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<document id="9C4136273C180B575C60350FD8E2FF01" ID-DOI="10.1080/00222933.2017.1326640" ID-ISSN="1464-5262" ID-Zenodo-Dep="5181243" ID-ZooBank="BD40ACCF-3EEB-43C3-BC54-0D2BB88F679F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628689236996" checkinUser="felipe" docAuthor="Frolov, Andrey V., Akhmetova, Lilia A. &amp; Vaz-de-Mello, Fernando Z." docDate="2017" docId="03E7473FFFC9FFBAFEADFB7676782735" docLanguage="en" docName="JNatHist.51.17-18.995-1014.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 51 (17 - 18)" docSource="http://dx.doi.org/10.1080/00222933.2017.1326640" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Paraegidium howdeni Frolov, Akhmetova" docType="treatment" docVersion="4" lastPageNumber="1012" masterDocId="FFDE3F47FFD9FFA8FFAFFFE0726C265C" masterDocTitle="Revision of the South American genus Paraegidium Vulcano et al. (Coleoptera: Scarabaeidae: Orphninae)" masterLastPageNumber="1014" masterPageNumber="995" pageNumber="1010" updateTime="1699049738498" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="B5BC7E80568E9DEE20F05472854C16A9">Revision of the South American genus Paraegidium Vulcano et al. (Coleoptera: Scarabaeidae: Orphninae)</mods:title>
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<mods:namePart id="BE180A2120E0004B7F78D4F7FFE82508">Frolov, Andrey V.</mods:namePart>
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<mods:namePart id="74186C281DB231A91ECBC35337272C2F">Akhmetova, Lilia A.</mods:namePart>
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<mods:namePart id="C3BB0174BB8CBB2CC7F84601BDCEA1E9">Vaz-de-Mello, Fernando Z.</mods:namePart>
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<mods:title id="697B12671EE24BF6BF175B319373F915">Journal of Natural History</mods:title>
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<mods:number id="71CE6442ED32F8031610CC42BE595172">2017-05-25</mods:number>
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<treatment id="03E7473FFFC9FFBAFEADFB7676782735" ID-DOI="http://doi.org/10.5281/zenodo.5190712" ID-Zenodo-Dep="5190712" LSID="urn:lsid:plazi:treatment:03E7473FFFC9FFBAFEADFB7676782735" httpUri="http://treatment.plazi.org/id/03E7473FFFC9FFBAFEADFB7676782735" lastPageId="18" lastPageNumber="1012" pageId="16" pageNumber="1010">
<subSubSection id="C354A5A2FFC9FFB8FEADFB76764E22EC" box="[258,1058,1174,1200]" pageId="16" pageNumber="1010" type="nomenclature">
<paragraph id="8BF1F629FFC9FFB8FEADFB76764E22EC" blockId="16.[258,1058,1174,1235]" box="[258,1058,1174,1200]" pageId="16" pageNumber="1010">
<heading id="D0B94145FFC9FFB8FEADFB76764E22EC" box="[258,1058,1174,1200]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="1010" reason="5">
<taxonomicName id="4C4E8DAAFFC9FFB8FEADFB76709522EC" authority="Frolov, Akhmetova" authorityName="Frolov, Akhmetova" box="[258,761,1174,1200]" class="Insecta" family="Hybosoridae" genus="Paraegidium" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1010" phylum="Arthropoda" rank="species" species="howdeni">
<emphasis id="B93A2A3BFFC9FFB8FEADFB76707D22EC" bold="true" box="[258,529,1174,1200]" italics="true" pageId="16" pageNumber="1010">Paraegidium howdeni</emphasis>
Frolov, Akhmetova
</taxonomicName>
&amp; Vaz-de-Mello, sp. nov.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C354A5A2FFC9FFB8FD88FB5A7090228F" box="[551,764,1210,1235]" pageId="16" pageNumber="1010" type="description">
<paragraph id="8BF1F629FFC9FFB8FD88FB5A7090228F" blockId="16.[258,1058,1174,1235]" box="[551,764,1210,1235]" pageId="16" pageNumber="1010">
(
<figureCitation id="1375EAACFFC9FFB8FD80FB5A70F0228F" box="[559,668,1210,1235]" captionStart="Figure 5" captionStartId="9.[160,225,1398,1421]" captionTargetBox="[163,1153,143,1367]" captionTargetId="figure-19@9.[163,1153,142,1367]" captionTargetPageId="9" captionText="Figure 5. Paraegidium howdeni, holotype male (a, f, g); paratype female (b); paratype males (ce, h). Habitus in dorsal view (a, b); pronotum and head (ce); aedeagus in lateral view (f); parameres in dorsal view (g); endophallus (h)." figureDoi="http://doi.org/10.5281/zenodo.5181253" httpUri="https://zenodo.org/record/5181253/files/figure.png" pageId="16" pageNumber="1010">Figures 5</figureCitation>
(ah), 7)
</paragraph>
</subSubSection>
<subSubSection id="C354A5A2FFC9FFB9FF0FFAE376362415" lastPageId="17" lastPageNumber="1011" pageId="16" pageNumber="1010" type="materials_examined">
<paragraph id="8BF1F629FFC9FFB8FF0FFAE3735B2340" blockId="16.[160,672,1283,1342]" box="[160,311,1283,1308]" pageId="16" pageNumber="1010">
<emphasis id="B93A2A3BFFC9FFB8FF0FFAE3735B2340" bold="true" box="[160,311,1283,1308]" italics="true" pageId="16" pageNumber="1010">Type locality</emphasis>
</paragraph>
<paragraph id="8BF1F629FFC9FFB8FF0FFAC670CC2362" blockId="16.[160,672,1283,1342]" box="[160,672,1318,1342]" pageId="16" pageNumber="1010">
<collectingCountry id="F359B6B9FFC9FFB8FF0FFAC672BA2362" box="[160,214,1318,1342]" name="Peru" pageId="16" pageNumber="1010">Peru</collectingCountry>
,
<collectingRegion id="498A38CBFFC9FFB8FF4EFAC673E72362" box="[225,395,1318,1342]" country="Peru" name="Madre de Dios" pageId="16" pageNumber="1010">Madre de Dios</collectingRegion>
, Rio Palma Real Grande.
</paragraph>
<paragraph id="8BF1F629FFC9FFB8FF0FFA8E73AE23DB" blockId="16.[160,1156,1390,1727]" box="[160,450,1390,1415]" pageId="16" pageNumber="1010">
<emphasis id="B93A2A3BFFC9FFB8FF0FFA8E73AE23DB" bold="true" box="[160,450,1390,1415]" italics="true" pageId="16" pageNumber="1010">Type material examined</emphasis>
</paragraph>
<paragraph id="8BF1F629FFC9FFB8FF0FFA72700723B3" blockId="16.[160,1156,1390,1727]" pageId="16" pageNumber="1010">
<materialsCitation id="3B26FC74FFC9FFB8FF0FFA72700723B3" collectingDate="1999-10" collectionCode="CMNC" collectorName="T. Larsen &amp; Ottawa &amp; Canada'" country="Peru" county="Male" elevation="220" latitude="-12.538889" location="Limon Camp" longLatPrecision="21" longitude="-68.86111" municipality="Rio Palma Real Grande" pageId="16" pageNumber="1010" specimenCount="1" stateProvince="Madre de Dios" typeStatus="holotype">
<typeStatus id="54F5488BFFC9FFB8FF0FFA72736523F6" box="[160,265,1426,1450]" pageId="16" pageNumber="1010" type="holotype">Holotype</typeStatus>
.
<collectingCounty id="62908EA5FFC9FFB8FEB9FA72732023F6" box="[278,332,1426,1450]" pageId="16" pageNumber="1010">Male</collectingCounty>
at
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labelled
<collectingCountry id="F359B6B9FFC9FFB8FD85FA72700423F6" box="[554,616,1426,1450]" name="Peru" pageId="16" pageNumber="1010">PERU</collectingCountry>
,
<collectingRegion id="498A38CBFFC9FFB8FDDBFA72717223F6" box="[628,798,1426,1450]" country="Peru" name="Madre de Dios" pageId="16" pageNumber="1010">Madre de Dios</collectingRegion>
:
<collectingMunicipality id="6B956C53FFC9FFB8FC85FA72765D23F6" box="[810,1073,1426,1450]" pageId="16" pageNumber="1010">Rio Palma Real Grande</collectingMunicipality>
,
<location id="8E91A0F2FFC9FFB8FB92FA72728E2391" LSID="urn:lsid:plazi:treatment:03E7473FFFC9FFBAFEADFB7676782735:8E91A0F2FFC9FFB8FB92FA72728E2391" country="Peru" county="Male" latitude="-12.538889" longLatPrecision="21" longitude="-68.86111" municipality="Rio Palma Real Grande" name="Limon Camp" pageId="16" pageNumber="1010" stateProvince="Madre de Dios">Limon Camp</location>
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12°32
<emphasis id="B93A2A3BFFC9FFB8FE9FFA54735A2390" bold="true" box="[304,310,1460,1484]" italics="true" pageId="16" pageNumber="1010">ʹ</emphasis>
20S
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<geoCoordinate id="EE7A90EEFFC9FFB8FEC2FA55738F2391" box="[365,483,1460,1485]" degrees="68" direction="west" minutes="51" orientation="longitude" pageId="16" pageNumber="1010" precision="15" seconds="40" value="-68.86111">
68°51
<emphasis id="B93A2A3BFFC9FFB8FE03FA5473DE2390" bold="true" box="[428,434,1460,1484]" italics="true" pageId="16" pageNumber="1010">ʹ</emphasis>
40W
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<collectingMethod id="520F8E3EFFC9FFB8FE5DFA5570892391" box="[498,741,1461,1485]" pageId="16" pageNumber="1010">Flight intercept trap</collectingMethod>
,
<quantity id="4CB65BCCFFC9FFB8FD58FA5571282391" box="[759,836,1461,1485]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="16" pageNumber="1010" unit="m" value="220.0">
<elevation id="0063111AFFC9FFB8FD58FA5571282391" box="[759,836,1461,1485]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="16" pageNumber="1010" unit="m" value="220.0">220 m</elevation>
</quantity>
<collectorName id="26BB93FFFFC9FFB8FCFCFA5571AD2391" box="[851,961,1461,1485]" pageId="16" pageNumber="1010">T. Larsen</collectorName>
,
<date id="FFF0D0E9FFC9FFB8FC7DFA5576492391" box="[978,1061,1461,1485]" pageId="16" pageNumber="1010" value="1999-10">
<collectingDate id="EFB42901FFC9FFB8FC7DFA5576492391" box="[978,1061,1461,1485]" pageId="16" pageNumber="1010" value="1999-10">X-1999</collectingDate>
</date>
/H. &amp; A. HOWDEN COLLECTION
<collectorName id="26BB93FFFFC9FFB8FE03FA37706C23B3" box="[428,512,1495,1519]" pageId="16" pageNumber="1010">Ottawa</collectorName>
,
<collectorName id="26BB93FFFFC9FFB8FDA4FA37700923B3" box="[523,613,1495,1519]" pageId="16" pageNumber="1010">Canada</collectorName>
.
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</paragraph>
<paragraph id="8BF1F629FFC9FFB9FF6FFA1A76042708" blockId="16.[160,1156,1390,1727]" lastBlockId="17.[160,1156,142,340]" lastPageId="17" lastPageNumber="1011" pageId="16" pageNumber="1010">
<materialsCitation id="3B26FC74FFC9FFB8FF6FFA1A7310204E" box="[192,380,1530,1554]" country="Peru" location="PERU" pageId="16" pageNumber="1010" specimenCount="1" typeStatus="paratype">
<typeStatus id="54F5488BFFC9FFB8FF6FFA1A735C204E" box="[192,304,1530,1554]" pageId="16" pageNumber="1010" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="F359B6B9FFC9FFB8FE91FA1A7310204E" box="[318,380,1530,1554]" name="Peru" pageId="16" pageNumber="1010">PERU</collectingCountry>
</materialsCitation>
.
<materialsCitation id="3B26FC74FFC9FFB8FE26FA1A737D2069" collectionCode="CMNC" country="Peru" location="Madre de Dios" pageId="16" pageNumber="1010" specimenCount="1" specimenCount-male="1" stateProvince="Madre de Dios" typeStatus="holotype">
<collectingRegion id="498A38CBFFC9FFB8FE26FA1A705B204E" box="[393,567,1530,1554]" country="Peru" name="Madre de Dios" pageId="16" pageNumber="1010">Madre de Dios</collectingRegion>
:
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at
<collectionCode id="ED5F6EECFFC9FFB8FD57FA1A7152204E" box="[760,830,1530,1554]" pageId="16" pageNumber="1010">CMNC</collectionCode>
with the same label as the
<typeStatus id="54F5488BFFC9FFB8FF0FF9FD737D2069" box="[160,273,1565,1589]" pageId="16" pageNumber="1010" type="holotype">holotypes</typeStatus>
</materialsCitation>
.
<collectingCountry id="F359B6B9FFC9FFB8FEB0F9FD731D2069" box="[287,369,1565,1589]" name="Brazil" pageId="16" pageNumber="1010">BRAZIL</collectingCountry>
.
<materialsCitation id="3B26FC74FFC9FFB8FED0F9FD71162069" box="[383,890,1564,1589]" collectionCode="CEMT" country="Peru" location="Mato Grosso" pageId="16" pageNumber="1010" specimenCount="1" specimenCount-male="1" stateProvince="Mato Grosso" typeStatus="paratype">
<collectingRegion id="498A38CBFFC9FFB8FED0F9FD707E2069" box="[383,530,1565,1589]" country="Brazil" name="Mato Grosso" pageId="16" pageNumber="1010">Mato Grosso</collectingRegion>
:
<specimenCount id="9D483DA0FFC9FFB8FD8FF9FD70E12069" box="[544,653,1565,1589]" pageId="16" pageNumber="1010" type="male">one male</specimenCount>
at
<collectionCode id="ED5F6EECFFC9FFB8FD17F9FD709B2069" box="[696,759,1565,1589]" pageId="16" pageNumber="1010">CEMT</collectionCode>
labelled V
</materialsCitation>
.VERA, M.
<materialsCitation id="3B26FC74FFC9FFB8FC4FF9FD72DD200B" country="Brazil" location="Grosso" pageId="16" pageNumber="1010" specimenCount="1" typeStatus="paratype">
<location id="8E91A0F2FFC9FFB8FC4FF9FD76582069" LSID="urn:lsid:plazi:treatment:03E7473FFFC9FFBAFEADFB7676782735:8E91A0F2FFC9FFB8FC4FF9FD76582069" box="[992,1076,1565,1589]" country="Brazil" name="Grosso" pageId="16" pageNumber="1010">Grosso</location>
,
<collectingCountry id="F359B6B9FFC9FFB8FBEDF9FD76EC2069" box="[1090,1152,1565,1589]" name="Brazil" pageId="16" pageNumber="1010">Brasil</collectingCountry>
,
<date id="FFF0D0E9FFC9FFB8FF0FF9DF72DD200B" box="[160,177,1599,1623]" pageId="16" pageNumber="1010" value="1973-10">X</date>
</materialsCitation>
.1973 M.
<materialsCitation id="3B26FC74FFC9FFB8FEB3F9DF7173200B" box="[284,799,1599,1623]" country="Brazil" latitude="-12.766666" location="Lon" longLatPrecision="1292" longitude="-55.6" municipality="Alvarenga" pageId="16" pageNumber="1010" specimenCount="1" typeStatus="paratype">
<collectingMunicipality id="6B956C53FFC9FFB8FEB3F9DF73FD200B" box="[284,401,1599,1623]" pageId="16" pageNumber="1010">Alvarenga</collectingMunicipality>
/
<location id="8E91A0F2FFC9FFB8FE36F9DF73AD200B" LSID="urn:lsid:plazi:treatment:03E7473FFFC9FFBAFEADFB7676782735:8E91A0F2FFC9FFB8FE36F9DF73AD200B" box="[409,449,1599,1623]" country="Brazil" latitude="-12.766666" longLatPrecision="1292" longitude="-55.6" municipality="Alvarenga" name="Lon" pageId="16" pageNumber="1010">Lon</location>
<geoCoordinate id="EE7A90EEFFC9FFB8FE64F9DF7059200B" box="[459,565,1599,1623]" degrees="55" direction="west" minutes="36" orientation="longitude" pageId="16" pageNumber="1010" precision="925" value="-55.6">
55 36
<emphasis id="B93A2A3BFFC9FFB8FDA2F9DF707F200B" bold="true" box="[525,531,1599,1623]" italics="true" pageId="16" pageNumber="1010">ʹ</emphasis>
W
</geoCoordinate>
<geoCoordinate id="EE7A90EEFFC9FFB8FD90F9DF70CC200B" box="[575,672,1599,1623]" degrees="12" direction="south" minutes="46" orientation="latitude" pageId="16" pageNumber="1010" precision="925" value="-12.766666">
12 46
<emphasis id="B93A2A3BFFC9FFB8FD2EF9DF70EB200B" bold="true" box="[641,647,1599,1623]" italics="true" pageId="16" pageNumber="1010">ʹ</emphasis>
S
</geoCoordinate>
/coleção M
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.
<materialsCitation id="3B26FC74FFC9FFB8FC81F9DF71C8200B" box="[814,932,1599,1623]" country="Brazil" location="Alvarenga'" pageId="16" pageNumber="1010" specimenCount="1" typeStatus="paratype">
<location id="8E91A0F2FFC9FFB8FC81F9DF71C8200B" LSID="urn:lsid:plazi:treatment:03E7473FFFC9FFBAFEADFB7676782735:8E91A0F2FFC9FFB8FC81F9DF71C8200B" box="[814,932,1599,1623]" country="Brazil" name="Alvarenga'" pageId="16" pageNumber="1010">Alvarenga</location>
</materialsCitation>
.
<materialsCitation id="3B26FC74FFC9FFB8FC1AF9DF70912026" collectionCode="CMNC" collectorName="Itaituba Maller-leg" country="Brazil" location="Para" pageId="16" pageNumber="1010" specimenCount="1" specimenCount-male="1" stateProvince="Para" typeStatus="paratype">
<collectingRegion id="498A38CBFFC9FFB8FC1AF9DF7184200B" box="[949,1000,1599,1623]" country="Brazil" name="Para" pageId="16" pageNumber="1010">Pará</collectingRegion>
:
<specimenCount id="9D483DA0FFC9FFB8FC59F9DF760F200B" box="[1014,1123,1599,1623]" pageId="16" pageNumber="1010" type="male">one male</specimenCount>
at
<collectionCode id="ED5F6EECFFC9FFB8FF0FF982728A2026" box="[160,230,1634,1658]" pageId="16" pageNumber="1010">CMNC</collectionCode>
labelled
<collectingCountry id="F359B6B9FFC9FFB8FEF0F98273C02026" box="[351,428,1634,1658]" name="Brazil" pageId="16" pageNumber="1010">BRASIL</collectingCountry>
<collectingRegion id="498A38CBFFC9FFB8FE72F98270762026" box="[477,538,1634,1658]" country="Brazil" name="Para" pageId="16" pageNumber="1010">PARA</collectingRegion>
<collectorName id="26BB93FFFFC9FFB8FD88F98270912026" box="[551,765,1634,1658]" pageId="16" pageNumber="1010">Itaituba Maller-leg</collectorName>
</materialsCitation>
.
<materialsCitation id="3B26FC74FFC9FFB8FCA2F98271562026" box="[781,826,1634,1658]" country="Brazil" location="Coll" pageId="16" pageNumber="1010" specimenCount="1" stateProvince="Para" typeStatus="paratype">
<location id="8E91A0F2FFC9FFB8FCA2F98271562026" LSID="urn:lsid:plazi:treatment:03E7473FFFC9FFBAFEADFB7676782735:8E91A0F2FFC9FFB8FCA2F98271562026" box="[781,826,1634,1658]" country="Brazil" name="Coll" pageId="16" pageNumber="1010" stateProvince="Para">Coll</location>
</materialsCitation>
. Martinez Nov. 963/H. &amp; A. HOWDEN COLLECTION ex. A. Martinez coll
<materialsCitation id="3B26FC74FFC9FFB8FD2CF965734D20E3" collectionCode="CEMT" country="Brazil" location="BRASIL" pageId="16" pageNumber="1010" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
.;
<specimenCount id="9D483DA0FFC9FFB8FD36F965716F20C1" box="[665,771,1669,1693]" pageId="16" pageNumber="1010" type="male">one male</specimenCount>
and
<specimenCount id="9D483DA0FFC9FFB8FC91F96571D120C1" box="[830,957,1669,1693]" pageId="16" pageNumber="1010" type="female">one female</specimenCount>
at
<collectionCode id="ED5F6EECFFC9FFB8FC4CF965764E20C1" box="[995,1058,1669,1693]" pageId="16" pageNumber="1010">CEMT</collectionCode>
labelled
<collectingCountry id="F359B6B9FFC9FFB8FF09F947729A20E3" box="[166,246,1703,1727]" name="Brazil" pageId="16" pageNumber="1010">BRASIL</collectingCountry>
: PA
</materialsCitation>
.
<materialsCitation id="3B26FC74FFC9FFB8FE84F947703920E3" box="[299,597,1703,1727]" country="Brazil" location="Redencao Pinkaiti" pageId="16" pageNumber="1010" specimenCount="1" typeStatus="paratype">
<location id="8E91A0F2FFC9FFB8FE84F947739F20E3" LSID="urn:lsid:plazi:treatment:03E7473FFFC9FFBAFEADFB7676782735:8E91A0F2FFC9FFB8FE84F947739F20E3" box="[299,499,1703,1727]" country="Brazil" name="Redencao Pinkaiti" pageId="16" pageNumber="1010">Redenção Pinkaiti</location>
AIK
<date id="FFF0D0E9FFC9FFB8FD93F947703920E3" box="[572,597,1703,1727]" pageId="16" pageNumber="1010" value="1998-10-31">31</date>
</materialsCitation>
<materialsCitation id="3B26FC74FFC9FFB8FDFAF947700620E3" box="[597,618,1703,1727]" country="Brazil" isTailEnumeration="true" location="Redencao Pinkaiti" pageId="16" pageNumber="1010" specimenCount="1" typeStatus="paratype">.X</materialsCitation>
.1998
<geoCoordinate id="EE7A90EEFFC9FFB8FD1FF947716E20E3" box="[688,770,1703,1727]" degrees="07" direction="south" minutes="46" orientation="latitude" pageId="16" pageNumber="1010" precision="925" value="-7.7666664">
07°46
<emphasis id="B93A2A3BFFC9FFB8FD40F947709920E3" bold="true" box="[751,757,1703,1727]" italics="true" pageId="16" pageNumber="1010">ʹ</emphasis>
S
</geoCoordinate>
<geoCoordinate id="EE7A90EEFFC9FFB8FCA6F947710920E3" box="[777,869,1703,1727]" degrees="51" direction="west" minutes="58" orientation="longitude" pageId="16" pageNumber="1010" precision="925" value="-51.966667">
51°58
<emphasis id="B93A2A3BFFC9FFB8FCE6F947712320E3" bold="true" box="[841,847,1703,1727]" italics="true" pageId="16" pageNumber="1010">ʹ</emphasis>
W
</geoCoordinate>
P.Y.
<materialsCitation id="3B26FC74FFC9FFB9FC33F947717226FA" collectionCode="CEMT, EPGC" country="Brazil" lastPageId="17" lastPageNumber="1011" location="Scheffler'" pageId="16" pageNumber="1010" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<location id="8E91A0F2FFC9FFB8FC33F947766E20E3" LSID="urn:lsid:plazi:treatment:03E7473FFFC9FFBAFEADFB7676782735:8E91A0F2FFC9FFB8FC33F947766E20E3" box="[924,1026,1703,1727]" country="Brazil" name="Scheffler'" pageId="16" pageNumber="1010">Scheffler</location>
;
<specimenCount id="9D483DA0FFC9FFB8FBA0F94776EF20E3" box="[1039,1155,1703,1727]" pageId="16" pageNumber="1010" type="male">two males</specimenCount>
and
<specimenCount id="9D483DA0FFC8FFB9FF7DFF6E733126FA" box="[210,349,142,166]" pageId="17" pageNumber="1011" type="female">four females</specimenCount>
at
<collectionCode id="ED5F6EECFFC8FFB9FE2DFF6E73AD26FA" box="[386,449,142,166]" pageId="17" pageNumber="1011">CEMT</collectionCode>
and
<collectionCode id="ED5F6EECFFC8FFB9FE55FF6E705A26FA" box="[506,566,142,166]" pageId="17" pageNumber="1011">EPGC</collectionCode>
labelled
<collectingCountry id="F359B6B9FFC8FFB9FD0CFF6E709F26FA" box="[675,755,142,166]" name="Brazil" pageId="17" pageNumber="1011">BRASIL</collectingCountry>
: PA
</materialsCitation>
.
<materialsCitation id="3B26FC74FFC8FFB9FC87FF6E762526FA" box="[808,1097,142,166]" country="Brazil" location="Redencao Pinkaiti" pageId="17" pageNumber="1011" specimenCount="1" typeStatus="paratype">
<location id="8E91A0F2FFC8FFB9FC87FF6E719C26FA" LSID="urn:lsid:plazi:treatment:03E7473FFFC9FFBAFEADFB7676782735:8E91A0F2FFC8FFB9FC87FF6E719C26FA" box="[808,1008,142,166]" country="Brazil" name="Redencao Pinkaiti" pageId="17" pageNumber="1011">Redenção Pinkaiti</location>
AIK
<date id="FFF0D0E9FFC8FFB9FB97FF6E762526FA" box="[1080,1097,142,166]" pageId="17" pageNumber="1011" value="1999-10">X</date>
</materialsCitation>
.1999
<geoCoordinate id="EE7A90EEFFC8FFB9FF0FFF51729E2695" box="[160,242,177,201]" degrees="07" direction="south" minutes="46" orientation="latitude" pageId="17" pageNumber="1011" precision="925" value="-7.7666664">
07°46
<emphasis id="B93A2A3BFFC8FFB9FF70FF5172892695" bold="true" box="[223,229,177,201]" italics="true" pageId="17" pageNumber="1011">ʹ</emphasis>
S
</geoCoordinate>
<geoCoordinate id="EE7A90EEFFC8FFB9FF50FF5173302695" box="[255,348,177,201]" degrees="51" direction="west" minutes="58" orientation="longitude" pageId="17" pageNumber="1011" precision="925" value="-51.966667">
51°58
<emphasis id="B93A2A3BFFC8FFB9FE90FF5173292695" bold="true" box="[319,325,177,201]" italics="true" pageId="17" pageNumber="1011">ʹ</emphasis>
W
</geoCoordinate>
P.Y.
<materialsCitation id="3B26FC74FFC8FFB9FE32FF5176682695" box="[413,1028,177,201]" collectionCode="CEMT" country="Brazil" location="Scheffler'" pageId="17" pageNumber="1011" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<location id="8E91A0F2FFC8FFB9FE32FF5170682695" LSID="urn:lsid:plazi:treatment:03E7473FFFC9FFBAFEADFB7676782735:8E91A0F2FFC8FFB9FE32FF5170682695" box="[413,516,177,201]" country="Brazil" name="Scheffler'" pageId="17" pageNumber="1011">Scheffler</location>
;
<specimenCount id="9D483DA0FFC8FFB9FDB8FF5170F72695" box="[535,667,177,201]" pageId="17" pageNumber="1011" type="female">one female</specimenCount>
at
<collectionCode id="ED5F6EECFFC8FFB9FD64FF5171662695" box="[715,778,177,201]" pageId="17" pageNumber="1011">CEMT</collectionCode>
labelled
<collectingCountry id="F359B6B9FFC8FFB9FC2CFF5171BF2695" box="[899,979,177,201]" name="Brazil" pageId="17" pageNumber="1011">BRASIL</collectingCountry>
: PA
</materialsCitation>
.
<materialsCitation id="3B26FC74FFC8FFB9FBBAFF51707126B0" country="Brazil" latitude="-7.7666664" location="Redencao Pinkaiti" longLatPrecision="1303" longitude="-51.966667" pageId="17" pageNumber="1011" specimenCount="1" typeStatus="paratype">
<location id="8E91A0F2FFC8FFB9FBBAFF51729E26B0" LSID="urn:lsid:plazi:treatment:03E7473FFFC9FFBAFEADFB7676782735:8E91A0F2FFC8FFB9FBBAFF51729E26B0" country="Brazil" latitude="-7.7666664" longLatPrecision="1303" longitude="-51.966667" name="Redencao Pinkaiti" pageId="17" pageNumber="1011">Redenção Pinkaiti</location>
AIK
<geoCoordinate id="EE7A90EEFFC8FFB9FEE6FF3473F726B0" box="[329,411,211,236]" degrees="07" direction="south" minutes="46" orientation="latitude" pageId="17" pageNumber="1011" precision="925" value="-7.7666664">
07°46
<emphasis id="B93A2A3BFFC8FFB9FE27FF3373E226B7" bold="true" box="[392,398,211,235]" italics="true" pageId="17" pageNumber="1011">ʹ</emphasis>
S
</geoCoordinate>
<geoCoordinate id="EE7A90EEFFC8FFB9FE08FF34706F26B0" box="[423,515,211,236]" degrees="51" direction="west" minutes="58" orientation="longitude" pageId="17" pageNumber="1011" precision="925" value="-51.966667">
51°58
<emphasis id="B93A2A3BFFC8FFB9FE48FF33738126B7" bold="true" box="[487,493,211,235]" italics="true" pageId="17" pageNumber="1011">ʹ</emphasis>
W
</geoCoordinate>
F
</materialsCitation>
.T.T.
<date id="FFF0D0E9FFC8FFB9FDFCFF3470C926B0" box="[595,677,212,236]" pageId="17" pageNumber="1011" value="1998-11">XI.1998</date>
P&amp;T Scheffler.
<materialsCitation id="3B26FC74FFC8FFB9FCCDFF34763F2752" collectionCode="CEMT" collectorName="Porto Velho BR" country="Brazil" location="Rondonia" pageId="17" pageNumber="1011" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Rondonia" typeStatus="paratype">
<collectingRegion id="498A38CBFFC8FFB9FCCDFF3471BC26B0" box="[866,976,212,236]" country="Brazil" name="Rondonia" pageId="17" pageNumber="1011">Rondônia</collectingRegion>
:
<specimenCount id="9D483DA0FFC8FFB9FC70FF34762126B0" box="[991,1101,212,236]" pageId="17" pageNumber="1011" type="male">one male</specimenCount>
and
<specimenCount id="9D483DA0FFC8FFB9FF0FFF16734F2752" box="[160,291,246,270]" pageId="17" pageNumber="1011" type="female">one female</specimenCount>
at
<collectionCode id="ED5F6EECFFC8FFB9FEFEFF1673FC2752" box="[337,400,246,270]" pageId="17" pageNumber="1011">CEMT</collectionCode>
labelled
<collectingCountry id="F359B6B9FFC8FFB9FDA7FF1670342752" box="[520,600,246,270]" name="Brazil" pageId="17" pageNumber="1011">BRASIL</collectingCountry>
,
<collectingRegion id="498A38CBFFC8FFB9FDC8FF1670B92752" box="[615,725,246,270]" country="Brazil" name="Rondonia" pageId="17" pageNumber="1011">Rondonia</collectingRegion>
:
<collectorName id="26BB93FFFFC8FFB9FD4BFF1671FA2752" box="[740,918,246,270]" pageId="17" pageNumber="1011">Porto Velho BR</collectorName>
364, km 2030
</materialsCitation>
.
<date id="FFF0D0E9FFC8FFB9FBCDFF167280276D" pageId="17" pageNumber="1011" value="1980-10-24">24. X.1980</date>
Eq.
<materialsCitation id="3B26FC74FFC8FFB9FEB4FEF971B9276D" box="[283,981,281,305]" collectionCode="MNRJ" collectorName="Vilhena" country="Brazil" pageId="17" pageNumber="1011" specimenCount="1" specimenCount-male="1" stateProvince="Rondonia" typeStatus="paratype">
INPA;
<specimenCount id="9D483DA0FFC8FFB9FECBFEF973A1276D" box="[356,461,281,305]" pageId="17" pageNumber="1011" type="male">one male</specimenCount>
at
<collectionCode id="ED5F6EECFFC8FFB9FE5FFEF97043276D" box="[496,559,281,305]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="17" pageNumber="1011">MNRJ</collectionCode>
labelled BRASIL-Rondônia
<collectorName id="26BB93FFFFC8FFB9FCCCFEF971DA276D" box="[867,950,281,305]" pageId="17" pageNumber="1011">Vilhena</collectorName>
<date id="FFF0D0E9FFC8FFB9FC11FEF971B9276D" box="[958,981,281,305]" pageId="17" pageNumber="1011" value="1987-11">XI</date>
</materialsCitation>
.1987 O. Roppa, P.
<materialsCitation id="3B26FC74FFC8FFB9FF12FEDC713C2708" box="[189,848,316,340]" country="Brazil" location="Museu Nacional" municipality="Magno" pageId="17" pageNumber="1011" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="paratype">
<collectingMunicipality id="6B956C53FFC8FFB9FF12FEDC73602708" box="[189,268,316,340]" pageId="17" pageNumber="1011">Magno</collectingMunicipality>
/
<location id="8E91A0F2FFC8FFB9FEBCFEDC73A52708" LSID="urn:lsid:plazi:treatment:03E7473FFFC9FFBAFEADFB7676782735:8E91A0F2FFC8FFB9FEBCFEDC73A52708" box="[275,457,316,340]" country="Brazil" municipality="Magno" name="Museu Nacional" pageId="17" pageNumber="1011" stateProvince="Rio de Janeiro">Museu Nacional</location>
de
<collectingRegion id="498A38CBFFC8FFB9FE58FEDC70F62708" box="[503,666,316,340]" country="Brazil" name="Rio de Janeiro" pageId="17" pageNumber="1011">Rio de Janeiro</collectingRegion>
/
<taxonomicName id="4C4E8DAAFFC8FFB9FD0DFEDC713C2708" box="[674,848,316,340]" class="Insecta" family="Hybosoridae" genus="Paraegidium" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1011" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B93A2A3BFFC8FFB9FD0DFEDC71402708" box="[674,812,316,340]" italics="true" pageId="17" pageNumber="1011">Paraegidium</emphasis>
sp
</taxonomicName>
</materialsCitation>
. Vaz-de-Mello det 2003.
</paragraph>
<paragraph id="8BF1F629FFC8FFB9FF0FFE63706927C0" blockId="17.[160,1157,387,585]" box="[160,517,387,412]" pageId="17" pageNumber="1011">
<emphasis id="B93A2A3BFFC8FFB9FF0FFE63706927C0" bold="true" box="[160,517,387,412]" italics="true" pageId="17" pageNumber="1011">Additional material examined</emphasis>
</paragraph>
<paragraph id="8BF1F629FFC8FFB9FF0FFE4776362415" blockId="17.[160,1157,387,585]" pageId="17" pageNumber="1011">
<materialsCitation id="3B26FC74FFC8FFB9FF0FFE4771A227E3" box="[160,974,423,447]" collectingDate="2009-10-20" collectorName="La Convencion &amp; Echarate &amp; Timpia" country="Peru" elevation="429" location="Cusco" pageId="17" pageNumber="1011" specimenCount="1" stateProvince="Cusco">
<collectingCountry id="F359B6B9FFC8FFB9FF0FFE4772B127E3" box="[160,221,423,447]" name="Peru" pageId="17" pageNumber="1011">PERU</collectingCountry>
.
<collectingRegion id="498A38CBFFC8FFB9FF49FE47734727E3" box="[230,299,423,447]" country="Peru" name="Cusco" pageId="17" pageNumber="1011">Cusco</collectingRegion>
:
<collectorName id="26BB93FFFFC8FFB9FE9BFE4773B527E3" box="[308,473,423,447]" pageId="17" pageNumber="1011">La Convención</collectorName>
,
<collectorName id="26BB93FFFFC8FFB9FE4DFE47702F27E3" box="[482,579,423,447]" pageId="17" pageNumber="1011">Echarate</collectorName>
,
<collectorName id="26BB93FFFFC8FFB9FDE3FE4770F527E3" box="[588,665,423,447]" pageId="17" pageNumber="1011">Timpia</collectorName>
,
<quantity id="4CB65BCCFFC8FFB9FD0DFE47708627E3" box="[674,746,423,447]" metricMagnitude="2" metricUnit="m" metricValue="4.29" pageId="17" pageNumber="1011" unit="m" value="429.0">
<elevation id="0063111AFFC8FFB9FD0DFE47708627E3" box="[674,746,423,447]" metricMagnitude="2" metricUnit="m" metricValue="4.29" pageId="17" pageNumber="1011" unit="m" value="429.0">429 m</elevation>
</quantity>
,
<date id="FFF0D0E9FFC8FFB9FD5BFE4771C127E3" box="[756,941,423,447]" pageId="17" pageNumber="1011" value="2009-10-20">
<collectingDate id="EFB42901FFC8FFB9FD5BFE4771C127E3" box="[756,941,423,447]" pageId="17" pageNumber="1011" value="2009-10-20">20 October 2009</collectingDate>
</date>
, M
</materialsCitation>
<materialsCitation id="3B26FC74FFC8FFB9FC61FE4776EC27E3" box="[974,1152,423,447]" collectingDate="2009-10-20" collectorName="La Convencion &amp; Echarate &amp; Timpia" country="Peru" elevation="429" isTailEnumeration="true" location="Alvarado" pageId="17" pageNumber="1011" specimenCount="1" stateProvince="Cusco">
.
<location id="8E91A0F2FFC8FFB9FC77FE47765527E3" LSID="urn:lsid:plazi:treatment:03E7473FFFC9FFBAFEADFB7676782735:8E91A0F2FFC8FFB9FC77FE47765527E3" box="[984,1081,423,447]" country="Peru" name="Alvarado" pageId="17" pageNumber="1011">Alvarado</location>
and E
</materialsCitation>
. Rázuri leg
<materialsCitation id="3B26FC74FFC8FFB9FEA1FE2972DB2458" collectingDate="2009-10-22" collectionCode="UNMSM" collectorName="La Convencion &amp; Echarate &amp; Timpia" country="Peru" elevation="371" pageId="17" pageNumber="1011" specimenCount="1" specimenCount-female="1">
.,
<specimenCount id="9D483DA0FFC8FFB9FEB4FE2973FA27BD" box="[283,406,457,481]" pageId="17" pageNumber="1011" type="female">one female</specimenCount>
(
<collectionCode id="ED5F6EECFFC8FFB9FE0FFE29739327BD" box="[416,511,457,481]" pageId="17" pageNumber="1011">UNMSM</collectionCode>
);
<collectorName id="26BB93FFFFC8FFB9FDA1FE2970DE27BD" box="[526,690,457,481]" pageId="17" pageNumber="1011">La Convención</collectorName>
,
<collectorName id="26BB93FFFFC8FFB9FD15FE29717727BD" box="[698,795,457,481]" pageId="17" pageNumber="1011">Echarate</collectorName>
,
<collectorName id="26BB93FFFFC8FFB9FC8BFE29711D27BD" box="[804,881,457,481]" pageId="17" pageNumber="1011">Timpia</collectorName>
,
<quantity id="4CB65BCCFFC8FFB9FCD6FE2971D327BD" box="[889,959,457,481]" metricMagnitude="2" metricUnit="m" metricValue="3.71" pageId="17" pageNumber="1011" unit="m" value="371.0">
<elevation id="0063111AFFC8FFB9FCD6FE2971D327BD" box="[889,959,457,481]" metricMagnitude="2" metricUnit="m" metricValue="3.71" pageId="17" pageNumber="1011" unit="m" value="371.0">371 m</elevation>
</quantity>
,
<date id="FFF0D0E9FFC8FFB9FC67FE2976EC27BD" box="[968,1152,457,481]" pageId="17" pageNumber="1011" value="2009-10-22">
<collectingDate id="EFB42901FFC8FFB9FC67FE2976EC27BD" box="[968,1152,457,481]" pageId="17" pageNumber="1011" value="2009-10-22">22 October 2009</collectingDate>
</date>
, M
</materialsCitation>
<materialsCitation id="3B26FC74FFC8FFB9FF18FE0C731B2458" box="[183,375,492,516]" collectingDate="2009-10-22" collectionCode="UNMSM" collectorName="La Convencion &amp; Echarate &amp; Timpia" country="Peru" elevation="371" isTailEnumeration="true" location="Alvarado" pageId="17" pageNumber="1011" specimenCount="1">
.
<location id="8E91A0F2FFC8FFB9FF69FE0C734B2458" LSID="urn:lsid:plazi:treatment:03E7473FFFC9FFBAFEADFB7676782735:8E91A0F2FFC8FFB9FF69FE0C734B2458" box="[198,295,492,516]" country="Peru" name="Alvarado" pageId="17" pageNumber="1011">Alvarado</location>
and E
</materialsCitation>
. Rázuri leg
<materialsCitation id="3B26FC74FFC8FFB9FE56FE0C73B3247B" collectingDate="2009-10-16" collectionCode="UNMSM" collectorName="La Convencion &amp; Echarate &amp; Timpia" country="Peru" elevation="442" pageId="17" pageNumber="1011" specimenCount="1" specimenCount-female="1">
.,
<specimenCount id="9D483DA0FFC8FFB9FDA5FE0C70E62458" box="[522,650,492,516]" pageId="17" pageNumber="1011" type="female">one female</specimenCount>
(
<collectionCode id="ED5F6EECFFC8FFB9FD34FE0C70962458" box="[667,762,492,516]" pageId="17" pageNumber="1011">UNMSM</collectionCode>
);
<collectorName id="26BB93FFFFC8FFB9FCA1FE0C71DB2458" box="[782,951,492,516]" pageId="17" pageNumber="1011">La Convención</collectorName>
,
<collectorName id="26BB93FFFFC8FFB9FC6AFE0C764A2458" box="[965,1062,492,516]" pageId="17" pageNumber="1011">Echarate</collectorName>
,
<collectorName id="26BB93FFFFC8FFB9FB9CFE0C76EC2458" box="[1075,1152,492,516]" pageId="17" pageNumber="1011">Timpia</collectorName>
,
<quantity id="4CB65BCCFFC8FFB9FF0FFDEF7287247B" box="[160,235,527,551]" metricMagnitude="2" metricUnit="m" metricValue="4.42" pageId="17" pageNumber="1011" unit="m" value="442.0">
<elevation id="0063111AFFC8FFB9FF0FFDEF7287247B" box="[160,235,527,551]" metricMagnitude="2" metricUnit="m" metricValue="4.42" pageId="17" pageNumber="1011" unit="m" value="442.0">442 m</elevation>
</quantity>
,
<date id="FFF0D0E9FFC8FFB9FF56FDEF73D7247B" box="[249,443,527,551]" pageId="17" pageNumber="1011" value="2009-10-16">
<collectingDate id="EFB42901FFC8FFB9FF56FDEF73D7247B" box="[249,443,527,551]" pageId="17" pageNumber="1011" value="2009-10-16">16 October 2009</collectingDate>
</date>
, M
</materialsCitation>
<materialsCitation id="3B26FC74FFC8FFB9FE4FFDEF70F1247B" box="[480,669,527,551]" collectingDate="2009-10-16" collectionCode="UNMSM" collectorName="La Convencion &amp; Echarate &amp; Timpia" country="Peru" elevation="442" isTailEnumeration="true" location="Alvarado" pageId="17" pageNumber="1011" specimenCount="1">
.
<location id="8E91A0F2FFC8FFB9FE42FDEF7022247B" LSID="urn:lsid:plazi:treatment:03E7473FFFC9FFBAFEADFB7676782735:8E91A0F2FFC8FFB9FE42FDEF7022247B" box="[493,590,527,551]" country="Peru" name="Alvarado" pageId="17" pageNumber="1011">Alvarado</location>
and E
</materialsCitation>
. Rázuri leg
<materialsCitation id="3B26FC74FFC8FFB9FCB1FDEF7648247B" box="[798,1060,527,551]" collectionCode="UNMSM" country="Peru" pageId="17" pageNumber="1011" specimenCount="1" specimenCount-female="1">
.,
<specimenCount id="9D483DA0FFC8FFB9FC80FDEF71C2247B" box="[815,942,527,551]" pageId="17" pageNumber="1011" type="female">one female</specimenCount>
(
<collectionCode id="ED5F6EECFFC8FFB9FC10FDEF7672247B" box="[959,1054,527,551]" pageId="17" pageNumber="1011">UNMSM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B26FC74FFC8FFB9FB9EFDEF71E52415" collectingDate="1996-09" collectorName="Pacaraima &amp; Ribeiro &amp; Vaz-de-Mello" country="Brazil" location="Roraima" pageId="17" pageNumber="1011" specimenCount="1" stateProvince="Roraima">
<collectingCountry id="F359B6B9FFC8FFB9FB9EFDEF76ED247B" box="[1073,1153,527,551]" name="Brazil" pageId="17" pageNumber="1011">BRAZIL</collectingCountry>
.
<collectingRegion id="498A38CBFFC8FFB9FF0FFDD172902415" box="[160,252,561,585]" country="Brazil" name="Roraima" pageId="17" pageNumber="1011">Roraima</collectingRegion>
:
<collectorName id="26BB93FFFFC8FFB9FEA8FDD173162415" box="[263,378,561,585]" pageId="17" pageNumber="1011">Pacaraima</collectorName>
,
<date id="FFF0D0E9FFC8FFB9FE2BFDD170502415" box="[388,572,561,585]" pageId="17" pageNumber="1011" value="1996-09">
<collectingDate id="EFB42901FFC8FFB9FE2BFDD170502415" box="[388,572,561,585]" pageId="17" pageNumber="1011" value="1996-09">September 1996</collectingDate>
</date>
,
<collectorName id="26BB93FFFFC8FFB9FDE8FDD170F82415" box="[583,660,561,585]" pageId="17" pageNumber="1011">Ribeiro</collectorName>
and
<collectorName id="26BB93FFFFC8FFB9FD62FDD171302415" box="[717,860,561,585]" pageId="17" pageNumber="1011">Vaz-de-Mello</collectorName>
leg
</materialsCitation>
<materialsCitation id="3B26FC74FFC8FFB9FC26FDD1763A2415" box="[905,1110,561,585]" collectionCode="CEMT" country="Brazil" pageId="17" pageNumber="1011" specimenCount="1" specimenCount-male="1">
.,
<specimenCount id="9D483DA0FFC8FFB9FC38FDD171932415" box="[919,1023,561,585]" pageId="17" pageNumber="1011" type="male">one male</specimenCount>
(
<collectionCode id="ED5F6EECFFC8FFB9FBA3FDD1763D2415" box="[1036,1105,561,585]" pageId="17" pageNumber="1011">CEMT</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C354A5A2FFC8FFB9FF0FFD9970AC2540" pageId="17" pageNumber="1011" type="diagnosis">
<paragraph id="8BF1F629FFC8FFB9FF0FFD99737824CE" blockId="17.[160,1156,633,796]" box="[160,276,633,658]" pageId="17" pageNumber="1011">
<emphasis id="B93A2A3BFFC8FFB9FF0FFD99737824CE" bold="true" box="[160,276,633,658]" italics="true" pageId="17" pageNumber="1011">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BF1F629FFC8FFB9FF0FFD7C70AC2540" blockId="17.[160,1156,633,796]" pageId="17" pageNumber="1011">
In the shape of the anterior frontoclypeal processes, larger males of
<taxonomicName id="4C4E8DAAFFC8FFB9FC04FD7C764F24E8" authority="Frolov &amp; Akhmetova &amp; Vaz-de-Mello, 2017" authorityName="Frolov &amp; Akhmetova &amp; Vaz-de-Mello" authorityYear="2017" box="[939,1059,668,692]" class="Insecta" family="Hybosoridae" genus="Paraegidium" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1011" phylum="Arthropoda" rank="species" species="howdeni" status="sp. nov.">
<emphasis id="B93A2A3BFFC8FFB9FC04FD7C764F24E8" box="[939,1059,668,692]" italics="true" pageId="17" pageNumber="1011">P. howdeni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2099740FFC8FFB9FB83FD7C76E824E8" box="[1068,1156,668,692]" pageId="17" pageNumber="1011" rank="species">sp. nov.</taxonomicNameLabel>
are similar to
<taxonomicName id="4C4E8DAAFFC8FFB9FE96FD5F73B5248B" authority="Frolov &amp; Akhmetova &amp; Vaz-de-Mello, 2017" authorityName="Frolov &amp; Akhmetova &amp; Vaz-de-Mello" authorityYear="2017" box="[313,473,703,727]" class="Insecta" family="Hybosoridae" genus="Paraegidium" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1011" phylum="Arthropoda" rank="species" species="monneorum" status="sp. nov.">
<emphasis id="B93A2A3BFFC8FFB9FE96FD5F73B5248B" box="[313,473,703,727]" italics="true" pageId="17" pageNumber="1011">P. monneorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2099740FFC8FFB9FE4FFD5F7055248B" box="[480,569,703,727]" pageId="17" pageNumber="1011" rank="species">sp. nov.</taxonomicNameLabel>
, while smaller males are similar to the smaller males of
<taxonomicName id="4C4E8DAAFFC8FFB9FF6EFD02735824A6" authority="Frolov &amp; Akhmetova &amp; Vaz-de-Mello, 2017" authorityName="Frolov &amp; Akhmetova &amp; Vaz-de-Mello" authorityYear="2017" box="[193,308,738,762]" class="Insecta" family="Hybosoridae" genus="Paraegidium" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1011" phylum="Arthropoda" rank="species" species="oliveirai" status="sp. nov.">
<emphasis id="B93A2A3BFFC8FFB9FF6EFD02735824A6" box="[193,308,738,762]" italics="true" pageId="17" pageNumber="1011">P. oliveirai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2099740FFC8FFB9FE91FD0273F424A6" box="[318,408,738,762]" pageId="17" pageNumber="1011" rank="species">sp. nov.</taxonomicNameLabel>
From both species
<taxonomicName id="4C4E8DAAFFC8FFB9FD2CFD02709124A6" authority="Frolov &amp; Akhmetova &amp; Vaz-de-Mello, 2017" authorityName="Frolov &amp; Akhmetova &amp; Vaz-de-Mello" authorityYear="2017" box="[643,765,738,762]" class="Insecta" family="Hybosoridae" genus="Paraegidium" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1011" phylum="Arthropoda" rank="species" species="howdeni" status="sp. nov.">
<emphasis id="B93A2A3BFFC8FFB9FD2CFD02709124A6" box="[643,765,738,762]" italics="true" pageId="17" pageNumber="1011">P. howdeni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2099740FFC8FFB9FCA7FD02710F24A6" box="[776,867,738,762]" pageId="17" pageNumber="1011" rank="species">sp. nov.</taxonomicNameLabel>
differs in the shape and relative length of the parameres (
<figureCitation id="1375EAACFFC8FFB9FD8FFCE470132540" box="[544,639,772,796]" captionStart="Figure 5" captionStartId="9.[160,225,1398,1421]" captionTargetBox="[163,1153,143,1367]" captionTargetId="figure-19@9.[163,1153,142,1367]" captionTargetPageId="9" captionText="Figure 5. Paraegidium howdeni, holotype male (a, f, g); paratype female (b); paratype males (ce, h). Habitus in dorsal view (a, b); pronotum and head (ce); aedeagus in lateral view (f); parameres in dorsal view (g); endophallus (h)." figureDoi="http://doi.org/10.5281/zenodo.5181253" httpUri="https://zenodo.org/record/5181253/files/figure.png" pageId="17" pageNumber="1011">Figure 5</figureCitation>
(f, g)).
</paragraph>
</subSubSection>
<subSubSection id="C354A5A2FFC8FFB9FF0FFCAC700323A6" pageId="17" pageNumber="1011" type="description">
<paragraph id="8BF1F629FFC8FFB9FF0FFCAC734A2539" blockId="17.[160,1156,844,1354]" box="[160,294,844,869]" pageId="17" pageNumber="1011">
<emphasis id="B93A2A3BFFC8FFB9FF0FFCAC734A2539" bold="true" box="[160,294,844,869]" italics="true" pageId="17" pageNumber="1011">Description</emphasis>
</paragraph>
<paragraph id="8BF1F629FFC8FFB9FF0FFC8F707B25DB" blockId="17.[160,1156,844,1354]" box="[160,535,879,903]" pageId="17" pageNumber="1011">
<typeStatus id="54F5488BFFC8FFB9FF0FFC8F736525DB" box="[160,265,879,903]" pageId="17" pageNumber="1011" type="holotype">Holotype</typeStatus>
, male (
<figureCitation id="1375EAACFFC8FFB9FEF3FC8F73D625DB" box="[348,442,879,903]" captionStart="Figure 5" captionStartId="9.[160,225,1398,1421]" captionTargetBox="[163,1153,143,1367]" captionTargetId="figure-19@9.[163,1153,142,1367]" captionTargetPageId="9" captionText="Figure 5. Paraegidium howdeni, holotype male (a, f, g); paratype female (b); paratype males (ce, h). Habitus in dorsal view (a, b); pronotum and head (ce); aedeagus in lateral view (f); parameres in dorsal view (g); endophallus (h)." figureDoi="http://doi.org/10.5281/zenodo.5181253" httpUri="https://zenodo.org/record/5181253/files/figure.png" pageId="17" pageNumber="1011">Figure 5</figureCitation>
(a, f, g)).
</paragraph>
<paragraph id="8BF1F629FFC8FFB9FF6FFC7270A325F6" blockId="17.[160,1156,844,1354]" box="[192,719,914,938]" pageId="17" pageNumber="1011">
Body length
<quantity id="4CB65BCCFFC8FFB9FEFCFC7273C025F6" box="[339,428,914,938]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="17" pageNumber="1011" unit="mm" value="7.6">7.6 mm</quantity>
. Colour uniformly brown.
</paragraph>
<paragraph id="8BF1F629FFC8FFB9FF6FFC55761C25B3" blockId="17.[160,1156,844,1354]" pageId="17" pageNumber="1011">Anterior frontoclypeal process long and wide, slightly narrower than head, with angulate sinuation medially, with slightly asymmetric, apically truncated lateral lobes.</paragraph>
<paragraph id="8BF1F629FFC8FFB9FF6FFC1A71F42226" blockId="17.[160,1156,844,1354]" pageId="17" pageNumber="1011">Pronotum narrower than elytra, deeply excavated in middle. Lateral pronotal processes depressed, somewhat ridge-shaped, their hind margins with carinae reaching base of pronotum. Pronotum coarsely punctate to rugose except for very apices of lateral pronotal processes and a narrow area near anterior margin.</paragraph>
<paragraph id="8BF1F629FFC8FFB9FF6FFB65707B22BE" blockId="17.[160,1156,844,1354]" pageId="17" pageNumber="1011">
Aedeagus with short parameres: ratio phallobase width/paramere width 2.5. Apices of parameres acute in lateral view and wide in dorsal view (
<figureCitation id="1375EAACFFC8FFB9FCB1FB47711722E3" box="[798,891,1191,1215]" captionStart="Figure 5" captionStartId="9.[160,225,1398,1421]" captionTargetBox="[163,1153,143,1367]" captionTargetId="figure-19@9.[163,1153,142,1367]" captionTargetPageId="9" captionText="Figure 5. Paraegidium howdeni, holotype male (a, f, g); paratype female (b); paratype males (ce, h). Habitus in dorsal view (a, b); pronotum and head (ce); aedeagus in lateral view (f); parameres in dorsal view (g); endophallus (h)." figureDoi="http://doi.org/10.5281/zenodo.5181253" httpUri="https://zenodo.org/record/5181253/files/figure.png" pageId="17" pageNumber="1011">Figure 5</figureCitation>
(f, g)). Endophallus with a group of spinules (
<figureCitation id="1375EAACFFC8FFB9FE21FB2A738122BE" box="[398,493,1226,1250]" captionStart="Figure 5" captionStartId="9.[160,225,1398,1421]" captionTargetBox="[163,1153,143,1367]" captionTargetId="figure-19@9.[163,1153,142,1367]" captionTargetPageId="9" captionText="Figure 5. Paraegidium howdeni, holotype male (a, f, g); paratype female (b); paratype males (ce, h). Habitus in dorsal view (a, b); pronotum and head (ce); aedeagus in lateral view (f); parameres in dorsal view (g); endophallus (h)." figureDoi="http://doi.org/10.5281/zenodo.5181253" httpUri="https://zenodo.org/record/5181253/files/figure.png" pageId="17" pageNumber="1011">Figure 5</figureCitation>
(h)).
</paragraph>
<paragraph id="8BF1F629FFC8FFB9FF6FFB0D733F2316" blockId="17.[160,1156,844,1354]" pageId="17" pageNumber="1011">
<emphasis id="B93A2A3BFFC8FFB9FF6FFB0D73622359" box="[192,270,1261,1285]" italics="true" pageId="17" pageNumber="1011">Female</emphasis>
(
<figureCitation id="1375EAACFFC8FFB9FE8EFB0D73EE2359" box="[289,386,1261,1285]" captionStart="Figure 5" captionStartId="9.[160,225,1398,1421]" captionTargetBox="[163,1153,143,1367]" captionTargetId="figure-19@9.[163,1153,142,1367]" captionTargetPageId="9" captionText="Figure 5. Paraegidium howdeni, holotype male (a, f, g); paratype female (b); paratype males (ce, h). Habitus in dorsal view (a, b); pronotum and head (ce); aedeagus in lateral view (f); parameres in dorsal view (g); endophallus (h)." figureDoi="http://doi.org/10.5281/zenodo.5181253" httpUri="https://zenodo.org/record/5181253/files/figure.png" pageId="17" pageNumber="1011">Figure 5</figureCitation>
(b)) differs from male in having a distinct protibial spur, relatively smaller pronotum without processes and excavations, and in the absence of the frontoclypeal process.
</paragraph>
<paragraph id="8BF1F629FFC8FFB9FF0FFA99736223CE" blockId="17.[160,1155,1401,1530]" box="[160,270,1401,1426]" pageId="17" pageNumber="1011">
<emphasis id="B93A2A3BFFC8FFB9FF0FFA99736223CE" bold="true" box="[160,270,1401,1426]" italics="true" pageId="17" pageNumber="1011">Variation</emphasis>
</paragraph>
<paragraph id="8BF1F629FFC8FFB9FF0FFA7D700323A6" blockId="17.[160,1155,1401,1530]" pageId="17" pageNumber="1011">
Body length of the
<typeStatus id="54F5488BFFC8FFB9FED2FA7D738723E9" box="[381,491,1437,1461]" pageId="17" pageNumber="1011" type="paratype">paratypes</typeStatus>
varies from
<quantity id="4CB65BCCFFC8FFB9FDD7FA7D717923E9" box="[632,789,1437,1461]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" metricValueMax="7.5" metricValueMin="6.3" pageId="17" pageNumber="1011" unit="mm" value="6.9" valueMax="7.5" valueMin="6.3">6.3 to 7.5 mm</quantity>
(males) and from 7.0 to
<quantity id="4CB65BCCFFC8FFB9FB81FA7D76EF23E9" box="[1070,1155,1437,1461]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="17" pageNumber="1011" unit="mm" value="7.5">7.5 mm</quantity>
(females). The
<typeStatus id="54F5488BFFC8FFB9FEE6FA2073DB2384" box="[329,439,1472,1496]" pageId="17" pageNumber="1011" type="paratype">paratypes</typeStatus>
have less developed anterior frontoclypeal and pronotal processes (
<figureCitation id="1375EAACFFC8FFB9FF58FA02733823A6" box="[247,340,1506,1530]" captionStart="Figure 5" captionStartId="9.[160,225,1398,1421]" captionTargetBox="[163,1153,143,1367]" captionTargetId="figure-19@9.[163,1153,142,1367]" captionTargetPageId="9" captionText="Figure 5. Paraegidium howdeni, holotype male (a, f, g); paratype female (b); paratype males (ce, h). Habitus in dorsal view (a, b); pronotum and head (ce); aedeagus in lateral view (f); parameres in dorsal view (g); endophallus (h)." figureDoi="http://doi.org/10.5281/zenodo.5181253" httpUri="https://zenodo.org/record/5181253/files/figure.png" pageId="17" pageNumber="1011">Figure 5</figureCitation>
(ce)) than the
<typeStatus id="54F5488BFFC8FFB9FDABFA02700723A6" box="[516,619,1506,1530]" pageId="17" pageNumber="1011" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C354A5A2FFC8FFBAFF0FF9CA713A26B0" lastPageId="18" lastPageNumber="1012" pageId="17" pageNumber="1011" type="distribution">
<paragraph id="8BF1F629FFC8FFB9FF0FF9CA7340201F" blockId="17.[160,1156,1578,1707]" box="[160,300,1578,1603]" pageId="17" pageNumber="1011">
<emphasis id="B93A2A3BFFC8FFB9FF0FF9CA7340201F" bold="true" box="[160,300,1578,1603]" italics="true" pageId="17" pageNumber="1011">Distribution</emphasis>
</paragraph>
<paragraph id="8BF1F629FFC8FFBAFF0FF9AD713A26B0" blockId="17.[160,1156,1578,1707]" lastBlockId="18.[160,1155,142,236]" lastPageId="18" lastPageNumber="1012" pageId="17" pageNumber="1011">
The species as treated here has a wide range in central
<collectingCountry id="F359B6B9FFC8FFB9FCA3F9AD71242039" box="[780,840,1613,1637]" name="Brazil" pageId="17" pageNumber="1011">Brazil</collectingCountry>
and eastern
<collectingCountry id="F359B6B9FFC8FFB9FC76F9AD76602039" box="[985,1036,1613,1637]" name="Peru" pageId="17" pageNumber="1011">Peru</collectingCountry>
(
<figureCitation id="1375EAACFFC8FFB9FBB4F9AD761A2039" box="[1051,1142,1613,1637]" captionStart="Figure 7" captionStartId="11.[160,225,909,932]" captionTargetBox="[178,1138,142,879]" captionTargetId="figure-227@11.[178,1138,142,879]" captionTargetPageId="11" captionText="Figure 7. Paraegidium spp. locality map." figureDoi="http://doi.org/10.5281/zenodo.5181257" httpUri="https://zenodo.org/record/5181257/files/figure.png" pageId="17" pageNumber="1011">Figure 7</figureCitation>
). However, the specimens from the two outermost localities, Echarate (Peruvian Yungas, the westernmost locality for the genus) and Pacaraima (Guiana Shield, northernmost locality for the genus), are tentatively identified as
<taxonomicName id="4C4E8DAAFFCBFFBAFD4BFF6E713126FA" authority="Frolov &amp; Akhmetova &amp; Vaz-de-Mello, 2017" authorityName="Frolov &amp; Akhmetova &amp; Vaz-de-Mello" authorityYear="2017" box="[740,861,142,166]" class="Insecta" family="Hybosoridae" genus="Paraegidium" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="1012" phylum="Arthropoda" rank="species" species="howdeni" status="sp. nov.">
<emphasis id="B93A2A3BFFCBFFBAFD4BFF6E713126FA" box="[740,861,142,166]" italics="true" pageId="18" pageNumber="1012">P. howdeni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2099740FFCBFFBAFCCAFF6E71D226FA" box="[869,958,142,166]" pageId="18" pageNumber="1012" rank="species">sp. nov.</taxonomicNameLabel>
and thus are not included in the
<typeStatus id="54F5488BFFCBFFBAFEFEFF5173EF2695" box="[337,387,177,201]" pageId="18" pageNumber="1012">type</typeStatus>
series. They may belong to other, yet undescribed species, however additional material is needed for more reliable identification.
</paragraph>
</subSubSection>
<subSubSection id="C354A5A2FFCBFFBAFF0FFECD76782735" box="[160,1044,301,361]" pageId="18" pageNumber="1012" type="etymology">
<paragraph id="8BF1F629FFCBFFBAFF0FFECD7371271A" blockId="18.[160,1044,301,361]" box="[160,285,301,326]" pageId="18" pageNumber="1012">
<emphasis id="B93A2A3BFFCBFFBAFF0FFECD7371271A" bold="true" box="[160,285,301,326]" italics="true" pageId="18" pageNumber="1012">Etymology</emphasis>
</paragraph>
<paragraph id="8BF1F629FFCBFFBAFF0FFEB176782735" blockId="18.[160,1044,301,361]" box="[160,1044,337,361]" pageId="18" pageNumber="1012">The species is named after scarabaeoidologist Henry F. Howden (19252014).</paragraph>
</subSubSection>
</treatment>
</document>