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<mods:title id="E65A14E59171C41DD035BE11497142F6">An overview of the extant genera and subgenera of the order Scolopendromorpha (Chilopoda): a new identification key and updated diagnoses</mods:title>
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<treatment id="03DE092DFFE3D719FF13FC85287ADD7A" ID-DOI="http://doi.org/10.5281/zenodo.4455401" ID-GBIF-Taxon="177020718" ID-Zenodo-Dep="4455401" LSID="urn:lsid:plazi:treatment:03DE092DFFE3D719FF13FC85287ADD7A" httpUri="http://treatment.plazi.org/id/03DE092DFFE3D719FF13FC85287ADD7A" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18">
<subSubSection id="C36DEBB0FFE3D71BFF13FC852FC8DF61" box="[151,653,868,895]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="8BC8B83BFFE3D71BFF13FC852FC8DF61" blockId="17.[151,653,868,895]" box="[151,653,868,895]" pageId="17" pageNumber="18">
<heading id="D0800F57FFE3D71BFF13FC852FC8DF61" bold="true" box="[151,653,868,895]" fontSize="11" level="2" pageId="17" pageNumber="18" reason="4">
<taxonomicName id="4C77C3B8FFE3D71BFF13FC852FC8DF61" ID-CoL="8CLF" authority="Verhoeff, 1906" authorityName="Verhoeff" authorityYear="1906" box="[151,653,868,895]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subGenus" subGenus="Chromatanops">
<emphasis id="B9036429FFE3D71BFF13FC852FC8DF61" bold="true" box="[151,653,868,895]" pageId="17" pageNumber="18">
<emphasis id="B9036429FFE3D71BFF13FC852C46DF61" bold="true" box="[151,259,868,894]" italics="true" pageId="17" pageNumber="18">Cryptops</emphasis>
(
<emphasis id="B9036429FFE3D71BFE90FC852C82DF60" bold="true" box="[276,455,868,895]" italics="true" pageId="17" pageNumber="18">Chromatanops</emphasis>
) Verhoeff, 1906
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36DEBB0FFE3D71BFF13FC482E65DFDE" box="[151,800,937,961]" pageId="17" pageNumber="18" type="type_taxon">
<paragraph id="8BC8B83BFFE3D71BFF13FC482E65DFDE" blockId="17.[151,800,937,961]" box="[151,800,937,961]" pageId="17" pageNumber="18">
<emphasis id="B9036429FFE3D71BFF13FC482C66DFDE" bold="true" box="[151,291,937,961]" pageId="17" pageNumber="18">
<typeStatus id="54CC0699FFE3D71BFF13FC482D89DFDE" box="[151,204,937,961]" pageId="17" pageNumber="18">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C77C3B8FFE3D71BFEAEFC482F39DFDE" ID-CoL="6BLXM" authority="Pocock, 1893" authorityName="Pocock" authorityYear="1893" box="[298,636,937,961]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bivittatus">
<emphasis id="B9036429FFE3D71BFEAEFC482CAEDFDE" box="[298,491,937,961]" italics="true" pageId="17" pageNumber="18">Cryptops bivittatus</emphasis>
<bibRefCitation id="EFE6C5CAFFE3D71BFE76FC482F39DFDE" author="Pocock, R. I." box="[498,636,937,961]" pageId="17" pageNumber="18" pagination="454 - 473" refId="ref49778" refString="Pocock, R. I. (1893) Contributions to our knowledge of the arthropod fauna of the West Indies. - Part II. Chilopoda. Zoological Journal of the Linnean Society, 24 (156), 454 - 473." type="journal article" year="1893">Pocock, 1893</bibRefCitation>
</taxonomicName>
(by monotypy).
</paragraph>
</subSubSection>
<subSubSection id="C36DEBB0FFE3D71BFF13FC0E2CD2D816" box="[151,407,1007,1033]" pageId="17" pageNumber="18" type="description">
<paragraph id="8BC8B83BFFE3D71BFF13FC0E2CD2D816" blockId="17.[151,1437,1007,2042]" box="[151,407,1007,1033]" pageId="17" pageNumber="18">
<emphasis id="B9036429FFE3D71BFF13FC0E2C3ED816" bold="true" box="[151,379,1007,1033]" pageId="17" pageNumber="18">Number of species:</emphasis>
1.
</paragraph>
</subSubSection>
<subSubSection id="C36DEBB0FFE3D719FF43FBF2287ADD7A" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18" type="discussion">
<paragraph id="8BC8B83BFFE3D71BFF43FBF2285BD869" blockId="17.[151,1437,1007,2042]" pageId="17" pageNumber="18">
<emphasis id="B9036429FFE3D71BFF43FBF22C7ED832" bold="true" box="[199,315,1043,1069]" pageId="17" pageNumber="18">Remarks.</emphasis>
Treated as a subgenus in
<bibRefCitation id="EFE6C5CAFFE3D71BFDDEFBF22E9AD832" author="Edgecombe, G. D. &amp; Bonato, L." box="[602,991,1043,1070]" pageId="17" pageNumber="18" refId="ref47498" refString="Edgecombe, G. D. &amp; Bonato, L. (2011) Chilopoda-taxonomic overview. Order Scolopendromorpha. In: Minelli, A. (Ed.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 1. Brill, Leiden, pp. 392 - 407." type="book" year="2011">Edgecombe &amp; Bonato (2011: 393)</bibRefCitation>
,
<bibRefCitation id="EFE6C5CAFFE3D71BFC6EFBF2298CD831" author="Lewis, J. G. E." box="[1002,1225,1043,1070]" pageId="17" pageNumber="18" pagination="575 - 584" refId="ref49251" refString="Lewis, J. G. E. (2016 a) On the Cryptops subgenus Trichocryptops Verhoeff, 1937, with a discussion of the problems of differentiating Cryptops species (Chilopoda: Scolopendromorpha: Cryptopidae). Zootaxa, 4139 (4), 575 - 584. https: // doi. org / 10.11646 / zootaxa. 4139.4.9" type="journal article" year="2016">Lewis (2016a: 575)</bibRefCitation>
. Both Chagas-Jr
<emphasis id="B9036429FFE3D71BFA0CFBF52DE8D84E" italics="true" pageId="17" pageNumber="18">et al</emphasis>
. (2014: 142) and
<bibRefCitation id="EFE6C5CAFFE3D71BFEF1FBD92F1DD84D" author="Bonato, L. &amp; Chagas-Junior, A. &amp; Edgecombe, G. D. &amp; Lewis, J. G. E. &amp; Minelli, A. &amp; Pereira, L. A. &amp; Shelley, R. M. &amp; Stoev, P. &amp; Zapparoli, M." box="[373,600,1079,1106]" pageId="17" pageNumber="18" refId="ref45701" refString="Bonato, L., Chagas-Junior, A., Edgecombe, G. D., Lewis, J. G. E., Minelli, A., Pereira, L. A., Shelley, R. M., Stoev, P. &amp; Zapparoli, M. (2016) ChiloBase 2.0 - A World Catalogue of Centipedes (Chilopoda). Available from: http: // chilobase. biologia. unipd. it (accessed 2 August 2016)" type="url" year="2016">
Bonato
<emphasis id="B9036429FFE3D71BFE49FBD92F43D84E" box="[461,518,1079,1105]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
(2016)
</bibRefCitation>
regarded
<taxonomicName id="4C77C3B8FFE3D71BFD4CFBD62E14D84E" authorityName="Pocock" authorityYear="1893" box="[712,849,1079,1105]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bivittatus">
<emphasis id="B9036429FFE3D71BFD4CFBD62E14D84E" box="[712,849,1079,1105]" italics="true" pageId="17" pageNumber="18">C. bivittatus</emphasis>
</taxonomicName>
(the only species of
<taxonomicName id="4C77C3B8FFE3D71BFBBEFBD6299AD84E" authorityName="Verhoeff" authorityYear="1906" box="[1082,1247,1079,1105]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subGenus" subGenus="Chromatanops">
<emphasis id="B9036429FFE3D71BFBBEFBD6299AD84E" box="[1082,1247,1079,1105]" italics="true" pageId="17" pageNumber="18">Chromatanops</emphasis>
</taxonomicName>
) as a member of
<taxonomicName id="4C77C3B8FFE3D71BFF13FBBA2DBBD86A" authorityName="Leach" authorityYear="1814" box="[151,254,1115,1141]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFE3D71BFF13FBBA2DBBD86A" box="[151,254,1115,1141]" italics="true" pageId="17" pageNumber="18">Cryptops</emphasis>
</taxonomicName>
, but at the same time
<bibRefCitation id="EFE6C5CAFFE3D71BFE70FBBD2F93D869" author="Bonato, L. &amp; Chagas-Junior, A. &amp; Edgecombe, G. D. &amp; Lewis, J. G. E. &amp; Minelli, A. &amp; Pereira, L. A. &amp; Shelley, R. M. &amp; Stoev, P. &amp; Zapparoli, M." box="[500,726,1115,1142]" pageId="17" pageNumber="18" refId="ref45701" refString="Bonato, L., Chagas-Junior, A., Edgecombe, G. D., Lewis, J. G. E., Minelli, A., Pereira, L. A., Shelley, R. M., Stoev, P. &amp; Zapparoli, M. (2016) ChiloBase 2.0 - A World Catalogue of Centipedes (Chilopoda). Available from: http: // chilobase. biologia. unipd. it (accessed 2 August 2016)" type="url" year="2016">
Bonato
<emphasis id="B9036429FFE3D71BFDC8FBBD2FC0D86A" box="[588,645,1115,1141]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
(2016)
</bibRefCitation>
included
<taxonomicName id="4C77C3B8FFE3D71BFCC0FBBA2EACD86A" authorityName="Verhoeff" authorityYear="1906" box="[836,1001,1115,1141]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subGenus" subGenus="Chromatanops">
<emphasis id="B9036429FFE3D71BFCC0FBBA2EACD86A" box="[836,1001,1115,1141]" italics="true" pageId="17" pageNumber="18">Chromatanops</emphasis>
</taxonomicName>
as a subgenus of
<taxonomicName id="4C77C3B8FFE3D71BFB34FBBA2852D86A" authorityName="Leach" authorityYear="1814" box="[1200,1303,1115,1141]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFE3D71BFB34FBBA2852D86A" box="[1200,1303,1115,1141]" italics="true" pageId="17" pageNumber="18">Cryptops</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC8B83BFFE3D71BFF43FB9E290BDB5A" blockId="17.[151,1437,1007,2042]" pageId="17" pageNumber="18">
<bibRefCitation id="EFE6C5CAFFE3D71BFF43FB9E2CECD885" author="Pocock, R. I." box="[199,425,1151,1178]" pageId="17" pageNumber="18" pagination="454 - 473" refId="ref49778" refString="Pocock, R. I. (1893) Contributions to our knowledge of the arthropod fauna of the West Indies. - Part II. Chilopoda. Zoological Journal of the Linnean Society, 24 (156), 454 - 473." type="journal article" year="1893">Pocock (1893: 462)</bibRefCitation>
described
<taxonomicName id="4C77C3B8FFE3D71BFDA2FB9E2FBBD886" authorityName="Pocock" authorityYear="1893" box="[550,766,1151,1177]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bivittatus">
<emphasis id="B9036429FFE3D71BFDA2FB9E2FBBD886" box="[550,766,1151,1177]" italics="true" pageId="17" pageNumber="18">Cryptops bivittatus</emphasis>
</taxonomicName>
from “
<collectingCountry id="F360F8ABFFE3D71BFCD7FB9E2E90D886" box="[851,981,1151,1177]" name="Saint Vincent and the Grenadines" pageId="17" pageNumber="18">St. Vincent</collectingCountry>
” by the only[?] specimen with “Length about 15.5 mm”. The original description is rather short and unillustrated. No information is available on clypeal setose plates, either the pretarsus or dorsal brush of maxillae 2, or length of the tarsungula. According to the original description (p. 462) the anterior margin of the forcipular coxosternite is “straight and furnished with 6 bristles” and sternites are “furnished with the ordinary cross-shaped sulci [=sutures]”. In 1903 Kraepelin (who listed two “Exemplare aus
<collectingCountry id="F360F8ABFFE3D71BFE87FAD22C38D952" box="[259,381,1331,1357]" name="Costa Rica" pageId="17" pageNumber="18">Costa Rica</collectingCountry>
und Columbien” at hand) confirmed Pococks description in virtually all details; in particular he wrote (p. 50) that sternites are with “starken Längs- and schwachen Querfurchen”. In 1906 Verhoeff established the subgenus
<taxonomicName id="4C77C3B8FFE3D71BFEB5FA9A2C93D98A" authorityName="Verhoeff" authorityYear="1906" box="[305,470,1403,1429]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subGenus" subGenus="Chromatanops">
<emphasis id="B9036429FFE3D71BFEB5FA9A2C93D98A" box="[305,470,1403,1429]" italics="true" pageId="17" pageNumber="18">Chromatanops</emphasis>
</taxonomicName>
(for
<taxonomicName id="4C77C3B8FFE3D71BFD94FA9A2FDFD98A" authorityName="Pocock" authorityYear="1893" box="[528,666,1403,1429]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bivittatus">
<emphasis id="B9036429FFE3D71BFD94FA9A2FDFD98A" box="[528,666,1403,1429]" italics="true" pageId="17" pageNumber="18">C. bivittatus</emphasis>
</taxonomicName>
only) herewith he surprisingly noted an absence [!] of sternal transverse thickening between coxae of legs (“Hauptsternite ohne innere Verdickungsleisten”; pp 289290), in contrast to Pococks and Kraepelins data on presence of the sternal transverse suture (see above). He also noted an absence of sternal trigonal sutures (“Keine Sternitdreiecke”) and not definitely bordered endosternites (“Endosternite nicht sharf abgegrenzt”) (p. 290). In his short (and unillustrated) diagnosis Verhoeff gave no important morphological details that may be used for separation of his “new subgenus”. However he copied Pococks and Kraepelins data on presence of tergal accumulations of dark pigment (“Körper mit schwartzen, in Längsstreifen angeordneten Pigmentmassen”; p. 390), a character he apparently regarded as diagnostic for his new taxon (this idea is confirmed by the name “
<taxonomicName id="4C77C3B8FFE3D71BFEB2F97A2C9EDAAA" authorityName="Verhoeff" authorityYear="1906" box="[310,475,1691,1717]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subGenus" subGenus="Chromatanops">
<emphasis id="B9036429FFE3D71BFEB2F97A2C9EDAAA" box="[310,475,1691,1717]" italics="true" pageId="17" pageNumber="18">Chromatanops</emphasis>
</taxonomicName>
”). Other data on this mysterious
<taxonomicName id="4C77C3B8FFE3D71BFCD0F97A2EFEDAAA" authorityName="Leach" authorityYear="1814" box="[852,955,1691,1717]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFE3D71BFCD0F97A2EFEDAAA" box="[852,955,1691,1717]" italics="true" pageId="17" pageNumber="18">Cryptops</emphasis>
</taxonomicName>
are scant, contained in
<bibRefCitation id="EFE6C5CAFFE3D71BFB43F97A2DF5DAC6" author="Chamberlin, R. V." pageId="17" pageNumber="18" pagination="1 - 17" refId="ref46670" refString="Chamberlin, R. V. (1922) Centipeds of Central America. Proceedings of the United States National Museum, 60 (7), 1 - 17." type="journal article" year="1922">Chamberlin (1922: 3)</bibRefCitation>
,
<bibRefCitation id="EFE6C5CAFFE3D71BFF39F95E2CDADAC6" author="Attems, C." box="[189,415,1727,1753]" pageId="17" pageNumber="18" refId="ref45522" refString="Attems, C. (1930) Myriapoda 2. Scolopendromorpha. Das Tierreich. Vol. 54. Walter de Gruyter et Co. Publ., Berlin, 308 pp." type="book" year="1930">Attems (1930: 243)</bibRefCitation>
,
<bibRefCitation id="EFE6C5CAFFE3D71BFE28F95E2FD3DAC6" author="Bucherl, W." box="[428,662,1727,1753]" pageId="17" pageNumber="18" pagination="251 - 372" refId="ref45866" refString="Bucherl, W. (1942) Catalogo dos quilopodos da zona neotropica. Memorias do Instituto Butantan, 15, 251 - 372." type="journal article" year="1942">Bücherl (1942: 324)</bibRefCitation>
,
<bibRefCitation id="EFE6C5CAFFE3D71BFD20F9212E30DAC6" author="Kraus, O." box="[676,885,1727,1754]" pageId="17" pageNumber="18" pagination="359 - 404" refId="ref48477" refString="Kraus, O. (1957) Myriapoden aus Peru VI. Chilopoden. Senckenbergiana Biologica, 38, 359 - 404." type="journal article" year="1957">Kraus (1957: 387)</bibRefCitation>
and Chagas-Jr
<emphasis id="B9036429FFE3D71BFBADF9212919DAC6" box="[1065,1116,1727,1753]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
. (2014: 142). Of these only Attems gave a short morphological account, albeit reproducing data of three previous authors. Original data on the geographic distribution of “
<taxonomicName id="4C77C3B8FFE3D71BFE57F8E62E29DB3E" authorityName="Pocock" authorityYear="1893" box="[467,876,1799,1826]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bivittatus" subGenus="Chromatanops">
<emphasis id="B9036429FFE3D71BFE57F8E62F7FDB3E" box="[467,570,1799,1825]" italics="true" pageId="17" pageNumber="18">Cryptops</emphasis>
(
<emphasis id="B9036429FFE3D71BFDC8F8E62FB4DB3E" box="[588,753,1799,1825]" italics="true" pageId="17" pageNumber="18">Chromatanops</emphasis>
)
<emphasis id="B9036429FFE3D71BFC80F8E62E29DB3E" box="[772,876,1799,1825]" italics="true" pageId="17" pageNumber="18">bivittatus</emphasis>
</taxonomicName>
” (6 new localities from
<collectingCountry id="F360F8ABFFE3D71BFB03F8E929F9DB3D" box="[1159,1212,1800,1826]" name="Peru" pageId="17" pageNumber="18">Peru</collectingCountry>
) were presented by
<bibRefCitation id="EFE6C5CAFFE3D71BFF13F8CD2C6BDB59" author="Kraus, O." box="[151,302,1835,1862]" pageId="17" pageNumber="18" pagination="359 - 404" refId="ref48477" refString="Kraus, O. (1957) Myriapoden aus Peru VI. Chilopoden. Senckenbergiana Biologica, 38, 359 - 404." type="journal article" year="1957">Kraus (1957)</bibRefCitation>
, whereas the other three authors just copied the previous faunistic data.
</paragraph>
<paragraph id="8BC8B83BFFE3D719FF43F8AE28D5DCE6" blockId="17.[151,1437,1007,2042]" lastBlockId="19.[151,1437,150,357]" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18">
As for Verhoeffs (1906) “diagnostic” dark coloration of his “
<taxonomicName id="4C77C3B8FFE3D71BFC15F8AE2973DB76" authorityName="Verhoeff" authorityYear="1906" box="[913,1078,1871,1897]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subGenus" subGenus="Chromatanops">
<emphasis id="B9036429FFE3D71BFC15F8AE2973DB76" box="[913,1078,1871,1897]" italics="true" pageId="17" pageNumber="18">Chromatanops</emphasis>
</taxonomicName>
”, we can state that it is of minor taxonomic value in
<taxonomicName id="4C77C3B8FFE3D71BFE2BF8922F53DB92" authorityName="Leach" authorityYear="1814" box="[431,534,1907,1933]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFE3D71BFE2BF8922F53DB92" box="[431,534,1907,1933]" italics="true" pageId="17" pageNumber="18">Cryptops</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFE6C5CAFFE3D71BFDACF8922E5EDB92" author="Schileyko, A." box="[552,795,1907,1933]" pageId="17" pageNumber="18" pagination="71 - 95" refId="ref50073" refString="Schileyko, A. (2007) The scolopendromorph centipedes (Chilopoda) of Vietnam, with contributions to the faunas of Cambodia and Laos. Part 3. Arthropoda Selecta, 16 (2), 71 - 95." type="journal article" year="2007">Schileyko (2007: 88)</bibRefCitation>
wrote about pigmentation in Vietnamese material (
<specimenCount id="9D7173B2FFE3D71BFAF5F8922C48DBAD" pageId="17" pageNumber="18" type="generic">150 specimens</specimenCount>
) of
<taxonomicName id="4C77C3B8FFE3D71BFEBEF8762F00DBAE" authority="Pocock, 1891" authorityName="Pocock" authorityYear="1891" box="[314,581,1943,1969]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="doriae">
<emphasis id="B9036429FFE3D71BFEBEF8762CE6DBAE" box="[314,419,1943,1969]" italics="true" pageId="17" pageNumber="18">C. doriae</emphasis>
Pocock, 1891
</taxonomicName>
: “… about 40 % of specimens showing inner accumulations of dark pigment … This dark pattern is much better visible dorsally, sometimes also laterally and, much more rarely, ventrally”. Schileykos data were confirmed by
<bibRefCitation id="EFE6C5CAFFE3D71BFDC1F83E2F9ADBE6" author="Lewis, J. G. E." box="[581,735,2015,2041]" pageId="17" pageNumber="18" pagination="505 - 518" refId="ref48813" refString="Lewis, J. G. E. (2009) A review of some characters used in the taxonomy of Cryptops (subgenus Cryptops) (Chilopoda: Scolopendromorpha: Cryptopidae). Soil Organisms, 81, 505 - 518." type="journal article" year="2009">Lewis (2009)</bibRefCitation>
who analysed in detail a variability of some morphological characters, traditionally used in taxonomy of
<taxonomicName id="4C77C3B8FFE1D719FD18FF762EC1DCAE" authority="(Cryptops)" authorityName="Leach" authorityYear="1814" baseAuthorityName="Cryptops" box="[668,900,151,177]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="20" phylum="Arthropoda" rank="subGenus" subGenus="Cryptops">
<emphasis id="B9036429FFE1D719FD18FF762E46DCAE" box="[668,771,151,177]" italics="true" pageId="19" pageNumber="20">Cryptops</emphasis>
(
<emphasis id="B9036429FFE1D719FC90FF762E3EDCAE" box="[788,891,151,177]" italics="true" pageId="19" pageNumber="20">Cryptops</emphasis>
)
</taxonomicName>
(p. 506): “To summarise, some species always exhibit the [dark] pigmentation; some may or may not exhibit it and others … are not pigmented”. Thus, this character may vary in
<taxonomicName id="4C77C3B8FFE1D719FEE5FF3E2C8DDCE6" authorityName="Leach" authorityYear="1814" box="[353,456,223,249]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFE1D719FEE5FF3E2C8DDCE6" box="[353,456,223,249]" italics="true" pageId="19" pageNumber="20">Cryptops</emphasis>
</taxonomicName>
even intraspecifically, so it cannot be used for separation of any subtaxa in this genus.
</paragraph>
<caption id="DF08E8B3FFE0D718FF13F9302920DBFA" ID-DOI="http://doi.org/10.5281/zenodo.4402155" ID-Zenodo-Dep="4402155" httpUri="https://zenodo.org/record/4402155/files/figure.png" pageId="18" pageNumber="19" startId="18.[151,264,1745,1769]" targetBox="[264,1325,199,1702]" targetPageId="18">
<paragraph id="8BC8B83BFFE0D718FF13F9302920DBFA" blockId="18.[151,1437,1745,2021]" pageId="18" pageNumber="19">
<emphasis id="B9036429FFE0D718FF13F9302C13DAF6" bold="true" box="[151,342,1745,1769]" pageId="18" pageNumber="19">FIGURES 2025.</emphasis>
<taxonomicName id="4C77C3B8FFE0D718FEDBF9332FEBDAF6" authority="(Pocock, 1896)" baseAuthorityName="Pocock" baseAuthorityYear="1896" box="[351,686,1745,1769]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="stolli">
<emphasis id="B9036429FFE0D718FEDBF9332F40DAF6" box="[351,517,1745,1769]" italics="true" pageId="18" pageNumber="19">Newportia stolli</emphasis>
(Pocock, 1896)
</taxonomicName>
; Ad. Rc 7235 (?)
<emphasis id="B9036429FFE0D718FCEFF9302ED4DAF6" bold="true" box="[875,913,1745,1769]" pageId="18" pageNumber="19">20a</emphasis>
ultimate LBS + ultimate legs dorsally;
<taxonomicName id="4C77C3B8FFE0D718FAB5F9332CFCDB12" authority="Chamberlin, 1922" authorityName="Chamberlin" authorityYear="1922" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="divergens">
<emphasis id="B9036429FFE0D718FAB5F9332DBCDB12" italics="true" pageId="18" pageNumber="19">Newportia divergens</emphasis>
<bibRefCitation id="EFE6C5CAFFE0D718FE85F9142CFCDB12" author="Chamberlin, R. V." box="[257,441,1781,1805]" pageId="18" pageNumber="19" pagination="1 - 17" refId="ref46670" refString="Chamberlin, R. V. (1922) Centipeds of Central America. Proceedings of the United States National Museum, 60 (7), 1 - 17." type="journal article" year="1922">Chamberlin, 1922</bibRefCitation>
</taxonomicName>
<emphasis id="B9036429FFE0D718FE45F9142CACDB12" bold="true" box="[449,489,1781,1805]" pageId="18" pageNumber="19">20b</emphasis>
right ultimate leg (schematic);
<taxonomicName id="4C77C3B8FFE0D718FCABF917284BDB12" authority="Lewis, 1989" authorityName="Lewis" authorityYear="1989" box="[815,1294,1781,1805]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="19" phylum="Arthropoda" rank="subSpecies" species="longitarsis" subSpecies="virginensis">
<emphasis id="B9036429FFE0D718FCABF91729C1DB12" box="[815,1156,1781,1805]" italics="true" pageId="18" pageNumber="19">Newportia longitarsis virginensis</emphasis>
Lewis, 1989
</taxonomicName>
; Ad. Rc 7470
<emphasis id="B9036429FFE0D718FF13F8F82DF5DB2E" bold="true" box="[151,176,1817,1841]" pageId="18" pageNumber="19">21</emphasis>
head and forcipular segment ventrally;
<taxonomicName id="4C77C3B8FFE0D718FDC9F8FB296FDB2E" authority="(Kraepelin, 1903)" baseAuthorityName="Kraepelin" baseAuthorityYear="1903" box="[589,1066,1817,1841]" class="Chilopoda" family="Scolopocryptopidae" genus="Newportia" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="collaris" subGenus="Tidops">
<emphasis id="B9036429FFE0D718FDC9F8FB2FFDDB2E" box="[589,696,1818,1841]" italics="true" pageId="18" pageNumber="19">Newportia</emphasis>
(
<emphasis id="B9036429FFE0D718FD4CF8F82E49DB2E" box="[712,780,1817,1841]" italics="true" pageId="18" pageNumber="19">Tidops</emphasis>
)
<emphasis id="B9036429FFE0D718FC98F8F82E2CDB2E" box="[796,873,1817,1841]" italics="true" pageId="18" pageNumber="19">collaris</emphasis>
(
<bibRefCitation id="EFE6C5CAFFE0D718FCFCF8F82964DB2E" author="Kraepelin, K." box="[888,1057,1817,1841]" pageId="18" pageNumber="19" pagination="1 - 276" refId="ref48450" refString="Kraepelin, K. (1903) Revision der Scolopendriden. Mitteilungen aus dem Naturhistorischen Museum Hamburg, 20 (2), 1 - 276." type="journal article" year="1903">Kraepelin, 1903</bibRefCitation>
)
</taxonomicName>
; MCZ 32933
<emphasis id="B9036429FFE0D718FB3FF8F82991DB2E" bold="true" box="[1211,1236,1817,1841]" pageId="18" pageNumber="19">22</emphasis>
forcipular segment ventrally
<emphasis id="B9036429FFE0D718FF7EF8DC2C56DB4A" bold="true" box="[250,275,1853,1877]" pageId="18" pageNumber="19">23</emphasis>
anterior half of head ventrally
<emphasis id="B9036429FFE0D718FDE5F8DC2F3FDB4A" bold="true" box="[609,634,1853,1877]" pageId="18" pageNumber="19">24</emphasis>
apical part of maxilla 2 laterally
<emphasis id="B9036429FFE0D718FC5BF8DC2EBDDB4A" bold="true" box="[991,1016,1853,1877]" pageId="18" pageNumber="19">25</emphasis>
right pleuron of mid body LBS laterally; (
<emphasis id="B9036429FFE0D718FF47F8802DA0DB66" bold="true" box="[195,229,1889,1913]" pageId="18" pageNumber="19">am</emphasis>
)—anterior margin of forcipular coxosternite, (
<emphasis id="B9036429FFE0D718FD4DF8802F9EDB66" bold="true" box="[713,731,1889,1913]" pageId="18" pageNumber="19">cl</emphasis>
)—clypeus, (
<emphasis id="B9036429FFE0D718FCE6F8802EC5DB66" bold="true" box="[866,896,1889,1913]" pageId="18" pageNumber="19">cst</emphasis>
)— forcipular coxosternite, (
<emphasis id="B9036429FFE0D718FB2DF88029F4DB66" bold="true" box="[1193,1201,1889,1913]" pageId="18" pageNumber="19">f</emphasis>
)—femur, (
<emphasis id="B9036429FFE0D718FAA1F8802869DB66" bold="true" box="[1317,1324,1889,1913]" pageId="18" pageNumber="19">l</emphasis>
)—labrum, (
<emphasis id="B9036429FFE0D718FF24F8642DFEDB82" bold="true" box="[160,187,1925,1949]" pageId="18" pageNumber="19">p2</emphasis>
)—pectinate pretarsus of maxilla 2, (
<emphasis id="B9036429FFE0D718FDBDF8642F11DB82" bold="true" box="[569,596,1925,1949]" pageId="18" pageNumber="19">pa</emphasis>
)—blunt projection of anterior margin of forcipular coxosternite, (
<emphasis id="B9036429FFE0D718FA86F864285CDB82" bold="true" box="[1282,1305,1925,1949]" pageId="18" pageNumber="19">pf</emphasis>
)—prefemur, (
<emphasis id="B9036429FFE0D718FF24F8482DEFDBDE" bold="true" box="[160,170,1961,1985]" pageId="18" pageNumber="19">s</emphasis>
)—open spiracle with atrium, (
<emphasis id="B9036429FFE0D718FE65F8482CB2DBDE" bold="true" box="[481,503,1961,1985]" pageId="18" pageNumber="19">t1</emphasis>
)—tarsus 1, (
<emphasis id="B9036429FFE0D718FDFFF8482FD2DBDE" bold="true" box="[635,663,1961,1985]" pageId="18" pageNumber="19">t2i</emphasis>
)—tarsus 2 with irregularly divided secondary articles, (
<emphasis id="B9036429FFE0D718FB49F84829A1DBDE" bold="true" box="[1229,1252,1961,1985]" pageId="18" pageNumber="19">th</emphasis>
)—trichoid sensillae, (
<emphasis id="B9036429FFE0D718FF4EF82C2D9CDBFA" bold="true" box="[202,217,1997,2021]" pageId="18" pageNumber="19">ti</emphasis>
)—tibia, (
<emphasis id="B9036429FFE0D718FEB9F82C2C14DBFA" bold="true" box="[317,337,1997,2021]" pageId="18" pageNumber="19">tr</emphasis>
)—tarsungula, (
<emphasis id="B9036429FFE0D718FE75F82C2F56DBFA" bold="true" box="[497,531,1997,2021]" pageId="18" pageNumber="19">trp</emphasis>
)—forcipular trochantero-prefemur, (
<emphasis id="B9036429FFE0D718FC08F82C2EE6DBFA" bold="true" box="[908,931,1997,2021]" pageId="18" pageNumber="19">ut</emphasis>
)—ultimate tergite.
</paragraph>
</caption>
<paragraph id="8BC8B83BFFE1D719FF43FEE2287ADD7A" blockId="19.[151,1437,150,357]" pageId="19" pageNumber="20">
Summing up, as there are no reliable diagnostic characters to confirm
<taxonomicName id="4C77C3B8FFE1D719FC4DFEE2292BDD02" authorityName="Verhoeff" authorityYear="1906" box="[969,1134,259,285]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="18" phylum="Arthropoda" rank="subGenus" subGenus="Chromatanops">
<emphasis id="B9036429FFE1D719FC4DFEE2292BDD02" box="[969,1134,259,285]" italics="true" pageId="19" pageNumber="20">Chromatanops</emphasis>
</taxonomicName>
as a subgenus, so
<taxonomicName id="4C77C3B8FFE1D719FAB2FEE22CBADD5E" authority="Verhoeff, 1906" authorityName="Verhoeff" authorityYear="1906" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="20" phylum="Arthropoda" rank="subGenus" subGenus="Chromatanops">
<emphasis id="B9036429FFE1D719FAB2FEE228D8DD02" box="[1334,1437,259,285]" italics="true" pageId="19" pageNumber="20">Cryptops</emphasis>
(
<emphasis id="B9036429FFE1D719FF25FEC62C03DD5E" box="[161,326,295,321]" italics="true" pageId="19" pageNumber="20">Chromatanops</emphasis>
) Verhoeff, 1906
</taxonomicName>
is considered a junior synonym of
<taxonomicName id="4C77C3B8FFE1D719FC16FEC629C9DD5E" authority="Leach, 1814" authorityName="Leach" authorityYear="1814" box="[914,1164,295,321]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFE1D719FC16FEC62EBCDD5E" box="[914,1017,295,321]" italics="true" pageId="19" pageNumber="20">Cryptops</emphasis>
Leach, 1814
</taxonomicName>
<emphasis id="B9036429FFE1D719FB10FEC629B8DD5E" bold="true" box="[1172,1277,295,321]" pageId="19" pageNumber="20">syn. nov.</emphasis>
and
<taxonomicName id="4C77C3B8FFE1D719FAB2FEC62F1BDD7A" authority="Pocock, 1893" authorityName="Pocock" authorityYear="1893" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="bivittatus" subGenus="Chromatanops">
<emphasis id="B9036429FFE1D719FAB2FEC628D8DD5E" box="[1334,1437,295,321]" italics="true" pageId="19" pageNumber="20">Cryptops</emphasis>
(
<emphasis id="B9036429FFE1D719FF25FEAA2C03DD7A" box="[161,326,331,357]" italics="true" pageId="19" pageNumber="20">Chromatanops</emphasis>
)
<emphasis id="B9036429FFE1D719FED2FEAA2CFBDD7A" box="[342,446,331,357]" italics="true" pageId="19" pageNumber="20">bivittatus</emphasis>
<bibRefCitation id="EFE6C5CAFFE1D719FE41FEAA2F1BDD7A" author="Pocock, R. I." box="[453,606,331,357]" pageId="19" pageNumber="20" pagination="454 - 473" refId="ref49778" refString="Pocock, R. I. (1893) Contributions to our knowledge of the arthropod fauna of the West Indies. - Part II. Chilopoda. Zoological Journal of the Linnean Society, 24 (156), 454 - 473." type="journal article" year="1893">Pocock, 1893</bibRefCitation>
</taxonomicName>
should be
<taxonomicName id="4C77C3B8FFE1D719FD5CFEAA298BDD7A" authority="Pocock, 1893" authorityName="Pocock" authorityYear="1893" box="[728,1230,331,357]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="bivittatus" status="stat. nov." subGenus="Cryptops">
<emphasis id="B9036429FFE1D719FD5CFEAA2E7ADD7A" box="[728,831,331,357]" italics="true" pageId="19" pageNumber="20">Cryptops</emphasis>
(
<emphasis id="B9036429FFE1D719FCCBFEAA2EF3DD7A" box="[847,950,331,357]" italics="true" pageId="19" pageNumber="20">Cryptops</emphasis>
)
<emphasis id="B9036429FFE1D719FC42FEAA296BDD7A" box="[966,1070,331,357]" italics="true" pageId="19" pageNumber="20">bivittatus</emphasis>
<bibRefCitation id="EFE6C5CAFFE1D719FBB1FEAA298BDD7A" author="Pocock, R. I." box="[1077,1230,331,357]" pageId="19" pageNumber="20" pagination="454 - 473" refId="ref49778" refString="Pocock, R. I. (1893) Contributions to our knowledge of the arthropod fauna of the West Indies. - Part II. Chilopoda. Zoological Journal of the Linnean Society, 24 (156), 454 - 473." type="journal article" year="1893">Pocock, 1893</bibRefCitation>
</taxonomicName>
<emphasis id="B9036429FFE1D719FB51FEAA287ADD7A" bold="true" box="[1237,1343,331,357]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="A230D952FFE1D719FB51FEAA287ADD7A" box="[1237,1343,331,357]" pageId="19" pageNumber="20">stat. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>