300 lines
53 KiB
XML
300 lines
53 KiB
XML
<document id="9038BE23D54B99EF516430C3DC099E8F" ID-CLB-Dataset="8703" ID-DOI="10.11646/zootaxa.4825.1.1" ID-GBIF-Dataset="5ab5f5c8-481e-4d1a-8643-21e72c367278" ID-ISSN="1175-5326" ID-Zenodo-Dep="4402145" ID-ZooBank="F230F199-1C94-4E2E-9CE4-5F56212C015F" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1609341789210" checkinUser="plazi" docAuthor="Schileyko, Arkady A., Vahtera, Varpu & Edgecombe, Gregory D." docDate="2020" docId="03DE092DFFE2D71AFF13FF762C93DB3B" docLanguage="en" docName="zootaxa.4825.1.1.pdf" docOrigin="Zootaxa 4825 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Cryptopidae Kohlrausch 1881" docType="treatment" docVersion="7" lastPageNumber="17" masterDocId="FFE77155FFF2D70AFF84FFE12D45DC1F" masterDocTitle="An overview of the extant genera and subgenera of the order Scolopendromorpha (Chilopoda): a new identification key and updated diagnoses" masterLastPageNumber="64" masterPageNumber="1" pageNumber="17" updateTime="1698852004925" updateUser="ExternalLinkService">
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<mods:titleInfo id="F7D500BA8773DB2EE8048A0C4B91FBE6">
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<mods:title id="8F9DEFA8D8858E6FB6EC27CD514C0F27">An overview of the extant genera and subgenera of the order Scolopendromorpha (Chilopoda): a new identification key and updated diagnoses</mods:title>
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<mods:name id="333CF5884F576B203CFB91CCB423D66E" type="personal">
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<mods:namePart id="20FC8D24A8392CFB360DE51D802226F6">Schileyko, Arkady A.</mods:namePart>
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<mods:nameIdentifier id="F587DEDB0BEAC16C799607131063A3CB" type="email">schileyko1965@gmail.com</mods:nameIdentifier>
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<mods:namePart id="D1832309254495293F16E912E00B9728">Vahtera, Varpu</mods:namePart>
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<mods:nameIdentifier id="85849CCF927979EBCF9816AFD8E3BEFC" type="email">varpu.vahtera@gmail.com</mods:nameIdentifier>
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<mods:namePart id="6CB10D4A1A6E87284BDBC81BFDA547F7">Edgecombe, Gregory D.</mods:namePart>
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<mods:nameIdentifier id="F49C741DF78FD6A80263076F0D2DA124" type="email">schileyko1965@gmail.com</mods:nameIdentifier>
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<mods:title id="CD3D08D1CB0FB0DBB5351616F1F8206D">Zootaxa</mods:title>
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<mods:date id="B459320F8BAC076492354C785B77B8DC">2020</mods:date>
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<mods:number id="12CF9E5EC12B872E8D15D829C8B2C399">2020-08-10</mods:number>
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<mods:number id="5097A6B290FC08874802731B30506B72">4825</mods:number>
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<treatment id="03DE092DFFE2D71AFF13FF762C93DB3B" ID-DOI="http://doi.org/10.5281/zenodo.4455397" ID-GBIF-Taxon="177020714" ID-Zenodo-Dep="4455397" LSID="urn:lsid:plazi:treatment:03DE092DFFE2D71AFF13FF762C93DB3B" httpUri="http://treatment.plazi.org/id/03DE092DFFE2D71AFF13FF762C93DB3B" lastPageNumber="17" pageId="16" pageNumber="17">
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<subSubSection id="C36DEBB0FFE2D71AFF13FF762F3DDCAE" box="[151,632,151,177]" pageId="16" pageNumber="17" type="nomenclature">
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<paragraph id="8BC8B83BFFE2D71AFF13FF762F3DDCAE" blockId="16.[151,632,151,177]" box="[151,632,151,177]" pageId="16" pageNumber="17">
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<heading id="D0800F57FFE2D71AFF13FF762F3DDCAE" bold="true" box="[151,632,151,177]" fontSize="11" level="2" pageId="16" pageNumber="17" reason="3">
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<emphasis id="B9036429FFE2D71AFF13FF762F3DDCAE" bold="true" box="[151,632,151,177]" pageId="16" pageNumber="17">
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Family
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<taxonomicName id="4C77C3B8FFE2D71AFF73FF762F3DDCAE" ID-CoL="8R3" ID-ENA="7543" authority="Kohlrausch, 1881" authorityName="Kohlrausch" authorityYear="1881" box="[247,632,151,177]" class="Chilopoda" family="Cryptopidae" kingdom="Animalia" order="Scolopendromorpha" pageId="16" pageNumber="17" phylum="Arthropoda" rank="family">Cryptopidae Kohlrausch, 1881</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36DEBB0FFE2D71AFF13FF3D2FC3DCEB" box="[151,646,219,244]" pageId="16" pageNumber="17" type="synonymic_list">
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<paragraph id="8BC8B83BFFE2D71AFF13FF3D2FC3DCEB" blockId="16.[151,646,219,244]" box="[151,646,219,244]" pageId="16" pageNumber="17">
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Synonyms.
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<taxonomicName id="4C77C3B8FFE2D71AFE8BFF3D2FC3DCEB" authority="in Machado (1953: 83)" authorityName="in Machado" authorityPageNumber="83" authorityYear="1953" box="[271,646,219,244]" class="Chilopoda" family="Cryptopsidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="16" pageNumber="17" phylum="Arthropoda" rank="family">
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Cryptopsidae in
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<bibRefCitation id="EFE6C5CAFFE2D71AFE32FF3D2FC3DCEB" author="Machado, A." box="[438,646,219,244]" pageId="16" pageNumber="17" pagination="77 - 92" refId="ref49486" refString="Machado, A. (1953) Alguns Miriapodes de Espanha (colheitas de J. Mateu). Archivos Instituto de Aclimaticion, 1, 77 - 92." type="journal article" year="1953">Machado (1953: 83)</bibRefCitation>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36DEBB0FFE2D71AFF13FEC029A6DFDB" pageId="16" pageNumber="17" type="diagnosis">
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<paragraph id="8BC8B83BFFE2D71AFF13FEC029A6DFDB" blockId="16.[151,1437,289,1828]" pageId="16" pageNumber="17">
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<emphasis id="B9036429FFE2D71AFF13FEC02C57DD24" bold="true" box="[151,274,289,315]" pageId="16" pageNumber="17">Diagnosis.</emphasis>
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Eyes absent. Labrum with a single median tooth (only in a few species of
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<taxonomicName id="4C77C3B8FFE2D71AFBC9FEC32876DD23" authority="(Cryptops)" authorityName="Leach" authorityYear="1814" baseAuthorityName="Cryptops" box="[1101,1331,290,316]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subGenus" subGenus="Cryptops">
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<emphasis id="B9036429FFE2D71AFBC9FEC329F1DD23" box="[1101,1204,290,316]" italics="true" pageId="16" pageNumber="17">Cryptops</emphasis>
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(
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<emphasis id="B9036429FFE2D71AFB47FEC3286FDD23" box="[1219,1322,290,316]" italics="true" pageId="16" pageNumber="17">Cryptops</emphasis>
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)
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</taxonomicName>
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with two additional lateral teeth;
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<figureCitation id="134CA4BEFFE2D71AFE21FEA72CBCDD7F" box="[421,505,326,352]" captionStart="FIGURES 26–32" captionStartId="20.[151,264,1597,1621]" captionTargetBox="[372,1216,196,1556]" captionTargetId="figure-19@20.[355,1232,181,1573]" captionTargetPageId="20" captionText="FIGURES 26–32. Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. paratype CIRAD P8A 26 head, forcipular segment + LBS 1–3 ventrally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 27 forcipular segment ventrally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. holotype MNHN 459 28 right ultimate leg medially; Cryptops (C.) caucasius Verhoeff, 1934; Ad. Rc 8003 29 cephalic plate + anterior margin of forcipular coxosternite ventrally; Cryptops australis (Newport, 1845); AM KS 30 apical part of maxilla 2 laterally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 31 apical part of of maxilla 2 laterally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. holotype MNHN 459 32 LBS 3 ventrally; (am)—anterior margin of forcipular coxosternite, (cl)—clypeus, (cs)—clypeal setose plate, (cst)—forcipular coxosternite, (db)—apical setae of dorsal brush, (es)—enlarged (spiniform) setae, (est)—endosternite, (l)—labrum, (lp)—lateral projection of endosternite, (ls)—long setae, (lsc)—sternal median longitudinal sulcus, (pa)—short rounded projection of anterior margin of forcipular coxosternite, (pf)—prefemur, (sl)—sternal median longitudinal suture, (sr)—sternal transverse ridge, (st)—saw teeth, (sts)—sternal transverse suture, (tgs)—sternal trigonal sutures, (ti)—tibia, (tr)—tarsungula, (vp)—ventral projection of pretarsus of maxilla 2." figureDoi="http://doi.org/10.5281/zenodo.4402157" httpUri="https://zenodo.org/record/4402157/files/figure.png" pageId="16" pageNumber="17">Fig. 29</figureCitation>
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). Slender pretarsus of maxillae 2 not pectinate, more or less curved (
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<figureCitation id="134CA4BEFFE2D71AFA8EFEA728CBDD7F" box="[1290,1422,326,352]" captionStart="FIGURES 26–32" captionStartId="20.[151,264,1597,1621]" captionTargetBox="[372,1216,196,1556]" captionTargetId="figure-19@20.[355,1232,181,1573]" captionTargetPageId="20" captionText="FIGURES 26–32. Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. paratype CIRAD P8A 26 head, forcipular segment + LBS 1–3 ventrally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 27 forcipular segment ventrally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. holotype MNHN 459 28 right ultimate leg medially; Cryptops (C.) caucasius Verhoeff, 1934; Ad. Rc 8003 29 cephalic plate + anterior margin of forcipular coxosternite ventrally; Cryptops australis (Newport, 1845); AM KS 30 apical part of maxilla 2 laterally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 31 apical part of of maxilla 2 laterally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. holotype MNHN 459 32 LBS 3 ventrally; (am)—anterior margin of forcipular coxosternite, (cl)—clypeus, (cs)—clypeal setose plate, (cst)—forcipular coxosternite, (db)—apical setae of dorsal brush, (es)—enlarged (spiniform) setae, (est)—endosternite, (l)—labrum, (lp)—lateral projection of endosternite, (ls)—long setae, (lsc)—sternal median longitudinal sulcus, (pa)—short rounded projection of anterior margin of forcipular coxosternite, (pf)—prefemur, (sl)—sternal median longitudinal suture, (sr)—sternal transverse ridge, (st)—saw teeth, (sts)—sternal transverse suture, (tgs)—sternal trigonal sutures, (ti)—tibia, (tr)—tarsungula, (vp)—ventral projection of pretarsus of maxilla 2." figureDoi="http://doi.org/10.5281/zenodo.4402157" httpUri="https://zenodo.org/record/4402157/files/figure.png" pageId="16" pageNumber="17">Figs 30, 31</figureCitation>
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), pointed (figs 3C, 6C in
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<bibRefCitation id="EFE6C5CAFFE2D71AFE2CFE8A2FE8DD9B" author="De Azara, L. N. & Ferreira, R. L." box="[424,685,362,390]" pageId="16" pageNumber="17" pagination="432 - 444" refId="ref45581" refString="De Azara, L. N. & Ferreira, R. L. (2013) The first troglobitic Cryptops (Trigonocryptops) (Chilopoda: Scolopendromorpha) from South America and the description of a non-troglobitic species from Brazil. Zootaxa, 3709 (5), 432 - 444. https: // doi. org / 10.11646 / zootaxa. 3709.5.2" type="journal article" year="2013">Ázara & Ferreira 2013</bibRefCitation>
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) or lobe-shaped (fig.
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<bibRefCitation id="EFE6C5CAFFE2D71AFC63FE8B29C8DD9B" author="Verhoeff, K. W." box="[999,1165,362,388]" pageId="16" pageNumber="17" pagination="1 - 112" refId="ref51555" refString="Verhoeff, K. W. (1934) Beitrage zur Systematik und Geographie der Chilopoden. Zoologische Jahrbucher, 66 (1 / 2), 1 - 112." type="journal article" year="1934">Verhoeff 1934</bibRefCitation>
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). Maxillary pretarsus in most species is accompanied by a ventral projection (“flange” sensu
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<bibRefCitation id="EFE6C5CAFFE2D71AFC25FE6F2998DDB7" author="Edgecombe, G. D. & Bonato, L." box="[929,1245,398,424]" pageId="16" pageNumber="17" refId="ref47498" refString="Edgecombe, G. D. & Bonato, L. (2011) Chilopoda-taxonomic overview. Order Scolopendromorpha. In: Minelli, A. (Ed.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 1. Brill, Leiden, pp. 392 - 407." type="book" year="2011">Edgecombe & Bonato 2011</bibRefCitation>
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,
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<figureCitation id="134CA4BEFFE2D71AFB6DFE6F282DDDB7" box="[1257,1384,398,424]" captionStart="FIGURES 26–32" captionStartId="20.[151,264,1597,1621]" captionTargetBox="[372,1216,196,1556]" captionTargetId="figure-19@20.[355,1232,181,1573]" captionTargetPageId="20" captionText="FIGURES 26–32. Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. paratype CIRAD P8A 26 head, forcipular segment + LBS 1–3 ventrally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 27 forcipular segment ventrally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. holotype MNHN 459 28 right ultimate leg medially; Cryptops (C.) caucasius Verhoeff, 1934; Ad. Rc 8003 29 cephalic plate + anterior margin of forcipular coxosternite ventrally; Cryptops australis (Newport, 1845); AM KS 30 apical part of maxilla 2 laterally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 31 apical part of of maxilla 2 laterally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. holotype MNHN 459 32 LBS 3 ventrally; (am)—anterior margin of forcipular coxosternite, (cl)—clypeus, (cs)—clypeal setose plate, (cst)—forcipular coxosternite, (db)—apical setae of dorsal brush, (es)—enlarged (spiniform) setae, (est)—endosternite, (l)—labrum, (lp)—lateral projection of endosternite, (ls)—long setae, (lsc)—sternal median longitudinal sulcus, (pa)—short rounded projection of anterior margin of forcipular coxosternite, (pf)—prefemur, (sl)—sternal median longitudinal suture, (sr)—sternal transverse ridge, (st)—saw teeth, (sts)—sternal transverse suture, (tgs)—sternal trigonal sutures, (ti)—tibia, (tr)—tarsungula, (vp)—ventral projection of pretarsus of maxilla 2." figureDoi="http://doi.org/10.5281/zenodo.4402157" httpUri="https://zenodo.org/record/4402157/files/figure.png" pageId="16" pageNumber="17">Figs 30, 31</figureCitation>
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) but never by accessory spine(s). Anterior margin of forcipular coxosternite in most species bilobed, with some long enlarged marginal setae (
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<figureCitation id="134CA4BEFFE2D71AFE34FE372F41DDEE" box="[432,516,470,497]" captionStart="FIGURES 26–32" captionStartId="20.[151,264,1597,1621]" captionTargetBox="[372,1216,196,1556]" captionTargetId="figure-19@20.[355,1232,181,1573]" captionTargetPageId="20" captionText="FIGURES 26–32. Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. paratype CIRAD P8A 26 head, forcipular segment + LBS 1–3 ventrally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 27 forcipular segment ventrally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. holotype MNHN 459 28 right ultimate leg medially; Cryptops (C.) caucasius Verhoeff, 1934; Ad. Rc 8003 29 cephalic plate + anterior margin of forcipular coxosternite ventrally; Cryptops australis (Newport, 1845); AM KS 30 apical part of maxilla 2 laterally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 31 apical part of of maxilla 2 laterally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. holotype MNHN 459 32 LBS 3 ventrally; (am)—anterior margin of forcipular coxosternite, (cl)—clypeus, (cs)—clypeal setose plate, (cst)—forcipular coxosternite, (db)—apical setae of dorsal brush, (es)—enlarged (spiniform) setae, (est)—endosternite, (l)—labrum, (lp)—lateral projection of endosternite, (ls)—long setae, (lsc)—sternal median longitudinal sulcus, (pa)—short rounded projection of anterior margin of forcipular coxosternite, (pf)—prefemur, (sl)—sternal median longitudinal suture, (sr)—sternal transverse ridge, (st)—saw teeth, (sts)—sternal transverse suture, (tgs)—sternal trigonal sutures, (ti)—tibia, (tr)—tarsungula, (vp)—ventral projection of pretarsus of maxilla 2." figureDoi="http://doi.org/10.5281/zenodo.4402157" httpUri="https://zenodo.org/record/4402157/files/figure.png" pageId="16" pageNumber="17">Fig. 26</figureCitation>
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); rarely is this margin additionally chitinised or with short rounded projections (in
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<taxonomicName id="4C77C3B8FFE2D71AFF13FE1B2CF5DE0B" authority="" authorityName="Pocock" authorityYear="1891" box="[151,432,506,532]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subGenus" subGenus="Paracryptops">
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<emphasis id="B9036429FFE2D71AFF13FE1B2DBBDE0B" box="[151,254,506,532]" italics="true" pageId="16" pageNumber="17">Cryptops</emphasis>
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(
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<emphasis id="B9036429FFE2D71AFE89FE1A2CE0DE0B" box="[269,421,507,532]" italics="true" pageId="16" pageNumber="17">Paracryptops</emphasis>
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)
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</taxonomicName>
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,
|
||
<figureCitation id="134CA4BEFFE2D71AFE38FE1B2F48DE0B" box="[444,525,506,532]" captionStart="FIGURES 26–32" captionStartId="20.[151,264,1597,1621]" captionTargetBox="[372,1216,196,1556]" captionTargetId="figure-19@20.[355,1232,181,1573]" captionTargetPageId="20" captionText="FIGURES 26–32. Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. paratype CIRAD P8A 26 head, forcipular segment + LBS 1–3 ventrally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 27 forcipular segment ventrally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. holotype MNHN 459 28 right ultimate leg medially; Cryptops (C.) caucasius Verhoeff, 1934; Ad. Rc 8003 29 cephalic plate + anterior margin of forcipular coxosternite ventrally; Cryptops australis (Newport, 1845); AM KS 30 apical part of maxilla 2 laterally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 31 apical part of of maxilla 2 laterally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. holotype MNHN 459 32 LBS 3 ventrally; (am)—anterior margin of forcipular coxosternite, (cl)—clypeus, (cs)—clypeal setose plate, (cst)—forcipular coxosternite, (db)—apical setae of dorsal brush, (es)—enlarged (spiniform) setae, (est)—endosternite, (l)—labrum, (lp)—lateral projection of endosternite, (ls)—long setae, (lsc)—sternal median longitudinal sulcus, (pa)—short rounded projection of anterior margin of forcipular coxosternite, (pf)—prefemur, (sl)—sternal median longitudinal suture, (sr)—sternal transverse ridge, (st)—saw teeth, (sts)—sternal transverse suture, (tgs)—sternal trigonal sutures, (ti)—tibia, (tr)—tarsungula, (vp)—ventral projection of pretarsus of maxilla 2." figureDoi="http://doi.org/10.5281/zenodo.4402157" httpUri="https://zenodo.org/record/4402157/files/figure.png" pageId="16" pageNumber="17">Fig. 27</figureCitation>
|
||
), but never with tooth-plates. Forcipular trochantero-prefemur never with process. Sternites never with paramedian sutures, but usually with “cruciform” sutures (
|
||
<figureCitation id="134CA4BEFFE2D71AFB9BFDFF2931DE27" box="[1055,1140,542,568]" captionStart="FIGURES 26–32" captionStartId="20.[151,264,1597,1621]" captionTargetBox="[372,1216,196,1556]" captionTargetId="figure-19@20.[355,1232,181,1573]" captionTargetPageId="20" captionText="FIGURES 26–32. Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. paratype CIRAD P8A 26 head, forcipular segment + LBS 1–3 ventrally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 27 forcipular segment ventrally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. holotype MNHN 459 28 right ultimate leg medially; Cryptops (C.) caucasius Verhoeff, 1934; Ad. Rc 8003 29 cephalic plate + anterior margin of forcipular coxosternite ventrally; Cryptops australis (Newport, 1845); AM KS 30 apical part of maxilla 2 laterally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 31 apical part of of maxilla 2 laterally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. holotype MNHN 459 32 LBS 3 ventrally; (am)—anterior margin of forcipular coxosternite, (cl)—clypeus, (cs)—clypeal setose plate, (cst)—forcipular coxosternite, (db)—apical setae of dorsal brush, (es)—enlarged (spiniform) setae, (est)—endosternite, (l)—labrum, (lp)—lateral projection of endosternite, (ls)—long setae, (lsc)—sternal median longitudinal sulcus, (pa)—short rounded projection of anterior margin of forcipular coxosternite, (pf)—prefemur, (sl)—sternal median longitudinal suture, (sr)—sternal transverse ridge, (st)—saw teeth, (sts)—sternal transverse suture, (tgs)—sternal trigonal sutures, (ti)—tibia, (tr)—tarsungula, (vp)—ventral projection of pretarsus of maxilla 2." figureDoi="http://doi.org/10.5281/zenodo.4402157" httpUri="https://zenodo.org/record/4402157/files/figure.png" pageId="16" pageNumber="17">Fig. 32</figureCitation>
|
||
), i.e. with well-developed transverse suture (sometimes with skeletal transverse ridge/thickening at its place) and with a single median longitudinal suture (more rarely sulcus) developed to varying degrees. 21 LBS; spiracles on macrosegments except for LBS 7; spiracles open, lacking flaps (fig. 4 A in
|
||
<bibRefCitation id="EFE6C5CAFFE2D71AFD33FD6B2EDADEBA" author="Vahtera, V. & Edgecombe, G. D. & Giribet, G." box="[695,927,650,677]" pageId="16" pageNumber="17" pagination="225 - 248" refId="ref51336" refString="Vahtera, V., Edgecombe, G. D. & Giribet, G. (2012 b) Spiracle structure in scolopendromorph centipedes (Chilopoda: Scolopendromorpha) and its contribution to phylogenetics. Zoomorphology, 131, 225 - 248. https: // doi. org / 10.1007 / s 00435 - 012 - 0157 - 0" type="journal article" year="2012">
|
||
Vahtera
|
||
<emphasis id="B9036429FFE2D71AFC90FD6A2E03DEBB" box="[788,838,650,676]" italics="true" pageId="16" pageNumber="17">et al</emphasis>
|
||
. 2012b
|
||
</bibRefCitation>
|
||
). Ultimate LBS considerably shorter than the penultimate one; coxopleuron without a process. Tarsal articles of locomotory legs (
|
||
<figureCitation id="134CA4BEFFE2D71AFBB3FD4F29C2DED6" box="[1079,1159,686,713]" captionStart="FIGURES 26–32" captionStartId="20.[151,264,1597,1621]" captionTargetBox="[372,1216,196,1556]" captionTargetId="figure-19@20.[355,1232,181,1573]" captionTargetPageId="20" captionText="FIGURES 26–32. Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. paratype CIRAD P8A 26 head, forcipular segment + LBS 1–3 ventrally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 27 forcipular segment ventrally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. holotype MNHN 459 28 right ultimate leg medially; Cryptops (C.) caucasius Verhoeff, 1934; Ad. Rc 8003 29 cephalic plate + anterior margin of forcipular coxosternite ventrally; Cryptops australis (Newport, 1845); AM KS 30 apical part of maxilla 2 laterally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 31 apical part of of maxilla 2 laterally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. holotype MNHN 459 32 LBS 3 ventrally; (am)—anterior margin of forcipular coxosternite, (cl)—clypeus, (cs)—clypeal setose plate, (cst)—forcipular coxosternite, (db)—apical setae of dorsal brush, (es)—enlarged (spiniform) setae, (est)—endosternite, (l)—labrum, (lp)—lateral projection of endosternite, (ls)—long setae, (lsc)—sternal median longitudinal sulcus, (pa)—short rounded projection of anterior margin of forcipular coxosternite, (pf)—prefemur, (sl)—sternal median longitudinal suture, (sr)—sternal transverse ridge, (st)—saw teeth, (sts)—sternal transverse suture, (tgs)—sternal trigonal sutures, (ti)—tibia, (tr)—tarsungula, (vp)—ventral projection of pretarsus of maxilla 2." figureDoi="http://doi.org/10.5281/zenodo.4402157" httpUri="https://zenodo.org/record/4402157/files/figure.png" pageId="16" pageNumber="17">Fig. 26</figureCitation>
|
||
) in most species fused in a solid tarsus (with or without visible traces of an articulation between them); legs lack both tibial and tarsal spurs. Relatively short ultimate legs (
|
||
<figureCitation id="134CA4BEFFE2D71AFE75FD172F07DF0F" box="[497,578,758,784]" captionStart="FIGURES 26–32" captionStartId="20.[151,264,1597,1621]" captionTargetBox="[372,1216,196,1556]" captionTargetId="figure-19@20.[355,1232,181,1573]" captionTargetPageId="20" captionText="FIGURES 26–32. Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. paratype CIRAD P8A 26 head, forcipular segment + LBS 1–3 ventrally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 27 forcipular segment ventrally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. holotype MNHN 459 28 right ultimate leg medially; Cryptops (C.) caucasius Verhoeff, 1934; Ad. Rc 8003 29 cephalic plate + anterior margin of forcipular coxosternite ventrally; Cryptops australis (Newport, 1845); AM KS 30 apical part of maxilla 2 laterally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 31 apical part of of maxilla 2 laterally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. holotype MNHN 459 32 LBS 3 ventrally; (am)—anterior margin of forcipular coxosternite, (cl)—clypeus, (cs)—clypeal setose plate, (cst)—forcipular coxosternite, (db)—apical setae of dorsal brush, (es)—enlarged (spiniform) setae, (est)—endosternite, (l)—labrum, (lp)—lateral projection of endosternite, (ls)—long setae, (lsc)—sternal median longitudinal sulcus, (pa)—short rounded projection of anterior margin of forcipular coxosternite, (pf)—prefemur, (sl)—sternal median longitudinal suture, (sr)—sternal transverse ridge, (st)—saw teeth, (sts)—sternal transverse suture, (tgs)—sternal trigonal sutures, (ti)—tibia, (tr)—tarsungula, (vp)—ventral projection of pretarsus of maxilla 2." figureDoi="http://doi.org/10.5281/zenodo.4402157" httpUri="https://zenodo.org/record/4402157/files/figure.png" pageId="16" pageNumber="17">Fig. 28</figureCitation>
|
||
) of “pocket knife” shape (sensu
|
||
<bibRefCitation id="EFE6C5CAFFE2D71AFC36FD172926DF0F" author="Schileyko, A." box="[946,1123,758,784]" pageId="16" pageNumber="17" pagination="519 - 530" refId="ref50114" refString="Schileyko, A. (2009) Ectonocryptoides sandrops - a new scolopendromorph centipede from Belize. Soil Organisms, 81 (3), 519 - 530." type="journal article" year="2009">Schileyko 2009</bibRefCitation>
|
||
) forming a kind of clasping apparatus, their femur (not always), tibia and tarsus
|
||
<quantity id="4C8F15DEFFE2D71AFD66FCFB2E48DF2B" box="[738,781,794,820]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="16" pageNumber="17" unit="in" value="1.0">1 in</quantity>
|
||
overwhelming majority of species with characteristic saw teeth, pretarsus not enlarged. Prefemur of the ultimate legs without processes (spinous ones or saw teeth), sometimes with short enlarged setae (
|
||
<figureCitation id="134CA4BEFFE2D71AFE7BFC832F16DF63" box="[511,595,866,892]" captionStart="FIGURES 26–32" captionStartId="20.[151,264,1597,1621]" captionTargetBox="[372,1216,196,1556]" captionTargetId="figure-19@20.[355,1232,181,1573]" captionTargetPageId="20" captionText="FIGURES 26–32. Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. paratype CIRAD P8A 26 head, forcipular segment + LBS 1–3 ventrally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 27 forcipular segment ventrally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. holotype MNHN 459 28 right ultimate leg medially; Cryptops (C.) caucasius Verhoeff, 1934; Ad. Rc 8003 29 cephalic plate + anterior margin of forcipular coxosternite ventrally; Cryptops australis (Newport, 1845); AM KS 30 apical part of maxilla 2 laterally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 31 apical part of of maxilla 2 laterally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. holotype MNHN 459 32 LBS 3 ventrally; (am)—anterior margin of forcipular coxosternite, (cl)—clypeus, (cs)—clypeal setose plate, (cst)—forcipular coxosternite, (db)—apical setae of dorsal brush, (es)—enlarged (spiniform) setae, (est)—endosternite, (l)—labrum, (lp)—lateral projection of endosternite, (ls)—long setae, (lsc)—sternal median longitudinal sulcus, (pa)—short rounded projection of anterior margin of forcipular coxosternite, (pf)—prefemur, (sl)—sternal median longitudinal suture, (sr)—sternal transverse ridge, (st)—saw teeth, (sts)—sternal transverse suture, (tgs)—sternal trigonal sutures, (ti)—tibia, (tr)—tarsungula, (vp)—ventral projection of pretarsus of maxilla 2." figureDoi="http://doi.org/10.5281/zenodo.4402157" httpUri="https://zenodo.org/record/4402157/files/figure.png" pageId="16" pageNumber="17">Fig. 28</figureCitation>
|
||
).
|
||
<bibRefCitation id="EFE6C5CAFFE2D71AFDE2FC832EA2DF63" author="Edgecombe, G. D. & Bonato, L." box="[614,999,866,892]" pageId="16" pageNumber="17" refId="ref47498" refString="Edgecombe, G. D. & Bonato, L. (2011) Chilopoda-taxonomic overview. Order Scolopendromorpha. In: Minelli, A. (Ed.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 1. Brill, Leiden, pp. 392 - 407." type="book" year="2011">Edgecombe & Bonato (2011: 393)</bibRefCitation>
|
||
also wrote: “Median cluster of sensilla coeloconica on clypeal part of epipharynx rhomboid, with lids covering the distal edge of the sensilla… Gizzard with stiff anteriorly-directed projections; projections without a distinct kink near their midlength”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36DEBB0FFE2D71AFF43FC2F2FE8DFF7" box="[199,685,973,1000]" pageId="16" pageNumber="17" type="discussion">
|
||
<paragraph id="8BC8B83BFFE2D71AFF43FC2F2FE8DFF7" blockId="16.[151,1437,289,1828]" box="[199,685,973,1000]" pageId="16" pageNumber="17">
|
||
<emphasis id="B9036429FFE2D71AFF43FC2F2CF1DFF7" bold="true" box="[199,436,973,1000]" pageId="16" pageNumber="17">Number of subtaxa.</emphasis>
|
||
1 genus, 4 subgenera.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36DEBB0FFE2D71AFF43FC102F71D813" box="[199,564,1009,1036]" pageId="16" pageNumber="17" type="description">
|
||
<paragraph id="8BC8B83BFFE2D71AFF43FC102F71D813" blockId="16.[151,1437,289,1828]" box="[199,564,1009,1036]" pageId="16" pageNumber="17">
|
||
<emphasis id="B9036429FFE2D71AFF43FC102CF2D814" bold="true" box="[199,439,1009,1035]" pageId="16" pageNumber="17">Sexual dimorphism.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36DEBB0FFE2D71AFF43FBF72E32D82F" box="[199,887,1046,1072]" pageId="16" pageNumber="17" type="distribution">
|
||
<paragraph id="8BC8B83BFFE2D71AFF43FBF72E32D82F" blockId="16.[151,1437,289,1828]" box="[199,887,1046,1072]" pageId="16" pageNumber="17">
|
||
<emphasis id="B9036429FFE2D71AFF43FBF72C5FD82F" bold="true" box="[199,282,1046,1072]" pageId="16" pageNumber="17">Range.</emphasis>
|
||
All tropical, subtropical and warm temperate regions.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36DEBB0FFE2D71AFF43FBDB2C93DB3B" pageId="16" pageNumber="17" type="discussion">
|
||
<paragraph id="8BC8B83BFFE2D71AFF43FBDB29E7D867" blockId="16.[151,1437,289,1828]" pageId="16" pageNumber="17">
|
||
<emphasis id="B9036429FFE2D71AFF43FBDB2C7ED84B" bold="true" box="[199,315,1082,1108]" pageId="16" pageNumber="17">Remarks.</emphasis>
|
||
Treated as a family in
|
||
<bibRefCitation id="EFE6C5CAFFE2D71AFDB3FBDB2EFDD84B" author="Edgecombe, G. D. & Bonato, L." box="[567,952,1082,1108]" pageId="16" pageNumber="17" refId="ref47498" refString="Edgecombe, G. D. & Bonato, L. (2011) Chilopoda-taxonomic overview. Order Scolopendromorpha. In: Minelli, A. (Ed.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 1. Brill, Leiden, pp. 392 - 407." type="book" year="2011">Edgecombe & Bonato (2011: 393)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE6C5CAFFE2D71AFC46FBDB298AD84A" author="Vahtera, V. & Edgecombe, G. D. & Giribet, G." box="[962,1231,1082,1109]" pageId="16" pageNumber="17" pagination="4 - 20" refId="ref51271" refString="Vahtera, V., Edgecombe, G. D. & Giribet, G. (2012 a) Evolution of blindness in scolopendromorph centipedes (Chilopoda: Scolopendromorpha): insight from an expanded sampling of molecular data. Cladistics, 28, 4 - 20. https: // doi. org / 10.1111 / j. 1096 - 0031.2011.00361. x" type="journal article" year="2012">
|
||
Vahtera
|
||
<emphasis id="B9036429FFE2D71AFB9AFBDA2910D84B" box="[1054,1109,1082,1108]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
|
||
(2012a: 6)
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE6C5CAFFE2D71AFB5FFBDB2C58D867" author="Edgecombe, G. D. & Vahtera, V. & Stock, S. R. & Kallonen, A. & Xiao, X. & Rack, A. & Giribet, G." pageId="16" pageNumber="17" pagination="768 - 786" refId="ref47554" refString="Edgecombe, G. D., Vahtera, V., Stock, S. R., Kallonen, A., Xiao, X., Rack, A. & Giribet, G. (2012) A scolopocryptopid centipede (Chilopoda: Scolopendromorpha) from Mexican amber: synchrotron microtomography and phylogenetic placement using a combined morphological and molecular data set. Zoological Journal of the Linnean Society, 166, 768 - 786. https: // doi. org / 10.1111 / j. 1096 - 3642.2012.00860. x" type="journal article" year="2012">
|
||
Edgecombe
|
||
<emphasis id="B9036429FFE2D71AFAE1FBDA28D8D84B" box="[1381,1437,1082,1108]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
|
||
(2012: 770)
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE6C5CAFFE2D71AFEADFBBF2C99D867" author="Lewis, J. G. E." box="[297,476,1118,1144]" pageId="16" pageNumber="17" pagination="1 - 34" refId="ref49041" refString="Lewis, J. G. E. (2013) A review of the species of the genus Cryptops Leach, 1815 from the Old World and Australasian region related to Cryptops (Cryptops) doriae Pocock, 1891 (Chilopoda: Scolopendromorpha: Cryptopidae). Zootaxa, 3683 (1), 1 - 34. https: // doi. org / 10.11646 / zootaxa. 3683.1.1" type="journal article" year="2013">Lewis (2013: 1)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE6C5CAFFE2D71AFE6DFBBF2F82D866" author="Lewis, J. G. E." box="[489,711,1118,1145]" pageId="16" pageNumber="17" pagination="575 - 584" refId="ref49251" refString="Lewis, J. G. E. (2016 a) On the Cryptops subgenus Trichocryptops Verhoeff, 1937, with a discussion of the problems of differentiating Cryptops species (Chilopoda: Scolopendromorpha: Cryptopidae). Zootaxa, 4139 (4), 575 - 584. https: // doi. org / 10.11646 / zootaxa. 4139.4.9" type="journal article" year="2016">Lewis (2016a: 575)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE6C5CAFFE2D71AFD57FBBF29D8D867" author="Stojanovic, D. Z. & Mitic, B. M. & Makarov, S. E." box="[723,1181,1118,1144]" pageId="16" pageNumber="17" pagination="21 - 25" refId="ref51141" refString="Stojanovic, D. Z., Mitic, B. M. & Makarov, S. E. (2019) Bifurcation of an ultimate leg in Cryptops parisi Brolemann, 1920 (Chilopoda: Scolopendromorpha: Cryptopidae). Arthropoda Selecta, 28 (1), 21 - 25. https: // doi. org / 10.15298 / arthsel. 28.1.03" type="journal article" year="2019">Stojanović, Mitić & Makarov (2019: 21)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC8B83BFFE2D71AFF43FB632F68D94E" blockId="16.[151,1437,289,1828]" pageId="16" pageNumber="17">
|
||
<bibRefCitation id="EFE6C5CAFFE2D71AFF43FB632F19D883" author="Edgecombe, G. D. & Bonato, L." box="[199,604,1154,1180]" pageId="16" pageNumber="17" refId="ref47498" refString="Edgecombe, G. D. & Bonato, L. (2011) Chilopoda-taxonomic overview. Order Scolopendromorpha. In: Minelli, A. (Ed.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 1. Brill, Leiden, pp. 392 - 407." type="book" year="2011">Edgecombe and Bonato (2011: 393)</bibRefCitation>
|
||
divided this family into two closely related genera—
|
||
<taxonomicName id="4C77C3B8FFE2D71AFB35FB63285DD883" authorityName="Leach" authorityYear="1814" box="[1201,1304,1154,1180]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFE2D71AFB35FB63285DD883" box="[1201,1304,1154,1180]" italics="true" pageId="16" pageNumber="17">Cryptops</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C77C3B8FFE2D71AFACAFB622DAED8DF" class="Chilopoda" family="Cryptopidae" genus="Paracryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFE2D71AFACAFB622DAED8DF" italics="true" pageId="16" pageNumber="17">Paracryptops</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<bibRefCitation id="EFE6C5CAFFE2D71AFEA8FB472F14D8DF" author="Vahtera, V. & Edgecombe, G. D. & Giribet, G." box="[300,593,1190,1216]" pageId="16" pageNumber="17" pagination="4 - 20" refId="ref51271" refString="Vahtera, V., Edgecombe, G. D. & Giribet, G. (2012 a) Evolution of blindness in scolopendromorph centipedes (Chilopoda: Scolopendromorpha): insight from an expanded sampling of molecular data. Cladistics, 28, 4 - 20. https: // doi. org / 10.1111 / j. 1096 - 0031.2011.00361. x" type="journal article" year="2012">
|
||
Vahtera
|
||
<emphasis id="B9036429FFE2D71AFE0EFB462C80D8DF" box="[394,453,1190,1216]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
|
||
(2012a: 13)
|
||
</bibRefCitation>
|
||
suggested (without formalizing; see below) the latter to be a synonym of the former; so at the moment this family is monotypic. According to both
|
||
<bibRefCitation id="EFE6C5CAFFE2D71AFC60FB2B282ED8FB" author="Edgecombe, G. D. & Bonato, L." box="[996,1387,1226,1252]" pageId="16" pageNumber="17" refId="ref47498" refString="Edgecombe, G. D. & Bonato, L. (2011) Chilopoda-taxonomic overview. Order Scolopendromorpha. In: Minelli, A. (Ed.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 1. Brill, Leiden, pp. 392 - 407." type="book" year="2011">Edgecombe & Bonato (2011: 393)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFE6C5CAFFE2D71AFF13FB0F2C34D916" author="Lewis, J. G. E." box="[151,369,1262,1289]" pageId="16" pageNumber="17" pagination="575 - 584" refId="ref49251" refString="Lewis, J. G. E. (2016 a) On the Cryptops subgenus Trichocryptops Verhoeff, 1937, with a discussion of the problems of differentiating Cryptops species (Chilopoda: Scolopendromorpha: Cryptopidae). Zootaxa, 4139 (4), 575 - 584. https: // doi. org / 10.11646 / zootaxa. 4139.4.9" type="journal article" year="2016">Lewis (2016a: 575)</bibRefCitation>
|
||
the genus
|
||
<taxonomicName id="4C77C3B8FFE2D71AFE63FB0F2F0BD917" authorityName="Leach" authorityYear="1814" box="[487,590,1262,1288]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFE2D71AFE63FB0F2F0BD917" box="[487,590,1262,1288]" italics="true" pageId="16" pageNumber="17">Cryptops</emphasis>
|
||
</taxonomicName>
|
||
comprises four subgenera:
|
||
<taxonomicName id="4C77C3B8FFE2D71AFC05FB0F292CD917" authority="(Cryptops)" authorityName="Leach" authorityYear="1814" baseAuthorityName="Cryptops" box="[897,1129,1262,1288]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subGenus" subGenus="Cryptops">
|
||
<emphasis id="B9036429FFE2D71AFC05FB0F2EADD917" box="[897,1000,1262,1288]" italics="true" pageId="16" pageNumber="17">Cryptops</emphasis>
|
||
(
|
||
<emphasis id="B9036429FFE2D71AFC73FB0F291BD917" box="[1015,1118,1262,1288]" italics="true" pageId="16" pageNumber="17">Cryptops</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B9036429FFE2D71AFBF0FB0E2868D917" box="[1140,1325,1263,1288]" italics="true" pageId="16" pageNumber="17">Trigonocryptops</emphasis>
|
||
Verhoeff, 1906,
|
||
<taxonomicName id="4C77C3B8FFE2D71AFF5AFAF32F70D933" authority="Verhoeff, 1934" authorityName="Verhoeff" authorityYear="1934" box="[222,565,1298,1324]" class="Chilopoda" family="Cryptopidae" genus="Haplocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="16" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFE2D71AFF5AFAF32CC1D933" box="[222,388,1298,1324]" italics="true" pageId="16" pageNumber="17">Haplocryptops</emphasis>
|
||
<bibRefCitation id="EFE6C5CAFFE2D71AFE08FAF32F70D933" author="Verhoeff, K. W." box="[396,565,1298,1324]" pageId="16" pageNumber="17" pagination="1 - 112" refId="ref51555" refString="Verhoeff, K. W. (1934) Beitrage zur Systematik und Geographie der Chilopoden. Zoologische Jahrbucher, 66 (1 / 2), 1 - 112." type="journal article" year="1934">Verhoeff, 1934</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C77C3B8FFE2D71AFDE9FAF32E83D933" authority="Verhoeff, 1906" authorityName="Verhoeff" authorityYear="1906" box="[621,966,1298,1324]" class="Chilopoda" family="Cryptopidae" genus="Chromatanops" kingdom="Animalia" order="Scolopendromorpha" pageId="16" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFE2D71AFDE9FAF32E57D933" box="[621,786,1298,1324]" italics="true" pageId="16" pageNumber="17">Chromatanops</emphasis>
|
||
Verhoeff, 1906
|
||
</taxonomicName>
|
||
, but we regard the latter to be a synonym of
|
||
<taxonomicName id="4C77C3B8FFE2D71AFF32FAD72CD9D94F" authority="(Cryptops)" authorityName="Leach" authorityYear="1814" baseAuthorityName="Cryptops" box="[182,412,1334,1360]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subGenus" subGenus="Cryptops">
|
||
<emphasis id="B9036429FFE2D71AFF32FAD72C58D94F" box="[182,285,1334,1360]" italics="true" pageId="16" pageNumber="17">Cryptops</emphasis>
|
||
(
|
||
<emphasis id="B9036429FFE2D71AFEA9FAD72CD1D94F" box="[301,404,1334,1360]" italics="true" pageId="16" pageNumber="17">Cryptops</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
(see below).
|
||
</paragraph>
|
||
<paragraph id="8BC8B83BFFE2D71AFF43FABB2C93DB3B" blockId="16.[151,1437,289,1828]" pageId="16" pageNumber="17">
|
||
There is certain confusion concerning structure of the pretarsus of maxillae
|
||
<quantity id="4C8F15DEFFE2D71AFB90FABB297AD96B" box="[1044,1087,1370,1396]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="16" pageNumber="17" unit="in" value="2.0">2 in</quantity>
|
||
this family (i.e. in genus
|
||
<taxonomicName id="4C77C3B8FFE2D71AFADFFABB2D83D987" authorityName="Leach" authorityYear="1814" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFE2D71AFADFFABB2D83D987" italics="true" pageId="16" pageNumber="17">Cryptops</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<bibRefCitation id="EFE6C5CAFFE2D71AFF58FA9F2CD9D987" author="Attems, C." box="[220,412,1406,1432]" pageId="16" pageNumber="17" refId="ref45522" refString="Attems, C. (1930) Myriapoda 2. Scolopendromorpha. Das Tierreich. Vol. 54. Walter de Gruyter et Co. Publ., Berlin, 308 pp." type="book" year="1930">Attems (1930: 7)</bibRefCitation>
|
||
only wrote that this structure is “schlank” (slim/slender) and gave a drawing (fig. 285) of a slightly[!] curved and apically pointed claw accompanied by a well-developed and apically rounded ventral projection. In 1934 Verhoeff, describing the new subgenus
|
||
<emphasis id="B9036429FFE2D71AFD6CFA272E47D9FF" box="[744,770,1478,1504]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
|
||
(
|
||
<taxonomicName id="4C77C3B8FFE2D71AFC96FA272EFDD9FF" authorityName="Verhoeff" authorityYear="1934" box="[786,952,1478,1504]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="16" pageNumber="22" phylum="Arthropoda" rank="subGenus" subGenus="Haplocryptops">
|
||
<emphasis id="B9036429FFE2D71AFC96FA272EFDD9FF" box="[786,952,1478,1504]" italics="true" pageId="16" pageNumber="17">Haplocryptops</emphasis>
|
||
</taxonomicName>
|
||
), noted two
|
||
<typeStatus id="54CC0699FFE2D71AFBC3FA2729C7D9FF" box="[1095,1154,1478,1504]" pageId="16" pageNumber="17">types</typeStatus>
|
||
of maxillary pretarsus in
|
||
<taxonomicName id="4C77C3B8FFE2D71AFF13FA0B2DBBDA1B" authorityName="Leach" authorityYear="1814" box="[151,254,1514,1540]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFE2D71AFF13FA0B2DBBDA1B" box="[151,254,1514,1540]" italics="true" pageId="16" pageNumber="17">Cryptops</emphasis>
|
||
</taxonomicName>
|
||
(see below). Recent authors (
|
||
<bibRefCitation id="EFE6C5CAFFE2D71AFDCCFA0B2EE2DA1B" author="Edgecombe, G. D. & Koch, M." box="[584,935,1514,1540]" pageId="16" pageNumber="17" pagination="872 - 901" refId="ref47403" refString="Edgecombe, G. D. & Koch, M. (2008) Phylogeny of scolopendromorph centipedes (Chilopoda): morphological analysis featuring characters from the peristomatic area. Cladistics, 24, 872 - 901. https: // doi. org / 10.1111 / j. 1096 - 0031.2008.00220. x" type="journal article" year="2008">Edgecombe & Koch 2008: 883</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE6C5CAFFE2D71AFC37FA0B29DCDA1B" author="Koch, M. & Edgecombe, G. D. & Shelley, R. M." box="[947,1177,1514,1540]" pageId="16" pageNumber="17" pagination="51 - 81" refId="ref48332" refString="Koch, M., Edgecombe, G. D. & Shelley, R. M. (2010) Anatomy of Ectonocryptoides (Scolopocryptopidae: Ectonocryptopinae) and the phylogeny of blind Scolopendromorpha (Chilopoda). International Journal of Myriapodology, 3, 51 - 81. https: // doi. org / 10.1163 / 187525410 X 12578602960344" type="journal article" year="2010">
|
||
Koch et
|
||
<emphasis id="B9036429FFE2D71AFB95FA0B2962DA1B" box="[1041,1063,1514,1540]" italics="true" pageId="16" pageNumber="17">al</emphasis>
|
||
. 2010: 77
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE6C5CAFFE2D71AFB20FA0B2C4EDA37" author="Edgecombe, G. D. & Bonato, L." pageId="16" pageNumber="17" refId="ref47498" refString="Edgecombe, G. D. & Bonato, L. (2011) Chilopoda-taxonomic overview. Order Scolopendromorpha. In: Minelli, A. (Ed.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 1. Brill, Leiden, pp. 392 - 407." type="book" year="2011">Edgecombe & Bonato 2011: 393</bibRefCitation>
|
||
) describe this pretarsus only as a “hook-like and flanged”, confirming these terms by corresponding drawings or SEMs (
|
||
<figureCitation id="134CA4BEFFE2D71AFEF9F9D32F46DA53" box="[381,515,1586,1612]" captionStart="FIGURES 26–32" captionStartId="20.[151,264,1597,1621]" captionTargetBox="[372,1216,196,1556]" captionTargetId="figure-19@20.[355,1232,181,1573]" captionTargetPageId="20" captionText="FIGURES 26–32. Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. paratype CIRAD P8A 26 head, forcipular segment + LBS 1–3 ventrally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 27 forcipular segment ventrally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. holotype MNHN 459 28 right ultimate leg medially; Cryptops (C.) caucasius Verhoeff, 1934; Ad. Rc 8003 29 cephalic plate + anterior margin of forcipular coxosternite ventrally; Cryptops australis (Newport, 1845); AM KS 30 apical part of maxilla 2 laterally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 31 apical part of of maxilla 2 laterally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. holotype MNHN 459 32 LBS 3 ventrally; (am)—anterior margin of forcipular coxosternite, (cl)—clypeus, (cs)—clypeal setose plate, (cst)—forcipular coxosternite, (db)—apical setae of dorsal brush, (es)—enlarged (spiniform) setae, (est)—endosternite, (l)—labrum, (lp)—lateral projection of endosternite, (ls)—long setae, (lsc)—sternal median longitudinal sulcus, (pa)—short rounded projection of anterior margin of forcipular coxosternite, (pf)—prefemur, (sl)—sternal median longitudinal suture, (sr)—sternal transverse ridge, (st)—saw teeth, (sts)—sternal transverse suture, (tgs)—sternal trigonal sutures, (ti)—tibia, (tr)—tarsungula, (vp)—ventral projection of pretarsus of maxilla 2." figureDoi="http://doi.org/10.5281/zenodo.4402157" httpUri="https://zenodo.org/record/4402157/files/figure.png" pageId="16" pageNumber="17">Figs 30, 31</figureCitation>
|
||
). We note, however, that this character is not stable in
|
||
<taxonomicName id="4C77C3B8FFE2D71AFBF8F9D329A6DA53" authorityName="Leach" authorityYear="1814" box="[1148,1251,1586,1612]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFE2D71AFBF8F9D329A6DA53" box="[1148,1251,1586,1612]" italics="true" pageId="16" pageNumber="17">Cryptops</emphasis>
|
||
</taxonomicName>
|
||
, not even being subgenus-specific as five adults of
|
||
<taxonomicName id="4C77C3B8FFE2D71AFDA4F9B72EC2DA6F" authority="Verhoef, 1934" authorityName="Verhoeff" authorityYear="1934" box="[544,903,1622,1648]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="caucasius" subGenus="Cryptops">
|
||
<emphasis id="B9036429FFE2D71AFDA4F9B72F7FDA6F" box="[544,570,1622,1648]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
|
||
(
|
||
<emphasis id="B9036429FFE2D71AFDCEF9B72F21DA6F" box="[586,612,1622,1648]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
|
||
)
|
||
<emphasis id="B9036429FFE2D71AFDF0F9B62FA7DA6F" box="[628,738,1623,1648]" italics="true" pageId="16" pageNumber="17">caucasius</emphasis>
|
||
Verhoef, 1934
|
||
</taxonomicName>
|
||
(Rc 6430, 8004) and two adults of
|
||
<taxonomicName id="4C77C3B8FFE2D71AFA91F9B72CDCDA8B" authority="Newport, 1844" authorityName="Newport" authorityYear="1844" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="anomalans" subGenus="Cryptops">
|
||
<emphasis id="B9036429FFE2D71AFA91F9B7286ADA6F" box="[1301,1327,1622,1648]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
|
||
(
|
||
<emphasis id="B9036429FFE2D71AFABBF9B7281CDA6F" box="[1343,1369,1622,1648]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
|
||
)
|
||
<emphasis id="B9036429FFE2D71AFAEDF9B62DADDA8B" italics="true" pageId="16" pageNumber="17">anomalans</emphasis>
|
||
<collectingRegion id="49B376D9FFE2D71AFF74F99B2C10DA8B" box="[240,341,1658,1684]" country="United Kingdom" name="Newport" pageId="16" pageNumber="17">Newport</collectingRegion>
|
||
, 1844
|
||
</taxonomicName>
|
||
(Rc 7450) show this pretarsus to be slightly curved (i.e. not hooked), apically pointed and lacking any ventral projection (see also Remarks to
|
||
<emphasis id="B9036429FFE2D71AFD6EF97F2E41DAA7" box="[746,772,1694,1720]" italics="true" pageId="16" pageNumber="17">C.</emphasis>
|
||
(
|
||
<taxonomicName id="4C77C3B8FFE2D71AFC92F97E2EEBDAA7" box="[790,942,1695,1720]" class="Chilopoda" family="Cryptopidae" genus="Paracryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFE2D71AFC92F97E2EEBDAA7" box="[790,942,1695,1720]" italics="true" pageId="16" pageNumber="17">Paracryptops</emphasis>
|
||
</taxonomicName>
|
||
) below). Thus there should be at least four
|
||
<typeStatus id="54CC0699FFE2D71AFF13F9232D97DAC3" box="[151,210,1730,1756]" pageId="16" pageNumber="17">types</typeStatus>
|
||
of this pretarsus in
|
||
<taxonomicName id="4C77C3B8FFE2D71AFE3EF9232F0CDAC3" authorityName="Kohlrausch" authorityYear="1881" box="[442,585,1730,1756]" class="Chilopoda" family="Cryptopidae" kingdom="Animalia" order="Scolopendromorpha" pageId="16" pageNumber="17" phylum="Arthropoda" rank="family">Cryptopidae</taxonomicName>
|
||
: 1. slightly curved, apically pointed, without ventral projection, 2. slightly curved, apically pointed, with ventral projection, 3. slightly curved, lobe-shaped, with ventral projection, 4. hooklike, with ventral projection.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |