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<taxonomicName id="4C77A3F0FF85ED2884FFFCC2FD79FCD2" authority="A. MASSAL., 1858" box="[251,644,871,895]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" isUncertain="true" kingdom="Plantae" order="Fagales" pageId="14" pageNumber="171" phylum="Tracheophyta" rank="species" species="gussonii">
<emphasis id="B9030461FF85ED2884FFFCC2FD79FCD2" bold="true" box="[251,644,871,895]" pageId="14" pageNumber="171">
<emphasis id="B9030461FF85ED2884FFFCC2FEBCFCD3" bold="true" box="[251,321,871,894]" italics="true" pageId="14" pageNumber="171">Fagus</emphasis>
cf.
<emphasis id="B9030461FF85ED28856DFCC2FE38FCD2" bold="true" box="[361,453,871,895]" italics="true" pageId="14" pageNumber="171">gussonii</emphasis>
A.MASSAL., 1858
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<paragraph id="8BC8D873FF85ED28855BFC2DFDDDFC0D" blockId="14.[351,544,904,928]" box="[351,544,904,928]" pageId="14" pageNumber="171">Pl. 3, Figs 8a, b, 9</paragraph>
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<paragraph id="8BC8D873FF85ED2884A2FC1DFDD6FB9F" blockId="14.[129,768,952,1074]" pageId="14" pageNumber="171">M a t e r i a l. An almost complete leaf (MGPTPU141027) and a fragmentary one (MGPT-PU141099) from GLA20, other two almost complete leaves from GLAz (MGPT-PU141026, MGPT-PU141100).</paragraph>
<paragraph id="8BC8D873FF85ED2884A2FBEFFEBAFA25" blockId="14.[129,768,1097,1416]" pageId="14" pageNumber="171">
D e s c r i p t i o n. Specimen figured on Pl. 3, Fig. 9. Simple leaf petiolate, petiole fragmentary, lamina elliptic, l × w about 50 ×
<quantity id="4C8F7596FF85ED288551FB2EFE5AFB0F" box="[341,423,1163,1187]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="14" pageNumber="171" unit="mm" value="25.0">25 mm</quantity>
, ratio l/w about 2, base shape slightly convex, base angle almost 90°, apex not preserved; margin entire (?), not well preserved; midvein slender, slightly s-shaped; secondaries probably craspedodromous, slender, very regularly spaced, 10 pairs preserved, arising at moderately steep angles, near the base at first slightly converging towards midvein; tertiaries probably percurrent, obtuse to midvein.
</paragraph>
<paragraph id="8BC8D873FF85ED2884A2FA05FEF2F788" blockId="14.[128,768,1440,2085]" pageId="14" pageNumber="171">
R e m a r k s. The leaves from Govone fall within the variability of the abundant Messinian leaves from the surroundings of Alba (
<bibRefCitation id="EFE6A582FF85ED288573FA44FD97FA54" author="Martinetto, E. &amp; Rista, C. &amp; Tarabra, E." box="[375,618,1505,1529]" pageId="14" pageNumber="171" pagination="3 - 30" refId="ref21378" refString="Martinetto, E., Rista, C., Tarabra, E. (2000): Classificazione e ordinamento museologico di foglie di Angiosperme messiniane dell'Albese [Classification and museum organisation of Messinian angiosperm leaves of the Albese region]. - Alba Pompeia, nuova serie, 21 (1): 3 - 30. (in Italian)" type="journal article" year="2000">Martinetto et al. 2000</bibRefCitation>
,
<bibRefCitation id="EFE6A582FF85ED288673FA44FD09FA54" author="Denk, T." box="[631,756,1505,1529]" pageId="14" pageNumber="171" pagination="1 - 72" refId="ref18327" refString="Denk, T. (2004): Revision of Fagus from the Cenozoic of Europe and southwestern Asia and its phylogenetic implications. - Documenta naturae, 150: 1 - 72." type="journal article" year="2004">Denk 2004</bibRefCitation>
).
<bibRefCitation id="EFE6A582FF85ED288485F9A7FEFAF9B7" author="Denk, T." box="[129,263,1538,1562]" pageId="14" pageNumber="171" pagination="1 - 72" refId="ref18327" refString="Denk, T. (2004): Revision of Fagus from the Cenozoic of Europe and southwestern Asia and its phylogenetic implications. - Documenta naturae, 150: 1 - 72." type="journal article" year="2004">Denk (2004)</bibRefCitation>
treated this assemblage as
<taxonomicName id="4C77A3F0FF85ED288622F9A7FF2FF997" authority="KOVATS" authorityName="KOVATS" class="Magnoliopsida" family="Fagaceae" genus="Fagus" isUncertain="true" kingdom="Plantae" order="Fagales" pageId="14" pageNumber="171" phylum="Tracheophyta" rank="species" species="haidingeri">
<emphasis id="B9030461FF85ED288622F9A7FD94F9B4" box="[550,617,1538,1561]" italics="true" pageId="14" pageNumber="171">Fagus</emphasis>
cf.
<emphasis id="B9030461FF85ED288694F9A7FD02F9B7" box="[656,767,1538,1562]" italics="true" pageId="14" pageNumber="171">haidingeri</emphasis>
KOVÁTS
</taxonomicName>
, but considered it to be in the “hybrid zone between
<taxonomicName id="4C77A3F0FF85ED288484F9E1FEEDF9F6" box="[128,272,1603,1627]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="14" pageNumber="171" phylum="Tracheophyta" rank="species" species="haidingeri">
<emphasis id="B9030461FF85ED288484F9E1FF6DF9F6" box="[128,144,1604,1627]" italics="true" pageId="14" pageNumber="171">F</emphasis>
.
<emphasis id="B9030461FF85ED2884A5F9E6FEEDF9F6" box="[161,272,1603,1627]" italics="true" pageId="14" pageNumber="171">haidingeri</emphasis>
</taxonomicName>
/
<taxonomicName id="4C77A3F0FF85ED288513F9E1FE6DF9F6" box="[279,400,1603,1627]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="14" pageNumber="171" phylum="Tracheophyta" rank="species" species="gussonii">
<emphasis id="B9030461FF85ED288513F9E1FEDAF9F6" box="[279,295,1604,1627]" italics="true" pageId="14" pageNumber="171">F</emphasis>
.
<emphasis id="B9030461FF85ED288533F9E1FE6DF9F6" box="[311,400,1604,1627]" italics="true" pageId="14" pageNumber="171">gussonii</emphasis>
</taxonomicName>
”. Later,
<bibRefCitation id="EFE6A582FF85ED2885F7F9E6FD06F9F6" author="Teodoridis, V. &amp; Kvacek, Z. &amp; Sami, M. &amp; Utescher, T. &amp; Martinetto, E." box="[499,763,1603,1627]" pageId="14" pageNumber="171" pagination="249 - 292" refId="ref23735" refString="Teodoridis, V., Kvacek, Z., Sami, M., Utescher, T., Martinetto, E. (2015 b): Palaeoenvironmental analysis of the Messinian macrofossil floras of Tossignano and Monte Tondo (Vena del Gesso basin, Romagna Apennines, northern Italy). - Acta Musei Nationalis Pragae, Series B - Historia Naturalis, 71 (3 - 4): 249 - 292." type="journal article" year="2015">Teodoridis et al. (2015b</bibRefCitation>
,
<bibRefCitation id="EFE6A582FF85ED288484F9C1FF4AF9D1" author="Teodoridis, V. &amp; Kvacek, Z. &amp; Sami, M." box="[128,183,1636,1660]" pageId="14" pageNumber="171" pagination="221 - 237" refId="ref23812" refString="Teodoridis, V., Kvacek, Z., Sami, M. (2017): Revision of the Messinian flora of Polenta (Romagna Apennines, Northern Italy). - Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen, 283: 221 - 237. https: // doi. org / 10.1127 / njgpa / 2017 / 0638" type="journal article" year="2017">2017</bibRefCitation>
) assigned Messinian specimens from Romagna (northern
<collectingCountry id="F36098E3FF85ED2884EBF921FEDCF931" box="[239,289,1668,1692]" name="Italy" pageId="14" pageNumber="171">Italy</collectingCountry>
) to
<taxonomicName id="4C77A3F0FF85ED288557F920FE33F931" box="[339,462,1669,1692]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="14" pageNumber="171" phylum="Tracheophyta" rank="species" species="gussonii">
<emphasis id="B9030461FF85ED288557F920FE9EF931" box="[339,355,1669,1692]" italics="true" pageId="14" pageNumber="171">F</emphasis>
.
<emphasis id="B9030461FF85ED288571F920FE33F931" box="[373,462,1669,1692]" italics="true" pageId="14" pageNumber="171">gussonii</emphasis>
</taxonomicName>
. As we do not see definite differences between
<taxonomicName id="4C77A3F0FF85ED288564F903FE5EF910" box="[352,419,1702,1725]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="14" pageNumber="171" phylum="Tracheophyta" rank="genus">
<emphasis id="B9030461FF85ED288564F903FE5EF910" box="[352,419,1702,1725]" italics="true" pageId="14" pageNumber="171">Fagus</emphasis>
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leaves from the
<typeStatus id="54CC66D1FF85ED28865FF903FD74F913" box="[603,649,1702,1726]" pageId="14" pageNumber="171">type</typeStatus>
locality of
<taxonomicName id="4C77A3F0FF85ED288484F963FE86F973" authority="(Senigallia)" baseAuthorityName="Senigallia" box="[128,379,1734,1758]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="14" pageNumber="171" phylum="Tracheophyta" rank="species" species="gussonii">
<emphasis id="B9030461FF85ED288484F963FF6DF970" box="[128,144,1734,1757]" italics="true" pageId="14" pageNumber="171">F</emphasis>
.
<emphasis id="B9030461FF85ED28849AF963FF0AF970" box="[158,247,1734,1757]" italics="true" pageId="14" pageNumber="171">gussonii</emphasis>
(Senigallia)
</taxonomicName>
, and those of the Romagna and Alba areas, we propose the use of the name
<taxonomicName id="4C77A3F0FF85ED288613F942FD27F953" box="[535,730,1766,1790]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" isUncertain="true" kingdom="Plantae" order="Fagales" pageId="14" pageNumber="171" phylum="Tracheophyta" rank="species" species="gussonii">
<emphasis id="B9030461FF85ED288613F942FDA7F953" box="[535,602,1767,1790]" italics="true" pageId="14" pageNumber="171">Fagus</emphasis>
cf.
<emphasis id="B9030461FF85ED288685F942FD27F953" box="[641,730,1767,1790]" italics="true" pageId="14" pageNumber="171">gussonii</emphasis>
</taxonomicName>
for those from the Alba region. Rich occurrences of
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<emphasis id="B9030461FF85ED288687F8ADFD6EF8B2" box="[643,659,1800,1823]" italics="true" pageId="14" pageNumber="171">F</emphasis>
.
<emphasis id="B9030461FF85ED2886A4F8ADFD04F8B2" box="[672,761,1800,1823]" italics="true" pageId="14" pageNumber="171">gussonii</emphasis>
</taxonomicName>
, e.g., in
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(
<bibRefCitation id="EFE6A582FF85ED288530F88DFE06F8ED" author="Guner, T. H. &amp; Bouchal, J. M. &amp; Kose, N. &amp; Goktas, F. &amp; Mayda, S. &amp; Denk, T." box="[308,507,1832,1856]" pageId="14" pageNumber="171" pagination="113 - 171" refId="ref19443" refString="Guner, T. H., Bouchal, J. M., Kose, N., Goktas, F., Mayda, S., Denk, T. (2017): Landscape heterogeneity in the Yatagan Basin (southwestern Turkey) during the middle Miocene inferred from plant macrofossils. - Palaeontographica, Abt. B, 296 (1 - 6): 113 - 171. https: // doi. org / 10.1127 / palb / 296 / 2017 / 113" type="journal article" year="2017">Güner et al. 2017</bibRefCitation>
) show wide variability, including leaves identical to those from Govone, and from northern
<collectingCountry id="F36098E3FF85ED2884E4F8CCFEECF82C" box="[224,273,1897,1921]" name="Italy" pageId="14" pageNumber="171">Italy</collectingCountry>
in general. The variation in leaf morphology of
<taxonomicName id="4C77A3F0FF85ED288484F82EFF3EF80F" box="[128,195,1931,1954]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="14" pageNumber="171" phylum="Tracheophyta" rank="genus">
<emphasis id="B9030461FF85ED288484F82EFF3EF80F" box="[128,195,1931,1954]" italics="true" pageId="14" pageNumber="171">Fagus</emphasis>
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of the rich northwestern Italian Pliocene assemblages (
<bibRefCitation id="EFE6A582FF85ED28848CF80EFEC0F86E" author="Martinetto, E." box="[136,317,1963,1987]" pageId="14" pageNumber="171" pagination="11 - 75" refId="ref21082" refString="Martinetto, E. (2003): Leaves of terrestrial plants from the shallow marine and transitional Pliocene deposits of Asti (Piedmont, NW Italy). - Bollettino della Societa Paleontologica Italiana, 42: 11 - 75." type="journal article" year="2003">Martinetto 2003</bibRefCitation>
,
<bibRefCitation id="EFE6A582FF85ED28854FF80EFE3BF86E" author="Denk, T." box="[331,454,1963,1987]" pageId="14" pageNumber="171" pagination="1 - 72" refId="ref18327" refString="Denk, T. (2004): Revision of Fagus from the Cenozoic of Europe and southwestern Asia and its phylogenetic implications. - Documenta naturae, 150: 1 - 72." type="journal article" year="2004">Denk 2004</bibRefCitation>
) differs from the Messinian ones. Therefore, it appears justified to apply a different name to the Pliocene beech leaves, i.e.,
<taxonomicName id="4C77A3F0FF85ED28866DF848FEF1F788" authority="(Denk 2004)" baseAuthorityName="Denk" baseAuthorityYear="2004" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="14" pageNumber="171" phylum="Tracheophyta" rank="species" species="haidingeri">
<emphasis id="B9030461FF85ED28866DF848FD84F7A9" box="[617,633,2029,2052]" italics="true" pageId="14" pageNumber="171">F</emphasis>
.
<emphasis id="B9030461FF85ED288694F849FD02F7A9" box="[656,767,2028,2052]" italics="true" pageId="14" pageNumber="171">haidingeri</emphasis>
(
<bibRefCitation id="EFE6A582FF85ED28848CF7A8FEF9F788" author="Denk, T." box="[136,260,2061,2085]" pageId="14" pageNumber="171" pagination="1 - 72" refId="ref18327" refString="Denk, T. (2004): Revision of Fagus from the Cenozoic of Europe and southwestern Asia and its phylogenetic implications. - Documenta naturae, 150: 1 - 72." type="journal article" year="2004">Denk 2004</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
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