266 lines
31 KiB
XML
266 lines
31 KiB
XML
<document id="55E431FB4ACC7C5E47807528CAE5EBD1" ID-CLB-Dataset="8703" ID-DOI="10.11646/zootaxa.4825.1.1" ID-GBIF-Dataset="5ab5f5c8-481e-4d1a-8643-21e72c367278" ID-ISSN="1175-5326" ID-Zenodo-Dep="4402145" ID-ZooBank="F230F199-1C94-4E2E-9CE4-5F56212C015F" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1609341789210" checkinUser="plazi" docAuthor="Schileyko, Arkady A., Vahtera, Varpu & Edgecombe, Gregory D." docDate="2020" docId="03DE092DFFE5D713FF13F97C2F4CDD02" docLanguage="en" docName="zootaxa.4825.1.1.pdf" docOrigin="Zootaxa 4825 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Theatops Newport 1844" docType="treatment" docVersion="7" lastPageNumber="25" masterDocId="FFE77155FFF2D70AFF84FFE12D45DC1F" masterDocTitle="An overview of the extant genera and subgenera of the order Scolopendromorpha (Chilopoda): a new identification key and updated diagnoses" masterLastPageNumber="64" masterPageNumber="1" pageNumber="24" updateTime="1698852004925" updateUser="ExternalLinkService">
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<mods:titleInfo id="538E652BB078420D865E5B4DFAD278F4">
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<mods:title id="239AF601389E3C64A57B3F60C6AFEE6E">An overview of the extant genera and subgenera of the order Scolopendromorpha (Chilopoda): a new identification key and updated diagnoses</mods:title>
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<mods:name id="CC73A315702992D000A1F75E664BF7E6" type="personal">
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<mods:namePart id="278F73F81A9661BD8D9A8B654D9EBBB8">Schileyko, Arkady A.</mods:namePart>
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<mods:nameIdentifier id="A63020178C893CFF10DF3D0BBD990009" type="email">schileyko1965@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="B36D5ADE32FA3E98EA3054C86A9FCC15">Author</mods:roleTerm>
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<mods:namePart id="7239E4E166CA6CD74E11F9F2DBBC1CC1">Vahtera, Varpu</mods:namePart>
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<mods:nameIdentifier id="2931258134D1C739F7EAD1B131BA5523" type="email">varpu.vahtera@gmail.com</mods:nameIdentifier>
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<mods:namePart id="B897D970DADCFEED334936182B5D3075">Edgecombe, Gregory D.</mods:namePart>
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<mods:nameIdentifier id="D13B5DF5A61A197E56E6CEDD3BB28FDA" type="email">schileyko1965@gmail.com</mods:nameIdentifier>
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<mods:title id="85AA3256D4CC70AAE442B57B2B3CD59F">Zootaxa</mods:title>
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<mods:date id="6123B16EF576E572630709E37610AAB6">2020</mods:date>
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<mods:number id="6D7E9EC57329526AFD57CB8961FE5143">4825</mods:number>
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<treatment id="03DE092DFFE5D713FF13F97C2F4CDD02" ID-DOI="http://doi.org/10.5281/zenodo.4455413" ID-GBIF-Taxon="177020716" ID-Zenodo-Dep="4455413" LSID="urn:lsid:plazi:treatment:03DE092DFFE5D713FF13F97C2F4CDD02" httpUri="http://treatment.plazi.org/id/03DE092DFFE5D713FF13F97C2F4CDD02" lastPageId="25" lastPageNumber="25" pageId="23" pageNumber="24">
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<subSubSection id="C36DEBB0FFE5D71DFF13F97C2C50DAC4" pageId="23" pageNumber="24" type="discussion">
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<subSubSection id="C36DEBB0FFE5D71DFF13F97C2CACDAA8" box="[151,489,1693,1720]" pageId="23" pageNumber="24" type="nomenclature">
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<paragraph id="8BC8B83BFFE5D71DFF13F97C2CACDAA8" blockId="23.[151,489,1693,1755]" box="[151,489,1693,1720]" pageId="23" pageNumber="24">
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<emphasis id="B9036429FFE5D71DFF13F97C2CACDAA8" bold="true" box="[151,489,1693,1720]" pageId="23" pageNumber="24">
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(!)
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<taxonomicName id="4C77C3B8FFE5D71DFF38F97C2CACDAA8" ID-CoL="646L5" ID-ENA="60165" authority="Newport, 1844" authorityName="Newport" authorityYear="1844" box="[188,489,1693,1720]" class="Chilopoda" family="Cryptopidae" genus="Theatops" kingdom="Animalia" order="Scolopendromorpha" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
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<emphasis id="B9036429FFE5D71DFF38F97C2C6EDAA7" bold="true" box="[188,299,1693,1720]" italics="true" pageId="23" pageNumber="24">Theatops</emphasis>
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<collectingRegion id="49B376D9FFE5D71DFEB6F97C2CE4DAA8" box="[306,417,1693,1719]" country="United Kingdom" name="Newport" pageId="23" pageNumber="24">Newport</collectingRegion>
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, 1844
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</taxonomicName>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36DEBB0FFE5D71DFF13F9202C50DAC4" box="[151,277,1729,1755]" pageId="23" pageNumber="24" type="description">
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<paragraph id="8BC8B83BFFE5D71DFF13F9202C50DAC4" blockId="23.[151,489,1693,1755]" box="[151,277,1729,1755]" pageId="23" pageNumber="24">
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<figureCitation id="134CA4BEFFE5D71DFF13F9202C50DAC4" box="[151,277,1729,1755]" captionStart="FIGURES 33–39" captionStartId="24.[151,264,1627,1651]" captionTargetBox="[172,1415,293,1582]" captionTargetId="figure-53@24.[151,1436,272,1603]" captionTargetPageId="24" captionText="FIGURES 33–39. Theatops posticus (Say, 1821); Sad. Rc 6488; 33 ultimate and penultimate LBS + ultimate legs dorsally; Theatops spinicaudus Wood, 1862; Ad. Rc 6489 34 head + LBS 1 dorsally; Theatops erythrocephalus (C. L. Koch, 1847); ZMB 3393 35 apical part of maxilla 2 laterally; Theatops spinicaudus Wood, 1862; Ad. Rc 6489 36 head + forcipular segment ventrally; Theatops posticus (Say, 1821); Sad. Rc 6488 37 LBS 21 + ultimate legs ventrally; Tonkinodentus lestes Schileyko, 1992; Holotype ad. Rc 6358 38 ultimate + penultimate LBS dorsally 39 cephalic plate + LBS 1 dorsally; (cst)—forcipular coxosternite, (cx)—coxopleuron, (ds)—depigmented eye spot, (p2)—non pectinate pretarsus of maxilla 2, (pf)—prefemur, (pr)—trochantero-prefemural process, (pts)—pretarsus, (T1)—tergite 1, (t2)—tarsus 2, (tp)—forcipular tooth-plate, (ust)—ultimate sternite, (ut)—ultimate tergite, (vp)—short ventral projection of pretarsus of maxilla 2." figureDoi="http://doi.org/10.5281/zenodo.4402161" httpUri="https://zenodo.org/record/4402161/files/figure.png" pageId="23" pageNumber="24">Figs 33–37</figureCitation>
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</paragraph>
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</subSubSection>
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</subSubSection>
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<subSubSection id="C36DEBB0FFE5D71DFF13F8E72E31DB01" box="[151,884,1798,1822]" pageId="23" pageNumber="24" type="type_taxon">
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<paragraph id="8BC8B83BFFE5D71DFF13F8E72E31DB01" blockId="23.[151,884,1798,1822]" box="[151,884,1798,1822]" pageId="23" pageNumber="24">
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<emphasis id="B9036429FFE5D71DFF13F8E72C66DB01" bold="true" box="[151,291,1798,1822]" pageId="23" pageNumber="24">
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<typeStatus id="54CC0699FFE5D71DFF13F8E72D89DB01" box="[151,204,1798,1822]" pageId="23" pageNumber="24">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C77C3B8FFE5D71DFEAEF8E72F0FDB01" authority="Say, 1821" authorityName="Say" authorityYear="1821" box="[298,586,1798,1822]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="posticus">
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<emphasis id="B9036429FFE5D71DFEAEF8E72CA5DB02" box="[298,480,1798,1821]" italics="true" pageId="23" pageNumber="24">Cryptops posticus</emphasis>
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Say, 1821
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</taxonomicName>
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(by subsequent designation).
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</paragraph>
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</subSubSection>
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<subSubSection id="C36DEBB0FFE5D71DFF13F8AD2CC6DBE8" pageId="23" pageNumber="24" type="diagnosis">
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<paragraph id="8BC8B83BFFE5D71DFF13F8AD2CC6DBE8" blockId="23.[151,1436,1868,2039]" pageId="23" pageNumber="24">
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<emphasis id="B9036429FFE5D71DFF13F8AD2C57DB79" bold="true" box="[151,274,1868,1894]" pageId="23" pageNumber="24">Diagnosis.</emphasis>
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Spiracles (figs 1AB in
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<bibRefCitation id="EFE6C5CAFFE5D71DFD9AF8AD2E43DB79" author="Vahtera, V. & Edgecombe, G. D. & Giribet, G." box="[542,774,1868,1895]" pageId="23" pageNumber="24" pagination="225 - 248" refId="ref51336" refString="Vahtera, V., Edgecombe, G. D. & Giribet, G. (2012 b) Spiracle structure in scolopendromorph centipedes (Chilopoda: Scolopendromorpha) and its contribution to phylogenetics. Zoomorphology, 131, 225 - 248. https: // doi. org / 10.1007 / s 00435 - 012 - 0157 - 0" type="journal article" year="2012">
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Vahtera
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<emphasis id="B9036429FFE5D71DFDF8F8AC2FEADB79" box="[636,687,1868,1894]" italics="true" pageId="23" pageNumber="24">et al</emphasis>
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||
. 2012b
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||
</bibRefCitation>
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||
, fig. 5 F in
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<bibRefCitation id="EFE6C5CAFFE5D71DFC09F8AC2927DB79" author="Bonato, L. & Orlando, M. & Zapparoli, M. & Fusco, G. & Bortolin, F." box="[909,1122,1868,1895]" pageId="23" pageNumber="24" pagination="1 - 23" refId="ref45789" refString="Bonato, L., Orlando, M., Zapparoli, M., Fusco, G. & Bortolin, F. (2017) New insights into Plutonium, one of the largest and least known European centipedes (Chilopoda): distribution, evolution and morphology. Zoological Journal of the Linnean Society, 20, 1 - 23. https: // doi. org / 10.1093 / zoolinnean / zlw 026" type="journal article" year="2017">
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Bonato
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<emphasis id="B9036429FFE5D71DFC62F8AC295CDB79" box="[998,1049,1868,1894]" italics="true" pageId="23" pageNumber="24">et al</emphasis>
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. 2017
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</bibRefCitation>
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) on macrosegments, LBS 7 with or without spiracles. Coxopleuron (
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<figureCitation id="134CA4BEFFE5D71DFDDAF8912FF5DB95" box="[606,688,1904,1930]" captionStart="FIGURES 33–39" captionStartId="24.[151,264,1627,1651]" captionTargetBox="[172,1415,293,1582]" captionTargetId="figure-53@24.[151,1436,272,1603]" captionTargetPageId="24" captionText="FIGURES 33–39. Theatops posticus (Say, 1821); Sad. Rc 6488; 33 ultimate and penultimate LBS + ultimate legs dorsally; Theatops spinicaudus Wood, 1862; Ad. Rc 6489 34 head + LBS 1 dorsally; Theatops erythrocephalus (C. L. Koch, 1847); ZMB 3393 35 apical part of maxilla 2 laterally; Theatops spinicaudus Wood, 1862; Ad. Rc 6489 36 head + forcipular segment ventrally; Theatops posticus (Say, 1821); Sad. Rc 6488 37 LBS 21 + ultimate legs ventrally; Tonkinodentus lestes Schileyko, 1992; Holotype ad. Rc 6358 38 ultimate + penultimate LBS dorsally 39 cephalic plate + LBS 1 dorsally; (cst)—forcipular coxosternite, (cx)—coxopleuron, (ds)—depigmented eye spot, (p2)—non pectinate pretarsus of maxilla 2, (pf)—prefemur, (pr)—trochantero-prefemural process, (pts)—pretarsus, (T1)—tergite 1, (t2)—tarsus 2, (tp)—forcipular tooth-plate, (ust)—ultimate sternite, (ut)—ultimate tergite, (vp)—short ventral projection of pretarsus of maxilla 2." figureDoi="http://doi.org/10.5281/zenodo.4402161" httpUri="https://zenodo.org/record/4402161/files/figure.png" pageId="23" pageNumber="24">Fig. 37</figureCitation>
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) usually with a spine at the place of corresponding process. Prefemur and femur of the ultimate legs (
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<figureCitation id="134CA4BEFFE5D71DFDB0F8752FF3DBB1" box="[564,694,1940,1966]" captionStart="FIGURES 33–39" captionStartId="24.[151,264,1627,1651]" captionTargetBox="[172,1415,293,1582]" captionTargetId="figure-53@24.[151,1436,272,1603]" captionTargetPageId="24" captionText="FIGURES 33–39. Theatops posticus (Say, 1821); Sad. Rc 6488; 33 ultimate and penultimate LBS + ultimate legs dorsally; Theatops spinicaudus Wood, 1862; Ad. Rc 6489 34 head + LBS 1 dorsally; Theatops erythrocephalus (C. L. Koch, 1847); ZMB 3393 35 apical part of maxilla 2 laterally; Theatops spinicaudus Wood, 1862; Ad. Rc 6489 36 head + forcipular segment ventrally; Theatops posticus (Say, 1821); Sad. Rc 6488 37 LBS 21 + ultimate legs ventrally; Tonkinodentus lestes Schileyko, 1992; Holotype ad. Rc 6358 38 ultimate + penultimate LBS dorsally 39 cephalic plate + LBS 1 dorsally; (cst)—forcipular coxosternite, (cx)—coxopleuron, (ds)—depigmented eye spot, (p2)—non pectinate pretarsus of maxilla 2, (pf)—prefemur, (pr)—trochantero-prefemural process, (pts)—pretarsus, (T1)—tergite 1, (t2)—tarsus 2, (tp)—forcipular tooth-plate, (ust)—ultimate sternite, (ut)—ultimate tergite, (vp)—short ventral projection of pretarsus of maxilla 2." figureDoi="http://doi.org/10.5281/zenodo.4402161" httpUri="https://zenodo.org/record/4402161/files/figure.png" pageId="23" pageNumber="24">Figs 33, 37</figureCitation>
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) with or without spines; pretarsus not shorter than corresponding tarsus 2 (in
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<taxonomicName id="4C77C3B8FFE5D71DFEA5F8582F63DBCD" authority="(Say, 1821)" baseAuthorityName="Say" baseAuthorityYear="1821" box="[289,550,1976,2002]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="posticus">
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<emphasis id="B9036429FFE5D71DFEA5F8582CDFDBCD" box="[289,410,1977,2002]" italics="true" pageId="23" pageNumber="24">T. posticus</emphasis>
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(Say, 1821)
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</taxonomicName>
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longer than the two tarsal articles together), circular in cross section (i.e. not blade-like ventrally).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C36DEBB0FFEAD713FF43FF772F4CDD02" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="discussion">
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<paragraph id="8BC8B83BFFEAD712FF43FF772FEBDCAE" blockId="24.[151,1437,150,249]" box="[199,686,150,177]" pageId="24" pageNumber="25">
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||
<emphasis id="B9036429FFEAD712FF43FF772CECDCAF" bold="true" box="[199,425,150,176]" pageId="24" pageNumber="25">Number of species.</emphasis>
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||
6 (
|
||
<bibRefCitation id="EFE6C5CAFFEAD712FE49FF772FE7DCAE" author="Bonato, L. & Chagas-Junior, A. & Edgecombe, G. D. & Lewis, J. G. E. & Minelli, A. & Pereira, L. A. & Shelley, R. M. & Stoev, P. & Zapparoli, M." box="[461,674,150,177]" pageId="24" pageNumber="25" refId="ref45701" refString="Bonato, L., Chagas-Junior, A., Edgecombe, G. D., Lewis, J. G. E., Minelli, A., Pereira, L. A., Shelley, R. M., Stoev, P. & Zapparoli, M. (2016) ChiloBase 2.0 - A World Catalogue of Centipedes (Chilopoda). Available from: http: // chilobase. biologia. unipd. it (accessed 2 August 2016)" type="url" year="2016">
|
||
Bonato
|
||
<emphasis id="B9036429FFEAD712FDA2FF762F1BDCAF" box="[550,606,150,176]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
|
||
2016
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC8B83BFFEAD712FF43FF5B2C87DCE7" blockId="24.[151,1437,150,249]" pageId="24" pageNumber="25">
|
||
<emphasis id="B9036429FFEAD712FF43FF5B2C7EDCCB" bold="true" box="[199,315,186,212]" pageId="24" pageNumber="25">Remarks.</emphasis>
|
||
Treated as a genus in
|
||
<bibRefCitation id="EFE6C5CAFFEAD712FDB1FF5B2E4ADCCA" author="Di, Z. & Cao, Z. & Wu, Y. & Yin, S. & Edgecombe, G. D. & Li, W." box="[565,783,186,213]" pageId="24" pageNumber="25" pagination="51 - 63" refId="ref47108" refString="Di, Z., Cao, Z., Wu, Y., Yin, S., Edgecombe, G. D. & Li, W. (2010) Discovery of the centipede family Plutoniumidae (Chilopoda) in Asia: a new species of Theatops from China, and the taxonomic value of spiracle distributions in Scolopendromorpha. Zootaxa, 2667 (1), 51 - 63. https: // doi. org / 10.11646 / zootaxa. 2667.1.4" type="journal article" year="2010">
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Di
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||
<emphasis id="B9036429FFEAD712FDDCFF5A2FD5DCCB" box="[600,656,186,212]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
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||
(2010: 51)
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||
</bibRefCitation>
|
||
,
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||
<bibRefCitation id="EFE6C5CAFFEAD712FC9FFF5B29DADCCA" author="Edgecombe, G. D. & Bonato, L." box="[795,1183,186,213]" pageId="24" pageNumber="25" refId="ref47498" refString="Edgecombe, G. D. & Bonato, L. (2011) Chilopoda-taxonomic overview. Order Scolopendromorpha. In: Minelli, A. (Ed.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 1. Brill, Leiden, pp. 392 - 407." type="book" year="2011">Edgecombe & Bonato (2011: 395)</bibRefCitation>
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,
|
||
<bibRefCitation id="EFE6C5CAFFEAD712FB2FFF5B2DF4DCE6" author="Vahtera, V. & Edgecombe, G. D. & Giribet, G." pageId="24" pageNumber="25" pagination="4 - 20" refId="ref51271" refString="Vahtera, V., Edgecombe, G. D. & Giribet, G. (2012 a) Evolution of blindness in scolopendromorph centipedes (Chilopoda: Scolopendromorpha): insight from an expanded sampling of molecular data. Cladistics, 28, 4 - 20. https: // doi. org / 10.1111 / j. 1096 - 0031.2011.00361. x" type="journal article" year="2012">
|
||
Vahtera
|
||
<emphasis id="B9036429FFEAD712FA83FF5A2805DCCB" box="[1287,1344,186,212]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
|
||
(2012a: 5)
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE6C5CAFFEAD712FF39FF3E2CF8DCE7" author="Bonato, L. & Orlando, M. & Zapparoli, M. & Fusco, G. & Bortolin, F." box="[189,445,222,249]" pageId="24" pageNumber="25" pagination="1 - 23" refId="ref45789" refString="Bonato, L., Orlando, M., Zapparoli, M., Fusco, G. & Bortolin, F. (2017) New insights into Plutonium, one of the largest and least known European centipedes (Chilopoda): distribution, evolution and morphology. Zoological Journal of the Linnean Society, 20, 1 - 23. https: // doi. org / 10.1093 / zoolinnean / zlw 026" type="journal article" year="2017">
|
||
Bonato
|
||
<emphasis id="B9036429FFEAD712FE90FF3E2C08DCE7" box="[276,333,222,248]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
|
||
(2017: 1)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF08E8B3FFEAD712FF13F9BA294EDB70" ID-DOI="http://doi.org/10.5281/zenodo.4402161" ID-Zenodo-Dep="4402161" httpUri="https://zenodo.org/record/4402161/files/figure.png" pageId="24" pageNumber="25" startId="24.[151,264,1627,1651]" targetBox="[172,1415,293,1582]" targetPageId="24">
|
||
<paragraph id="8BC8B83BFFEAD712FF13F9BA294EDB70" blockId="24.[151,1436,1627,1903]" pageId="24" pageNumber="25">
|
||
<emphasis id="B9036429FFEAD712FF13F9BA2C13DA6C" bold="true" box="[151,342,1627,1651]" pageId="24" pageNumber="25">FIGURES 33–39.</emphasis>
|
||
<taxonomicName id="4C77C3B8FFEAD712FEDAF9BA2FDDDA6C" authority="(Say, 1821)" baseAuthorityName="Say" baseAuthorityYear="1821" box="[350,664,1627,1651]" class="Chilopoda" family="Cryptopidae" genus="Theatops" kingdom="Animalia" order="Scolopendromorpha" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="posticus">
|
||
<emphasis id="B9036429FFEAD712FEDAF9BA2F53DA6C" box="[350,534,1627,1651]" italics="true" pageId="24" pageNumber="25">Theatops posticus</emphasis>
|
||
(Say, 1821)
|
||
</taxonomicName>
|
||
; Sad. Rc 6488;
|
||
<emphasis id="B9036429FFEAD712FCC4F9BA2E1CDA6C" bold="true" box="[832,857,1627,1651]" pageId="24" pageNumber="25">33</emphasis>
|
||
ultimate and penultimate LBS + ultimate legs dorsally;
|
||
<taxonomicName id="4C77C3B8FFEAD712FF13F99E2CB3DA88" authority="Wood, 1862" authorityName="Wood" authorityYear="1862" box="[151,502,1663,1687]" class="Chilopoda" family="Cryptopidae" genus="Theatops" kingdom="Animalia" order="Scolopendromorpha" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="spinicaudus">
|
||
<emphasis id="B9036429FFEAD712FF13F99E2C36DA88" box="[151,371,1663,1687]" italics="true" pageId="24" pageNumber="25">Theatops spinicaudus</emphasis>
|
||
Wood, 1862
|
||
</taxonomicName>
|
||
; Ad. Rc 6489
|
||
<emphasis id="B9036429FFEAD712FD06F99E2FDEDA88" bold="true" box="[642,667,1663,1687]" pageId="24" pageNumber="25">34</emphasis>
|
||
head + LBS 1 dorsally;
|
||
<taxonomicName id="4C77C3B8FFEAD712FC0BF99E2819DA88" authority="(C. L. Koch, 1847)" baseAuthorityName="C. L. Koch" baseAuthorityYear="1847" box="[911,1372,1663,1687]" class="Chilopoda" family="Cryptopidae" genus="Theatops" kingdom="Animalia" order="Scolopendromorpha" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="erythrocephalus">
|
||
<emphasis id="B9036429FFEAD712FC0BF99E29D0DA88" box="[911,1173,1663,1687]" italics="true" pageId="24" pageNumber="25">Theatops erythrocephalus</emphasis>
|
||
(C. L. Koch, 1847)
|
||
</taxonomicName>
|
||
; ZMB 3393
|
||
<emphasis id="B9036429FFEAD712FF55F9422DAFDAA4" bold="true" box="[209,234,1699,1723]" pageId="24" pageNumber="25">35</emphasis>
|
||
apical part of maxilla 2 laterally;
|
||
<taxonomicName id="4C77C3B8FFEAD712FDC3F9422EEFDAA4" authority="Wood, 1862" authorityName="Wood" authorityYear="1862" box="[583,938,1699,1723]" class="Chilopoda" family="Cryptopidae" genus="Theatops" kingdom="Animalia" order="Scolopendromorpha" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="spinicaudus">
|
||
<emphasis id="B9036429FFEAD712FDC3F9422E61DAA4" box="[583,804,1699,1723]" italics="true" pageId="24" pageNumber="25">Theatops spinicaudus</emphasis>
|
||
Wood, 1862
|
||
</taxonomicName>
|
||
; Ad. Rc 6489
|
||
<emphasis id="B9036429FFEAD712FBB8F9422910DAA4" bold="true" box="[1084,1109,1699,1723]" pageId="24" pageNumber="25">36</emphasis>
|
||
head + forcipular segment ventrally;
|
||
<taxonomicName id="4C77C3B8FFEAD712FF5EF9262F48DAC0" authority="(Say, 1821)" baseAuthorityName="Say" baseAuthorityYear="1821" box="[218,525,1735,1759]" class="Chilopoda" family="Cryptopidae" genus="Theatops" kingdom="Animalia" order="Scolopendromorpha" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="posticus">
|
||
<emphasis id="B9036429FFEAD712FF5EF9262CCADAC0" box="[218,399,1735,1759]" italics="true" pageId="24" pageNumber="25">Theatops posticus</emphasis>
|
||
(Say, 1821)
|
||
</taxonomicName>
|
||
; Sad. Rc 6488
|
||
<emphasis id="B9036429FFEAD712FD20F9262FF8DAC0" bold="true" box="[676,701,1735,1759]" pageId="24" pageNumber="25">37</emphasis>
|
||
LBS 21 + ultimate legs ventrally;
|
||
<taxonomicName id="4C77C3B8FFEAD712FB9EF92628DCDAC0" authority="Schileyko, 1992" authorityName="Schileyko" authorityYear="1992" box="[1050,1433,1735,1759]" class="Chilopoda" family="Cryptopidae" genus="Tonkinodentus" kingdom="Animalia" order="Scolopendromorpha" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="lestes">
|
||
<emphasis id="B9036429FFEAD712FB9EF92629AEDAC0" box="[1050,1259,1735,1759]" italics="true" pageId="24" pageNumber="25">Tonkinodentus lestes</emphasis>
|
||
<bibRefCitation id="EFE6C5CAFFEAD712FB75F92628DCDAC0" author="Schileyko" box="[1265,1433,1735,1759]" pageId="24" pageNumber="25" pagination="5 - 19" refId="ref49894" refString="Schileyko, Ark. A. (1992) Scolopenders of Vietnam and some aspects of the system of Scolopendromorpha (Chilopoda Epimorpha) Part 1. Arthropoda Selecta, 1 (1), 5 - 19." type="journal article" year="1992">Schileyko, 1992</bibRefCitation>
|
||
</taxonomicName>
|
||
; Holotype ad. Rc 6358
|
||
<emphasis id="B9036429FFEAD712FEFAF90A2CD2DB1C" bold="true" box="[382,407,1771,1795]" pageId="24" pageNumber="25">38</emphasis>
|
||
ultimate + penultimate LBS dorsally
|
||
<emphasis id="B9036429FFEAD712FC9EF90A2E76DB1C" bold="true" box="[794,819,1771,1795]" pageId="24" pageNumber="25">39</emphasis>
|
||
cephalic plate + LBS 1 dorsally; (
|
||
<emphasis id="B9036429FFEAD712FB10F90A29F7DB1C" bold="true" box="[1172,1202,1771,1795]" pageId="24" pageNumber="25">cst</emphasis>
|
||
)—forcipular coxosternite, (
|
||
<emphasis id="B9036429FFEAD712FF57F8EE2DAEDB38" bold="true" box="[211,235,1807,1831]" pageId="24" pageNumber="25">cx</emphasis>
|
||
)—coxopleuron, (
|
||
<emphasis id="B9036429FFEAD712FE26F8EE2CFFDB38" bold="true" box="[418,442,1807,1831]" pageId="24" pageNumber="25">ds</emphasis>
|
||
)—depigmented eye spot, (
|
||
<emphasis id="B9036429FFEAD712FD54F8EE2FAEDB38" bold="true" box="[720,747,1807,1831]" pageId="24" pageNumber="25">p2</emphasis>
|
||
)—non pectinate pretarsus of maxilla 2, (
|
||
<emphasis id="B9036429FFEAD712FB16F8EE29ECDB38" bold="true" box="[1170,1193,1807,1831]" pageId="24" pageNumber="25">pf</emphasis>
|
||
)—prefemur, (
|
||
<emphasis id="B9036429FFEAD712FAB9F8EE2813DB38" bold="true" box="[1341,1366,1807,1831]" pageId="24" pageNumber="25">pr</emphasis>
|
||
)—trochantero-prefemural process, (
|
||
<emphasis id="B9036429FFEAD712FE50F8D22CB1DB54" bold="true" box="[468,500,1843,1867]" pageId="24" pageNumber="25">pts</emphasis>
|
||
)—pretarsus, (
|
||
<emphasis id="B9036429FFEAD712FD0FF8D22FEDDB54" bold="true" box="[651,680,1843,1867]" pageId="24" pageNumber="25">T1</emphasis>
|
||
)—tergite 1, (
|
||
<emphasis id="B9036429FFEAD712FCBDF8D22E0ADB54" bold="true" box="[825,847,1843,1867]" pageId="24" pageNumber="25">t2</emphasis>
|
||
)—tarsus 2, (
|
||
<emphasis id="B9036429FFEAD712FC5FF8D22EB7DB54" bold="true" box="[987,1010,1843,1867]" pageId="24" pageNumber="25">tp</emphasis>
|
||
)—forcipular tooth-plate, (
|
||
<emphasis id="B9036429FFEAD712FA83F8D22862DB54" bold="true" box="[1287,1319,1843,1867]" pageId="24" pageNumber="25">ust</emphasis>
|
||
)—ultimate sternite, (
|
||
<emphasis id="B9036429FFEAD712FF73F8B62C4BDB70" bold="true" box="[247,270,1879,1903]" pageId="24" pageNumber="25">ut</emphasis>
|
||
)—ultimate tergite, (
|
||
<emphasis id="B9036429FFEAD712FE5AF8B62CBCDB70" bold="true" box="[478,505,1879,1903]" pageId="24" pageNumber="25">vp</emphasis>
|
||
)—short ventral projection of pretarsus of maxilla 2.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC8B83BFFEAD713FF43F87C2F4CDD02" blockId="24.[151,1436,1949,2012]" lastBlockId="25.[151,1437,151,285]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
|
||
<bibRefCitation id="EFE6C5CAFFEAD712FF43F87C2F7CDBA8" author="Edgecombe, G. D. & Koch, M." box="[199,569,1949,1975]" pageId="24" pageNumber="25" pagination="872 - 901" refId="ref47403" refString="Edgecombe, G. D. & Koch, M. (2008) Phylogeny of scolopendromorph centipedes (Chilopoda): morphological analysis featuring characters from the peristomatic area. Cladistics, 24, 872 - 901. https: // doi. org / 10.1111 / j. 1096 - 0031.2008.00220. x" type="journal article" year="2008">Edgecombe & Koch (2008: 894)</bibRefCitation>
|
||
described the pretarsus of maxillae
|
||
<quantity id="4C8F15DEFFEAD712FC55F87C2EB9DBA8" box="[977,1020,1949,1975]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="24" pageNumber="25" unit="in" value="2.0">2 in</quantity>
|
||
both
|
||
<taxonomicName id="4C77C3B8FFEAD712FBB9F87F2D95DBC4" authority="(C. L. Koch" baseAuthorityName="C. L. Koch" baseAuthorityYear="1847" class="Chilopoda" family="Cryptopidae" genus="Theatops" kingdom="Animalia" order="Scolopendromorpha" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="erythrocephalus">
|
||
<emphasis id="B9036429FFEAD712FBB9F87F284BDBA8" box="[1085,1294,1949,1975]" italics="true" pageId="24" pageNumber="25">T. erythrocephalus</emphasis>
|
||
(C.L. Koch, 1847
|
||
</taxonomicName>
|
||
) and
|
||
<taxonomicName id="4C77C3B8FFEAD712FE95F8232CCFDBC4" baseAuthorityName="Say" baseAuthorityYear="1821" box="[273,394,1986,2011]" class="Chilopoda" family="Cryptopidae" genus="Theatops" kingdom="Animalia" order="Scolopendromorpha" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="posticus">
|
||
<emphasis id="B9036429FFEAD712FE95F8232CCFDBC4" box="[273,394,1986,2011]" italics="true" pageId="24" pageNumber="25">T. posticus</emphasis>
|
||
</taxonomicName>
|
||
as “two curved processes, one above the other”. The most recent account on
|
||
<taxonomicName id="4C77C3B8FFEAD712FB7AF8202820DBC4" authorityName="Newport" authorityYear="1844" box="[1278,1381,1985,2011]" class="Chilopoda" family="Cryptopidae" genus="Theatops" kingdom="Animalia" order="Scolopendromorpha" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFEAD712FB7AF8202820DBC4" box="[1278,1381,1985,2011]" italics="true" pageId="24" pageNumber="25">Theatops</emphasis>
|
||
</taxonomicName>
|
||
—
|
||
<bibRefCitation id="EFE6C5CAFFEAD713FA05F8202C67DCAD" author="Di, Z. & Cao, Z. & Wu, Y. & Yin, S. & Edgecombe, G. D. & Li, W." lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" pagination="51 - 63" refId="ref47108" refString="Di, Z., Cao, Z., Wu, Y., Yin, S., Edgecombe, G. D. & Li, W. (2010) Discovery of the centipede family Plutoniumidae (Chilopoda) in Asia: a new species of Theatops from China, and the taxonomic value of spiracle distributions in Scolopendromorpha. Zootaxa, 2667 (1), 51 - 63. https: // doi. org / 10.11646 / zootaxa. 2667.1.4" type="journal article" year="2010">
|
||
Di
|
||
<emphasis id="B9036429FFEBD713FF13FF762D94DCAE" box="[151,209,151,177]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
|
||
(2010)
|
||
</bibRefCitation>
|
||
—contains no information on structure of maxillae 2. A re-studied adult (Rc 6489) of
|
||
<taxonomicName id="4C77C3B8FFEBD713FB7DFF762C64DCCA" authority="Wood, 1862" authorityName="Wood" authorityYear="1862" class="Chilopoda" family="Cryptopidae" genus="Theatops" kingdom="Animalia" order="Scolopendromorpha" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="spinicaudus">
|
||
<emphasis id="B9036429FFEBD713FB7DFF7628D9DCAE" box="[1273,1436,151,177]" italics="true" pageId="25" pageNumber="26">T. spinicaudus</emphasis>
|
||
Wood, 1862
|
||
</taxonomicName>
|
||
demonstrates not “two curved processes” but a pretarsus and a much shorter ventral projection (see also Diagnosis of
|
||
<taxonomicName id="4C77C3B8FFEBD713FEE4FF3E2F46DCE6" authorityName="Bollman" authorityYear="1893" box="[352,515,223,249]" class="Chilopoda" family="Plutoniumidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="25" pageNumber="26" phylum="Arthropoda" rank="family">Plutoniumidae</taxonomicName>
|
||
above) whereas in two adults (Rc 6488, 6493) of
|
||
<taxonomicName id="4C77C3B8FFEBD713FBB4FF3E29E2DCE7" baseAuthorityName="Say" baseAuthorityYear="1821" box="[1072,1191,223,248]" class="Chilopoda" family="Cryptopidae" genus="Theatops" kingdom="Animalia" order="Scolopendromorpha" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="posticus">
|
||
<emphasis id="B9036429FFEBD713FBB4FF3E29E2DCE7" box="[1072,1191,223,248]" italics="true" pageId="25" pageNumber="26">T. posticus</emphasis>
|
||
</taxonomicName>
|
||
the ventral projection is nearly as long as the pretarsus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |