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<document id="562156995DED2947D854D5DA2EDBFA3A" ID-CLB-Dataset="20167" ID-DOI="10.5852/ejt.2022.803.1691" ID-GBIF-Dataset="af09c8e1-b481-4de0-b9d0-83ba26bf9876" ID-ISSN="2118-9773" ID-Zenodo-Dep="6359066" ID-ZooBank="8B66C8AE-F00A-42F6-9641-26B0ECC49F78" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1647350252755" checkinUser="felipe" docAuthor="Enghoff, Henrik" docDate="2022" docId="03F15C39D6440655FD9CFAD6C1F5FDFB" docLanguage="en" docName="ejt-803.1-136.pdf" docOrigin="European Journal of Taxonomy 803" docSource="http://dx.doi.org/10.5852/ejt.2022.803.1691" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Uncodrama Enghoff 2022, gen. nov." docType="treatment" docUuid="8ECD78AE-280D-4480-A354-E0C15CFB024F" docUuidSource="ZooBank" docVersion="7" lastPageNumber="99" masterDocId="FFC82441D6250637FFF3FFE3C52CFFD8" masterDocTitle="Mountains of millipedes. The family Odontopygidae in the Eastern Arc Mountains of Tanzania (Diplopoda, Spirostreptida)" masterLastPageNumber="136" masterPageNumber="1" pageNumber="98" updateTime="1699318071475" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="D080260DF90A10B29660E77DD55B6FCF">Mountains of millipedes. The family Odontopygidae in the Eastern Arc Mountains of Tanzania (Diplopoda, Spirostreptida)</mods:title>
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<mods:namePart id="F4C0738A6B06A7D1A8ABD6899EA65BF2">Enghoff, Henrik</mods:namePart>
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<paragraph id="8BE7ED2FD6440656FD9CFAD6C6E9FA97" blockId="97.[363,1224,1332,1396]" box="[623,965,1332,1359]" pageId="97" pageNumber="98">
<heading id="D0AF5A43D6440656FD9CFAD6C6E9FA97" box="[623,965,1332,1359]" centered="true" fontSize="11" level="2" pageId="97" pageNumber="98" reason="2">
Genus
<taxonomicName id="4C5896ACD6440656FD31FAD7C67FFA97" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[706,851,1332,1359]" class="Diplopoda" family="Odontopygidae" genus="Uncodrama" kingdom="Animalia" order="Spirostreptida" pageId="97" pageNumber="98" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B92C313DD6440656FD31FAD7C67FFA97" bold="true" box="[706,851,1332,1359]" italics="true" pageId="97" pageNumber="98">Uncodrama</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F8C46D6440656FCA9FAD6C6E9FA97" box="[858,965,1333,1359]" pageId="97" pageNumber="98" rank="genus">gen. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C342BEA4D6440656FE98FABAC1E4FAAC" box="[363,1224,1369,1396]" pageId="97" pageNumber="98" type="description">
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<uri id="FFC9E12DD6440656FE98FABAC1E4FAAC" box="[363,1224,1369,1396]" pageId="97" pageNumber="98">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
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<subSubSection id="C342BEA4D6440656FF4EFA78C75DFA3D" pageId="97" pageNumber="98" type="type_taxon">
<paragraph id="8BE7ED2FD6440656FF4EFA78C474FA6D" blockId="97.[189,344,1435,1461]" box="[189,344,1435,1461]" pageId="97" pageNumber="98">
<heading id="D0AF5A43D6440656FF4EFA78C474FA6D" bold="true" box="[189,344,1435,1461]" fontSize="11" level="3" pageId="97" pageNumber="98" reason="3">
<emphasis id="B92C313DD6440656FF4EFA78C474FA6D" bold="true" box="[189,344,1435,1461]" pageId="97" pageNumber="98">
<typeStatus id="54E3538DD6440656FF4EFA78C5D6FA6D" box="[189,250,1435,1461]" pageId="97" pageNumber="98">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE7ED2FD6440656FF4EFA28C75DFA3D" blockId="97.[189,625,1483,1509]" box="[189,625,1483,1509]" pageId="97" pageNumber="98">
<taxonomicName id="4C5896ACD6440656FF4EFA28C495FA3C" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[189,441,1483,1509]" class="Diplopoda" family="Odontopygidae" genus="Uncodrama" kingdom="Animalia" order="Spirostreptida" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="coronata" status="gen. et sp. nov.">
<emphasis id="B92C313DD6440656FF4EFA28C495FA3C" box="[189,441,1483,1509]" italics="true" pageId="97" pageNumber="98">Uncodrama coronata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F8C46D6440656FE32FA28C75DFA3D" box="[449,625,1483,1509]" pageId="97" pageNumber="98" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="C342BEA4D6440656FF4EF9EEC1C3F941" pageId="97" pageNumber="98" type="reference_group">
<paragraph id="8BE7ED2FD6440656FF4EF9EEC7ABF9FF" blockId="97.[189,647,1549,1575]" box="[189,647,1549,1575]" pageId="97" pageNumber="98">
<heading id="D0AF5A43D6440656FF4EF9EEC7ABF9FF" bold="true" box="[189,647,1549,1575]" fontSize="11" level="3" pageId="97" pageNumber="98" reason="3">
<emphasis id="B92C313DD6440656FF4EF9EEC7ABF9FF" bold="true" box="[189,647,1549,1575]" pageId="97" pageNumber="98">Other provisionally included species</emphasis>
</heading>
</paragraph>
<paragraph id="8BE7ED2FD6440656FF4EF9DFC02BF98F" blockId="97.[189,1287,1596,1623]" box="[189,1287,1596,1623]" pageId="97" pageNumber="98">
<taxonomicName id="4C5896ACD6440656FF4EF9DFC79EF98F" authority="(Chamberlin, 1927)" authorityName="Enghoff" authorityYear="2022" baseAuthorityName="Chamberlin" baseAuthorityYear="1927" box="[189,690,1596,1623]" class="Diplopoda" family="Odontopygidae" genus="Uncodrama" kingdom="Animalia" order="Spirostreptida" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="medjensis" status="comb. nov.">
<emphasis id="B92C313DD6440656FF4EF9DFC4E8F98E" box="[189,452,1596,1622]" italics="true" pageId="97" pageNumber="98">Uncodrama medjensis</emphasis>
(
<bibRefCitation id="EFC990DED6440656FE20F9DFC786F98F" author="Chamberlin R. V." box="[467,682,1596,1623]" pageId="97" pageNumber="98" pagination="177 - 249" refId="ref46758" refString="Chamberlin R. V. 1927. The Chilopoda and Diplopoda collected by the American Museum of Natural History Congo Expedition (1909 - 1915), with notes on some other African species. Bulletin of the American Museum of Natural History 57 (4): 177 - 249." type="journal article" year="1927">Chamberlin, 1927</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A21F8C46D6440656FD49F9DFC610F98F" box="[698,828,1596,1623]" pageId="97" pageNumber="98" rank="species">comb. nov.</taxonomicNameLabel>
ex
<emphasis id="B92C313DD6440656FC95F9DFC6D3F98E" box="[870,1023,1596,1622]" italics="true" pageId="97" pageNumber="98">Hapthysanus</emphasis>
Attems, 1909.
<collectingCountry id="F34FADBFD6440656FB42F9DEC02FF98F" box="[1201,1283,1597,1623]" name="Democratic Republic of the Congo" pageId="97" pageNumber="98">Congo</collectingCountry>
.
</paragraph>
<paragraph id="8BE7ED2FD6440656FF4EF99CC1C3F941" blockId="97.[189,1263,1663,1689]" box="[189,1263,1663,1689]" pageId="97" pageNumber="98">
<taxonomicName id="4C5896ACD6440656FF4EF99CC75EF941" authority="(Karsch, 1881)" authorityName="Enghoff" authorityYear="2022" baseAuthorityName="Karsch" baseAuthorityYear="1881" box="[189,626,1663,1689]" class="Diplopoda" family="Odontopygidae" genus="Uncodrama" kingdom="Animalia" order="Spirostreptida" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="tumidens" status="comb. nov.">
<emphasis id="B92C313DD6440656FF4EF99CC496F941" box="[189,442,1663,1689]" italics="true" pageId="97" pageNumber="98">Uncodrama tumidens</emphasis>
(Karsch, 1881)
</taxonomicName>
<taxonomicNameLabel id="A21F8C46D6440656FD8AF99CC7D7F941" box="[633,763,1663,1689]" pageId="97" pageNumber="98" rank="species">comb. nov.</taxonomicNameLabel>
ex.
<emphasis id="B92C313DD6440656FCDFF99CC6EAF941" box="[812,966,1663,1689]" italics="true" pageId="97" pageNumber="98">Spirostreptus</emphasis>
(
<taxonomicName id="4C5896ACD6440656FC24F99CC14AF941" ID-CoL="7PQP5" box="[983,1126,1663,1689]" class="Diplopoda" family="Odontopygidae" genus="Odontopyge" kingdom="Animalia" order="Spirostreptida" pageId="97" pageNumber="98" phylum="Arthropoda" rank="genus">
<emphasis id="B92C313DD6440656FC24F99CC14AF941" box="[983,1126,1663,1689]" italics="true" pageId="97" pageNumber="98">Odontopyge</emphasis>
</taxonomicName>
). S.
<collectingCountry id="F34FADBFD6440656FB6DF99CC1C7F941" box="[1182,1259,1663,1689]" name="Sudan" pageId="97" pageNumber="98">Sudan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C342BEA4D6440656FF4EF922C195F88A" pageId="97" pageNumber="98" type="diagnosis">
<paragraph id="8BE7ED2FD6440656FF4EF922C41AF903" blockId="97.[189,310,1729,1755]" box="[189,310,1729,1755]" pageId="97" pageNumber="98">
<heading id="D0AF5A43D6440656FF4EF922C41AF903" bold="true" box="[189,310,1729,1755]" fontSize="11" level="3" pageId="97" pageNumber="98" reason="3">
<emphasis id="B92C313DD6440656FF4EF922C41AF903" bold="true" box="[189,310,1729,1755]" pageId="97" pageNumber="98">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BE7ED2FD6440656FF4EF913C195F88A" blockId="97.[189,1399,1776,1875]" pageId="97" pageNumber="98">
Differs from other genera of
<taxonomicName id="4C5896ACD6440656FDE9F913C7ECF8D2" authorityName="Hoffman" authorityYear="1991" box="[538,704,1776,1802]" family="Odontopygidae" kingdom="Animalia" order="Spirostreptida" pageId="97" pageNumber="14" rank="tribe" tribe="Prionopetalini">Prionopetalini</taxonomicName>
by the combination of a subrectangular distal lobe on the proplica, a wide mesal opening between pro- and metaplica (gonocoel), a simple whiplike solenomere and a simple, twisted boat-shaped telomere with a distal, complicated antlerlike lobe.
</paragraph>
</subSubSection>
<subSubSection id="C342BEA4D6440656FF4EF898C7F0F830" pageId="97" pageNumber="98" type="etymology">
<paragraph id="8BE7ED2FD6440656FF4EF898C468F84D" blockId="97.[189,324,1915,1941]" box="[189,324,1915,1941]" pageId="97" pageNumber="98">
<heading id="D0AF5A43D6440656FF4EF898C468F84D" bold="true" box="[189,324,1915,1941]" fontSize="11" level="3" pageId="97" pageNumber="98" reason="3">
<emphasis id="B92C313DD6440656FF4EF898C468F84D" bold="true" box="[189,324,1915,1941]" pageId="97" pageNumber="98">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BE7ED2FD6440656FF4EF849C7F0F830" blockId="97.[189,1399,1962,2025]" pageId="97" pageNumber="98">
The genus name (gender feminine) is an anagram of
<emphasis id="B92C313DD6440656FCDAF848C699F81C" box="[809,949,1963,1988]" italics="true" pageId="97" pageNumber="98">Damacornu</emphasis>
Enghoff, 2018, another genus with an antlerlike structure on the gonopod telopodite.
</paragraph>
</subSubSection>
<subSubSection id="C342BEA4D6470655FF4EFEEAC1F5FDFB" pageId="98" pageNumber="99" type="discussion">
<paragraph id="8BE7ED2FD6470655FF4EFEEAC403FEFB" blockId="98.[189,303,265,291]" box="[189,303,265,291]" pageId="98" pageNumber="99">
<heading id="D0AF5A43D6470655FF4EFEEAC403FEFB" bold="true" box="[189,303,265,291]" fontSize="11" level="3" pageId="98" pageNumber="99" reason="3">
<emphasis id="B92C313DD6470655FF4EFEEAC403FEFB" bold="true" box="[189,303,265,291]" pageId="98" pageNumber="99">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BE7ED2FD6470655FF4EFEDBC1F5FDFB" blockId="98.[189,1399,311,547]" pageId="98" pageNumber="99">
In Kraus (1966) key to genera of “
<taxonomicName id="4C5896ACD6470655FD81FED4C61CFE89" authorityName="Attems" authorityYear="1909" box="[626,816,311,337]" pageId="98" pageNumber="13" rank="subFamily" subFamily="Odontopyginae">Odontopyginae</taxonomicName>
”,
<taxonomicName id="4C5896ACD6470655FCBAFED4C166FE89" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[841,1098,311,337]" class="Diplopoda" family="Odontopygidae" genus="Uncodrama" kingdom="Animalia" order="Spirostreptida" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="coronata" status="gen. et sp. nov.">
<emphasis id="B92C313DD6470655FCBAFED4C166FE89" box="[841,1098,311,337]" italics="true" pageId="98" pageNumber="99">Uncodrama coronata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F8C46D6470655FBA6FEDBC03DFE8A" box="[1109,1297,312,338]" pageId="98" pageNumber="99" rank="species">gen. et sp. nov.</taxonomicNameLabel>
runs all the way through to
<taxonomicName id="4C5896ACD6470655FE42FEB9C76CFEAC" box="[433,576,346,372]" class="Diplopoda" family="Odontopygidae" genus="Odontopyge" kingdom="Animalia" order="Spirostreptida" pageId="98" pageNumber="99" phylum="Arthropoda" rank="genus">
<emphasis id="B92C313DD6470655FE42FEB9C76CFEAC" box="[433,576,346,372]" italics="true" pageId="98" pageNumber="99">Odontopyge</emphasis>
</taxonomicName>
, a name which was mis-applied until
<bibRefCitation id="EFC990DED6470655FBEAFEB9C1F0FEAC" author="Hoffman R. L." box="[1049,1244,346,372]" pageId="98" pageNumber="99" pagination="65 - 73" refId="ref48407" refString="Hoffman R. L. 1991. What is Odontopyge? A solution to a long-standing nomenclatorial enigma in the Diplopoda (Spirostreptida Odontopygidae). Tropical Zoology 4: 65 - 73. https: // doi. org / 10.1080 / 03946975.1991.10539475" type="journal article" year="1991">Hoffman (1991)</bibRefCitation>
clarified the situation.
<bibRefCitation id="EFC990DED6470655FEC0FE9EC4D4FE40" author="Enghoff H." box="[307,504,381,408]" pageId="98" pageNumber="99" pagination="1 - 19" refId="ref47336" refString="Enghoff H. 2016 a. A mountain of millipedes III: A new genus for three new species from the Udzungwa mountains and surroundings, Tanzania, as well as several ' orphaned' species previously assigned to Odontopyge Brandt, 1841 (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 177: 1 - 19. https: // doi. org / 10.5852 / ejt. 2016.177" type="journal article" year="2016">Enghoff (2016a)</bibRefCitation>
further addressed the “
<taxonomicName id="4C5896ACD6470655FCE6FE9EC688FE4F" box="[789,932,381,407]" class="Diplopoda" family="Odontopygidae" genus="Odontopyge" kingdom="Animalia" order="Spirostreptida" pageId="98" pageNumber="99" phylum="Arthropoda" rank="genus">
<emphasis id="B92C313DD6470655FCE6FE9EC688FE4F" box="[789,932,381,407]" italics="true" pageId="98" pageNumber="99">Odontopyge</emphasis>
</taxonomicName>
problem”. Of the “orphaned” nominal
<taxonomicName id="4C5896ACD6470655FF4EFE43C487FE62" box="[189,427,416,442]" pageId="98" pageNumber="99">
<emphasis id="B92C313DD6470655FF4EFE43C460FE62" box="[189,332,416,442]" italics="true" pageId="98" pageNumber="99">Odontopyge</emphasis>
species
</taxonomicName>
listed by
<bibRefCitation id="EFC990DED6470655FDD6FE43C7C0FE63" author="Enghoff H." box="[549,748,416,443]" pageId="98" pageNumber="99" pagination="1 - 19" refId="ref47336" refString="Enghoff H. 2016 a. A mountain of millipedes III: A new genus for three new species from the Udzungwa mountains and surroundings, Tanzania, as well as several ' orphaned' species previously assigned to Odontopyge Brandt, 1841 (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 177: 1 - 19. https: // doi. org / 10.5852 / ejt. 2016.177" type="journal article" year="2016">Enghoff (2016a)</bibRefCitation>
, one may provisionally be re-allocated to
<taxonomicName id="4C5896ACD6470655FB19FE43C05AFE62" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[1258,1398,416,442]" class="Diplopoda" family="Odontopygidae" genus="Uncodrama" kingdom="Animalia" order="Spirostreptida" pageId="98" pageNumber="99" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B92C313DD6470655FB19FE43C05AFE62" box="[1258,1398,416,442]" italics="true" pageId="98" pageNumber="99">Uncodrama</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F8C46D6470655FF4EFE20C403FE05" box="[189,303,451,477]" pageId="98" pageNumber="99" rank="genus">gen. nov.</taxonomicNameLabel>
, viz,
<taxonomicName id="4C5896ACD6470655FE85FE20C621FE05" authority="(Chamberlin, 1927)" authorityName="Enghoff" authorityYear="2022" baseAuthorityName="Chamberlin" baseAuthorityYear="1927" box="[374,781,451,477]" class="Diplopoda" family="Odontopygidae" genus="Uncodrama" kingdom="Animalia" order="Spirostreptida" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="medjensis">
<emphasis id="B92C313DD6470655FE85FE20C73EFE05" box="[374,530,451,477]" italics="true" pageId="98" pageNumber="99">O. medjensis</emphasis>
(
<bibRefCitation id="EFC990DED6470655FDD5FE20C62FFE05" author="Chamberlin R. V." box="[550,771,451,477]" pageId="98" pageNumber="99" pagination="177 - 249" refId="ref46758" refString="Chamberlin R. V. 1927. The Chilopoda and Diplopoda collected by the American Museum of Natural History Congo Expedition (1909 - 1915), with notes on some other African species. Bulletin of the American Museum of Natural History 57 (4): 177 - 249." type="journal article" year="1927">Chamberlin, 1927</bibRefCitation>
)
</taxonomicName>
, judged from the original description (
<bibRefCitation id="EFC990DED6470655FB19FE20C5D6FDD8" author="Chamberlin R. V." pageId="98" pageNumber="99" pagination="177 - 249" refId="ref46758" refString="Chamberlin R. V. 1927. The Chilopoda and Diplopoda collected by the American Museum of Natural History Congo Expedition (1909 - 1915), with notes on some other African species. Bulletin of the American Museum of Natural History 57 (4): 177 - 249." type="journal article" year="1927">Chamberlin 1927</bibRefCitation>
, as
<emphasis id="B92C313DD6470655FED4FE05C4D7FE27" box="[295,507,486,512]" italics="true" pageId="98" pageNumber="99">
<taxonomicName id="4C5896ACD6470655FED4FE05C4FBFDD8" authorityName="Attems" authorityYear="1909" box="[295,471,486,512]" class="Diplopoda" family="Odontopygidae" genus="Haplothysanus" kingdom="Animalia" order="Spirostreptida" pageId="98" pageNumber="99" phylum="Arthropoda" rank="genus">Haplothysanus</taxonomicName>
m.
</emphasis>
) and the re-description by
<bibRefCitation id="EFC990DED6470655FCC9FE05C6F5FDD8" author="Kraus O." box="[826,985,485,512]" pageId="98" pageNumber="99" pagination="1 - 207" refId="ref48861" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae-Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." type="journal article" year="1960">Kraus (1960)</bibRefCitation>
. The same applies, with even more reservation, to
<taxonomicName id="4C5896ACD6470655FE9FFDEAC798FDFB" authority="(Karsch, 1881)" authorityName="Enghoff" authorityYear="2022" baseAuthorityName="Karsch" baseAuthorityYear="1881" box="[364,692,520,547]" class="Diplopoda" family="Odontopygidae" genus="Uncodrama" kingdom="Animalia" order="Spirostreptida" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="tumidens">
<emphasis id="B92C313DD6470655FE9FFDEAC4D6FDFA" box="[364,506,520,547]" italics="true" pageId="98" pageNumber="99">O. tumidens</emphasis>
(Karsch, 1881)
</taxonomicName>
, based on the re-description by
<bibRefCitation id="EFC990DED6470655FBD4FDEAC1F9FDFB" author="Attems C. G." box="[1063,1237,521,547]" pageId="98" pageNumber="99" pagination="1 - 233" refId="ref46256" refString="Attems C. G. 1914. Afrikanische Spirostreptiden nebst Uberblick uber die Spirostreptiden orbis terrarum. Zoologica Stuttgart 65 - 66: 1 - 233." type="journal article" year="1914">Attems (1914)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>