71" box="[151,305,1272,1294]" gridcol="0" gridrow="8" pageId="7" pageNumber="44">IND</th>
<td id="9943AA61FF95BED7FE5EFB1ECF761D71" box="[412,550,1272,1294]" gridcol="1" gridrow="8" pageId="7" pageNumber="44">3.1 ± 0.2</td>
<td id="9943AA61FF95BED7FD62FB1ECE5B1D71" box="[672,779,1272,1294]" gridcol="2" gridrow="8" pageId="7" pageNumber="44">2.8–3.3</td>
<td id="9943AA61FF95BED7FC59FB1EC9171D71" box="[923,1095,1272,1294]" gridcol="3" gridrow="8" pageId="7" pageNumber="44">3.6 ± 0.1</td>
<td id="9943AA61FF95BED7FB67FB1EC8CB1D71" box="[1189,1435,1272,1294]" gridcol="4" gridrow="8" pageId="7" pageNumber="44">3.3–3.8</td>
</tr>
<tr id="DA92C31DFF95BED7FF55FAC4C8CB1D1E" box="[151,1435,1314,1377]" gridrow="9" pageId="7" pageNumber="44">
<th id="9943AA61FF95BED7FF55FAC4CC611D1E" box="[151,305,1314,1377]" gridcol="0" gridrow="9" pageId="7" pageNumber="44">EN ED</th>
<td id="9943AA61FF95BED7FE5EFAC4CF761D1E" box="[412,550,1314,1377]" gridcol="1" gridrow="9" pageId="7" pageNumber="44">2.1 ± 0.1 2.7 ± 0.1</td>
<td id="9943AA61FF95BED7FD62FAC4CE5B1D1E" box="[672,779,1314,1377]" gridcol="2" gridrow="9" pageId="7" pageNumber="44">2.0–2.3 2.5–2.9</td>
<td id="9943AA61FF95BED7FC59FAC4C9171D1E" box="[923,1095,1314,1377]" gridcol="3" gridrow="9" pageId="7" pageNumber="44">2.4 ± 0.2 3.0 ± 0.3</td>
<td id="9943AA61FF95BED7FB67FAC4C8CB1D1E" box="[1189,1435,1314,1377]" gridcol="4" gridrow="9" pageId="7" pageNumber="44">2.2–2.8 2.6–3.6</td>
</tr>
<tr id="DA92C31DFF95BED7FF55FA90C8CB1DF3" box="[151,1435,1398,1420]" gridrow="10" pageId="7" pageNumber="44">
<th id="9943AA61FF95BED7FF55FA90CC611DF3" box="[151,305,1398,1420]" gridcol="0" gridrow="10" pageId="7" pageNumber="44">TD</th>
<td id="9943AA61FF95BED7FE5EFA90CF761DF3" box="[412,550,1398,1420]" gridcol="1" gridrow="10" pageId="7" pageNumber="44">1.2 ± 0.1</td>
<td id="9943AA61FF95BED7FD62FA90CE5B1DF3" box="[672,779,1398,1420]" gridcol="2" gridrow="10" pageId="7" pageNumber="44">1.1–1.5</td>
<td id="9943AA61FF95BED7FC59FA90C9171DF3" box="[923,1095,1398,1420]" gridcol="3" gridrow="10" pageId="7" pageNumber="44">1.4 ± 0.1</td>
<td id="9943AA61FF95BED7FB67FA90C8CB1DF3" box="[1189,1435,1398,1420]" gridcol="4" gridrow="10" pageId="7" pageNumber="44">1.2–1.6</td>
</tr>
<tr id="DA92C31DFF95BED7FF55FA46C8CB1D9F" box="[151,1435,1440,1504]" gridrow="11" pageId="7" pageNumber="44">
<th id="9943AA61FF95BED7FF55FA46CC611D9F" box="[151,305,1440,1504]" gridcol="0" gridrow="11" pageId="7" pageNumber="44">ETS F3D</th>
<td id="9943AA61FF95BED7FE5EFA46CF761D9F" box="[412,550,1440,1504]" gridcol="1" gridrow="11" pageId="7" pageNumber="44">3.8 ± 0.4 0.8 ± 0.1</td>
<td id="9943AA61FF95BED7FD62FA46CE5B1D9F" box="[672,779,1440,1504]" gridcol="2" gridrow="11" pageId="7" pageNumber="44">3.3–4.7 0.6–0.8</td>
<td id="9943AA61FF95BED7FC59FA46C9171D9F" box="[923,1095,1440,1504]" gridcol="3" gridrow="11" pageId="7" pageNumber="44">4.1 ± 0.3 1.0 ± 0.4</td>
<td id="9943AA61FF95BED7FB67FA46C8CB1D9F" box="[1189,1435,1440,1504]" gridcol="4" gridrow="11" pageId="7" pageNumber="44">3.5–4.9 0.7–1.8</td>
</tr>
<tr id="DA92C31DFF95BED7FF55FA12C8CB1E76" box="[151,1435,1524,1545]" gridrow="12" pageId="7" pageNumber="44">
<th id="9943AA61FF95BED7FF55FA12CC611E76" box="[151,305,1524,1545]" gridcol="0" gridrow="12" pageId="7" pageNumber="44">T4D</th>
<td id="9943AA61FF95BED7FE5EFA12CF761E76" box="[412,550,1524,1545]" gridcol="1" gridrow="12" pageId="7" pageNumber="44">0.9 ± 0.1</td>
<td id="9943AA61FF95BED7FD62FA12CE5B1E76" box="[672,779,1524,1545]" gridcol="2" gridrow="12" pageId="7" pageNumber="44">0.8–1.0</td>
<td id="9943AA61FF95BED7FC59FA12C9171E76" box="[923,1095,1524,1545]" gridcol="3" gridrow="12" pageId="7" pageNumber="44">1.0 ± 0.3</td>
<td id="9943AA61FF95BED7FB67FA12C8CB1E76" box="[1189,1435,1524,1545]" gridcol="4" gridrow="12" pageId="7" pageNumber="44">0.8–1.9</td>
</tr>
<tr id="DA92C31DFF95BED7FF55F9F8C8CB1E4B" box="[151,1435,1566,1588]" gridrow="13" pageId="7" pageNumber="44">
<th id="9943AA61FF95BED7FF55F9F8CC611E4B" box="[151,305,1566,1588]" gridcol="0" gridrow="13" pageId="7" pageNumber="44">1FiL</th>
<td id="9943AA61FF95BED7FE5EF9F8CF761E4B" box="[412,550,1566,1588]" gridcol="1" gridrow="13" pageId="7" pageNumber="44">3.4 ± 0.1</td>
<td id="9943AA61FF95BED7FD62F9F8CE5B1E4B" box="[672,779,1566,1588]" gridcol="2" gridrow="13" pageId="7" pageNumber="44">3.2–3.6</td>
<td id="9943AA61FF95BED7FC59F9F8C9171E4B" box="[923,1095,1566,1588]" gridcol="3" gridrow="13" pageId="7" pageNumber="44">3.8 ± 0.4</td>
<td id="9943AA61FF95BED7FB67F9F8C8CB1E4B" box="[1189,1435,1566,1588]" gridcol="4" gridrow="13" pageId="7" pageNumber="44">3.1–4.5</td>
</tr>
<tr id="DA92C31DFF95BED7FF55F9AEC8CB1E21" box="[151,1435,1608,1630]" gridrow="14" pageId="7" pageNumber="44">
<th id="9943AA61FF95BED7FF55F9AECC611E21" box="[151,305,1608,1630]" gridcol="0" gridrow="14" pageId="7" pageNumber="44">2FiL</th>
<td id="9943AA61FF95BED7FE5EF9AECF761E21" box="[412,550,1608,1630]" gridcol="1" gridrow="14" pageId="7" pageNumber="44">4.0 ± 0.3</td>
<td id="9943AA61FF95BED7FD62F9AECE5B1E21" box="[672,779,1608,1630]" gridcol="2" gridrow="14" pageId="7" pageNumber="44">3.7–4.7</td>
<td id="9943AA61FF95BED7FC59F9AEC9171E21" box="[923,1095,1608,1630]" gridcol="3" gridrow="14" pageId="7" pageNumber="44">4.4 ± 0.4</td>
<td id="9943AA61FF95BED7FB67F9AEC8CB1E21" box="[1189,1435,1608,1630]" gridcol="4" gridrow="14" pageId="7" pageNumber="44">3.6–5.1</td>
</tr>
</table>
</paragraph>
<subSubSection id="2CB892D4FF954123FF07F941C8C31AB0" lastPageId="11" lastPageNumber="48" pageId="7" pageNumber="44" type="distribution">
<paragraph id="641DC15FFF95412FFF07F941CC611FC3" blockId="7.[151,1436,1703,1980]" pageId="7" pageNumber="44">
<emphasis id="56D61D4DFF95412FFF07F941CFE41EBF" bold="true" box="[197,692,1703,1728]" pageId="7" pageNumber="44">Distribution, habitat and natural history.</emphasis>
<taxonomicName id="A3A2BADCFF95412FFD7EF94ECEC61EBF" box="[700,918,1704,1728]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="7" pageNumber="44" phylum="Chordata" rank="species" species="tokuko">
<emphasis id="56D61D4DFF95412FFD7EF94ECEC61EBF" box="[700,918,1704,1728]" italics="true" pageId="7" pageNumber="44">Aromobates tokuko</emphasis>
</taxonomicName>
is only known from four localities in the Río 
<taxonomicName id="A3A2BADCFF95412FFF55F92ACDBC1E9B" box="[151,236,1740,1764]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="7" pageNumber="48" phylum="Chordata" rank="species" species="tokuko">Tokuko</taxonomicName>
basin (Kiriponsa, near Ipika, upstream of the Kusare waterfall, and Terakibu) on the eastern versant of the Sierra de Perijá, northwestern 
<collectingCountry id="1CB581CFFF95412FFE31F909CF391F78" box="[499,617,1775,1799]" name="Venezuela" pageId="7" pageNumber="44">Venezuela</collectingCountry>
, between elevations of 419 and 
<quantity id="A35A6CBAFF95412FFC19F909C9621F78" box="[987,1074,1775,1799]" metricMagnitude="3" metricUnit="m" metricValue="1.005" pageId="7" pageNumber="44" unit="m" value="1005.0">1005 m</quantity>
(
<figureCitation id="FC99DDDAFF95412FFB81F909C9DB1F78" box="[1091,1163,1775,1799]" captionStart="FIGURE 4" captionStartId="9.[151,250,1841,1864]" captionTargetBox="[151,1436,193,1819]" captionTargetId="figure@9.[151,1436,193,1820]" captionTargetPageId="9" captionText="FIGURE 4. (top) relief map of the Río Tukuko basin showing the four localities where the type series of Aromobates tokuko sp. nov. was collected; 1. Kiriponsa; 2. Ipika; 3. Upstream Kusare waterfall; 4. Terakibu. (bottom) map of NE Colombia and NW Venezuela showing major physiographic elements, from west to east: Sierra de Santa Marta, Sierra de Perijá, Maracaibo Lake, and Cordillera de Mérida; red frame indicates the relative position of the Río Tokuko basin in the Sierra de Perijá." httpUri="https://zenodo.org/record/277913/files/figure.png" pageId="7" pageNumber="44">Fig. 4</figureCitation>
). The vegetation of this altitudinal belt was described by 
<bibRefCitation id="0033BCAEFF95412FFDC7F8F2CE491F53" author="Huber" box="[517,793,1812,1836]" pageId="7" pageNumber="44" refString="Huber, O. &amp; Alarcon, C. (1988) Mapa de vegetacion de Venezuela. 1: 2.000.000." type="book chapter" year="1988">Huber &amp; Alarcón (1988)</bibRefCitation>
as seasonal semideciduous ombrofilous basimontane forest (from 
<quantity id="A35A6CBAFF95412FFEC9F8D1CCDC1F2F" box="[267,396,1847,1872]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="8.0" metricValueMin="1.0" pageId="7" pageNumber="44" unit="m" value="450.0" valueMax="800.0" valueMin="100.0">100–800 m</quantity>
asl), and ombrofilous submontane/montane evergreen forest (from 
<quantity id="A35A6CBAFF95412FFB5EF8D1C87C1F2F" box="[1180,1324,1847,1872]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="2.5" metricValueMin="0.8" pageId="7" pageNumber="44" unit="m" value="1650.0" valueMax="2500.0" valueMin="800.0">800–2500 m</quantity>
asl). The region has a biseasonal climatic regime, with a dry period from December–April, and a rainy period from May– November, with maximum rain peaks in May and October, and minimum rain in January and July (
<bibRefCitation id="0033BCAEFF95412FFB29F899CC751FC3" author="Masciangioli" pageId="7" pageNumber="44" refString="Masciangioli, P. &amp; Febres, G. (2000) Climatologia de la cuenca de Maracaibo. Pp 21 - 32. In: Rodriguez, G. (ed.). El sistema de Maracaibo, Biologia y Ambiente. Second edition. Instituto Venezolano de Investigaciones Cientificas, Caracas." type="book chapter" year="2000">Masciangioli &amp; Febres 2000</bibRefCitation>
).
</paragraph>
<caption id="30DD91D7FF9A4120FF55F8A2CC041FE7" httpUri="https://zenodo.org/record/277911/files/figure.png" pageId="8" pageNumber="45" targetBox="[151,1436,193,1838]" targetPageId="8">
<paragraph id="641DC15FFF9A4120FF55F8A2CC041FE7" blockId="8.[151,1436,1860,1944]" pageId="8" pageNumber="45">
<emphasis id="56D61D4DFF9A4120FF55F8A2CC431F25" bold="true" box="[151,275,1860,1883]" pageId="8" pageNumber="45">FIGURE 3.</emphasis>
Color pattern variation in 
<taxonomicName id="A3A2BADCFF9A4120FDE6F8A3CFB71F24" box="[548,743,1861,1883]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="8" pageNumber="45" phylum="Chordata" rank="species" species="tokuko" status="sp. nov.">
<emphasis id="56D61D4DFF9A4120FDE6F8A3CFB71F24" box="[548,743,1861,1883]" italics="true" pageId="8" pageNumber="45">Aromobates tokuko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4DE5A036FF9A4120FD2DF8A3CE101F25" box="[751,832,1861,1882]" pageId="8" pageNumber="45" rank="species">
<emphasis id="56D61D4DFF9A4120FD2DF8A3CE101F25" bold="true" box="[751,832,1861,1882]" pageId="8" pageNumber="45">sp. nov.</emphasis>
</taxonomicNameLabel>
(a) male paratype (MHNLS 18499) with well-defined dorsolateral stripes and dorsal pattern; (b) female paratype (MHNLS 18490) with uniform dorsal and lateral color, and incomplete dorsolateral stripe.
</paragraph>
</caption>
<caption id="30DD91D7FF9B4121FF55F8D7C8081FDC" httpUri="https://zenodo.org/record/277913/files/figure.png" pageId="9" pageNumber="46" targetBox="[151,1436,193,1819]" targetPageId="9">
<paragraph id="641DC15FFF9B4121FF55F8D7C8081FDC" blockId="9.[151,1436,1841,1955]" pageId="9" pageNumber="46">
<emphasis id="56D61D4DFF9B4121FF55F8D7CC431F38" bold="true" box="[151,275,1841,1864]" pageId="9" pageNumber="46">FIGURE 4.</emphasis>
(top) relief map of the Río Tukuko basin showing the four localities where the type series of 
<taxonomicName id="A3A2BADCFF9B4121FB1AF8D7C8CC1F38" box="[1240,1436,1841,1863]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="9" pageNumber="46" phylum="Chordata" rank="species" species="tokuko" status="sp. nov.">
<emphasis id="56D61D4DFF9B4121FB1AF8D7C8CC1F38" box="[1240,1436,1841,1863]" italics="true" pageId="9" pageNumber="46">Aromobates tokuko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4DE5A036FF9B4121FF55F8B7CDB81F19" box="[151,232,1873,1894]" pageId="9" pageNumber="46" rank="species">
<emphasis id="56D61D4DFF9B4121FF55F8B7CDB81F19" bold="true" box="[151,232,1873,1894]" pageId="9" pageNumber="46">sp. nov.</emphasis>
</taxonomicNameLabel>
was collected; 1. Kiriponsa; 2. Ipika; 3. Upstream Kusare waterfall; 4. Terakibu. (bottom) map of NE Colombia and NW Venezuela showing major physiographic elements, from west to east: Sierra de Santa Marta, Sierra de Perijá, Maracaibo Lake, and Cordillera de Mérida; red frame indicates the relative position of the Río 
<taxonomicName id="A3A2BADCFF9B4121FC24F86BC9631FDC" box="[998,1075,1933,1955]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="9" pageNumber="48" phylum="Chordata" rank="species" species="tokuko">Tokuko</taxonomicName>
basin in the Sierra de Perijá.
</paragraph>
</caption>
<caption id="30DD91D7FF984122FF55F861CF9D1FC3" httpUri="https://zenodo.org/record/277914/files/figure.png" pageId="10" pageNumber="47" targetBox="[151,1436,193,1904]" targetPageId="10">
<paragraph id="641DC15FFF984122FF55F861CF9D1FC3" blockId="10.[151,1436,1927,1981]" pageId="10" pageNumber="47">
<emphasis id="56D61D4DFF984122FF55F861CC441FE2" bold="true" box="[151,276,1927,1950]" pageId="10" pageNumber="47">FIGURE 5.</emphasis>
View of the stream of Kusare waterfall (tributary of Río 
<taxonomicName id="A3A2BADCFF984122FCA4F86ECEE71FE1" box="[870,951,1928,1950]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="10" pageNumber="48" phylum="Chordata" rank="species" species="tokuko">Tokuko</taxonomicName>
), one of the localities inhabited by 
<taxonomicName id="A3A2BADCFF984122FAE1F86ECD8B1FC2" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="10" pageNumber="47" phylum="Chordata" rank="species" species="tokuko" status="sp. nov.">
<emphasis id="56D61D4DFF984122FAE1F86ECD8B1FC2" italics="true" pageId="10" pageNumber="47">Aromobates tokuko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4DE5A036FF984122FF23F841CC611FC3" box="[225,305,1959,1980]" pageId="10" pageNumber="47" rank="species">
<emphasis id="56D61D4DFF984122FF23F841CC611FC3" bold="true" box="[225,305,1959,1980]" pageId="10" pageNumber="47">sp. nov.</emphasis>
</taxonomicNameLabel>
All other localities are similar in aspect.
</paragraph>
</caption>
<paragraph id="641DC15FFF994123FF07FF7EC91D19AD" blockId="11.[151,1437,151,791]" pageId="11" pageNumber="48">
<taxonomicName id="A3A2BADCFF994123FF07FF7ECCCD18CF" box="[197,413,152,176]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="11" pageNumber="48" phylum="Chordata" rank="species" species="tokuko">
<emphasis id="56D61D4DFF994123FF07FF7ECCCD18CF" box="[197,413,152,176]" italics="true" pageId="11" pageNumber="48">Aromobates tokuko</emphasis>
</taxonomicName>
is a diurnal and cryptic riparian frog. Even though its presence in the streams can be readily detected by its vocalization, it is very hard to detect visually because males call from underneath rocks or logs, or from hollows and shelters, and exhibit a cryptic dorsal color pattern. The eleven specimens (three males, four females and four juveniles) collected near Ipika on 
<date id="101CE79FFF994123FD17FEE2CEDB1963" box="[725,907,260,284]" pageId="11" pageNumber="48" value="2006-11">November 2006</date>
were found under rocks and rotten logs around a small quiet pool, in an area of about 
<quantity id="A35A6CBAFF994123FD81FECCCF3D193D" box="[579,621,298,322]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="48" unit="m" value="4.0">4 m</quantity>
2, along a small intermittent stream permanently shaded by dense canopy in the gallery forest. During the collecting effort (1400–1500 h), several males were heard calling from hidden sites along the stream, but we failed to record the vocalizations. Several tadpoles of 
<taxonomicName id="A3A2BADCFF994123FBD5FE95C9D319F4" box="[1047,1155,371,395]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="11" pageNumber="48" phylum="Chordata" rank="species" species="tokuko">
<emphasis id="56D61D4DFF994123FBD5FE95C9D319F4" box="[1047,1155,371,395]" italics="true" pageId="11" pageNumber="48">A. tokuko</emphasis>
</taxonomicName>
in different larval stages were found in the pool but no adult specimen carrying tadpoles were observed. Several tadpoles, and two adults of 
<taxonomicName id="A3A2BADCFF994123FF55FE5DCCEB19AC" box="[151,443,442,467]" class="Amphibia" family="Leptodactylidae" genus="Leptodactylus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="48" phylum="Chordata" rank="species" species="wagneri">
<emphasis id="56D61D4DFF994123FF55FE5DCC6419AC" box="[151,308,443,467]" italics="true" pageId="11" pageNumber="48">Leptodactylus</emphasis>
cf. 
<emphasis id="56D61D4DFF994123FE9DFE5ACCEB19AC" box="[351,443,444,467]" italics="true" pageId="11" pageNumber="48">wagneri</emphasis>
</taxonomicName>
were also found in syntopy with the tadpoles of 
<taxonomicName id="A3A2BADCFF994123FC1EFE5DC91719AC" box="[988,1095,443,467]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="11" pageNumber="48" phylum="Chordata" rank="species" species="tokuko">
<emphasis id="56D61D4DFF994123FC1EFE5DC91719AC" box="[988,1095,443,467]" italics="true" pageId="11" pageNumber="48">A. tokuko</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="641DC15FFF994123FF07FE39CCD21A1D" blockId="11.[151,1437,151,791]" pageId="11" pageNumber="48">
On 
<date id="101CE79FFF994123FF2CFE39CCF31988" box="[238,419,479,503]" pageId="11" pageNumber="48" value="2006-12">December 2006</date>
, two males (MHNLS 18498 and 18499) were collected while vocalizing at 1140 h, hidden under rocks along a shaded section of a fast moving stream, upstream from Kusare waterfall (
<figureCitation id="FC99DDDAFF994123FB02FDE4C8551A65" box="[1216,1285,514,538]" captionStart="FIGURE 5" captionStartId="10.[151,250,1927,1950]" captionTargetBox="[151,1436,193,1904]" captionTargetId="figure@10.[151,1436,193,1906]" captionTargetPageId="10" captionText="FIGURE 5. View of the stream of Kusare waterfall (tributary of Río Tokuko), one of the localities inhabited by Aromobates tokuko sp. nov. All other localities are similar in aspect." httpUri="https://zenodo.org/record/277914/files/figure.png" pageId="11" pageNumber="48">Fig. 5</figureCitation>
). Both males were about 
<quantity id="A35A6CBAFF994123FEDEFDC1CC0D1A40" box="[284,349,551,575]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="48" unit="m" value="1.5">1.5 m</quantity>
apart while calling. At this locality, 
<taxonomicName id="A3A2BADCFF994123FD3CFDC1CE391A40" box="[766,873,551,575]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="11" pageNumber="48" phylum="Chordata" rank="species" species="tokuko">
<emphasis id="56D61D4DFF994123FD3CFDC1CE391A40" box="[766,873,551,575]" italics="true" pageId="11" pageNumber="48">A. tokuko</emphasis>
</taxonomicName>
is syntopic with 
<taxonomicName id="A3A2BADCFF994123FBEFFDC1C84A1A40" box="[1069,1306,551,575]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="11" pageNumber="48" phylum="Chordata" rank="species" species="haematiticus">
<emphasis id="56D61D4DFF994123FBEFFDC1C84A1A40" box="[1069,1306,551,575]" italics="true" pageId="11" pageNumber="48">Rhaebo haematiticus</emphasis>
</taxonomicName>
and 
<taxonomicName id="A3A2BADCFF994123FA91FDCECC2B1A1C" class="Amphibia" family="Leptodactylidae" genus="Leptodactylus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="48" phylum="Chordata" rank="species" species="wagneri">
<emphasis id="56D61D4DFF994123FA91FDCECDA41A1C" italics="true" pageId="11" pageNumber="48">Leptodactylus</emphasis>
cf. 
<emphasis id="56D61D4DFF994123FEDDFDAACC2B1A1C" box="[287,379,588,611]" italics="true" pageId="11" pageNumber="48">wagneri</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="641DC15FFF994123FF07FD89C8C31AB0" blockId="11.[151,1437,151,791]" pageId="11" pageNumber="48">Finally, the specimens found at Kiriponsa (a female and three males) were collected at about 1600 h under rocks and in crevices of a large stone along a fast-moving stream. Numerous males were heard calling at this hour. The stream of Kiriponsa runs alongside an indigenous community and is surrounded by a shady coffee plantation.</paragraph>
</subSubSection>
<subSubSection id="2CB892D4FF994123FF07FD3DCC4A1B68" pageId="11" pageNumber="48" type="etymology">
<paragraph id="641DC15FFF994123FF07FD3DCC4A1B68" blockId="11.[151,1437,151,791]" pageId="11" pageNumber="48">
<emphasis id="56D61D4DFF994123FF07FD3DCC1C1A8C" bold="true" box="[197,332,731,755]" pageId="11" pageNumber="48">Etymology.</emphasis>
The specific epithet “ 
<taxonomicName id="A3A2BADCFF994123FD82FD3DCFDB1A8C" box="[576,651,731,755]" class="Amphibia" family="Aromobatidae" genus="Aromobates" kingdom="Animalia" order="Anura" pageId="11" pageNumber="48" phylum="Chordata" rank="species" species="tokuko">
<emphasis id="56D61D4DFF994123FD82FD3DCFDB1A8C" box="[576,651,731,755]" italics="true" pageId="11" pageNumber="48">tokuko</emphasis>
</taxonomicName>
” is a noun in apposition and refers to the river drainage where the species occurs.
</paragraph>
</subSubSection>
</treatment>
</document>