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<mods:title id="C93961CD373BD32327F4C032445D8E75">An overview of the extant genera and subgenera of the order Scolopendromorpha (Chilopoda): a new identification key and updated diagnoses</mods:title>
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<mods:namePart id="B1C437256629A9DCF111D8C75D482F89">Schileyko, Arkady A.</mods:namePart>
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<mods:namePart id="CC81EF5C07F382C1F9DB730758CD93EE">Vahtera, Varpu</mods:namePart>
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<mods:namePart id="75DDFA50251C737DCD64CDA190F8F7EC">Edgecombe, Gregory D.</mods:namePart>
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<treatment id="03DE092DFFE1D71FFF13F9CD2CFDDB58" ID-DOI="http://doi.org/10.5281/zenodo.4402201" ID-GBIF-Taxon="177020722" ID-Zenodo-Dep="4402201" LSID="urn:lsid:plazi:treatment:03DE092DFFE1D71FFF13F9CD2CFDDB58" httpUri="http://treatment.plazi.org/id/03DE092DFFE1D71FFF13F9CD2CFDDB58" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20">
<subSubSection id="C36DEBB0FFE1D719FF13F9CD2FCFDA59" box="[151,650,1579,1606]" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="8BC8B83BFFE1D719FF13F9CD2FCFDA59" blockId="19.[151,650,1579,1606]" box="[151,650,1579,1606]" pageId="19" pageNumber="20">
<heading id="D0800F57FFE1D719FF13F9CD2FCFDA59" bold="true" box="[151,650,1579,1606]" fontSize="11" level="2" pageId="19" pageNumber="20" reason="4">
<taxonomicName id="4C77C3B8FFE1D719FF13F9CD2FCFDA59" ID-CoL="8DKV" authority="Verhoeff, 1934" authorityName="Verhoeff" authorityYear="1934" box="[151,650,1579,1606]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="20" phylum="Arthropoda" rank="subGenus" subGenus="Haplocryptops">
<emphasis id="B9036429FFE1D719FF13F9CD2FCFDA59" bold="true" box="[151,650,1579,1606]" pageId="19" pageNumber="20">
<emphasis id="B9036429FFE1D719FF13F9CD2C46DA59" bold="true" box="[151,259,1580,1606]" italics="true" pageId="19" pageNumber="20">Cryptops</emphasis>
(
<emphasis id="B9036429FFE1D719FE90F9CA2C81DA59" bold="true" box="[276,452,1579,1606]" italics="true" pageId="19" pageNumber="20">Haplocryptops</emphasis>
)
<bibRefCitation id="EFE6C5CAFFE1D719FE50F9CD2FCFDA59" author="Verhoeff, K. W." box="[468,650,1580,1606]" pageId="19" pageNumber="20" pagination="1 - 112" refId="ref51555" refString="Verhoeff, K. W. (1934) Beitrage zur Systematik und Geographie der Chilopoden. Zoologische Jahrbucher, 66 (1 / 2), 1 - 112." type="journal article" year="1934">Verhoeff, 1934</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36DEBB0FFE1D719FF13F9902945DA96" box="[151,1024,1648,1673]" pageId="19" pageNumber="20" type="type_taxon">
<paragraph id="8BC8B83BFFE1D719FF13F9902945DA96" blockId="19.[151,1024,1648,1673]" box="[151,1024,1648,1673]" pageId="19" pageNumber="20">
<emphasis id="B9036429FFE1D719FF13F9902C66DA97" bold="true" box="[151,291,1648,1673]" pageId="19" pageNumber="20">
<typeStatus id="54CC0699FFE1D719FF13F9902D89DA96" box="[151,204,1649,1673]" pageId="19" pageNumber="20">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C77C3B8FFE1D719FEAEF9902E1EDA96" ID-CoL="ZZ94" authority="Verhoeff, 1934" authorityName="Verhoeff" authorityYear="1934" box="[298,859,1648,1673]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="acapulcensis" subGenus="Haplocryptops">
<emphasis id="B9036429FFE1D719FEAEF9902CC2DA97" box="[298,391,1649,1672]" italics="true" pageId="19" pageNumber="20">Cryptops</emphasis>
(
<emphasis id="B9036429FFE1D719FE11F9912F69DA97" box="[405,556,1648,1672]" italics="true" pageId="19" pageNumber="20">Haplocryptops</emphasis>
)
<emphasis id="B9036429FFE1D719FDBFF9912FFBDA97" box="[571,702,1648,1672]" italics="true" pageId="19" pageNumber="20">acapulcensis</emphasis>
<bibRefCitation id="EFE6C5CAFFE1D719FD40F9912E1EDA96" author="Verhoeff, K. W." box="[708,859,1648,1673]" pageId="19" pageNumber="20" pagination="1 - 112" refId="ref51555" refString="Verhoeff, K. W. (1934) Beitrage zur Systematik und Geographie der Chilopoden. Zoologische Jahrbucher, 66 (1 / 2), 1 - 112." type="journal article" year="1934">Verhoeff, 1934</bibRefCitation>
</taxonomicName>
(by monotypy).
</paragraph>
</subSubSection>
<subSubSection id="C36DEBB0FFE1D719FF13F9572C80DB7E" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="8BC8B83BFFE1D719FF13F9572E4EDB22" blockId="19.[151,1437,1718,1997]" pageId="19" pageNumber="20">
<emphasis id="B9036429FFE1D719FF13F9572C57DACF" bold="true" box="[151,274,1718,1744]" pageId="19" pageNumber="20">Diagnosis.</emphasis>
Thin pretarsus of maxillae 2 slightly curved and pointed apically, without ventral projection (figs
<quantity id="4C8F15DEFFE1D719FAD3F9562DE8DAEA" metricMagnitude="0" metricUnit="m" metricValue="1.64211" pageId="19" pageNumber="20" unit="in" value="64.65">64, 65 in</quantity>
<bibRefCitation id="EFE6C5CAFFE1D719FF37F93A2C13DAEA" author="Verhoeff, K. W." box="[179,342,1755,1781]" pageId="19" pageNumber="20" pagination="1 - 112" refId="ref51555" refString="Verhoeff, K. W. (1934) Beitrage zur Systematik und Geographie der Chilopoden. Zoologische Jahrbucher, 66 (1 / 2), 1 - 112." type="journal article" year="1934">Verhoeff 1934</bibRefCitation>
); dorsal brush of maxilla 2 composed of very short setae being considerably shorter than pretarsus. Anterior margin of forcipular coxosternite virtually straight (fig.
<quantity id="4C8F15DEFFE1D719FCECF91E2EE5DB06" box="[872,928,1791,1817]" metricMagnitude="0" metricUnit="m" metricValue="1.6764" pageId="19" pageNumber="20" unit="in" value="66.0">66 in</quantity>
<bibRefCitation id="EFE6C5CAFFE1D719FC22F91E290CDB06" author="Verhoeff, K. W." box="[934,1097,1791,1817]" pageId="19" pageNumber="20" pagination="1 - 112" refId="ref51555" refString="Verhoeff, K. W. (1934) Beitrage zur Systematik und Geographie der Chilopoden. Zoologische Jahrbucher, 66 (1 / 2), 1 - 112." type="journal article" year="1934">Verhoeff 1934</bibRefCitation>
); tarsungula long, overlapping each other by at least 1/3 of their length when adducted.
</paragraph>
<paragraph id="8BC8B83BFFE1D719FF43F8A62C80DB7E" blockId="19.[151,1437,1718,1997]" box="[199,453,1862,1889]" pageId="19" pageNumber="20">
<emphasis id="B9036429FFE1D719FF43F8A62CECDB7F" bold="true" box="[199,425,1862,1889]" pageId="19" pageNumber="20">Number of species.</emphasis>
1.
</paragraph>
</subSubSection>
<subSubSection id="C36DEBB0FFE1D71FFF43F88A2C51D82A" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20" type="discussion">
<paragraph id="8BC8B83BFFE1D719FF43F88A2F44DBD2" blockId="19.[151,1437,1718,1997]" pageId="19" pageNumber="20">
<emphasis id="B9036429FFE1D719FF43F88A2C7EDB9A" bold="true" box="[199,315,1899,1925]" pageId="19" pageNumber="20">Remarks.</emphasis>
Treated as a subgenus in
<bibRefCitation id="EFE6C5CAFFE1D719FDF4F88A2947DB9A" author="Edgecombe, G. D. &amp; Bonato, L." box="[624,1026,1899,1925]" pageId="19" pageNumber="20" refId="ref47498" refString="Edgecombe, G. D. &amp; Bonato, L. (2011) Chilopoda-taxonomic overview. Order Scolopendromorpha. In: Minelli, A. (Ed.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 1. Brill, Leiden, pp. 392 - 407." type="book" year="2011">Edgecombe &amp; Bonato (2011: 393)</bibRefCitation>
,
<bibRefCitation id="EFE6C5CAFFE1D719FB96F88A29B3DB9A" author="Lewis, J. G. E." box="[1042,1270,1898,1925]" pageId="19" pageNumber="20" pagination="575 - 584" refId="ref49251" refString="Lewis, J. G. E. (2016 a) On the Cryptops subgenus Trichocryptops Verhoeff, 1937, with a discussion of the problems of differentiating Cryptops species (Chilopoda: Scolopendromorpha: Cryptopidae). Zootaxa, 4139 (4), 575 - 584. https: // doi. org / 10.11646 / zootaxa. 4139.4.9" type="journal article" year="2016">Lewis (2016a: 575)</bibRefCitation>
,
<bibRefCitation id="EFE6C5CAFFE1D719FA81F88A2DA0DBB5" author="Bonato, L. &amp; Chagas-Junior, A. &amp; Edgecombe, G. D. &amp; Lewis, J. G. E. &amp; Minelli, A. &amp; Pereira, L. A. &amp; Shelley, R. M. &amp; Stoev, P. &amp; Zapparoli, M." pageId="19" pageNumber="20" refId="ref45701" refString="Bonato, L., Chagas-Junior, A., Edgecombe, G. D., Lewis, J. G. E., Minelli, A., Pereira, L. A., Shelley, R. M., Stoev, P. &amp; Zapparoli, M. (2016) ChiloBase 2.0 - A World Catalogue of Centipedes (Chilopoda). Available from: http: // chilobase. biologia. unipd. it (accessed 2 August 2016)" type="url" year="2016">
Bonato
<emphasis id="B9036429FFE1D719FAE5F88A28D8DB9A" box="[1377,1437,1899,1925]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
(2016)
</bibRefCitation>
. No information is available on clypeal setose plates, the configuration of sternal sutures/sulci, or the presence and shape of endosternites.
</paragraph>
<caption id="DF08E8B3FFE6D71EFF13F9DC283BDBFE" ID-DOI="http://doi.org/10.5281/zenodo.4402157" ID-Zenodo-Dep="4402157" httpUri="https://zenodo.org/record/4402157/files/figure.png" pageId="20" pageNumber="21" startId="20.[151,264,1597,1621]" targetBox="[372,1216,196,1556]" targetPageId="20">
<paragraph id="8BC8B83BFFE6D71EFF13F9DC283BDBFE" blockId="20.[151,1437,1597,2017]" pageId="20" pageNumber="21">
<emphasis id="B9036429FFE6D71EFF13F9DC2C13DA4A" bold="true" box="[151,342,1597,1621]" pageId="20" pageNumber="21">FIGURES 2632.</emphasis>
<taxonomicName id="4C77C3B8FFE6D71EFED9F9DF2927DA4A" authority="Schileyko, Iorio &amp; Coulis, 2018" authorityName="Schileyko, Iorio &amp; Coulis" authorityYear="2018" box="[349,1122,1597,1621]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="martinicensis" subGenus="Trigonocryptops">
<emphasis id="B9036429FFE6D71EFED9F9DF2CFFDA4A" box="[349,442,1598,1621]" italics="true" pageId="20" pageNumber="21">Cryptops</emphasis>
(
<emphasis id="B9036429FFE6D71EFE4FF9DF2F37DA4A" box="[459,626,1598,1621]" italics="true" pageId="20" pageNumber="21">Trigonocryptops</emphasis>
)
<emphasis id="B9036429FFE6D71EFD07F9DF2E49DA4A" box="[643,780,1598,1621]" italics="true" pageId="20" pageNumber="21">martinicensis</emphasis>
<bibRefCitation id="EFE6C5CAFFE6D71EFC97F9DC2927DA4A" author="Schileyko, A. &amp; Iorio, E. &amp; Coulis, M." box="[787,1122,1597,1621]" pageId="20" pageNumber="21" pagination="559 - 574" refId="ref50439" refString="Schileyko, A., Iorio, E. &amp; Coulis, M. (2018) A contribution to the knowledge of scolopendromorph centipedes of Martinique Island, with descriptions of two new species (Chilopoda: Scolopendromorpha). Zootaxa, 4486 (4), 559 - 574. https: // doi. org / 10.11646 / zootaxa. 4486.4.9" type="journal article" year="2018">Schileyko, Iorio &amp; Coulis, 2018</bibRefCitation>
</taxonomicName>
; Ad. paratype CIRAD P8A
<emphasis id="B9036429FFE6D71EFA07F9DC28D9DA4A" bold="true" box="[1411,1436,1597,1621]" pageId="20" pageNumber="21">26</emphasis>
head, forcipular segment + LBS 13 ventrally;
<taxonomicName id="4C77C3B8FFE6D71EFDF3F9832910DA66" authority="Pocock, 1891" authorityName="Pocock" authorityYear="1891" box="[631,1109,1633,1657]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="weberi" subGenus="Paracryptops">
<emphasis id="B9036429FFE6D71EFDF3F9832F91DA66" box="[631,724,1634,1657]" italics="true" pageId="20" pageNumber="21">Cryptops</emphasis>
(
<emphasis id="B9036429FFE6D71EFD67F9832E28DA66" box="[739,877,1634,1657]" italics="true" pageId="20" pageNumber="21">Paracryptops</emphasis>
)
<emphasis id="B9036429FFE6D71EFCF8F9802E84DA66" box="[892,961,1633,1657]" italics="true" pageId="20" pageNumber="21">weberi</emphasis>
Pocock, 1891
</taxonomicName>
; MCZ DNA 102459
<emphasis id="B9036429FFE6D71EFAAFF9802801DA66" bold="true" box="[1323,1348,1633,1657]" pageId="20" pageNumber="21">27</emphasis>
forcipular segment ventrally;
<taxonomicName id="4C77C3B8FFE6D71EFE04F96729C2DA82" authority="Schileyko, Iorio &amp; Coulis, 2018" authorityName="Schileyko, Iorio &amp; Coulis" authorityYear="2018" box="[384,1159,1669,1693]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="martinicensis" subGenus="Trigonocryptops">
<emphasis id="B9036429FFE6D71EFE04F9672C98DA82" box="[384,477,1670,1693]" italics="true" pageId="20" pageNumber="21">Cryptops</emphasis>
(
<emphasis id="B9036429FFE6D71EFE6AF9672FD0DA82" box="[494,661,1670,1693]" italics="true" pageId="20" pageNumber="21">Trigonocryptops</emphasis>
)
<emphasis id="B9036429FFE6D71EFD22F9672E6ADA82" box="[678,815,1670,1693]" italics="true" pageId="20" pageNumber="21">martinicensis</emphasis>
<bibRefCitation id="EFE6C5CAFFE6D71EFCB3F96429C2DA82" author="Schileyko, A. &amp; Iorio, E. &amp; Coulis, M." box="[823,1159,1669,1693]" pageId="20" pageNumber="21" pagination="559 - 574" refId="ref50439" refString="Schileyko, A., Iorio, E. &amp; Coulis, M. (2018) A contribution to the knowledge of scolopendromorph centipedes of Martinique Island, with descriptions of two new species (Chilopoda: Scolopendromorpha). Zootaxa, 4486 (4), 559 - 574. https: // doi. org / 10.11646 / zootaxa. 4486.4.9" type="journal article" year="2018">Schileyko, Iorio &amp; Coulis, 2018</bibRefCitation>
</taxonomicName>
; Ad. holotype MNHN 459
<emphasis id="B9036429FFE6D71EFF13F9482DF5DADE" bold="true" box="[151,176,1705,1729]" pageId="20" pageNumber="21">28</emphasis>
right ultimate leg medially;
<taxonomicName id="4C77C3B8FFE6D71EFE58F94B2E3FDADE" authority="Verhoeff, 1934" authorityName="Verhoeff" authorityYear="1934" box="[476,890,1705,1729]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="caucasius" subGenus="Cryptops">
<emphasis id="B9036429FFE6D71EFE58F94B2F90DADE" box="[476,725,1705,1729]" italics="true" pageId="20" pageNumber="21">Cryptops (C.) caucasius</emphasis>
<bibRefCitation id="EFE6C5CAFFE6D71EFD59F9482E3FDADE" author="Verhoeff, K. W." box="[733,890,1705,1729]" pageId="20" pageNumber="21" pagination="1 - 112" refId="ref51555" refString="Verhoeff, K. W. (1934) Beitrage zur Systematik und Geographie der Chilopoden. Zoologische Jahrbucher, 66 (1 / 2), 1 - 112." type="journal article" year="1934">Verhoeff, 1934</bibRefCitation>
</taxonomicName>
; Ad. Rc 8003
<emphasis id="B9036429FFE6D71EFB96F948296EDADE" bold="true" box="[1042,1067,1705,1729]" pageId="20" pageNumber="21">29</emphasis>
cephalic plate + anterior margin of forcipular coxosternite ventrally;
<taxonomicName id="4C77C3B8FFE6D71EFE6BF92F2E27DAFA" authority="(Newport, 1845)" baseAuthorityName="Newport" baseAuthorityYear="1845" box="[495,866,1741,1765]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="B9036429FFE6D71EFE6BF92F2FE8DAFA" box="[495,685,1741,1765]" italics="true" pageId="20" pageNumber="21">Cryptops australis</emphasis>
(Newport, 1845)
</taxonomicName>
; AM KS
<emphasis id="B9036429FFE6D71EFC40F92C2E98DAFA" bold="true" box="[964,989,1741,1765]" pageId="20" pageNumber="21">30</emphasis>
apical part of maxilla 2 laterally;
<taxonomicName id="4C77C3B8FFE6D71EFABBF92F2F5EDB16" authority="Pocock, 1891" authorityName="Pocock" authorityYear="1891" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="weberi" subGenus="Paracryptops">
<emphasis id="B9036429FFE6D71EFABBF92F28D9DAFA" box="[1343,1436,1742,1765]" italics="true" pageId="20" pageNumber="21">Cryptops</emphasis>
(
<emphasis id="B9036429FFE6D71EFF24F9132C6FDB16" box="[160,298,1778,1801]" italics="true" pageId="20" pageNumber="21">Paracryptops</emphasis>
)
<emphasis id="B9036429FFE6D71EFEB8F9102CC4DB16" box="[316,385,1777,1801]" italics="true" pageId="20" pageNumber="21">weberi</emphasis>
Pocock, 1891
</taxonomicName>
; MCZ DNA 102459
<emphasis id="B9036429FFE6D71EFD7BF9102E5DDB16" bold="true" box="[767,792,1777,1801]" pageId="20" pageNumber="21">31</emphasis>
apical part of of maxilla 2 laterally;
<taxonomicName id="4C77C3B8FFE6D71EFB23F9132FE8DB32" authority="Schileyko, Iorio &amp; Coulis, 2018" authorityName="Schileyko, Iorio &amp; Coulis" authorityYear="2018" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="martinicensis" subGenus="Trigonocryptops">
<emphasis id="B9036429FFE6D71EFB23F9132841DB16" box="[1191,1284,1778,1801]" italics="true" pageId="20" pageNumber="21">Cryptops</emphasis>
(
<emphasis id="B9036429FFE6D71EFA93F9132D84DB32" italics="true" pageId="20" pageNumber="21">Trigonocryptops</emphasis>
)
<emphasis id="B9036429FFE6D71EFF55F8F72C1FDB32" box="[209,346,1814,1837]" italics="true" pageId="20" pageNumber="21">martinicensis</emphasis>
<bibRefCitation id="EFE6C5CAFFE6D71EFEE5F8F42FE8DB32" author="Schileyko, A. &amp; Iorio, E. &amp; Coulis, M." box="[353,685,1813,1837]" pageId="20" pageNumber="21" pagination="559 - 574" refId="ref50439" refString="Schileyko, A., Iorio, E. &amp; Coulis, M. (2018) A contribution to the knowledge of scolopendromorph centipedes of Martinique Island, with descriptions of two new species (Chilopoda: Scolopendromorpha). Zootaxa, 4486 (4), 559 - 574. https: // doi. org / 10.11646 / zootaxa. 4486.4.9" type="journal article" year="2018">Schileyko, Iorio &amp; Coulis, 2018</bibRefCitation>
</taxonomicName>
; Ad. holotype MNHN 459
<emphasis id="B9036429FFE6D71EFC40F8F42E98DB32" bold="true" box="[964,989,1813,1837]" pageId="20" pageNumber="21">32</emphasis>
LBS 3 ventrally; (
<emphasis id="B9036429FFE6D71EFB1BF8F42984DB32" bold="true" box="[1183,1217,1813,1837]" pageId="20" pageNumber="21">am</emphasis>
)—anterior margin of forcipular coxosternite, (
<emphasis id="B9036429FFE6D71EFE12F8D82CEDDB4E" bold="true" box="[406,424,1849,1873]" pageId="20" pageNumber="21">cl</emphasis>
)—clypeus, (
<emphasis id="B9036429FFE6D71EFDABF8D82F01DB4E" bold="true" box="[559,580,1849,1873]" pageId="20" pageNumber="21">cs</emphasis>
)—clypeal setose plate, (
<emphasis id="B9036429FFE6D71EFCC1F8D82E26DB4E" bold="true" box="[837,867,1849,1873]" pageId="20" pageNumber="21">cst</emphasis>
)—forcipular coxosternite, (
<emphasis id="B9036429FFE6D71EFB07F8D829DADB4E" bold="true" box="[1155,1183,1849,1873]" pageId="20" pageNumber="21">db</emphasis>
)—apical setae of dorsal brush, (
<emphasis id="B9036429FFE6D71EFF63F8BC2DB9DB6A" bold="true" box="[231,252,1885,1909]" pageId="20" pageNumber="21">es</emphasis>
)—enlarged (spiniform) setae, (
<emphasis id="B9036429FFE6D71EFDC7F8BC2F24DB6A" bold="true" box="[579,609,1885,1909]" pageId="20" pageNumber="21">est</emphasis>
)—endosternite, (
<emphasis id="B9036429FFE6D71EFC92F8BC2E58DB6A" bold="true" box="[790,797,1885,1909]" pageId="20" pageNumber="21">l</emphasis>
)—labrum, (
<emphasis id="B9036429FFE6D71EFC1AF8BC2EF6DB6A" bold="true" box="[926,947,1885,1909]" pageId="20" pageNumber="21">lp</emphasis>
)—lateral projection of endosternite, (
<emphasis id="B9036429FFE6D71EFAB9F8BC280BDB6A" bold="true" box="[1341,1358,1885,1909]" pageId="20" pageNumber="21">ls</emphasis>
)—long setae, (
<emphasis id="B9036429FFE6D71EFF64F8602DB9DB86" bold="true" box="[224,252,1921,1945]" pageId="20" pageNumber="21">lsc</emphasis>
)—sternal median longitudinal sulcus, (
<emphasis id="B9036429FFE6D71EFD16F8602FE8DB86" bold="true" box="[658,685,1921,1945]" pageId="20" pageNumber="21">pa</emphasis>
)—short rounded projection of anterior margin of forcipular coxosternite, (
<emphasis id="B9036429FFE6D71EFF24F8442DF2DBA2" bold="true" box="[160,183,1957,1981]" pageId="20" pageNumber="21">pf</emphasis>
)—prefemur, (
<emphasis id="B9036429FFE6D71EFECEF8442C1EDBA2" bold="true" box="[330,347,1957,1981]" pageId="20" pageNumber="21">sl</emphasis>
)—sternal median longitudinal suture, (
<emphasis id="B9036429FFE6D71EFD76F8442E42DBA2" bold="true" box="[754,775,1957,1981]" pageId="20" pageNumber="21">sr</emphasis>
)—sternal transverse ridge, (
<emphasis id="B9036429FFE6D71EFBAAF8442905DBA2" bold="true" box="[1070,1088,1957,1981]" pageId="20" pageNumber="21">st</emphasis>
)—saw teeth, (
<emphasis id="B9036429FFE6D71EFB5DF84429B0DBA2" bold="true" box="[1241,1269,1957,1981]" pageId="20" pageNumber="21">sts</emphasis>
)—sternal transverse suture, (
<emphasis id="B9036429FFE6D71EFEA2F8282C00DBFE" bold="true" box="[294,325,1993,2017]" pageId="20" pageNumber="21">tgs</emphasis>
)—sternal trigonal sutures, (
<emphasis id="B9036429FFE6D71EFDE7F8282F37DBFE" bold="true" box="[611,626,1993,2017]" pageId="20" pageNumber="21">ti</emphasis>
)—tibia, (
<emphasis id="B9036429FFE6D71EFD52F8282FAFDBFE" bold="true" box="[726,746,1993,2017]" pageId="20" pageNumber="21">tr</emphasis>
)—tarsungula, (
<emphasis id="B9036429FFE6D71EFC0EF8282EE0DBFE" bold="true" box="[906,933,1993,2017]" pageId="20" pageNumber="21">vp</emphasis>
)—ventral projection of pretarsus of maxilla 2.
</paragraph>
</caption>
<paragraph id="8BC8B83BFFE7D71FFF43FF762F7ADE9A" blockId="21.[151,1437,151,933]" pageId="21" pageNumber="22">
At the moment the subgenus
<taxonomicName id="4C77C3B8FFE7D71FFD8CFF762FEBDCAE" authorityName="Verhoeff" authorityYear="1934" box="[520,686,151,177]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subGenus" subGenus="Haplocryptops">
<emphasis id="B9036429FFE7D71FFD8CFF762FEBDCAE" box="[520,686,151,177]" italics="true" pageId="21" pageNumber="22">Haplocryptops</emphasis>
</taxonomicName>
is known only from the
<typeStatus id="54CC0699FFE7D71FFC3DFF76295EDCAE" box="[953,1051,151,177]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C77C3B8FFE7D71FFBBAFF76282ADCAE" authorityName="Verhoeff" authorityYear="1934" box="[1086,1391,151,177]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="acapulcensis" subGenus="Haplocryptops">
<emphasis id="B9036429FFE7D71FFBBAFF7629E0DCAE" box="[1086,1189,151,177]" italics="true" pageId="21" pageNumber="22">Cryptops</emphasis>
(
<emphasis id="B9036429FFE7D71FFB37FF76298ADCAF" box="[1203,1231,151,176]" italics="true" pageId="21" pageNumber="22">H.</emphasis>
)
<emphasis id="B9036429FFE7D71FFB5AFF76282ADCAE" box="[1246,1391,151,177]" italics="true" pageId="21" pageNumber="22">acapulcensis</emphasis>
</taxonomicName>
and an incomplete (without ultimate legs) specimen from
<collectingRegion id="49B376D9FFE7D71FFD68FF5A2E7DDCCA" box="[748,824,187,213]" country="Mexico" name="Jalisco" pageId="21" pageNumber="22">Jalisco</collectingRegion>
(also
<collectingCountry id="F360F8ABFFE7D71FFCFFFF5A2E97DCC9" box="[891,978,187,214]" name="Mexico" pageId="21" pageNumber="22">Mexico</collectingCountry>
) assigned to
<emphasis id="B9036429FFE7D71FFBE0FF5A2932DCCA" box="[1124,1143,187,213]" italics="true" pageId="21" pageNumber="22">C</emphasis>
. (
<emphasis id="B9036429FFE7D71FFB09FF5A29E4DCCB" box="[1165,1185,187,212]" italics="true" pageId="21" pageNumber="22">H</emphasis>
.) cf.
<emphasis id="B9036429FFE7D71FFB5FFF5A283FDCCA" box="[1243,1402,187,213]" italics="true" pageId="21" pageNumber="22">acalpuncensis</emphasis>
by
<bibRefCitation id="EFE6C5CAFFE7D71FFF13FF3E2CD3DCE6" author="Cupul-Magana, F. G." box="[151,406,223,249]" pageId="21" pageNumber="22" pagination="1 - 17" refId="ref46934" refString="Cupul-Magana, F. G. (2012) Los ciempies escutigeromorfos (Scutigeromorpha), escolopendromorfos (Scolopendromorpha) y geofilomorfos (Geophilomorpha) de la selva tropical caducifolia de la reserva de Chamela, Jalisco, Mexico. Insecta Mundi, 0208, 1 - 17." type="book chapter" year="2012">Cupul-Magaña (2012)</bibRefCitation>
. The subgeneric diagnosis provided by
<bibRefCitation id="EFE6C5CAFFE7D71FFCD8FF3E29FBDCE6" author="Edgecombe, G. D. &amp; Bonato, L." box="[860,1214,223,249]" pageId="21" pageNumber="22" refId="ref47498" refString="Edgecombe, G. D. &amp; Bonato, L. (2011) Chilopoda-taxonomic overview. Order Scolopendromorpha. In: Minelli, A. (Ed.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 1. Brill, Leiden, pp. 392 - 407." type="book" year="2011">Edgecombe &amp; Bonatos (2011)</bibRefCitation>
attempted to distill characters regarded as diagnosis of
<taxonomicName id="4C77C3B8FFE7D71FFDA8FEE22F97DD02" authorityName="Verhoeff" authorityYear="1934" box="[556,722,259,285]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subGenus" subGenus="Haplocryptops">
<emphasis id="B9036429FFE7D71FFDA8FEE22F97DD02" box="[556,722,259,285]" italics="true" pageId="21" pageNumber="22">Haplocryptops</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFE6C5CAFFE7D71FFD7BFEE22EF3DD02" author="Verhoeff, K. W." box="[767,950,259,285]" pageId="21" pageNumber="22" pagination="1 - 112" refId="ref51555" refString="Verhoeff, K. W. (1934) Beitrage zur Systematik und Geographie der Chilopoden. Zoologische Jahrbucher, 66 (1 / 2), 1 - 112." type="journal article" year="1934">Verhoeff (1934)</bibRefCitation>
. It states (p. 393): “Second maxillary claw [=pretarsus] simple, pointed rather than with a ventral flange; dorsal brush on second maxillae composed of very short setae. Forcipular coxosternite lacking median suture”. The last sentence is not diagnostic because a bulk of species of the nominate subgenus do not exhibit this suture at all (
<figureCitation id="134CA4BEFFE7D71FFC01FE8E2E9EDD96" box="[901,987,367,393]" captionStart="FIGURES 2632" captionStartId="20.[151,264,1597,1621]" captionTargetBox="[372,1216,196,1556]" captionTargetId="figure-19@20.[355,1232,181,1573]" captionTargetPageId="20" captionText="FIGURES 2632. Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio &amp; Coulis, 2018; Ad. paratype CIRAD P8A 26 head, forcipular segment + LBS 13 ventrally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 27 forcipular segment ventrally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio &amp; Coulis, 2018; Ad. holotype MNHN 459 28 right ultimate leg medially; Cryptops (C.) caucasius Verhoeff, 1934; Ad. Rc 8003 29 cephalic plate + anterior margin of forcipular coxosternite ventrally; Cryptops australis (Newport, 1845); AM KS 30 apical part of maxilla 2 laterally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 31 apical part of of maxilla 2 laterally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio &amp; Coulis, 2018; Ad. holotype MNHN 459 32 LBS 3 ventrally; (am)—anterior margin of forcipular coxosternite, (cl)—clypeus, (cs)—clypeal setose plate, (cst)—forcipular coxosternite, (db)—apical setae of dorsal brush, (es)—enlarged (spiniform) setae, (est)—endosternite, (l)—labrum, (lp)—lateral projection of endosternite, (ls)—long setae, (lsc)—sternal median longitudinal sulcus, (pa)—short rounded projection of anterior margin of forcipular coxosternite, (pf)—prefemur, (sl)—sternal median longitudinal suture, (sr)—sternal transverse ridge, (st)—saw teeth, (sts)—sternal transverse suture, (tgs)—sternal trigonal sutures, (ti)—tibia, (tr)—tarsungula, (vp)—ventral projection of pretarsus of maxilla 2." figureDoi="http://doi.org/10.5281/zenodo.4402157" httpUri="https://zenodo.org/record/4402157/files/figure.png" pageId="21" pageNumber="22">Fig. 29</figureCitation>
). According to
<bibRefCitation id="EFE6C5CAFFE7D71FFB0DFE8E2814DD96" author="Verhoeff, K. W." box="[1161,1361,367,393]" pageId="21" pageNumber="22" pagination="1 - 112" refId="ref51555" refString="Verhoeff, K. W. (1934) Beitrage zur Systematik und Geographie der Chilopoden. Zoologische Jahrbucher, 66 (1 / 2), 1 - 112." type="journal article" year="1934">Verhoeffs (1934)</bibRefCitation>
figure 65 the pretarsus of maxilla
<quantity id="4C8F15DEFFE7D71FFE4CFE722CB6DDB2" box="[456,499,403,429]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="21" pageNumber="22" unit="in" value="2.0">2 in</quantity>
<taxonomicName id="4C77C3B8FFE7D71FFE7EFE722FA5DDB2" authorityName="Verhoeff" authorityYear="1934" box="[506,736,403,429]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="acapulcensis" subGenus="Haplocryptops">
<emphasis id="B9036429FFE7D71FFE7EFE722F56DDB2" box="[506,531,403,429]" italics="true" pageId="21" pageNumber="22">C.</emphasis>
(
<emphasis id="B9036429FFE7D71FFDA7FE722F7ADDB3" box="[547,575,403,428]" italics="true" pageId="21" pageNumber="22">H.</emphasis>
)
<emphasis id="B9036429FFE7D71FFDCBFE722FA5DDB2" box="[591,736,403,429]" italics="true" pageId="21" pageNumber="22">acapulcensis</emphasis>
</taxonomicName>
is actually pointed apically (not rounded as in some species of nominotypical subgenus) and has no rounded ventral process (“ventral flange”). This process (fig.
<quantity id="4C8F15DEFFE7D71FFB7DFE562876DDCE" box="[1273,1331,439,465]" metricMagnitude="0" metricUnit="m" metricValue="1.7526" pageId="21" pageNumber="22" unit="in" value="69.0">69 in</quantity>
<bibRefCitation id="EFE6C5CAFFE7D71FFABFFE562D8ADDEA" author="Verhoeff, K. W." pageId="21" pageNumber="22" pagination="1 - 112" refId="ref51555" refString="Verhoeff, K. W. (1934) Beitrage zur Systematik und Geographie der Chilopoden. Zoologische Jahrbucher, 66 (1 / 2), 1 - 112." type="journal article" year="1934">Verhoeff 1934</bibRefCitation>
and fig.
<quantity id="4C8F15DEFFE7D71FFEB4FE3B2C33DDEA" box="[304,374,474,501]" metricMagnitude="0" metricUnit="m" metricValue="7.239" pageId="21" pageNumber="22" unit="in" value="285.0">285 in</quantity>
<bibRefCitation id="EFE6C5CAFFE7D71FFEFFFE3A2F4EDDEA" author="Attems, C." box="[379,523,475,501]" pageId="21" pageNumber="22" refId="ref45522" refString="Attems, C. (1930) Myriapoda 2. Scolopendromorpha. Das Tierreich. Vol. 54. Walter de Gruyter et Co. Publ., Berlin, 308 pp." type="book" year="1930">Attems 1930</bibRefCitation>
) should be present in Palearctic species studied by
<bibRefCitation id="EFE6C5CAFFE7D71FFBC6FE3A29B2DDEA" author="Verhoeff, K. W." box="[1090,1271,475,501]" pageId="21" pageNumber="22" pagination="1 - 112" refId="ref51555" refString="Verhoeff, K. W. (1934) Beitrage zur Systematik und Geographie der Chilopoden. Zoologische Jahrbucher, 66 (1 / 2), 1 - 112." type="journal article" year="1934">Verhoeff (1934)</bibRefCitation>
; who wrote (p. 40) “Diese Putzapparat fand ich bei allen unsern paläarctischen Arten in derselben Weise ausgebildet, nich dagegen bei der in Betracht kommenden mexicanischen Art …”. However as this structure is not characteristic for all species of
<taxonomicName id="4C77C3B8FFE7D71FFF32FDA62CD9DE7E" authority="(Cryptops)" authorityName="Leach" authorityYear="1814" baseAuthorityName="Cryptops" box="[182,412,583,609]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subGenus" subGenus="Cryptops">
<emphasis id="B9036429FFE7D71FFF32FDA62C58DE7E" box="[182,285,583,609]" italics="true" pageId="21" pageNumber="22">Cryptops</emphasis>
(
<emphasis id="B9036429FFE7D71FFEA9FDA62CD1DE7E" box="[301,404,583,609]" italics="true" pageId="21" pageNumber="22">Cryptops</emphasis>
)
</taxonomicName>
and since it is also present in some species of
<emphasis id="B9036429FFE7D71FFC23FDA62925DE7F" box="[935,1120,583,608]" italics="true" pageId="21" pageNumber="22">Trigonocryptops</emphasis>
(see above) it should not be used for separation of
<taxonomicName id="4C77C3B8FFE7D71FFE16FD8A2F7DDE9A" authorityName="Verhoeff" authorityYear="1934" box="[402,568,619,645]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subGenus" subGenus="Haplocryptops">
<emphasis id="B9036429FFE7D71FFE16FD8A2F7DDE9A" box="[402,568,619,645]" italics="true" pageId="21" pageNumber="22">Haplocryptops</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC8B83BFFE7D71FFF43FD6E2CD9DFBA" blockId="21.[151,1437,151,933]" pageId="21" pageNumber="22">
Also, according to the Verhoeff s (1934) figures 64 and 65, the maxillary 2 dorsal brush of
<taxonomicName id="4C77C3B8FFE7D71FFB72FD6E28D9DEB6" authorityName="Verhoeff" authorityYear="1934" box="[1270,1436,655,681]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subGenus" subGenus="Haplocryptops">
<emphasis id="B9036429FFE7D71FFB72FD6E28D9DEB6" box="[1270,1436,655,681]" italics="true" pageId="21" pageNumber="22">Haplocryptops</emphasis>
</taxonomicName>
seems to consist of remarkably short setae, the apical setae being much shorter than the pretarsus. However, as the length of the dorsal brush varies considerably among species of
<taxonomicName id="4C77C3B8FFE7D71FFCFAFD362EA0DEEE" authorityName="Leach" authorityYear="1814" box="[894,997,727,753]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFE7D71FFCFAFD362EA0DEEE" box="[894,997,727,753]" italics="true" pageId="21" pageNumber="22">Cryptops</emphasis>
</taxonomicName>
(see Verhoeffs and Attems drawings mentioned above) this condition cannot guarantee the definite separation of
<taxonomicName id="4C77C3B8FFE7D71FFC6BFD1A29D0DF0A" authorityName="Verhoeff" authorityYear="1934" box="[1007,1173,763,789]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subGenus" subGenus="Haplocryptops">
<emphasis id="B9036429FFE7D71FFC6BFD1A29D0DF0A" box="[1007,1173,763,789]" italics="true" pageId="21" pageNumber="22">Haplocryptops</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFE6C5CAFFE7D71FFB20FD1A2DEBDF26" author="Cupul-Magana, F. G." pageId="21" pageNumber="22" pagination="1 - 17" refId="ref46934" refString="Cupul-Magana, F. G. (2012) Los ciempies escutigeromorfos (Scutigeromorpha), escolopendromorfos (Scolopendromorpha) y geofilomorfos (Geophilomorpha) de la selva tropical caducifolia de la reserva de Chamela, Jalisco, Mexico. Insecta Mundi, 0208, 1 - 17." type="book chapter" year="2012">Cupul-Magaña (2012: 4)</bibRefCitation>
gave no data on structures/details of maxillae
<quantity id="4C8F15DEFFE7D71FFD46FCFE2FABDF26" box="[706,750,799,825]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="21" pageNumber="22" unit="in" value="2.0">2 in</quantity>
his specimen of
<taxonomicName id="4C77C3B8FFE7D71FFC35FCFE29DFDF26" authorityName="Verhoeff" authorityYear="1934" box="[945,1178,799,825]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="acapulcensis" subGenus="Haplocryptops">
<emphasis id="B9036429FFE7D71FFC35FCFE2E8EDF26" box="[945,971,799,825]" italics="true" pageId="21" pageNumber="22">C.</emphasis>
(
<emphasis id="B9036429FFE7D71FFC58FCFE2EBDDF27" box="[988,1016,799,824]" italics="true" pageId="21" pageNumber="22">H.</emphasis>
)
<emphasis id="B9036429FFE7D71FFB8DFCFE29DFDF26" box="[1033,1178,799,825]" italics="true" pageId="21" pageNumber="22">acapulcensis</emphasis>
</taxonomicName>
. Summing up, the differences between
<taxonomicName id="4C77C3B8FFE7D71FFEE6FCA22F4DDF42" authorityName="Verhoeff" authorityYear="1934" box="[354,520,835,861]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subGenus" subGenus="Haplocryptops">
<emphasis id="B9036429FFE7D71FFEE6FCA22F4DDF42" box="[354,520,835,861]" italics="true" pageId="21" pageNumber="22">Haplocryptops</emphasis>
</taxonomicName>
and the nominate subspecies seem to be minor and not significant, leaving us to doubt the validity of the former. However, we prefer to keep this subgenus until representative material from the
<typeStatus id="54CC0699FFE7D71FFF13FC6A2D82DFBA" box="[151,199,907,933]" pageId="21" pageNumber="22">type</typeStatus>
locality is studied.
</paragraph>
<paragraph id="8BC8B83BFFE7D71FFF13FC172E46D80E" blockId="21.[151,771,1014,1077]" box="[151,771,1014,1041]" pageId="21" pageNumber="22">
<emphasis id="B9036429FFE7D71FFF13FC172E46D80E" bold="true" box="[151,771,1014,1041]" pageId="21" pageNumber="22">
(!)
<taxonomicName id="4C77C3B8FFE7D71FFF38FC162FC8D80F" authority="Pocock, 1891" authorityName="Pocock" authorityYear="1891" box="[188,653,1014,1041]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subGenus" subGenus="Paracryptops">
<emphasis id="B9036429FFE7D71FFF38FC162C6DD80E" bold="true" box="[188,296,1015,1041]" italics="true" pageId="21" pageNumber="22">Cryptops</emphasis>
(
<emphasis id="B9036429FFE7D71FFEBCFC162C9DD80E" bold="true" box="[312,472,1015,1041]" italics="true" pageId="21" pageNumber="22">Paracryptops</emphasis>
) Pocock, 1891
</taxonomicName>
stat. nov.
</emphasis>
</paragraph>
<paragraph id="8BC8B83BFFE7D71FFF13FBFA2C51D82A" blockId="21.[151,771,1014,1077]" box="[151,276,1051,1077]" pageId="21" pageNumber="22">
<figureCitation id="134CA4BEFFE7D71FFF13FBFA2C51D82A" box="[151,276,1051,1077]" captionStart="FIGURES 2632" captionStartId="20.[151,264,1597,1621]" captionTargetBox="[372,1216,196,1556]" captionTargetId="figure-19@20.[355,1232,181,1573]" captionTargetPageId="20" captionText="FIGURES 2632. Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio &amp; Coulis, 2018; Ad. paratype CIRAD P8A 26 head, forcipular segment + LBS 13 ventrally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 27 forcipular segment ventrally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio &amp; Coulis, 2018; Ad. holotype MNHN 459 28 right ultimate leg medially; Cryptops (C.) caucasius Verhoeff, 1934; Ad. Rc 8003 29 cephalic plate + anterior margin of forcipular coxosternite ventrally; Cryptops australis (Newport, 1845); AM KS 30 apical part of maxilla 2 laterally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 31 apical part of of maxilla 2 laterally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio &amp; Coulis, 2018; Ad. holotype MNHN 459 32 LBS 3 ventrally; (am)—anterior margin of forcipular coxosternite, (cl)—clypeus, (cs)—clypeal setose plate, (cst)—forcipular coxosternite, (db)—apical setae of dorsal brush, (es)—enlarged (spiniform) setae, (est)—endosternite, (l)—labrum, (lp)—lateral projection of endosternite, (ls)—long setae, (lsc)—sternal median longitudinal sulcus, (pa)—short rounded projection of anterior margin of forcipular coxosternite, (pf)—prefemur, (sl)—sternal median longitudinal suture, (sr)—sternal transverse ridge, (st)—saw teeth, (sts)—sternal transverse suture, (tgs)—sternal trigonal sutures, (ti)—tibia, (tr)—tarsungula, (vp)—ventral projection of pretarsus of maxilla 2." figureDoi="http://doi.org/10.5281/zenodo.4402157" httpUri="https://zenodo.org/record/4402157/files/figure.png" pageId="21" pageNumber="22">Figs 27, 31</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36DEBB0FFE7D71FFF13FBBE2E71D868" box="[151,820,1119,1143]" pageId="21" pageNumber="22" type="type_taxon">
<paragraph id="8BC8B83BFFE7D71FFF13FBBE2E71D868" blockId="21.[151,820,1119,1143]" box="[151,820,1119,1143]" pageId="21" pageNumber="22">
<emphasis id="B9036429FFE7D71FFF13FBBE2C66D868" bold="true" box="[151,291,1119,1143]" pageId="21" pageNumber="22">
<typeStatus id="54CC0699FFE7D71FFF13FBBE2D89D868" box="[151,204,1119,1143]" pageId="21" pageNumber="22">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C77C3B8FFE7D71FFEAEFB812FD5D868" ID-CoL="4CRPM" authority="Pocock, 1891" authorityName="Pocock" authorityYear="1891" box="[298,656,1119,1143]" class="Chilopoda" family="Cryptopidae" genus="Paracryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="weberi">
<emphasis id="B9036429FFE7D71FFEAEFB812CBAD868" box="[298,511,1119,1143]" italics="true" pageId="21" pageNumber="22">Paracryptops weberi</emphasis>
Pocock, 1891
</taxonomicName>
(by monotypy).
</paragraph>
</subSubSection>
<subSubSection id="C36DEBB0FFE7D71FFF13FB442969D9FF" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="8BC8B83BFFE7D71FFF13FB442921D9A4" blockId="21.[151,1437,1189,1863]" pageId="21" pageNumber="22">
<emphasis id="B9036429FFE7D71FFF13FB442C57D8A0" bold="true" box="[151,274,1189,1215]" pageId="21" pageNumber="22">Diagnosis.</emphasis>
Clypeus without setose plates. Pretarsus of maxillae 2 slender, from slightly to strongly hooked apically, with (
<figureCitation id="134CA4BEFFE7D71FFF5EFB282C68D8FC" box="[218,301,1225,1251]" captionStart="FIGURES 2632" captionStartId="20.[151,264,1597,1621]" captionTargetBox="[372,1216,196,1556]" captionTargetId="figure-19@20.[355,1232,181,1573]" captionTargetPageId="20" captionText="FIGURES 2632. Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio &amp; Coulis, 2018; Ad. paratype CIRAD P8A 26 head, forcipular segment + LBS 13 ventrally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 27 forcipular segment ventrally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio &amp; Coulis, 2018; Ad. holotype MNHN 459 28 right ultimate leg medially; Cryptops (C.) caucasius Verhoeff, 1934; Ad. Rc 8003 29 cephalic plate + anterior margin of forcipular coxosternite ventrally; Cryptops australis (Newport, 1845); AM KS 30 apical part of maxilla 2 laterally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 31 apical part of of maxilla 2 laterally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio &amp; Coulis, 2018; Ad. holotype MNHN 459 32 LBS 3 ventrally; (am)—anterior margin of forcipular coxosternite, (cl)—clypeus, (cs)—clypeal setose plate, (cst)—forcipular coxosternite, (db)—apical setae of dorsal brush, (es)—enlarged (spiniform) setae, (est)—endosternite, (l)—labrum, (lp)—lateral projection of endosternite, (ls)—long setae, (lsc)—sternal median longitudinal sulcus, (pa)—short rounded projection of anterior margin of forcipular coxosternite, (pf)—prefemur, (sl)—sternal median longitudinal suture, (sr)—sternal transverse ridge, (st)—saw teeth, (sts)—sternal transverse suture, (tgs)—sternal trigonal sutures, (ti)—tibia, (tr)—tarsungula, (vp)—ventral projection of pretarsus of maxilla 2." figureDoi="http://doi.org/10.5281/zenodo.4402157" httpUri="https://zenodo.org/record/4402157/files/figure.png" pageId="21" pageNumber="22">Fig. 31</figureCitation>
) or without ventral projection. Very dense dorsal brush (
<figureCitation id="134CA4BEFFE7D71FFC32FB28294CD8FC" box="[950,1033,1225,1251]" captionStart="FIGURES 2632" captionStartId="20.[151,264,1597,1621]" captionTargetBox="[372,1216,196,1556]" captionTargetId="figure-19@20.[355,1232,181,1573]" captionTargetPageId="20" captionText="FIGURES 2632. Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio &amp; Coulis, 2018; Ad. paratype CIRAD P8A 26 head, forcipular segment + LBS 13 ventrally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 27 forcipular segment ventrally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio &amp; Coulis, 2018; Ad. holotype MNHN 459 28 right ultimate leg medially; Cryptops (C.) caucasius Verhoeff, 1934; Ad. Rc 8003 29 cephalic plate + anterior margin of forcipular coxosternite ventrally; Cryptops australis (Newport, 1845); AM KS 30 apical part of maxilla 2 laterally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 31 apical part of of maxilla 2 laterally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio &amp; Coulis, 2018; Ad. holotype MNHN 459 32 LBS 3 ventrally; (am)—anterior margin of forcipular coxosternite, (cl)—clypeus, (cs)—clypeal setose plate, (cst)—forcipular coxosternite, (db)—apical setae of dorsal brush, (es)—enlarged (spiniform) setae, (est)—endosternite, (l)—labrum, (lp)—lateral projection of endosternite, (ls)—long setae, (lsc)—sternal median longitudinal sulcus, (pa)—short rounded projection of anterior margin of forcipular coxosternite, (pf)—prefemur, (sl)—sternal median longitudinal suture, (sr)—sternal transverse ridge, (st)—saw teeth, (sts)—sternal transverse suture, (tgs)—sternal trigonal sutures, (ti)—tibia, (tr)—tarsungula, (vp)—ventral projection of pretarsus of maxilla 2." figureDoi="http://doi.org/10.5281/zenodo.4402157" httpUri="https://zenodo.org/record/4402157/files/figure.png" pageId="21" pageNumber="22">Fig. 31</figureCitation>
) visibly longer than (or as long as) pretarsus, consisting of tiny and virtually transparent setae. Anterior margin of forcipular coxosternite with short, blunt, apically slightly rounded lobes which have no chitinised margin; a few long setae placed at bases of these projections; sharply pointed tarsungula very short (
<figureCitation id="134CA4BEFFE7D71FFD5CFAD42E69D950" box="[728,812,1333,1359]" captionStart="FIGURES 2632" captionStartId="20.[151,264,1597,1621]" captionTargetBox="[372,1216,196,1556]" captionTargetId="figure-19@20.[355,1232,181,1573]" captionTargetPageId="20" captionText="FIGURES 2632. Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio &amp; Coulis, 2018; Ad. paratype CIRAD P8A 26 head, forcipular segment + LBS 13 ventrally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 27 forcipular segment ventrally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio &amp; Coulis, 2018; Ad. holotype MNHN 459 28 right ultimate leg medially; Cryptops (C.) caucasius Verhoeff, 1934; Ad. Rc 8003 29 cephalic plate + anterior margin of forcipular coxosternite ventrally; Cryptops australis (Newport, 1845); AM KS 30 apical part of maxilla 2 laterally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 31 apical part of of maxilla 2 laterally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio &amp; Coulis, 2018; Ad. holotype MNHN 459 32 LBS 3 ventrally; (am)—anterior margin of forcipular coxosternite, (cl)—clypeus, (cs)—clypeal setose plate, (cst)—forcipular coxosternite, (db)—apical setae of dorsal brush, (es)—enlarged (spiniform) setae, (est)—endosternite, (l)—labrum, (lp)—lateral projection of endosternite, (ls)—long setae, (lsc)—sternal median longitudinal sulcus, (pa)—short rounded projection of anterior margin of forcipular coxosternite, (pf)—prefemur, (sl)—sternal median longitudinal suture, (sr)—sternal transverse ridge, (st)—saw teeth, (sts)—sternal transverse suture, (tgs)—sternal trigonal sutures, (ti)—tibia, (tr)—tarsungula, (vp)—ventral projection of pretarsus of maxilla 2." figureDoi="http://doi.org/10.5281/zenodo.4402157" httpUri="https://zenodo.org/record/4402157/files/figure.png" pageId="21" pageNumber="22">Fig. 27</figureCitation>
), barely (or even not) overlapping each other when adducted. Sternites with well-developed transverse thickening between coxae of legs and with very wide and shallow incomplete longitudinal depression (not sulcus) at the place of median suture. Short endosternite well bordered by transverse suture, but in most LBS not visible, being covered by the following sternite.
</paragraph>
<paragraph id="8BC8B83BFFE7D71FFF43FA242969D9FF" blockId="21.[151,1437,1189,1863]" box="[199,1068,1477,1504]" pageId="21" pageNumber="22">
<emphasis id="B9036429FFE7D71FFF43FA242CECD9C0" bold="true" box="[199,425,1477,1503]" pageId="21" pageNumber="22">Number of species.</emphasis>
5 (
<bibRefCitation id="EFE6C5CAFFE7D71FFE48FA242E04D9FF" author="Edgecombe, G. D. &amp; Bonato, L." box="[460,833,1477,1504]" pageId="21" pageNumber="22" refId="ref47498" refString="Edgecombe, G. D. &amp; Bonato, L. (2011) Chilopoda-taxonomic overview. Order Scolopendromorpha. In: Minelli, A. (Ed.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 1. Brill, Leiden, pp. 392 - 407." type="book" year="2011">Edgecombe &amp; Bonato 2011: 395</bibRefCitation>
,
<bibRefCitation id="EFE6C5CAFFE7D71FFCC8FA272965D9FF" author="Bonato, L. &amp; Chagas-Junior, A. &amp; Edgecombe, G. D. &amp; Lewis, J. G. E. &amp; Minelli, A. &amp; Pereira, L. A. &amp; Shelley, R. M. &amp; Stoev, P. &amp; Zapparoli, M." box="[844,1056,1477,1504]" pageId="21" pageNumber="22" refId="ref45701" refString="Bonato, L., Chagas-Junior, A., Edgecombe, G. D., Lewis, J. G. E., Minelli, A., Pereira, L. A., Shelley, R. M., Stoev, P. &amp; Zapparoli, M. (2016) ChiloBase 2.0 - A World Catalogue of Centipedes (Chilopoda). Available from: http: // chilobase. biologia. unipd. it (accessed 2 August 2016)" type="url" year="2016">
Bonato
<emphasis id="B9036429FFE7D71FFC20FA272E99D9C0" box="[932,988,1477,1503]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
2016
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36DEBB0FFE7D71FFF43FA082CFDDB58" pageId="21" pageNumber="22" type="discussion">
<paragraph id="8BC8B83BFFE7D71FFF43FA0829A8DAA8" blockId="21.[151,1437,1189,1863]" pageId="21" pageNumber="22">
<emphasis id="B9036429FFE7D71FFF43FA082C7EDA1C" bold="true" box="[199,315,1513,1539]" pageId="21" pageNumber="22">Remarks.</emphasis>
The most recent morphological accounts on
<taxonomicName id="4C77C3B8FFE7D71FFCCEFA0B2EA7DA1C" box="[842,994,1514,1539]" class="Chilopoda" family="Cryptopidae" genus="Paracryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFE7D71FFCCEFA0B2EA7DA1C" box="[842,994,1514,1539]" italics="true" pageId="21" pageNumber="22">Paracryptops</emphasis>
</taxonomicName>
were given by Chagas-Jr and Shelley (2004: 3) and
<bibRefCitation id="EFE6C5CAFFE7D71FFEBDF9EC2F6FDA38" author="Schileyko, A." box="[313,554,1549,1575]" pageId="21" pageNumber="22" pagination="71 - 95" refId="ref50073" refString="Schileyko, A. (2007) The scolopendromorph centipedes (Chilopoda) of Vietnam, with contributions to the faunas of Cambodia and Laos. Part 3. Arthropoda Selecta, 16 (2), 71 - 95." type="journal article" year="2007">Schileyko (2007: 91)</bibRefCitation>
. Treated as a genus in
<bibRefCitation id="EFE6C5CAFFE7D71FFCAAF9EC29FDDA37" author="Edgecombe, G. D. &amp; Bonato, L." box="[814,1208,1549,1576]" pageId="21" pageNumber="22" refId="ref47498" refString="Edgecombe, G. D. &amp; Bonato, L. (2011) Chilopoda-taxonomic overview. Order Scolopendromorpha. In: Minelli, A. (Ed.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 1. Brill, Leiden, pp. 392 - 407." type="book" year="2011">Edgecombe &amp; Bonato (2011: 395)</bibRefCitation>
. Based on both molecular and morphological results
<bibRefCitation id="EFE6C5CAFFE7D71FFDA5F9D02E08DA54" author="Vahtera, V. &amp; Edgecombe, G. D. &amp; Giribet, G." box="[545,845,1585,1611]" pageId="21" pageNumber="22" pagination="4 - 20" refId="ref51271" refString="Vahtera, V., Edgecombe, G. D. &amp; Giribet, G. (2012 a) Evolution of blindness in scolopendromorph centipedes (Chilopoda: Scolopendromorpha): insight from an expanded sampling of molecular data. Cladistics, 28, 4 - 20. https: // doi. org / 10.1111 / j. 1096 - 0031.2011.00361. x" type="journal article" year="2012">
Vahtera
<emphasis id="B9036429FFE7D71FFD05F9D32FF8DA54" box="[641,701,1585,1611]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
(2012a: 13)
</bibRefCitation>
wrote that
<taxonomicName id="4C77C3B8FFE7D71FFC52F9D3292BDA54" box="[982,1134,1586,1611]" class="Chilopoda" family="Cryptopidae" genus="Paracryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFE7D71FFC52F9D3292BDA54" box="[982,1134,1586,1611]" italics="true" pageId="21" pageNumber="22">Paracryptops</emphasis>
</taxonomicName>
“nested within
<taxonomicName id="4C77C3B8FFE7D71FFAADF9D328D5DA53" authorityName="Leach" authorityYear="1814" box="[1321,1424,1586,1612]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFE7D71FFAADF9D328D5DA53" box="[1321,1424,1586,1612]" italics="true" pageId="21" pageNumber="22">Cryptops</emphasis>
</taxonomicName>
” without formalizing the new taxonomic stratus of the latter, although in their subsequent work (2012b) they treated
<taxonomicName id="4C77C3B8FFE7D71FFF13F99B2C6ADA8C" box="[151,303,1658,1683]" class="Chilopoda" family="Cryptopidae" genus="Paracryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFE7D71FFF13F99B2C6ADA8C" box="[151,303,1658,1683]" italics="true" pageId="21" pageNumber="22">Paracryptops</emphasis>
</taxonomicName>
as a genus again. Taking into consideration the unequivocal results obtained by
<bibRefCitation id="EFE6C5CAFFE7D71FFB22F99828DDDA8C" author="Vahtera, V. &amp; Edgecombe, G. D. &amp; Giribet, G." box="[1190,1432,1657,1683]" pageId="21" pageNumber="22" pagination="4 - 20" refId="ref51271" refString="Vahtera, V., Edgecombe, G. D. &amp; Giribet, G. (2012 a) Evolution of blindness in scolopendromorph centipedes (Chilopoda: Scolopendromorpha): insight from an expanded sampling of molecular data. Cladistics, 28, 4 - 20. https: // doi. org / 10.1111 / j. 1096 - 0031.2011.00361. x" type="journal article" year="2012">
Vahtera
<emphasis id="B9036429FFE7D71FFA85F99B287DDA8C" box="[1281,1336,1657,1683]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
(2012a)
</bibRefCitation>
, we propose this taxon to be a subgenus of
<taxonomicName id="4C77C3B8FFE7D71FFDF6F97F2F9CDAA7" authorityName="Leach" authorityYear="1814" box="[626,729,1694,1720]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFE7D71FFDF6F97F2F9CDAA7" box="[626,729,1694,1720]" italics="true" pageId="21" pageNumber="22">Cryptops</emphasis>
</taxonomicName>
, i.e.
<emphasis id="B9036429FFE7D71FFC94F97F2E6FDAA7" box="[784,810,1694,1720]" italics="true" pageId="21" pageNumber="22">C.</emphasis>
(
<taxonomicName id="4C77C3B8FFE7D71FFCBEF97F2E97DAA8" box="[826,978,1694,1719]" class="Chilopoda" family="Cryptopidae" genus="Paracryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFE7D71FFCBEF97F2E97DAA8" box="[826,978,1694,1719]" italics="true" pageId="21" pageNumber="22">Paracryptops</emphasis>
</taxonomicName>
) Pocock, 1891
<emphasis id="B9036429FFE7D71FFB06F97C29A8DAA8" bold="true" box="[1154,1261,1693,1719]" pageId="21" pageNumber="22">stat. nov.</emphasis>
</paragraph>
<paragraph id="8BC8B83BFFE7D71FFF43F9202CFDDB58" blockId="21.[151,1437,1189,1863]" pageId="21" pageNumber="22">
Seven adult specimens from
<collectingCountry id="F360F8ABFFE7D71FFD89F9202F29DAC4" box="[525,620,1729,1755]" name="Vietnam" pageId="21" pageNumber="22">Vietnam</collectingCountry>
(Rc 6535, 6658, 7130, 7383, 7433) of
<taxonomicName id="4C77C3B8FFE7D71FFBACF9232830DAC4" authority="Silvestri, 1924" authorityName="Silvestri" authorityYear="1924" box="[1064,1397,1729,1756]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="indicus" subGenus="Paracryptops">
<emphasis id="B9036429FFE7D71FFBACF9232907DAC3" box="[1064,1090,1730,1756]" italics="true" pageId="21" pageNumber="22">C.</emphasis>
(
<emphasis id="B9036429FFE7D71FFBD7F923292DDAC4" box="[1107,1128,1730,1755]" italics="true" pageId="21" pageNumber="22">P.</emphasis>
)
<emphasis id="B9036429FFE7D71FFBFCF920298CDAC4" box="[1144,1225,1729,1755]" italics="true" pageId="21" pageNumber="22">indicus</emphasis>
Silvestri, 1924
</taxonomicName>
restudied by light microscopy show the pretarsus of maxillae 2 being slightly curved and lacking ventral projection(s) and sternal median suture being absent (an uncommon conditionin
<taxonomicName id="4C77C3B8FFE7D71FFC1DF8E8296DDB3B" authorityName="Kohlrausch" authorityYear="1881" box="[921,1064,1801,1828]" class="Chilopoda" family="Cryptopidae" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="22" phylum="Arthropoda" rank="family">Cryptopidae</taxonomicName>
), sometimes replaced by poorlydeveloped median sulcus.
</paragraph>
</subSubSection>
</treatment>
</document>