245 lines
26 KiB
XML
245 lines
26 KiB
XML
<document id="2B67DFB8AA52E7264A2F5E8AFF7B3C8C" ID-DOI="10.11646/zootaxa.3948.1.2" ID-GBIF-Dataset="9e109059-3d75-4e2c-997e-6f4217d201d0" ID-ISSN="1175-5326" ID-Zenodo-Dep="244003" ID-ZooBank="9A0330AD-EEF7-4A80-8D80-1B6116BD856F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461042933937" checkinUser="plazi" docAuthor="Filippini, Valentina, Galante, Eduardo & Micó, Estefanía" docDate="2015" docId="03E1981EB53AFFE9FF59FE2C0295EF65" docLanguage="en" docName="zt03948p040.pdf" docOrigin="Zootaxa 3948 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anomala pincelada Filippini, Galante" docType="treatment" docVersion="7" masterDocId="FFD8E066B530FFE2FFCEFFD3014CEC5D" masterDocTitle="Description of six new species of Anomalini from Costa Rica (Coleoptera: Scarabaeidae: Rutelinae)" updateTime="1698576078296" updateUser="plazi">
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<mods:mods id="B3D594CF435CDE655C8682098AF9FDC7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="D3B49F772ADF80021D6A59698339DF56">
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<mods:title id="6FA8810FC4DF9031AF79B96197638E8D">Description of six new species of Anomalini from Costa Rica (Coleoptera: Scarabaeidae: Rutelinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="6639E5830E127ED81424E5D63324ECFC" type="personal">
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<mods:role id="CD0F44A8398B1F36EE9902CC51092164">
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<mods:roleTerm id="2F10512C29DDDCC3E948654487637FE8">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3EC0954C40272F4EBE25C43CB7D1F878">Filippini, Valentina</mods:namePart>
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</mods:name>
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<mods:name id="B388BFA610D69C6F18DD5060F2CDCAD5" type="personal">
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<mods:role id="73E44CD10AFF68DF6991472431CD17BC">
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<mods:roleTerm id="C1816C9F049F384D45E5AC4C83DE870C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D7D33C1CEB333EB42561732829C2A2F1">Galante, Eduardo</mods:namePart>
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</mods:name>
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<mods:name id="50BEAFC28D5C52571D7986FEE7FFB871" type="personal">
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<mods:role id="AE3BD3718F9A21E0C0BFD06E866D05EA">
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<mods:roleTerm id="9B2B581923430E4D6C0F8BC60E1DC049">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D0BFC12D043FAC9F1BB80295D7B4001D">Micó, Estefanía</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="1117E653973EBF7C59F700F5E59BF7CA">text</mods:typeOfResource>
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<mods:relatedItem id="530FE7DABF638D3CF73A2D515694EC72" type="host">
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<mods:titleInfo id="0E5E26F0279FD3080412BFAAF6944B6E">
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<mods:title id="926B4CAD49293A03F3BFF3A314F7DE57">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="758EED526DEC1FB8E5B7F6C1D0719008">
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<mods:date id="759EDF6B4ABEB72AD3A902C1FA4D6D17">2015</mods:date>
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<mods:detail id="95FAF0AB92BC044CC8C8C9E78BD68AA2" type="volume">
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<mods:number id="E2723CA8122AF7C7565F599B6A07467C">3948</mods:number>
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</mods:detail>
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<mods:detail id="01C40BD974A9FD4A31091DD2DCDCE417" type="issue">
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<mods:number id="AD8ECB76D7A6A2C8410CBE01B0DAE1AA">1</mods:number>
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</mods:detail>
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</mods:part>
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<mods:classification id="38DD611FAF549B62779C229F72AB8179">journal volume</mods:classification>
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<mods:identifier id="89D0D76C518F44ED6A21107795E4EB45" type="DOI">10.11646/zootaxa.3948.1.2</mods:identifier>
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<mods:identifier id="4DEF22C4AF8103430BB86D50A15AFA37" type="GBIF-Dataset">9e109059-3d75-4e2c-997e-6f4217d201d0</mods:identifier>
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<mods:identifier id="BC1A881D63E838CCDC846FA3AC9243DC" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="EA4B0195414560018D97B170932E2505" type="Zenodo-Dep">244003</mods:identifier>
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<mods:identifier id="988BD6EAF872CDCC67755C64446FA993" type="ZooBank">9A0330AD-EEF7-4A80-8D80-1B6116BD856F</mods:identifier>
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</mods:mods>
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<treatment id="03E1981EB53AFFE9FF59FE2C0295EF65" ID-DOI="http://doi.org/10.5281/zenodo.6122706" ID-GBIF-Taxon="119609990" ID-Zenodo-Dep="6122706" LSID="urn:lsid:plazi:treatment:03E1981EB53AFFE9FF59FE2C0295EF65" httpUri="http://treatment.plazi.org/id/03E1981EB53AFFE9FF59FE2C0295EF65" lastPageId="11" pageId="10">
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<subSubSection id="C3527A83B53AFFE8FF59FE2C0195EE66" pageId="10" type="nomenclature">
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<paragraph id="8BF72908B53AFFE8FF59FE2C023FEE47" blockId="10.[151,883,511,571]" box="[151,883,511,538]" pageId="10">
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<heading id="D0BF9E64B53AFFE8FF59FE2C023FEE47" bold="true" box="[151,883,511,538]" fontSize="11" level="1" pageId="10" reason="1">
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<emphasis id="B93CF51AB53AFFE8FF59FE2C023FEE47" bold="true" box="[151,883,511,538]" pageId="10">
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<taxonomicName id="4C48528BB53AFFE8FF59FE2C0325EE47" authority="Filippini, Galante" authorityName="Filippini, Galante" box="[151,617,511,538]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="pincelada">
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<emphasis id="B93CF51AB53AFFE8FF59FE2C00CEEE44" bold="true" box="[151,386,511,537]" italics="true" pageId="10">Anomala pincelada</emphasis>
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<collectingCountry id="F35F6998B53AFFE8FE47FDD300BAEE47" box="[393,502,512,538]" name="Philippines" pageId="10">Filippini</collectingCountry>
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, Galante
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</taxonomicName>
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, & Micó, new species
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF72908B53AFFE8FF59FDF10195EE66" blockId="10.[151,883,511,571]" box="[151,217,546,571]" pageId="10">
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<figureCitation id="1373358DB53AFFE8FF59FDF10195EE66" box="[151,217,546,571]" captionStart="FIGURES 1 – 6" captionStartId="4.[151,264,1977,2000]" captionTargetBox="[186,1400,705,1952]" captionTargetId="figure@4.[179,1408,697,1957]" captionTargetPageId="4" captionText="FIGURES 1 – 6. Habitus. 1: Anomala aglaos (holotype). 2: A. estrella (holotype). 3: A. inbio (holotype). 4: A. pincelada (holotype). 5: Callistethus ruteloides (holotype). 6: C. yalizo (holotype). Scale = 5 mm." httpUri="https://zenodo.org/record/244004/files/figure.png" pageId="10">Fig. 4</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3527A83B53AFFE9FF59FDB90239ECA5" lastPageId="11" pageId="10" type="materials_examined">
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<paragraph id="8BF72908B53AFFE8FF59FDB9058EEEFA" blockId="10.[151,1437,618,2011]" pageId="10">
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<emphasis id="B93CF51AB53AFFE8FF59FDB900CDEEDE" bold="true" box="[151,385,618,643]" pageId="10">Material examined.</emphasis>
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<typeStatus id="54F397AAB53AFFE8FE47FDB900BAEEDE" box="[393,502,618,643]" pageId="10" type="holotype">Holotype</typeStatus>
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: ♂"Fca. Jenny,
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<quantity id="4CB084EDB53AFFE8FD69FDB803A3EEDE" box="[679,751,618,643]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="10" unit="km" value="30.0">30 km</quantity>
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N de
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<collectingCountry id="F35F6998B53AFFE8FCF8FDB902C6EEDE" box="[822,906,618,643]" name="Liberia" pageId="10">Liberia</collectingCountry>
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, P.N. Guanacaste, Prov. Guan.,
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<collectingCountry id="F35F6998B53AFFE8FB3FFDB804D5EEDE" box="[1265,1433,619,643]" name="Costa Rica" pageId="10">COSTA RICA</collectingCountry>
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.
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<date id="FFF60FC8B53AFFE8FF59FD5D0001EEFA" box="[151,333,654,679]" pageId="10" value="1993-04-18" valueMax="1993-04-25" valueMin="1993-04-18">18-25 Abr 1993</date>
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, E. Araya, L- N 316200_364400 / INBIOCRI001167801" deposited in
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<taxonomicName id="4C48528BB53AFFE8FBA3FD5C05F2EEFA" box="[1133,1214,655,679]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="inbio">
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<collectionCode id="ED59B1CDB53AFFE8FBA3FD5C05F2EEFA" box="[1133,1214,655,679]" pageId="10">INBIO</collectionCode>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BF72908B53AFFE8FF09FD6002C5E86E" blockId="10.[151,1437,618,2011]" pageId="10">
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<typeStatus id="54F397AAB53AFFE8FF09FD600078EE96" box="[199,308,691,715]" pageId="10" type="paratype">Paratypes</typeStatus>
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(9):
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<specimenCount id="9D4EE281B53AFFE8FEBFFD6000D8EE96" box="[369,404,691,715]" pageId="10" type="generic">1♂</specimenCount>
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||
"3 Km este de Cuajiniquil,
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||
<quantity id="4CB084EDB53AFFE8FD2CFD600269EE96" box="[738,805,691,715]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="10" unit="m" value="300.0">300m</quantity>
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, Prov. Guanacaste,
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<collectingCountry id="F35F6998B53AFFE8FBC5FD6005C7EE96" box="[1035,1163,690,715]" name="Costa Rica" pageId="10">Costa Rica</collectingCountry>
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,
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<date id="FFF60FC8B53AFFE8FB54FD610466EE96" box="[1178,1322,690,715]" pageId="10" value="1992-06-25">25 jun 1992</date>
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, III curso Parataxon. L-N 325600_355200 / INBIOCRI000863924";
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<specimenCount id="9D4EE281B53AFFE8FC9FFD040238EEB2" box="[849,884,727,751]" pageId="10" type="generic">1♂</specimenCount>
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"Santa Rosa National Park Guanacaste Prov.
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<collectingCountry id="F35F6998B53AFFE8FF59FD280072EF4E" box="[151,318,763,787]" name="Costa Rica" pageId="10">COSTA RICA</collectingCountry>
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.
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<date id="FFF60FC8B53AFFE8FE89FD2800BEEF4E" box="[327,498,763,787]" pageId="10" value="1980-05-09" valueMax="1980-05-11" valueMin="1980-05-09">9-11 May 1980</date>
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DH Janzen & W Hallwachs / INBIOCRI001116980";
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<specimenCount id="9D4EE281B53AFFE8FB9DFD280534EF4E" box="[1107,1144,763,787]" pageId="10" type="generic">1♂</specimenCount>
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, 1♀ "Fca. Jenny,
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||
<quantity id="4CB084EDB53AFFE8FAF2FD2804CEEF4E" box="[1340,1410,762,787]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="10" unit="km" value="30.0">30 km</quantity>
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N de
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<collectingCountry id="F35F6998B53AFFE8FF71FCCD005FEF6A" box="[191,275,798,823]" name="Liberia" pageId="10">Liberia</collectingCountry>
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||
, P.N. Guanacaste, Prov. Guan.,
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||
<collectingCountry id="F35F6998B53AFFE8FD5BFCCC020FEF6A" box="[661,835,799,823]" name="Costa Rica" pageId="10">COSTA RICA</collectingCountry>
|
||
.
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<date id="FFF60FC8B53AFFE8FC9DFCCC0543EF6A" box="[851,1039,799,823]" pageId="10" value="1993-05-09" valueMax="1993-05-14" valueMin="1993-05-09">9-14 May 1993</date>
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. E. Araya, L-N 316200_364400" INBIOCRI001183402 and INBIOCRI001183321; 1♀ "Tierras Morenas, Rio San Lorenzo, Tenorio, Prov. Guana,
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||
<collectingCountry id="F35F6998B53AFFE8FF59FCB40071EF22" box="[151,317,871,895]" name="Costa Rica" pageId="10">COSTA RICA</collectingCountry>
|
||
,
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||
<quantity id="4CB084EDB53AFFE8FE89FCB500D5EF22" box="[327,409,870,895]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="10" unit="m" value="1050.0">1050m</quantity>
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||
.
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<date id="FFF60FC8B53AFFE8FE6DFCB4035BEF22" box="[419,535,871,895]" pageId="10" value="1993-05">May 1993</date>
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||
. G. Rodriguez, L S 283950_424500 # 2118 / INBIOCRI001180936";
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||
<specimenCount id="9D4EE281B53AFFE8FAEEFCB4040FEF22" box="[1312,1347,871,895]" pageId="10" type="generic">1♂</specimenCount>
|
||
"Sector Las Pailas, P. N. Guanacaste, A. C. Guanacaste, Prov. Guana,
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<collectingCountry id="F35F6998B53AFFE8FC86FC5802A3EFFE" box="[840,1007,907,931]" name="Costa Rica" pageId="10">COSTA RICA</collectingCountry>
|
||
.
|
||
<specimenCount id="9D4EE281B53AFFE8FC37FC58050AEFFF" box="[1017,1094,907,931]" pageId="10" type="generic">
|
||
<quantity id="4CB084EDB53AFFE8FC37FC58050EEFFF" box="[1017,1090,907,931]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="10" unit="m" value="800.0">800 m</quantity>
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.
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</specimenCount>
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<date id="FFF60FC8B53AFFE8FB83FC5905A2EFFE" box="[1101,1262,906,931]" pageId="10" value="1994-06-06" valueMax="1994-06-26" valueMin="1994-06-06">6-26 Jun 1994</date>
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, K. Taylor, L N 309500_389500 # 3063 / INBIOCRI001909257"; 2♀ "Finca Jenny,
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<quantity id="4CB084EDB53AFFE8FC6BFC7C02A2EF9A" box="[933,1006,942,967]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="10" unit="km" value="30.0">30 km</quantity>
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N de
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<collectingCountry id="F35F6998B53AFFE8FBF6FC7D05C0EF9A" box="[1080,1164,942,967]" name="Liberia" pageId="10">Liberia</collectingCountry>
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, P.N. Guanacaste, Prov. Guan.,
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<collectingCountry id="F35F6998B53AFFE8FF38FC0000E7EFB6" box="[246,427,979,1003]" name="Costa Rica" pageId="10">COSTA RICA</collectingCountry>
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||
.
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||
<date id="FFF60FC8B53AFFE8FE0CFC0103DFEFB6" box="[450,659,978,1003]" pageId="10" value="1993-04-18" valueMax="1993-04-25" valueMin="1993-04-18">18-25 Abr 1993</date>
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||
, E. Araya, L- N 316200_364400" INBIOCRI001167798 and
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||
<collectionCode id="ED59B1CDB53AFFE8FF59FC2501A9E852" box="[151,229,1014,1039]" pageId="10">CEUA</collectionCode>
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||
00106161;
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||
<specimenCount id="9D4EE281B53AFFE8FEAFFC2400C8E852" box="[353,388,1015,1039]" pageId="10" type="generic">1♂</specimenCount>
|
||
"Est. Murcielago,
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||
<quantity id="4CB084EDB53AFFE8FD98FC2403C2E852" box="[598,654,1014,1039]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="10" unit="km" value="8.0">8 km</quantity>
|
||
S.
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||
<collectingCountry id="F35F6998B53AFFE8FD64FC24038FE852" box="[682,707,1015,1039]" name="Iceland" pageId="10">O</collectingCountry>
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||
. de Cuajiniquil, Prov. Guana,
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||
<collectingCountry id="F35F6998B53AFFE8FBDEFC2405FBE852" box="[1040,1207,1015,1039]" name="Costa Rica" pageId="10">COSTA RICA</collectingCountry>
|
||
.
|
||
<specimenCount id="9D4EE281B53AFFE8FB0FFC240444E852" box="[1217,1288,1015,1039]" pageId="10" type="generic">
|
||
<quantity id="4CB084EDB53AFFE8FB0FFC240448E852" box="[1217,1284,1015,1039]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="10" unit="m" value="100.0">100m</quantity>
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||
.
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</specimenCount>
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<date id="FFF60FC8B53AFFE8FAC3FC25019EE86E" pageId="10" value="1993-06-16" valueMax="1993-07-04" valueMin="1993-06-16">16 Jun- 4 Jul 1993</date>
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||
. F. Quesada, L N 320300_347200 # 2177 /
|
||
<collectionCode id="ED59B1CDB53AFFE8FD74FBC90245E86E" box="[698,777,1050,1075]" pageId="10">CEUA</collectionCode>
|
||
00106160".
|
||
</paragraph>
|
||
<paragraph id="8BF72908B53AFFE8FF09FBED053BE80B" blockId="10.[151,1437,618,2011]" box="[199,1143,1086,1111]" pageId="10">
|
||
<emphasis id="B93CF51AB53AFFE8FF09FBED0344E80A" bold="true" box="[199,520,1086,1111]" pageId="10">
|
||
<typeStatus id="54F397AAB53AFFE8FF09FBED0078E80A" box="[199,308,1086,1111]" pageId="10" type="holotype">Holotype</typeStatus>
|
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male description.
|
||
</emphasis>
|
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Body shape elongate. Length
|
||
<quantity id="4CB084EDB53AFFE8FC93FBEC0288E80B" box="[861,964,1087,1111]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="10" unit="mm" value="13.2">13.2 mm</quantity>
|
||
. Width
|
||
<quantity id="4CB084EDB53AFFE8FBD4FBEC053FE80B" box="[1050,1139,1087,1111]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="10" unit="mm" value="7.9">7.9 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF72908B53AFFE8FF09FBB10190E956" blockId="10.[151,1437,618,2011]" pageId="10">Color. Head reddish brown. Pronotum ochre with an irregular median brown macula and small maculae on lateral foveae. Scutellum dark brown. Elytra ochre with a dark brown macula at each side of scutellum and dark stripes on apical third of first interstice, from basal to apical calli and near lateral margins. Pygidium ochre. Legs ochre with base and apex of tibiae dark brown; protarsi and mesotarsi reddish brown, metatarsi dark brown. Venter ochre.</paragraph>
|
||
<paragraph id="8BF72908B53AFFE8FF09FAC50024E9C6" blockId="10.[151,1437,618,2011]" pageId="10">Clypeus trapezoidal, with anterior angles widely curved; surface densely punctate-reticulate, flat. Anterior margin straight, clypeus anteriorly thin. Clypeus ratio width/length 2.1. Frons densely punctate, flat. Ocular canthum long, thin, with acute apex. Interocular ratio (interocular width/width of eye): 2.6. Antenna: ratio funiculus/club 0.8.</paragraph>
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<paragraph id="8BF72908B53AFFE8FF09FA74001EEA5B" blockId="10.[151,1437,618,2011]" pageId="10">Pronotum subtrapezoidal, ratio width/length 1.7, lateral margins regularly convex. Anterior angles quadrate and blunt. Posterior angles obtuse and blunt. Basal margin sinuate, subapical bead complete. Surface with fine, dense punctures.</paragraph>
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<paragraph id="8BF72908B53AFFE8FF09F9C101B1EA13" blockId="10.[151,1437,618,2011]" pageId="10">Scutellum. Shape subtriangular, with rounded sides and blunt apex. Ratio width/length 1.4. Surface densely punctate.</paragraph>
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<paragraph id="8BF72908B53AFFE8FF09F98900BEEACB" blockId="10.[151,1437,618,2011]" pageId="10">Elytra with costae defined by regular rows of punctures, subsutural interstices with 2–3 rows of punctures; maximum width towards apex.</paragraph>
|
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<paragraph id="8BF72908B53AFFE8FF09F971059CEAE6" blockId="10.[151,1437,618,2011]" box="[199,1232,1698,1723]" pageId="10">Pygidium with coalescing punctures and long setae at apical margin. Ratio width/length 1.6.</paragraph>
|
||
<paragraph id="8BF72908B53AFFE8FF09F9140286EB7A" blockId="10.[151,1437,618,2011]" pageId="10">Space between the mesocoxae narrow, flat. Mesometasternal suture well defined, at base of mesocoxae. Abdominal sternites with 1 row of setae and 2–3 rows of sparse, fine punctures per sternite. Terminal sternite punctate with subapical bead narrow and sinuate, apical membrane wide.</paragraph>
|
||
<paragraph id="8BF72908B53AFFE8FF09F8E10007EB86" blockId="10.[151,1437,618,2011]" pageId="10">Protibia (Fig. 10) tridentate; apical tooth long and curved; middle tooth below internal apex of protibia, triangular in shape, acute. Metatibia stout, fusiform, ratio length/width 3.0. First external carina consists of a row of sparse setae, surface punctate above second external carina and rugose below. Protarsal claws: internal claw bifurcate, with upper branch of same length and half the width of the lower branch. Inferior margin with a sharp angle near base.</paragraph>
|
||
<paragraph id="8BF72908B53BFFE9FF09FF440239ECA5" blockId="11.[151,1436,151,824]" pageId="11">
|
||
Aedeagus (
|
||
<figureCitation id="1373358DB53BFFE9FE85FF4400EAECED" box="[331,422,151,176]" captionStart="FIGURES 13 – 18" captionStartId="6.[151,264,1885,1908]" captionTargetBox="[153,1429,1020,1837]" captionTargetId="figure@6.[151,1436,1010,1864]" captionTargetPageId="6" captionText="FIGURES 13 – 18. Shape of aedeagus, lateral view (top), dorsal view (bottom left), ventral view (bottom right). 13: Anomala aglaos (paratype, Isla Bonita, Costa Rica). 14: A. estrella (holotype). 15: A. inbio (holotype). 16: A. pincelada (holotype). 17: Callistethus ruteloides (holotype). 18: C. yalizo (Cerro Chompipe, Heredia). Scale = 1 mm." httpUri="https://zenodo.org/record/244005/files/figure.png" pageId="11">Fig. 16</figureCitation>
|
||
): parameres deeply and widely sinuate anteriorly in a lateral view, becoming nearly bifurcate. Endophallus (
|
||
<figureCitation id="1373358DB53BFFE9FE6DFF6F00B8EC89" box="[419,500,188,213]" captionStart="FIGURES 19 – 22" captionStartId="7.[151,264,1709,1732]" captionTargetBox="[155,1421,200,1673]" captionTargetId="figure@7.[151,1436,193,1688]" captionTargetPageId="7" captionText="FIGURES 19 – 22. Endophallus of 19: Anomala aglaos, (a) particular in frontal view, (b) lateral view (paratype, Isla Bonita, Costa Rica). 20: A. estrella (paratype, Hacienda Tiquires, Costa Rica). 21: A. inbio (holotype). 22: A. pincelada (paratype, Cuajiniquil, Costa Rica). Scale = 1 mm." httpUri="https://zenodo.org/record/244006/files/figure.png" pageId="11">Fig. 22</figureCitation>
|
||
) with 1 long lateral sacculus tapering at apex; ejaculatory duct opening on the other side, at base; 1 small diverticle with long spines ventrally at base.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3527A83B53BFFE9FF09FED7039DEDA8" pageId="11" type="description">
|
||
<paragraph id="8BF72908B53BFFE9FF09FED70582EDD5" blockId="11.[151,1436,151,824]" pageId="11">
|
||
<emphasis id="B93CF51AB53BFFE9FF09FED70077ED40" bold="true" box="[199,315,260,285]" pageId="11">Variation.</emphasis>
|
||
Macula on pronotal disc of different size, from a subpentagonal macula not reaching posterior margin to covering most of the surface with ochre sides; a median narrow, faint, lighter band may be present; maculae on elytra of variable length; metatibia from ochre to red with darker apex. Female similar to male, protibia (Fig. 10) with longer and wider apical tooth; internal protarsal claw with curved inferior margin.
|
||
</paragraph>
|
||
<paragraph id="8BF72908B53BFFE9FF09FE47039DEDA8" blockId="11.[151,1436,151,824]" pageId="11">
|
||
Body length
|
||
<quantity id="4CB084EDB53BFFE9FEACFE470340EDF1" box="[354,524,404,429]" metricMagnitude="-2" metricUnit="m" metricValue="1.465" metricValueMax="1.61" metricValueMin="1.3199999999999998" pageId="11" unit="mm" value="14.65" valueMax="16.1" valueMin="13.2">13.2–16.1 mm</quantity>
|
||
, body width
|
||
<quantity id="4CB084EDB53BFFE9FD63FE470270EDF1" box="[685,828,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="8.55" metricValueMax="9.2" metricValueMin="7.9" pageId="11" unit="mm" value="8.55" valueMax="9.2" valueMin="7.9">7.9–9.2 mm</quantity>
|
||
. Clypeus w/l: 2.0–2.3. Interocular ratio (interocular width/width of eye): 2.5–2.9. Antenna: ratio funiculus/club 0.7–0.8. Pronotum w/l: 1.7–1.8. Scutellum w/l: 1.4– 1.6. Pygidium w/l: 1.5–1.6. Metatibia l/w: 2.5–3.0.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3527A83B53BFFE9FF09FE2C0365EE90" pageId="11" type="diagnosis">
|
||
<paragraph id="8BF72908B53BFFE9FF09FE2C0365EE90" blockId="11.[151,1436,151,824]" pageId="11">
|
||
<emphasis id="B93CF51AB53BFFE9FF09FE2C000DEE45" bold="true" box="[199,321,511,536]" pageId="11">Diagnosis.</emphasis>
|
||
Externally nearly identical to
|
||
<taxonomicName id="4C48528BB53BFFE9FD50FDD3054EEE45" authority="Burmeister, 1844" authorityName="Burmeister" authorityYear="1844" box="[670,1026,511,536]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="inconstans">
|
||
<emphasis id="B93CF51AB53BFFE9FD50FDD3027BEE4A" box="[670,823,512,535]" italics="true" pageId="11">A. inconstans</emphasis>
|
||
Burmeister, 1844
|
||
</taxonomicName>
|
||
(
|
||
<collectingCountry id="F35F6998B53BFFE9FBDDFE2C05CAEE45" box="[1043,1158,511,536]" name="Colombia" pageId="11">Colombia</collectingCountry>
|
||
, MLUH), it is separated from this species by the shape of male genitalia, which has bifurcated apex of parameres, whereas
|
||
<taxonomicName id="4C48528BB53BFFE9FACDFDF704D0EE61" box="[1283,1436,548,572]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="inconstans">
|
||
<emphasis id="B93CF51AB53BFFE9FACDFDF704D0EE61" box="[1283,1436,548,572]" italics="true" pageId="11">A. inconstans</emphasis>
|
||
</taxonomicName>
|
||
has parameres with thin and curved apex, blunt and wide ventral angle, and a frontal lobe-like expansion on anterior part of ventral margin (
|
||
<figureCitation id="1373358DB53BFFE9FDC1FDBF032BEED9" box="[527,615,620,645]" captionStart="FIGURES 27 – 33. 27" captionStartId="14.[151,264,1539,1562]" captionTargetBox="[156,1433,641,1510]" captionTargetId="figure@14.[151,1436,630,1517]" captionTargetPageId="14" captionText="FIGURES 27 – 33. 27: Aedeagus of Anomala obovata, lateral view (Quebrada Kuisa, Costa Rica, INBIO). 28: Endophallus of A. obovata (Cerro Chompipe, Costa Rica, INBIO). 29: Aedeagus of A. jansoni, lateral view (Monte Rotondo, Costa Rica, syntype, MNHUB). 30: Aedeagus of A. antica, lateral view (Estación Palo Verde, Costa Rica, INBIO). 31: Aedeagus of A. luciae, lateral view (Santa Lucia, Antilles, lectotype, MNHN). 32: Aedeagus of A. quiche, lateral view (Estación Maritza, Costa Rica, INBIO). 33: Aedeagus of A. inconstans, (a) lateral view, (b) other side of parameres (Bogotá, Colombia, syntype, MLUH). Scale = 1 mm." httpUri="https://zenodo.org/record/244009/files/figure.png" pageId="11">Fig. 33</figureCitation>
|
||
). Specimens from
|
||
<collectingCountry id="F35F6998B53BFFE9FC88FDBF02D2EED8" box="[838,926,620,645]" name="Mexico" pageId="11">Mexico</collectingCountry>
|
||
,
|
||
<collectingCountry id="F35F6998B53BFFE9FC60FDBE055BEED8" box="[942,1047,621,645]" name="Paraguay" pageId="11">Paraguay</collectingCountry>
|
||
,
|
||
<collectingCountry id="F35F6998B53BFFE9FBE9FDBF05EFEED8" box="[1063,1187,620,645]" name="Guatemala" pageId="11">Guatemala</collectingCountry>
|
||
, and
|
||
<collectingCountry id="F35F6998B53BFFE9FB29FDBF0416EED8" box="[1255,1370,620,645]" name="Venezuela" pageId="11">Venezuela</collectingCountry>
|
||
from MNHUB collection, also similar to
|
||
<emphasis id="B93CF51AB53BFFE9FDEAFD43038EEEFA" box="[548,706,656,679]" italics="true" pageId="11">
|
||
<taxonomicName id="4C48528BB53BFFE9FDEAFD4303F2EEFA" box="[548,702,656,679]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="inconstans">A. inconstans</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
all show different aedeagi, so these species are part of a large and widely distributed species complex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3527A83B53BFFE9FF09FD0400B8EF48" pageId="11" type="etymology">
|
||
<paragraph id="8BF72908B53BFFE9FF09FD0400B8EF48" blockId="11.[151,1436,151,824]" pageId="11">
|
||
<emphasis id="B93CF51AB53BFFE9FF09FD040002EEAD" bold="true" box="[199,334,727,752]" pageId="11">Etymology.</emphasis>
|
||
Latinized from Spanish noun
|
||
<taxonomicName id="4C48528BB53BFFE9FD6CFD0B025CEEAD" box="[674,784,728,752]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="pincelada">
|
||
<emphasis id="B93CF51AB53BFFE9FD6CFD0B025CEEAD" box="[674,784,728,752]" italics="true" pageId="11">pincelada</emphasis>
|
||
</taxonomicName>
|
||
, brush stroke, for the elytral pattern. This name should be treated as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3527A83B53BFFE9FF09FCCC0295EF65" box="[199,985,799,824]" pageId="11" type="distribution">
|
||
<paragraph id="8BF72908B53BFFE9FF09FCCC0295EF65" blockId="11.[151,1436,151,824]" box="[199,985,799,824]" pageId="11">
|
||
<emphasis id="B93CF51AB53BFFE9FF09FCCC002CEF65" bold="true" box="[199,352,799,824]" pageId="11">Distribution.</emphasis>
|
||
Guanacaste mountain range, from
|
||
<quantity id="4CB084EDB53BFFE9FD27FCCC0224EF6A" box="[745,872,799,824]" metricMagnitude="2" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="11" unit="m" value="400.0" valueMax="600.0" valueMin="200.0">200–600 m</quantity>
|
||
(
|
||
<figureCitation id="1373358DB53BFFE9FCB6FCCC0280EF65" box="[888,972,799,824]" captionStart="FIGURE 34 – 37" captionStartId="15.[151,250,1947,1969]" captionTargetBox="[158,1427,933,1891]" captionTargetId="figure@15.[151,1436,925,1898]" captionTargetPageId="15" captionText="FIGURE 34 – 37. Distribution of 34: Anomala aglaos and A. estrella. 35: A. inbio and A. pincelada. 36: Callistethus ruteloides. 37: C. yalizo." httpUri="https://zenodo.org/record/244010/files/figure.png" pageId="11">Fig. 35</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |