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<document id="ADBA27A158D36F35CB06BD5FDB8D197A" ID-CLB-Dataset="25508" ID-DOI="10.11646/zootaxa.4671.3.2" ID-GBIF-Dataset="43f393b8-6c05-4861-9b7c-2f7f1a169d10" ID-ISSN="1175-5326" ID-Zenodo-Dep="3442512" ID-ZooBank="99459D5F-3C35-4F7D-9768-D70616676851" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1568800771299" checkinUser="plazi" docAuthor="Watson, Charlotte, Tilic, Ekin &amp; Rouse, Greg W." docDate="2019" docId="771987EF77733A2D2AFEC1D1FEDEDA15" docLanguage="en" docName="zootaxa.4671.3.2.pdf" docOrigin="Zootaxa 4671 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hyalopale angeliensis Watson &amp; Tilic &amp; Rouse 2019, sp. nov." docType="treatment" docUuid="9106A06A-97A8-4D5D-A208-0F9C91152008" docUuidSource="ZooBank" docVersion="10" lastPageNumber="356" masterDocId="8B20FF9777633A3C2A69C53DFFF9D83B" masterDocTitle="Revision of Hyalopale (Chrysopetalidae; Phyllodocida; Annelida): an amphi-Atlantic Hyalopale bispinosa species complex and five new species from reefs of the Caribbean Sea and Indo-Pacific Oceans" masterLastPageNumber="368" masterPageNumber="339" pageNumber="355" updateTime="1698755304942" updateUser="ExternalLinkService">
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<mods:title id="376F617DD0F7E2E73F6F059DD271C133">Revision of Hyalopale (Chrysopetalidae; Phyllodocida; Annelida): an amphi-Atlantic Hyalopale bispinosa species complex and five new species from reefs of the Caribbean Sea and Indo-Pacific Oceans</mods:title>
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<mods:namePart id="44EF8EE33C00919EF046F554A4E4BF36">Watson, Charlotte</mods:namePart>
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<mods:namePart id="A76CC690ACA8674CD726CE5FE7C7A7C5">Tilic, Ekin</mods:namePart>
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<mods:namePart id="22BAC7305DCDB350BF978E11657DA68F">Rouse, Greg W.</mods:namePart>
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<treatment id="771987EF77733A2D2AFEC1D1FEDEDA15" ID-GBIF-Taxon="159608542" LSID="urn:lsid:zoobank.org:act:9106A06A-97A8-4D5D-A208-0F9C91152008" httpUri="http://treatment.plazi.org/id/771987EF77733A2D2AFEC1D1FEDEDA15" lastPageId="17" lastPageNumber="356" pageId="16" pageNumber="355">
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<paragraph id="FF0F36F977733A2C2AFEC1D1FDFEDD3C" blockId="16.[151,960,1260,1359]" box="[151,519,1260,1287]" pageId="16" pageNumber="355">
<heading id="A447819577733A2C2AFEC1D1FDFEDD3C" bold="true" box="[151,519,1260,1287]" fontSize="11" level="1" pageId="16" pageNumber="355" reason="1">
<emphasis id="CDC4EAEB77733A2C2AFEC1D1FDFEDD3C" bold="true" box="[151,519,1260,1287]" pageId="16" pageNumber="355">
<taxonomicName id="38B04D7A77733A2C2AFEC1D1FE58DD3C" ID-CoL="85RF9" authority="Watson &amp; Tilic &amp; Rouse, 2019" authorityName="Watson &amp; Tilic &amp; Rouse" authorityYear="2019" box="[151,417,1260,1287]" class="Polychaeta" family="Chrysopetalidae" genus="Hyalopale" kingdom="Animalia" order="Phyllodocida" pageId="16" pageNumber="355" phylum="Annelida" rank="species" species="angeliensis" status="sp. nov.">
<emphasis id="CDC4EAEB77733A2C2AFEC1D1FE58DD3C" bold="true" box="[151,417,1260,1287]" italics="true" pageId="16" pageNumber="355">Hyalopale angeliensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D6F7579077733A2C2BC0C1D0FDFEDD3C" box="[425,519,1261,1287]" pageId="16" pageNumber="355" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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urn:lsid:zoobank.org:act:
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<subSubSection id="B7AA657277733A2C2AFEC008FEB8DD74" box="[151,321,1333,1359]" pageId="16" pageNumber="355" type="description">
<paragraph id="FF0F36F977733A2C2AFEC008FEB8DD74" blockId="16.[151,960,1260,1359]" box="[151,321,1333,1359]" pageId="16" pageNumber="355">
<figureCitation id="678B2A7C77733A2C2AFEC008FF09DD74" box="[151,240,1333,1359]" captionStart="FIGURE 1" captionStartId="6.[151,250,1860,1885]" captionTargetBox="[222,1365,166,1840]" captionTargetId="figure@6.[222,1365,166,1840]" captionTargetPageId="6" captionText="FIGURE 1. Composite plate comparing notochaetal main paleae in Hyalopale species. A. Hyalopale bispinosa s.s. Florida, W. Atlantic; B. H. cf. bispinosa Crete, Mediterranean Sea; C. H. leslieae sp. nov. Florida Keys, W. Atlantic; D. H. furfuricula sp. nov. Mozambique, W. Indian Ocean; E. H. angeliensis sp. nov. Western Australia, E. Indian Ocean; F. H. zerofskii sp. nov. California, E. Pacific; G. H. sapphiriglancyorum sp. nov. E. Indonesia, W. Pacific. Note: correct number of raised and internal ribs figured." figureDoi="http://doi.org/10.5281/zenodo.3442516" httpUri="https://zenodo.org/record/3442516/files/figure.png" pageId="16" pageNumber="355">Figs 1E</figureCitation>
;
<figureCitation id="678B2A7C77733A2C2A95C008FEF0DD74" box="[252,265,1333,1359]" captionStart="FIGURE 9" captionStartId="17.[151,250,1930,1955]" captionTargetBox="[173,1412,579,1895]" captionTargetId="figure@17.[160,1427,572,1911]" captionTargetPageId="17" captionText="FIGURE 9. H. angeliensis sp. nov. A,C. Dampier, Western Australia, Holotype, AM W.36166 A. detail middle group palea, C. detail of midline-most main paleae; B,D. Seychelles, USNM 1076948. B. detail middle group palea, D. neurochaetal falcigerous types from mid-body segment, left to right: superior very long bladed, mid-superior long bladed, mid-group broader bladed with larger serrations, inferior group short bladed." figureDoi="http://doi.org/10.5281/zenodo.3442532" httpUri="https://zenodo.org/record/3442532/files/figure.png" pageId="16" pageNumber="355">9</figureCitation>
AD
</paragraph>
</subSubSection>
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<paragraph id="FF0F36F977733A2C2AFEC041FAD2DD80" blockId="16.[151,1437,1404,2043]" pageId="16" pageNumber="355">
<emphasis id="CDC4EAEB77733A2C2AFEC041FE79DDAC" bold="true" box="[151,384,1404,1431]" pageId="16" pageNumber="355">Material examined.</emphasis>
<materialsCitation id="4FD83CA477733A2C2BEFC040FAD2DD80" ID-GBIF-Occurrence="2413161059" collectingDate="2000-08-01" collectionCode="AM" collectorName="P. A. Hutchings" county="Eastern Indian Ocean" latitude="-20.8" location="Angel Island" municipality="Dampier" pageId="16" pageNumber="355" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<emphasis id="CDC4EAEB77733A2C2BEFC040FE0ADDAC" bold="true" box="[390,499,1405,1431]" pageId="16" pageNumber="355">
<typeStatus id="200B885B77733A2C2BEFC040FE0ADDAC" box="[390,499,1405,1431]" pageId="16" pageNumber="355" type="holotype">Holotype</typeStatus>
</emphasis>
.
<collectionCode id="99A1AE3C77733A2C2B96C040FDD5DDAC" box="[511,556,1405,1431]" country="Australia" name="Australian Museum" pageId="16" pageNumber="355" type="Museum">AM</collectionCode>
W.3166,
<collectingCounty id="166E4E7577733A2C28FFC040FC71DDAC" box="[662,904,1405,1431]" pageId="16" pageNumber="355">Eastern Indian Ocean</collectingCounty>
,
<collectingRegion id="3D74F81B77733A2C29FAC040FBA4DDAC" box="[915,1117,1405,1431]" country="Australia" name="Western Australia" pageId="16" pageNumber="355">Western Australia</collectingRegion>
,
<collectingMunicipality id="1F6BAC8377733A2C2E0EC040FB32DDAC" box="[1127,1227,1405,1431]" pageId="16" pageNumber="355">Dampier</collectingMunicipality>
,
<location id="FA6F602277733A2C2EBAC040FA9CDDAC" LSID="urn:lsid:plazi:treatment:771987EF77733A2D2AFEC1D1FEDEDA15:FA6F602277733A2C2EBAC040FA9CDDAC" box="[1235,1381,1405,1431]" county="Eastern Indian Ocean" latitude="-20.8" municipality="Dampier" name="Angel Island" pageId="16" pageNumber="355" stateProvince="Western Australia">Angel Island</location>
, Stn. WA639,
<geoCoordinate id="9A84503E77733A2C2A92C09CFE9DDD80" box="[251,356,1441,1467]" degrees="20" direction="south" minutes="48" orientation="latitude" pageId="16" pageNumber="355" precision="925" value="-20.8">20° 48 S</geoCoordinate>
,
<geoCoordinate id="9A84503E77733A2C2B18C09DFE10DD80" box="[369,489,1440,1467]" pageId="16" pageNumber="355">116° 81 E</geoCoordinate>
,
<date id="8B0E103977733A2C2B9CC09CFD82DD80" box="[501,635,1441,1467]" pageId="16" pageNumber="355" value="2000-08-01">
<collectingDate id="9B4AE9D177733A2C2B9CC09CFD82DD80" box="[501,635,1441,1467]" pageId="16" pageNumber="355" value="2000-08-01">1 Aug 2000</collectingDate>
</date>
, coll.
<collectorName id="5245532F77733A2C28D7C09CFC96DD80" box="[702,879,1441,1467]" pageId="16" pageNumber="355">P. A. Hutchings</collectorName>
, L. Avery, 13E, L:
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, W:
<quantity id="38489B1C77733A2C2EBCC09CFADEDD80" box="[1237,1319,1441,1467]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="16" pageNumber="355" unit="mm" value="0.4">0.4mm</quantity>
.
</materialsCitation>
</paragraph>
<paragraph id="FF0F36F977733A2C2AAEC0F9FB5DDE38" blockId="16.[151,1437,1404,2043]" pageId="16" pageNumber="355">
<emphasis id="CDC4EAEB77733A2C2AAEC0F9FE43DDE4" bold="true" box="[199,442,1476,1503]" pageId="16" pageNumber="355">Additional material.</emphasis>
<materialsCitation id="4FD83CA477733A2C2BA8C0F8FB58DE38" ID-GBIF-Occurrence="2413161052" collectingDate="1975-12" collectionCode="USNM" collectorName="K. Buchanan" country="Seychelles" latitude="-19.75" location="Western Indian Ocean" longLatPrecision="1062" longitude="1.13" municipality="Round Island" pageId="16" pageNumber="355" specimenCode="USNM 1076948" specimenCount="1">
<specimenCode id="AF169E8277733A2C2BA8C0F8FD87DDE5" box="[449,638,1476,1503]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="355" type="Museum">USNM 1076948</specimenCode>
,
<location id="FA6F602277733A2C28E0C0F8FC7DDDE4" LSID="urn:lsid:plazi:treatment:771987EF77733A2D2AFEC1D1FEDEDA15:FA6F602277733A2C28E0C0F8FC7DDDE4" box="[649,900,1477,1503]" country="Seychelles" latitude="-19.75" longLatPrecision="1062" longitude="1.13" municipality="Round Island" name="Western Indian Ocean" pageId="16" pageNumber="355">Western Indian Ocean</location>
,
<collectingCountry id="87A7766977733A2C29F9C0F8FBF2DDE4" box="[912,1035,1477,1503]" name="Seychelles" pageId="16" pageNumber="355">Seychelles</collectingCountry>
,
<collectingMunicipality id="1F6BAC8377733A2C2E7CC0F8FB57DDE4" box="[1045,1198,1477,1503]" pageId="16" pageNumber="355">Round Island</collectingMunicipality>
,
<geoCoordinate id="9A84503E77733A2C2ED3C0F8FADDDDE4" box="[1210,1316,1476,1503]" degrees="19" direction="south" minutes="45" orientation="latitude" pageId="16" pageNumber="355" precision="925" value="-19.75">19° 45 S</geoCoordinate>
,
<geoCoordinate id="9A84503E77733A2C2F59C0F9FA61DDE4" box="[1328,1432,1476,1503]" degrees="57" direction="east" minutes="50" orientation="longitude" pageId="16" pageNumber="355" precision="925" value="57.833332">57° 50E</geoCoordinate>
, Sta. 14, intertidal from sponge, coll.
<collectorName id="5245532F77733A2C285BC0D4FD3EDE38" box="[562,711,1513,1539]" pageId="16" pageNumber="355">K. Buchanan</collectorName>
,
<date id="8B0E103977733A2C28BBC0D4FCB8DE38" box="[722,833,1512,1539]" pageId="16" pageNumber="355" value="1975-12">
<collectingDate id="9B4AE9D177733A2C28BBC0D4FCB8DE38" box="[722,833,1512,1539]" pageId="16" pageNumber="355" value="1975-12">Dec 1975</collectingDate>
</date>
, 1, 13E, L:
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, W:
<quantity id="38489B1C77733A2C2E56C0D5FB58DE38" box="[1087,1185,1512,1539]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="16" pageNumber="355" unit="mm" value="0.55">0.55mm</quantity>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B7AA657277733A2C2AAEC331FA7ADE1C" box="[199,1411,1548,1575]" pageId="16" pageNumber="355" type="diagnosis">
<paragraph id="FF0F36F977733A2C2AAEC331FA7ADE1C" blockId="16.[151,1437,1404,2043]" box="[199,1411,1548,1575]" pageId="16" pageNumber="355">
<emphasis id="CDC4EAEB77733A2C2AAEC331FEBBDE1C" bold="true" box="[199,322,1548,1575]" pageId="16" pageNumber="355">Diagnosis.</emphasis>
Mid-body main paleae narrow with well-defined apices, (16)1822 ribs, 46 shallow raised ribs.
</paragraph>
</subSubSection>
<subSubSection id="B7AA657277733A2C2AAEC30DFB49DF8F" pageId="16" pageNumber="355" type="description">
<paragraph id="FF0F36F977733A2C2AAEC30DFCE6DE77" blockId="16.[151,1437,1404,2043]" box="[199,799,1584,1612]" pageId="16" pageNumber="355">
<emphasis id="CDC4EAEB77733A2C2AAEC30DFE9BDE70" bold="true" box="[199,354,1584,1611]" pageId="16" pageNumber="355">Description (</emphasis>
based on
<typeStatus id="200B885B77733A2C2BA5C30CFDD7DE70" box="[460,558,1585,1611]" pageId="16" pageNumber="355" type="holotype">holotype</typeStatus>
except where noted).
</paragraph>
<paragraph id="FF0F36F977733A2C2AAEC368FB49DF8F" blockId="16.[151,1437,1404,2043]" pageId="16" pageNumber="355">
Slightly wasted entire individual; body pale yellow, three pairs of melded dark red eyes; cirri intact, some neurochaetal
<typeStatus id="200B885B77733A2C2B6FC344FEB8DEA8" box="[262,321,1657,1683]" pageId="16" pageNumber="355">types</typeStatus>
loosening in posterior body. Segment II with 4 short, curved notochaetal spines; segment III with lateral spine, 8 main paleae. Mid-body notochaetal fan with single lateral spine, 10 main paleae with 1822 internal ribs, 46 patchy shallow raised ribs; midline spines absent. Main paleae slender with sloping brow, small blunt apices, inner margin may be slightly folded with minute serrations, convex margin with widely spaced serrations mid-way, becoming finer on brow leading to apex (
<figureCitation id="678B2A7C77733A2C28BCC234FCD7DF18" box="[725,814,1801,1827]" captionStart="FIGURE 9" captionStartId="17.[151,250,1930,1955]" captionTargetBox="[173,1412,579,1895]" captionTargetId="figure@17.[160,1427,572,1911]" captionTargetPageId="17" captionText="FIGURE 9. H. angeliensis sp. nov. A,C. Dampier, Western Australia, Holotype, AM W.36166 A. detail middle group palea, C. detail of midline-most main paleae; B,D. Seychelles, USNM 1076948. B. detail middle group palea, D. neurochaetal falcigerous types from mid-body segment, left to right: superior very long bladed, mid-superior long bladed, mid-group broader bladed with larger serrations, inferior group short bladed." figureDoi="http://doi.org/10.5281/zenodo.3442532" httpUri="https://zenodo.org/record/3442532/files/figure.png" pageId="16" pageNumber="355">Fig. 9A</figureCitation>
, Western Australia;
<figureCitation id="678B2A7C77733A2C2E67C234FB9DDF18" box="[1038,1124,1801,1827]" captionStart="FIGURE 9" captionStartId="17.[151,250,1930,1955]" captionTargetBox="[173,1412,579,1895]" captionTargetId="figure@17.[160,1427,572,1911]" captionTargetPageId="17" captionText="FIGURE 9. H. angeliensis sp. nov. A,C. Dampier, Western Australia, Holotype, AM W.36166 A. detail middle group palea, C. detail of midline-most main paleae; B,D. Seychelles, USNM 1076948. B. detail middle group palea, D. neurochaetal falcigerous types from mid-body segment, left to right: superior very long bladed, mid-superior long bladed, mid-group broader bladed with larger serrations, inferior group short bladed." figureDoi="http://doi.org/10.5281/zenodo.3442532" httpUri="https://zenodo.org/record/3442532/files/figure.png" pageId="16" pageNumber="355">Fig. 9B</figureCitation>
,
<collectingCountry id="87A7766977733A2C2E19C234FB14DF1F" box="[1136,1261,1801,1828]" name="Seychelles" pageId="16" pageNumber="355">Seychelles</collectingCountry>
). Narrower, lateral-most main with 1617 ribs and 23 smaller, more symmetrical-shaped, midline-most main with lower number of ribs (
<figureCitation id="678B2A7C77733A2C2A84C26CFEBADF50" box="[237,323,1873,1899]" captionStart="FIGURE 9" captionStartId="17.[151,250,1930,1955]" captionTargetBox="[173,1412,579,1895]" captionTargetId="figure@17.[160,1427,572,1911]" captionTargetPageId="17" captionText="FIGURE 9. H. angeliensis sp. nov. A,C. Dampier, Western Australia, Holotype, AM W.36166 A. detail middle group palea, C. detail of midline-most main paleae; B,D. Seychelles, USNM 1076948. B. detail middle group palea, D. neurochaetal falcigerous types from mid-body segment, left to right: superior very long bladed, mid-superior long bladed, mid-group broader bladed with larger serrations, inferior group short bladed." figureDoi="http://doi.org/10.5281/zenodo.3442532" httpUri="https://zenodo.org/record/3442532/files/figure.png" pageId="16" pageNumber="355">Fig. 9C</figureCitation>
, see 1611 ribs). Neurochaetae mid-body comprise: four superior group very long-bladed; three mid superior group long-bladed; 8 mid-group with slightly shorter-blades, basal serration; 45 inferior group slightly shorter bladed. Mid-body neuropodia with broader mid-group falcigers (
<figureCitation id="678B2A7C77733A2C29AAC2A4FBE2DF88" box="[963,1051,1945,1971]" captionStart="FIGURE 9" captionStartId="17.[151,250,1930,1955]" captionTargetBox="[173,1412,579,1895]" captionTargetId="figure@17.[160,1427,572,1911]" captionTargetPageId="17" captionText="FIGURE 9. H. angeliensis sp. nov. A,C. Dampier, Western Australia, Holotype, AM W.36166 A. detail middle group palea, C. detail of midline-most main paleae; B,D. Seychelles, USNM 1076948. B. detail middle group palea, D. neurochaetal falcigerous types from mid-body segment, left to right: superior very long bladed, mid-superior long bladed, mid-group broader bladed with larger serrations, inferior group short bladed." figureDoi="http://doi.org/10.5281/zenodo.3442532" httpUri="https://zenodo.org/record/3442532/files/figure.png" pageId="16" pageNumber="355">Fig. 9D</figureCitation>
,
<collectingCountry id="87A7766977733A2C2E4EC2A4FB5DDF8F" box="[1063,1188,1945,1972]" name="Seychelles" pageId="16" pageNumber="355">Seychelles</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="B7AA657277733A2D2AAEC280FC80D9D3" lastPageId="17" lastPageNumber="356" pageId="16" pageNumber="355" type="discussion">
<paragraph id="FF0F36F977733A2D2AAEC280FC80D9D3" blockId="16.[151,1437,1404,2043]" lastBlockId="17.[151,1437,153,558]" lastPageId="17" lastPageNumber="356" pageId="16" pageNumber="355">
<emphasis id="CDC4EAEB77733A2C2AAEC280FEC2DFEC" bold="true" box="[199,315,1981,2007]" pageId="16" pageNumber="355">Remarks.</emphasis>
The two specimens of
<taxonomicName id="38B04D7A77733A2C282AC280FCC7DFEC" authority="Watson &amp; Tilic &amp; Rouse, 2019" authorityName="Watson &amp; Tilic &amp; Rouse" authorityYear="2019" box="[579,830,1981,2007]" class="Polychaeta" family="Chrysopetalidae" genus="Hyalopale" kingdom="Animalia" order="Phyllodocida" pageId="16" pageNumber="355" phylum="Annelida" rank="species" species="angeliensis" status="sp. nov.">
<emphasis id="CDC4EAEB77733A2C282AC280FCC7DFEC" box="[579,830,1981,2007]" italics="true" pageId="16" pageNumber="355">Hyalopale angeliensis</emphasis>
</taxonomicName>
<emphasis id="CDC4EAEB77733A2C292FC280FC59DFEC" bold="true" box="[838,928,1981,2007]" pageId="16" pageNumber="355">
<taxonomicNameLabel id="D6F7579077733A2C292FC280FC59DFEC" box="[838,928,1981,2007]" pageId="16" pageNumber="355" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are found at localities occurring on a similar parallel at ~
<geoCoordinate id="9A84503E77733A2C2B43C2DCFE93DFC0" box="[298,362,2017,2043]" degrees="20" direction="south" orientation="latitude" pageId="16" pageNumber="355" precision="55555" value="-20.0">20° S</geoCoordinate>
in the Eastern and mid-Indian Ocean.
<taxonomicName id="38B04D7A77733A2C2943C2DCFBDFDFC0" authority="Watson &amp; Tilic &amp; Rouse, 2019" authorityName="Watson &amp; Tilic &amp; Rouse" authorityYear="2019" box="[810,1062,2017,2043]" class="Polychaeta" family="Chrysopetalidae" genus="Hyalopale" kingdom="Animalia" order="Phyllodocida" pageId="16" pageNumber="355" phylum="Annelida" rank="species" species="angeliensis" status="sp. nov.">
<emphasis id="CDC4EAEB77733A2C2943C2DCFBDFDFC0" box="[810,1062,2017,2043]" italics="true" pageId="16" pageNumber="355">Hyalopale angeliensis</emphasis>
</taxonomicName>
<emphasis id="CDC4EAEB77733A2C2E46C2DCFB73DFC0" bold="true" box="[1071,1162,2017,2043]" pageId="16" pageNumber="355">
<taxonomicNameLabel id="D6F7579077733A2C2E46C2DCFB73DFC0" box="[1071,1162,2017,2043]" pageId="16" pageNumber="355" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is a small species, pos- sessing a relatively low number of paleal ribs; material examined from north-western Western Australia and the
<collectingCountry id="87A7766977723A2D2AFEC586FEF6D8EE" box="[151,271,187,213]" name="Seychelles" pageId="17" pageNumber="356">Seychelles</collectingCountry>
Islands exhibit very similar, relatively narrow main paleae shape (respectively
<figureCitation id="678B2A7C77723A2D2EE7C586FB06D8EE" box="[1166,1279,187,213]" captionStart="FIGURE 9" captionStartId="17.[151,250,1930,1955]" captionTargetBox="[173,1412,579,1895]" captionTargetId="figure@17.[160,1427,572,1911]" captionTargetPageId="17" captionText="FIGURE 9. H. angeliensis sp. nov. A,C. Dampier, Western Australia, Holotype, AM W.36166 A. detail middle group palea, C. detail of midline-most main paleae; B,D. Seychelles, USNM 1076948. B. detail middle group palea, D. neurochaetal falcigerous types from mid-body segment, left to right: superior very long bladed, mid-superior long bladed, mid-group broader bladed with larger serrations, inferior group short bladed." figureDoi="http://doi.org/10.5281/zenodo.3442532" httpUri="https://zenodo.org/record/3442532/files/figure.png" pageId="17" pageNumber="356">Fig. 9A,B</figureCitation>
) with slightly upswept apices, 1822 ribs and 46 shallow raised ribs. Both individuals also consistently possess 23 smaller, midline-most main paleae (
<figureCitation id="678B2A7C77723A2D2BA4C43DFDDFD921" box="[461,550,256,282]" captionStart="FIGURE 9" captionStartId="17.[151,250,1930,1955]" captionTargetBox="[173,1412,579,1895]" captionTargetId="figure@17.[160,1427,572,1911]" captionTargetPageId="17" captionText="FIGURE 9. H. angeliensis sp. nov. A,C. Dampier, Western Australia, Holotype, AM W.36166 A. detail middle group palea, C. detail of midline-most main paleae; B,D. Seychelles, USNM 1076948. B. detail middle group palea, D. neurochaetal falcigerous types from mid-body segment, left to right: superior very long bladed, mid-superior long bladed, mid-group broader bladed with larger serrations, inferior group short bladed." figureDoi="http://doi.org/10.5281/zenodo.3442532" httpUri="https://zenodo.org/record/3442532/files/figure.png" pageId="17" pageNumber="356">Fig. 9C</figureCitation>
); other
<taxonomicName id="38B04D7A77723A2D2817C43DFD0AD921" authorityName="Perkins" authorityYear="1985" box="[638,755,256,282]" class="Polychaeta" family="Chrysopetalidae" genus="Hyalopale" kingdom="Animalia" order="Phyllodocida" pageId="17" pageNumber="356" phylum="Annelida" rank="genus">
<emphasis id="CDC4EAEB77723A2D2817C43DFD0AD921" box="[638,755,256,282]" italics="true" pageId="17" pageNumber="356">Hyalopale</emphasis>
</taxonomicName>
species may only have 12 (rarely 3) smaller, midline-most main paleae.
<taxonomicName id="38B04D7A77723A2D2B47C41FFDD0D907" authority="Watson &amp; Tilic &amp; Rouse, 2019" authorityName="Watson &amp; Tilic &amp; Rouse" authorityYear="2019" box="[302,553,290,316]" class="Polychaeta" family="Chrysopetalidae" genus="Hyalopale" kingdom="Animalia" order="Phyllodocida" pageId="17" pageNumber="356" phylum="Annelida" rank="species" species="angeliensis" status="sp. nov.">
<emphasis id="CDC4EAEB77723A2D2B47C41FFDD0D907" box="[302,553,290,316]" italics="true" pageId="17" pageNumber="356">Hyalopale angeliensis</emphasis>
</taxonomicName>
<emphasis id="CDC4EAEB77723A2D2858C41EFD75D906" bold="true" box="[561,652,291,317]" pageId="17" pageNumber="356">
<taxonomicNameLabel id="D6F7579077723A2D2858C41EFD75D906" box="[561,652,291,317]" pageId="17" pageNumber="356" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="38B04D7A77723A2D28A2C41EFC2BD907" authority="Watson &amp; Tilic &amp; Rouse, 2019" authorityName="Watson &amp; Tilic &amp; Rouse" authorityYear="2019" box="[715,978,290,316]" class="Polychaeta" family="Chrysopetalidae" genus="Hyalopale" kingdom="Animalia" order="Phyllodocida" pageId="17" pageNumber="356" phylum="Annelida" rank="species" species="sapphiriglancyorum" status="sp. nov.">
<emphasis id="CDC4EAEB77723A2D28A2C41EFC2BD907" box="[715,978,290,316]" italics="true" pageId="17" pageNumber="356">H. sapphiriglancyorum</emphasis>
</taxonomicName>
<emphasis id="CDC4EAEB77723A2D29B3C41EFBCDD906" bold="true" box="[986,1076,291,317]" pageId="17" pageNumber="356">
<taxonomicNameLabel id="D6F7579077723A2D29B3C41EFBCDD906" box="[986,1076,291,317]" pageId="17" pageNumber="356" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
have a general similarity in the narrow- shaped main paleae (cf
<figureCitation id="678B2A7C77723A2D286EC478FD73D964" box="[519,650,325,351]" captionStart="FIGURE 1" captionStartId="6.[151,250,1860,1885]" captionTargetBox="[222,1365,166,1840]" captionTargetId="figure@6.[222,1365,166,1840]" captionTargetPageId="6" captionText="FIGURE 1. Composite plate comparing notochaetal main paleae in Hyalopale species. A. Hyalopale bispinosa s.s. Florida, W. Atlantic; B. H. cf. bispinosa Crete, Mediterranean Sea; C. H. leslieae sp. nov. Florida Keys, W. Atlantic; D. H. furfuricula sp. nov. Mozambique, W. Indian Ocean; E. H. angeliensis sp. nov. Western Australia, E. Indian Ocean; F. H. zerofskii sp. nov. California, E. Pacific; G. H. sapphiriglancyorum sp. nov. E. Indonesia, W. Pacific. Note: correct number of raised and internal ribs figured." figureDoi="http://doi.org/10.5281/zenodo.3442516" httpUri="https://zenodo.org/record/3442516/files/figure.png" pageId="17" pageNumber="356">Fig. 1 E, G</figureCitation>
), but the former has more distinct upswept apices, a slightly higher range of rib numbers and absence of midline spines, compared to the latter. The absence of midline spines seen in
<taxonomicName id="38B04D7A77723A2D2AFEC4B7FE6BD99F" authority="Watson &amp; Tilic &amp; Rouse, 2019" authorityName="Watson &amp; Tilic &amp; Rouse" authorityYear="2019" box="[151,402,394,420]" class="Polychaeta" family="Chrysopetalidae" genus="Hyalopale" kingdom="Animalia" order="Phyllodocida" pageId="17" pageNumber="356" phylum="Annelida" rank="species" species="angeliensis" status="sp. nov.">
<emphasis id="CDC4EAEB77723A2D2AFEC4B7FE6BD99F" box="[151,402,394,420]" italics="true" pageId="17" pageNumber="356">Hyalopale angeliensis</emphasis>
</taxonomicName>
<emphasis id="CDC4EAEB77723A2D2BF0C4B7FE0DD99F" bold="true" box="[409,500,394,420]" pageId="17" pageNumber="356">
<taxonomicNameLabel id="D6F7579077723A2D2BF0C4B7FE0DD99F" box="[409,500,394,420]" pageId="17" pageNumber="356" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Eastern Indian Ocean) is also observed in
<taxonomicName id="38B04D7A77723A2D298BC4B6FB87D99F" authority="Watson &amp; Tilic &amp; Rouse, 2019" authorityName="Watson &amp; Tilic &amp; Rouse" authorityYear="2019" box="[994,1150,394,420]" class="Polychaeta" family="Chrysopetalidae" genus="Hyalopale" kingdom="Animalia" order="Phyllodocida" pageId="17" pageNumber="356" phylum="Annelida" rank="species" species="furfuricula" status="sp. nov.">
<emphasis id="CDC4EAEB77723A2D298BC4B6FB87D99F" box="[994,1150,394,420]" italics="true" pageId="17" pageNumber="356">H. furfuricula</emphasis>
</taxonomicName>
<emphasis id="CDC4EAEB77723A2D2EEFC4B7FB18D99F" bold="true" box="[1158,1249,394,420]" pageId="17" pageNumber="356">
<taxonomicNameLabel id="D6F7579077723A2D2EEFC4B7FB18D99F" box="[1158,1249,394,420]" pageId="17" pageNumber="356" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Western Indian Ocean) but both species possess very different main paleae shape and raised rib patterns (cf
<figureCitation id="678B2A7C77723A2D2ECCC491FADFD9FC" box="[1189,1318,428,455]" captionStart="FIGURE 1" captionStartId="6.[151,250,1860,1885]" captionTargetBox="[222,1365,166,1840]" captionTargetId="figure@6.[222,1365,166,1840]" captionTargetPageId="6" captionText="FIGURE 1. Composite plate comparing notochaetal main paleae in Hyalopale species. A. Hyalopale bispinosa s.s. Florida, W. Atlantic; B. H. cf. bispinosa Crete, Mediterranean Sea; C. H. leslieae sp. nov. Florida Keys, W. Atlantic; D. H. furfuricula sp. nov. Mozambique, W. Indian Ocean; E. H. angeliensis sp. nov. Western Australia, E. Indian Ocean; F. H. zerofskii sp. nov. California, E. Pacific; G. H. sapphiriglancyorum sp. nov. E. Indonesia, W. Pacific. Note: correct number of raised and internal ribs figured." figureDoi="http://doi.org/10.5281/zenodo.3442516" httpUri="https://zenodo.org/record/3442516/files/figure.png" pageId="17" pageNumber="356">Fig. 1 E, D</figureCitation>
). Unfortunately, no DNA sequences could be acquired for this new species.
</paragraph>
</subSubSection>
<subSubSection id="B7AA657277723A2D2AAEC4CCFEDEDA15" pageId="17" pageNumber="356" type="etymology">
<paragraph id="FF0F36F977723A2D2AAEC4CCFEDEDA15" blockId="17.[151,1437,153,558]" pageId="17" pageNumber="356">
<emphasis id="CDC4EAEB77723A2D2AAEC4CCFEB7DA30" bold="true" box="[199,334,497,523]" pageId="17" pageNumber="356">Etymology.</emphasis>
The species name is named after the remote Angel Island off Dampier on the northwestern Australian coast.
</paragraph>
</subSubSection>
</treatment>
</document>