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<document id="BFFB06014A7E81C98C63674CFA3CB685" ID-CLB-Dataset="7413" ID-DOI="10.11646/zootaxa.4951.1.2" ID-GBIF-Dataset="d7a40f10-4145-499c-97be-3f9bc5088d2f" ID-ISSN="1175-5326" ID-Zenodo-Dep="4655625" ID-ZooBank="557825A0-714A-426A-917F-1C9AB7372C30" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617265349256" checkinUser="plazi" docAuthor="Archibald, S. Bruce &amp; Makarkin, Vladimir N." docDate="2021" docId="03DE878EFFE6C45FFF67FB21CE4982CD" docLanguage="en" docName="zootaxa.4951.1.2.pdf" docOrigin="Zootaxa 4951 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Archiinocellia oligoneura Handlirsch 1910" docType="treatment" docVersion="7" lastPageNumber="58" masterDocId="FFE7FFF6FFF6C44EFFF0FFA7CC6F8000" masterDocTitle="Early Eocene snakeflies (Raphidioptera) of western North America from the Okanagan Highlands and Green River Formation" masterLastPageNumber="79" masterPageNumber="41" pageNumber="57" updateTime="1698935824968" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B1BB61EE887152320DFBEEC4CB3A86C6">Early Eocene snakeflies (Raphidioptera) of western North America from the Okanagan Highlands and Green River Formation</mods:title>
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<mods:name id="5B6A0F7E62E3E809509A822934CAE400" type="personal">
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<mods:namePart id="05017C8C769020B96608700D6628B1FF">Archibald, S. Bruce</mods:namePart>
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<mods:affiliation id="E9481FB2BD639E9BD840B148B34B9BBE">Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V 5 A 1 S 6, Canada. https: // orcid. org / 0000 - 0002 - 4397 - 2497 &amp; Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America &amp; Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada</mods:affiliation>
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<mods:namePart id="8EDBCF4BD98FF316077DFEF09AFD3BF5">Makarkin, Vladimir N.</mods:namePart>
<mods:affiliation id="4896CD67FFF597C7901B37A0FC3A1126">Federal Scientific Centre of the East Asia Terrestrial Biodiversity, Far Eastern Branch of the Russian Academy of Sciences, Vladivo-</mods:affiliation>
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<mods:title id="F538312FB14969F293CA23B10B83CAA3">Zootaxa</mods:title>
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<mods:date id="28D91074FA2194BB13A25A1F3811C9CB">2021</mods:date>
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<treatment id="03DE878EFFE6C45FFF67FB21CE4982CD" ID-DOI="http://doi.org/10.5281/zenodo.4681529" ID-GBIF-Taxon="180999353" ID-Zenodo-Dep="4681529" LSID="urn:lsid:plazi:treatment:03DE878EFFE6C45FFF67FB21CE4982CD" httpUri="http://treatment.plazi.org/id/03DE878EFFE6C45FFF67FB21CE4982CD" lastPageId="17" lastPageNumber="58" pageId="16" pageNumber="57">
<subSubSection id="C36D6513FFE6C45EFF67FB21CEC384A0" box="[151,684,1158,1185]" pageId="16" pageNumber="57" type="nomenclature">
<paragraph id="8BC83698FFE6C45EFF67FB21CEC384A0" blockId="16.[151,684,1158,1221]" box="[151,684,1158,1185]" pageId="16" pageNumber="57">
<heading id="D08081F4FFE6C45EFF67FB21CEC384A0" bold="true" box="[151,684,1158,1185]" fontSize="11" level="1" pageId="16" pageNumber="57" reason="1">
<taxonomicName id="4C774D1BFFE6C45EFF67FB21CEC384A0" ID-CoL="G93B" authority="Handlirsch, 1910" authorityName="Handlirsch" authorityYear="1910" box="[151,684,1158,1185]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="oligoneura">
<emphasis id="B903EA8AFFE6C45EFF67FB21CEC384A0" bold="true" box="[151,684,1158,1185]" pageId="16" pageNumber="57">
<emphasis id="B903EA8AFFE6C45EFF67FB21CDA084A1" bold="true" box="[151,463,1158,1185]" italics="true" pageId="16" pageNumber="57">Archiinocellia oligoneura</emphasis>
<bibRefCitation id="EFE64B69FFE6C45EFE27FB21CEC384A0" author="Handlirsch, A." box="[471,684,1158,1184]" pageId="16" pageNumber="57" pagination="93 - 129" refId="ref23134" refString="Handlirsch, A. (1910) Contributions to Canadian Palaeontology. Vol. II. Part III. Canadian fossil insects. V. Insects from the Tertiary lake deposits of the Southern Interior of British Columbia collected by Mr. Lawrence Lambe, in 1906. Memoirs of the Geological Survey of Canada, 12, 93 - 129. https: // doi. org / 10.4095 / 100486" type="journal article" year="1910">Handlirsch, 1910</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C36D6513FFE6C45EFF67FB0CCC8984C5" box="[151,230,1195,1221]" pageId="16" pageNumber="57" type="description">
<paragraph id="8BC83698FFE6C45EFF67FB0CCC8984C5" blockId="16.[151,684,1158,1221]" box="[151,230,1195,1221]" pageId="16" pageNumber="57">
<figureCitation id="134C2A1DFFE6C45EFF67FB0CCC8984C5" box="[151,230,1195,1221]" captionStart="FIGURE 11" captionStartId="17.[151,250,1553,1577]" captionTargetBox="[159,1431,741,1526]" captionTargetId="figure-267@17.[151,1436,741,1529]" captionTargetPageId="17" captionText="FIGURE 11. Archiinocellia oligoneura Handlirsch, 1910. A, holotype GSC 7250; B, forewing venation; C, hind wing venation. Venation drawn in B and C overlap on the right half of the photograph, venation on the left side could be of either wing. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.4655667" httpUri="https://zenodo.org/record/4655667/files/figure.png" pageId="16" pageNumber="57">Fig. 11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36D6513FFE6C45EFF67FB54CD7B857A" pageId="16" pageNumber="57" type="materials_examined">
<paragraph id="8BC83698FFE6C45EFF67FB54C8908531" blockId="16.[151,1437,1266,2013]" pageId="16" pageNumber="57">
<emphasis id="B903EA8AFFE6C45EFF67FB54CD28850C" bold="true" box="[151,327,1266,1293]" pageId="16" pageNumber="57">Type material.</emphasis>
<materialsCitation id="3B1F3CC5FFE6C45EFEBFFB54C8908531" ID-GBIF-Occurrence="3071122306" collectingDate="1906-07-21" collectionCode="GSC" country="Canada" location="Canada" pageId="16" pageNumber="57" specimenCode="GSC 7250" specimenCount="1" typeStatus="holotype">
<typeStatus id="54CC883AFFE6C45EFEBFFB54CDD8850D" box="[335,439,1267,1293]" pageId="16" pageNumber="57" type="holotype">Holotype</typeStatus>
<specimenCode id="DBD19EE3FFE6C45EFE30FB54CE55850D" box="[448,570,1266,1293]" collectionCode="GSC" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="16" pageNumber="57">GSC 7250</specimenCode>
, collected by L[awrence]. M. L[ambe],
<date id="FFC91058FFE6C45EFBF3FB54C8EA850D" box="[1027,1157,1267,1293]" pageId="16" pageNumber="57" value="1906-07-21">
<collectingDate id="EF8DE9B0FFE6C45EFBF3FB54C8EA850D" box="[1027,1157,1267,1293]" pageId="16" pageNumber="57" value="1906-07-21">21.vii.1906</collectingDate>
</date>
, deposited in the collections of the
<collectionCode id="ED66AE5DFFE6C45EFEECFAB0CD3A8531" box="[284,341,1303,1329]" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="16" pageNumber="57">GSC</collectionCode>
, Ottawa. Portions of overlapped fore- and hind wings covered with
<collectingCountry id="F3607608FFE6C45EFBBEFAB0C8CD8531" box="[1102,1186,1303,1329]" name="Canada" pageId="16" pageNumber="57">Canada</collectingCountry>
balsam.
</materialsCitation>
</paragraph>
<paragraph id="8BC83698FFE6C45EFF37FA9CCD7B857A" blockId="16.[151,1437,1266,2013]" pageId="16" pageNumber="57">
<materialsCitation id="3B1F3CC5FFE6C45EFF37FA9CCD7F857A" ID-GBIF-Occurrence="3071122305" country="Canada" location="Horsefly" municipality="Eocene" pageId="16" pageNumber="57" specimenCount="1" stateProvince="British Columbia" typeStatus="holotype">
<emphasis id="B903EA8AFFE6C45EFF37FA9CCE658554" bold="true" box="[199,522,1338,1365]" pageId="16" pageNumber="57">
<typeStatus id="54CC883AFFE6C45EFF37FA9CCD6D8555" box="[199,258,1339,1365]" pageId="16" pageNumber="57">Type</typeStatus>
locality and horizon.
</emphasis>
“Opposite Horsefly mine”,
<location id="8EA86043FFE6C45EFCA6FA9CCFD68555" LSID="urn:lsid:plazi:treatment:03DE878EFFE6C45FFF67FB21CE4982CD:8EA86043FFE6C45EFCA6FA9CCFD68555" box="[854,953,1339,1365]" country="Canada" municipality="Eocene" name="Horsefly" pageId="16" pageNumber="57" stateProvince="British Columbia">Horsefly</location>
,
<collectingRegion id="49B3F87AFFE6C45EFC38FA9CC8FF8555" box="[968,1168,1339,1365]" country="Canada" name="British Columbia" pageId="16" pageNumber="57">British Columbia</collectingRegion>
,
<collectingCountry id="F3607608FFE6C45EFB50FA9CC8978555" box="[1184,1272,1339,1365]" name="Canada" pageId="16" pageNumber="57">Canada</collectingCountry>
; early
<collectingMunicipality id="6BACACE2FFE6C45EFABAFA9CC9F38555" box="[1354,1436,1339,1365]" pageId="16" pageNumber="57">Eocene</collectingMunicipality>
(Ypresian)
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.
</paragraph>
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<subSubSection id="C36D6513FFE6C45EFF37FA25CE9385C1" pageId="16" pageNumber="57" type="diagnosis">
<paragraph id="8BC83698FFE6C45EFF37FA25CE9385C1" blockId="16.[151,1437,1266,2013]" pageId="16" pageNumber="57">
<emphasis id="B903EA8AFFE6C45EFF37FA25CD2D859C" bold="true" box="[199,322,1410,1436]" pageId="16" pageNumber="57">Diagnosis.</emphasis>
May be distinguished from
<taxonomicName id="4C774D1BFFE6C45EFD70FA24CF2C859D" authorityName="Archibald &amp; Makarkin" authorityYear="2021" baseAuthorityName="Engel" baseAuthorityYear="2011" box="[640,835,1411,1437]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="protomaculata">
<emphasis id="B903EA8AFFE6C45EFD70FA24CEFE859D" box="[640,657,1411,1437]" italics="true" pageId="16" pageNumber="57">A</emphasis>
.
<emphasis id="B903EA8AFFE6C45EFD6FFA24CF2C859D" box="[671,835,1411,1437]" italics="true" pageId="16" pageNumber="57">protomaculata</emphasis>
</taxonomicName>
by clearly wider wings (forewing length/width
<quantity id="4C8F9B7DFFE6C45EFAADFA24CC8A85C1" metricMagnitude="-2" metricUnit="m" metricValue="7.3279" metricValueMax="7.8232" metricValueMin="6.8326" pageId="16" pageNumber="57" unit="in" value="2.885" valueMax="3.08" valueMin="2.69">2.69 3.08 in</quantity>
<taxonomicName id="4C774D1BFFE6C45EFF1CFA00CDE985C1" authorityName="Handlirsch" authorityYear="1910" box="[236,390,1447,1473]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="oligoneura">
<emphasis id="B903EA8AFFE6C45EFF1CFA00CC9285C1" box="[236,253,1447,1473]" italics="true" pageId="16" pageNumber="57">A</emphasis>
.
<emphasis id="B903EA8AFFE6C45EFEFBFA00CDE985C1" box="[267,390,1447,1473]" italics="true" pageId="16" pageNumber="57">oligoneura</emphasis>
</taxonomicName>
;
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<taxonomicName id="4C774D1BFFE6C45EFDD9FA00CE8385C1" authorityName="Archibald &amp; Makarkin" authorityYear="2021" baseAuthorityName="Engel" baseAuthorityYear="2011" box="[553,748,1447,1473]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="protomaculata">
<emphasis id="B903EA8AFFE6C45EFDD9FA00CE5585C1" box="[553,570,1447,1473]" italics="true" pageId="16" pageNumber="57">A</emphasis>
.
<emphasis id="B903EA8AFFE6C45EFDB8FA00CE8385C1" box="[584,748,1447,1473]" italics="true" pageId="16" pageNumber="57">protomaculata</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C36D6513FFE6C45FFF37FA6DCD8A8285" lastPageId="17" lastPageNumber="58" pageId="16" pageNumber="57" type="description">
<paragraph id="8BC83698FFE6C45EFF37FA6DCFFA87DD" blockId="16.[151,1437,1266,2013]" pageId="16" pageNumber="57">
<emphasis id="B903EA8AFFE6C45EFF37FA6DCD1A85E4" bold="true" box="[199,373,1482,1508]" pageId="16" pageNumber="57">Redescription.</emphasis>
Forewing
<quantity id="4C8F9B7DFFE6C45EFE1EFA6CCE2485E5" box="[494,587,1483,1509]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="16" pageNumber="57" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">78 mm</quantity>
long (estimated),
<quantity id="4C8F9B7DFFE6C45EFCE7FA6CCF0285E5" box="[791,877,1483,1509]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="16" pageNumber="57" unit="mm" value="2.6">2.6 mm</quantity>
wide. Costal space not preserved. ScP very short, terminating on costal margin strongly distad crossvein 2ra-rp; length from termination of ScP to pterostigma slightly greater than length of pterostigma. Distal part of subcostal space broad, with distal crossvein 2scp-r nearly straight. Pterostigma elongate (4.6 times longer than width measured along to costal margin), lightly pigmented. At least two branches of RA: one within pterostigma, strongly inclined, in middle of pterostigma; other forming distal margin of pterostigma; no branch distad pterostigma discernible. RA space with two crossveins forming two radial cells nearly equal in length: 2ra-rp located about mid-way between pterostigma, termination of ScP; 3ra-rp located in distal part of pterostigma along RA. RP originating at obtuse angle, with three pectinate branches: RP1 deeply forked, with anterior branch simple, posterior forked; other branches (RP2, RP3) simple. One long intraradial crossvein rp1-rp2 between stems of RP, RP1. Two preserved between RP, MA (1r-m, 2r-m) connecting stem of RP, MA. M forked well proximad origin of RP, probably fused with CuA for short distance. MA deeply, dichotomously forked twice. MP strongly zigzagged, anterior trace with two simple branches. Two intramedian crossveins form two
<emphasis id="B903EA8AFFE6C45EFB0FF8F0C94C8771" box="[1279,1315,1879,1905]" italics="true" pageId="16" pageNumber="57">doi</emphasis>
(distal
<emphasis id="B903EA8AFFE6C45EFA88F8F0C9F38771" box="[1400,1436,1879,1905]" italics="true" pageId="16" pageNumber="57">doi</emphasis>
markedly longer): 2im very long, joins posterior branch of MA. Anterior trace CuA strongly zigzagged, fused with MP for long distance to form part of posterior margin of basal
<emphasis id="B903EA8AFFE6C45EFC95F838CFE687B9" box="[869,905,1951,1977]" italics="true" pageId="16" pageNumber="57">doi</emphasis>
. Crossvein icu between CuA, CuP long. Basal portion, CuP, anal veins not preserved or not clearly understandable.
</paragraph>
<paragraph id="8BC83698FFE7C45FFF37FF30CD8A8285" blockId="17.[151,1437,150,717]" pageId="17" pageNumber="58">
Hind wing about
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long (estimated),
<quantity id="4C8F9B7DFFE7C45FFD4FFF31CF4D80B1" box="[703,802,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" pageId="17" pageNumber="58" unit="mm" value="2.25">2.25 mm</quantity>
wide. Costal space and ScP not preserved. One crossvein detected in subcostal space, 2scp-r, forming basal margin of pterostigma. Distal part of subcostal space broad, with distal crossvein 2scp-r nearly straight. Pterostigma elongate (4.7 times longer than width measured along to costal margin), lightly pigmented. At least two branches of RA: one within pterostigma, strongly inclined, in middle of pterostigma; other forming distal margin of pterostigma; no branches distad pterostigma discernible. RA space with two preserved crossveins: 2ra-rp distad termination of ScP; 3ra-rp located in distal part of pterostigma along RA. RP with three pectinate branches: RP1 deeply forked, with anterior branch simple, posterior forked; other branches (RP2, RP3) simple. One long intraradial crossvein rp1-rp2 between stems of RP, RP1. Two preserved between RP, MA: 2r-m connects stem of RP, MA; 3r-m connects stem of RP1, MA. M probably forked proximad origin of RP. MA deeply dichotomously forked twice. MP: anterior trace zigzagged, simple, with three simple, pectinate branches (alternatively: anterior trace forked, with two simple, pectinate branches). Two intramedian crossveins form two
<emphasis id="B903EA8AFFE7C45FFF67FD84CCD4823D" box="[151,187,547,573]" italics="true" pageId="17" pageNumber="58">doi</emphasis>
, proximal much shorter than distal. One preserved crossvein between M, Cu (2m-cu) connecting MP, CuA. Anterior trace of CuA simple distally, with one preserved simple branch. Basal portion, CuP, anal veins not preserved or not clearly understandable.
</paragraph>
</subSubSection>
<subSubSection id="C36D6513FFE7C45FFF37FD28CE4982CD" pageId="17" pageNumber="58" type="discussion">
<paragraph id="8BC83698FFE7C45FFF37FD28CE4982CD" blockId="17.[151,1437,150,717]" pageId="17" pageNumber="58">
<emphasis id="B903EA8AFFE7C45FFF37FD28CD5B82A9" bold="true" box="[199,308,655,681]" pageId="17" pageNumber="58">Remarks</emphasis>
.
<bibRefCitation id="EFE64B69FFE7C45FFEB3FD28CE7F82A9" author="Handlirsch, A." box="[323,528,655,681]" pageId="17" pageNumber="58" pagination="93 - 129" refId="ref23134" refString="Handlirsch, A. (1910) Contributions to Canadian Palaeontology. Vol. II. Part III. Canadian fossil insects. V. Insects from the Tertiary lake deposits of the Southern Interior of British Columbia collected by Mr. Lawrence Lambe, in 1906. Memoirs of the Geological Survey of Canada, 12, 93 - 129. https: // doi. org / 10.4095 / 100486" type="journal article" year="1910">Handlirsch (1910)</bibRefCitation>
incorrectly believed that his
<figureCitation id="134C2A1DFFE7C45FFCA8FD28CFF582A9" box="[856,922,655,681]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[182,1407,190,1909]" captionTargetId="figure-26@6.[165,1423,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Wing venation of Megaraphidia ootsa sp. nov., holotype RBCM P1556. A, right forewing, B, left forewing; C, right hind wing; D, left hind wing. Scale bar = 3 mm (all to scale)." figureDoi="http://doi.org/10.5281/zenodo.4655637" httpUri="https://zenodo.org/record/4655637/files/figure.png" pageId="17" pageNumber="58">Fig. 4</figureCitation>
depicted the forewing and his
<figureCitation id="134C2A1DFFE7C45FFB07FD28C95682A9" box="[1271,1337,654,681]" captionStart="FIGURE 5" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[187,1401,181,1900]" captionTargetId="figure-19@9.[187,1401,181,1919]" captionTargetPageId="9" captionText="FIGURE 5. Megaraphidia klondika sp. nov., holotype. A, counterpart UWBM 97073; B, part SR03-01-01; C, forewing venation; D, hind wing venation. Scale bar = 3 mm (all to scale)." figureDoi="http://doi.org/10.5281/zenodo.4655641" httpUri="https://zenodo.org/record/4655641/files/figure.png" pageId="17" pageNumber="58">Fig. 5</figureCitation>
the hind wing, however, the opposite is true.
</paragraph>
</subSubSection>
</treatment>
</document>