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<document id="98320CD9E4AFE81A10BD0D6BBDDE0671" ID-CLB-Dataset="20167" ID-DOI="10.5852/ejt.2022.803.1691" ID-GBIF-Dataset="af09c8e1-b481-4de0-b9d0-83ba26bf9876" ID-ISSN="2118-9773" ID-Zenodo-Dep="6359066" ID-ZooBank="8B66C8AE-F00A-42F6-9641-26B0ECC49F78" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1647350252755" checkinUser="felipe" docAuthor="Enghoff, Henrik" docDate="2022" docId="03F15C39D6760661FDA8FC39C048FA79" docLanguage="en" docName="ejt-803.1-136.pdf" docOrigin="European Journal of Taxonomy 803" docSource="http://dx.doi.org/10.5852/ejt.2022.803.1691" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Raduliverpa donatellae Enghoff 2022, sp. nov." docType="treatment" docUuid="640AD516-04DF-433C-8229-7F450593BF5A" docUuidSource="ZooBank" docVersion="8" lastPageNumber="87" masterDocId="FFC82441D6250637FFF3FFE3C52CFFD8" masterDocTitle="Mountains of millipedes. The family Odontopygidae in the Eastern Arc Mountains of Tanzania (Diplopoda, Spirostreptida)" masterLastPageNumber="136" masterPageNumber="1" pageNumber="84" updateTime="1699318071475" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="349EC35FDAF13B9775D190827C1A9D29">Mountains of millipedes. The family Odontopygidae in the Eastern Arc Mountains of Tanzania (Diplopoda, Spirostreptida)</mods:title>
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<mods:namePart id="8878FD5C7BFBBFFA80B13DAF6073C033">Enghoff, Henrik</mods:namePart>
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<mods:affiliation id="224899C6287D15BD7CAF077A30F12251">Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100 Copenhagen Ø, Denmark.</mods:affiliation>
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<mods:title id="23781A30E7090606FE173FE33500EFF7">European Journal of Taxonomy</mods:title>
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<mods:date id="5A43936BAA04279A94100D1620DBDD8A">2022</mods:date>
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<treatment id="03F15C39D6760661FDA8FC39C048FA79" ID-DOI="http://doi.org/10.5281/zenodo.6373715" ID-GBIF-Taxon="194181578" ID-Zenodo-Dep="6373715" LSID="urn:lsid:zoobank.org:act:640AD516-04DF-433C-8229-7F450593BF5A" httpUri="http://treatment.plazi.org/id/03F15C39D6760661FDA8FC39C048FA79" lastPageId="86" lastPageNumber="87" pageId="83" pageNumber="84">
<subSubSection id="C342BEA4D6760664FDA8FC39C6F5FC2D" box="[603,985,986,1013]" pageId="83" pageNumber="84" type="nomenclature">
<paragraph id="8BE7ED2FD6760664FDA8FC39C6F5FC2D" blockId="83.[373,1216,986,1085]" box="[603,985,986,1013]" pageId="83" pageNumber="84">
<heading id="D0AF5A43D6760664FDA8FC39C6F5FC2D" box="[603,985,986,1013]" centered="true" fontSize="11" level="2" pageId="83" pageNumber="84" reason="2">
<taxonomicName id="4C5896ACD6760664FDA8FC39C65BFC2D" ID-CoL="98ZHJ" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[603,887,986,1013]" class="Diplopoda" family="Odontopygidae" genus="Raduliverpa" kingdom="Animalia" order="Spirostreptida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="donatellae" status="sp. nov.">
<emphasis id="B92C313DD6760664FDA8FC39C65BFC2D" bold="true" box="[603,887,986,1013]" italics="true" pageId="83" pageNumber="84">Raduliverpa donatellae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F8C46D6760664FC8DFC38C6F5FC2D" box="[894,985,987,1013]" pageId="83" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C342BEA4D6760664FE86FC1DC677FBE5" pageId="83" pageNumber="84" type="description">
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<uri id="FFC9E12DD6760664FE86FC1DC1ECFBC1" box="[373,1216,1022,1049]" pageId="83" pageNumber="84">
urn:lsid:zoobank.org:act:
<uuid id="FFFED7FAD6760664FD62FC1DC1ECFBC1" box="[657,1216,1022,1049]" pageId="83" pageNumber="84">640AD516-04DF-433C-8229-7F450593BF5A</uuid>
</uri>
</paragraph>
<paragraph id="8BE7ED2FD6760664FD2BFBC0C677FBE5" blockId="83.[373,1216,986,1085]" box="[728,859,1058,1085]" pageId="83" pageNumber="84">
<figureCitation id="1363F1AAD6760664FD2BFBC0C677FBE5" box="[728,859,1058,1085]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="84.[189,232,1788,1814]" captionStartId-1="85.[189,232,1722,1748]" captionTargetBox-0="[260,1328,266,1762]" captionTargetBox-1="[189,1398,265,1669]" captionTargetId-0="figure-117@84.[260,1328,266,1762]" captionTargetId-1="figure-99@85.[189,1398,265,1669]" captionTargetPageId-0="84" captionTargetPageId-1="85" captionText-0="Fig. 52. Raduliverpa donatellae sp. nov. AC. Holotype, ♂ (NHMD 621759). Left gonopod coxa. A. posterior view. B. Mesal view. C. Anterior view. D. Midbody dorsal limbus. EG. Paratype, ♂ (NHMD 621760), first and second pair of legs (right first legs misshapen/diminished). E. Sublateral view. F. Anterior view. G. Ventral view. Abbreviations: APS= mesapical prefemoral setae; CXS =coxosternal setae; LCS= lateral coxalprocess;LPS = lateral prefemoralsetae;MF= metaplical flange; MP=metaplica; msp = metaplical spinelike process; mtp =metaplical triangular process; PP = proplica. Scale bars: AC, EG =0.1 mm; D =0.01 mm." captionText-1="Fig. 53. Raduliverpa donatellae sp. nov., holotype, ♂ (NHMD 621759), gonopod telopodites. A. Right telopodite, posterior view. B. Left telopodite, subanterior view. C. Right telopodite, disto-mesal-basal view. D. Close-up of right solenomere tip (also with part of post-torsal spine) to show the genuscharacteristic striation. E. Left telopodite, anterior-basal view. Abbreviations: PTS=post-torsal spine; SLM = solenomere; TL =torsotope lobe; TM = telomere; tlm = telomere lamellae; TT =torsotope. Scale bars: AC, E =0.1 mm; D =0.02 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6359231" figureDoi-1="http://doi.org/10.5281/zenodo.6359233" httpUri-0="https://zenodo.org/record/6359231/files/figure.png" httpUri-1="https://zenodo.org/record/6359233/files/figure.png" pageId="83" pageNumber="84">Figs 5253</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C342BEA4D6760664FF4EFB87C167FAE7" pageId="83" pageNumber="84" type="diagnosis">
<paragraph id="8BE7ED2FD6760664FF4EFB87C41AFBA6" blockId="83.[189,310,1124,1150]" box="[189,310,1124,1150]" pageId="83" pageNumber="84">
<heading id="D0AF5A43D6760664FF4EFB87C41AFBA6" bold="true" box="[189,310,1124,1150]" fontSize="11" level="3" pageId="83" pageNumber="84" reason="3">
<emphasis id="B92C313DD6760664FF4EFB87C41AFBA6" bold="true" box="[189,310,1124,1150]" pageId="83" pageNumber="84">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BE7ED2FD6760664FF4EFB77C167FAE7" blockId="83.[189,1399,1172,1343]" pageId="83" pageNumber="84">
Differs from other species of
<taxonomicName id="4C5896ACD6760664FDDBFB77C795FB76" authorityName="Frederiksen &amp; Enghoff" authorityYear="2015" box="[552,697,1172,1198]" class="Diplopoda" family="Odontopygidae" genus="Raduliverpa" kingdom="Animalia" order="Spirostreptida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B92C313DD6760664FDDBFB77C795FB76" box="[552,697,1172,1198]" italics="true" pageId="83" pageNumber="84">Raduliverpa</emphasis>
</taxonomicName>
by having the limbus only very shallowly lobed, and the metaplical spinelike process (
<emphasis id="B92C313DD6760664FDD9FB5AC775FB0A" box="[554,601,1209,1234]" italics="true" pageId="83" pageNumber="84">msp</emphasis>
) very strongly curved, almost like a pig-tail, as opposed to the serrated limbus and much less curving
<emphasis id="B92C313DD6760664FD75FB3EC799FB2E" box="[646,693,1245,1270]" italics="true" pageId="83" pageNumber="84">msp</emphasis>
seen in other species, see
<bibRefCitation id="EFC990DED6760664FC1AFB3FC061FB2F" author="Frederiksen S. B. &amp; Enghoff H." box="[1001,1357,1244,1271]" pageId="83" pageNumber="84" pagination="541 - 560" refId="ref48260" refString="Frederiksen S. B. &amp; Enghoff H. 2015. East African odontopygid millipedes 4: A restricted redefinition of the genus Rhamphidarpoides Kraus, 1960, a related new genus, five new species, and notes on solenomere function (Diplopoda; Spirostreptida; Odontopygidae). Zootaxa 3926: 541 - 560. https: // doi. org / 10.11646 / zootaxa. 3926.4.5" type="journal article" year="2015">Frederiksen &amp; Enghoff (2015)</bibRefCitation>
for
<taxonomicName id="4C5896ACD6760664FF4EFAE2C7F7FAC3" authority="Frederiksen &amp; Enghoff, 2015" authorityName="Frederiksen &amp; Enghoff" authorityYear="2015" box="[189,731,1280,1307]" class="Diplopoda" family="Odontopygidae" genus="Raduliverpa" kingdom="Animalia" order="Spirostreptida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="serpentispina">
<emphasis id="B92C313DD6760664FF4EFAE2C457FAC2" box="[189,379,1281,1306]" italics="true" pageId="83" pageNumber="84">R. serpentispina</emphasis>
<bibRefCitation id="EFC990DED6760664FE70FAE3C7F7FAC3" author="Frederiksen S. B. &amp; Enghoff H." box="[387,731,1280,1307]" pageId="83" pageNumber="84" pagination="541 - 560" refId="ref48260" refString="Frederiksen S. B. &amp; Enghoff H. 2015. East African odontopygid millipedes 4: A restricted redefinition of the genus Rhamphidarpoides Kraus, 1960, a related new genus, five new species, and notes on solenomere function (Diplopoda; Spirostreptida; Odontopygidae). Zootaxa 3926: 541 - 560. https: // doi. org / 10.11646 / zootaxa. 3926.4.5" type="journal article" year="2015">Frederiksen &amp; Enghoff, 2015</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C5896ACD6760664FCE0FAE2C1FCFAC3" authority="Frederiksen &amp; Enghoff, 2015" authorityName="Frederiksen &amp; Enghoff" authorityYear="2015" box="[787,1232,1280,1307]" class="Diplopoda" family="Odontopygidae" genus="Raduliverpa" kingdom="Animalia" order="Spirostreptida" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="mitis">
<emphasis id="B92C313DD6760664FCE0FAE2C641FAC2" box="[787,877,1281,1306]" italics="true" pageId="83" pageNumber="84">R. mitis</emphasis>
<bibRefCitation id="EFC990DED6760664FC87FAE3C1FCFAC3" author="Frederiksen S. B. &amp; Enghoff H." box="[884,1232,1280,1307]" pageId="83" pageNumber="84" pagination="541 - 560" refId="ref48260" refString="Frederiksen S. B. &amp; Enghoff H. 2015. East African odontopygid millipedes 4: A restricted redefinition of the genus Rhamphidarpoides Kraus, 1960, a related new genus, five new species, and notes on solenomere function (Diplopoda; Spirostreptida; Odontopygidae). Zootaxa 3926: 541 - 560. https: // doi. org / 10.11646 / zootaxa. 3926.4.5" type="journal article" year="2015">Frederiksen &amp; Enghoff, 2015</bibRefCitation>
</taxonomicName>
,
<bibRefCitation id="EFC990DED6760664FB28FAE3C05AFAC3" author="Kraus O." box="[1243,1398,1280,1307]" pageId="83" pageNumber="84" pagination="1 - 207" refId="ref48861" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae-Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." type="journal article" year="1960">Kraus (1960)</bibRefCitation>
for
<emphasis id="B92C313DD6760664FF14FAC6C470FAE6" box="[231,348,1317,1342]" italics="true" pageId="83" pageNumber="84">R. serrata</emphasis>
(
<bibRefCitation id="EFC990DED6760664FE98FAC7C4D2FAE7" author="Kraus O." box="[363,510,1316,1343]" pageId="83" pageNumber="84" pagination="1 - 207" refId="ref48861" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae-Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." type="journal article" year="1960">Kraus, 1960</bibRefCitation>
) and
<bibRefCitation id="EFC990DED6760664FDCDFAC7C7C4FAE7" author="Attems C." box="[574,744,1316,1343]" pageId="83" pageNumber="84" pagination="158 - 201" refId="ref46316" refString="Attems C. 1935. Diplopoden des Belgischen Congo II. Spirostreptoidea und Spiroboloidea. Revue de Zoologie et de Botanique africaines 26: 98 - 11 + 158 - 201 + 327 - 396." type="journal article" year="1935">Attems (1935)</bibRefCitation>
for
<emphasis id="B92C313DD6760664FCEAFAC6C6A6FAE6" box="[793,906,1317,1342]" italics="true" pageId="83" pageNumber="84">R. sicaria</emphasis>
(
<bibRefCitation id="EFC990DED6760664FC69FAC7C112FAE7" author="Attems C." box="[922,1086,1316,1343]" pageId="83" pageNumber="84" pagination="158 - 201" refId="ref46316" refString="Attems C. 1935. Diplopoden des Belgischen Congo II. Spirostreptoidea und Spiroboloidea. Revue de Zoologie et de Botanique africaines 26: 98 - 11 + 158 - 201 + 327 - 396." type="journal article" year="1935">Attems, 1935</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C342BEA4D6760664FF4EFA85C1F8FA77" box="[189,1236,1382,1455]" pageId="83" pageNumber="84" type="etymology">
<paragraph id="8BE7ED2FD6760664FF4EFA85C468FA58" blockId="83.[189,324,1382,1408]" box="[189,324,1382,1408]" pageId="83" pageNumber="84">
<heading id="D0AF5A43D6760664FF4EFA85C468FA58" bold="true" box="[189,324,1382,1408]" fontSize="11" level="3" pageId="83" pageNumber="84" reason="3">
<emphasis id="B92C313DD6760664FF4EFA85C468FA58" bold="true" box="[189,324,1382,1408]" pageId="83" pageNumber="84">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BE7ED2FD6760664FF4EFA76C1F8FA77" blockId="83.[189,1236,1429,1455]" box="[189,1236,1429,1455]" pageId="83" pageNumber="84">After Donatella Foddai, Italian myriapodologist, former resident of Hotel Dolichoiulus.</paragraph>
</subSubSection>
<subSubSection id="C342BEA4D6760664FF4EFA34C130F916" pageId="83" pageNumber="84" type="materials_examined">
<paragraph id="8BE7ED2FD6760664FF4EFA34C76FFA2A" blockId="83.[189,579,1495,1522]" box="[189,579,1495,1522]" pageId="83" pageNumber="84">
<emphasis id="B92C313DD6760664FF4EFA34C486FA29" bold="true" box="[189,426,1495,1521]" pageId="83" pageNumber="84">Material examined</emphasis>
(total
<specimenCount id="9D5E26A6D6760664FE04FA34C716FA2A" box="[503,570,1495,1522]" pageId="83" pageNumber="84" type="male">2 ♂♂</specimenCount>
)
</paragraph>
<paragraph id="8BE7ED2FD6760664FF19F9E4C67AF9B1" blockId="83.[189,1399,1543,1641]" pageId="83" pageNumber="84">
<materialsCitation id="3B30E772D6760664FF19F9E4C67AF9B1" ID-GBIF-Occurrence="3712026349" collectingDate="1996-09" collectionCode="NHMD" collectorName="Mgambo Proposed FR &amp; Norbert Cordeiro" country="Tanzania" elevation="350" location="Muheza District" municipality="East Usambara Mts" pageId="83" pageNumber="84" specimenCode="NHMD 621759" specimenCount="1" specimenCount-male="1" stateProvince="Tanga Region" typeStatus="holotype">
<emphasis id="B92C313DD6760664FF19F9E4C470F9F9" bold="true" box="[234,348,1543,1569]" pageId="83" pageNumber="84">
<typeStatus id="54E3538DD6760664FF19F9E4C470F9F9" box="[234,348,1543,1569]" pageId="83" pageNumber="84" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F34FADBFD6760664FF4EF9C8C47EF99D" box="[189,338,1579,1605]" name="Tanzania" pageId="83" pageNumber="84">TANZANIA</collectingCountry>
<specimenCount id="9D5E26A6D6760664FE9FF9C8C4A9F99D" box="[364,389,1579,1605]" pageId="83" pageNumber="84" type="male"></specimenCount>
;
<collectingMunicipality id="6B837755D6760664FE60F9C8C751F99D" box="[403,637,1578,1605]" pageId="83" pageNumber="84">East Usambara Mts</collectingMunicipality>
,
<collectingRegion id="499C23CDD6760664FD79F9C8C61EF99C" box="[650,818,1578,1605]" country="Tanzania" name="Tanga" pageId="83" pageNumber="84">Tanga Region</collectingRegion>
,
<location id="8E87BBF4D6760664FCCCF9C9C12DF99C" LSID="urn:lsid:plazi:treatment:03F15C39D6760661FDA8FC39C048FA79:8E87BBF4D6760664FCCCF9C9C12DF99C" box="[831,1025,1578,1604]" country="Tanzania" municipality="East Usambara Mts" name="Muheza District" pageId="83" pageNumber="84" stateProvince="Tanga Region">Muheza District</location>
,
<collectorName id="26AD88F9D6760664FBFDF9C9C031F99D" box="[1038,1309,1578,1605]" pageId="83" pageNumber="84">Mgambo Proposed FR</collectorName>
;
<elevation id="00750A1CD6760664FAD8F9C9C5DAF9B0" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="83" pageNumber="84" unit="m" value="350.0">
<quantity id="4CA040CAD6760664FAD8F9C9C05BF99D" box="[1323,1399,1578,1605]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="83" pageNumber="84" unit="m" value="350.0">350 m</quantity>
a.s.l.
</elevation>
;
<date id="FFE6CBEFD6760664FEF0F9ACC450F9B0" box="[259,380,1614,1641]" pageId="83" pageNumber="84" value="1996-09">
<collectingDate id="EFA23207D6760664FEF0F9ACC450F9B0" box="[259,380,1614,1641]" pageId="83" pageNumber="84" value="1996-09">Sep. 1996</collectingDate>
</date>
;
<collectorName id="26AD88F9D6760664FE7BF9ADC77EF9B0" box="[392,594,1614,1640]" pageId="83" pageNumber="84">Norbert Cordeiro</collectorName>
leg.;
<specimenCode id="DBFE4554D6760664FD66F9ACC67EF9B1" box="[661,850,1614,1641]" collectionCode="NHMD" pageId="83" pageNumber="84">NHMD 621759</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BE7ED2FD6760664FF19F972C130F916" blockId="83.[189,1052,1681,1743]" pageId="83" pageNumber="84">
<materialsCitation id="3B30E772D6760664FF19F972C134F916" ID-GBIF-Occurrence="3712026351" collectingDate="1996-09" collectionCode="NHMD" collectorName="Mgambo Proposed FR &amp; Norbert Cordeiro" country="Tanzania" elevation="350" location="TANZANIA" municipality="East Usambara Mts" pageId="83" pageNumber="84" specimenCode="NHMD 621760" specimenCount="1" specimenCount-male="1" stateProvince="Tanga" typeStatus="paratype">
<emphasis id="B92C313DD6760664FF19F972C470F973" bold="true" box="[234,348,1681,1707]" pageId="83" pageNumber="84">
<typeStatus id="54E3538DD6760664FF19F972C470F973" box="[234,348,1681,1707]" pageId="83" pageNumber="84" type="paratype">Paratype</typeStatus>
</emphasis>
<collectingCountry id="F34FADBFD6760664FF4EF956C47EF917" box="[189,338,1717,1743]" name="Tanzania" pageId="83" pageNumber="84">TANZANIA</collectingCountry>
<specimenCount id="9D5E26A6D6760664FE9AF956C4B4F917" box="[361,408,1717,1743]" pageId="83" pageNumber="84" type="male">1 ♂</specimenCount>
; same collection data as for holotype;
<specimenCode id="DBFE4554D6760664FCA8F956C134F916" box="[859,1048,1716,1743]" collectionCode="NHMD" pageId="83" pageNumber="84">NHMD 621760</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C342BEA4D6760661FF4EF915C477FAC8" lastPageId="86" lastPageNumber="87" pageId="83" pageNumber="84" type="description">
<paragraph id="8BE7ED2FD6760664FF4EF915C48EF8CA" blockId="83.[189,418,1782,1810]" box="[189,418,1782,1810]" pageId="83" pageNumber="84">
<emphasis id="B92C313DD6760664FF4EF915C462F8C8" bold="true" box="[189,334,1782,1808]" pageId="83" pageNumber="84">Description</emphasis>
(male)
</paragraph>
<paragraph id="8BE7ED2FD6760664FF4EF8C5C05BF898" blockId="83.[189,1399,1830,1857]" box="[189,1399,1830,1857]" pageId="83" pageNumber="84">
SIZE. Length
<quantity id="4CA040CAD6760664FEA5F8C5C4FBF899" box="[342,471,1830,1857]" metricMagnitude="-2" metricUnit="m" metricValue="5.65" metricValueMax="5.7" metricValueMin="5.6" pageId="83" pageNumber="84" unit="mm" value="56.5" valueMax="57.0" valueMin="56.0">5657 mm</quantity>
, diameter 2.93.0 mm, 7071 podous rings, no apodous rings in front of telson.
</paragraph>
<paragraph id="8BE7ED2FD6760664FF4EF88BC42BF87E" blockId="83.[189,1399,1896,1958]" pageId="83" pageNumber="84">COLOUR. Strongly faded after 22 years in alcohol, but faint indications of a medium-broad light dorsal stripe.</paragraph>
<paragraph id="8BE7ED2FD6760664FF4EF82DC4CDF831" blockId="83.[189,481,1998,2025]" box="[189,481,1998,2025]" pageId="83" pageNumber="84">SUPRALABRAL SETAE. 45.</paragraph>
<caption id="DF27BDA7D6710663FF4EF91FC73DF832" ID-DOI="http://doi.org/10.5281/zenodo.6359231" ID-Zenodo-Dep="6359231" httpUri="https://zenodo.org/record/6359231/files/figure.png" pageId="84" pageNumber="85" startId="84.[189,232,1788,1814]" targetBox="[260,1328,266,1762]" targetPageId="84">
<paragraph id="8BE7ED2FD6710663FF4EF91FC73DF832" blockId="84.[185,1399,1788,2026]" pageId="84" pageNumber="85">
<emphasis id="B92C313DD6710663FF4EF91FC432F8CE" bold="true" box="[189,286,1788,1814]" pageId="84" pageNumber="85">Fig. 52.</emphasis>
<taxonomicName id="4C5896ACD6710663FEDAF91FC768F8CE" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[297,580,1788,1814]" class="Diplopoda" family="Odontopygidae" genus="Raduliverpa" kingdom="Animalia" order="Spirostreptida" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="donatellae" status="sp. nov.">
<emphasis id="B92C313DD6710663FEDAF91FC768F8CE" box="[297,580,1788,1814]" italics="true" pageId="84" pageNumber="85">Raduliverpa donatellae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F8C46D6710663FDA3F91EC783F8CF" box="[592,687,1789,1815]" pageId="84" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B92C313DD6710663FD49F91FC7DFF8CE" bold="true" box="[698,755,1788,1814]" pageId="84" pageNumber="85">AC</emphasis>
. Holotype, ♂ (NHMD 621759). Left gonopod coxa. A. posterior view. B. Mesal view. C. Anterior view.
<emphasis id="B92C313DD6710663FCB6F8C3C676F8E2" bold="true" box="[837,858,1824,1850]" pageId="84" pageNumber="85">D</emphasis>
. Midbody dorsal limbus.
<emphasis id="B92C313DD6710663FB69F8FCC1FFF8E1" bold="true" box="[1178,1235,1823,1849]" pageId="84" pageNumber="85">EG</emphasis>
. Paratype, ♂ (NHMD 621760), first and second pair of legs (right first legs misshapen/diminished). E. Sublateral view. F. Anterior view. G. Ventral view. Abbreviations:
<emphasis id="B92C313DD6710663FCFBF885C617F858" box="[776,827,1894,1920]" italics="true" pageId="84" pageNumber="85">APS</emphasis>
= mesapical prefemoral setae;
<emphasis id="B92C313DD6710663FB6CF885C1FFF858" box="[1183,1235,1894,1920]" italics="true" pageId="84" pageNumber="85">CXS</emphasis>
=coxosternal setae;
<emphasis id="B92C313DD6710663FEF7F869C41BF87C" box="[260,311,1930,1956]" italics="true" pageId="84" pageNumber="85">LCS</emphasis>
= lateral coxal process;
<emphasis id="B92C313DD6710663FDB3F869C75DF87C" box="[576,625,1930,1956]" italics="true" pageId="84" pageNumber="85">LPS</emphasis>
= lateral prefemoral setae;
<emphasis id="B92C313DD6710663FC56F869C6FCF87B" box="[933,976,1930,1955]" italics="true" pageId="84" pageNumber="85">MF</emphasis>
= metaplical flange;
<emphasis id="B92C313DD6710663FB49F869C1C9F87B" box="[1210,1253,1930,1955]" italics="true" pageId="84" pageNumber="85">MP</emphasis>
=metaplica;
<emphasis id="B92C313DD6710663FF4AF84EC5C5F81E" box="[185,233,1965,1990]" italics="true" pageId="84" pageNumber="85">msp</emphasis>
= metaplical spinelike process;
<emphasis id="B92C313DD6710663FD93F84EC7A0F81E" box="[608,652,1965,1990]" italics="true" pageId="84" pageNumber="85">mtp</emphasis>
=metaplical triangular process;
<emphasis id="B92C313DD6710663FBF0F84EC10BF81E" box="[1027,1063,1965,1990]" italics="true" pageId="84" pageNumber="85">PP</emphasis>
= proplica. Scale bars: AC, EG =0.1 mm; D =0.01 mm.
</paragraph>
</caption>
<caption id="DF27BDA7D6700662FF4EF959C75EF85D" ID-DOI="http://doi.org/10.5281/zenodo.6359233" ID-Zenodo-Dep="6359233" httpUri="https://zenodo.org/record/6359233/files/figure.png" pageId="85" pageNumber="86" startId="85.[189,232,1722,1748]" targetBox="[189,1398,265,1669]" targetPageId="85">
<paragraph id="8BE7ED2FD6700662FF4EF959C75EF85D" blockId="85.[185,1399,1722,1925]" pageId="85" pageNumber="86">
<emphasis id="B92C313DD6700662FF4EF959C436F90C" bold="true" box="[189,282,1722,1748]" pageId="85" pageNumber="86">Fig. 53.</emphasis>
<taxonomicName id="4C5896ACD6700662FED2F959C714F90C" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[289,568,1722,1748]" class="Diplopoda" family="Odontopygidae" genus="Raduliverpa" kingdom="Animalia" order="Spirostreptida" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="donatellae" status="sp. nov.">
<emphasis id="B92C313DD6700662FED2F959C714F90C" box="[289,568,1722,1748]" italics="true" pageId="85" pageNumber="86">Raduliverpa donatellae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F8C46D6700662FDCCF958C7B0F90D" box="[575,668,1723,1749]" pageId="85" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (NHMD 621759), gonopod telopodites.
<emphasis id="B92C313DD6700662FAE2F959C00AF90C" bold="true" box="[1297,1318,1722,1748]" pageId="85" pageNumber="86">A</emphasis>
. Right telopodite, posterior view.
<emphasis id="B92C313DD6700662FE0EF93DC723F920" bold="true" box="[509,527,1758,1784]" pageId="85" pageNumber="86">B</emphasis>
. Left telopodite, subanterior view.
<emphasis id="B92C313DD6700662FC46F93EC6E5F92F" bold="true" box="[949,969,1757,1783]" pageId="85" pageNumber="86">C</emphasis>
. Right telopodite, disto-mesal-basal view.
<emphasis id="B92C313DD6700662FEF4F8E2C430F8C3" bold="true" box="[263,284,1793,1819]" pageId="85" pageNumber="86">D</emphasis>
. Close-up of right solenomere tip (also with part of post-torsal spine) to show the genuscharacteristic striation.
<emphasis id="B92C313DD6700662FE25F8C7C4CBF8E6" bold="true" box="[470,487,1828,1854]" pageId="85" pageNumber="86">E</emphasis>
. Left telopodite, anterior-basal view. Abbreviations:
<emphasis id="B92C313DD6700662FB97F8C7C1B9F8E6" box="[1124,1173,1828,1854]" italics="true" pageId="85" pageNumber="86">PTS</emphasis>
=post-torsal spine;
<emphasis id="B92C313DD6700662FF4AF8A4C5DDF8B9" box="[185,241,1863,1889]" italics="true" pageId="85" pageNumber="86">SLM</emphasis>
= solenomere;
<emphasis id="B92C313DD6700662FE50F8ABC4E8F8B9" box="[419,452,1864,1889]" italics="true" pageId="85" pageNumber="86">TL</emphasis>
=torsotope lobe;
<emphasis id="B92C313DD6700662FD6AF8ABC7EEF8B9" box="[665,706,1864,1889]" italics="true" pageId="85" pageNumber="86">TM</emphasis>
= telomere;
<emphasis id="B92C313DD6700662FCA0F8A4C655F8B9" box="[851,889,1863,1889]" italics="true" pageId="85" pageNumber="86">tlm</emphasis>
= telomere lamellae;
<emphasis id="B92C313DD6700662FB8CF8ABC18CF8B9" box="[1151,1184,1864,1889]" italics="true" pageId="85" pageNumber="86">TT</emphasis>
=torsotope. Scale bars: AC, E =0.1 mm; D =0.02 mm.
</paragraph>
</caption>
<paragraph id="8BE7ED2FD6730661FF4EFEEAC4F7FE9F" blockId="86.[189,1399,265,327]" pageId="86" pageNumber="87">MANDIBULAR STIPES. Ventrodistally produced in strong triangular, slightly mesad process, distal margin very shallowly concave.</paragraph>
<paragraph id="8BE7ED2FD6730661FF4EFE8EC7B2FE71" blockId="86.[189,1398,365,426]" pageId="86" pageNumber="87">ANAL VALVES. With with moderate dorsal spine, a protruding ventral corner, margins raised, each with three setae on poorly developed ravelins.</paragraph>
<paragraph id="8BE7ED2FD6730661FF4EFE32C65AFE32" blockId="86.[189,886,464,491]" box="[189,886,464,491]" pageId="86" pageNumber="87">
LIMBUS (
<figureCitation id="1363F1AAD6730661FED2FE33C4A5FE33" box="[289,393,464,491]" captionStart="Fig" captionStartId="84.[189,232,1788,1814]" captionTargetBox="[260,1328,266,1762]" captionTargetId="figure-117@84.[260,1328,266,1762]" captionTargetPageId="84" captionText="Fig. 52. Raduliverpa donatellae sp. nov. AC. Holotype, ♂ (NHMD 621759). Left gonopod coxa. A. posterior view. B. Mesal view. C. Anterior view. D. Midbody dorsal limbus. EG. Paratype, ♂ (NHMD 621760), first and second pair of legs (right first legs misshapen/diminished). E. Sublateral view. F. Anterior view. G. Ventral view. Abbreviations: APS= mesapical prefemoral setae; CXS =coxosternal setae; LCS= lateral coxalprocess;LPS = lateral prefemoralsetae;MF= metaplical flange; MP=metaplica; msp = metaplical spinelike process; mtp =metaplical triangular process; PP = proplica. Scale bars: AC, EG =0.1 mm; D =0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.6359231" httpUri="https://zenodo.org/record/6359231/files/figure.png" pageId="86" pageNumber="87">Fig. 52D</figureCitation>
). Striate, margin with very shallow lobes.
</paragraph>
<paragraph id="8BE7ED2FD6730661FF4EFDF2C64BFD95" blockId="86.[189,1398,529,589]" pageId="86" pageNumber="87">LEGS. With postfemoral and tibial pads, covering whole length of podomeres, from leg-pair ca 6, diminishing and eventually disappearing toward hind end.</paragraph>
<paragraph id="8BE7ED2FD6730661FF4EFD97C181FD0B" blockId="86.[189,1399,628,724]" pageId="86" pageNumber="87">
FIRST PAIR OF LEGS (
<figureCitation id="1363F1AAD6730661FE52FD97C71CFD57" box="[417,560,628,655]" captionStart="Fig" captionStartId="84.[189,232,1788,1814]" captionTargetBox="[260,1328,266,1762]" captionTargetId="figure-117@84.[260,1328,266,1762]" captionTargetPageId="84" captionText="Fig. 52. Raduliverpa donatellae sp. nov. AC. Holotype, ♂ (NHMD 621759). Left gonopod coxa. A. posterior view. B. Mesal view. C. Anterior view. D. Midbody dorsal limbus. EG. Paratype, ♂ (NHMD 621760), first and second pair of legs (right first legs misshapen/diminished). E. Sublateral view. F. Anterior view. G. Ventral view. Abbreviations: APS= mesapical prefemoral setae; CXS =coxosternal setae; LCS= lateral coxalprocess;LPS = lateral prefemoralsetae;MF= metaplical flange; MP=metaplica; msp = metaplical spinelike process; mtp =metaplical triangular process; PP = proplica. Scale bars: AC, EG =0.1 mm; D =0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.6359231" httpUri="https://zenodo.org/record/6359231/files/figure.png" pageId="86" pageNumber="87">Fig. 52EG</figureCitation>
). Prefemoral lobes with rounded-triangular in ventral view. Five to six coxosternal seta (
<emphasis id="B92C313DD6730661FE7DFD74C4EEFD69" box="[398,450,663,689]" italics="true" pageId="86" pageNumber="87">CXS</emphasis>
) close to lateral margin of coxosternum, well separated from prefemoral lobes. Prefemora with 2 of short mesapical setae (
<emphasis id="B92C313DD6730661FD49FD5AC7C1FD0B" box="[698,749,697,723]" italics="true" pageId="86" pageNumber="87">APS</emphasis>
) and 512 peglike lateral setae (
<emphasis id="B92C313DD6730661FB98FD5AC1B0FD0B" box="[1131,1180,697,723]" italics="true" pageId="86" pageNumber="87">LPS</emphasis>
).
</paragraph>
<paragraph id="8BE7ED2FD6730661FF4EFD19C6ABFCCC" blockId="86.[189,903,761,788]" box="[189,903,761,788]" pageId="86" pageNumber="87">STERNUM 9. Transversely rounded-triangular (not illustrated).</paragraph>
<paragraph id="8BE7ED2FD6730661FF4EFCD9C47EFBD8" blockId="86.[189,1398,826,1024]" pageId="86" pageNumber="87">
GONOPOD COXA (
<figureCitation id="1363F1AAD6730661FE76FCD9C736FC8D" box="[389,538,826,853]" captionStart="Fig" captionStartId="84.[189,232,1788,1814]" captionTargetBox="[260,1328,266,1762]" captionTargetId="figure-117@84.[260,1328,266,1762]" captionTargetPageId="84" captionText="Fig. 52. Raduliverpa donatellae sp. nov. AC. Holotype, ♂ (NHMD 621759). Left gonopod coxa. A. posterior view. B. Mesal view. C. Anterior view. D. Midbody dorsal limbus. EG. Paratype, ♂ (NHMD 621760), first and second pair of legs (right first legs misshapen/diminished). E. Sublateral view. F. Anterior view. G. Ventral view. Abbreviations: APS= mesapical prefemoral setae; CXS =coxosternal setae; LCS= lateral coxalprocess;LPS = lateral prefemoralsetae;MF= metaplical flange; MP=metaplica; msp = metaplical spinelike process; mtp =metaplical triangular process; PP = proplica. Scale bars: AC, EG =0.1 mm; D =0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.6359231" httpUri="https://zenodo.org/record/6359231/files/figure.png" pageId="86" pageNumber="87">Fig. 52AC</figureCitation>
). Slender, straight, almost parallel-sided, distolaterally with an almost equilateral triangular lateral process (
<emphasis id="B92C313DD6730661FD84FCBEC786FCAF" box="[631,682,861,887]" italics="true" pageId="86" pageNumber="87">LCS</emphasis>
). Proplica (
<emphasis id="B92C313DD6730661FCC4FCBEC677FCAE" box="[823,859,861,886]" italics="true" pageId="86" pageNumber="87">PP</emphasis>
) simple, proplical lobe hibben behind fold of metaplica. Metaplica (
<emphasis id="B92C313DD6730661FE3BFC63C4DEFC41" box="[456,498,896,921]" italics="true" pageId="86" pageNumber="87">MP</emphasis>
) with a poorly delimited mesad flange (
<emphasis id="B92C313DD6730661FC24FC63C12DFC41" box="[983,1025,896,921]" italics="true" pageId="86" pageNumber="87">MF</emphasis>
) basally, ca at mid-length with small, mesad triangular, process (
<emphasis id="B92C313DD6730661FDA0FC41C753FC63" box="[595,639,930,955]" italics="true" pageId="86" pageNumber="87">mtp</emphasis>
), distally with long, slender, strongly curved spinelike process (
<emphasis id="B92C313DD6730661FF34FC26C5DAFC06" box="[199,246,965,990]" italics="true" pageId="86" pageNumber="87">msp</emphasis>
), process first directed basad then making a smooth 180° turn and projecting distad beyond gonopod tip.
</paragraph>
<paragraph id="8BE7ED2FD6730661FF4EFBC4C477FAC8" blockId="86.[189,1399,1063,1296]" pageId="86" pageNumber="87">
GONOPOD TELOPODITE (
<figureCitation id="1363F1AAD6730661FE4EFBC4C73FFB9A" box="[445,531,1063,1090]" captionStart="Fig" captionStartId="85.[189,232,1722,1748]" captionTargetBox="[189,1398,265,1669]" captionTargetId="figure-99@85.[189,1398,265,1669]" captionTargetPageId="85" captionText="Fig. 53. Raduliverpa donatellae sp. nov., holotype, ♂ (NHMD 621759), gonopod telopodites. A. Right telopodite, posterior view. B. Left telopodite, subanterior view. C. Right telopodite, disto-mesal-basal view. D. Close-up of right solenomere tip (also with part of post-torsal spine) to show the genuscharacteristic striation. E. Left telopodite, anterior-basal view. Abbreviations: PTS=post-torsal spine; SLM = solenomere; TL =torsotope lobe; TM = telomere; tlm = telomere lamellae; TT =torsotope. Scale bars: AC, E =0.1 mm; D =0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.6359233" httpUri="https://zenodo.org/record/6359233/files/figure.png" pageId="86" pageNumber="87">Fig. 53</figureCitation>
). Arculus 80º. Torsotope (
<emphasis id="B92C313DD6730661FCB7FBCBC649FB99" box="[836,869,1064,1089]" italics="true" pageId="86" pageNumber="87">
<collectingRegion id="499C23CDD6730661FCB7FBCBC649FB99" box="[836,869,1064,1089]" country="Mozambique" name="Tete" pageId="86" pageNumber="87">TT</collectingRegion>
</emphasis>
) extended, with angular lobe (
<emphasis id="B92C313DD6730661FB39FBCBC1C7FB99" box="[1226,1259,1064,1089]" italics="true" pageId="86" pageNumber="87">TL</emphasis>
) fitting into torsion. A long, stout post-torsal spine (
<emphasis id="B92C313DD6730661FD61FBA9C7EFFBBC" box="[658,707,1098,1124]" italics="true" pageId="86" pageNumber="87">PTS</emphasis>
) making a full turn around post-torsal narrowing which is hence not evident. Solenomere and telomere originating near mesalmost part of telopodite, flexed back in front of and distal to torsotope. Solenomere (
<emphasis id="B92C313DD6730661FD1AFB6CC60CFB71" box="[745,800,1167,1193]" italics="true" pageId="86" pageNumber="87">SLM</emphasis>
) not very long, relatively stout, partially concealed within telomere, terminally flattened, pointed and transversely striate (
<figureCitation id="1363F1AAD6730661FC1FFB52C174FB14" box="[1004,1112,1201,1228]" captionStart="Fig" captionStartId="85.[189,232,1722,1748]" captionTargetBox="[189,1398,265,1669]" captionTargetId="figure-99@85.[189,1398,265,1669]" captionTargetPageId="85" captionText="Fig. 53. Raduliverpa donatellae sp. nov., holotype, ♂ (NHMD 621759), gonopod telopodites. A. Right telopodite, posterior view. B. Left telopodite, subanterior view. C. Right telopodite, disto-mesal-basal view. D. Close-up of right solenomere tip (also with part of post-torsal spine) to show the genuscharacteristic striation. E. Left telopodite, anterior-basal view. Abbreviations: PTS=post-torsal spine; SLM = solenomere; TL =torsotope lobe; TM = telomere; tlm = telomere lamellae; TT =torsotope. Scale bars: AC, E =0.1 mm; D =0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.6359233" httpUri="https://zenodo.org/record/6359233/files/figure.png" pageId="86" pageNumber="87">Fig. 53D</figureCitation>
). Telomere (TM) shaped like an irregular bowl, smooth on the outside and along margins, with several contorted lamellae (
<emphasis id="B92C313DD6730661FAB4FB30C040FB35" box="[1351,1388,1235,1261]" italics="true" pageId="86" pageNumber="87">tml</emphasis>
) on the inside.
</paragraph>
</subSubSection>
<subSubSection id="C342BEA4D6730661FF4EFAD5C048FA79" pageId="86" pageNumber="87" type="distribution">
<paragraph id="8BE7ED2FD6730661FF4EFAD5C4C2FA88" blockId="86.[189,494,1334,1360]" box="[189,494,1334,1360]" pageId="86" pageNumber="87">
<heading id="D0AF5A43D6730661FF4EFAD5C4C2FA88" bold="true" box="[189,494,1334,1360]" fontSize="11" level="3" pageId="86" pageNumber="87" reason="3">
<emphasis id="B92C313DD6730661FF4EFAD5C4C2FA88" bold="true" box="[189,494,1334,1360]" pageId="86" pageNumber="87">Distribution and habitat</emphasis>
</heading>
</paragraph>
<paragraph id="8BE7ED2FD6730661FF4EFA86C048FA79" blockId="86.[189,1399,1380,1441]" pageId="86" pageNumber="87">
Known only from Mgambo FR (formerly: Mgambo Proposed forest Reserve) in the north-east of the East Usambara Mts, altitude
<quantity id="4CA040CAD6730661FDE0FA65C771FA79" box="[531,605,1414,1441]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="86" pageNumber="87" unit="m" value="350.0">350 m</quantity>
a.s.l. See
<bibRefCitation id="EFC990DED6730661FD21FA65C698FA79" author="Oliver S. A. &amp; Bracebridge C. E. &amp; Fanning E. &amp; Howell K. M. &amp; Technical Paper No." box="[722,948,1414,1441]" pageId="86" pageNumber="87" refId="ref49055" refString="Oliver S. A., Bracebridge C. E., Fanning E. &amp; Howell K. M. (eds) 2002. Mgambo Forest Reserve: A Biodiversity Survey. East Usambara Conservation Area Management Programme, Technical Paper No. 59. Frontier Tanzania; Forestry and Beekeeping Division, Dar es Salaam, Tanzania and Indufor / Metsahallitus Group, Vantaa, Finland. Available from https: // www. yumpu. com / en / document / view / 8227386 / mgambo-forest-reserve-abiodiversity-survey-eastern-arc- [accessed 22 Feb. 2022]." type="url" year="2002">
Oliver
<emphasis id="B92C313DD6730661FCD7FA64C673FA78" box="[804,863,1414,1440]" italics="true" pageId="86" pageNumber="87">et al.</emphasis>
(2002)
</bibRefCitation>
for information on the Mgambo FR.
</paragraph>
</subSubSection>
</treatment>
</document>