327 lines
49 KiB
XML
327 lines
49 KiB
XML
<document id="97EB862FDC0FA3BB99562B1D0CA61EFF" ID-CLB-Dataset="8703" ID-DOI="10.11646/zootaxa.4825.1.1" ID-GBIF-Dataset="5ab5f5c8-481e-4d1a-8643-21e72c367278" ID-ISSN="1175-5326" ID-Zenodo-Dep="4402145" ID-ZooBank="F230F199-1C94-4E2E-9CE4-5F56212C015F" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1609341789210" checkinUser="plazi" docAuthor="Schileyko, Arkady A., Vahtera, Varpu & Edgecombe, Gregory D." docDate="2020" docId="03DE092DFFDED727FF13FCFF294EDE22" docLanguage="en" docName="zootaxa.4825.1.1.pdf" docOrigin="Zootaxa 4825 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Digitipes Attems 1930" docType="treatment" docVersion="5" lastPageNumber="46" masterDocId="FFE77155FFF2D70AFF84FFE12D45DC1F" masterDocTitle="An overview of the extant genera and subgenera of the order Scolopendromorpha (Chilopoda): a new identification key and updated diagnoses" masterLastPageNumber="64" masterPageNumber="1" pageNumber="45" updateTime="1698852004925" updateUser="ExternalLinkService">
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<mods:titleInfo id="48A56665E169A77346E4632073A3BCF3">
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<mods:title id="8574AAC68BC1945C8949ED8FA7237189">An overview of the extant genera and subgenera of the order Scolopendromorpha (Chilopoda): a new identification key and updated diagnoses</mods:title>
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<mods:name id="ED7482D626DE0A9D233299254E493CBE" type="personal">
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<mods:namePart id="92BEDFBD37F5087EF181F5AC05177B00">Schileyko, Arkady A.</mods:namePart>
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<mods:nameIdentifier id="7FDDD526691FC5B630017620D6F0BB44" type="email">schileyko1965@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="0FE3281F6D731157974AEE7EB354EA44">Author</mods:roleTerm>
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<mods:namePart id="66C61C2EC74986E93C631D932A205542">Vahtera, Varpu</mods:namePart>
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<mods:nameIdentifier id="C0478FB93A8B112CFF26988D90522DCF" type="email">varpu.vahtera@gmail.com</mods:nameIdentifier>
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<mods:namePart id="33CFA4151CF596B1516F0B963B0F7DA7">Edgecombe, Gregory D.</mods:namePart>
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<mods:nameIdentifier id="2C2D617618B5BE2D63A87D5EC99E6AE3" type="email">schileyko1965@gmail.com</mods:nameIdentifier>
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<mods:title id="BAE336DFB159FD5E164B87984604A5A9">Zootaxa</mods:title>
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<mods:date id="6A5CB8D1FDB949CCB6C0DADC7CEA6DB9">2020</mods:date>
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<mods:number id="1728C59DCBBFD64E726D9425E55E2DF5">2020-08-10</mods:number>
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<mods:number id="63740505A738B58B64299974D72F38A2">4825</mods:number>
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<treatment id="03DE092DFFDED727FF13FCFF294EDE22" ID-DOI="http://doi.org/10.5281/zenodo.4457013" ID-GBIF-Taxon="177020727" ID-Zenodo-Dep="4457013" LSID="urn:lsid:plazi:treatment:03DE092DFFDED727FF13FCFF294EDE22" httpUri="http://treatment.plazi.org/id/03DE092DFFDED727FF13FCFF294EDE22" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
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<subSubSection id="C36DEBB0FFDED726FF13FCFF2C51DF42" pageId="44" pageNumber="45" type="discussion">
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<subSubSection id="C36DEBB0FFDED726FF13FCFF2C95DF27" box="[151,464,798,825]" pageId="44" pageNumber="45" type="nomenclature">
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<paragraph id="8BC8B83BFFDED726FF13FCFF2C95DF27" blockId="44.[151,464,798,861]" box="[151,464,798,825]" pageId="44" pageNumber="45">
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<emphasis id="B9036429FFDED726FF13FCFF2C95DF27" bold="true" box="[151,464,798,825]" pageId="44" pageNumber="45">
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(!)
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<taxonomicName id="4C77C3B8FFDED726FF38FCFE2C95DF27" ID-CoL="45DM" ID-ENA="1096233" authority="Attems, 1930" authorityName="Attems" authorityYear="1930" box="[188,464,798,825]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
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<emphasis id="B9036429FFDED726FF38FCFE2C6DDF26" bold="true" box="[188,296,799,825]" italics="true" pageId="44" pageNumber="45">Digitipes</emphasis>
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<bibRefCitation id="EFE6C5CAFFDED726FEA9FCFF2C95DF27" author="Attems, C." box="[301,464,798,824]" pageId="44" pageNumber="45" refId="ref45522" refString="Attems, C. (1930) Myriapoda 2. Scolopendromorpha. Das Tierreich. Vol. 54. Walter de Gruyter et Co. Publ., Berlin, 308 pp." type="book" year="1930">Attems, 1930</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C36DEBB0FFDED726FF13FCA22C51DF42" box="[151,276,835,861]" pageId="44" pageNumber="45" type="description">
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<paragraph id="8BC8B83BFFDED726FF13FCA22C51DF42" blockId="44.[151,464,798,861]" box="[151,276,835,861]" pageId="44" pageNumber="45">
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<figureCitation id="134CA4BEFFDED726FF13FCA22C51DF42" box="[151,276,835,861]" captionStart="FIGURES 89–96" captionStartId="47.[151,264,1723,1747]" captionTargetBox="[181,1399,207,1674]" captionTargetId="figure-17@47.[166,1421,181,1700]" captionTargetPageId="47" captionText="FIGURES 89–96. Digitipes coonoorensis Jangi & Dass, 1984; Male CES 091088 89 femur of right ultimate leg medially; Digitipes sp.; Ad. Rc 7565 90 clypeal part of head ventrally 91 forcipular segment ventrally 92 LBS 13–15 ventrally; Otostigmus (O.) multidens nordicus Schileyko, 1995; Ad. Rc 6545 93 LBS 21 ventrally; Ad. Rc 6747 94 anterior part of head ventrally; Otostigmus (O.) astenus (Kohlrausch, 1881); MCZ DNA 102463 95 pretarsus of maxilla 2; Sad. Rc 7507 96 forcipular segment ventrally; (ap)—apical spines of coxopleural process, (as)—accessory spines, (bs)—basal suture of forcipular toothplate, (cl)—clypeus, (cx)—coxopleuron, (df)—conical distomedial process of ultimate femur, (f)—ultimate femur, (l)—labrum, (lc)—lateral spine of coxopleural process, (lsc)—sternal median longitudinal sulcus, (md)—medial longitudinal depression of ultimate femur, (p2)—pretarsus of maxilla 2, (pd)—sternal posterior depression, (pr)—trochantero-prefemoral process, (psl)—sternal paramedian sulci, (tp)—forcipular tooth-plate, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402185" httpUri="https://zenodo.org/record/4402185/files/figure.png" pageId="44" pageNumber="45">Figs 89–92</figureCitation>
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</paragraph>
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</subSubSection>
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</subSubSection>
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<subSubSection id="C36DEBB0FFDED726FF13FC662E6ADF80" box="[151,815,903,927]" pageId="44" pageNumber="45" type="type_taxon">
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<paragraph id="8BC8B83BFFDED726FF13FC662E6ADF80" blockId="44.[151,815,903,927]" box="[151,815,903,927]" pageId="44" pageNumber="45">
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<emphasis id="B9036429FFDED726FF13FC662C66DF80" bold="true" box="[151,291,903,927]" pageId="44" pageNumber="45">
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<typeStatus id="54CC0699FFDED726FF13FC662D89DF80" box="[151,204,903,927]" pageId="44" pageNumber="45">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C77C3B8FFDED726FEAEFC692FCFDF80" ID-CoL="363VX" authority="Attems, 1930" authorityName="Attems" authorityYear="1930" box="[298,650,903,927]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="verdascens">
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<emphasis id="B9036429FFDED726FEAEFC692CB8DF80" box="[298,509,903,927]" italics="true" pageId="44" pageNumber="45">Digitipes verdascens</emphasis>
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<bibRefCitation id="EFE6C5CAFFDED726FD86FC662FCFDF80" author="Attems, C." box="[514,650,903,927]" pageId="44" pageNumber="45" refId="ref45522" refString="Attems, C. (1930) Myriapoda 2. Scolopendromorpha. Das Tierreich. Vol. 54. Walter de Gruyter et Co. Publ., Berlin, 308 pp." type="book" year="1930">Attems, 1930</bibRefCitation>
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</taxonomicName>
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(by monotypy).
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</paragraph>
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</subSubSection>
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<subSubSection id="C36DEBB0FFDED726FF13FC2C2E2ED933" pageId="44" pageNumber="45" type="diagnosis">
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<paragraph id="8BC8B83BFFDED726FF13FC2C2E2ED933" blockId="44.[151,1437,973,2008]" pageId="44" pageNumber="45">
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<emphasis id="B9036429FFDED726FF13FC2C2C57DFF8" bold="true" box="[151,274,973,999]" pageId="44" pageNumber="45">Diagnosis.</emphasis>
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Median tooth of labrum very small, much shorter than lateral labral lobes (
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<figureCitation id="134CA4BEFFDED726FBF5FC2C2982DFF8" box="[1137,1223,973,999]" captionStart="FIGURES 89–96" captionStartId="47.[151,264,1723,1747]" captionTargetBox="[181,1399,207,1674]" captionTargetId="figure-17@47.[166,1421,181,1700]" captionTargetPageId="47" captionText="FIGURES 89–96. Digitipes coonoorensis Jangi & Dass, 1984; Male CES 091088 89 femur of right ultimate leg medially; Digitipes sp.; Ad. Rc 7565 90 clypeal part of head ventrally 91 forcipular segment ventrally 92 LBS 13–15 ventrally; Otostigmus (O.) multidens nordicus Schileyko, 1995; Ad. Rc 6545 93 LBS 21 ventrally; Ad. Rc 6747 94 anterior part of head ventrally; Otostigmus (O.) astenus (Kohlrausch, 1881); MCZ DNA 102463 95 pretarsus of maxilla 2; Sad. Rc 7507 96 forcipular segment ventrally; (ap)—apical spines of coxopleural process, (as)—accessory spines, (bs)—basal suture of forcipular toothplate, (cl)—clypeus, (cx)—coxopleuron, (df)—conical distomedial process of ultimate femur, (f)—ultimate femur, (l)—labrum, (lc)—lateral spine of coxopleural process, (lsc)—sternal median longitudinal sulcus, (md)—medial longitudinal depression of ultimate femur, (p2)—pretarsus of maxilla 2, (pd)—sternal posterior depression, (pr)—trochantero-prefemoral process, (psl)—sternal paramedian sulci, (tp)—forcipular tooth-plate, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402185" httpUri="https://zenodo.org/record/4402185/files/figure.png" pageId="44" pageNumber="45">Fig. 90</figureCitation>
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). Forcipular toothplates present, trochantero-prefemur with well-developed process (
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<figureCitation id="134CA4BEFFDED726FC1CFC102EAAD814" box="[920,1007,1009,1035]" captionStart="FIGURES 89–96" captionStartId="47.[151,264,1723,1747]" captionTargetBox="[181,1399,207,1674]" captionTargetId="figure-17@47.[166,1421,181,1700]" captionTargetPageId="47" captionText="FIGURES 89–96. Digitipes coonoorensis Jangi & Dass, 1984; Male CES 091088 89 femur of right ultimate leg medially; Digitipes sp.; Ad. Rc 7565 90 clypeal part of head ventrally 91 forcipular segment ventrally 92 LBS 13–15 ventrally; Otostigmus (O.) multidens nordicus Schileyko, 1995; Ad. Rc 6545 93 LBS 21 ventrally; Ad. Rc 6747 94 anterior part of head ventrally; Otostigmus (O.) astenus (Kohlrausch, 1881); MCZ DNA 102463 95 pretarsus of maxilla 2; Sad. Rc 7507 96 forcipular segment ventrally; (ap)—apical spines of coxopleural process, (as)—accessory spines, (bs)—basal suture of forcipular toothplate, (cl)—clypeus, (cx)—coxopleuron, (df)—conical distomedial process of ultimate femur, (f)—ultimate femur, (l)—labrum, (lc)—lateral spine of coxopleural process, (lsc)—sternal median longitudinal sulcus, (md)—medial longitudinal depression of ultimate femur, (p2)—pretarsus of maxilla 2, (pd)—sternal posterior depression, (pr)—trochantero-prefemoral process, (psl)—sternal paramedian sulci, (tp)—forcipular tooth-plate, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402185" httpUri="https://zenodo.org/record/4402185/files/figure.png" pageId="44" pageNumber="45">Fig. 91</figureCitation>
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). Rarely some posterior tergites with well-developed longitudinal keels (for example in
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<taxonomicName id="4C77C3B8FFDED726FD49FBF7291DD830" authority="Jangi and Dass, 1984" authorityName="Jangi and Dass" authorityYear="1984" box="[717,1112,1045,1072]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="barnabasi">
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<emphasis id="B9036429FFDED726FD49FBF72E24D830" box="[717,865,1045,1071]" italics="true" pageId="44" pageNumber="45">D. barnabasi</emphasis>
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Jangi and Dass, 1984
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</taxonomicName>
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; see fig. 10 of
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<bibRefCitation id="EFE6C5CAFFDED726FB78FBF42C6DD84C" author="Joshi, J. & Edgecombe, G. D." pageId="44" pageNumber="45" pagination="99 - 145" refId="ref47850" refString="Joshi, J. & Edgecombe, G. D. (2013) Revision of the scolopendrid centipede Digitipes Attems, 1930, from India (Chilopoda: Scolopendromorpha): reconciling molecular and morphological estimates of species diversity. Zootaxa, 3626 (1), 99 - 145. https: // doi. org / 10.11646 / zootaxa. 3626.1.5" type="journal article" year="2013">Joshi & Edgecombe 2013</bibRefCitation>
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), usually these keels lacking or very poorly developed. Sternites may bear paramedian sutures (or sulci) as well as a median longitudinal sulcus and posterior depression (
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<figureCitation id="134CA4BEFFDED726FC3DFBBC294BD868" box="[953,1038,1117,1143]" captionStart="FIGURES 89–96" captionStartId="47.[151,264,1723,1747]" captionTargetBox="[181,1399,207,1674]" captionTargetId="figure-17@47.[166,1421,181,1700]" captionTargetPageId="47" captionText="FIGURES 89–96. Digitipes coonoorensis Jangi & Dass, 1984; Male CES 091088 89 femur of right ultimate leg medially; Digitipes sp.; Ad. Rc 7565 90 clypeal part of head ventrally 91 forcipular segment ventrally 92 LBS 13–15 ventrally; Otostigmus (O.) multidens nordicus Schileyko, 1995; Ad. Rc 6545 93 LBS 21 ventrally; Ad. Rc 6747 94 anterior part of head ventrally; Otostigmus (O.) astenus (Kohlrausch, 1881); MCZ DNA 102463 95 pretarsus of maxilla 2; Sad. Rc 7507 96 forcipular segment ventrally; (ap)—apical spines of coxopleural process, (as)—accessory spines, (bs)—basal suture of forcipular toothplate, (cl)—clypeus, (cx)—coxopleuron, (df)—conical distomedial process of ultimate femur, (f)—ultimate femur, (l)—labrum, (lc)—lateral spine of coxopleural process, (lsc)—sternal median longitudinal sulcus, (md)—medial longitudinal depression of ultimate femur, (p2)—pretarsus of maxilla 2, (pd)—sternal posterior depression, (pr)—trochantero-prefemoral process, (psl)—sternal paramedian sulci, (tp)—forcipular tooth-plate, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402185" httpUri="https://zenodo.org/record/4402185/files/figure.png" pageId="44" pageNumber="45">Fig. 92</figureCitation>
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). LBS 7 lacking spiracles, the latter with atrium. Legs with tarsal spur(s), the only exception—
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<taxonomicName id="4C77C3B8FFDED726FCABFB63290CD884" authority="(Pocock, 1890)" baseAuthorityName="Pocock" baseAuthorityYear="1890" box="[815,1097,1153,1179]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="nudus">
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<emphasis id="B9036429FFDED726FCABFB632ED0D884" box="[815,917,1153,1179]" italics="true" pageId="44" pageNumber="45">D. nudus</emphasis>
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(Pocock, 1890)
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</taxonomicName>
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(former
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<taxonomicName id="4C77C3B8FFDED726FB2AFB632C3AD8A0" authority="Joshi & Edgecombe, 2013" authorityName="Joshi & Edgecombe" authorityYear="2013" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="periyarensis">
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<emphasis id="B9036429FFDED726FB2AFB63281FD884" box="[1198,1370,1154,1179]" italics="true" pageId="44" pageNumber="45">D. periyarensis</emphasis>
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<bibRefCitation id="EFE6C5CAFFDED726FAE7FB602C3AD8A0" author="Joshi, J. & Edgecombe, G. D." pageId="44" pageNumber="45" pagination="99 - 145" refId="ref47850" refString="Joshi, J. & Edgecombe, G. D. (2013) Revision of the scolopendrid centipede Digitipes Attems, 1930, from India (Chilopoda: Scolopendromorpha): reconciling molecular and morphological estimates of species diversity. Zootaxa, 3626 (1), 99 - 145. https: // doi. org / 10.11646 / zootaxa. 3626.1.5" type="journal article" year="2013">Joshi & Edgecombe, 2013</bibRefCitation>
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</taxonomicName>
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) (see below). Coxopleural process from short to moderately long, with apical spines. Ultimate prefemur with spines but lacking corner spine, pretarsus well-developed. Femur of males with well-developed (but incomplete) medial longitudinal depression leading to an obtuse conical distomedial process (
|
||
<figureCitation id="134CA4BEFFDED726FB4DFB0C2859D918" box="[1225,1308,1261,1287]" captionStart="FIGURES 89–96" captionStartId="47.[151,264,1723,1747]" captionTargetBox="[181,1399,207,1674]" captionTargetId="figure-17@47.[166,1421,181,1700]" captionTargetPageId="47" captionText="FIGURES 89–96. Digitipes coonoorensis Jangi & Dass, 1984; Male CES 091088 89 femur of right ultimate leg medially; Digitipes sp.; Ad. Rc 7565 90 clypeal part of head ventrally 91 forcipular segment ventrally 92 LBS 13–15 ventrally; Otostigmus (O.) multidens nordicus Schileyko, 1995; Ad. Rc 6545 93 LBS 21 ventrally; Ad. Rc 6747 94 anterior part of head ventrally; Otostigmus (O.) astenus (Kohlrausch, 1881); MCZ DNA 102463 95 pretarsus of maxilla 2; Sad. Rc 7507 96 forcipular segment ventrally; (ap)—apical spines of coxopleural process, (as)—accessory spines, (bs)—basal suture of forcipular toothplate, (cl)—clypeus, (cx)—coxopleuron, (df)—conical distomedial process of ultimate femur, (f)—ultimate femur, (l)—labrum, (lc)—lateral spine of coxopleural process, (lsc)—sternal median longitudinal sulcus, (md)—medial longitudinal depression of ultimate femur, (p2)—pretarsus of maxilla 2, (pd)—sternal posterior depression, (pr)—trochantero-prefemoral process, (psl)—sternal paramedian sulci, (tp)—forcipular tooth-plate, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402185" httpUri="https://zenodo.org/record/4402185/files/figure.png" pageId="44" pageNumber="45">Fig. 89</figureCitation>
|
||
) that bears scattered setae (see also figs 9, 21,
|
||
<quantity id="4C8F15DEFFDED726FDA4FAF02FC6D934" box="[544,643,1297,1324]" metricMagnitude="0" metricUnit="m" metricValue="1.0805159999999998" pageId="44" pageNumber="45" unit="in" value="42.54">42, 54 in</quantity>
|
||
<bibRefCitation id="EFE6C5CAFFDED726FD0EFAF02E1AD933" author="Siriwut, W. & Edgecombe, G. D. & Sutcharit, C. & Tongkerd, P. & Panha, S." box="[650,863,1297,1324]" pageId="44" pageNumber="45" pagination="71 - 87" refId="ref50894" refString="Siriwut, W., Edgecombe, G. D., Sutcharit, C., Tongkerd, P. & Panha, S. (2015) First record of the African-Indian centipede genus Digitipes Attems, 1930 (Scolopendromorpha: Otostigminae) from Myanmar, and the systematic position of a new species based on molecular phylogenetics. Zootaxa, 3931 (1), 71 - 87. https: // doi. org / 10.11646 / zootaxa. 3931.1.5" type="journal article" year="2015">
|
||
Siriwut
|
||
<emphasis id="B9036429FFDED726FD67FAF32E51D934" box="[739,788,1297,1323]" italics="true" pageId="44" pageNumber="45">et al</emphasis>
|
||
. 2015
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36DEBB0FFDED726FF43FAD42F5DD987" pageId="44" pageNumber="45" type="description">
|
||
<paragraph id="8BC8B83BFFDED726FF43FAD42D81D96B" blockId="44.[151,1437,973,2008]" pageId="44" pageNumber="45">
|
||
<emphasis id="B9036429FFDED726FF43FAD42CEBD950" bold="true" box="[199,430,1333,1359]" pageId="44" pageNumber="45">Number of species.</emphasis>
|
||
9 (
|
||
<bibRefCitation id="EFE6C5CAFFDED726FE53FAD42E1DD950" author="Edgecombe, G. D. & Bonato, L." box="[471,856,1333,1360]" pageId="44" pageNumber="45" refId="ref47498" refString="Edgecombe, G. D. & Bonato, L. (2011) Chilopoda-taxonomic overview. Order Scolopendromorpha. In: Minelli, A. (Ed.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 1. Brill, Leiden, pp. 392 - 407." type="book" year="2011">Edgecombe & Bonato 2011: 402</bibRefCitation>
|
||
), 11 (
|
||
<bibRefCitation id="EFE6C5CAFFDED726FC1EFAD42932D94F" author="Bonato, L. & Chagas-Junior, A. & Edgecombe, G. D. & Lewis, J. G. E. & Minelli, A. & Pereira, L. A. & Shelley, R. M. & Stoev, P. & Zapparoli, M." box="[922,1143,1333,1360]" pageId="44" pageNumber="45" refId="ref45701" refString="Bonato, L., Chagas-Junior, A., Edgecombe, G. D., Lewis, J. G. E., Minelli, A., Pereira, L. A., Shelley, R. M., Stoev, P. & Zapparoli, M. (2016) ChiloBase 2.0 - A World Catalogue of Centipedes (Chilopoda). Available from: http: // chilobase. biologia. unipd. it (accessed 2 August 2016)" type="url" year="2016">
|
||
Bonato
|
||
<emphasis id="B9036429FFDED726FC72FAD7296FD950" box="[1014,1066,1333,1359]" italics="true" pageId="44" pageNumber="45">et al</emphasis>
|
||
. 2016
|
||
</bibRefCitation>
|
||
), 10 (
|
||
<bibRefCitation id="EFE6C5CAFFDED726FB3EFAD42DF2D96B" author="Siriwut, W. & Edgecombe, G. D. & Sutcharit, C. & Tongkerd, P. & Panha, S." pageId="44" pageNumber="45" pagination="71 - 87" refId="ref50894" refString="Siriwut, W., Edgecombe, G. D., Sutcharit, C., Tongkerd, P. & Panha, S. (2015) First record of the African-Indian centipede genus Digitipes Attems, 1930 (Scolopendromorpha: Otostigminae) from Myanmar, and the systematic position of a new species based on molecular phylogenetics. Zootaxa, 3931 (1), 71 - 87. https: // doi. org / 10.11646 / zootaxa. 3931.1.5" type="journal article" year="2015">
|
||
Siriwut
|
||
<emphasis id="B9036429FFDED726FA9CFAD72816D950" box="[1304,1363,1333,1359]" italics="true" pageId="44" pageNumber="45">et al.</emphasis>
|
||
2015: 76
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC8B83BFFDED726FF43FA9C2F5DD987" blockId="44.[151,1437,973,2008]" box="[199,536,1405,1432]" pageId="44" pageNumber="45">
|
||
<emphasis id="B9036429FFDED726FF43FA9C2CF2D988" bold="true" box="[199,439,1405,1431]" pageId="44" pageNumber="45">Sexual dimorphism.</emphasis>
|
||
Present.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36DEBB0FFDED727FF43FA40294EDE22" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45" type="discussion">
|
||
<subSubSection id="C36DEBB0FFDED726FF43FA402DB3DAA8" pageId="44" pageNumber="45" type="discussion">
|
||
<paragraph id="8BC8B83BFFDED726FF43FA402DB3DAA8" blockId="44.[151,1437,973,2008]" pageId="44" pageNumber="45">
|
||
<emphasis id="B9036429FFDED726FF43FA402C7ED9A4" bold="true" box="[199,315,1441,1467]" pageId="44" pageNumber="45">Remarks.</emphasis>
|
||
Treated as a genus in
|
||
<bibRefCitation id="EFE6C5CAFFDED726FDB6FA402EF3D9A4" author="Edgecombe, G. D. & Bonato, L." box="[562,950,1441,1468]" pageId="44" pageNumber="45" refId="ref47498" refString="Edgecombe, G. D. & Bonato, L. (2011) Chilopoda-taxonomic overview. Order Scolopendromorpha. In: Minelli, A. (Ed.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 1. Brill, Leiden, pp. 392 - 407." type="book" year="2011">Edgecombe & Bonato (2011: 402)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE6C5CAFFDED726FC45FA402864D9A4" author="Joshi, J. & Edgecombe, G. D." box="[961,1313,1441,1467]" pageId="44" pageNumber="45" pagination="99 - 145" refId="ref47850" refString="Joshi, J. & Edgecombe, G. D. (2013) Revision of the scolopendrid centipede Digitipes Attems, 1930, from India (Chilopoda: Scolopendromorpha): reconciling molecular and morphological estimates of species diversity. Zootaxa, 3626 (1), 99 - 145. https: // doi. org / 10.11646 / zootaxa. 3626.1.5" type="journal article" year="2013">Joshi & Edgecombe (2013: 99)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE6C5CAFFDED726FAA8FA402C07D9FF" author="Vahtera, V. & Edgecombe, G. D. & Giribet, G." pageId="44" pageNumber="45" pagination="578 - 602" refId="ref51397" refString="Vahtera, V., Edgecombe, G. D. & Giribet, G. (2013). Phylogenetics of scolopendromorph centipedes: can denser taxon sampling improve an artificial classification? Invertebrate Systematics, 27 (5), 578 - 602. https: // doi. org / 10.1071 / IS 13035" type="journal article" year="2013">
|
||
Vahtera
|
||
<emphasis id="B9036429FFDED726FA0CFA432DE8D9C0" italics="true" pageId="44" pageNumber="45">et al</emphasis>
|
||
. (2013: 594)
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE6C5CAFFDED726FECAFA242F8ED9C0" author="Joshi, J. & Edgecombe, G. D." box="[334,715,1477,1503]" pageId="44" pageNumber="45" pagination="1316 - 1335" refId="ref47976" refString="Joshi, J. & Edgecombe, G. D. (2018) Molecular phylogeny and systematics of the centipede genus Ethmostigmus Pocock (Chilopoda: Scolopendromorpha) from peninsular India. Invertebrate Systematics, 32, 1316 - 1335. https: // doi. org / 10.1071 / IS 18030" type="journal article" year="2018">Joshi & Edgecombe (2018: 1318)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE6C5CAFFDED726FD53FA24296ED9C0" author="Joshi, J. & Edgecombe, G. D." box="[727,1067,1477,1503]" pageId="44" pageNumber="45" pagination="1 - 10" refId="ref48029" refString="Joshi, J. & Edgecombe, G. D. (2019) Evolutionary biogeography of the centipede genus Ethmostigmus from Peninsular India: testing an ancient vicariance hypothesis for Old World tropical diversity. BMC Evolutionary Biology, 19 (41), 1 - 10. https: // doi. org / 10.1186 / s 12862 - 019 - 1367 - 6" type="journal article" year="2019">Joshi & Edgecombe (2019: 3)</bibRefCitation>
|
||
.
|
||
<bibRefCitation id="EFE6C5CAFFDED726FBB3FA242859D9FF" author="Siriwut, W. & Edgecombe, G. D. & Sutcharit, C. & Tongkerd, P. & Panha, S." box="[1079,1308,1477,1504]" pageId="44" pageNumber="45" pagination="71 - 87" refId="ref50894" refString="Siriwut, W., Edgecombe, G. D., Sutcharit, C., Tongkerd, P. & Panha, S. (2015) First record of the African-Indian centipede genus Digitipes Attems, 1930 (Scolopendromorpha: Otostigminae) from Myanmar, and the systematic position of a new species based on molecular phylogenetics. Zootaxa, 3931 (1), 71 - 87. https: // doi. org / 10.11646 / zootaxa. 3931.1.5" type="journal article" year="2015">
|
||
Siriwut
|
||
<emphasis id="B9036429FFDED726FB15FA27298FD9C0" box="[1169,1226,1477,1503]" italics="true" pageId="44" pageNumber="45">et al.</emphasis>
|
||
(2015)
|
||
</bibRefCitation>
|
||
considered it as a genus but wrote (p. 83): “Within the
|
||
<taxonomicName id="4C77C3B8FFDED726FD0DFA082E66DA1C" authorityName="Kraepelin" authorityYear="1903" box="[649,803,1513,1539]" class="Chilopoda" family="Scolopendridae" kingdom="Animalia" order="Scolopendromorpha" pageId="44" pageNumber="45" phylum="Arthropoda" rank="subFamily" subFamily="Otostigminae">Otostigminae</taxonomicName>
|
||
, members of
|
||
<taxonomicName id="4C77C3B8FFDED726FC39FA0B2961DA1C" authorityName="Attems" authorityYear="1930" box="[957,1060,1514,1539]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFDED726FC39FA0B2961DA1C" box="[957,1060,1514,1539]" italics="true" pageId="44" pageNumber="45">Digitipes</emphasis>
|
||
</taxonomicName>
|
||
were separated into two clades”.
|
||
<bibRefCitation id="EFE6C5CAFFDED726FF13F9EC2C95DA38" author="Joshi, J. & Edgecombe, G. D." box="[151,464,1549,1575]" pageId="44" pageNumber="45" pagination="393 - 408" refId="ref47914" refString="Joshi, J. & Edgecombe, G. D. (2017) Tracking the variability of phenotypic traits on a molecular phylogeny: an example from scolopendrid centipedes in peninsular India. Organisms Diversity & Evolution, 17, 393 - 408. https: // doi. org / 10.1007 / s 13127 - 017 - 0323 - 9" type="journal article" year="2017">Joshi & Edgecombe (2017)</bibRefCitation>
|
||
analyzed in detail the Indian species of this genus. More densely sampled molecular phylogenies by
|
||
<bibRefCitation id="EFE6C5CAFFDED726FECCF9D02F6FDA54" author="Siriwut, W. & Edgecombe, G. D. & Sutcharit, C. & Tongkerd, P. & Panha, S." box="[328,554,1585,1611]" pageId="44" pageNumber="45" pagination="1005 - 1049" refId="ref51057" refString="Siriwut, W., Edgecombe, G. D., Sutcharit, C., Tongkerd, P. & Panha, S. (2018) Systematic revision and phylogenetic reassessment of the centipede genera Rhysida Wood, 1862 and Alluropus Silvestri, 1912 (Chilopoda: Scolopendromorpha) in Southeast Asia, with further discussion of the subfamily Otostigminae. Invertebrate Systematics, 32, 1005 - 1049. https: // doi. org / 10.1071 / IS 17081" type="journal article" year="2018">
|
||
Siriwut
|
||
<emphasis id="B9036429FFDED726FE25F9D32C9CDA54" box="[417,473,1585,1611]" italics="true" pageId="44" pageNumber="45">et al.</emphasis>
|
||
(2018)
|
||
</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFE6C5CAFFDED726FDE4F9D02E6CDA54" author="Joshi, J. & Karanth, P. K. & Edgecombe, G. D." box="[608,809,1585,1612]" pageId="44" pageNumber="45" pagination="828 - 861" refId="ref48095" refString="Joshi, J., Karanth, P. K. & Edgecombe, G. D. (2020) The Out-of-India hypothesis: Evidence from an ancient centipede genus, Rhysida (Chilopoda: Scolopendromorpha) from the Oriental Region, and systematics of Indian species. Zoological Journal of the Linnean Society, 189, 828 - 861." type="journal article" year="2020">
|
||
Joshi
|
||
<emphasis id="B9036429FFDED726FD24F9D32F94DA54" box="[672,721,1585,1611]" italics="true" pageId="44" pageNumber="45">et al</emphasis>
|
||
. (2020)
|
||
</bibRefCitation>
|
||
depicted
|
||
<taxonomicName id="4C77C3B8FFDED726FC11F9D32EB9DA54" authorityName="Attems" authorityYear="1930" box="[917,1020,1586,1611]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFDED726FC11F9D32EB9DA54" box="[917,1020,1586,1611]" italics="true" pageId="44" pageNumber="45">Digitipes</emphasis>
|
||
</taxonomicName>
|
||
as monophyletic apart from the nesting of
|
||
<taxonomicName id="4C77C3B8FFDED726FF5AF9B72F06DA70" authority="Pocock, 1890" authorityName="Pocock" authorityYear="1890" box="[222,579,1621,1648]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="nudus">
|
||
<emphasis id="B9036429FFDED726FF5AF9B72CE3DA70" box="[222,422,1621,1648]" italics="true" pageId="44" pageNumber="45">Otostigmus nudus</emphasis>
|
||
Pocock, 1890
|
||
</taxonomicName>
|
||
(former
|
||
<taxonomicName id="4C77C3B8FFDED726FD20F9B72E0BDA70" authorityName="Joshi & Edgecombe" authorityYear="2013" box="[676,846,1622,1647]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="periyarensis">
|
||
<emphasis id="B9036429FFDED726FD20F9B72E0BDA70" box="[676,846,1622,1647]" italics="true" pageId="44" pageNumber="45">D. periyarensis</emphasis>
|
||
</taxonomicName>
|
||
) within it. However, more material should be studied (using both traditional and molecular approaches) to be sure to which genus (
|
||
<emphasis id="B9036429FFDED726FB87F99B2939DA8B" box="[1027,1148,1658,1684]" italics="true" pageId="44" pageNumber="45">Otistigmus</emphasis>
|
||
or
|
||
<taxonomicName id="4C77C3B8FFDED726FB27F99B284FDA8C" authorityName="Attems" authorityYear="1930" box="[1187,1290,1658,1683]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFDED726FB27F99B284FDA8C" box="[1187,1290,1658,1683]" italics="true" pageId="44" pageNumber="45">Digitipes</emphasis>
|
||
</taxonomicName>
|
||
) this species belongs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36DEBB0FFDED726FF43F9232950DBAC" pageId="44" pageNumber="45" type="description">
|
||
<paragraph id="8BC8B83BFFDED726FF43F9232EECDB73" blockId="44.[151,1437,973,2008]" pageId="44" pageNumber="45">
|
||
An incomplete (it lacks some posterior LBS) specimen of
|
||
<taxonomicName id="4C77C3B8FFDED726FCE1F9232E89DAC4" authorityName="Attems" authorityYear="1930" box="[869,972,1730,1755]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFDED726FCE1F9232E89DAC4" box="[869,972,1730,1755]" italics="true" pageId="44" pageNumber="45">Digitipes</emphasis>
|
||
</taxonomicName>
|
||
sp. (Rc 7565 from E
|
||
<collectingCountry id="F360F8ABFFDED726FB4EF9202853DAC3" box="[1226,1302,1729,1756]" name="Gabon" pageId="44" pageNumber="45">Gabon</collectingCountry>
|
||
) has a very small median tooth of the labrum (
|
||
<figureCitation id="134CA4BEFFDED726FDADF9042F3ADAE0" box="[553,639,1765,1791]" captionStart="FIGURES 89–96" captionStartId="47.[151,264,1723,1747]" captionTargetBox="[181,1399,207,1674]" captionTargetId="figure-17@47.[166,1421,181,1700]" captionTargetPageId="47" captionText="FIGURES 89–96. Digitipes coonoorensis Jangi & Dass, 1984; Male CES 091088 89 femur of right ultimate leg medially; Digitipes sp.; Ad. Rc 7565 90 clypeal part of head ventrally 91 forcipular segment ventrally 92 LBS 13–15 ventrally; Otostigmus (O.) multidens nordicus Schileyko, 1995; Ad. Rc 6545 93 LBS 21 ventrally; Ad. Rc 6747 94 anterior part of head ventrally; Otostigmus (O.) astenus (Kohlrausch, 1881); MCZ DNA 102463 95 pretarsus of maxilla 2; Sad. Rc 7507 96 forcipular segment ventrally; (ap)—apical spines of coxopleural process, (as)—accessory spines, (bs)—basal suture of forcipular toothplate, (cl)—clypeus, (cx)—coxopleuron, (df)—conical distomedial process of ultimate femur, (f)—ultimate femur, (l)—labrum, (lc)—lateral spine of coxopleural process, (lsc)—sternal median longitudinal sulcus, (md)—medial longitudinal depression of ultimate femur, (p2)—pretarsus of maxilla 2, (pd)—sternal posterior depression, (pr)—trochantero-prefemoral process, (psl)—sternal paramedian sulci, (tp)—forcipular tooth-plate, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402185" httpUri="https://zenodo.org/record/4402185/files/figure.png" pageId="44" pageNumber="45">Fig. 90</figureCitation>
|
||
), which is also characteristic within
|
||
<taxonomicName id="4C77C3B8FFDED726FBA2F90429F8DAE0" authorityName="Kraepelin" authorityYear="1903" box="[1062,1213,1765,1791]" class="Chilopoda" family="Scolopendridae" kingdom="Animalia" order="Scolopendromorpha" pageId="44" pageNumber="45" phylum="Arthropoda" rank="subFamily" subFamily="Otostigminae">Otostigminae</taxonomicName>
|
||
for
|
||
<emphasis id="B9036429FFDED726FB74F9072C66DB3C" italics="true" pageId="44" pageNumber="45">
|
||
<taxonomicName id="4C77C3B8FFDED726FB74F90728C5DB1F" authorityName="Attems" authorityYear="1934" box="[1264,1408,1766,1792]" class="Chilopoda" family="Scolopendridae" genus="Sterropristes" kingdom="Animalia" order="Scolopendromorpha" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">Sterropristes</taxonomicName>
|
||
+
|
||
<taxonomicName id="4C77C3B8FFDED726FF13F8E82C66DB3C" authorityName="Tomosvary" authorityYear="1882" box="[151,291,1801,1827]" class="Chilopoda" family="Scolopendridae" genus="Edentistoma" kingdom="Animalia" order="Scolopendromorpha" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">Edentistoma</taxonomicName>
|
||
</emphasis>
|
||
(see below), and an unusually wide process of the forcipular trochantero-prefemur (
|
||
<figureCitation id="134CA4BEFFDED726FB5FF8E8286ADB3C" box="[1243,1327,1801,1827]" captionStart="FIGURES 89–96" captionStartId="47.[151,264,1723,1747]" captionTargetBox="[181,1399,207,1674]" captionTargetId="figure-17@47.[166,1421,181,1700]" captionTargetPageId="47" captionText="FIGURES 89–96. Digitipes coonoorensis Jangi & Dass, 1984; Male CES 091088 89 femur of right ultimate leg medially; Digitipes sp.; Ad. Rc 7565 90 clypeal part of head ventrally 91 forcipular segment ventrally 92 LBS 13–15 ventrally; Otostigmus (O.) multidens nordicus Schileyko, 1995; Ad. Rc 6545 93 LBS 21 ventrally; Ad. Rc 6747 94 anterior part of head ventrally; Otostigmus (O.) astenus (Kohlrausch, 1881); MCZ DNA 102463 95 pretarsus of maxilla 2; Sad. Rc 7507 96 forcipular segment ventrally; (ap)—apical spines of coxopleural process, (as)—accessory spines, (bs)—basal suture of forcipular toothplate, (cl)—clypeus, (cx)—coxopleuron, (df)—conical distomedial process of ultimate femur, (f)—ultimate femur, (l)—labrum, (lc)—lateral spine of coxopleural process, (lsc)—sternal median longitudinal sulcus, (md)—medial longitudinal depression of ultimate femur, (p2)—pretarsus of maxilla 2, (pd)—sternal posterior depression, (pr)—trochantero-prefemoral process, (psl)—sternal paramedian sulci, (tp)—forcipular tooth-plate, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402185" httpUri="https://zenodo.org/record/4402185/files/figure.png" pageId="44" pageNumber="45">Fig. 91</figureCitation>
|
||
). In addition, this specimen has no legs with tarsal spurs (a very rare condition in
|
||
<taxonomicName id="4C77C3B8FFDED726FC39F8CC291CDB57" authorityName="Kraepelin" authorityYear="1903" box="[957,1113,1837,1864]" class="Chilopoda" family="Scolopendridae" kingdom="Animalia" order="Scolopendromorpha" pageId="44" pageNumber="45" phylum="Arthropoda" rank="subFamily" subFamily="Otostigminae">Otostigminae</taxonomicName>
|
||
), so it is similar to the former
|
||
<taxonomicName id="4C77C3B8FFDED726FF13F8B32CCBDB74" authorityName="Joshi & Edgecombe" authorityYear="2013" box="[151,398,1874,1899]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="periyarensis">
|
||
<emphasis id="B9036429FFDED726FF13F8B32CCBDB74" box="[151,398,1874,1899]" italics="true" pageId="44" pageNumber="45">Digitipes periyarensis</emphasis>
|
||
</taxonomicName>
|
||
in this (and some other) morphological aspects.
|
||
</paragraph>
|
||
<paragraph id="8BC8B83BFFDED726FF43F8972950DBAC" blockId="44.[151,1437,973,2008]" pageId="44" pageNumber="45">
|
||
An obtuse conical distomedial process of the femur of male
|
||
<taxonomicName id="4C77C3B8FFDED726FCE2F8972E88DB90" authorityName="Attems" authorityYear="1930" box="[870,973,1910,1935]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFDED726FCE2F8972E88DB90" box="[870,973,1910,1935]" italics="true" pageId="44" pageNumber="45">Digitipes</emphasis>
|
||
</taxonomicName>
|
||
is (with the exception of being spineless) similar to the standard corner spine of the ultimate prefemur of
|
||
<taxonomicName id="4C77C3B8FFDED726FCDBF8782954DBAC" authorityName="Leach" authorityYear="1814" box="[863,1041,1945,1971]" class="Chilopoda" family="Scolopendridae" kingdom="Animalia" order="Scolopendromorpha" pageId="44" pageNumber="45" phylum="Arthropoda" rank="family">Scolopendridae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36DEBB0FFDED727FF43F85F294EDE22" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45" type="discussion">
|
||
<paragraph id="8BC8B83BFFDED727FF43F85F2C75DD7A" blockId="44.[151,1437,973,2008]" lastBlockId="45.[151,1437,151,573]" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
|
||
We consider this genus as problematic since the femoral characters are restricted to mature males such that the females of
|
||
<taxonomicName id="4C77C3B8FFDFD727FEBBFF762CE3DCAF" authorityName="Attems" authorityYear="1930" box="[319,422,151,176]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFDFD727FEBBFF762CE3DCAF" box="[319,422,151,176]" italics="true" pageId="45" pageNumber="46">Digitipes</emphasis>
|
||
</taxonomicName>
|
||
cannot be morphologically separated from those of
|
||
<taxonomicName id="4C77C3B8FFDFD727FC77FF76284BDCAE" authority="(Otostigmus)" authorityName="Porat" authorityYear="1876" baseAuthorityName="Otostigmus" box="[1011,1294,151,177]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="45" pageNumber="46" phylum="Arthropoda" rank="subGenus" subGenus="Otostigmus">
|
||
<emphasis id="B9036429FFDFD727FC77FF762936DCAE" box="[1011,1139,151,177]" italics="true" pageId="45" pageNumber="46">Otostigmus</emphasis>
|
||
(
|
||
<emphasis id="B9036429FFDFD727FB07FF762846DCAE" box="[1155,1283,151,177]" italics="true" pageId="45" pageNumber="46">Otostigmus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
. In addition, “Specimens that possess the femoral process make up a small percentage of our total sample of
|
||
<taxonomicName id="4C77C3B8FFDFD727FB5AFF5A2800DCCB" authority="(Joshi & Edgecombe 2013: 101)" authorityName="Attems" authorityYear="1930" baseAuthorityName="Joshi & Edgecombe" baseAuthorityPageNumber="101" baseAuthorityYear="2013" box="[1246,1349,187,212]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFDFD727FB5AFF5A2800DCCB" box="[1246,1349,187,212]" italics="true" pageId="45" pageNumber="46">Digitipes</emphasis>
|
||
</taxonomicName>
|
||
” (
|
||
<bibRefCitation id="EFE6C5CAFFDFD727FAE5FF5A2CF6DCE6" author="Joshi, J. & Edgecombe, G. D." pageId="45" pageNumber="46" pagination="99 - 145" refId="ref47850" refString="Joshi, J. & Edgecombe, G. D. (2013) Revision of the scolopendrid centipede Digitipes Attems, 1930, from India (Chilopoda: Scolopendromorpha): reconciling molecular and morphological estimates of species diversity. Zootaxa, 3626 (1), 99 - 145. https: // doi. org / 10.11646 / zootaxa. 3626.1.5" type="journal article" year="2013">Joshi & Edgecombe 2013: 101</bibRefCitation>
|
||
). Analyzing the monophyly of
|
||
<taxonomicName id="4C77C3B8FFDFD727FC8AFF3E29DFDCE6" authority="Vahtera et al. (2013: 595)" authorityName="Vahtera" authorityPageNumber="595" authorityYear="2013" box="[782,1178,222,249]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFDFD727FC8AFF3E2E30DCE7" box="[782,885,223,248]" italics="true" pageId="45" pageNumber="46">Digitipes</emphasis>
|
||
<bibRefCitation id="EFE6C5CAFFDFD727FCFFFF3E29DFDCE6" author="Vahtera, V. & Edgecombe, G. D. & Giribet, G." box="[891,1178,222,249]" pageId="45" pageNumber="46" pagination="578 - 602" refId="ref51397" refString="Vahtera, V., Edgecombe, G. D. & Giribet, G. (2013). Phylogenetics of scolopendromorph centipedes: can denser taxon sampling improve an artificial classification? Invertebrate Systematics, 27 (5), 578 - 602. https: // doi. org / 10.1071 / IS 13035" type="journal article" year="2013">
|
||
Vahtera
|
||
<emphasis id="B9036429FFDFD727FC5CFF3E294CDCE6" box="[984,1033,223,249]" italics="true" pageId="45" pageNumber="46">et al</emphasis>
|
||
. (2013: 595)
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
wrote: “… the clade is sensitive to parameter set variation as it is resolved as monophyletic only under two of the six explored parameter sets”, although the data of Siriwut
|
||
<emphasis id="B9036429FFDFD727FD9BFEC62F16DD5E" box="[543,595,295,321]" italics="true" pageId="45" pageNumber="46">at al</emphasis>
|
||
. (2018) and
|
||
<bibRefCitation id="EFE6C5CAFFDFD727FD60FEC62EF5DD5E" author="Joshi, J. & Karanth, P. K. & Edgecombe, G. D." box="[740,944,295,321]" pageId="45" pageNumber="46" pagination="828 - 861" refId="ref48095" refString="Joshi, J., Karanth, P. K. & Edgecombe, G. D. (2020) The Out-of-India hypothesis: Evidence from an ancient centipede genus, Rhysida (Chilopoda: Scolopendromorpha) from the Oriental Region, and systematics of Indian species. Zoological Journal of the Linnean Society, 189, 828 - 861." type="journal article" year="2020">
|
||
Joshi
|
||
<emphasis id="B9036429FFDFD727FCA1FEC62E12DD5E" box="[805,855,295,321]" italics="true" pageId="45" pageNumber="46">et al</emphasis>
|
||
. (2020)
|
||
</bibRefCitation>
|
||
strengthened the case for monophyly using more species.
|
||
</paragraph>
|
||
<paragraph id="8BC8B83BFFDFD727FF43FE8E294EDE22" blockId="45.[151,1437,151,573]" pageId="45" pageNumber="46">
|
||
All diagnostic features of
|
||
<taxonomicName id="4C77C3B8FFDFD727FE6CFE8E2F0ADD97" box="[488,591,367,392]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFDFD727FE6CFE8E2F0ADD97" box="[488,591,367,392]" italics="true" pageId="45" pageNumber="46">Digitipes</emphasis>
|
||
</taxonomicName>
|
||
listed by
|
||
<bibRefCitation id="EFE6C5CAFFDFD727FD3FFE8E295ADD96" author="Joshi, J. & Edgecombe, G. D." box="[699,1055,367,393]" pageId="45" pageNumber="46" pagination="99 - 145" refId="ref47850" refString="Joshi, J. & Edgecombe, G. D. (2013) Revision of the scolopendrid centipede Digitipes Attems, 1930, from India (Chilopoda: Scolopendromorpha): reconciling molecular and morphological estimates of species diversity. Zootaxa, 3626 (1), 99 - 145. https: // doi. org / 10.11646 / zootaxa. 3626.1.5" type="journal article" year="2013">Joshi & Edgecombe (2013:100)</bibRefCitation>
|
||
except for the male’s longitudinal groove and distomedial process of the ultimate femur are shared by all representatives of the nominotypical subgenus of
|
||
<taxonomicName id="4C77C3B8FFDFD727FF66FE562C27DDCE" box="[226,354,439,465]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFDFD727FF66FE562C27DDCE" box="[226,354,439,465]" italics="true" pageId="45" pageNumber="46">Otostigmus</emphasis>
|
||
</taxonomicName>
|
||
. The “composite description” of
|
||
<taxonomicName id="4C77C3B8FFDFD727FD55FE562E7DDDCF" box="[721,824,439,464]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFDFD727FD55FE562E7DDDCF" box="[721,824,439,464]" italics="true" pageId="45" pageNumber="46">Digitipes</emphasis>
|
||
</taxonomicName>
|
||
provided by Siriwut
|
||
<emphasis id="B9036429FFDFD727FBA6FE562911DDCE" box="[1058,1108,439,465]" italics="true" pageId="45" pageNumber="46">at al</emphasis>
|
||
. (2015: 75) also perfectly fits for
|
||
<taxonomicName id="4C77C3B8FFDFD727FF3BFE3A2C9FDDEA" authority="(Otostigmus)" authorityName="Porat" authorityYear="1876" baseAuthorityName="Otostigmus" box="[191,474,475,501]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="45" pageNumber="46" phylum="Arthropoda" rank="subGenus" subGenus="Otostigmus">
|
||
<emphasis id="B9036429FFDFD727FF3BFE3A2C7ADDEA" box="[191,319,475,501]" italics="true" pageId="45" pageNumber="46">Otostigmus</emphasis>
|
||
(
|
||
<emphasis id="B9036429FFDFD727FED4FE3A2C95DDEA" box="[336,464,475,501]" italics="true" pageId="45" pageNumber="46">Otostigmus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
, apart from the aforementioned male characters. Taking into consideration the virtual morphological identity of these two genera, and the fact that the monophyly of
|
||
<taxonomicName id="4C77C3B8FFDFD727FB98FE1E29C6DE07" box="[1052,1155,511,536]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFDFD727FB98FE1E29C6DE07" box="[1052,1155,511,536]" italics="true" pageId="45" pageNumber="46">Digitipes</emphasis>
|
||
</taxonomicName>
|
||
is not well supported by the morphological data, we consider this genus to require further investigation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |