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<document id="F7226434475AFB3A9C8201B28ACC7A60" ID-DOI="10.24823/EJB.2021.364" ID-ISSN="1747-0036" ID-Zenodo-Dep="10515418" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1704823505549" checkinUser="felipe" docAuthor="Atkins, H. J. &amp; Kartonegoro, A." docDate="2021" docId="AA395B43AC0CFFC3BC69BEEAFC97FC00" docLanguage="en" docName="EdinbJBot.78.364.1-122.pdf" docOrigin="Edinburgh Journal of Botany 78 (364)" docSource="http://dx.doi.org/10.24823/ejb.2021.364" docStyle="DocumentStyle:73DE8CF451BB5BCBD336E07A8276E690.2:EdinbJBot.2021-.journal_article" docStyleId="73DE8CF451BB5BCBD336E07A8276E690" docStyleName="EdinbJBot.2021-.journal_article" docStyleVersion="2" docTitle="Cyrtandra hekensis Karton. &amp; H. J. Atkins" docType="treatment" docVersion="4" lastPageNumber="51" masterDocId="5600233BAC3CFFF1BC10BF13FF94FFB3" masterDocTitle="A TAXONOMIC REVISION OF CYRTANDRA (GESNERIACEAE) IN SULAWESI, INDONESIA" masterLastPageNumber="122" masterPageNumber="1" pageNumber="49" updateTime="1705363298848" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="71EE1417006662352724EC8D1333D58C">A TAXONOMIC REVISION OF CYRTANDRA (GESNERIACEAE) IN SULAWESI, INDONESIA</mods:title>
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<mods:namePart id="639DF0D73971B7BC3D93EA591537D802">Atkins, H. J.</mods:namePart>
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<mods:namePart id="147A604582B677F7A4EA9613DA478AF4">Kartonegoro, A.</mods:namePart>
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<mods:title id="D5573A48667F8C9A9A9141836EDC75C5">Edinburgh Journal of Botany</mods:title>
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<mods:part id="7D7EC0D76E64FF33B9A5068728B32CA6">
<mods:date id="3B5F55B867080F78944185F449632411">2021</mods:date>
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<mods:number id="015568053FA7D0470DA9DA4195E8D153">2021-09-08</mods:number>
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<mods:classification id="03AE98AFF7B73A08C466EE3C3EDB5099">journal article</mods:classification>
<mods:identifier id="A99494570F05EE2D47F1EF019FE03E2C" type="DOI">10.24823/EJB.2021.364</mods:identifier>
<mods:identifier id="8D46F93FFF2345D8ADBEDA9D599B4D11" type="ISSN">1747-0036</mods:identifier>
<mods:identifier id="13019A06E0BF3151B1486394AD3B2925" type="Zenodo-Dep">10515418</mods:identifier>
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<treatment id="AA395B43AC0CFFC3BC69BEEAFC97FC00" ID-DOI="http://doi.org/10.5281/zenodo.10515613" ID-Zenodo-Dep="10515613" LSID="urn:lsid:plazi:treatment:AA395B43AC0CFFC3BC69BEEAFC97FC00" httpUri="http://treatment.plazi.org/id/AA395B43AC0CFFC3BC69BEEAFC97FC00" lastPageId="50" lastPageNumber="51" pageId="48" pageNumber="49">
<subSubSection id="6A8AB9DEAC0CFFC1BC69BEEAFB9EFDA1" box="[121,1034,505,530]" pageId="48" pageNumber="49" type="nomenclature">
<paragraph id="222FEA55AC0CFFC1BC69BEEAFB9EFDA1" blockId="48.[121,1141,505,604]" box="[121,1034,505,530]" pageId="48" pageNumber="49">
<heading id="79675D39AC0CFFC1BC69BEEAFB9EFDA1" box="[121,1034,505,530]" fontSize="36" level="2" pageId="48" pageNumber="49" reason="2">
<emphasis id="10E43647AC0CFFC1BC69BEEAFF09FDA1" bold="true" box="[121,157,505,530]" pageId="48" pageNumber="49">13.</emphasis>
<taxonomicName id="E59091D6AC0CFFC1BCB4BEEAFDF8FDA1" ID-CoL="33KD5" authority="Karton. &amp; H. J. Atkins" authorityName="Karton. &amp; H. J. Atkins" box="[164,620,505,530]" class="Magnoliopsida" family="Gesneriaceae" genus="Cyrtandra" kingdom="Plantae" order="Lamiales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="species" species="hekensis">
<treatmentCitation id="A331CC44AC0CFFC1BCB4BEEAFEEBFDA1" author="Karton. &amp; H. J. Atkins" box="[164,383,505,530]" issue="2" journal="Edinburgh J. Bot." page="182" pageId="48" pageNumber="49" score="11.793866417724223" volume="75" year="2018">
<emphasis id="10E43647AC0CFFC1BCB4BEEAFEEBFDA1" bold="true" box="[164,383,505,530]" pageId="48" pageNumber="49">Cyrtandra hekensis</emphasis>
</treatmentCitation>
<materialsCitation id="92F8E008AC0CFFC1BD96BEEAFDF8FDA1" box="[390,620,505,530]" collectorName="J. Bot." location="Edinburgh" pageId="48" pageNumber="49" specimenCount="1" stateProvince="City of Edinburgh">Karton. &amp; H.J.Atkins</materialsCitation>
</taxonomicName>
,
<bibCitation id="F7BAE239AC0CFFC1BE66BEEAFB93FDA1" box="[630,1031,505,530]" issue="2" journal="Edinburgh J. Bot." page="182" pageId="48" pageNumber="49" score="11.793866417724223" volume="75" year="2018">
<collectingRegion id="E05424B7AC0CFFC1BE66BEEAFD73FDA1" box="[630,743,505,530]" country="United Kingdom" name="City of Edinburgh" pageId="48" pageNumber="49">Edinburgh</collectingRegion>
<collectorName id="8F658F83AC0CFFC1BEFEBEEAFCA3FDA1" box="[750,823,505,530]" pageId="48" pageNumber="49">J. Bot.</collectorName>
75(2): 182 (2018)
</bibCitation>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6A8AB9DEAC0CFFC1B800BEEAFE26FDEF" pageId="48" pageNumber="49" type="materials_examined">
<paragraph id="222FEA55AC0CFFC1B800BEEAFE26FDEF" blockId="48.[121,1141,505,604]" pageId="48" pageNumber="49">
–
<typeStatus id="FD2B54F7AC0CFFC1B837BEEAFBF4FDA1" box="[1063,1120,505,530]" pageId="48" pageNumber="49">Type</typeStatus>
:
<materialsCitation id="92F8E008AC0CFFC1BC87BD0DFF2AFDEF" collectingDate="2008-04-09" collectionCode="BO" collectorName="Thomas &amp; Ardi" country="Indonesia" location="Mt Hek" pageId="48" pageNumber="49" specimenCount="1" stateProvince="Sulawesi" typeStatus="holotype">
<collectingCountry id="5A87AAC5AC0CFFC1BC87BD0DFE93FD84" box="[151,263,542,567]" name="Indonesia" pageId="48" pageNumber="49">Indonesia</collectingCountry>
,
<collectingRegion id="E05424B7AC0CFFC1BD01BD0DFEECFD84" box="[273,376,542,567]" country="Indonesia" name="Sulawesi Utara" pageId="48" pageNumber="49">Sulawesi</collectingRegion>
,
<collectingRegion id="E05424B7AC0CFFC1BD92BD0DFDD4FD84" box="[386,576,542,567]" country="Indonesia" name="Sulawesi Tengah" pageId="48" pageNumber="49">Central Sulawesi</collectingRegion>
,
<location id="274FBC8EAC0CFFC1BE59BD0DFD08FD84" LSID="urn:lsid:plazi:treatment:AA395B43AC0CFFC3BC69BEEAFC97FC00:274FBC8EAC0CFFC1BE59BD0DFD08FD84" box="[585,668,542,567]" country="Indonesia" name="Mt Hek" pageId="48" pageNumber="49" stateProvince="Sulawesi">Mt Hek</location>
,
<date id="562ECC95AC0CFFC1BEB7BD0DFC84FD84" box="[679,784,542,567]" pageId="48" pageNumber="49" value="2008-04-09">
<collectingDate id="466A357DAC0CFFC1BEB7BD0DFC84FD84" box="[679,784,542,567]" pageId="48" pageNumber="49" value="2008-04-09">9 iv 2008</collectingDate>
</date>
,
<emphasis id="10E43647AC0CFFC1BF0ABD0DFC29FD84" box="[794,957,542,567]" italics="true" pageId="48" pageNumber="49">
<collectorName id="8F658F83AC0CFFC1BF0ABD0DFCE1FD84" box="[794,885,542,567]" pageId="48" pageNumber="49">Thomas</collectorName>
&amp;
<collectorName id="8F658F83AC0CFFC1BF81BD0DFC29FD84" box="[913,957,542,567]" pageId="48" pageNumber="49">Ardi</collectorName>
</emphasis>
08-26 (
<typeStatus id="FD2B54F7AC0CFFC1B801BD0DFBE1FD84" box="[1041,1141,542,567]" pageId="48" pageNumber="49" type="holotype">holotype</typeStatus>
<collectionCode id="44817290AC0CFFC1BC88BD50FF2EFDEF" box="[152,186,579,604]" pageId="48" pageNumber="49">BO</collectionCode>
,
</materialsCitation>
<materialsCitation id="92F8E008AC0CFFC1BCD4BD50FE3FFDEF" box="[196,427,579,604]" collectingDate="2008-04-09" collectionCode="E" collectorName="Thomas &amp; Ardi" country="Indonesia" location="Mt Hek" pageId="48" pageNumber="49" specimenCount="1" stateProvince="Sulawesi Utara" typeStatus="isotype">
<typeStatus id="FD2B54F7AC0CFFC1BCD4BD50FE82FDEF" box="[196,278,579,604]" pageId="48" pageNumber="49" type="isotype">isotype</typeStatus>
<collectionCode id="44817290AC0CFFC1BD0CBD50FEBAFDEF" box="[284,302,579,604]" pageId="48" pageNumber="49">E</collectionCode>
).
<figureCitation id="BAABF6D0AC0CFFC1BD51BD50FE3FFDEF" box="[321,427,579,604]" captionStart="Figure 26" captionStartId="49.[121,185,1551,1573]" captionTargetBox="[237,1047,268,1530]" captionTargetId="figure-6@49.[225,1061,265,1533]" captionTargetPageId="49" captionText="Figure 26. Cyrtandra hekensis. a, Habit; b, opened corolla, showing position of stamens and staminodes; c, opened calyx, showing inner surface; d, side view of corolla; e, gynoecium, showing disc and top of pedicel; f, fruit with persistent calyx partially removed; g, bracts. Drawn by Claire Banks from Hendrian et al. 901. Reproduced from Kartonegoro et al. (2018) with the permission of the Trustees of the Royal Botanic Garden Edinburgh." figureDoi="http://doi.org/10.5281/zenodo.10515472" httpUri="https://zenodo.org/record/10515472/files/figure.png" pageId="48" pageNumber="49">
<emphasis id="10E43647AC0CFFC1BD51BD50FE3FFDEF" bold="true" box="[321,427,579,604]" pageId="48" pageNumber="49">Figure 26</emphasis>
</figureCitation>
</materialsCitation>
<emphasis id="10E43647AC0CFFC1BDBBBD50FE26FDEF" bold="true" box="[427,434,579,604]" pageId="48" pageNumber="49">.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="6A8AB9DEAC0CFFC1BC69BD6BFBDFFA75" pageId="48" pageNumber="49" type="description">
<paragraph id="222FEA55AC0CFFC1BC69BD6BFBDFFA75" blockId="48.[121,1155,632,1478]" pageId="48" pageNumber="49">
Herb to
<quantity id="E56847B0AC0CFFC1BCC3BD6BFE8EFD22" box="[211,282,632,657]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" pageId="48" pageNumber="49" unit="cm" value="25.0">25 cm</quantity>
in height.
<emphasis id="10E43647AC0CFFC1BD80BD6BFE42FD22" box="[400,470,632,657]" italics="true" pageId="48" pageNumber="49">Stems</emphasis>
ridged, striate, upper stem appearing reddish, from dense, coarse indumentum on young growth, older stems glabrous.
<emphasis id="10E43647AC0CFFC1BF38BD8EFCE2FD05" box="[808,886,669,694]" italics="true" pageId="48" pageNumber="49">Leaves</emphasis>
opposite, slightly unequal; those of a pair well developed but one slightly smaller; petioles
<quantity id="E56847B0AC0CFFC1BFBBBDD1FB83FD68" box="[939,1047,706,731]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="2.0" metricValueMin="0.8" pageId="48" pageNumber="49" unit="cm" value="1.4" valueMax="2.0" valueMin="0.8">0.8–2 cm</quantity>
long, densely, coarse-hairy on young growth becoming glabrous on older growth; blades 7–7.5 ×
<quantity id="E56847B0AC0CFFC1BC9DBC1EFF6DFC95" box="[141,249,781,806]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" metricValueMax="3.0" metricValueMin="1.6" pageId="48" pageNumber="49" unit="cm" value="2.3" valueMax="3.0" valueMin="1.6">1.6–3 cm</quantity>
(larger leaves), 3.5–5 ×
<quantity id="E56847B0AC0CFFC1BE1ABC1EFD1FFC95" box="[522,651,781,806]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.7" metricValueMin="1.3" pageId="48" pageNumber="49" unit="cm" value="1.5" valueMax="1.7" valueMin="1.3">1.3–1.7 cm</quantity>
(smaller leaves), narrow elliptic or oblong to somewhat oblanceolate, base decurrent, only narrowly winged at base, margin serrate, apex acuminate; 4–6 pairs of lateral veins and reticulate tertiary venation; dark green and mostly glabrous above with some hairs towards the base, paler green and with scattered hairs below with coarse hairs on the midrib and veins.
<emphasis id="10E43647AC0CFFC1BEF0BCB1FC16FC08" box="[736,898,930,955]" italics="true" pageId="48" pageNumber="49">Inflorescences</emphasis>
axillary, subsessile to shortly pedunculate, 1- or 2-flowered; peduncles red,
<quantity id="E56847B0AC0CFFC1BEF9BCDBFCDDFC52" box="[745,841,968,993]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="48" pageNumber="49" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
long, hairy; bracts green, 9 ×
<quantity id="E56847B0AC0CFFC1BCB2BCFEFE90FBB5" box="[162,260,1005,1030]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="48" pageNumber="49" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
, ovate, connate c.
<quantity id="E56847B0AC0CFFC1BDDDBCFEFD84FBB5" box="[461,528,1005,1030]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="48" pageNumber="49" unit="mm" value="4.0">4 mm</quantity>
from base, with some coarse hairs on midrib and margins, margin slightly serrate; bracteoles paired, green, 6–8 ×
<quantity id="E56847B0AC0CFFC1BF5ABB01FC1AFB98" box="[842,910,1042,1067]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="48" pageNumber="49" unit="mm" value="3.0">3 mm</quantity>
, oblong, with coarse hairs on midrib and margins; pedicels
<quantity id="E56847B0AC0CFFC1BE38BB2BFD1CFBE2" box="[552,648,1080,1105]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="48" pageNumber="49" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
long, glabrous.
<emphasis id="10E43647AC0CFFC1BF2BBB2BFCE3FBE2" box="[827,887,1080,1105]" italics="true" pageId="48" pageNumber="49">Calyx</emphasis>
tubular, pale green,
<quantity id="E56847B0AC0CFFC1BC6ABB4EFF62FBC5" box="[122,246,1117,1142]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="48" pageNumber="49" unit="mm" value="10.5" valueMax="11.0" valueMin="10.0">10–11 mm</quantity>
long, upper lobes c.
<quantity id="E56847B0AC0CFFC1BDC8BB4EFD8EFBC5" box="[472,538,1117,1142]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="48" pageNumber="49" unit="mm" value="2.0">2 mm</quantity>
long, lower lobes c.
<quantity id="E56847B0AC0CFFC1BEEABB4EFCA8FBC5" box="[762,828,1117,1142]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="48" pageNumber="49" unit="mm" value="3.0">3 mm</quantity>
long, acuminate, glabrous except for a few tufts of hair at apex.
<emphasis id="10E43647AC0CFFC1BE32BB91FDE4FB28" box="[546,624,1154,1179]" italics="true" pageId="48" pageNumber="49">Corolla</emphasis>
white with greenish yellow markings in throat, c.
<quantity id="E56847B0AC0CFFC1BC9BBBBBFF49FB72" box="[139,221,1192,1217]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="48" pageNumber="49" unit="mm" value="15.0">15 mm</quantity>
long, tube narrow in lower 1/3, widening to mouth in apical 2/3, hairy externally in the apical 1/3, particularly densely so towards the mouth, upper lobes rounded, 3–4 ×
<quantity id="E56847B0AC0CFFC1BC6ABBE1FF29FAB8" box="[122,189,1266,1291]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="48" pageNumber="49" unit="mm" value="3.0">3 mm</quantity>
, lower and lateral lobes rounded, c.3 ×
<quantity id="E56847B0AC0CFFC1BE63BBE1FD2CFAB8" box="[627,696,1266,1291]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="48" pageNumber="49" unit="mm" value="3.0">3 mm</quantity>
.
<emphasis id="10E43647AC0CFFC1BED2BBE1FCB0FAB8" box="[706,804,1266,1291]" italics="true" pageId="48" pageNumber="49">Stamens</emphasis>
with filaments c.
<quantity id="E56847B0AC0CFFC1BFF3BBE1FBB1FAB8" box="[995,1061,1266,1291]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="48" pageNumber="49" unit="mm" value="5.0">5 mm</quantity>
long, attached c.
<quantity id="E56847B0AC0CFFC1BCE5BA0BFEACFA82" box="[245,312,1304,1329]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="48" pageNumber="49" unit="mm" value="6.0">6 mm</quantity>
from base of corolla, glabrous; anthers c.
<quantity id="E56847B0AC0CFFC1BF02BA0BFCDBFA82" box="[786,847,1304,1329]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="48" pageNumber="49" unit="mm" value="1.0">1 mm</quantity>
long, coherent at apices; staminodes 2, c.
<quantity id="E56847B0AC0CFFC1BD20BA2EFE1DFAE5" box="[304,393,1341,1366]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="48" pageNumber="49" unit="mm" value="0.5">0.5 mm</quantity>
long.
<emphasis id="10E43647AC0CFFC1BDDEBA2EFDDEFAE5" box="[462,586,1341,1366]" italics="true" pageId="48" pageNumber="49">Gynoecium</emphasis>
c.
<quantity id="E56847B0AC0CFFC1BE73BA2EFD20FAE5" box="[611,692,1341,1366]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="48" pageNumber="49" unit="mm" value="10.0">10 mm</quantity>
long; disc cupular with lobed margin, glabrous externally, c.
<quantity id="E56847B0AC0CFFC1BD7BBA71FE57FAC8" box="[363,451,1378,1403]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="48" pageNumber="49" unit="mm" value="1.5">1.5 mm</quantity>
long; ovary glabrous, style eglandular hairy towards apex; stigma slightly bilobed, lobes small, c.
<quantity id="E56847B0AC0CFFC1BE38BA9BFDF2FA12" box="[552,614,1416,1441]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="48" pageNumber="49" unit="mm" value="1.0">1 mm</quantity>
across.
<emphasis id="10E43647AC0CFFC1BED7BA9BFC92FA12" box="[711,774,1416,1441]" italics="true" pageId="48" pageNumber="49">Fruits</emphasis>
ovoid, glabrous, verrucose, drying light brown, 6 ×
<quantity id="E56847B0AC0CFFC1BD3DBABEFEE4FA75" box="[301,368,1453,1478]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="48" pageNumber="49" unit="mm" value="4.0">4 mm</quantity>
; base of style and calyx persistent, bracts sometimes persistent.
</paragraph>
</subSubSection>
<subSubSection id="6A8AB9DEAC0CFFC3BC69BAF1FC97FC00" lastPageId="50" lastPageNumber="51" pageId="48" pageNumber="49" type="distribution">
<paragraph id="222FEA55AC0CFFC1BC69BAF1FDEEFA48" blockId="48.[121,634,1506,1531]" box="[121,634,1506,1531]" pageId="48" pageNumber="49">
<emphasis id="10E43647AC0CFFC1BC69BAF1FF6CFA48" box="[121,248,1506,1531]" italics="true" pageId="48" pageNumber="49">Distribution</emphasis>
.
<collectingRegion id="E05424B7AC0CFFC1BD16BAF1FE56FA48" box="[262,450,1506,1531]" country="Indonesia" name="Sulawesi Tengah" pageId="48" pageNumber="49">Central Sulawesi</collectingRegion>
(see
<figureCitation id="BAABF6D0AC0CFFC1BE11BAF1FDFFFA48" box="[513,619,1506,1531]" captionStart="Figure 29" captionStartId="55.[121,185,1256,1278]" captionTargetBox="[246,1039,269,1233]" captionTargetId="figure-117@55.[243,1042,265,1237]" captionTargetPageId="55" captionText="Figure 29. Distribution of Cyrtandra gorontaloensis (black diamonds), C. hekensis and C. hendrianii (green star) and C. hispidula (black triangle)." figureDoi="http://doi.org/10.5281/zenodo.10515480" httpUri="https://zenodo.org/record/10515480/files/figure.png" pageId="48" pageNumber="49">
<emphasis id="10E43647AC0CFFC1BE11BAF1FDFFFA48" bold="true" box="[513,619,1506,1531]" pageId="48" pageNumber="49">Figure 29</emphasis>
</figureCitation>
).
</paragraph>
<paragraph id="222FEA55AC0CFFC1BC69B905FC88F99C" blockId="48.[121,796,1558,1583]" box="[121,796,1558,1583]" pageId="48" pageNumber="49">
<emphasis id="10E43647AC0CFFC1BC69B905FEC3F99C" box="[121,343,1558,1583]" italics="true" pageId="48" pageNumber="49">Habitat and ecology</emphasis>
. Hill forest at an altitude of
<quantity id="E56847B0AC0CFFC1BE82B905FC8CF99C" box="[658,792,1558,1583]" metricMagnitude="2" metricUnit="m" metricValue="5.4" metricValueMax="6.6" metricValueMin="4.2" pageId="48" pageNumber="49" unit="m" value="540.0" valueMax="660.0" valueMin="420.0">
420–
<specimenCount id="349621DCAC0CFFC1BEDBB905FC8CF99C" box="[715,792,1558,1583]" count="660" pageId="48" pageNumber="49" type="generic">660 m</specimenCount>
</quantity>
.
</paragraph>
<paragraph id="222FEA55AC0CFFC1BC69B959FE0BF93A" blockId="48.[121,1011,1610,1673]" pageId="48" pageNumber="49">
<emphasis id="10E43647AC0CFFC1BC69B959FF79F9D0" box="[121,237,1610,1635]" italics="true" pageId="48" pageNumber="49">Etymology</emphasis>
. This species is named after the mountain on which it was collected (
<bibRefCitation id="460197A4AC0CFFC1BC6FB963FE04F93A" author="Kartonegoro A &amp; Bone RE &amp; Atkins HJ" box="[127,400,1648,1673]" pageId="48" pageNumber="49" pagination="173 - 204" refId="ref50599" refString="Kartonegoro A, Bone RE, Atkins HJ. 2018. Eleven new species of Cyrtandra (Gesneriaceae) from Sulawesi, Indonesia. Edinburgh Journal of Botany. 75 (2): 173 - 204." type="journal article" year="2018">
Kartonegoro
<emphasis id="10E43647AC0CFFC1BD05B963FEDFF93A" box="[277,331,1648,1673]" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
, 2018
</bibRefCitation>
).
</paragraph>
<caption id="76EFBADDAC0DFFC0BC69B91CFE7FF918" ID-DOI="http://doi.org/10.5281/zenodo.10515472" ID-Zenodo-Dep="10515472" httpUri="https://zenodo.org/record/10515472/files/figure.png" pageId="49" pageNumber="50" startId="49.[121,185,1551,1573]" targetBox="[237,1047,268,1530]" targetPageId="49" targetType="figure">
<paragraph id="222FEA55AC0DFFC0BC69B91CFE7FF918" blockId="49.[121,1158,1551,1707]" pageId="49" pageNumber="50">
<emphasis id="10E43647AC0DFFC0BC69B91CFF4DF996" bold="true" box="[121,217,1551,1573]" pageId="49" pageNumber="50">Figure 26</emphasis>
.
<taxonomicName id="E59091D6AC0DFFC0BCF5B91CFE36F996" box="[229,418,1551,1573]" class="Magnoliopsida" family="Gesneriaceae" genus="Cyrtandra" kingdom="Plantae" order="Lamiales" pageId="49" pageNumber="50" phylum="Tracheophyta" rank="species" species="hekensis">
<emphasis id="10E43647AC0DFFC0BCF5B91CFE36F996" box="[229,418,1551,1573]" italics="true" pageId="49" pageNumber="50">Cyrtandra hekensis</emphasis>
</taxonomicName>
. a, Habit; b, opened corolla, showing position of stamens and staminodes; c, opened calyx, showing inner surface; d, side view of corolla; e, gynoecium, showing disc and top of pedicel; f, fruit with persistent calyx partially removed; g, bracts. Drawn by Claire Banks from
<emphasis id="10E43647AC0DFFC0BC6AB960FF40F93A" box="[122,212,1651,1673]" italics="true" pageId="49" pageNumber="50">Hendrian</emphasis>
et al. 901. Reproduced from
<bibRefCitation id="460197A4AC0DFFC0BDE8B960FD61F93A" author="Kartonegoro A &amp; Bone RE &amp; Atkins HJ" box="[504,757,1651,1673]" pageId="49" pageNumber="50" pagination="173 - 204" refId="ref50599" refString="Kartonegoro A, Bone RE, Atkins HJ. 2018. Eleven new species of Cyrtandra (Gesneriaceae) from Sulawesi, Indonesia. Edinburgh Journal of Botany. 75 (2): 173 - 204." type="journal article" year="2018">
Kartonegoro
<emphasis id="10E43647AC0DFFC0BE6CB960FD3AF93A" box="[636,686,1651,1673]" italics="true" pageId="49" pageNumber="50">et al.</emphasis>
(2018)
</bibRefCitation>
with the permission of the Trustees of the Royal Botanic Garden Edinburgh.
</paragraph>
</caption>
<paragraph id="222FEA55AC0EFFC3BC69BE19FE7FFD0E" blockId="50.[121,1160,266,701]" pageId="50" pageNumber="51">
<emphasis id="10E43647AC0EFFC3BC69BE19FDB0FE90" box="[121,548,266,291]" italics="true" pageId="50" pageNumber="51">Proposed IUCN conservation category</emphasis>
. The low number of collections for
<taxonomicName id="E59091D6AC0EFFC3BFABBE19FF48FEFB" class="Magnoliopsida" family="Gesneriaceae" genus="Cyrtandra" kingdom="Plantae" order="Lamiales" pageId="50" pageNumber="51" phylum="Tracheophyta" rank="species" species="hekensis">
<emphasis id="10E43647AC0EFFC3BFABBE19FF48FEFB" italics="true" pageId="50" pageNumber="51">Cyrtandra hekensis</emphasis>
</taxonomicName>
means that it is not possible to calculate an EOO, but it has an AOO of
<quantity id="E56847B0AC0EFFC3B819BE3CFBD6FEFB" box="[1033,1090,303,328]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="50" pageNumber="51" unit="km" value="4.0">4 km</quantity>
<superScript id="D5E5471DAC0EFFC3B852BE3DFBDEFE8E" attach="left" box="[1090,1098,302,317]" fontSize="6" pageId="50" pageNumber="51">2</superScript>
, based on a 2 ×
<quantity id="E56847B0AC0EFFC3BD39BE47FEF5FEDE" box="[297,353,340,365]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="50" pageNumber="51" unit="km" value="2.0">2 km</quantity>
grid cell size, under the B criteria (
<bibRefCitation id="460197A4AC0EFFC3BEFABE47FC49FEDE" author="Bachman S &amp; Moat J &amp; Hill AW &amp; de la Torre J &amp; Scott B." box="[746,989,340,365]" pageId="50" pageNumber="51" pagination="117 - 126" refId="ref49715" refString="Bachman S, Moat J, Hill AW, de la Torre J, Scott B. 2011. Supporting Red List threat assessments with GeoCAT: Geospatial Conservation Assessment Tool. In: Smith V, Penev L, editors. E-Infrastructures for Data Publishing in Biodiversity Science. Zookeys. 150: 117 - 126 (version BETA)." type="journal article" year="2011">
Bachman
<emphasis id="10E43647AC0EFFC3BF71BE47FC03FEDE" box="[865,919,340,365]" italics="true" pageId="50" pageNumber="51">et al.</emphasis>
, 2011
</bibRefCitation>
). The species is known only from Mount Hek, Banggai Regency, where it was collected at relatively low altitude during expeditions in 2004 and 2008. This forest
<typeStatus id="FD2B54F7AC0EFFC3BF25BE8CFCF1FE0B" box="[821,869,415,440]" pageId="50" pageNumber="51">type</typeStatus>
is one of the most threatened in
<collectingRegion id="E05424B7AC0EFFC3BD07BED7FEE9FE6E" box="[279,381,452,477]" country="Indonesia" name="Sulawesi Utara" pageId="50" pageNumber="51">Sulawesi</collectingRegion>
(
<bibRefCitation id="460197A4AC0EFFC3BD9ABED7FDF3FE6E" author="Cannon CH &amp; Summers M &amp; Harting JR &amp; Kessler PJ" box="[394,615,452,477]" pageId="50" pageNumber="51" pagination="747 - 759" refId="ref50009" refString="Cannon CH, Summers M, Harting JR, Kessler PJ. 2007. Developing conservation priorities based on forest type, condition, and threats in a poorly known ecoregion: Sulawesi, Indonesia. Biotropica 39 (6): 747 - 759." type="journal article" year="2007">
Cannon
<emphasis id="10E43647AC0EFFC3BDFBBED7FDB6FE6E" box="[491,546,452,477]" italics="true" pageId="50" pageNumber="51">et al.</emphasis>
, 2007
</bibRefCitation>
) and Mount Hek is in an area with no formal protection (UNEP-WCMC &amp;
<bibRefCitation id="460197A4AC0EFFC3BDA2BEF9FDA6FDB0" author="IUCN" box="[434,562,490,515]" pageId="50" pageNumber="51" refId="ref51358" refString="UNEP-WCMC, IUCN. 2019. Protected Planet: the World Database on Protected Areas (WDPA). Cambridge: UNEP-WCMC and IUCN. https: // www. protectedplanet. net [Accessed 1 November 2019.]" type="url" year="2019">IUCN, 2019</bibRefCitation>
). It is, however, in one of the most under-collected areas of
<collectingRegion id="E05424B7AC0EFFC3BCCFBD1CFED1FD9B" box="[223,325,527,552]" country="Indonesia" name="Sulawesi Utara" pageId="50" pageNumber="51">Sulawesi</collectingRegion>
(
<bibRefCitation id="460197A4AC0EFFC3BD41BD1CFDB8FD9B" author="Kessler PJA &amp; Bos MN &amp; Sierra Daza SEC &amp; Kop A &amp; Willemse LPM &amp; Pitopang R &amp; Gradstein SR" box="[337,556,527,552]" pageId="50" pageNumber="51" pagination="1 - 160" refId="ref50637" refString="Kessler PJA, Bos MN, Sierra Daza SEC, Kop A, Willemse LPM, Pitopang R, Gradstein SR. 2002. Checklist of the woody plants of Sulawesi, Indonesia. Blumea Suppl. 14: 1 - 160" type="journal article" year="2002">
Kessler
<emphasis id="10E43647AC0EFFC3BDBFBD1CFE72FD9B" box="[431,486,527,552]" italics="true" pageId="50" pageNumber="51">et al.</emphasis>
, 2002
</bibRefCitation>
;
<bibRefCitation id="460197A4AC0EFFC3BE28BD1CFC80FD9B" author="Cannon CH &amp; Summers M &amp; Harting JR &amp; Kessler PJ" box="[568,788,527,552]" pageId="50" pageNumber="51" pagination="747 - 759" refId="ref50009" refString="Cannon CH, Summers M, Harting JR, Kessler PJ. 2007. Developing conservation priorities based on forest type, condition, and threats in a poorly known ecoregion: Sulawesi, Indonesia. Biotropica 39 (6): 747 - 759." type="journal article" year="2007">
Cannon
<emphasis id="10E43647AC0EFFC3BE87BD1CFD5AFD9B" box="[663,718,527,552]" italics="true" pageId="50" pageNumber="51">et al.</emphasis>
, 2007
</bibRefCitation>
), and it is possible that there are more populations of this species. To reflect its narrow distribution in a single location in a threatened forest
<typeStatus id="FD2B54F7AC0EFFC3BD65BD49FE31FDC0" box="[373,421,602,627]" pageId="50" pageNumber="51">type</typeStatus>
and lack of formal protection (
<bibRefCitation id="460197A4AC0EFFC3BF16BD49FC70FDC0" author="Cannon CH &amp; Summers M &amp; Harting JR &amp; Kessler PJ" box="[774,996,602,627]" pageId="50" pageNumber="51" pagination="747 - 759" refId="ref50009" refString="Cannon CH, Summers M, Harting JR, Kessler PJ. 2007. Developing conservation priorities based on forest type, condition, and threats in a poorly known ecoregion: Sulawesi, Indonesia. Biotropica 39 (6): 747 - 759." type="journal article" year="2007">
Cannon
<emphasis id="10E43647AC0EFFC3BF78BD49FC0BFDC0" box="[872,927,602,627]" italics="true" pageId="50" pageNumber="51">et al.</emphasis>
, 2007
</bibRefCitation>
),
<taxonomicName id="E59091D6AC0EFFC3BFE9BD49FF48FD2B" class="Magnoliopsida" family="Gesneriaceae" genus="Cyrtandra" kingdom="Plantae" order="Lamiales" pageId="50" pageNumber="51" phylum="Tracheophyta" rank="species" species="hekensis">
<emphasis id="10E43647AC0EFFC3BFE9BD49FF48FD2B" italics="true" pageId="50" pageNumber="51">Cyrtandra hekensis</emphasis>
</taxonomicName>
is assessed as Critically Endangered (CR), B2ab(iii). Habitat protection and
<emphasis id="10E43647AC0EFFC3B82EBD6CFB1CFD2B" box="[1086,1160,639,664]" italics="true" pageId="50" pageNumber="51">ex situ</emphasis>
conservation are recommended.
</paragraph>
<paragraph id="222FEA55AC0EFFC3BC69BDC4FD85FCBD" blockId="50.[121,1114,727,782]" pageId="50" pageNumber="51">
<emphasis id="10E43647AC0EFFC3BC69BDC4FE3DFD5E" box="[121,425,727,749]" italics="true" pageId="50" pageNumber="51">Additional specimen examined</emphasis>
.
<materialsCitation id="92F8E008AC0EFFC3BDA6BDC4FD99FCBD" collectingDate="2004-02-26" collectorName="Hendrian &amp; Newman &amp; Scott &amp; Saleh &amp; Supriadi" country="Indonesia" county="Banggai" location="Mt Hek" pageId="50" pageNumber="51" specimenCount="1" stateProvince="Sulawesi Tengah">
<collectingCountry id="5A87AAC5AC0EFFC3BDA6BDC4FD8FFD5E" box="[438,539,727,749]" name="Indonesia" pageId="50" pageNumber="51">INDONESIA</collectingCountry>
.
<emphasis id="10E43647AC0EFFC3BE37BDC4FD45FD5E" bold="true" box="[551,721,727,749]" pageId="50" pageNumber="51">
<collectingRegion id="E05424B7AC0EFFC3BE37BDC4FD45FD5E" box="[551,721,727,749]" country="Indonesia" name="Sulawesi Tengah" pageId="50" pageNumber="51">Central Sulawesi</collectingRegion>
</emphasis>
.
<emphasis id="10E43647AC0EFFC3BECDBDC4FCBAFD5E" bold="true" box="[733,814,727,749]" italics="true" pageId="50" pageNumber="51">
<collectingCounty id="CB4E92D9AC0EFFC3BECDBDC4FCBAFD5E" box="[733,814,727,749]" pageId="50" pageNumber="51">Banggai</collectingCounty>
</emphasis>
:
<location id="274FBC8EAC0EFFC3BF2ABDC4FC10FD5E" LSID="urn:lsid:plazi:treatment:AA395B43AC0CFFC3BC69BEEAFC97FC00:274FBC8EAC0EFFC3BF2ABDC4FC10FD5E" box="[826,900,727,749]" country="Indonesia" county="Banggai" name="Mt Hek" pageId="50" pageNumber="51" stateProvince="Sulawesi Tengah">Mt Hek</location>
,
<date id="562ECC95AC0EFFC3BF9DBDC4FC67FD5E" box="[909,1011,727,749]" pageId="50" pageNumber="51" value="2004-02-26">
<collectingDate id="466A357DAC0EFFC3BF9DBDC4FC67FD5E" box="[909,1011,727,749]" pageId="50" pageNumber="51" value="2004-02-26">26 ii 2004</collectingDate>
</date>
,
<emphasis id="10E43647AC0EFFC3BFECBDC4FE28FCBD" italics="true" pageId="50" pageNumber="51">
<collectorName id="8F658F83AC0EFFC3BFECBDC4FBC2FD5E" box="[1020,1110,727,749]" pageId="50" pageNumber="51">Hendrian</collectorName>
,
<collectorName id="8F658F83AC0EFFC3BC69BDEBFF46FCBD" box="[121,210,760,782]" pageId="50" pageNumber="51">Newman</collectorName>
,
<collectorName id="8F658F83AC0EFFC3BCCBBDEBFE84FCBD" box="[219,272,760,782]" pageId="50" pageNumber="51">Scott</collectorName>
,
<collectorName id="8F658F83AC0EFFC3BD0ABDEBFEC5FCBD" box="[282,337,760,782]" pageId="50" pageNumber="51">Saleh</collectorName>
&amp;
<collectorName id="8F658F83AC0EFFC3BD7BBDEBFE28FCBD" box="[363,444,760,782]" pageId="50" pageNumber="51">Supriadi</collectorName>
</emphasis>
901 (E)
</materialsCitation>
.
</paragraph>
<paragraph id="222FEA55AC0EFFC3BC69BC39FC97FC00" blockId="50.[121,1151,810,947]" pageId="50" pageNumber="51">
<taxonomicName id="E59091D6AC0EFFC3BC69BC39FED9FCF0" box="[121,333,810,835]" class="Magnoliopsida" family="Gesneriaceae" genus="Cyrtandra" kingdom="Plantae" order="Lamiales" pageId="50" pageNumber="51" phylum="Tracheophyta" rank="species" species="hekensis">
<emphasis id="10E43647AC0EFFC3BC69BC39FED9FCF0" box="[121,333,810,835]" italics="true" pageId="50" pageNumber="51">Cyrtandra hekensis</emphasis>
</taxonomicName>
is most similar to the widespread
<taxonomicName id="E59091D6AC0EFFC3BEC3BC39FCC9FCF0" box="[723,861,810,835]" class="Magnoliopsida" family="Gesneriaceae" genus="Cyrtandra" kingdom="Plantae" order="Lamiales" pageId="50" pageNumber="51" phylum="Tracheophyta" rank="species" species="polyneura">
<emphasis id="10E43647AC0EFFC3BEC3BC39FCC9FCF0" box="[723,861,810,835]" italics="true" pageId="50" pageNumber="51">C. polyneura</emphasis>
</taxonomicName>
in its subequal, decurrent leaves, bracteate inflorescence, and white flowers with greenish yellow markings in the throat but can be distinguished by its much smaller, narrower leaves, fewer flowers per inflorescence, and coarser indumentum on young growth.
</paragraph>
</subSubSection>
</treatment>
</document>
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