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<document id="2FCDADA396CB6924B9137BE14F589866" ID-DOI="10.11646/zootaxa.4006.3.10" ID-GBIF-Dataset="43667874-dbfa-4674-a2b8-970cfc9afbc0" ID-ISSN="1175-5326" ID-Zenodo-Dep="243983" ID-ZooBank="5169CFEE-071C-40F1-9A16-4E898975EA33" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461215483631" checkinUser="plazi" docAuthor="Soulier-Perkins, Adeline &amp; Stroiński, Adam" docDate="2015" docId="03E1BE04F038FFFDFF7DB59FFD63F82C" docLanguage="en" docName="zt04006p600.pdf" docOrigin="Zootaxa 4006 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Binaluana Soulier-Perkins &amp; Stroiński, 2015, gen. nov." docType="treatment" docVersion="8" lastPageNumber="593" masterDocId="FFD8C67CF039FFFAFFEAB07DFFFBFFC4" masterDocTitle="A new lophopid genus as another piece in the biogeographical history puzzle of the family in the Sunda Shelf (Hemiptera: Fulgoromorpha: Lophopidae)" masterLastPageNumber="600" masterPageNumber="586" pageNumber="587" updateTime="1698602547519" updateUser="plazi">
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<mods:title id="A058A6048538BE8C17E83D296E041758">A new lophopid genus as another piece in the biogeographical history puzzle of the family in the Sunda Shelf (Hemiptera: Fulgoromorpha: Lophopidae)</mods:title>
</mods:titleInfo>
<mods:name id="42EB439F1A7C2F52D8C3D1154DA72A2F" type="personal">
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<mods:namePart id="DB52CD750DB7ACE190C1467418BB7960">Soulier-Perkins, Adeline</mods:namePart>
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<mods:namePart id="84E9FCFDAE55E36F3D504A0B421A1036">Stroiński, Adam</mods:namePart>
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<mods:title id="BC3EBB63C65E014480AEED1CBAF53342">Zootaxa</mods:title>
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<mods:part id="76052C70C8B97B1358275211D2872072">
<mods:date id="C37886E9FB332AB7B3629BA3C03E3A16">2015</mods:date>
<mods:detail id="7B3677746AA6E3D7C6B51A862D97CA6A" type="volume">
<mods:number id="4C6EC3BD9EC9C0EBE0866B0CCA547B58">4006</mods:number>
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<mods:number id="5F535F0F22ACA1B73D5B71C2B696B21F">3</mods:number>
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<mods:classification id="F6C665F16D5A6A55943A1433F98DA7A7">journal article</mods:classification>
<mods:identifier id="1CF68448066B48B7268210EB9642ED64" type="DOI">10.11646/zootaxa.4006.3.10</mods:identifier>
<mods:identifier id="BF459404A6EAEBE4747374B73B717675" type="GBIF-Dataset">43667874-dbfa-4674-a2b8-970cfc9afbc0</mods:identifier>
<mods:identifier id="4B31C581AD8799FB26FEBD053D63E7DA" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03E1BE04F038FFFDFF7DB59FFD63F82C" ID-DOI="http://doi.org/10.5281/zenodo.6122419" ID-GBIF-Taxon="119630768" ID-Zenodo-Dep="6122419" LSID="urn:lsid:plazi:treatment:03E1BE04F038FFFDFF7DB59FFD63F82C" httpUri="http://treatment.plazi.org/id/03E1BE04F038FFFDFF7DB59FFD63F82C" lastPageId="7" lastPageNumber="593" pageId="1" pageNumber="587">
<subSubSection id="C3525C99F038FFFBFF7DB59FFEF7F9DF" pageId="1" pageNumber="587" type="nomenclature">
<paragraph id="8BF70F12F038FFFBFF7DB59FFE76FA39" blockId="1.[151,397,1506,1563]" box="[151,397,1506,1533]" pageId="1" pageNumber="587">
<heading id="D0BFB87EF038FFFBFF7DB59FFE76FA39" bold="true" box="[151,397,1506,1533]" fontSize="11" level="1" pageId="1" pageNumber="587" reason="1">
<emphasis id="B93CD300F038FFFBFF7DB59FFE76FA39" bold="true" box="[151,397,1506,1533]" pageId="1" pageNumber="587">
<taxonomicName id="4C487491F038FFFBFF7DB59FFEE3FA38" box="[151,280,1506,1532]" class="Insecta" family="Lophopidae" genus="Binaluana" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="587" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B93CD300F038FFFBFF7DB59FFEE3FA38" bold="true" box="[151,280,1506,1532]" italics="true" pageId="1" pageNumber="587">Binaluana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F6E7BF038FFFBFEF5B59EFE76FA39" box="[287,397,1507,1533]" pageId="1" pageNumber="587" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF70F12F038FFFBFF7DB678FEF7F9DF" blockId="1.[151,397,1506,1563]" box="[151,268,1541,1563]" pageId="1" pageNumber="587">
(
<figureCitation id="13731397F038FFFBFF74B678FEFFF9DF" box="[158,260,1541,1563]" captionStart-0="FIGURES 1 – 5" captionStart-1="FIGURES 6 – 11" captionStart-2="FIGURES 12 – 17" captionStart-3="FIGURES 18 – 23" captionStart-4="FIGURES 24 – 29" captionStart-5="FIGURES 30 – 35" captionStart-6="FIGURES 36 – 41" captionStart-7="FIGURES 42 – 47" captionStart-8="FIGURES 48 – 52" captionStart-9="FIGURES 53 – 60" captionStartId-0="2.[151,264,1906,1929]" captionStartId-1="3.[151,264,1914,1937]" captionStartId-2="4.[151,264,1978,2001]" captionStartId-3="5.[151,264,1684,1707]" captionStartId-4="6.[151,262,1951,1974]" captionStartId-5="7.[151,264,1689,1712]" captionStartId-6="8.[151,264,1647,1670]" captionStartId-7="9.[151,264,1640,1663]" captionStartId-8="10.[151,264,1781,1804]" captionStartId-9="11.[151,264,1848,1871]" captionTargetBox-0="[169,1428,491,1875]" captionTargetBox-1="[171,1425,203,1873]" captionTargetBox-2="[160,1420,523,1955]" captionTargetBox-3="[151,1436,193,1653]" captionTargetBox-4="[173,1407,514,1916]" captionTargetBox-5="[151,1436,193,1655]" captionTargetBox-6="[173,1420,194,1613]" captionTargetBox-7="[151,1436,159,1620]" captionTargetBox-8="[151,1436,256,1716]" captionTargetBox-9="[175,1403,205,1810]" captionTargetId-0="figure@2.[151,1436,481,1885]" captionTargetId-1="figure@3.[151,1436,193,1893]" captionTargetId-2="figure@4.[160,1420,523,1964]" captionTargetId-3="figure@5.[151,1436,193,1664]" captionTargetId-4="figure@6.[173,1407,514,1926]" captionTargetId-5="figure@7.[151,1436,193,1664]" captionTargetId-6="figure@8.[173,1420,194,1621]" captionTargetId-7="figure@9.[151,1436,159,1625]" captionTargetId-8="figure@10.[151,1436,256,1719]" captionTargetId-9="figure@11.[151,1436,193,1827]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionTargetPageId-6="8" captionTargetPageId-7="9" captionTargetPageId-8="10" captionTargetPageId-9="11" captionText-0="FIGURES 1 – 5. Binaluana emarginata gen. et sp. nov., female. (1) Habitus, lateral view; (2) same, dorsal view; (3) anterior part of body, lateral view; (4) anterior part, frontal view; (5) anterior part of body, dorsal view." captionText-1="FIGURES 6 – 11. Binaluana emarginata gen. et sp. nov., female. (6) Head, rostrum and legs, ventral view; (7) abdomen and hind legs, ventral view; (8) labels; (9 – 11) abdomen: (9) dorsal view, (10) ventral view, (11) posterior view." captionText-2="FIGURES 12 – 17. Binaluana emarginata gen. et sp. nov., female, SEM images. (12 – 13) Anterior part of body: (12) dorsal view, (13) dorso-lateral view; (14) vertex, dorsal view; (15) head, frontal view; (16 – 17) frons, frontal view." captionText-3="FIGURES 18 – 23. Binaluana emarginata gen. et sp. nov., female, SEM images. (18) Anterior part of body, lateral view; (19) head and pronotum, lateral view; (20) upper part of head, lateral view; (21) pronotum and mesonotum, lateral view; (22) lower part of head and pronotum, lateral view; (23) upper part of pronotum and mesonotum, lateral view." captionText-4="FIGURES 24 – 29. Binaluana emarginata gen. et sp. nov., female, SEM images. (24 – 27) Antenna: (24 – 25) dorsal view, (26) dorso-lateral view, (27) apical view; (28 – 29) plate organs." captionText-5="FIGURES 30 – 35. Binaluana emarginata gen. et sp. nov., female, SEM images. (30 – 35) Tegmina." captionText-6="FIGURES 36 – 41. Binaluana emarginata gen. et sp. nov., female, SEM images. (36) Fore femur and tibia, ventral view; (37 – 40) hind tibia and tarsal segments; (41) rostrum, ventral view." captionText-7="FIGURES 42 – 47. Binaluana emarginata gen. et sp. nov., female, SEM images. (42 – 43) Abdomen: (42) dorso-lateral view, (43) lateral view; (44 – 45) terminalia: (44) dorso-lateral view, (45) lateral view; (46) anal tube, posterior view; (47) median part of anal tube, dorso-lateral view." captionText-8="FIGURES 48 – 52. Binaluana emarginata gen. et sp. nov., female, SEM images. (48) Abdomen, ventral view; (50) terminalia, ventral view; (49) same, lateral view; (50 – 51) gonapophysis VIII: (50) ventral view, (51) posterior view; (52) gonoplac with wax secretion, lateral view." captionText-9="FIGURES 53 – 60. Binaluana emarginata gen. et sp. nov., female. (53 – 55) Pregenital sternite: (53) ventral view, (54) ventroposterior view, (55) posterior view; (56) anal tube, dorsal view; (57) gonoplac and gonapophysis VIII, external view (lateral); (58) bursa copulatrix, lateral view; (59 – 60) spermatheca." httpUri-0="https://zenodo.org/record/243984/files/figure.png" httpUri-1="https://zenodo.org/record/243985/files/figure.png" httpUri-2="https://zenodo.org/record/243986/files/figure.png" httpUri-3="https://zenodo.org/record/243987/files/figure.png" httpUri-4="https://zenodo.org/record/243988/files/figure.png" httpUri-5="https://zenodo.org/record/243989/files/figure.png" httpUri-6="https://zenodo.org/record/243990/files/figure.png" httpUri-7="https://zenodo.org/record/243991/files/figure.png" httpUri-8="https://zenodo.org/record/243992/files/figure.png" httpUri-9="https://zenodo.org/record/243993/files/figure.png" pageId="1" pageNumber="587">Figs 1–60</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3525C99F038FFFBFF7DB639FCEBF99D" box="[151,784,1603,1625]" pageId="1" pageNumber="587" type="materials_examined">
<paragraph id="8BF70F12F038FFFBFF7DB639FCEBF99D" blockId="1.[151,784,1603,1625]" box="[151,784,1603,1625]" pageId="1" pageNumber="587">
<typeStatus id="54F3B1B0F038FFFBFF7DB639FF32F99D" box="[151,201,1604,1625]" pageId="1" pageNumber="587">Type</typeStatus>
species:
<taxonomicName id="4C487491F038FFFBFECCB63EFDF1F99D" box="[294,522,1603,1625]" class="Insecta" family="Lophopidae" genus="Binaluana" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="587" phylum="Arthropoda" rank="species" species="emarginata" status="sp. nov.">
<emphasis id="B93CD300F038FFFBFECCB63EFDF1F99D" box="[294,522,1603,1625]" italics="true" pageId="1" pageNumber="587">Binaluana emarginata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F6E7BF038FFFBFDFAB639FD9AF99D" box="[528,609,1604,1625]" pageId="1" pageNumber="587" rank="species">sp. nov.</taxonomicNameLabel>
, here designated.
</paragraph>
</subSubSection>
<subSubSection id="C3525C99F038FFFBFF7DB6EBFDD1F917" pageId="1" pageNumber="587" type="etymology">
<paragraph id="8BF70F12F038FFFBFF7DB6EBFDD1F917" blockId="1.[151,1437,1686,2035]" pageId="1" pageNumber="587">
<emphasis id="B93CD300F038FFFBFF7DB6EBFEEDF96B" bold="true" box="[151,278,1686,1711]" pageId="1" pageNumber="587">Etymology</emphasis>
. Melichar intended to name this genus after the location where it was found, Binaluan. We kept his name
<taxonomicName id="4C487491F038FFFBFF30B6C7FEB4F916" box="[218,335,1722,1746]" class="Insecta" family="Lophopidae" genus="Binaluana" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="587" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B93CD300F038FFFBFF30B6C7FEB4F916" box="[218,335,1722,1746]" italics="true" pageId="1" pageNumber="587">Binaluana</emphasis>
</taxonomicName>
. Gender: feminine.
</paragraph>
</subSubSection>
<subSubSection id="C3525C99F038FFFBFF2DB6A3FB33F8FB" pageId="1" pageNumber="587" type="diagnosis">
<paragraph id="8BF70F12F038FFFBFF2DB6A3FB33F8FB" blockId="1.[151,1437,1686,2035]" pageId="1" pageNumber="587">
<emphasis id="B93CD300F038FFFBFF2DB6A3FEC1F933" bold="true" box="[199,314,1758,1783]" pageId="1" pageNumber="587">Diagnosis</emphasis>
.
<taxonomicName id="4C487491F038FFFBFEA0B6A3FE44F932" box="[330,447,1758,1782]" class="Insecta" family="Lophopidae" genus="Binaluana" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="587" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B93CD300F038FFFBFEA0B6A3FE44F932" box="[330,447,1758,1782]" italics="true" pageId="1" pageNumber="587">Binaluana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F6E7BF038FFFBFE23B6A2FDCAF932" box="[457,561,1759,1783]" pageId="1" pageNumber="587" rank="genus">gen. nov.</taxonomicNameLabel>
similar to
<taxonomicName id="4C487491F038FFFBFD5BB6A3FD0CF932" box="[689,759,1758,1782]" class="Insecta" family="Lophopidae" genus="Zeleja" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="587" phylum="Arthropoda" rank="genus">
<emphasis id="B93CD300F038FFFBFD5BB6A3FD0CF932" box="[689,759,1758,1782]" italics="true" pageId="1" pageNumber="587">Zeleja</emphasis>
</taxonomicName>
and
<taxonomicName id="4C487491F038FFFBFCD8B6A2FC83F932" box="[818,888,1759,1782]" class="Insecta" family="Lophopidae" genus="Bisma" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="587" phylum="Arthropoda" rank="genus">
<emphasis id="B93CD300F038FFFBFCD8B6A2FC83F932" box="[818,888,1759,1782]" italics="true" pageId="1" pageNumber="587">Bisma</emphasis>
</taxonomicName>
, but
<taxonomicName id="4C487491F038FFFBFC5CB6A2FC07F932" box="[950,1020,1759,1782]" class="Insecta" family="Lophopidae" genus="Bisma" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="587" phylum="Arthropoda" rank="genus">
<emphasis id="B93CD300F038FFFBFC5CB6A2FC07F932" box="[950,1020,1759,1782]" italics="true" pageId="1" pageNumber="587">Bisma</emphasis>
</taxonomicName>
presents some carinae on the frons, when frons in
<taxonomicName id="4C487491F038FFFBFEB8B77FFE3CF8DE" box="[338,455,1794,1818]" class="Insecta" family="Lophopidae" genus="Binaluana" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="587" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B93CD300F038FFFBFEB8B77FFE3CF8DE" box="[338,455,1794,1818]" italics="true" pageId="1" pageNumber="587">Binaluana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C487491F038FFFBFDE5B77FFDAEF8DE" box="[527,597,1794,1818]" class="Insecta" family="Lophopidae" genus="Zeleja" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="587" phylum="Arthropoda" rank="genus">
<emphasis id="B93CD300F038FFFBFDE5B77FFDAEF8DE" box="[527,597,1794,1818]" italics="true" pageId="1" pageNumber="587">Zeleja</emphasis>
</taxonomicName>
presents none.
<taxonomicName id="4C487491F038FFFBFCF4B77FFC68F8DE" box="[798,915,1794,1818]" class="Insecta" family="Lophopidae" genus="Binaluana" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="587" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B93CD300F038FFFBFCF4B77FFC68F8DE" box="[798,915,1794,1818]" italics="true" pageId="1" pageNumber="587">Binaluana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C487491F038FFFBFC31B77EFBDAF8DE" box="[987,1057,1795,1818]" class="Insecta" family="Lophopidae" genus="Bisma" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="587" phylum="Arthropoda" rank="genus">
<emphasis id="B93CD300F038FFFBFC31B77EFBDAF8DE" box="[987,1057,1795,1818]" italics="true" pageId="1" pageNumber="587">Bisma</emphasis>
</taxonomicName>
have a shorter prothorax than mesothorax, with a clear produced disc, when
<taxonomicName id="4C487491F038FFFBFD75B75BFD1EF8FA" box="[671,741,1830,1854]" class="Insecta" family="Lophopidae" genus="Zeleja" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="587" phylum="Arthropoda" rank="genus">
<emphasis id="B93CD300F038FFFBFD75B75BFD1EF8FA" box="[671,741,1830,1854]" italics="true" pageId="1" pageNumber="587">Zeleja</emphasis>
</taxonomicName>
has a pro- and mesothorax equal in length.
</paragraph>
</subSubSection>
<subSubSection id="C3525C99F038FFFDFF2DB737FD6BF80C" lastPageId="7" lastPageNumber="593" pageId="1" pageNumber="587" type="description">
<paragraph id="8BF70F12F038FFFBFF2DB737FCEBF8A7" blockId="1.[151,1437,1686,2035]" box="[199,784,1866,1891]" pageId="1" pageNumber="587">
<emphasis id="B93CD300F038FFFBFF2DB737FEAAF8A7" bold="true" box="[199,337,1866,1891]" pageId="1" pageNumber="587">Description</emphasis>
. Body elongate and narrow (
<figureCitation id="13731397F038FFFBFD7DB737FCF8F8A7" box="[663,771,1866,1891]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,264,1906,1929]" captionTargetBox="[169,1428,491,1875]" captionTargetId="figure@2.[151,1436,481,1885]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Binaluana emarginata gen. et sp. nov., female. (1) Habitus, lateral view; (2) same, dorsal view; (3) anterior part of body, lateral view; (4) anterior part, frontal view; (5) anterior part of body, dorsal view." httpUri="https://zenodo.org/record/243984/files/figure.png" pageId="1" pageNumber="587">Figs. 1–2</figureCitation>
).
</paragraph>
<paragraph id="8BF70F12F038FFF8FF2DB712FC02FE68" blockId="1.[151,1437,1686,2035]" lastBlockId="2.[151,1436,151,429]" lastPageId="2" lastPageNumber="588" pageId="1" pageNumber="587">
HEAD. Head with compound eyes narrower than prothorax (
<figureCitation id="13731397F038FFFBFC6AB713FC12F843" box="[896,1001,1902,1927]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,264,1906,1929]" captionTargetBox="[169,1428,491,1875]" captionTargetId="figure@2.[151,1436,481,1885]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Binaluana emarginata gen. et sp. nov., female. (1) Habitus, lateral view; (2) same, dorsal view; (3) anterior part of body, lateral view; (4) anterior part, frontal view; (5) anterior part of body, dorsal view." httpUri="https://zenodo.org/record/243984/files/figure.png" pageId="1" pageNumber="587">Figs 2, 5</figureCitation>
&amp;
<figureCitation id="13731397F038FFFBFBF9B712FBC9F843" box="[1043,1074,1903,1927]" captionStart="FIGURES 12 – 17" captionStartId="4.[151,264,1978,2001]" captionTargetBox="[160,1420,523,1955]" captionTargetId="figure@4.[160,1420,523,1964]" captionTargetPageId="4" captionText="FIGURES 12 – 17. Binaluana emarginata gen. et sp. nov., female, SEM images. (12 – 13) Anterior part of body: (12) dorsal view, (13) dorso-lateral view; (14) vertex, dorsal view; (15) head, frontal view; (16 – 17) frons, frontal view." httpUri="https://zenodo.org/record/243986/files/figure.png" pageId="1" pageNumber="587">12</figureCitation>
). Vertex longer than wide, flat, without median carina, lateral margins carinate, parallel and elevated, anterior margin obsolete (
<figureCitation id="13731397F038FFFBFB01B7EFFACCF86F" box="[1259,1335,1938,1963]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,264,1906,1929]" captionTargetBox="[169,1428,491,1875]" captionTargetId="figure@2.[151,1436,481,1885]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Binaluana emarginata gen. et sp. nov., female. (1) Habitus, lateral view; (2) same, dorsal view; (3) anterior part of body, lateral view; (4) anterior part, frontal view; (5) anterior part of body, dorsal view." httpUri="https://zenodo.org/record/243984/files/figure.png" pageId="1" pageNumber="587">Figs 4</figureCitation>
,
<figureCitation id="13731397F038FFFBFAACB7EFFA6BF86F" box="[1350,1424,1938,1963]" captionStart="FIGURES 12 – 17" captionStartId="4.[151,264,1978,2001]" captionTargetBox="[160,1420,523,1955]" captionTargetId="figure@4.[160,1420,523,1964]" captionTargetPageId="4" captionText="FIGURES 12 – 17. Binaluana emarginata gen. et sp. nov., female, SEM images. (12 – 13) Anterior part of body: (12) dorsal view, (13) dorso-lateral view; (14) vertex, dorsal view; (15) head, frontal view; (16 – 17) frons, frontal view." httpUri="https://zenodo.org/record/243986/files/figure.png" pageId="1" pageNumber="587">15–17</figureCitation>
). Frons generally flat with the exception of the protuberance just bellow the upper margin, no carinae, lateral margins carinate and continuous from the vertex to the frontoclypeal margin. Frons bottle-shape, with widest part along the frontoclypeal suture. In lateral view, top of the head higher than the surface of pro- and mesothorax, compound eyes rounded, flattened postero-ventrally, with margin cut out ventrally; ocelli present; foramen of antenna not touching the compound eyes; ocellar and genal carina absent (
<figureCitation id="13731397F03BFFF8FCB1B0A2FC39FF3C" box="[859,962,223,248]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,264,1906,1929]" captionTargetBox="[169,1428,491,1875]" captionTargetId="figure@2.[151,1436,481,1885]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Binaluana emarginata gen. et sp. nov., female. (1) Habitus, lateral view; (2) same, dorsal view; (3) anterior part of body, lateral view; (4) anterior part, frontal view; (5) anterior part of body, dorsal view." httpUri="https://zenodo.org/record/243984/files/figure.png" pageId="2" pageNumber="588">Figs 1, 3</figureCitation>
,
<figureCitation id="13731397F03BFFF8FC3AB09DFC15FF3C" box="[976,1006,224,248]" captionStart="FIGURES 12 – 17" captionStartId="4.[151,264,1978,2001]" captionTargetBox="[160,1420,523,1955]" captionTargetId="figure@4.[160,1420,523,1964]" captionTargetPageId="4" captionText="FIGURES 12 – 17. Binaluana emarginata gen. et sp. nov., female, SEM images. (12 – 13) Anterior part of body: (12) dorsal view, (13) dorso-lateral view; (14) vertex, dorsal view; (15) head, frontal view; (16 – 17) frons, frontal view." httpUri="https://zenodo.org/record/243986/files/figure.png" pageId="2" pageNumber="588">13</figureCitation>
,
<figureCitation id="13731397F03BFFF8FC11B09DFBBEFF3C" box="[1019,1093,224,248]" captionStart="FIGURES 18 – 23" captionStartId="5.[151,264,1684,1707]" captionTargetBox="[151,1436,193,1653]" captionTargetId="figure@5.[151,1436,193,1664]" captionTargetPageId="5" captionText="FIGURES 18 – 23. Binaluana emarginata gen. et sp. nov., female, SEM images. (18) Anterior part of body, lateral view; (19) head and pronotum, lateral view; (20) upper part of head, lateral view; (21) pronotum and mesonotum, lateral view; (22) lower part of head and pronotum, lateral view; (23) upper part of pronotum and mesonotum, lateral view." httpUri="https://zenodo.org/record/243987/files/figure.png" pageId="2" pageNumber="588">18–22</figureCitation>
). Carinated line starting at the edge of frontoclypeal suture and lateral frontal margin, running toward the ocellar and disappearing (
<figureCitation id="13731397F03BFFF8FAEBB179FA9DFED8" box="[1281,1382,260,285]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,264,1906,1929]" captionTargetBox="[169,1428,491,1875]" captionTargetId="figure@2.[151,1436,481,1885]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Binaluana emarginata gen. et sp. nov., female. (1) Habitus, lateral view; (2) same, dorsal view; (3) anterior part of body, lateral view; (4) anterior part, frontal view; (5) anterior part of body, dorsal view." httpUri="https://zenodo.org/record/243984/files/figure.png" pageId="2" pageNumber="588">Figs 1–2</figureCitation>
,
<figureCitation id="13731397F03BFFF8FA98B179FF4CFE84" captionStart="FIGURES 18 – 23" captionStartId="5.[151,264,1684,1707]" captionTargetBox="[151,1436,193,1653]" captionTargetId="figure@5.[151,1436,193,1664]" captionTargetPageId="5" captionText="FIGURES 18 – 23. Binaluana emarginata gen. et sp. nov., female, SEM images. (18) Anterior part of body, lateral view; (19) head and pronotum, lateral view; (20) upper part of head, lateral view; (21) pronotum and mesonotum, lateral view; (22) lower part of head and pronotum, lateral view; (23) upper part of pronotum and mesonotum, lateral view." httpUri="https://zenodo.org/record/243987/files/figure.png" pageId="2" pageNumber="588">18– 22</figureCitation>
). Pedicel of antenna twice longer than wide (
<figureCitation id="13731397F03BFFF8FD32B15AFCB9FE84" box="[728,834,295,320]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,264,1906,1929]" captionTargetBox="[169,1428,491,1875]" captionTargetId="figure@2.[151,1436,481,1885]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Binaluana emarginata gen. et sp. nov., female. (1) Habitus, lateral view; (2) same, dorsal view; (3) anterior part of body, lateral view; (4) anterior part, frontal view; (5) anterior part of body, dorsal view." httpUri="https://zenodo.org/record/243984/files/figure.png" pageId="2" pageNumber="588">Figs 4–5</figureCitation>
,
<figureCitation id="13731397F03BFFF8FCB9B155FC65FE84" box="[851,926,296,320]" captionStart="FIGURES 24 – 29" captionStartId="6.[151,262,1951,1974]" captionTargetBox="[173,1407,514,1916]" captionTargetId="figure@6.[173,1407,514,1926]" captionTargetPageId="6" captionText="FIGURES 24 – 29. Binaluana emarginata gen. et sp. nov., female, SEM images. (24 – 27) Antenna: (24 – 25) dorsal view, (26) dorso-lateral view, (27) apical view; (28 – 29) plate organs." httpUri="https://zenodo.org/record/243988/files/figure.png" pageId="2" pageNumber="588">24–27</figureCitation>
). Sensory plate organs multi-petals shaped, surrounded by strong denticules, present on whole pedicel, arranged in lines (
<figureCitation id="13731397F03BFFF8FC11B131FB85FEA0" box="[1019,1150,332,357]" captionStart="FIGURES 24 – 29" captionStartId="6.[151,262,1951,1974]" captionTargetBox="[173,1407,514,1916]" captionTargetId="figure@6.[173,1407,514,1926]" captionTargetPageId="6" captionText="FIGURES 24 – 29. Binaluana emarginata gen. et sp. nov., female, SEM images. (24 – 27) Antenna: (24 – 25) dorsal view, (26) dorso-lateral view, (27) apical view; (28 – 29) plate organs." httpUri="https://zenodo.org/record/243988/files/figure.png" pageId="2" pageNumber="588">Figs 24–29</figureCitation>
); trichoid sensilla present (
<typeStatus id="54F3B1B0F03BFFF8FF74B10DFF2BFE4C" box="[158,208,368,392]" pageId="2" pageNumber="588">type</typeStatus>
1 and 2). Clypeus strongly carinate laterally without median carina (
<figureCitation id="13731397F03BFFF8FBEBB112FBB5FE4C" box="[1025,1102,367,392]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,264,1906,1929]" captionTargetBox="[169,1428,491,1875]" captionTargetId="figure@2.[151,1436,481,1885]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Binaluana emarginata gen. et sp. nov., female. (1) Habitus, lateral view; (2) same, dorsal view; (3) anterior part of body, lateral view; (4) anterior part, frontal view; (5) anterior part of body, dorsal view." httpUri="https://zenodo.org/record/243984/files/figure.png" pageId="2" pageNumber="588">Figs 4</figureCitation>
,
<figureCitation id="13731397F03BFFF8FBB4B112FB97FE4C" box="[1118,1132,367,392]" captionStart="FIGURES 6 – 11" captionStartId="3.[151,264,1914,1937]" captionTargetBox="[171,1425,203,1873]" captionTargetId="figure@3.[151,1436,193,1893]" captionTargetPageId="3" captionText="FIGURES 6 – 11. Binaluana emarginata gen. et sp. nov., female. (6) Head, rostrum and legs, ventral view; (7) abdomen and hind legs, ventral view; (8) labels; (9 – 11) abdomen: (9) dorsal view, (10) ventral view, (11) posterior view." httpUri="https://zenodo.org/record/243985/files/figure.png" pageId="2" pageNumber="588">6</figureCitation>
&amp;
<figureCitation id="13731397F03BFFF8FB72B112FB4CFE4C" box="[1176,1207,367,392]" captionStart="FIGURES 12 – 17" captionStartId="4.[151,264,1978,2001]" captionTargetBox="[160,1420,523,1955]" captionTargetId="figure@4.[160,1420,523,1964]" captionTargetPageId="4" captionText="FIGURES 12 – 17. Binaluana emarginata gen. et sp. nov., female, SEM images. (12 – 13) Anterior part of body: (12) dorsal view, (13) dorso-lateral view; (14) vertex, dorsal view; (15) head, frontal view; (16 – 17) frons, frontal view." httpUri="https://zenodo.org/record/243986/files/figure.png" pageId="2" pageNumber="588">15</figureCitation>
). Rostrum reaching metacoxa, apical segment distinctly shorter than subapical one (
<figureCitation id="13731397F03BFFF8FC88B1E9FC53FE69" box="[866,936,404,429]" captionStart="FIGURES 6 – 11" captionStartId="3.[151,264,1914,1937]" captionTargetBox="[171,1425,203,1873]" captionTargetId="figure@3.[151,1436,193,1893]" captionTargetPageId="3" captionText="FIGURES 6 – 11. Binaluana emarginata gen. et sp. nov., female. (6) Head, rostrum and legs, ventral view; (7) abdomen and hind legs, ventral view; (8) labels; (9 – 11) abdomen: (9) dorsal view, (10) ventral view, (11) posterior view." httpUri="https://zenodo.org/record/243985/files/figure.png" pageId="2" pageNumber="588">Figs 6</figureCitation>
&amp;
<figureCitation id="13731397F03BFFF8FC26B1E9FC10FE68" box="[972,1003,404,428]" captionStart="FIGURES 36 – 41" captionStartId="8.[151,264,1647,1670]" captionTargetBox="[173,1420,194,1613]" captionTargetId="figure@8.[173,1420,194,1621]" captionTargetPageId="8" captionText="FIGURES 36 – 41. Binaluana emarginata gen. et sp. nov., female, SEM images. (36) Fore femur and tibia, ventral view; (37 – 40) hind tibia and tarsal segments; (41) rostrum, ventral view." httpUri="https://zenodo.org/record/243990/files/figure.png" pageId="2" pageNumber="588">41</figureCitation>
).
</paragraph>
<caption id="DF375F9AF03BFFF8FF7DB70FFBAAF863" httpUri="https://zenodo.org/record/243984/files/figure.png" pageId="2" pageNumber="588" targetBox="[169,1428,491,1875]" targetPageId="2">
<paragraph id="8BF70F12F03BFFF8FF7DB70FFBAAF863" blockId="2.[151,1436,1906,1959]" pageId="2" pageNumber="588">
<emphasis id="B93CD300F03BFFF8FF7DB70FFECEF84C" bold="true" box="[151,309,1906,1929]" pageId="2" pageNumber="588">FIGURES 1–5</emphasis>
.
<emphasis id="B93CD300F03BFFF8FEA8B70FFD2BF84C" bold="true" box="[322,720,1906,1929]" pageId="2" pageNumber="588">
<taxonomicName id="4C487491F03BFFF8FEA8B70FFDD4F84C" box="[322,559,1906,1929]" class="Insecta" family="Lophopidae" genus="Binaluana" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="588" phylum="Arthropoda" rank="species" species="emarginata">
<emphasis id="B93CD300F03BFFF8FEA8B70FFDD4F84C" bold="true" box="[322,559,1906,1929]" italics="true" pageId="2" pageNumber="588">Binaluana emarginata</emphasis>
</taxonomicName>
gen. et sp. nov
</emphasis>
., female. (1) Habitus, lateral view; (2) same, dorsal view; (3) anterior part of body, lateral view; (4) anterior part, frontal view; (5) anterior part of body, dorsal view.
</paragraph>
</caption>
<caption id="DF375F9AF03AFFF9FF7DB707FB35F874" httpUri="https://zenodo.org/record/243985/files/figure.png" pageId="3" pageNumber="589" targetBox="[171,1425,203,1873]" targetPageId="3">
<paragraph id="8BF70F12F03AFFF9FF7DB707FB35F874" blockId="3.[151,1435,1914,1968]" pageId="3" pageNumber="589">
<emphasis id="B93CD300F03AFFF9FF7DB707FEBBF854" bold="true" box="[151,320,1914,1937]" pageId="3" pageNumber="589">FIGURES 6–11</emphasis>
.
<emphasis id="B93CD300F03AFFF9FEA7B707FD27F854" bold="true" box="[333,732,1914,1937]" pageId="3" pageNumber="589">
<taxonomicName id="4C487491F03AFFF9FEA7B707FDC1F854" box="[333,570,1914,1937]" class="Insecta" family="Lophopidae" genus="Binaluana" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="589" phylum="Arthropoda" rank="species" species="emarginata">
<emphasis id="B93CD300F03AFFF9FEA7B707FDC1F854" bold="true" box="[333,570,1914,1937]" italics="true" pageId="3" pageNumber="589">Binaluana emarginata</emphasis>
</taxonomicName>
gen. et sp. nov
</emphasis>
., female. (6) Head, rostrum and legs, ventral view; (7) abdomen and hind legs, ventral view; (8) labels; (9–11) abdomen: (9) dorsal view, (10) ventral view, (11) posterior view.
</paragraph>
</caption>
<paragraph id="8BF70F12F03DFFFEFF2DB0E5FF15FE14" blockId="4.[151,1437,151,464]" pageId="4" pageNumber="590">
THORAX. Pronotum anteriorly rounded, median disc tricarinate, carinae connected anteriorly and reaching posterior margin (
<figureCitation id="13731397F03DFFFEFE82B0C1FE2DFF11" box="[360,470,188,213]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,264,1906,1929]" captionTargetBox="[169,1428,491,1875]" captionTargetId="figure@2.[151,1436,481,1885]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Binaluana emarginata gen. et sp. nov., female. (1) Habitus, lateral view; (2) same, dorsal view; (3) anterior part of body, lateral view; (4) anterior part, frontal view; (5) anterior part of body, dorsal view." httpUri="https://zenodo.org/record/243984/files/figure.png" pageId="4" pageNumber="590">Figs 2, 5</figureCitation>
,
<figureCitation id="13731397F03DFFFEFE0CB0C1FDCBFF10" box="[486,560,188,212]" captionStart="FIGURES 12 – 17" captionStartId="4.[151,264,1978,2001]" captionTargetBox="[160,1420,523,1955]" captionTargetId="figure@4.[160,1420,523,1964]" captionTargetPageId="4" captionText="FIGURES 12 – 17. Binaluana emarginata gen. et sp. nov., female, SEM images. (12 – 13) Anterior part of body: (12) dorsal view, (13) dorso-lateral view; (14) vertex, dorsal view; (15) head, frontal view; (16 – 17) frons, frontal view." httpUri="https://zenodo.org/record/243986/files/figure.png" pageId="4" pageNumber="590">12–13</figureCitation>
). Median disc elevated and flattened, slightly protruding anteriorly. Strong carina present on lateral part of pronotum (
<figureCitation id="13731397F03DFFFEFD9CB0A2FCB6FF3C" box="[630,845,223,248]" captionStart="FIGURES 18 – 23" captionStartId="5.[151,264,1684,1707]" captionTargetBox="[151,1436,193,1653]" captionTargetId="figure@5.[151,1436,193,1664]" captionTargetPageId="5" captionText="FIGURES 18 – 23. Binaluana emarginata gen. et sp. nov., female, SEM images. (18) Anterior part of body, lateral view; (19) head and pronotum, lateral view; (20) upper part of head, lateral view; (21) pronotum and mesonotum, lateral view; (22) lower part of head and pronotum, lateral view; (23) upper part of pronotum and mesonotum, lateral view." httpUri="https://zenodo.org/record/243987/files/figure.png" pageId="4" pageNumber="590">Figs 18–19, 21–22</figureCitation>
). Posterior margin widely rounded. Mesothorax with a single median carina starting on the anterior margin but not reaching the scutelum. Pro- and mesothoracic legs flattened, but not foliaceous, with femur and tibia of the same length, prothoracic leg more flattened than mesothoracic one (
<figureCitation id="13731397F03DFFFEFE9AB131FE40FEA1" box="[368,443,332,357]" captionStart="FIGURES 6 – 11" captionStartId="3.[151,264,1914,1937]" captionTargetBox="[171,1425,203,1873]" captionTargetId="figure@3.[151,1436,193,1893]" captionTargetPageId="3" captionText="FIGURES 6 – 11. Binaluana emarginata gen. et sp. nov., female. (6) Head, rostrum and legs, ventral view; (7) abdomen and hind legs, ventral view; (8) labels; (9 – 11) abdomen: (9) dorsal view, (10) ventral view, (11) posterior view." httpUri="https://zenodo.org/record/243985/files/figure.png" pageId="4" pageNumber="590">Figs 6</figureCitation>
,
<figureCitation id="13731397F03DFFFEFE20B131FE11FEA1" box="[458,490,332,357]" captionStart="FIGURES 36 – 41" captionStartId="8.[151,264,1647,1670]" captionTargetBox="[173,1420,194,1613]" captionTargetId="figure@8.[173,1420,194,1621]" captionTargetPageId="8" captionText="FIGURES 36 – 41. Binaluana emarginata gen. et sp. nov., female, SEM images. (36) Fore femur and tibia, ventral view; (37 – 40) hind tibia and tarsal segments; (41) rostrum, ventral view." httpUri="https://zenodo.org/record/243990/files/figure.png" pageId="4" pageNumber="590">36</figureCitation>
). Metathoracic leg with tibia longer than femur. Tibia with 2 lateral spines on the lower half, row of large apical teeth present, with median diastemma. First tarsal segment bearing apical spines organised in a triangular area, second tarsal segment without any spine reduced into a small rounded lobe (
<figureCitation id="13731397F03DFFFEFAA4B1E9FA6CFE68" box="[1358,1431,404,429]" captionStart="FIGURES 6 – 11" captionStartId="3.[151,264,1914,1937]" captionTargetBox="[171,1425,203,1873]" captionTargetId="figure@3.[151,1436,193,1893]" captionTargetPageId="3" captionText="FIGURES 6 – 11. Binaluana emarginata gen. et sp. nov., female. (6) Head, rostrum and legs, ventral view; (7) abdomen and hind legs, ventral view; (8) labels; (9 – 11) abdomen: (9) dorsal view, (10) ventral view, (11) posterior view." httpUri="https://zenodo.org/record/243985/files/figure.png" pageId="4" pageNumber="590">Figs 7</figureCitation>
,
<figureCitation id="13731397F03DFFFEFF7DB1C5FF1AFE14" box="[151,225,440,464]" captionStart="FIGURES 36 – 41" captionStartId="8.[151,264,1647,1670]" captionTargetBox="[173,1420,194,1613]" captionTargetId="figure@8.[173,1420,194,1621]" captionTargetPageId="8" captionText="FIGURES 36 – 41. Binaluana emarginata gen. et sp. nov., female, SEM images. (36) Fore femur and tibia, ventral view; (37 – 40) hind tibia and tarsal segments; (41) rostrum, ventral view." httpUri="https://zenodo.org/record/243990/files/figure.png" pageId="4" pageNumber="590">37–41</figureCitation>
).
</paragraph>
<caption id="DF375F9AF03DFFFEFF7DB7C7FB2FF834" httpUri="https://zenodo.org/record/243986/files/figure.png" pageId="4" pageNumber="590" targetBox="[160,1420,523,1955]" targetPageId="4">
<paragraph id="8BF70F12F03DFFFEFF7DB7C7FB2FF834" blockId="4.[151,1436,1978,2032]" pageId="4" pageNumber="590">
<emphasis id="B93CD300F03DFFFEFF7DB7C7FEB4F814" bold="true" box="[151,335,1978,2001]" pageId="4" pageNumber="590">FIGURES 12–17</emphasis>
.
<emphasis id="B93CD300F03DFFFEFEB7B7C7FD14F814" bold="true" box="[349,751,1978,2001]" pageId="4" pageNumber="590">
<taxonomicName id="4C487491F03DFFFEFEB7B7C7FDB0F814" box="[349,587,1978,2001]" class="Insecta" family="Lophopidae" genus="Binaluana" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="590" phylum="Arthropoda" rank="species" species="emarginata">
<emphasis id="B93CD300F03DFFFEFEB7B7C7FDB0F814" bold="true" box="[349,587,1978,2001]" italics="true" pageId="4" pageNumber="590">Binaluana emarginata</emphasis>
</taxonomicName>
gen. et sp. nov
</emphasis>
., female, SEM images. (12–13) Anterior part of body: (12) dorsal view, (13) dorso-lateral view; (14) vertex, dorsal view; (15) head, frontal view; (16–17) frons, frontal view.
</paragraph>
</caption>
<caption id="DF375F9AF03CFFFFFF7DB6E9FB7BF92C" httpUri="https://zenodo.org/record/243987/files/figure.png" pageId="5" pageNumber="591" targetBox="[151,1436,193,1653]" targetPageId="5">
<paragraph id="8BF70F12F03CFFFFFF7DB6E9FB7BF92C" blockId="5.[151,1436,1684,1768]" pageId="5" pageNumber="591">
<emphasis id="B93CD300F03CFFFFFF7DB6E9FEB5F96E" bold="true" box="[151,334,1684,1707]" pageId="5" pageNumber="591">FIGURES 18–23</emphasis>
.
<emphasis id="B93CD300F03CFFFFFEB0B6E9FD1EF96F" bold="true" box="[346,741,1684,1707]" pageId="5" pageNumber="591">
<taxonomicName id="4C487491F03CFFFFFEB0B6E9FDBCF96E" box="[346,583,1684,1707]" class="Insecta" family="Lophopidae" genus="Binaluana" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="591" phylum="Arthropoda" rank="species" species="emarginata">
<emphasis id="B93CD300F03CFFFFFEB0B6E9FDBCF96E" bold="true" box="[346,583,1684,1707]" italics="true" pageId="5" pageNumber="591">Binaluana emarginata</emphasis>
</taxonomicName>
gen. et sp. nov
</emphasis>
., female, SEM images. (18) Anterior part of body, lateral view; (19) head and pronotum, lateral view; (20) upper part of head, lateral view; (21) pronotum and mesonotum, lateral view; (22) lower part of head and pronotum, lateral view; (23) upper part of pronotum and mesonotum, lateral view.
</paragraph>
</caption>
<paragraph id="8BF70F12F03CFFFCFF2DB772FABDFF3D" blockId="5.[151,1437,1806,2007]" lastBlockId="6.[151,1437,151,464]" lastPageId="6" lastPageNumber="592" pageId="5" pageNumber="591">
TEGMINA (
<figureCitation id="13731397F03CFFFFFED5B773FE5FF8E3" box="[319,420,1806,1831]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,264,1906,1929]" captionTargetBox="[169,1428,491,1875]" captionTargetId="figure@2.[151,1436,481,1885]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Binaluana emarginata gen. et sp. nov., female. (1) Habitus, lateral view; (2) same, dorsal view; (3) anterior part of body, lateral view; (4) anterior part, frontal view; (5) anterior part of body, dorsal view." httpUri="https://zenodo.org/record/243984/files/figure.png" pageId="5" pageNumber="591">Figs 1–2</figureCitation>
,
<figureCitation id="13731397F03CFFFFFE58B773FE00F8E3" box="[434,507,1806,1831]" captionStart="FIGURES 30 – 35" captionStartId="7.[151,264,1689,1712]" captionTargetBox="[151,1436,193,1655]" captionTargetId="figure@7.[151,1436,193,1664]" captionTargetPageId="7" captionText="FIGURES 30 – 35. Binaluana emarginata gen. et sp. nov., female, SEM images. (30 – 35) Tegmina." httpUri="https://zenodo.org/record/243989/files/figure.png" pageId="5" pageNumber="591">30–35</figureCitation>
). Tegmen elongate, flattened, membranous, completely covered with setae, on and between veins. Costal margin almost parallel to postclaval margin. Costal area narrow at the base and getting wider toward the apex, reaching the level of claval apex, transversal veinlets present all along it. Postcostal cell with veinlets. Basal cell much narrower than long. ScP+R+MA leaving basal cell by a long common stem. Forking of the veins, leaving the basal cell, is done in the following order: ScP+R+MA, CuA and MP. ScP+RA forking almost at same level than secondary forking of
<collectingCountry id="F35F4F82F03CFFFFFDD0B7C2FDA7F812" box="[570,604,1983,2006]" name="Poland" pageId="5" pageNumber="591">RP</collectingCountry>
and both a little before the end of costal area. MP forking before ScP+RA forking. MP1+2 forking before MP3+4 and not forking any further. CuA forking a little later than fusion of Pcu+A1, all CuA terminals reaching postclaval margin. Transverse veinlets present in radial and median cells, but weakly visible. Few transverse veinlets present in apex of cubital cell. Transverse veinlets between CuP and Pcu and Pcu+A1.
</paragraph>
<paragraph id="8BF70F12F03FFFFCFF2DB175FE77FEE4" blockId="6.[151,1437,151,464]" box="[199,396,263,288]" pageId="6" pageNumber="592">MALE. Unknown.</paragraph>
<paragraph id="8BF70F12F03FFFFCFF2DB156FD84FE14" blockId="6.[151,1437,151,464]" pageId="6" pageNumber="592">
FEMALE TERMINALIA. Seventh abdominal sternite bearing a protuberance ventrally (
<figureCitation id="13731397F03FFFFCFB93B157FB15FE87" box="[1145,1262,298,323]" captionStart="FIGURES 6 – 11" captionStartId="3.[151,264,1914,1937]" captionTargetBox="[171,1425,203,1873]" captionTargetId="figure@3.[151,1436,193,1893]" captionTargetPageId="3" captionText="FIGURES 6 – 11. Binaluana emarginata gen. et sp. nov., female. (6) Head, rostrum and legs, ventral view; (7) abdomen and hind legs, ventral view; (8) labels; (9 – 11) abdomen: (9) dorsal view, (10) ventral view, (11) posterior view." httpUri="https://zenodo.org/record/243985/files/figure.png" pageId="6" pageNumber="592">Figs 7, 10</figureCitation>
,
<figureCitation id="13731397F03FFFFCFB16B157FABFFE87" box="[1276,1348,298,323]" captionStart="FIGURES 42 – 47" captionStartId="9.[151,264,1640,1663]" captionTargetBox="[151,1436,159,1620]" captionTargetId="figure@9.[151,1436,159,1625]" captionTargetPageId="9" captionText="FIGURES 42 – 47. Binaluana emarginata gen. et sp. nov., female, SEM images. (42 – 43) Abdomen: (42) dorso-lateral view, (43) lateral view; (44 – 45) terminalia: (44) dorso-lateral view, (45) lateral view; (46) anal tube, posterior view; (47) median part of anal tube, dorso-lateral view." httpUri="https://zenodo.org/record/243991/files/figure.png" pageId="6" pageNumber="592">44–45</figureCitation>
,
<figureCitation id="13731397F03FFFFCFABBB157FA96FE87" box="[1361,1389,298,323]" captionStart="FIGURES 48 – 52" captionStartId="10.[151,264,1781,1804]" captionTargetBox="[151,1436,256,1716]" captionTargetId="figure@10.[151,1436,256,1719]" captionTargetPageId="10" captionText="FIGURES 48 – 52. Binaluana emarginata gen. et sp. nov., female, SEM images. (48) Abdomen, ventral view; (50) terminalia, ventral view; (49) same, lateral view; (50 – 51) gonapophysis VIII: (50) ventral view, (51) posterior view; (52) gonoplac with wax secretion, lateral view." httpUri="https://zenodo.org/record/243992/files/figure.png" pageId="6" pageNumber="592">48</figureCitation>
–50,
<figureCitation id="13731397F03FFFFCFF7DB130FF25FEA2" box="[151,222,333,358]" captionStart="FIGURES 53 – 60" captionStartId="11.[151,264,1848,1871]" captionTargetBox="[175,1403,205,1810]" captionTargetId="figure@11.[151,1436,193,1827]" captionTargetPageId="11" captionText="FIGURES 53 – 60. Binaluana emarginata gen. et sp. nov., female. (53 – 55) Pregenital sternite: (53) ventral view, (54) ventroposterior view, (55) posterior view; (56) anal tube, dorsal view; (57) gonoplac and gonapophysis VIII, external view (lateral); (58) bursa copulatrix, lateral view; (59 – 60) spermatheca." httpUri="https://zenodo.org/record/243993/files/figure.png" pageId="6" pageNumber="592">53–55</figureCitation>
), anterior margin convex anteriorly and posterior margin with lateral corners protruding. Medio-ventral extension at the apex of the anal tube absent. Anal tube bearing two large lobes apically, oriented postero-ventrally and covering the gonoplacs (
<figureCitation id="13731397F03FFFFCFE35B1E9FDA9FE68" box="[479,594,404,429]" captionStart="FIGURES 6 – 11" captionStartId="3.[151,264,1914,1937]" captionTargetBox="[171,1425,203,1873]" captionTargetId="figure@3.[151,1436,193,1893]" captionTargetPageId="3" captionText="FIGURES 6 – 11. Binaluana emarginata gen. et sp. nov., female. (6) Head, rostrum and legs, ventral view; (7) abdomen and hind legs, ventral view; (8) labels; (9 – 11) abdomen: (9) dorsal view, (10) ventral view, (11) posterior view." httpUri="https://zenodo.org/record/243985/files/figure.png" pageId="6" pageNumber="592">Figs 9–11</figureCitation>
,
<figureCitation id="13731397F03FFFFCFD8AB1E9FD53FE68" box="[608,680,404,428]" captionStart="FIGURES 42 – 47" captionStartId="9.[151,264,1640,1663]" captionTargetBox="[151,1436,159,1620]" captionTargetId="figure@9.[151,1436,159,1625]" captionTargetPageId="9" captionText="FIGURES 42 – 47. Binaluana emarginata gen. et sp. nov., female, SEM images. (42 – 43) Abdomen: (42) dorso-lateral view, (43) lateral view; (44 – 45) terminalia: (44) dorso-lateral view, (45) lateral view; (46) anal tube, posterior view; (47) median part of anal tube, dorso-lateral view." httpUri="https://zenodo.org/record/243991/files/figure.png" pageId="6" pageNumber="592">42–47</figureCitation>
,
<figureCitation id="13731397F03FFFFCFD5CB1E9FD2EFE69" box="[694,725,404,429]" captionStart="FIGURES 53 – 60" captionStartId="11.[151,264,1848,1871]" captionTargetBox="[175,1403,205,1810]" captionTargetId="figure@11.[151,1436,193,1827]" captionTargetPageId="11" captionText="FIGURES 53 – 60. Binaluana emarginata gen. et sp. nov., female. (53 – 55) Pregenital sternite: (53) ventral view, (54) ventroposterior view, (55) posterior view; (56) anal tube, dorsal view; (57) gonoplac and gonapophysis VIII, external view (lateral); (58) bursa copulatrix, lateral view; (59 – 60) spermatheca." httpUri="https://zenodo.org/record/243993/files/figure.png" pageId="6" pageNumber="592">56</figureCitation>
). Small longitudinal carina along the ventral part of anal tube. Dense and long setae covering the anal tube.
</paragraph>
<caption id="DF375F9AF03FFFFCFF7DB7E2FD2EF81D" httpUri="https://zenodo.org/record/243988/files/figure.png" pageId="6" pageNumber="592" targetBox="[173,1407,514,1916]" targetPageId="6">
<paragraph id="8BF70F12F03FFFFCFF7DB7E2FD2EF81D" blockId="6.[151,1436,1951,2009]" pageId="6" pageNumber="592">
<emphasis id="B93CD300F03FFFFCFF7DB7E2FEB6F871" bold="true" box="[151,333,1951,1974]" pageId="6" pageNumber="592">FIGURES 24–29</emphasis>
.
<emphasis id="B93CD300F03FFFFCFEB1B7E2FD13F871" bold="true" box="[347,744,1951,1974]" pageId="6" pageNumber="592">
<taxonomicName id="4C487491F03FFFFCFEB1B7E2FDBEF871" box="[347,581,1951,1974]" class="Insecta" family="Lophopidae" genus="Binaluana" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="592" phylum="Arthropoda" rank="species" species="emarginata">
<emphasis id="B93CD300F03FFFFCFEB1B7E2FDBEF871" bold="true" box="[347,581,1951,1974]" italics="true" pageId="6" pageNumber="592">Binaluana emarginata</emphasis>
</taxonomicName>
gen. et sp. nov
</emphasis>
., female, SEM images. (24–27) Antenna: (24–25) dorsal view, (26) dorso-lateral view, (27) apical view; (28–29) plate organs.
</paragraph>
</caption>
<caption id="DF375F9AF03EFFFDFF7DB6E4FB76F974" box="[151,1165,1689,1712]" httpUri="https://zenodo.org/record/243989/files/figure.png" pageId="7" pageNumber="593" targetBox="[151,1436,193,1655]" targetPageId="7">
<paragraph id="8BF70F12F03EFFFDFF7DB6E4FB76F974" blockId="7.[151,1165,1689,1712]" box="[151,1165,1689,1712]" pageId="7" pageNumber="593">
<emphasis id="B93CD300F03EFFFDFF7DB6E4FEB6F96B" bold="true" box="[151,333,1689,1712]" pageId="7" pageNumber="593">FIGURES 30–35</emphasis>
.
<emphasis id="B93CD300F03EFFFDFEB0B6E4FD1FF96B" bold="true" box="[346,740,1689,1712]" pageId="7" pageNumber="593">
<taxonomicName id="4C487491F03EFFFDFEB0B6E4FDBDF96B" box="[346,582,1689,1712]" class="Insecta" family="Lophopidae" genus="Binaluana" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="593" phylum="Arthropoda" rank="species" species="emarginata">
<emphasis id="B93CD300F03EFFFDFEB0B6E4FDBDF96B" bold="true" box="[346,582,1689,1712]" italics="true" pageId="7" pageNumber="593">Binaluana emarginata</emphasis>
</taxonomicName>
gen. et sp. nov
</emphasis>
., female, SEM images. (30–35) Tegmina.
</paragraph>
</caption>
<paragraph id="8BF70F12F03EFFFDFF2DB6A8FF38F89C" blockId="7.[151,1437,1749,2024]" pageId="7" pageNumber="593">
Gonoplacs, unilobate elongate-rounded and bearing dense setae internally and externally (
<figureCitation id="13731397F03EFFFDFB53B6A8FAC1F92A" box="[1209,1338,1749,1774]" captionStart="FIGURES 42 – 47" captionStartId="9.[151,264,1640,1663]" captionTargetBox="[151,1436,159,1620]" captionTargetId="figure@9.[151,1436,159,1625]" captionTargetPageId="9" captionText="FIGURES 42 – 47. Binaluana emarginata gen. et sp. nov., female, SEM images. (42 – 43) Abdomen: (42) dorso-lateral view, (43) lateral view; (44 – 45) terminalia: (44) dorso-lateral view, (45) lateral view; (46) anal tube, posterior view; (47) median part of anal tube, dorso-lateral view." httpUri="https://zenodo.org/record/243991/files/figure.png" pageId="7" pageNumber="593">Figs 42–45</figureCitation>
,
<figureCitation id="13731397F03EFFFDFAACB6A8FA9FF92A" box="[1350,1380,1749,1774]" captionStart="FIGURES 48 – 52" captionStartId="10.[151,264,1781,1804]" captionTargetBox="[151,1436,256,1716]" captionTargetId="figure@10.[151,1436,256,1719]" captionTargetPageId="10" captionText="FIGURES 48 – 52. Binaluana emarginata gen. et sp. nov., female, SEM images. (48) Abdomen, ventral view; (50) terminalia, ventral view; (49) same, lateral view; (50 – 51) gonapophysis VIII: (50) ventral view, (51) posterior view; (52) gonoplac with wax secretion, lateral view." httpUri="https://zenodo.org/record/243992/files/figure.png" pageId="7" pageNumber="593">52</figureCitation>
,
<figureCitation id="13731397F03EFFFDFA9AB6A8FA6BF92A" box="[1392,1424,1749,1774]" captionStart="FIGURES 53 – 60" captionStartId="11.[151,264,1848,1871]" captionTargetBox="[175,1403,205,1810]" captionTargetId="figure@11.[151,1436,193,1827]" captionTargetPageId="11" captionText="FIGURES 53 – 60. Binaluana emarginata gen. et sp. nov., female. (53 – 55) Pregenital sternite: (53) ventral view, (54) ventroposterior view, (55) posterior view; (56) anal tube, dorsal view; (57) gonoplac and gonapophysis VIII, external view (lateral); (58) bursa copulatrix, lateral view; (59 – 60) spermatheca." httpUri="https://zenodo.org/record/243993/files/figure.png" pageId="7" pageNumber="593">57</figureCitation>
). Gonocoxae IX present, gonapophysis IX clearly visible when for the other Lophopid genera they are strongly reduced and hardly observable sometime (
<figureCitation id="13731397F03EFFFDFD96B761FD28F8F1" box="[636,723,1820,1845]" captionStart="FIGURES 53 – 60" captionStartId="11.[151,264,1848,1871]" captionTargetBox="[175,1403,205,1810]" captionTargetId="figure@11.[151,1436,193,1827]" captionTargetPageId="11" captionText="FIGURES 53 – 60. Binaluana emarginata gen. et sp. nov., female. (53 – 55) Pregenital sternite: (53) ventral view, (54) ventroposterior view, (55) posterior view; (56) anal tube, dorsal view; (57) gonoplac and gonapophysis VIII, external view (lateral); (58) bursa copulatrix, lateral view; (59 – 60) spermatheca." httpUri="https://zenodo.org/record/243993/files/figure.png" pageId="7" pageNumber="593">Fig. 57</figureCitation>
).
<emphasis id="B93CD300F03EFFFDFD02B760FC5CF8F0" box="[744,935,1820,1844]" italics="true" pageId="7" pageNumber="593">Bursa copulatrix</emphasis>
with a single pouch with visible cells (
<figureCitation id="13731397F03EFFFDFA84B761FF4CF89C" captionStart="FIGURES 53 – 60" captionStartId="11.[151,264,1848,1871]" captionTargetBox="[175,1403,205,1810]" captionTargetId="figure@11.[151,1436,193,1827]" captionTargetPageId="11" captionText="FIGURES 53 – 60. Binaluana emarginata gen. et sp. nov., female. (53 – 55) Pregenital sternite: (53) ventral view, (54) ventroposterior view, (55) posterior view; (56) anal tube, dorsal view; (57) gonoplac and gonapophysis VIII, external view (lateral); (58) bursa copulatrix, lateral view; (59 – 60) spermatheca." httpUri="https://zenodo.org/record/243993/files/figure.png" pageId="7" pageNumber="593">Fig. 58</figureCitation>
).
</paragraph>
<paragraph id="8BF70F12F03EFFFDFF2DB71FFD22F85A" blockId="7.[151,1437,1749,2024]" pageId="7" pageNumber="593">
Spermatheca well developed,
<emphasis id="B93CD300F03EFFFDFDD4B71EFCE3F8BF" box="[574,792,1891,1915]" italics="true" pageId="7" pageNumber="593">ductus receptaculi</emphasis>
distinctly longer than
<emphasis id="B93CD300F03EFFFDFBA2B71EFAC2F8BF" box="[1096,1337,1891,1915]" italics="true" pageId="7" pageNumber="593">diverticulum ductus,</emphasis>
ribbed.
<emphasis id="B93CD300F03EFFFDFF7DB7FBFE80F85A" box="[151,379,1926,1950]" italics="true" pageId="7" pageNumber="593">Diverticulum ductus</emphasis>
smooth, bulbous (
<figureCitation id="13731397F03EFFFDFDA3B7F8FD36F85A" box="[585,717,1925,1950]" captionStart="FIGURES 53 – 60" captionStartId="11.[151,264,1848,1871]" captionTargetBox="[175,1403,205,1810]" captionTargetId="figure@11.[151,1436,193,1827]" captionTargetPageId="11" captionText="FIGURES 53 – 60. Binaluana emarginata gen. et sp. nov., female. (53 – 55) Pregenital sternite: (53) ventral view, (54) ventroposterior view, (55) posterior view; (56) anal tube, dorsal view; (57) gonoplac and gonapophysis VIII, external view (lateral); (58) bursa copulatrix, lateral view; (59 – 60) spermatheca." httpUri="https://zenodo.org/record/243993/files/figure.png" pageId="7" pageNumber="593">Figs 59–60</figureCitation>
).
</paragraph>
<paragraph id="8BF70F12F03EFFFDFF7DB7CDFD6BF80C" blockId="7.[151,1437,1749,2024]" box="[151,656,1967,1992]" pageId="7" pageNumber="593">
As for all
<taxonomicName id="4C487491F03EFFFDFEE2B7D2FE6BF80C" box="[264,400,1967,1992]" class="Insecta" family="Lophopidae" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="593" phylum="Arthropoda" rank="family">Lophopidae</taxonomicName>
, gonospiculum absent.
</paragraph>
</subSubSection>
<subSubSection id="C3525C99F03EFFFDFF2DB7B2FD63F82C" box="[199,664,1999,2024]" pageId="7" pageNumber="593" type="distribution">
<paragraph id="8BF70F12F03EFFFDFF2DB7B2FD63F82C" blockId="7.[151,1437,1749,2024]" box="[199,664,1999,2024]" pageId="7" pageNumber="593">
<emphasis id="B93CD300F03EFFFDFF2DB7B2FEA2F82C" bold="true" box="[199,345,1999,2024]" pageId="7" pageNumber="593">Distribution</emphasis>
. Palawan (The
<collectingCountry id="F35F4F82F03EFFFDFDE3B7B2FD77F82C" box="[521,652,1999,2024]" name="Philippines" pageId="7" pageNumber="593">Philippines</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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