201 lines
37 KiB
XML
201 lines
37 KiB
XML
<document id="88AD06299DEC73814D239FBE561B372B" ID-CLB-Dataset="8703" ID-DOI="10.11646/zootaxa.4825.1.1" ID-GBIF-Dataset="5ab5f5c8-481e-4d1a-8643-21e72c367278" ID-ISSN="1175-5326" ID-Zenodo-Dep="4402145" ID-ZooBank="F230F199-1C94-4E2E-9CE4-5F56212C015F" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1609341789210" checkinUser="plazi" docAuthor="Schileyko, Arkady A., Vahtera, Varpu & Edgecombe, Gregory D." docDate="2020" docId="03DE092DFFDAD723FF13F9432DA3DE9A" docLanguage="en" docName="zootaxa.4825.1.1.pdf" docOrigin="Zootaxa 4825 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Otostigmini Kraepelin 1903" docType="treatment" docVersion="7" lastPageNumber="42" masterDocId="FFE77155FFF2D70AFF84FFE12D45DC1F" masterDocTitle="An overview of the extant genera and subgenera of the order Scolopendromorpha (Chilopoda): a new identification key and updated diagnoses" masterLastPageNumber="64" masterPageNumber="1" pageNumber="41" updateTime="1698852004925" updateUser="ExternalLinkService">
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<mods:mods id="1AB2D0405830274E254602D76740D2AE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="793F5CBF121CE2D9FCA483DE9E6451E2">
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<mods:title id="47B5DA361F59330FFE454402CC47341B">An overview of the extant genera and subgenera of the order Scolopendromorpha (Chilopoda): a new identification key and updated diagnoses</mods:title>
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</mods:titleInfo>
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<mods:name id="9B5C8BA46B8E4989D6E16167EEFA25E1" type="personal">
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<mods:roleTerm id="1FC394235DD91AEA04F00A7AD0A69123">Author</mods:roleTerm>
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<mods:namePart id="80F96622AE9351B860A7546146C470C9">Schileyko, Arkady A.</mods:namePart>
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<mods:nameIdentifier id="E2914335325792FD59002CC64E33C02C" type="email">schileyko1965@gmail.com</mods:nameIdentifier>
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<mods:name id="C2F226D422E8BD9DF8852A2CBC59B639" type="personal">
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<mods:roleTerm id="D009CBAF19F8F2090493525BB18678B5">Author</mods:roleTerm>
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<mods:namePart id="4874B827ED03E5CDF92B0952CA03522D">Vahtera, Varpu</mods:namePart>
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<mods:nameIdentifier id="C7C906CF7B7FFCCA7D2081BC827365B4" type="email">varpu.vahtera@gmail.com</mods:nameIdentifier>
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<mods:name id="B24E3D417BD0EDC25439240E9807A986" type="personal">
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<mods:namePart id="1FF6BEBC0D1D0E92B71588B2258A77C9">Edgecombe, Gregory D.</mods:namePart>
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<mods:nameIdentifier id="25E222292DC282727731B194E7321211" type="ORCID">0000-0002-9591-8011</mods:nameIdentifier>
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<mods:nameIdentifier id="D5C44F3DA084E98CC2C15CAAD8513ACB" type="email">schileyko1965@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="876B81393241D38AA5598088435A3511">text</mods:typeOfResource>
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<mods:title id="7F7D12F8853029EE5B9D04F6D72BA99D">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="7BB1F9CE262B6115B801A54585C5A94F">
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<mods:date id="755D7AD30A05341CF8AE54B189E63FFC">2020</mods:date>
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<mods:detail id="9F636B45226DB1A7861C39022A969A28" type="pubDate">
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<mods:number id="05375C4D89427C7C808737934B5E4963">2020-08-10</mods:number>
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<mods:detail id="13A1E5B4E358BBDD19D1A898688300DC" type="volume">
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<mods:number id="D78BB72D56AB613C3E26D9DFE124C266">4825</mods:number>
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<mods:classification id="D045BBCE4AB8E906CCE07AE0F079D310">journal article</mods:classification>
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<mods:identifier id="58B18F7B1781ABF4C963103BF77F94B5" type="CLB-Dataset">8703</mods:identifier>
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<mods:identifier id="C9654B22E290A44F2B993A1B246743C1" type="DOI">10.11646/zootaxa.4825.1.1</mods:identifier>
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<treatment id="03DE092DFFDAD723FF13F9432DA3DE9A" ID-DOI="http://doi.org/10.5281/zenodo.4455423" ID-GBIF-Taxon="177020729" ID-Zenodo-Dep="4455423" LSID="urn:lsid:plazi:treatment:03DE092DFFDAD723FF13F9432DA3DE9A" httpUri="http://treatment.plazi.org/id/03DE092DFFDAD723FF13F9432DA3DE9A" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
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<subSubSection id="C36DEBB0FFDAD722FF13F9432F00DAA3" box="[151,581,1698,1724]" pageId="40" pageNumber="41" type="nomenclature">
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<paragraph id="8BC8B83BFFDAD722FF13F9432F00DAA3" blockId="40.[151,581,1698,1724]" box="[151,581,1698,1724]" pageId="40" pageNumber="41">
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<heading id="D0800F57FFDAD722FF13F9432F00DAA3" bold="true" box="[151,581,1698,1724]" fontSize="11" level="2" pageId="40" pageNumber="41" reason="3">
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<emphasis id="B9036429FFDAD722FF13F9432F00DAA3" bold="true" box="[151,581,1698,1724]" pageId="40" pageNumber="41">
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Tribe
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<taxonomicName id="4C77C3B8FFDAD722FF66F9432F00DAA3" authority="Kraepelin, 1903" authorityName="Kraepelin" authorityYear="1903" box="[226,581,1698,1724]" class="Chilopoda" family="Scolopendridae" kingdom="Animalia" order="Scolopendromorpha" pageId="40" pageNumber="41" phylum="Arthropoda" rank="tribe" tribe="Otostigmini">
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Otostigmini
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<bibRefCitation id="EFE6C5CAFFDAD722FEFAF9432F00DAA3" author="Kraepelin, K." box="[382,581,1698,1724]" pageId="40" pageNumber="41" pagination="1 - 276" refId="ref48450" refString="Kraepelin, K. (1903) Revision der Scolopendriden. Mitteilungen aus dem Naturhistorischen Museum Hamburg, 20 (2), 1 - 276." type="journal article" year="1903">Kraepelin, 1903</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36DEBB0FFDAD722FF13F9062F5BDAE0" box="[151,542,1767,1791]" pageId="40" pageNumber="41" type="synonymic_list">
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<paragraph id="8BC8B83BFFDAD722FF13F9062F5BDAE0" blockId="40.[151,542,1767,1791]" box="[151,542,1767,1791]" pageId="40" pageNumber="41">
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Synonyms.
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<taxonomicName id="4C77C3B8FFDAD722FE89F9062F5BDAE0" authority="Attems, 1930" authorityName="Attems" authorityYear="1930" box="[269,542,1767,1791]" class="Chilopoda" family="Scolopendridae" kingdom="Animalia" order="Scolopendromorpha" pageId="40" pageNumber="53" phylum="Arthropoda" rank="tribe" tribe="Arrhabdotini">
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Arrhabdotini
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<bibRefCitation id="EFE6C5CAFFDAD722FE11F9062F5BDAE0" author="Attems, C." box="[405,542,1767,1791]" pageId="40" pageNumber="41" refId="ref45522" refString="Attems, C. (1930) Myriapoda 2. Scolopendromorpha. Das Tierreich. Vol. 54. Walter de Gruyter et Co. Publ., Berlin, 308 pp." type="book" year="1930">Attems, 1930</bibRefCitation>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36DEBB0FFDAD723FF13F8CC2F8ADD7A" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41" type="diagnosis">
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<paragraph id="8BC8B83BFFDAD723FF13F8CC2F8ADD7A" blockId="40.[151,1437,1837,2044]" lastBlockId="41.[151,1437,151,645]" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
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<emphasis id="B9036429FFDAD722FF13F8CC2C57DB58" bold="true" box="[151,274,1837,1863]" pageId="40" pageNumber="41">Diagnosis.</emphasis>
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Antenna relatively long (i.e. reaching at least to posterior margin of tergite 4 when reflexed), basal antennal articles cylindrical (not strongly flattened), definitely longer than wide. Median tooth of labrum (
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<figureCitation id="134CA4BEFFDAD722FA8EF8B02819DB73" box="[1290,1372,1873,1900]" captionStart="FIGURES 89–96" captionStartId="47.[151,264,1723,1747]" captionTargetBox="[181,1399,207,1674]" captionTargetId="figure-17@47.[166,1421,181,1700]" captionTargetPageId="47" captionText="FIGURES 89–96. Digitipes coonoorensis Jangi & Dass, 1984; Male CES 091088 89 femur of right ultimate leg medially; Digitipes sp.; Ad. Rc 7565 90 clypeal part of head ventrally 91 forcipular segment ventrally 92 LBS 13–15 ventrally; Otostigmus (O.) multidens nordicus Schileyko, 1995; Ad. Rc 6545 93 LBS 21 ventrally; Ad. Rc 6747 94 anterior part of head ventrally; Otostigmus (O.) astenus (Kohlrausch, 1881); MCZ DNA 102463 95 pretarsus of maxilla 2; Sad. Rc 7507 96 forcipular segment ventrally; (ap)—apical spines of coxopleural process, (as)—accessory spines, (bs)—basal suture of forcipular toothplate, (cl)—clypeus, (cx)—coxopleuron, (df)—conical distomedial process of ultimate femur, (f)—ultimate femur, (l)—labrum, (lc)—lateral spine of coxopleural process, (lsc)—sternal median longitudinal sulcus, (md)—medial longitudinal depression of ultimate femur, (p2)—pretarsus of maxilla 2, (pd)—sternal posterior depression, (pr)—trochantero-prefemoral process, (psl)—sternal paramedian sulci, (tp)—forcipular tooth-plate, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402185" httpUri="https://zenodo.org/record/4402185/files/figure.png" pageId="40" pageNumber="41">Fig. 95</figureCitation>
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) well developed (except for
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<taxonomicName id="4C77C3B8FFDAD722FE12F8942F67DB90" authorityName="Tomosvary" authorityYear="1882" box="[406,546,1909,1935]" class="Chilopoda" family="Scolopendridae" genus="Edentistoma" kingdom="Animalia" order="Scolopendromorpha" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
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<emphasis id="B9036429FFDAD722FE12F8942F67DB90" box="[406,546,1909,1935]" italics="true" pageId="40" pageNumber="41">Edentistoma</emphasis>
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</taxonomicName>
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, former tribe
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<taxonomicName id="4C77C3B8FFDAD722FD3BF8942E16DB90" authorityName="Attems" authorityYear="1930" box="[703,851,1909,1935]" class="Chilopoda" family="Scolopendridae" kingdom="Animalia" order="Scolopendromorpha" pageId="40" pageNumber="41" phylum="Arthropoda" rank="tribe" tribe="Arrhabdotini">Arrhabdotini</taxonomicName>
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; see below). Forcipular tooth-plates present (except for
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<taxonomicName id="4C77C3B8FFDAD722FF3BF8782C0EDBAC" authorityName="Tomosvary" authorityYear="1882" box="[191,331,1945,1971]" class="Chilopoda" family="Scolopendridae" genus="Edentistoma" kingdom="Animalia" order="Scolopendromorpha" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
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<emphasis id="B9036429FFDAD722FF3BF8782C0EDBAC" box="[191,331,1945,1971]" italics="true" pageId="40" pageNumber="41">Edentistoma</emphasis>
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</taxonomicName>
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), their basal sutures form a virtually straight line (
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<figureCitation id="134CA4BEFFDAD722FCFBF8782E97DBAB" box="[895,978,1945,1972]" captionStart="FIGURES 89–96" captionStartId="47.[151,264,1723,1747]" captionTargetBox="[181,1399,207,1674]" captionTargetId="figure-17@47.[166,1421,181,1700]" captionTargetPageId="47" captionText="FIGURES 89–96. Digitipes coonoorensis Jangi & Dass, 1984; Male CES 091088 89 femur of right ultimate leg medially; Digitipes sp.; Ad. Rc 7565 90 clypeal part of head ventrally 91 forcipular segment ventrally 92 LBS 13–15 ventrally; Otostigmus (O.) multidens nordicus Schileyko, 1995; Ad. Rc 6545 93 LBS 21 ventrally; Ad. Rc 6747 94 anterior part of head ventrally; Otostigmus (O.) astenus (Kohlrausch, 1881); MCZ DNA 102463 95 pretarsus of maxilla 2; Sad. Rc 7507 96 forcipular segment ventrally; (ap)—apical spines of coxopleural process, (as)—accessory spines, (bs)—basal suture of forcipular toothplate, (cl)—clypeus, (cx)—coxopleuron, (df)—conical distomedial process of ultimate femur, (f)—ultimate femur, (l)—labrum, (lc)—lateral spine of coxopleural process, (lsc)—sternal median longitudinal sulcus, (md)—medial longitudinal depression of ultimate femur, (p2)—pretarsus of maxilla 2, (pd)—sternal posterior depression, (pr)—trochantero-prefemoral process, (psl)—sternal paramedian sulci, (tp)—forcipular tooth-plate, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402185" httpUri="https://zenodo.org/record/4402185/files/figure.png" pageId="40" pageNumber="41">Fig. 96</figureCitation>
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), right angle (
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<figureCitation id="134CA4BEFFDAD722FBEBF8782985DBAC" box="[1135,1216,1945,1971]" captionStart="FIGURES 89–96" captionStartId="47.[151,264,1723,1747]" captionTargetBox="[181,1399,207,1674]" captionTargetId="figure-17@47.[166,1421,181,1700]" captionTargetPageId="47" captionText="FIGURES 89–96. Digitipes coonoorensis Jangi & Dass, 1984; Male CES 091088 89 femur of right ultimate leg medially; Digitipes sp.; Ad. Rc 7565 90 clypeal part of head ventrally 91 forcipular segment ventrally 92 LBS 13–15 ventrally; Otostigmus (O.) multidens nordicus Schileyko, 1995; Ad. Rc 6545 93 LBS 21 ventrally; Ad. Rc 6747 94 anterior part of head ventrally; Otostigmus (O.) astenus (Kohlrausch, 1881); MCZ DNA 102463 95 pretarsus of maxilla 2; Sad. Rc 7507 96 forcipular segment ventrally; (ap)—apical spines of coxopleural process, (as)—accessory spines, (bs)—basal suture of forcipular toothplate, (cl)—clypeus, (cx)—coxopleuron, (df)—conical distomedial process of ultimate femur, (f)—ultimate femur, (l)—labrum, (lc)—lateral spine of coxopleural process, (lsc)—sternal median longitudinal sulcus, (md)—medial longitudinal depression of ultimate femur, (p2)—pretarsus of maxilla 2, (pd)—sternal posterior depression, (pr)—trochantero-prefemoral process, (psl)—sternal paramedian sulci, (tp)—forcipular tooth-plate, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402185" httpUri="https://zenodo.org/record/4402185/files/figure.png" pageId="40" pageNumber="41">Fig. 91</figureCitation>
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) or an obtuse angle (
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<figureCitation id="134CA4BEFFDAD722FF24F85C2DB4DBC8" box="[160,241,1981,2007]" captionStart="FIGURES 84–88" captionStartId="46.[151,264,1784,1808]" captionTargetBox="[186,1400,200,1740]" captionTargetId="figure-19@46.[167,1420,181,1761]" captionTargetPageId="46" captionText="FIGURES 84–88. Alipes multicostis Imhoff, 1854; Live specimen CDT (photo by Dmitry Telnov) 84 general view dorsally; Alipes grandidieri Lucas, 1864; BMNH 1959.3.8 85 distal part of ultimate leg laterally; Alipes appendiculatus Pocock, 1896; BMNH1950.3.9.2 86 forcipular segment ventrally 87 LBS 4–10 dorsally; Alipes grandidieri Lucas, 1864; BMNH1959.3.8 88 LBS 21 + prefemora of ultimate legs ventrally; (bs)—basal suture of forcipular tooth-plate, (cp)—very short spineless coxopleural process, (cx)—coxopleuron, (pf)—ultimate prefemur, (pr)—trochantero-prefemoral process, (t1)—flattened tarsus 1, (t2)—flattened tarsus 2, (T5)—terite 5, (ti)—flattened tibia, (tk)—tergal longitudinal keels, (tp)—forcipular tooth-plate, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402183" httpUri="https://zenodo.org/record/4402183/files/figure.png" pageId="40" pageNumber="41">Fig. 86</figureCitation>
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,
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<figureCitation id="134CA4BEFFDAD722FF79F85C2C6CDBC8" box="[253,297,1981,2007]" captionStart="FIGURES 97–101" captionStartId="48.[151,264,1693,1717]" captionTargetBox="[171,1417,200,1647]" captionTargetId="figure-19@48.[151,1436,181,1669]" captionTargetPageId="48" captionText="FIGURES 97–101. Otostigmus (Parotostigmus) scabricauda (Humbert et Saussure, 1870); Ad. male Rc 6730 97 LBS 21 and prefemora + femora of ultimate legs ventrally; Otostigmus (P.) salticus Schileyko, Iorio & Coulis, 2018; Ad. holotype male MNHN 460 98 posterior margin of ultimate tergite + prefemora of ultimate legs dorsally; Otostigmus (P.) diringshofeni Bücherl, 1969; Ad. male Rc 7981 99 LBS 21 + prefemora of ultimate legs ventrally; Otostigmus (P.) salticus Schileyko, Iorio & Coulis, 2018; Ad. holotype male MNHN 460 100 anterior margin of forcipular coxosternite + maxillae 2 ventrally; Ethmostigmus rubripes spinosus (Newport, 1845); Juv. Rc 7174 101 left side of ultimate LBS + ultimate legs laterally; (as)—accessory spine, (bs)—basal suture of forcipular tooth-plate, (cp)— practically reduced spineless coxopleural process, (csp)—corner spine of ultimate prefemur, (cx)—coxopleuron, (dg)—digitiform process of ultimate prefemur, (p2)—pretarsus of maxilla 2, (pf)—ultimate prefemur, (pr)—trochantero-prefemoral process, (pst)—penultimate sternite, (pts)—ultimate pretarsus, (su)—ventral spines of ultimate prefemur, (tp)—forcipular tooth-plate, (ut)—ultimate tergite." figureDoi="http://doi.org/10.5281/zenodo.4402189" httpUri="https://zenodo.org/record/4402189/files/figure.png" pageId="40" pageNumber="41">100</figureCitation>
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,
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<figureCitation id="134CA4BEFFDAD722FEB1F85C2C26DBC7" box="[309,355,1981,2008]" captionStart="FIGURES 102–107" captionStartId="50.[151,264,1685,1709]" captionTargetBox="[274,1314,198,1647]" captionTargetId="figure-19@50.[257,1331,181,1661]" captionTargetPageId="50" captionText="FIGURES 102–107. Ethmostigmus rubripes spinosus (Newport, 1845); Juv. Rc 7174 102 forcipular segment ventrally; Ethmostigmus rubripes platycephalus (Newport, 1845); Ad. Rc 7514 103 left pleuron of LBS 3 laterally; Ad. Rc 7513 104 LBS 21 ventrally; Sad. СDT 105 coxopleural processes + prefemora of ultimate legs ventrally; Rhysida longipes (Newport, 1845); Ad. Rc 6683 106 anterior margin of forcipular coxosternite ventrally 107 coxopleural processes, legs 20 + ultimate legs laterally; (ap)—apical spine, (bs)—basal suture of forcipular tooth-plate, (cp)—coxopleural process, (csp)—corner spine of ultimate prefemur, (cx)—coxopleuron, (im)—intersclerite membrane, (lc)—lateral spines of coxopleural process, (pf)—ultimate prefemur, (pr)—trochantero-prefemoral process, (pts)—ultimate pretarsus, (rpf)—abnormally small (regenerated) right ultimate prefemur, (rpr)—rudiment of trochantero-prefemoral process, (s)—open spiracle without atrium, (scs)— spine at place of corner spine of ultimate prefemur, (su)—spines of ultimate prefemur, (t1)—ultimate tarsus 1, (tp)—forcipular tooth-plates, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402191" httpUri="https://zenodo.org/record/4402191/files/figure.png" pageId="40" pageNumber="41">106</figureCitation>
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); trochantero-prefemoral process present in most genera. Tarsungula long in most of genera (
|
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<figureCitation id="134CA4BEFFDAD722FAEBF85C2DF3DBE3" captionStart="FIGURES 89–96" captionStartId="47.[151,264,1723,1747]" captionTargetBox="[181,1399,207,1674]" captionTargetId="figure-17@47.[166,1421,181,1700]" captionTargetPageId="47" captionText="FIGURES 89–96. Digitipes coonoorensis Jangi & Dass, 1984; Male CES 091088 89 femur of right ultimate leg medially; Digitipes sp.; Ad. Rc 7565 90 clypeal part of head ventrally 91 forcipular segment ventrally 92 LBS 13–15 ventrally; Otostigmus (O.) multidens nordicus Schileyko, 1995; Ad. Rc 6545 93 LBS 21 ventrally; Ad. Rc 6747 94 anterior part of head ventrally; Otostigmus (O.) astenus (Kohlrausch, 1881); MCZ DNA 102463 95 pretarsus of maxilla 2; Sad. Rc 7507 96 forcipular segment ventrally; (ap)—apical spines of coxopleural process, (as)—accessory spines, (bs)—basal suture of forcipular toothplate, (cl)—clypeus, (cx)—coxopleuron, (df)—conical distomedial process of ultimate femur, (f)—ultimate femur, (l)—labrum, (lc)—lateral spine of coxopleural process, (lsc)—sternal median longitudinal sulcus, (md)—medial longitudinal depression of ultimate femur, (p2)—pretarsus of maxilla 2, (pd)—sternal posterior depression, (pr)—trochantero-prefemoral process, (psl)—sternal paramedian sulci, (tp)—forcipular tooth-plate, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402185" httpUri="https://zenodo.org/record/4402185/files/figure.png" pageId="40" pageNumber="41">Fig. 96</figureCitation>
|
||
), much elongated in
|
||
<taxonomicName id="4C77C3B8FFDAD722FE21F8002F74DBE4" authorityName="Tomosvary" authorityYear="1882" box="[421,561,2017,2043]" class="Chilopoda" family="Scolopendridae" genus="Edentistoma" kingdom="Animalia" order="Scolopendromorpha" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFDAD722FE21F8002F74DBE4" box="[421,561,2017,2043]" italics="true" pageId="40" pageNumber="41">Edentistoma</emphasis>
|
||
</taxonomicName>
|
||
and visibly shortened in
|
||
<taxonomicName id="4C77C3B8FFDAD722FCD5F8002EA4DBE4" authorityName="Attems" authorityYear="1934" box="[849,993,2017,2043]" class="Chilopoda" family="Scolopendridae" genus="Sterropristes" kingdom="Animalia" order="Scolopendromorpha" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFDAD722FCD5F8002EA4DBE4" box="[849,993,2017,2043]" italics="true" pageId="40" pageNumber="41">Sterropristes</emphasis>
|
||
</taxonomicName>
|
||
(compare
|
||
<figureCitation id="134CA4BEFFDAD722FBDFF80029FDDBE4" box="[1115,1208,2017,2043]" captionStart="FIGURES 113–118" captionStartId="54.[151,264,1746,1770]" captionTargetBox="[167,1419,198,1704]" captionTargetId="figure-19@54.[151,1436,181,1722]" captionTargetPageId="54" captionText="FIGURES 113–118. Edentistoma octosulcatum Tömösváry, 1882; Voucher ZMUT_Chi01 113 left ultimate leg laterally; ZMB 3618 114 forcipular segment ventrally; Voucher ZMUT_Chi01 115 right pleuron of LBS 7 laterally 116 LBS 13–21+ right ultimate leg dorsally 117 LBS 20–21+ right ultimate leg ventrally 118 left ultimate leg medially; (20)—leg 20, (bl)—low blade-like lobe in place of forcipular tooth-plate, (bs)—basal suture of blade-like lobe, (cp)—spineless coxopleural process, (csp)—spineless “corner spine(s)” of ultimate prefemur, (cx)—coxopleuron, (im)—intersclerite membrane, (lsc)—sternal median longitudinal sulcus, (pf)—relatively short ultimate prefemur, (pts)—pretarsus, (rpr)—rudiment of trochantero-prefemoral process, (s)—open spiracle without atrium, (sh)—spiracle hump, (t1)—tarsus 1, (t2)—tarsus 2, (tk)—tergal longitudinal keels, (tr)—visibly elongated tarsungula, (ust)—ultimate sternite, (ut)—ultimate tergite." figureDoi="http://doi.org/10.5281/zenodo.4402195" httpUri="https://zenodo.org/record/4402195/files/figure.png" pageId="40" pageNumber="41">Fig. 114</figureCitation>
|
||
and Fig. 120); their inner surface rounded or with 1 or 2 sharp longitudinal ridges, rarely saw-like (in
|
||
<taxonomicName id="4C77C3B8FFDBD723FBA5FF7629F4DCAE" authorityName="Attems" authorityYear="1934" box="[1057,1201,151,177]" class="Chilopoda" family="Scolopendridae" genus="Sterropristes" kingdom="Animalia" order="Scolopendromorpha" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFDBD723FBA5FF7629F4DCAE" box="[1057,1201,151,177]" italics="true" pageId="41" pageNumber="42">Sterropristes</emphasis>
|
||
</taxonomicName>
|
||
only). LBS 7 with or without spiracles. Pleuron (
|
||
<figureCitation id="134CA4BEFFDBD723FE4DFF5A2F5ADCCA" box="[457,543,187,213]" captionStart="FIGURES 78–83" captionStartId="42.[151,264,1708,1732]" captionTargetBox="[168,1421,193,1665]" captionTargetId="figure-19@42.[151,1436,181,1684]" captionTargetPageId="42" captionText="FIGURES 78–83. Asanada sokotrana Pocock, 1899; BMNH 1934.2.15.151–154 78 LBS 21 ventrally; Asanada indica Jangi & Dass, 1984; Smaller ad. Rc 7484 79 head + LBS 1 dorsally; Asanada brevicornis Meinert, 1886; BMNH 1889.7.15.21–23 80 apical part of maxilla 2 dorso-laterally; Asanada sokotrana Pocock, 1899; BMNH 1899.7.4.85–86 81 LBS 20–21 and ultimate legs dorsally; Otostigmus (O.) multidens Haase, 1887; Ad. Rc 7510 82 right pleuron of LBS 3 laterally; Otostigmus (O.) multidens nordicus Schileyko, 1995; Ad. Rc 6747 83 left side of ultimate LBS + right ultimate leg medially; (an)—antenna, (cx)—coxopleuron, (f)—ultimate femur, (im)—intersclerite membrane, (p2)—pretarsus of maxilla 2, (pf)—ultimate prefemur, (pts)—non enlarged ultimate pretarsus, (s)—open spiracle with atrium, (su)—spines of ultimate prefemur, (T1)—tergite 1, (t1)—tarsus 1, (t2)—tarsus 2, (ti)—tibia, (ust)—ultimate sternite, (ut)—ultimate tergite." figureDoi="http://doi.org/10.5281/zenodo.4402181" httpUri="https://zenodo.org/record/4402181/files/figure.png" pageId="41" pageNumber="42">Figs 82</figureCitation>
|
||
,
|
||
<figureCitation id="134CA4BEFFDBD723FDAFFF5A2F13DCCA" box="[555,598,187,213]" captionStart="FIGURES 102–107" captionStartId="50.[151,264,1685,1709]" captionTargetBox="[274,1314,198,1647]" captionTargetId="figure-19@50.[257,1331,181,1661]" captionTargetPageId="50" captionText="FIGURES 102–107. Ethmostigmus rubripes spinosus (Newport, 1845); Juv. Rc 7174 102 forcipular segment ventrally; Ethmostigmus rubripes platycephalus (Newport, 1845); Ad. Rc 7514 103 left pleuron of LBS 3 laterally; Ad. Rc 7513 104 LBS 21 ventrally; Sad. СDT 105 coxopleural processes + prefemora of ultimate legs ventrally; Rhysida longipes (Newport, 1845); Ad. Rc 6683 106 anterior margin of forcipular coxosternite ventrally 107 coxopleural processes, legs 20 + ultimate legs laterally; (ap)—apical spine, (bs)—basal suture of forcipular tooth-plate, (cp)—coxopleural process, (csp)—corner spine of ultimate prefemur, (cx)—coxopleuron, (im)—intersclerite membrane, (lc)—lateral spines of coxopleural process, (pf)—ultimate prefemur, (pr)—trochantero-prefemoral process, (pts)—ultimate pretarsus, (rpf)—abnormally small (regenerated) right ultimate prefemur, (rpr)—rudiment of trochantero-prefemoral process, (s)—open spiracle without atrium, (scs)— spine at place of corner spine of ultimate prefemur, (su)—spines of ultimate prefemur, (t1)—ultimate tarsus 1, (tp)—forcipular tooth-plates, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402191" httpUri="https://zenodo.org/record/4402191/files/figure.png" pageId="41" pageNumber="42">103</figureCitation>
|
||
) with intersclerite membrane well visible, without set of longitudinal pleurites anteriorly. Coxopleuron with (
|
||
<figureCitation id="134CA4BEFFDBD723FDA0FF3E2FF9DCE6" box="[548,700,222,249]" captionStart="FIGURES 102–107" captionStartId="50.[151,264,1685,1709]" captionTargetBox="[274,1314,198,1647]" captionTargetId="figure-19@50.[257,1331,181,1661]" captionTargetPageId="50" captionText="FIGURES 102–107. Ethmostigmus rubripes spinosus (Newport, 1845); Juv. Rc 7174 102 forcipular segment ventrally; Ethmostigmus rubripes platycephalus (Newport, 1845); Ad. Rc 7514 103 left pleuron of LBS 3 laterally; Ad. Rc 7513 104 LBS 21 ventrally; Sad. СDT 105 coxopleural processes + prefemora of ultimate legs ventrally; Rhysida longipes (Newport, 1845); Ad. Rc 6683 106 anterior margin of forcipular coxosternite ventrally 107 coxopleural processes, legs 20 + ultimate legs laterally; (ap)—apical spine, (bs)—basal suture of forcipular tooth-plate, (cp)—coxopleural process, (csp)—corner spine of ultimate prefemur, (cx)—coxopleuron, (im)—intersclerite membrane, (lc)—lateral spines of coxopleural process, (pf)—ultimate prefemur, (pr)—trochantero-prefemoral process, (pts)—ultimate pretarsus, (rpf)—abnormally small (regenerated) right ultimate prefemur, (rpr)—rudiment of trochantero-prefemoral process, (s)—open spiracle without atrium, (scs)— spine at place of corner spine of ultimate prefemur, (su)—spines of ultimate prefemur, (t1)—ultimate tarsus 1, (tp)—forcipular tooth-plates, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402191" httpUri="https://zenodo.org/record/4402191/files/figure.png" pageId="41" pageNumber="42">Fig. 104, 105</figureCitation>
|
||
) or without (in
|
||
<taxonomicName id="4C77C3B8FFDBD723FCEBFF3E29EEDCE6" authority="" authorityName="Pocock" authorityYear="1896" box="[879,1195,223,249]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="41" pageNumber="42" phylum="Arthropoda" rank="subGenus" subGenus="Parotostigmus">
|
||
<emphasis id="B9036429FFDBD723FCEBFF3E2EAADCE6" box="[879,1007,223,249]" italics="true" pageId="41" pageNumber="42">Otostigmus</emphasis>
|
||
(
|
||
<emphasis id="B9036429FFDBD723FB84FF3E29E7DCE7" box="[1024,1186,223,248]" italics="true" pageId="41" pageNumber="42">Parotostigmus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="134CA4BEFFDBD723FB3FFF3E2848DCE6" box="[1211,1293,223,249]" captionStart="FIGURES 97–101" captionStartId="48.[151,264,1693,1717]" captionTargetBox="[171,1417,200,1647]" captionTargetId="figure-19@48.[151,1436,181,1669]" captionTargetPageId="48" captionText="FIGURES 97–101. Otostigmus (Parotostigmus) scabricauda (Humbert et Saussure, 1870); Ad. male Rc 6730 97 LBS 21 and prefemora + femora of ultimate legs ventrally; Otostigmus (P.) salticus Schileyko, Iorio & Coulis, 2018; Ad. holotype male MNHN 460 98 posterior margin of ultimate tergite + prefemora of ultimate legs dorsally; Otostigmus (P.) diringshofeni Bücherl, 1969; Ad. male Rc 7981 99 LBS 21 + prefemora of ultimate legs ventrally; Otostigmus (P.) salticus Schileyko, Iorio & Coulis, 2018; Ad. holotype male MNHN 460 100 anterior margin of forcipular coxosternite + maxillae 2 ventrally; Ethmostigmus rubripes spinosus (Newport, 1845); Juv. Rc 7174 101 left side of ultimate LBS + ultimate legs laterally; (as)—accessory spine, (bs)—basal suture of forcipular tooth-plate, (cp)— practically reduced spineless coxopleural process, (csp)—corner spine of ultimate prefemur, (cx)—coxopleuron, (dg)—digitiform process of ultimate prefemur, (p2)—pretarsus of maxilla 2, (pf)—ultimate prefemur, (pr)—trochantero-prefemoral process, (pst)—penultimate sternite, (pts)—ultimate pretarsus, (su)—ventral spines of ultimate prefemur, (tp)—forcipular tooth-plate, (ut)—ultimate tergite." figureDoi="http://doi.org/10.5281/zenodo.4402189" httpUri="https://zenodo.org/record/4402189/files/figure.png" pageId="41" pageNumber="42">Fig. 99</figureCitation>
|
||
) and
|
||
<taxonomicName id="4C77C3B8FFDBD723FAC9FF3E28D6DCE6" box="[1357,1427,223,249]" class="Chilopoda" family="Scolopendridae" genus="Alipes" kingdom="Animalia" order="Scolopendromorpha" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFDBD723FAC9FF3E28D6DCE6" box="[1357,1427,223,249]" italics="true" pageId="41" pageNumber="42">Alipes</emphasis>
|
||
</taxonomicName>
|
||
) well-developed process. Ultimate leg pretarsus claw-shaped, well-developed in most genera.
|
||
<bibRefCitation id="EFE6C5CAFFDBD723FB21FEE22C5CDD5E" author="Edgecombe, G. D. & Bonato, L." pageId="41" pageNumber="42" refId="ref47498" refString="Edgecombe, G. D. & Bonato, L. (2011) Chilopoda-taxonomic overview. Order Scolopendromorpha. In: Minelli, A. (Ed.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 1. Brill, Leiden, pp. 392 - 407." type="book" year="2011">Edgecombe & Bonato (2011: 400)</bibRefCitation>
|
||
also wrote: “
|
||
<collectingRegion id="49B376D9FFDBD723FE2CFEC62CBCDD5E" box="[424,505,295,321]" country="Saudi Arabia" name="Al Hudud ash Shamaliyah" pageId="41" pageNumber="42">Border</collectingRegion>
|
||
between labral and clypeal part of epipharynx strongly curved forwards... Testicular vesicles oriented oblique to central deferens duct”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36DEBB0FFDBD723FF43FE8E2CA2DDB2" pageId="41" pageNumber="42" type="description">
|
||
<paragraph id="8BC8B83BFFDBD723FF43FE8E2CA2DDB2" blockId="41.[151,1437,151,645]" pageId="41" pageNumber="42">
|
||
<emphasis id="B9036429FFDBD723FF43FE8E2CF1DD96" bold="true" box="[199,436,366,393]" pageId="41" pageNumber="42">Number of subtaxa.</emphasis>
|
||
6 genera (3 subgenera in genus
|
||
<taxonomicName id="4C77C3B8FFDBD723FC9AFE8E2EDBDD96" box="[798,926,367,393]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFDBD723FC9AFE8E2EDBDD96" box="[798,926,367,393]" italics="true" pageId="41" pageNumber="42">Otostigmus</emphasis>
|
||
</taxonomicName>
|
||
). “Ca 200 species in six genera” sensu
|
||
<bibRefCitation id="EFE6C5CAFFDBD723FADDFE8E2CA6DDB2" author="Edgecombe, G. D. & Bonato, L." pageId="41" pageNumber="42" refId="ref47498" refString="Edgecombe, G. D. & Bonato, L. (2011) Chilopoda-taxonomic overview. Order Scolopendromorpha. In: Minelli, A. (Ed.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 1. Brill, Leiden, pp. 392 - 407." type="book" year="2011">Edgecombe & Bonato (2011: 400)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36DEBB0FFDBD723FF43FE562CF8DDCE" box="[199,445,439,465]" pageId="41" pageNumber="42" type="distribution">
|
||
<paragraph id="8BC8B83BFFDBD723FF43FE562CF8DDCE" blockId="41.[151,1437,151,645]" box="[199,445,439,465]" pageId="41" pageNumber="42">
|
||
<emphasis id="B9036429FFDBD723FF43FE562C5FDDCE" bold="true" box="[199,282,439,465]" pageId="41" pageNumber="42">Range.</emphasis>
|
||
As for family.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36DEBB0FFDBD723FF43FE3B2E74DDEA" box="[199,817,474,501]" pageId="41" pageNumber="42" type="description">
|
||
<paragraph id="8BC8B83BFFDBD723FF43FE3B2E74DDEA" blockId="41.[151,1437,151,645]" box="[199,817,474,501]" pageId="41" pageNumber="42">
|
||
<emphasis id="B9036429FFDBD723FF43FE3B2CF2DDEB" bold="true" box="[199,439,474,500]" pageId="41" pageNumber="42">Sexual dimorphism.</emphasis>
|
||
Present in 3 taxa of genus-group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36DEBB0FFDBD723FF43FE1E2DA3DE9A" pageId="41" pageNumber="42" type="discussion">
|
||
<paragraph id="8BC8B83BFFDBD723FF43FE1E2DA3DE9A" blockId="41.[151,1437,151,645]" pageId="41" pageNumber="42">
|
||
<emphasis id="B9036429FFDBD723FF43FE1E2C7EDE06" bold="true" box="[199,315,511,537]" pageId="41" pageNumber="42">Remarks.</emphasis>
|
||
Treated as a tribe in
|
||
<bibRefCitation id="EFE6C5CAFFDBD723FD9AFE1E2ED8DE06" author="Edgecombe, G. D. & Bonato, L." box="[542,925,511,537]" pageId="41" pageNumber="42" refId="ref47498" refString="Edgecombe, G. D. & Bonato, L. (2011) Chilopoda-taxonomic overview. Order Scolopendromorpha. In: Minelli, A. (Ed.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 1. Brill, Leiden, pp. 392 - 407." type="book" year="2011">Edgecombe & Bonato (2011: 400)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE6C5CAFFDBD723FC23FE1E29ECDE06" author="Vahtera, V. & Edgecombe, G. D. & Giribet, G." box="[935,1193,511,537]" pageId="41" pageNumber="42" pagination="4 - 20" refId="ref51271" refString="Vahtera, V., Edgecombe, G. D. & Giribet, G. (2012 a) Evolution of blindness in scolopendromorph centipedes (Chilopoda: Scolopendromorpha): insight from an expanded sampling of molecular data. Cladistics, 28, 4 - 20. https: // doi. org / 10.1111 / j. 1096 - 0031.2011.00361. x" type="journal article" year="2012">
|
||
Vahtera
|
||
<emphasis id="B9036429FFDBD723FB86FE1E297CDE06" box="[1026,1081,511,537]" italics="true" pageId="41" pageNumber="42">et al.</emphasis>
|
||
(2012a: 7
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE6C5CAFFDBD723FB30FE1E2870DE06" author="Vahtera, V. & Edgecombe, G. D. & Giribet, G." box="[1204,1333,510,537]" pageId="41" pageNumber="42" pagination="225 - 248" refId="ref51336" refString="Vahtera, V., Edgecombe, G. D. & Giribet, G. (2012 b) Spiracle structure in scolopendromorph centipedes (Chilopoda: Scolopendromorpha) and its contribution to phylogenetics. Zoomorphology, 131, 225 - 248. https: // doi. org / 10.1007 / s 00435 - 012 - 0157 - 0" type="journal article" year="2012">2012b: 235</bibRefCitation>
|
||
),
|
||
<bibRefCitation id="EFE6C5CAFFDBD723FAC3FE1E2C1BDE22" author="Vahtera, V. & Edgecombe, G. D. & Giribet, G." pageId="41" pageNumber="42" pagination="578 - 602" refId="ref51397" refString="Vahtera, V., Edgecombe, G. D. & Giribet, G. (2013). Phylogenetics of scolopendromorph centipedes: can denser taxon sampling improve an artificial classification? Invertebrate Systematics, 27 (5), 578 - 602. https: // doi. org / 10.1071 / IS 13035" type="journal article" year="2013">
|
||
Vahtera
|
||
<emphasis id="B9036429FFDBD723FF13FDC22D95DE22" box="[151,208,547,573]" italics="true" pageId="41" pageNumber="42">et al.</emphasis>
|
||
(2013: 578)
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE6C5CAFFDBD723FEEEFDC22F97DE22" author="Vahtera, V. & Edgecombe, G. D." box="[362,722,547,573]" pageId="41" pageNumber="42" pagination="1 - 8" refId="ref51207" refString="Vahtera, V. & Edgecombe, G. D. (2014) First Molecular Data and the Phylogenetic Position of the Millipede-Like Centipede Edentistoma octosulcatum Tomosvary, 1882 (Chilopoda: Scolopendromorpha: Scolopendridae). PLoS ONE, 9 (11), 1 - 8. https: // doi. org / 10.1371 / journal. pone. 0112461" type="journal article" year="2014">Vahtera & Edgecombe (2014: 2</bibRefCitation>
|
||
, 5, 7). In 2013 Vahtera
|
||
<emphasis id="B9036429FFDBD723FC59FDC22953DE22" box="[989,1046,547,573]" italics="true" pageId="41" pageNumber="42">et al.</emphasis>
|
||
wrote (p. 578): “The generic classification of
|
||
<taxonomicName id="4C77C3B8FFDBD723FEAEFDA62CF5DE7E" authorityName="Kraepelin" authorityYear="1903" box="[298,432,583,609]" class="Chilopoda" family="Scolopendridae" kingdom="Animalia" order="Scolopendromorpha" pageId="41" pageNumber="53" phylum="Arthropoda" rank="tribe" tribe="Otostigmini">Otostigmini</taxonomicName>
|
||
has a poor fit to phylogenetic relationships, although nodal support within this tribe is weak”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |