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<document id="C9B3579A78F650D2F65B74598FD892A3" ID-DOI="10.5281/zenodo.5169515" ID-ISSN="1942-1354" ID-Zenodo-Dep="5169515" ID-ZooBank="7F2A2366-B4E4-4F37-A5A5-45CB51D4D859" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628277797618" checkinUser="felipe" docAuthor="Rifkind, Jacques" docDate="2017" docId="03F1E067CC01FFF110CFFDFAFD43FE81" docLanguage="en" docName="InsectaMundi.0591.1-18.pdf" docOrigin="Insecta Mundi 2017 (591)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Quadrophenia mooni , Rifkind 2017, new species" docType="treatment" docVersion="3" lastPageNumber="7" masterDocId="FFC8981FCC06FFF9100FFF9EFF94FFDB" masterDocTitle="New genera and species of mimetic Cleridae from Mexico and Central America (Coleoptera: Cleroidea)" masterLastPageNumber="18" masterPageNumber="1" pageNumber="6" updateTime="1699039302695" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="5796DD05E86B38DDD2E4FD4D0645092B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="64700F171D2967A0F4DBD1BE9367004F">
<mods:title id="6370A56BA4CDF728065E410A115F505D">New genera and species of mimetic Cleridae from Mexico and Central America (Coleoptera: Cleroidea)</mods:title>
</mods:titleInfo>
<mods:name id="9A50E274F266CF9ED6284D220C9A067D" type="personal">
<mods:role id="9DD8C27AB88DD3275C0011778FF10BF1">
<mods:roleTerm id="0BF208AFD8AE6ABC585CBF825A9B28FD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3D91F7F033F0D25564491485F7CC14D6">Rifkind, Jacques</mods:namePart>
</mods:name>
<mods:typeOfResource id="E9EDF2E11B483AEB399A3E6C75564050">text</mods:typeOfResource>
<mods:relatedItem id="9BEE3575759C593316E83FE823B54D46" type="host">
<mods:titleInfo id="69C2581FF40C401765F80635DF46B0DC">
<mods:title id="95F6532CD5B67646367C8D1ADFA4D31D">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="A0874708CD950E6BB25AA016EF601AA2">
<mods:date id="D5CE6476965629DC6BD101CD3B96B731">2017</mods:date>
<mods:detail id="9840EDF2A136B5D449480527A00CAA84" type="pubDate">
<mods:number id="794D71CD164C638EDD87EDB671EA82B7">2017-12-29</mods:number>
</mods:detail>
<mods:detail id="B4F40ABA0DD0D3C92EAEBF48502E75FD" type="volume">
<mods:number id="CAC5AACD833FAE9049327ECB6E6213A1">2017</mods:number>
</mods:detail>
<mods:detail id="BE6C333C0476E8597F9B48E3234F8A6E" type="issue">
<mods:number id="010261558817C5BDE2EF0CD242A6990B">591</mods:number>
</mods:detail>
<mods:extent id="23B0676F27E2477A52E82C98EDCE98E3" unit="page">
<mods:start id="3B7D5EDDB4C28869806ABD671568B8A1">1</mods:start>
<mods:end id="4D319E1F3BC63F9F5FDEA5D0DA296762">18</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="2FC264DA9C4A08F6D45B615559192615">journal article</mods:classification>
<mods:identifier id="12A93D5040EE4E91CFCAC32B346070BE" type="DOI">10.5281/zenodo.5169515</mods:identifier>
<mods:identifier id="3D0BE95EE643AE35BF71AC63572E358B" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="A9B0AF471B307D88DF854BD808635BEA" type="Zenodo-Dep">5169515</mods:identifier>
<mods:identifier id="DB0D60C6D084A07DFBBB28775CFF3217" type="ZooBank">7F2A2366-B4E4-4F37-A5A5-45CB51D4D859</mods:identifier>
</mods:mods>
<treatment id="03F1E067CC01FFF110CFFDFAFD43FE81" LSID="urn:lsid:plazi:treatment:03F1E067CC01FFF110CFFDFAFD43FE81" httpUri="http://treatment.plazi.org/id/03F1E067CC01FFF110CFFDFAFD43FE81" lastPageId="8" lastPageNumber="7" pageId="7" pageNumber="6">
<subSubSection id="C34202FACC01FFFE10CFFDFAFC9EFDA7" box="[192,778,612,637]" pageId="7" pageNumber="6" type="nomenclature">
<paragraph id="8BE75171CC01FFFE10CFFDFAFC9EFDA7" blockId="7.[192,778,612,669]" box="[192,778,612,637]" pageId="7" pageNumber="6">
<heading id="D0AFE61DCC01FFFE10CFFDFAFC9EFDA7" bold="true" box="[192,778,612,637]" fontSize="10" level="2" pageId="7" pageNumber="6" reason="3">
<emphasis id="B92C8D63CC01FFFE10CFFDFAFC9EFDA7" bold="true" box="[192,778,612,637]" pageId="7" pageNumber="6">
<taxonomicName id="4C582AF2CC01FFFE10CFFDFAFDCCFDA7" authority="Rifkind" authorityName=", Rifkind" authorityYear="2017" box="[192,600,612,636]" class="Insecta" family="Melyridae" genus="Quadrophenia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="mooni" status="sp. nov.">
<emphasis id="B92C8D63CC01FFFE10CFFDFAFE71FDA7" bold="true" box="[192,485,612,636]" italics="true" pageId="7" pageNumber="6">Quadrophenia mooni</emphasis>
Rifkind
</taxonomicName>
,
<taxonomicNameLabel id="A21F3018CC01FFFE1269FDFBFC9EFDA7" box="[614,778,612,637]" pageId="7" pageNumber="6" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34202FACC01FFFE10CFFD1BFED0FD46" box="[192,324,645,669]" pageId="7" pageNumber="6" type="description">
<paragraph id="8BE75171CC01FFFE10CFFD1BFED0FD46" blockId="7.[192,778,612,669]" box="[192,324,645,669]" pageId="7" pageNumber="6">
(
<figureCitation id="13634DF4CC01FFFE10C6FD1BFE84FD46" box="[201,272,645,669]" captionStart="Figures 8–11" captionStartId="16.[192,288,1875,1897]" captionTargetBox="[326,1373,341,1753]" captionTargetId="figure-50@16.[192,1440,338,1773]" captionTargetPageId="16" captionText="Figures 8–11. Quadrophenia n.spp. 8) Quadrophenia mooni new species, habitus; 9) Quadrophenia entwistlei new species, habitus; 10) Quadrophenia entwistlei, lateral view; 11) Quadrophenia daltreyi new species, habitus." figureDoi="http://doi.org/10.5281/zenodo.5169521" httpUri="https://zenodo.org/record/5169521/files/figure.png" pageId="7" pageNumber="6">Fig. 8</figureCitation>
,
<figureCitation id="13634DF4CC01FFFE1111FD1BFEAFFD46" box="[286,315,645,669]" captionStart="Figures 17–21" captionStartId="19.[192,288,1875,1897]" captionText="Figures 17–21. Quadrophenia new genus, antennae of new species. 17) Q. townshendi, antenna; 18) Q. mooni, capitulum; 19) Q. entwistlei, capitulum; 20) Q. daltreyi, capitulum; 21) Q. sumidero, capitulum." pageId="7" pageNumber="6">18</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34202FACC01FFFE10CFFD5BFBD5FC64" pageId="7" pageNumber="6" type="materials_examined">
<paragraph id="8BE75171CC01FFFE10CFFD5BFBD5FC64" blockId="7.[190,1440,709,959]" pageId="7" pageNumber="6">
<emphasis id="B92C8D63CC01FFFE10CFFD5BFE1EFD06" bold="true" box="[192,394,709,733]" pageId="7" pageNumber="6">Type material.</emphasis>
<materialsCitation id="3B305B2CCC01FFFE119DFD5BFDE6FD26" collectingDate="2000-04-16" collectingDateMax="2000-04-19" collectingDateMin="2000-04-16" collectorName="J. &amp; A. Rifkind &amp; P. Gum" location="Monteverde" municipality="Costa Rica" pageId="7" pageNumber="6" specimenCount="1" stateProvince="Puntarenas" typeStatus="holotype">
<typeStatus id="54E3EFD3CC01FFFE119DFD5BFD90FD06" box="[402,516,709,733]" pageId="7" pageNumber="6" type="holotype">Holotype</typeStatus>
, female.
<collectingMunicipality id="6B83CB0BCC01FFFE127EFD5BFD6CFD06" box="[625,760,709,733]" pageId="7" pageNumber="6">Costa Rica</collectingMunicipality>
,
<collectingRegion id="499C9F93CC01FFFE130BFD58FC74FD06" box="[772,992,710,733]" country="Costa Rica" name="Puntarenas" pageId="7" pageNumber="6">Puntarenas Prov.</collectingRegion>
,
<location id="8E8707AACC01FFFE13E3FD5BFB16FD06" LSID="urn:lsid:plazi:treatment:03F1E067CC01FFF110CFFDFAFD43FE81:8E8707AACC01FFFE13E3FD5BFB16FD06" box="[1004,1154,709,733]" municipality="Costa Rica" name="Monteverde" pageId="7" pageNumber="6" stateProvince="Puntarenas">Monteverde</location>
, cloud forest,
<date id="FFE677B1CC01FFFE1523FD5BFF6AFD25" pageId="7" pageNumber="6" value="2000-04-16" valueMax="2000-04-19" valueMin="2000-04-16">
<collectingDate id="EFA28E59CC01FFFE1523FD5BFF6AFD25" pageId="7" pageNumber="6" value="2000-04-16" valueMax="2000-04-19" valueMin="2000-04-16">IV-16-19- 2000</collectingDate>
</date>
,
<collectorName id="26AD34A7CC01FFFE1106FD78FE56FD26" box="[265,450,741,766]" pageId="7" pageNumber="6">J. &amp; A. Rifkind</collectorName>
,
<collectorName id="26AD34A7CC01FFFE11C1FD78FDB9FD25" box="[462,557,742,766]" pageId="7" pageNumber="6">P. Gum</collectorName>
, colls
</materialsCitation>
.
<materialsCitation id="3B305B2CCC01FFFE1273FD7BFC4FFD25" box="[636,987,741,766]" collectionCode="CSCA" pageId="7" pageNumber="6" specimenCount="1" stateProvince="Puntarenas" typeStatus="holotype">
<typeStatus id="54E3EFD3CC01FFFE1273FD7BFD7FFD26" box="[636,747,741,765]" pageId="7" pageNumber="6" type="holotype">Holotype</typeStatus>
deposited in
<collectionCode id="ED49C9B4CC01FFFE1383FD78FC4FFD25" box="[908,987,742,766]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="7" pageNumber="6" type="Museum">CSCA</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B305B2CCC01FFFE13EAFD78FE71FCE5" collectingDate="1988-03" collectionCode="JNRC, INBIO" collectorName="Guanac. Pr. &amp; Biod." country="Costa Rica" elevation="1200" location="Volcan Cacao" municipality="Estac. Cacao" pageId="7" pageNumber="6" specimenCode="CR1000, 028563" specimenCount="1" typeStatus="paratype">
<typeStatus id="54E3EFD3CC01FFFE13EAFD78FBF2FD25" box="[997,1126,742,766]" pageId="7" pageNumber="6" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="F34F11E1CC01FFFE1460FD78FA89FD25" box="[1135,1309,742,766]" name="Costa Rica" pageId="7" pageNumber="6">COSTA RICA</collectingCountry>
: 1 (
<collectionCode id="ED49C9B4CC01FFFE154BFD78FA07FD25" box="[1348,1427,742,766]" pageId="7" pageNumber="6">JNRC</collectionCode>
),
<collectingMunicipality id="6B83CB0BCC01FFFE10B0FC98FEC9FCC5" box="[191,349,774,798]" pageId="7" pageNumber="6">Estac. Cacao</collectingMunicipality>
,
<quantity id="4CA0FC94CC01FFFE1168FC98FD9FFCC6" box="[359,523,774,798]" metricMagnitude="3" metricUnit="m" metricValue="1.2" metricValueMax="1.4" metricValueMin="1.0" pageId="7" pageNumber="6" unit="m" value="1200.0" valueMax="1400.0" valueMin="1000.0">
<elevation id="0075B642CC01FFFE1168FC98FD9FFCC6" box="[359,523,774,798]" metricMagnitude="3" metricUnit="m" metricValue="1.2" metricValueMax="1.4" metricValueMin="1.0" pageId="7" pageNumber="6" unit="m" value="1200.0" valueMax="1400.0" valueMin="1000.0">1000–1400 m</elevation>
</quantity>
, SW side
<location id="8E8707AACC01FFFE1272FC98FCB7FCC5" LSID="urn:lsid:plazi:treatment:03F1E067CC01FFF110CFFDFAFD43FE81:8E8707AACC01FFFE1272FC98FCB7FCC5" box="[637,803,774,798]" country="Costa Rica" municipality="Estac. Cacao" name="Volcan Cacao" pageId="7" pageNumber="6">Volcan Cacao</location>
,
<collectorName id="26AD34A7CC01FFFE1322FC98FC2BFCC6" box="[813,959,774,798]" pageId="7" pageNumber="6">Guanac. Pr.</collectorName>
,
<date id="FFE677B1CC01FFFE13C5FC98FBD5FCC5" box="[970,1089,774,798]" pageId="7" pageNumber="6" value="1988-03">
<collectingDate id="EFA28E59CC01FFFE13C5FC98FBD5FCC5" box="[970,1089,774,798]" pageId="7" pageNumber="6" value="1988-03">Mar 1988</collectingDate>
</date>
, GNP
<collectorName id="26AD34A7CC01FFFE1482FC98FB5BFCC5" box="[1165,1231,774,798]" pageId="7" pageNumber="6">Biod.</collectorName>
, 323300. 375700,
<collectionCode id="ED49C9B4CC01FFFE10B1FCB8FE85FCE5" box="[190,273,806,830]" pageId="7" pageNumber="6">INBIO</collectionCode>
<specimenCode id="DBFEF90ACC01FFFE1116FCB8FEEAFCE5" box="[281,382,806,830]" collectionCode="INBIO" pageId="7" pageNumber="6">CR1000</specimenCode>
,
<specimenCode id="DBFEF90ACC01FFFE1186FCB8FE71FCE5" box="[393,485,806,830]" collectionCode="INBIO" pageId="7" pageNumber="6">028563</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B305B2CCC01FFFE11FEFCB8FA9AFCE5" box="[497,1294,806,830]" collectingDate="1988-03" collectionCode="JNRC, INBIO" collectorName="Guanac. Pr. &amp; Biod." country="Costa Rica" elevation="1200" location="Volcan Cacao" municipality="Estac. Cacao" pageId="7" pageNumber="6" specimenCode="CR1000, 028339" specimenCount="1" typeStatus="paratype">
1 (
<collectionCode id="ED49C9B4CC01FFFE1200FCB8FDCAFCE5" box="[527,606,806,830]" pageId="7" pageNumber="6">JNRC</collectionCode>
), same data as previous, except
<collectionCode id="ED49C9B4CC01FFFE13E8FCB8FBAEFCE5" box="[999,1082,806,830]" pageId="7" pageNumber="6">INBIO</collectionCode>
<specimenCode id="DBFEF90ACC01FFFE144DFCB8FB33FCE5" box="[1090,1191,806,830]" collectionCode="INBIO" pageId="7" pageNumber="6">CR1000</specimenCode>
,
<specimenCode id="DBFEF90ACC01FFFE14BDFCB8FA9AFCE5" box="[1202,1294,806,830]" collectionCode="INBIO" pageId="7" pageNumber="6">028339</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B305B2CCC01FFFE1515FCB8FCC0FC85" collectingDate="1988-03" collectionCode="MUCR, INBIO" collectorName="Guanac. Pr. &amp; Biod." country="Costa Rica" elevation="1200" location="Volcan Cacao" municipality="Estac. Cacao" pageId="7" pageNumber="6" specimenCode="CR1000, 028586" specimenCount="1" typeStatus="paratype">
1 (
<collectionCode id="ED49C9B4CC01FFFE1538FCB8FA05FCE5" box="[1335,1425,806,830]" country="Costa Rica" httpUri="http://grbio.org/cool/ww9x-1v7b" name="Museo de Insectos" pageId="7" pageNumber="6">MUCR</collectionCode>
), same data as previous, except
<collectionCode id="ED49C9B4CC01FFFE1220FCD8FD16FC85" box="[559,642,838,862]" pageId="7" pageNumber="6">INBIO</collectionCode>
<specimenCode id="DBFEF90ACC01FFFE1286FCD8FD7AFC85" box="[649,750,838,862]" collectionCode="INBIO" pageId="7" pageNumber="6">CR1000</specimenCode>
,
<specimenCode id="DBFEF90ACC01FFFE12F7FCD8FCC0FC85" box="[760,852,838,862]" collectionCode="INBIO" pageId="7" pageNumber="6">028586</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B305B2CCC01FFFE1351FCD8FE1FFCA4" collectingDate="1988-03" collectionCode="CSCA, INBIO" collectorName="Guanac. Pr. &amp; Biod." country="Costa Rica" elevation="1200" location="Volcan Cacao" municipality="Estac. Cacao" pageId="7" pageNumber="6" specimenCode="CR1000, 028372" specimenCount="1" typeStatus="paratype">
1 (
<collectionCode id="ED49C9B4CC01FFFE1374FCD8FC5EFC85" box="[891,970,838,862]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="7" pageNumber="6" type="Museum">CSCA</collectionCode>
), same data as previous, except
<collectionCode id="ED49C9B4CC01FFFE1543FCD8FA0BFC85" box="[1356,1439,838,862]" pageId="7" pageNumber="6">INBIO</collectionCode>
<specimenCode id="DBFEF90ACC01FFFE10B0FCF9FEB0FCA4" box="[191,292,871,895]" collectionCode="INBIO" pageId="7" pageNumber="6">CR1000</specimenCode>
,
<specimenCode id="DBFEF90ACC01FFFE1120FCF9FE1FFCA4" box="[303,395,871,895]" collectionCode="INBIO" pageId="7" pageNumber="6">028372</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B305B2CCC01FFFE1197FCF9FBFAFC44" collectionCode="WOPC, INBIO" collectorName="Prov. Guan. &amp; Malaise &amp; Mar &amp; Ago" country="Costa Rica" elevation="1200" location="Lado Suroeste del Volcan Cacao" municipality="Est. Cacao" pageId="7" pageNumber="6" specimenCode="CR1000, 232988" specimenCount="1" typeStatus="paratype">
1 (
<collectionCode id="ED49C9B4CC01FFFE11B9FCF9FD9BFCA4" box="[438,527,871,895]" pageId="7" pageNumber="6">WOPC</collectionCode>
),
<collectingMunicipality id="6B83CB0BCC01FFFE122CFCF9FD3DFCA4" box="[547,681,871,895]" pageId="7" pageNumber="6">Est. Cacao</collectingMunicipality>
,
<quantity id="4CA0FC94CC01FFFE12BAFCF9FCC8FCA5" box="[693,860,871,895]" metricMagnitude="3" metricUnit="m" metricValue="1.2" metricValueMax="1.4" metricValueMin="1.0" pageId="7" pageNumber="6" unit="m" value="1200.0" valueMax="1400.0" valueMin="1000.0">
<elevation id="0075B642CC01FFFE12BAFCF9FCC8FCA5" box="[693,860,871,895]" metricMagnitude="3" metricUnit="m" metricValue="1.2" metricValueMax="1.4" metricValueMin="1.0" pageId="7" pageNumber="6" unit="m" value="1200.0" valueMax="1400.0" valueMin="1000.0">1000–1400 m</elevation>
</quantity>
,
<location id="8E8707AACC01FFFE1366FCF8FB63FCA4" LSID="urn:lsid:plazi:treatment:03F1E067CC01FFF110CFFDFAFD43FE81:8E8707AACC01FFFE1366FCF8FB63FCA4" box="[873,1271,870,895]" country="Costa Rica" municipality="Est. Cacao" name="Lado Suroeste del Volcan Cacao" pageId="7" pageNumber="6">Lado Suroeste del Volcan Cacao</location>
,
<collectorName id="26AD34A7CC01FFFE150CFCF9FA0EFCA4" box="[1283,1434,871,895]" pageId="7" pageNumber="6">Prov. Guan.</collectorName>
, tp
<collectorName id="26AD34A7CC01FFFE10EDFC19FED2FC44" box="[226,326,903,927]" pageId="7" pageNumber="6">Malaise</collectorName>
,
<collectorName id="26AD34A7CC01FFFE115CFC19FE13FC45" box="[339,391,903,926]" pageId="7" pageNumber="6">Mar</collectorName>
a
<collectorName id="26AD34A7CC01FFFE11A9FC19FE4CFC44" box="[422,472,903,927]" pageId="7" pageNumber="6">Ago</collectorName>
, 1990, L–N–323300, 375700,
<collectionCode id="ED49C9B4CC01FFFE134AFC19FC0CFC44" box="[837,920,903,927]" pageId="7" pageNumber="6">INBIO</collectionCode>
<specimenCode id="DBFEF90ACC01FFFE13AEFC19FB92FC44" box="[929,1030,903,927]" collectionCode="INBIO" pageId="7" pageNumber="6">CR1000</specimenCode>
,
<specimenCode id="DBFEF90ACC01FFFE141DFC19FBFAFC44" box="[1042,1134,903,927]" collectionCode="INBIO" pageId="7" pageNumber="6">232988</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B305B2CCC01FFFE1474FC19FBA9FC64" collectionCode="EMEC" collectorName="J. A. Chemsak" country="Costa Rica" elevation="1300" location="Monteverde" pageId="7" pageNumber="6" specimenCount="1" stateProvince="Puntarenas" typeStatus="paratype">
1 (
<collectionCode id="ED49C9B4CC01FFFE1495FC19FB65FC44" box="[1178,1265,903,927]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" lsid="urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="7" pageNumber="6" type="University or college">EMEC</collectionCode>
),
<location id="8E8707AACC01FFFE1509FC19FA08FC44" LSID="urn:lsid:plazi:treatment:03F1E067CC01FFF110CFFDFAFD43FE81:8E8707AACC01FFFE1509FC19FA08FC44" box="[1286,1436,903,927]" country="Costa Rica" name="Monteverde" pageId="7" pageNumber="6" stateProvince="Puntarenas">Monteverde</location>
,
<collectingRegion id="499C9F93CC01FFFE10CEFC39FE08FC65" box="[193,412,935,958]" country="Costa Rica" name="Puntarenas" pageId="7" pageNumber="6">Puntarenas Prov.</collectingRegion>
,
<quantity id="4CA0FC94CC01FFFE11A7FC39FD91FC65" box="[424,517,935,959]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="6" unit="m" value="1300.0">
<elevation id="0075B642CC01FFFE11A7FC39FD91FC65" box="[424,517,935,959]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="6" unit="m" value="1300.0">1300 m</elevation>
</quantity>
, V-17 to V-20-[19]85,
<collectorName id="26AD34A7CC01FFFE131FFC39FC52FC64" box="[784,966,935,959]" pageId="7" pageNumber="6">J. A. Chemsak</collectorName>
, collector
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34202FACC01FFFE10CFFC79FEBAFB5B" pageId="7" pageNumber="6" type="diagnosis">
<paragraph id="8BE75171CC01FFFE10CFFC79FEBAFB5B" blockId="7.[191,1440,999,1152]" pageId="7" pageNumber="6">
<emphasis id="B92C8D63CC01FFFE10CFFC79FEC4FC24" bold="true" box="[192,336,999,1023]" pageId="7" pageNumber="6">Diagnosis.</emphasis>
<taxonomicName id="4C582AF2CC01FFFE1159FC79FDCEFC24" authorityName=", Rifkind" authorityYear="2017" box="[342,602,999,1023]" class="Insecta" family="Melyridae" genus="Quadrophenia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="mooni">
<emphasis id="B92C8D63CC01FFFE1159FC79FDCEFC24" box="[342,602,999,1023]" italics="true" pageId="7" pageNumber="6">Quadrophenia mooni</emphasis>
</taxonomicName>
can be distinguished from congeners based on a unique combination of dorsal color and pattern. From
<taxonomicName id="4C582AF2CC01FFFE125BFB96FC97FBFB" authorityName=", Rifkind" authorityYear="2017" box="[596,771,1032,1056]" class="Insecta" family="Melyridae" genus="Quadrophenia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="townshendi">
<emphasis id="B92C8D63CC01FFFE125BFB96FC97FBFB" box="[596,771,1032,1056]" italics="true" pageId="7" pageNumber="6">Q. townshendi</emphasis>
</taxonomicName>
it can in most cases be separated by its flavotestaceous pronotal and elytral ground color. From specimens of the uncommon testaceous morph of
<taxonomicName id="4C582AF2CC01FFFE14FFFBB6FA0AFB9B" authorityName=", Rifkind" authorityYear="2017" box="[1264,1438,1064,1088]" class="Insecta" family="Melyridae" genus="Quadrophenia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="townshendi">
<emphasis id="B92C8D63CC01FFFE14FFFBB6FA0AFB9B" box="[1264,1438,1064,1088]" italics="true" pageId="7" pageNumber="6">Q. townshendi</emphasis>
</taxonomicName>
(and from
<taxonomicName id="4C582AF2CC01FFFE114FFBD6FE4DFBBB" authorityName=", Rifkind" authorityYear="2017" box="[320,473,1096,1120]" class="Insecta" family="Melyridae" genus="Quadrophenia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="entwistlei">
<emphasis id="B92C8D63CC01FFFE114FFBD6FE4DFBBB" box="[320,473,1096,1120]" italics="true" pageId="7" pageNumber="6">Q. entwistlei</emphasis>
</taxonomicName>
, described below) it can be distinguished by its uniformly testaceous antennae and legs.
</paragraph>
</subSubSection>
<subSubSection id="C34202FACC01FFFE10CFFB36FA34F8DE" pageId="7" pageNumber="6" type="description">
<paragraph id="8BE75171CC01FFFE10CFFB36FB20F998" blockId="7.[190,1442,1192,1603]" pageId="7" pageNumber="6">
<emphasis id="B92C8D63CC01FFFE10CFFB36FEFFFB1B" bold="true" box="[192,363,1192,1216]" pageId="7" pageNumber="6">Description.</emphasis>
(
<typeStatus id="54E3EFD3CC01FFFE1174FB37FE7BFB1A" box="[379,495,1193,1217]" pageId="7" pageNumber="6" type="holotype">Holotype</typeStatus>
). Length: 9.0 mm. Form: subflattened dorsally, expanded posteriorly. Color: flavotestaceous (pronotum slightly orange-testaceous at middle); mesepisternum, mesosternum later- ally, and metasternum, piceous; head, mandibles, a curvate V-shaped spot at pronotal anterior margin, and scutellum, black; elytra with four large black maculae arranged as in
<figureCitation id="13634DF4CC01FFFE1431FA97FB10FAF9" box="[1086,1156,1289,1314]" captionStart="Figures 8–11" captionStartId="16.[192,288,1875,1897]" captionTargetBox="[326,1373,341,1753]" captionTargetId="figure-50@16.[192,1440,338,1773]" captionTargetPageId="16" captionText="Figures 8–11. Quadrophenia n.spp. 8) Quadrophenia mooni new species, habitus; 9) Quadrophenia entwistlei new species, habitus; 10) Quadrophenia entwistlei, lateral view; 11) Quadrophenia daltreyi new species, habitus." figureDoi="http://doi.org/10.5281/zenodo.5169521" httpUri="https://zenodo.org/record/5169521/files/figure.png" pageId="7" pageNumber="6">Fig. 8</figureCitation>
; maculae not attaining epipleura laterally. Antennae: terminal antennomere with internal and external / distal emarginations exaggerated, apex prolonged, subacuminate (
<figureCitation id="13634DF4CC01FFFE12ECFAD4FCACFAB9" box="[739,824,1354,1378]" captionStart="Figures 17–21" captionStartId="19.[192,288,1875,1897]" captionText="Figures 17–21. Quadrophenia new genus, antennae of new species. 17) Q. townshendi, antenna; 18) Q. mooni, capitulum; 19) Q. entwistlei, capitulum; 20) Q. daltreyi, capitulum; 21) Q. sumidero, capitulum." pageId="7" pageNumber="6">Fig. 18</figureCitation>
). Pronotum: sparsely, coarsely but shallowly punctate posteriorly on either side of midline. Elytra: longer than broad (ratio of length to maximum width 9:7); expanded posteriorly; widest at posterior 1/3; surface shining, shallowly roughened, punctures shallow, mostly indistinct, somewhat larger and more densely arranged at sides; minute asetiferous punctations also present; vestiture composed of erect and suberect, rather robust black setae. Abdomen: ventrite 5 broad, hind angles gently rounded, posterior margin broadly, arcuately emarginate; ventrite 6 arcuately rounded posteriorly, very slightly, shallowly inflected at middle of hind margin; tergite 6 semicircularly rounded posteriorly, surpassing hind margin of ventrite 6.
</paragraph>
<paragraph id="8BE75171CC01FFFE10CFF9F2FA34F8DE" blockId="7.[190,1440,1644,1797]" pageId="7" pageNumber="6">
<emphasis id="B92C8D63CC01FFFE10CFF9F2FEDEF95F" bold="true" box="[192,330,1644,1668]" pageId="7" pageNumber="6">Variation.</emphasis>
Length of specimens ranges from 8.0–9.75 mm. The extent and placement of the dorsal black maculae is somewhat variable; the pronotum of some specimens is narrowly darkened posteriorly, and one Costa Rican specimen has the epipleural margin darkened along its length, and the elytral humeral and postmedian maculae connected by a narrow dark streak on either side of the sutural bead. The male pygidium has ventrite 6 with the hind margin rather shallowly, arcuately emarginate at middle.
</paragraph>
</subSubSection>
<subSubSection id="C34202FACC01FFFE10CFF8B3FCC1F85D" pageId="7" pageNumber="6" type="distribution">
<paragraph id="8BE75171CC01FFFE10CFF8B3FCC1F85D" blockId="7.[192,1440,1837,1926]" pageId="7" pageNumber="6">
<emphasis id="B92C8D63CC01FFFE10CFF8B3FEE0F89E" bold="true" box="[192,372,1837,1861]" pageId="7" pageNumber="6">Distribution.</emphasis>
This species is known from the Cordillera de
<collectingRegion id="499C9F93CC01FFFE13A5F8B3FBA8F89E" box="[938,1084,1837,1861]" country="Costa Rica" name="Guanacaste" pageId="7" pageNumber="6">Guanacaste</collectingRegion>
and Cordillera de Tilarán in
<collectingCountry id="F34F11E1CC01FFFE10CFF8D0FED3F8BD" box="[192,327,1870,1894]" name="Costa Rica" pageId="7" pageNumber="6">Costa Rica</collectingCountry>
. It has been collected in cloud forest in Monteverde by beating, and in Malaise traps set between
<quantity id="4CA0FC94CC01FFFE111EF8F0FEF8F85E" box="[273,364,1902,1926]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="6" unit="m" value="1000.0">1000 m</quantity>
and
<quantity id="4CA0FC94CC01FFFE11A6F8F0FD90F85E" box="[425,516,1902,1926]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="7" pageNumber="6" unit="m" value="1400.0">1400 m</quantity>
elevation on Volcán Cacao.
</paragraph>
</subSubSection>
<subSubSection id="C34202FACC0EFFF110CFFF5CFB67FF01" box="[192,1267,194,218]" pageId="8" pageNumber="7" type="etymology">
<paragraph id="8BE75171CC0EFFF110CFFF5CFB67FF01" blockId="8.[192,1267,194,218]" box="[192,1267,194,218]" pageId="8" pageNumber="7">
<emphasis id="B92C8D63CC0EFFF110CFFF5CFEC8FF01" bold="true" box="[192,348,194,218]" pageId="8" pageNumber="7">Etymology.</emphasis>
The new species is named for the late Keith Moon, drummer for The Who.
</paragraph>
</subSubSection>
<subSubSection id="C34202FACC0EFFF110CFFE9CFD43FE81" pageId="8" pageNumber="7" type="discussion">
<paragraph id="8BE75171CC0EFFF110CFFE9CFD43FE81" blockId="8.[190,1440,258,346]" pageId="8" pageNumber="7">
<emphasis id="B92C8D63CC0EFFF110CFFE9CFED0FEC1" bold="true" box="[192,324,258,282]" pageId="8" pageNumber="7">Remarks.</emphasis>
<taxonomicName id="4C582AF2CC0EFFF11146FE9CFDD9FEC2" authorityName=", Rifkind" authorityYear="2017" box="[329,589,258,282]" class="Insecta" family="Melyridae" genus="Quadrophenia" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="mooni">
<emphasis id="B92C8D63CC0EFFF11146FE9CFDD9FEC2" box="[329,589,258,282]" italics="true" pageId="8" pageNumber="7">Quadrophenia mooni</emphasis>
</taxonomicName>
is a strikingly similar mimic of several sympatric species of leaf beetle belonging to the genus
<taxonomicName id="4C582AF2CC0EFFF111EBFEBCFD7CFEE1" authority="Chevrolat" authorityName="Chevrolat" box="[484,744,290,314]" class="Insecta" family="Chrysomelidae" genus="Diabrotica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B92C8D63CC0EFFF111EBFEBCFDF2FEE1" box="[484,614,290,314]" italics="true" pageId="8" pageNumber="7">Diabrotica</emphasis>
Chevrolat
</taxonomicName>
(
<taxonomicName id="4C582AF2CC0EFFF112F6FEBCFC22FEE1" authorityName="Latreille" authorityYear="1802" box="[761,950,290,314]" class="Insecta" family="Chrysomelidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="family">Chrysomelidae</taxonomicName>
: Galerucinae) (e.g.
<taxonomicName id="4C582AF2CC0EFFF114A8FEBCFDCCFE81" authority="Derunkov, Prado, Tishechkin, Konstantinov" authorityName="Derunkov, Prado, Tishechkin, Konstantinov" class="Insecta" family="Chrysomelidae" genus="Diabrotica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="caveyi">
<emphasis id="B92C8D63CC0EFFF114A8FEBCFA83FEE2" box="[1191,1303,290,313]" italics="true" pageId="8" pageNumber="7">D. caveyi</emphasis>
Derunkov, Prado, Tishechkin, Konstantinov
</taxonomicName>
(Fig. 22)).
</paragraph>
</subSubSection>
</treatment>
</document>
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