299 lines
32 KiB
XML
299 lines
32 KiB
XML
<document id="2333FE90FF2BFAA31D98124B4DF4778E" ID-CLB-Dataset="5605" ID-DOI="10.11646/zootaxa.3795.5.2" ID-GBIF-Dataset="87d17125-337d-4679-bdcd-102f6b022ad2" ID-ISSN="1175-5326" ID-Zenodo-Dep="4915083" ID-ZooBank="9394634C-9836-4973-868B-BDEE414E4EA8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623198152603" checkinUser="felipe" docAuthor="Petzold, Alice, Vargas-Ramírez, Mario, Kehlmaier, Christian, Vamberger, Melita, Branch, William R., Preez, Louis Du, Hofmeyr, Margaretha D., Meyer, Leon, Schleicher, Alfred, Široký, Pavel & Fritz, Uwe" docDate="2014" docId="CB02879FF927FFD1FF74F9FEFE27FCE2" docLanguage="en" docName="zt03795p548.pdf" docOrigin="Zootaxa 3795 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pelomedusa schweinfurthi Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz 2014, sp. nov." docType="treatment" docVersion="7" lastPageNumber="540" masterDocId="373BFFE7F937FFC0FFE3FFB8FFC9FFBF" masterDocTitle="A revision of African helmeted terrapins (Testudines: Pelomedusidae: Pelomedusa), with descriptions of six new species" masterLastPageNumber="548" masterPageNumber="523" pageNumber="539" updateTime="1699006367747" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3732E62D9C2C4162C72645B4054B1126">A revision of African helmeted terrapins (Testudines: Pelomedusidae: Pelomedusa), with descriptions of six new species</mods:title>
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<mods:namePart id="EEF76CCDC7716E0200A97E47334FC08F">Petzold, Alice</mods:namePart>
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<mods:affiliation id="77A66AF38069556B1600E8E7A065C94D">Museum of Zoology, Senckenberg Dresden, A. B. Meyer Building, 01109 Dresden, Germany</mods:affiliation>
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<mods:namePart id="329D19C0657F7A4015B354E241079CC3">Vargas-Ramírez, Mario</mods:namePart>
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<mods:affiliation id="9AE14E0822678DFD9D6C9159EF5AD31C">Museum of Zoology, Senckenberg Dresden, A. B. Meyer Building, 01109 Dresden, Germany</mods:affiliation>
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<mods:affiliation id="B841ACC2251CCE89AD277684E16517A0">Museum of Zoology, Senckenberg Dresden, A. B. Meyer Building, 01109 Dresden, Germany</mods:affiliation>
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<mods:namePart id="6716806979746690442B367DAFE2FFDB">Vamberger, Melita</mods:namePart>
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<mods:namePart id="34AD7CE3E54CDEC6810A53F14F64B74D">Branch, William R.</mods:namePart>
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<mods:affiliation id="A90A53CA383CBEA88668FCAB95FBA00F">Department of Herpetology, Port Elizabeth Museum, P. O. Box 13147, Humewood 6013, South Africa & Department of Zoology, P. O. Box 77000, Nelson Mandela Metropolitan University, Port Elizabeth 6031, South Africa</mods:affiliation>
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<mods:namePart id="CE8FE61E139A6171232CB589E71FB51B">Preez, Louis Du</mods:namePart>
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<mods:affiliation id="B5880BB5E394DC5C5F42EB41056D8370">Unit for Environmental Science & Management, North-West University, Potchefstroom Campus, Private Bag X 6001, Potchefstroom 2520, South Africa</mods:affiliation>
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<mods:namePart id="CCAF94C4ACAFB3941D4097F3A9899645">Hofmeyr, Margaretha D.</mods:namePart>
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<mods:namePart id="D2B5BAEA20A9F7E89EC553DBB8153530">Meyer, Leon</mods:namePart>
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<mods:namePart id="CA5B31E43634EBA8990D85B447A9CFEA">Schleicher, Alfred</mods:namePart>
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<mods:affiliation id="E467B3F5844CD60BC49872D6BCD368F3">P. O. Box 30566, Windhoek, Namibia</mods:affiliation>
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<mods:namePart id="4708932856DA9B084F0D51CB814683D0">Široký, Pavel</mods:namePart>
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<mods:affiliation id="9634E6E0C6056C80ED9F7395C5B54A5E">Department of Biology and Wildlife Diseases, Faculty of Veterinary Hygiene and Ecology, University of Veterinary and Pharmaceutical Sciences, Palackého tř. 1 / 3, 612 42 Brno, Czech Republic</mods:affiliation>
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<mods:namePart id="DA14DA808130244F6DA2CE94FA5EFA17">Fritz, Uwe</mods:namePart>
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<mods:affiliation id="F609E405BD990825CC902B0DFC288D47">Museum of Zoology, Senckenberg Dresden, A. B. Meyer Building, 01109 Dresden, Germany</mods:affiliation>
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<mods:title id="D67BD6437263B0BA153949769D7A7508">Zootaxa</mods:title>
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<mods:date id="3E679E1E63F6ECAD2945D086590E1BF5">2014</mods:date>
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<mods:number id="5A6BD1469836D9A548B581BCAE31857C">2014-05-15</mods:number>
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<mods:number id="607DCC9879E8B38E066BB5E90C992674">3795</mods:number>
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<mods:identifier id="EA68F26F3388ADB89619783DAE65BEA6" type="DOI">10.11646/zootaxa.3795.5.2</mods:identifier>
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<treatment id="CB02879FF927FFD1FF74F9FEFE27FCE2" ID-DOI="http://doi.org/10.5281/zenodo.4915099" ID-GBIF-Taxon="183403859" ID-Zenodo-Dep="4915099" LSID="urn:lsid:plazi:treatment:CB02879FF927FFD1FF74F9FEFE27FCE2" httpUri="http://treatment.plazi.org/id/CB02879FF927FFD1FF74F9FEFE27FCE2" lastPageId="17" lastPageNumber="540" pageId="16" pageNumber="539">
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<heading id="185C81E5F927FFD0FF74F9FEFDF4F9DF" bold="true" box="[151,573,1606,1632]" fontSize="11" level="1" pageId="16" pageNumber="539" reason="1">
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<emphasis id="71DFEA9BF927FFD0FF74F9FEFDF4F9DF" bold="true" box="[151,573,1606,1632]" pageId="16" pageNumber="539">
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<taxonomicName id="84AB4D0AF927FFD0FF74F9FEFE11F9DF" authority="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz, 2014" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[151,472,1606,1632]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="539" phylum="Chordata" rank="species" species="schweinfurthi" status="sp. nov.">
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<emphasis id="71DFEA9BF927FFD0FF74F9FEFE11F9DF" bold="true" box="[151,472,1606,1632]" italics="true" pageId="16" pageNumber="539">Pelomedusa schweinfurthi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="6AEC57E0F927FFD0FE03F9FEFDF4F9DF" box="[480,573,1606,1632]" pageId="16" pageNumber="539" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<subSubSection id="0BB16502F927FFD0FF74F934FA9FF8C1" pageId="16" pageNumber="539" type="diagnosis">
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<paragraph id="43143689F927FFD0FF74F934FA9FF8C1" blockId="16.[151,1436,1676,1990]" pageId="16" pageNumber="539">
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<emphasis id="71DFEA9BF927FFD0FF74F934FEDAF91A" bold="true" box="[151,275,1676,1701]" pageId="16" pageNumber="539">Diagnosis:</emphasis>
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Small to medium-sized, dark-coloured helmeted terrapins with a known maximum straight carapacial length of
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. Pectoral scutes rectangular with wide midseam contact or triangular with narrow midseam contact. Temporal head scales large, mostly undivided. Two small barbels under chin. In adults, carapace and plastron rather dark; soft parts ventrally lighter than dorsally.
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<taxonomicName id="84AB4D0AF927FFD0FC8DF941FB53F8AD" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[878,1178,1785,1810]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="539" phylum="Chordata" rank="species" species="schweinfurthi">
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<emphasis id="71DFEA9BF927FFD0FC8DF941FB53F8AD" box="[878,1178,1785,1810]" italics="true" pageId="16" pageNumber="539">Pelomedusa schweinfurthi</emphasis>
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</taxonomicName>
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differs from all other
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<taxonomicName id="84AB4D0AF927FFD0FF74F8A5FEB2F889" box="[151,379,1821,1846]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="16" pageNumber="539" phylum="Chordata" rank="species" species="undetermined">
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<emphasis id="71DFEA9BF927FFD0FF74F8A5FEE9F889" box="[151,288,1821,1846]" italics="true" pageId="16" pageNumber="539">Pelomedusa</emphasis>
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species
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</taxonomicName>
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by the presence of thymine (T) instead of cytosine (C) at position 279, by the presence of guanine (G) instead of adenine (A) or a gap at position 297, and by the presence of adenine (A) instead of guanine (G) at position 305 of the 360-bp-long reference alignment of the 12S rRNA gene (Supporting Information).
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</paragraph>
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<subSubSection id="0BB16502F927FFD0FF24F831FC33F879" pageId="16" pageNumber="539" type="materials_examined">
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<materialsCitation id="F3C33CD4F927FFD0FF24F831FC33F879" ID-GBIF-Occurrence="3319794317" accessionNumber="SMF 56161" collectingDate="1955-03-03" collectionCode="SMF" collectorName="C. Scherpner" country="South Sudan" county="Liria" latitude="4.6443334" location="Senckenberg-Museum" longLatPrecision="13" longitude="32.0805" municipality="Frankfurt am Main" pageId="16" pageNumber="539" specimenCount="1" stateProvince="Central Equatoria" typeStatus="holotype">
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<emphasis id="71DFEA9BF927FFD0FF24F831FEF4F81E" bold="true" box="[199,317,1929,1953]" pageId="16" pageNumber="539">
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<typeStatus id="9C10882BF927FFD0FF24F831FEF1F81E" box="[199,312,1929,1953]" pageId="16" pageNumber="539" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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<location id="46746052F927FFD0FEA4F831FD8FF81D" LSID="urn:lsid:plazi:treatment:CB02879FF927FFD1FF74F9FEFE27FCE2:46746052F927FFD0FEA4F831FD8FF81D" box="[327,582,1929,1954]" country="South Sudan" county="Liria" latitude="4.6443334" longLatPrecision="13" longitude="32.0805" municipality="Frankfurt am Main" name="Senckenberg-Museum" pageId="16" pageNumber="539" stateProvince="Central Equatoria">Senckenberg-Museum</location>
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,
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<collectingMunicipality id="A370ACF3F927FFD0FDB0F831FCE7F81D" box="[595,814,1929,1954]" pageId="16" pageNumber="539">Frankfurt am Main</collectingMunicipality>
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(
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<collectionCode id="25BAAE4CF927FFD0FCA3F831FCB3F81D" box="[832,890,1929,1954]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="16" pageNumber="539">SMF</collectionCode>
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56161, female,
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,
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,
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<collectingCountry id="3BBC7619F927FFD0FAB9F831FF29F879" name="South Sudan" pageId="16" pageNumber="539">South Sudan</collectingCountry>
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,
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; leg.
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<collectorName id="EE5E535FF927FFD0FDE5F815FD53F879" box="[518,666,1965,1990]" pageId="16" pageNumber="539">C. Scherpner</collectorName>
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,
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|
||
<collectingDate id="2751E9A1F927FFD0FD45F815FC8FF879" box="[678,838,1965,1990]" pageId="16" pageNumber="539" value="1955-03-03">3 March 1955</collectingDate>
|
||
</date>
|
||
;
|
||
<figureCitation id="DB902A0CF927FFD0FCB2F815FC5AF879" box="[849,915,1965,1990]" captionStart="FIGURE 4" captionStartId="15.[151,250,1913,1935]" captionTargetBox="[187,1380,204,1862]" captionTargetId="figure-68@15.[151,1436,143,1882]" captionTargetPageId="15" captionText="FIGURE 4. Dorsal and ventral aspects of the holotype of Pelomedusa neumanni sp. nov. (NMP6V 74974, adult male, Kakamega, Kenya; top) and of Pelomedusa schweinfurthi sp. nov. (SMF 56161, female, Liria, Central Equatoria, South Sudan; bottom). Scale bars, 3 cm. Photos: P. Široký and E. Morawa." figureDoi="http://doi.org/10.5281/zenodo.4915093" httpUri="https://zenodo.org/record/4915093/files/figure.png" pageId="16" pageNumber="539">Fig. 4</figureCitation>
|
||
bottom).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BB16502F926FFD1FF24FF2FFF27FE37" pageId="17" pageNumber="540" type="description">
|
||
<paragraph id="43143689F926FFD1FF24FF2FFD5CFEA2" blockId="17.[151,1437,151,861]" pageId="17" pageNumber="540">
|
||
<emphasis id="71DFEA9BF926FFD1FF24FF2FFDE9FF0F" bold="true" box="[199,544,151,176]" pageId="17" pageNumber="540">
|
||
Description of the
|
||
<typeStatus id="9C10882BF926FFD1FE53FF20FDD2FF0F" box="[432,539,152,176]" pageId="17" pageNumber="540" type="holotype">holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
Straight carapacial length
|
||
<quantity id="84539B6CF926FFD1FCBDFF2FFC09FF10" box="[862,960,151,176]" metricMagnitude="-1" metricUnit="m" metricValue="1.23" pageId="17" pageNumber="540" unit="cm" value="12.3">12.3 cm</quantity>
|
||
, plastral length
|
||
<quantity id="84539B6CF926FFD1FB9CFF2FFB2BFF10" box="[1151,1250,151,176]" metricMagnitude="-1" metricUnit="m" metricValue="1.05" pageId="17" pageNumber="540" unit="cm" value="10.5">10.5 cm</quantity>
|
||
. Pectoral scutes rectangular with wide midseam contact. Temporal scale divided on left side, undivided on right side of head. Two small barbels under chin. Carapace and plastron dark brown. Dorsal side of soft parts and entire forelegs dark grey, ventral side of hind legs light grey. Tail short.
|
||
</paragraph>
|
||
<paragraph id="43143689F926FFD1FF24FE90FF27FE37" blockId="17.[151,1437,151,861]" pageId="17" pageNumber="540">
|
||
<materialsCitation id="F3C33CD4F926FFD1FF24FE90FF23FE37" ID-GBIF-Occurrence="3319794309" accessionNumber="SMF 56160, ZMB 15697" collectingDate="1955-03-03" collectionCode="SMF, ZMB" country="South Sudan" county="Liria" latitude="4.6443334" location="Frankfurt am Main" longLatPrecision="13" longitude="32.0805" municipality="Djur River" pageId="17" pageNumber="540" specimenCode="65162" specimenCount="1" stateProvince="Western Bahr el Ghazal" typeStatus="paratype">
|
||
<emphasis id="71DFEA9BF926FFD1FF24FE90FE81FEFF" bold="true" box="[199,328,296,320]" pageId="17" pageNumber="540">
|
||
<typeStatus id="9C10882BF926FFD1FF24FE90FE8AFEFF" box="[199,323,296,320]" pageId="17" pageNumber="540" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
Senckenberg-Museum,
|
||
<location id="46746052F926FFD1FDBDFE9FFCF1FEFF" LSID="urn:lsid:plazi:treatment:CB02879FF927FFD1FF74F9FEFE27FCE2:46746052F926FFD1FDBDFE9FFCF1FEFF" box="[606,824,295,320]" country="South Sudan" municipality="Djur River" name="Frankfurt am Main" pageId="17" pageNumber="540" stateProvince="Western Bahr el Ghazal">Frankfurt am Main</location>
|
||
(
|
||
<collectionCode id="25BAAE4CF926FFD1FCAAFE9FFC4AFEFF" box="[841,899,295,320]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="17" pageNumber="540">SMF</collectionCode>
|
||
56160,
|
||
<specimenCode id="130D9EF2F926FFD1FC00FE9FFBE5FEFF" box="[995,1068,295,320]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="17" pageNumber="540">65162</specimenCode>
|
||
, males, same data as holotype);
|
||
<collectorName id="EE5E535FF926FFD1FF74FEF4FF30FEDC" box="[151,249,332,355]" pageId="17" pageNumber="540">Museum</collectorName>
|
||
für
|
||
<collectorName id="EE5E535FF926FFD1FED1FEF4FE70FEDA" box="[306,441,332,357]" pageId="17" pageNumber="540">Naturkunde</collectorName>
|
||
,
|
||
<collectorName id="EE5E535FF926FFD1FE29FEF4FDD9FEDA" box="[458,528,332,357]" pageId="17" pageNumber="540">Berlin</collectorName>
|
||
(
|
||
<collectionCode id="25BAAE4CF926FFD1FDC7FEF4FDABFEDA" box="[548,610,332,357]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="17" pageNumber="540" type="Museum">ZMB</collectionCode>
|
||
15697, juvenile,
|
||
<collectorName id="EE5E535FF926FFD1FCD5FEF4FBD4FEDC" box="[822,1053,332,357]" pageId="17" pageNumber="540">
|
||
<collectingMunicipality id="A370ACF3F926FFD1FCD5FEF4FC7DFEDA" box="[822,948,332,357]" pageId="17" pageNumber="540">Djur River</collectingMunicipality>
|
||
W Wau
|
||
</collectorName>
|
||
,
|
||
<collectingRegion id="816FF86BF926FFD1FBCEFEF4FA83FEDA" box="[1069,1354,332,357]" country="South Sudan" name="Western Bahr el Ghazal" pageId="17" pageNumber="540">Western Bahr el Ghazal</collectingRegion>
|
||
,
|
||
<collectingCountry id="3BBC7619F926FFD1FAB8FEF4FF2BFE37" name="South Sudan" pageId="17" pageNumber="540">South Sudan</collectingCountry>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BB16502F926FFD1FF24FE2BFD0AFDA7" pageId="17" pageNumber="540" type="etymology">
|
||
<paragraph id="43143689F926FFD1FF24FE2BFD0AFDA7" blockId="17.[151,1437,151,861]" pageId="17" pageNumber="540">
|
||
<emphasis id="71DFEA9BF926FFD1FF24FE2BFE66FE13" bold="true" box="[199,431,403,428]" pageId="17" pageNumber="540">Derivatio nominis:</emphasis>
|
||
We dedicate this species to Georg August Schweinfurth (
|
||
<date id="37151049F926FFD1FB6AFE2CFABAFE12" box="[1161,1395,404,429]" pageId="17" pageNumber="540" value="1836-12-29">29 December 1836</date>
|
||
–
|
||
<date id="37151049F926FFD1FA61FE2CFE9DFE6F" pageId="17" pageNumber="540" value="1925-09-19">19 September 1925</date>
|
||
) who was the last of the great explorers of Africa. Schweinfurth collected the oldest of the
|
||
<typeStatus id="9C10882BF926FFD1FF74FE64FECBFE4B" box="[151,258,476,500]" pageId="17" pageNumber="540" type="paratype">paratypes</typeStatus>
|
||
(ZMB 15697) during his famous third Africa expedition (1868–1871) “inside the heart of Africa” (
|
||
<bibRefCitation id="273A4B78F926FFD1FF7DFE47FEBEFDA7" author="Schweinfurth, G." box="[158,375,511,536]" pageId="17" pageNumber="540" refId="ref17806" refString="Schweinfurth, G. (1874) Im Herzen von Afrika. Reisen und Entdeckungen im centralen Aequatorial-Afrika wahrend der Jahre 1868 bis 1871. Volume 1 and 2. F. A. Brockhaus, Leipzig, XIII + 599 pp. (Vol. 1), XII + 561 pp., 2 maps (Vol. 2)." type="book" year="1874">Schweinfurth 1874</bibRefCitation>
|
||
) in present-day
|
||
<collectingCountry id="3BBC7619F926FFD1FDCEFE47FD77FDA7" box="[557,702,511,536]" name="South Sudan" pageId="17" pageNumber="540">South Sudan</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BB16502F926FFD1FF24FD9BFAEBFD82" box="[199,1314,547,573]" pageId="17" pageNumber="540" type="distribution">
|
||
<paragraph id="43143689F926FFD1FF24FD9BFAEBFD82" blockId="17.[151,1437,151,861]" box="[199,1314,547,573]" pageId="17" pageNumber="540">
|
||
<emphasis id="71DFEA9BF926FFD1FF24FD9BFEABFD83" bold="true" box="[199,354,547,572]" pageId="17" pageNumber="540">Distribution:</emphasis>
|
||
<collectingCountry id="3BBC7619F926FFD1FE8AFD9CFD4DFD82" box="[361,644,548,573]" name="Central African Republic" pageId="17" pageNumber="540">Central African Republic</collectingCountry>
|
||
(
|
||
<bibRefCitation id="273A4B78F926FFD1FD70FD9CFC01FD82" author="Vargas-Ramirez, M. & Vences, M. & Branch, W. R. & Daniels, S. R. & Glaw, F. & Hofmeyr, M. D. & Kuchling, G. & Maran, J. & Papenfuss, T. J. & Siroky, P. & Vieites, D. R. & Fritz, U." box="[659,968,548,573]" pageId="17" pageNumber="540" pagination="428 - 440" refId="ref18768" refString="Vargas-Ramirez, M., Vences, M., Branch, W. R., Daniels, S. R., Glaw, F., Hofmeyr, M. D., Kuchling, G., Maran, J., Papenfuss, T. J., Siroky, P., Vieites, D. R. & Fritz, U. (2010) Deep genealogical lineages in the widely distributed African helmeted terrapin: evidence from mitochondrial and nuclear DNA (Testudines: Pelomedusidae: Pelomedusa subrufa). Molecular Phylogenetics and Evolution, 56, 428 - 440. http: // dx. doi. org / 10.1016 / j. ympev. 2010.03.019" type="journal article" year="2010">
|
||
Vargas-Ramírez
|
||
<emphasis id="71DFEA9BF926FFD1FCACFD9DFC41FD82" box="[847,904,548,573]" italics="true" pageId="17" pageNumber="540">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
) and
|
||
<collectingCountry id="3BBC7619F926FFD1FBE5FD9CFB5CFD82" box="[1030,1173,548,573]" name="South Sudan" pageId="17" pageNumber="540">South Sudan</collectingCountry>
|
||
(this study).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BB16502F926FFD1FF24FDF0FE27FCE2" pageId="17" pageNumber="540" type="discussion">
|
||
<paragraph id="43143689F926FFD1FF24FDF0FD2EFCAA" blockId="17.[151,1437,151,861]" pageId="17" pageNumber="540">
|
||
<emphasis id="71DFEA9BF926FFD1FF24FDF0FEF4FDDF" bold="true" box="[199,317,584,608]" pageId="17" pageNumber="540">Remarks:</emphasis>
|
||
<taxonomicName id="84AB4D0AF926FFD1FEABFDFFFDBCFDDF" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[328,629,583,608]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="17" pageNumber="540" phylum="Chordata" rank="species" species="schweinfurthi">
|
||
<emphasis id="71DFEA9BF926FFD1FEABFDFFFDBCFDDF" box="[328,629,583,608]" italics="true" pageId="17" pageNumber="540">Pelomedusa schweinfurthi</emphasis>
|
||
</taxonomicName>
|
||
corresponds to mtDNA lineage IV of
|
||
<bibRefCitation id="273A4B78F926FFD1FBA0FDFFFA5EFDDF" author="Vargas-Ramirez, M. & Vences, M. & Branch, W. R. & Daniels, S. R. & Glaw, F. & Hofmeyr, M. D. & Kuchling, G. & Maran, J. & Papenfuss, T. J. & Siroky, P. & Vieites, D. R. & Fritz, U." box="[1091,1431,583,608]" pageId="17" pageNumber="540" pagination="428 - 440" refId="ref18768" refString="Vargas-Ramirez, M., Vences, M., Branch, W. R., Daniels, S. R., Glaw, F., Hofmeyr, M. D., Kuchling, G., Maran, J., Papenfuss, T. J., Siroky, P., Vieites, D. R. & Fritz, U. (2010) Deep genealogical lineages in the widely distributed African helmeted terrapin: evidence from mitochondrial and nuclear DNA (Testudines: Pelomedusidae: Pelomedusa subrufa). Molecular Phylogenetics and Evolution, 56, 428 - 440. http: // dx. doi. org / 10.1016 / j. ympev. 2010.03.019" type="journal article" year="2010">
|
||
Vargas-Ramírez
|
||
<emphasis id="71DFEA9BF926FFD1FAE1FDF1FAF6FDDF" box="[1282,1343,583,608]" italics="true" pageId="17" pageNumber="540">et al.</emphasis>
|
||
(2010)
|
||
</bibRefCitation>
|
||
, which was previously only recorded from the
|
||
<collectingCountry id="3BBC7619F926FFD1FD5FFDD4FC2CFD3A" box="[700,997,620,645]" name="Central African Republic" pageId="17" pageNumber="540">Central African Republic</collectingCountry>
|
||
. Our new 12S sequences of the type specimens provide evidence for the occurrence in
|
||
<collectingCountry id="3BBC7619F926FFD1FD25FD37FC91FD17" box="[710,856,655,680]" name="South Sudan" pageId="17" pageNumber="540">South Sudan</collectingCountry>
|
||
. The temporal scale is divided on the left side of the head in the
|
||
<typeStatus id="9C10882BF926FFD1FEFEFD0CFE4BFD72" box="[285,386,692,717]" pageId="17" pageNumber="540" type="holotype">holotype</typeStatus>
|
||
, but undivided on the right side, as it is in all
|
||
<typeStatus id="9C10882BF926FFD1FC73FD0CFC36FD73" box="[912,1023,692,716]" pageId="17" pageNumber="540" type="paratype">paratypes</typeStatus>
|
||
. The largest specimen (SMF 56162) has a straight carapacial length of
|
||
<quantity id="84539B6CF926FFD1FDF1FD6FFDA6FD50" box="[530,623,727,752]" metricMagnitude="-1" metricUnit="m" metricValue="1.5699999999999998" pageId="17" pageNumber="540" unit="cm" value="15.7">15.7 cm</quantity>
|
||
. Unlike many other
|
||
<taxonomicName id="84AB4D0AF926FFD1FCB3FD6FFBFAFD4F" box="[848,1075,727,752]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="17" pageNumber="540" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="71DFEA9BF926FFD1FCB3FD6FFC10FD4F" box="[848,985,727,752]" italics="true" pageId="17" pageNumber="540">Pelomedusa</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="84AB4D0AF926FFD1FBDEFD61FB39FD4F" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[1085,1264,727,752]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="17" pageNumber="540" phylum="Chordata" rank="species" species="schweinfurthi">
|
||
<emphasis id="71DFEA9BF926FFD1FBDEFD61FB39FD4F" box="[1085,1264,727,752]" italics="true" pageId="17" pageNumber="540">P. schweinfurthi</emphasis>
|
||
</taxonomicName>
|
||
is overall quite dark coloured, with a predominantly brown plastron.
|
||
</paragraph>
|
||
<paragraph id="43143689F926FFD1FF24FCA7FE27FCE2" blockId="17.[151,1437,151,861]" pageId="17" pageNumber="540">
|
||
<taxonomicName id="84AB4D0AF926FFD1FF24FCA7FE38FC87" authorityName="Petzold & Vargas-Ramírez & Kehlmaier & Vamberger & Branch & Preez & Hofmeyr & Meyer & Schleicher & Široký & Fritz" authorityYear="2014" box="[199,497,799,824]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="17" pageNumber="540" phylum="Chordata" rank="species" species="schweinfurthi">
|
||
<emphasis id="71DFEA9BF926FFD1FF24FCA7FE38FC87" box="[199,497,799,824]" italics="true" pageId="17" pageNumber="540">Pelomedusa schweinfurthi</emphasis>
|
||
</taxonomicName>
|
||
belongs to the northern species group and is with high support sister to candidate species B from
|
||
<collectingCountry id="3BBC7619F926FFD1FEA5FCFCFE45FCE2" box="[326,396,836,861]" name="Sudan" pageId="17" pageNumber="540">Sudan</collectingCountry>
|
||
(
|
||
<figureCitation id="DB902A0CF926FFD1FE78FCFCFE15FCE2" box="[411,476,836,861]" captionStart="FIGURE 1" captionStartId="3.[151,250,999,1021]" captionTargetBox="[167,1426,219,969]" captionTargetId="figure-480@3.[151,1436,199,975]" captionTargetPageId="3" captionText="FIGURE 1. Maximum Likelihood tree for helmeted terrapins (Pelomedusa spp.) using 1848 bp of mitochondrial DNA (12S, cyt b, ND4+tRNAs), rooted with Pelusios sinuatus. Terminal clades collapsed to cartoons. The topology of the Bayesian 50% majority rule consensus tree was identical. Numbers along branches indicate bootstrap support and clade support under BI (posterior probabilities) greater than 50 or 0.95, respectively. Asterisks represent maximum support under both methods. Colours and symbol correspond to the map (Fig. 2). On the right, proposed species names indicated; in brackets, mtDNA lineages. For clade membership of individual samples, see Table S1. The questionable sample MTD T 5484 from Swellendam District (Western Cape), South Africa, is highlighted in red (see text). The shown terrapins are Pelomedusa neumanni (top; Kakamega, Kenya) and P. galeata (bottom; Port Elizabeth, South Africa—photos: H. Prokop and W.R. Branch)." figureDoi="http://doi.org/10.5281/zenodo.4915085" httpUri="https://zenodo.org/record/4915085/files/figure.png" pageId="17" pageNumber="540">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |