264 lines
37 KiB
XML
264 lines
37 KiB
XML
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<mods:mods id="2F6AF24C5FA277B7A0CBD131A0B558F2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="70C65F9049BFFF0D5EF6A08F85453CD1">
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<mods:title id="63FC5D6F510ED1F8136686ADE16D92DE">An overview of the extant genera and subgenera of the order Scolopendromorpha (Chilopoda): a new identification key and updated diagnoses</mods:title>
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<mods:name id="FEEC542F863BF00E61D9D6F021D0AB7F" type="personal">
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<mods:namePart id="0F35682D97FA9BAC51F7FE46A92AC7C7">Schileyko, Arkady A.</mods:namePart>
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<mods:nameIdentifier id="055783E1E12B6DDF969E0A072B3AD238" type="email">schileyko1965@gmail.com</mods:nameIdentifier>
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<mods:namePart id="B25A51D70E635BA38F3EC2420B01F243">Vahtera, Varpu</mods:namePart>
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<mods:nameIdentifier id="56C43D7E0FB3EA55981F703C171F7792" type="email">varpu.vahtera@gmail.com</mods:nameIdentifier>
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<mods:namePart id="10DCA2E4435D42298966EEE7D9F9DB7B">Edgecombe, Gregory D.</mods:namePart>
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<mods:title id="20FFF887488E14FABF628C2BD1AD126C">Zootaxa</mods:title>
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<mods:date id="8EE8B603F64F5FD7912FDF6CE49AF08E">2020</mods:date>
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<mods:number id="0E7DBAD20CA7FAE9CB1B5273A876EC69">2020-08-10</mods:number>
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<mods:number id="38884046D4DDA7442006465240B19F73">4825</mods:number>
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<treatment id="03DE092DFFCAD732FF13FF76283DD86D" ID-DOI="http://doi.org/10.5281/zenodo.4402210" ID-GBIF-Taxon="177020739" ID-Zenodo-Dep="4402210" LSID="urn:lsid:plazi:treatment:03DE092DFFCAD732FF13FF76283DD86D" httpUri="http://treatment.plazi.org/id/03DE092DFFCAD732FF13FF76283DD86D" lastPageNumber="57" pageId="56" pageNumber="57">
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<subSubSection id="C36DEBB0FFCAD732FF13FF762C6ADCCA" pageId="56" pageNumber="57" type="nomenclature">
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<paragraph id="8BC8B83BFFCAD732FF13FF762C9CDCAE" blockId="56.[151,473,151,213]" box="[151,473,151,177]" pageId="56" pageNumber="57">
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<heading id="D0800F57FFCAD732FF13FF762C9CDCAE" bold="true" box="[151,473,151,177]" fontSize="11" level="2" pageId="56" pageNumber="57" reason="4">
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<taxonomicName id="4C77C3B8FFCAD732FF13FF762C9CDCAE" ID-CoL="7NB8" ID-ENA="1418208" authority="Attems, 1934" authorityName="Attems" authorityYear="1934" box="[151,473,151,177]" class="Chilopoda" family="Scolopendridae" genus="Sterropristes" kingdom="Animalia" order="Scolopendromorpha" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
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<emphasis id="B9036429FFCAD732FF13FF762C9CDCAE" bold="true" box="[151,473,151,177]" pageId="56" pageNumber="57">
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<emphasis id="B9036429FFCAD732FF13FF762C75DCAE" bold="true" box="[151,304,151,177]" italics="true" pageId="56" pageNumber="57">Sterropristes</emphasis>
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<bibRefCitation id="EFE6C5CAFFCAD732FEB2FF762C9CDCAE" author="Attems, C." box="[310,473,151,177]" pageId="56" pageNumber="57" pagination="43 - 62" refId="ref45555" refString="Attems, C. (1934) Neue Myriopoden des Museums Basel. Verhandlungen der Naturforschenden Gesellschaft in Basel, 45, 43 - 62." type="journal article" year="1934">Attems, 1934</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BC8B83BFFCAD732FF13FF5A2C6ADCCA" blockId="56.[151,473,151,213]" box="[151,303,187,213]" pageId="56" pageNumber="57">Figs 119–123</paragraph>
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</subSubSection>
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<subSubSection id="C36DEBB0FFCAD732FF13FEE12F72DD07" box="[151,567,256,280]" pageId="56" pageNumber="57" type="description">
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<paragraph id="8BC8B83BFFCAD732FF13FEE12F72DD07" blockId="56.[151,869,256,310]" box="[151,567,256,280]" pageId="56" pageNumber="57">
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Synonyms.
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<taxonomicName id="4C77C3B8FFCAD732FE8BFEE12F72DD07" authority="Verhoeff, 1937" authorityName="Verhoeff" authorityYear="1937" box="[271,567,256,280]" class="Chilopoda" family="Scolopendridae" genus="Malaccolabis" kingdom="Animalia" order="Scolopendromorpha" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
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<emphasis id="B9036429FFCAD732FE8BFEE12CDCDD07" box="[271,409,256,280]" italics="true" pageId="56" pageNumber="57">Malaccolabis</emphasis>
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<bibRefCitation id="EFE6C5CAFFCAD732FE1BFEE12F72DD07" author="Verhoeff, K. W." box="[415,567,256,280]" pageId="56" pageNumber="57" pagination="198 - 238" refId="ref51586" refString="Verhoeff, K. W. (1937) Chilopoden aus Malacca, nach den Objecten des Raffles Museum in Singapore. 1. Teil. Scolopendromorpha und Geophilomorpha. Bulletin of the Raffles Museum, 13, 198 - 238." type="journal article" year="1937">Verhoeff, 1937</bibRefCitation>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36DEBB0FFCAD732FF13FEFF2E20DD29" box="[151,869,286,310]" pageId="56" pageNumber="57" type="type_taxon">
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<paragraph id="8BC8B83BFFCAD732FF13FEFF2E20DD29" blockId="56.[151,869,256,310]" box="[151,869,286,310]" pageId="56" pageNumber="57">
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<emphasis id="B9036429FFCAD732FF13FEFF2C66DD29" bold="true" box="[151,291,286,310]" pageId="56" pageNumber="57">
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<typeStatus id="54CC0699FFCAD732FF13FEFF2D89DD29" box="[151,204,286,310]" pageId="56" pageNumber="57">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C77C3B8FFCAD732FEAEFEFE2F85DD29" ID-CoL="6ZS9D" authority="Attems, 1934" authorityName="Attems" authorityYear="1934" box="[298,704,286,310]" class="Chilopoda" family="Scolopendridae" genus="Sterropristes" kingdom="Animalia" order="Scolopendromorpha" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="sarasinorum">
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<emphasis id="B9036429FFCAD732FEAEFEFE2F77DD29" box="[298,562,287,310]" italics="true" pageId="56" pageNumber="57">Sterropristes sarasinorum</emphasis>
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<bibRefCitation id="EFE6C5CAFFCAD732FDB3FEFF2F85DD29" author="Attems, C." box="[567,704,286,310]" pageId="56" pageNumber="57" pagination="43 - 62" refId="ref45555" refString="Attems, C. (1934) Neue Myriopoden des Museums Basel. Verhandlungen der Naturforschenden Gesellschaft in Basel, 45, 43 - 62." type="journal article" year="1934">Attems, 1934</bibRefCitation>
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</taxonomicName>
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(by monotypy).
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</paragraph>
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</subSubSection>
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<subSubSection id="C36DEBB0FFCAD732FF13FE852F80DDBC" pageId="56" pageNumber="57" type="diagnosis">
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<paragraph id="8BC8B83BFFCAD732FF13FE852CE4DD61" blockId="56.[151,1437,356,1139]" box="[151,417,356,383]" pageId="56" pageNumber="57">
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<emphasis id="B9036429FFCAD732FF13FE852C57DD61" bold="true" box="[151,274,356,382]" pageId="56" pageNumber="57">Diagnosis.</emphasis>
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As for tribe.
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</paragraph>
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<paragraph id="8BC8B83BFFCAD732FF43FE682F80DDBC" blockId="56.[151,1437,356,1139]" box="[199,709,392,419]" pageId="56" pageNumber="57">
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<emphasis id="B9036429FFCAD732FF43FE682CECDDBD" bold="true" box="[199,425,392,419]" pageId="56" pageNumber="57">Number of species.</emphasis>
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3 (
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<bibRefCitation id="EFE6C5CAFFCAD732FE48FE692FFFDDBC" author="Muadsub, S. & Sutcharit, C. & Pimvichai, P. & Enghoff, H. & Edgecombe, G. D. & Panha, S." box="[460,698,392,419]" pageId="56" pageNumber="57" pagination="35 - 52" refId="ref49624" refString="Muadsub, S., Sutcharit, C., Pimvichai, P., Enghoff, H., Edgecombe, G. D. & Panha, S. (2012) Revision of the rare centipede genus Sterropristes Attems, 1934, with description of a new species from Thailand (Chilopoda: Scolopendromorpha: Scolopendridae). Zootaxa, 3484 (1), 35 - 52. https: // doi. org / 10.11646 / zootaxa. 3484.1.2" type="journal article" year="2012">
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Muadsub
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<emphasis id="B9036429FFCAD732FDBAFE682F33DDBD" box="[574,630,392,418]" italics="true" pageId="56" pageNumber="57">et al.</emphasis>
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2012
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</bibRefCitation>
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).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36DEBB0FFCAD732FF43FE4C283DD86D" pageId="56" pageNumber="57" type="discussion">
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<paragraph id="8BC8B83BFFCAD732FF43FE4C2FF2DDF4" blockId="56.[151,1437,356,1139]" pageId="56" pageNumber="57">
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<emphasis id="B9036429FFCAD732FF43FE4C2C7EDDD8" bold="true" box="[199,315,429,455]" pageId="56" pageNumber="57">Remarks.</emphasis>
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Treated as a genus in
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<bibRefCitation id="EFE6C5CAFFCAD732FDBEFE4D2E86DDD8" author="Edgecombe, G. D. & Bonato, L." box="[570,963,428,455]" pageId="56" pageNumber="57" refId="ref47498" refString="Edgecombe, G. D. & Bonato, L. (2011) Chilopoda-taxonomic overview. Order Scolopendromorpha. In: Minelli, A. (Ed.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 1. Brill, Leiden, pp. 392 - 407." type="book" year="2011">Edgecombe & Bonato (2011: 403)</bibRefCitation>
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,
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<bibRefCitation id="EFE6C5CAFFCAD732FC4BFE4D29BFDDD8" author="Muadsub, S. & Sutcharit, C. & Pimvichai, P. & Enghoff, H. & Edgecombe, G. D. & Panha, S." box="[975,1274,428,455]" pageId="56" pageNumber="57" pagination="35 - 52" refId="ref49624" refString="Muadsub, S., Sutcharit, C., Pimvichai, P., Enghoff, H., Edgecombe, G. D. & Panha, S. (2012) Revision of the rare centipede genus Sterropristes Attems, 1934, with description of a new species from Thailand (Chilopoda: Scolopendromorpha: Scolopendridae). Zootaxa, 3484 (1), 35 - 52. https: // doi. org / 10.11646 / zootaxa. 3484.1.2" type="journal article" year="2012">
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Muadsub
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<emphasis id="B9036429FFCAD732FBC4FE4C293CDDD9" box="[1088,1145,428,454]" italics="true" pageId="56" pageNumber="57">et al.</emphasis>
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||
(2012: 36)
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</bibRefCitation>
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,
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<bibRefCitation id="EFE6C5CAFFCAD732FA82FE4D2C58DDF4" author="Vahtera, V. & Edgecombe, G. D. & Giribet, G." pageId="56" pageNumber="57" pagination="578 - 602" refId="ref51397" refString="Vahtera, V., Edgecombe, G. D. & Giribet, G. (2013). Phylogenetics of scolopendromorph centipedes: can denser taxon sampling improve an artificial classification? Invertebrate Systematics, 27 (5), 578 - 602. https: // doi. org / 10.1071 / IS 13035" type="journal article" year="2013">
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Vahtera
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<emphasis id="B9036429FFCAD732FAE7FE4C28D8DDD9" box="[1379,1437,428,454]" italics="true" pageId="56" pageNumber="57">et al.</emphasis>
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(2013: 594)
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||
</bibRefCitation>
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||
,
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||
<bibRefCitation id="EFE6C5CAFFCAD732FEACFE312FC8DDF4" author="Vahtera, V. & Edgecombe, G. D." box="[296,653,464,491]" pageId="56" pageNumber="57" pagination="1 - 8" refId="ref51207" refString="Vahtera, V. & Edgecombe, G. D. (2014) First Molecular Data and the Phylogenetic Position of the Millipede-Like Centipede Edentistoma octosulcatum Tomosvary, 1882 (Chilopoda: Scolopendromorpha: Scolopendridae). PLoS ONE, 9 (11), 1 - 8. https: // doi. org / 10.1371 / journal. pone. 0112461" type="journal article" year="2014">Vahtera & Edgecombe (2014: 2</bibRefCitation>
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, 7).
|
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</paragraph>
|
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<paragraph id="8BC8B83BFFCAD732FF43FE152864D818" blockId="56.[151,1437,356,1139]" pageId="56" pageNumber="57">
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Several “diagnostic” features of
|
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<taxonomicName id="4C77C3B8FFCAD732FDBCFE142F8DDE10" authorityName="Attems" authorityYear="1934" box="[568,712,501,527]" class="Chilopoda" family="Scolopendridae" genus="Sterropristes" kingdom="Animalia" order="Scolopendromorpha" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B9036429FFCAD732FDBCFE142F8DDE10" box="[568,712,501,527]" italics="true" pageId="56" pageNumber="57">Sterropristes</emphasis>
|
||
</taxonomicName>
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||
mentioned by
|
||
<bibRefCitation id="EFE6C5CAFFCAD732FCF7FE15292ADE10" author="Muadsub, S. & Sutcharit, C. & Pimvichai, P. & Enghoff, H. & Edgecombe, G. D. & Panha, S." box="[883,1135,500,527]" pageId="56" pageNumber="57" pagination="35 - 52" refId="ref49624" refString="Muadsub, S., Sutcharit, C., Pimvichai, P., Enghoff, H., Edgecombe, G. D. & Panha, S. (2012) Revision of the rare centipede genus Sterropristes Attems, 1934, with description of a new species from Thailand (Chilopoda: Scolopendromorpha: Scolopendridae). Zootaxa, 3484 (1), 35 - 52. https: // doi. org / 10.11646 / zootaxa. 3484.1.2" type="journal article" year="2012">
|
||
Muadsub
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||
<emphasis id="B9036429FFCAD732FC67FE142958DE11" box="[995,1053,500,526]" italics="true" pageId="56" pageNumber="57">et al.</emphasis>
|
||
(2012)
|
||
</bibRefCitation>
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||
are also shared with other genera of
|
||
<taxonomicName id="4C77C3B8FFCAD732FE81FDF92CDADE2D" authorityName="Kraepelin" authorityYear="1903" box="[261,415,536,562]" class="Chilopoda" family="Scolopendridae" kingdom="Animalia" order="Scolopendromorpha" pageId="56" pageNumber="57" phylum="Arthropoda" rank="subFamily" subFamily="Otostigminae">Otostigminae</taxonomicName>
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||
, an exception being the “saw-like internal margin of the forcipular tarsungula” which serves as the “unique distinguishing character of
|
||
<taxonomicName id="4C77C3B8FFCAD732FDF0FDDC2E41DE48" authority="(Muadsub et al. 2012: 36)" authorityName="Attems" authorityYear="1934" baseAuthorityName="Muadsub" baseAuthorityPageNumber="36" baseAuthorityYear="2012" box="[628,772,573,599]" class="Chilopoda" family="Scolopendridae" genus="Sterropristes" kingdom="Animalia" order="Scolopendromorpha" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFCAD732FDF0FDDC2E41DE48" box="[628,772,573,599]" italics="true" pageId="56" pageNumber="57">Sterropristes</emphasis>
|
||
</taxonomicName>
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||
” (
|
||
<bibRefCitation id="EFE6C5CAFFCAD732FC9BFDDD297CDE48" author="Muadsub, S. & Sutcharit, C. & Pimvichai, P. & Enghoff, H. & Edgecombe, G. D. & Panha, S." box="[799,1081,572,599]" pageId="56" pageNumber="57" pagination="35 - 52" refId="ref49624" refString="Muadsub, S., Sutcharit, C., Pimvichai, P., Enghoff, H., Edgecombe, G. D. & Panha, S. (2012) Revision of the rare centipede genus Sterropristes Attems, 1934, with description of a new species from Thailand (Chilopoda: Scolopendromorpha: Scolopendridae). Zootaxa, 3484 (1), 35 - 52. https: // doi. org / 10.11646 / zootaxa. 3484.1.2" type="journal article" year="2012">
|
||
Muadsub
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<emphasis id="B9036429FFCAD732FC15FDDC2E8FDE49" box="[913,970,572,598]" italics="true" pageId="56" pageNumber="57">et al.</emphasis>
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||
2012: 36
|
||
</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="EFE6C5CAFFCAD732FBC9FDDD2DEADE64" author="Vahtera, V. & Edgecombe, G. D." pageId="56" pageNumber="57" pagination="1 - 8" refId="ref51207" refString="Vahtera, V. & Edgecombe, G. D. (2014) First Molecular Data and the Phylogenetic Position of the Millipede-Like Centipede Edentistoma octosulcatum Tomosvary, 1882 (Chilopoda: Scolopendromorpha: Scolopendridae). PLoS ONE, 9 (11), 1 - 8. https: // doi. org / 10.1371 / journal. pone. 0112461" type="journal article" year="2014">Vahtera & Edgecombe (2014: 7)</bibRefCitation>
|
||
also wrote: “a close relationship between
|
||
<taxonomicName id="4C77C3B8FFCAD732FD19FD802E68DE64" authorityName="Attems" authorityYear="1934" box="[669,813,609,635]" class="Chilopoda" family="Scolopendridae" genus="Sterropristes" kingdom="Animalia" order="Scolopendromorpha" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFCAD732FD19FD802E68DE64" box="[669,813,609,635]" italics="true" pageId="56" pageNumber="57">Sterropristes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C77C3B8FFCAD732FCEEFD812EB3DE65" authorityName="Tomosvary" authorityYear="1882" box="[874,1014,608,634]" class="Chilopoda" family="Scolopendridae" genus="Edentistoma" kingdom="Animalia" order="Scolopendromorpha" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFCAD732FCEEFD812EB3DE65" box="[874,1014,608,634]" italics="true" pageId="56" pageNumber="57">Edentistoma</emphasis>
|
||
</taxonomicName>
|
||
cannot be definitely discounted”. In fact, within
|
||
<taxonomicName id="4C77C3B8FFCAD732FE99FD652CF1DE81" authorityName="Kraepelin" authorityYear="1903" box="[285,436,644,670]" class="Chilopoda" family="Scolopendridae" kingdom="Animalia" order="Scolopendromorpha" pageId="56" pageNumber="57" phylum="Arthropoda" rank="subFamily" subFamily="Otostigminae">Otostigminae</taxonomicName>
|
||
only two these genera (plus
|
||
<taxonomicName id="4C77C3B8FFCAD732FD77FD642E1FDE81" box="[755,858,645,670]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFCAD732FD77FD642E1FDE81" box="[755,858,645,670]" italics="true" pageId="56" pageNumber="57">Digitipes</emphasis>
|
||
</taxonomicName>
|
||
, at least in part; see above) have an unusually small median tooth of the labrum (in
|
||
<taxonomicName id="4C77C3B8FFCAD732FE71FD492FE3DEDD" box="[501,678,680,706]" class="Chilopoda" family="Scolopendridae" kingdom="Animalia" order="Scolopendromorpha" pageId="56" pageNumber="57" phylum="Arthropoda" rank="subFamily" subFamily="Scolopendrinae">Scolopendrinae</taxonomicName>
|
||
, also shared with species of
|
||
<taxonomicName id="4C77C3B8FFCAD732FC60FD49282EDEDC" authority="" authorityName="Ribaut" authorityYear="1923" box="[996,1387,680,707]" class="Chilopoda" family="Scolopendridae" genus="Cormocephalus" kingdom="Animalia" order="Scolopendromorpha" pageId="56" pageNumber="57" phylum="Arthropoda" rank="subGenus" subGenus="Campylostigmus">
|
||
<emphasis id="B9036429FFCAD732FC60FD4929D0DEDD" box="[996,1173,680,706]" italics="true" pageId="56" pageNumber="57">Cormocephalus</emphasis>
|
||
(
|
||
<emphasis id="B9036429FFCAD732FB22FD492824DEDD" box="[1190,1377,680,706]" italics="true" pageId="56" pageNumber="57">Campylostigmus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
; see
|
||
<bibRefCitation id="EFE6C5CAFFCAD732FF13FD2D2C84DEF8" author="Edgecombe, G. D. & Koch, M." box="[151,449,716,743]" pageId="56" pageNumber="57" pagination="872 - 901" refId="ref47403" refString="Edgecombe, G. D. & Koch, M. (2008) Phylogeny of scolopendromorph centipedes (Chilopoda): morphological analysis featuring characters from the peristomatic area. Cladistics, 24, 872 - 901. https: // doi. org / 10.1111 / j. 1096 - 0031.2008.00220. x" type="journal article" year="2008">Edgecombe & Koch 2008</bibRefCitation>
|
||
, fig. 10e) and very characteristic short and much enlarged ultimate legs (see above), in both cases having dorsal sulci on proximal podomeres (
|
||
<figureCitation id="134CA4BEFFCAD732FC83FD112E2CDF14" box="[775,873,752,779]" captionStart="FIGURES 113–118" captionStartId="54.[151,264,1746,1770]" captionTargetBox="[167,1419,198,1704]" captionTargetId="figure-19@54.[151,1436,181,1722]" captionTargetPageId="54" captionText="FIGURES 113–118. Edentistoma octosulcatum Tömösváry, 1882; Voucher ZMUT_Chi01 113 left ultimate leg laterally; ZMB 3618 114 forcipular segment ventrally; Voucher ZMUT_Chi01 115 right pleuron of LBS 7 laterally 116 LBS 13–21+ right ultimate leg dorsally 117 LBS 20–21+ right ultimate leg ventrally 118 left ultimate leg medially; (20)—leg 20, (bl)—low blade-like lobe in place of forcipular tooth-plate, (bs)—basal suture of blade-like lobe, (cp)—spineless coxopleural process, (csp)—spineless “corner spine(s)” of ultimate prefemur, (cx)—coxopleuron, (im)—intersclerite membrane, (lsc)—sternal median longitudinal sulcus, (pf)—relatively short ultimate prefemur, (pts)—pretarsus, (rpr)—rudiment of trochantero-prefemoral process, (s)—open spiracle without atrium, (sh)—spiracle hump, (t1)—tarsus 1, (t2)—tarsus 2, (tk)—tergal longitudinal keels, (tr)—visibly elongated tarsungula, (ust)—ultimate sternite, (ut)—ultimate tergite." figureDoi="http://doi.org/10.5281/zenodo.4402195" httpUri="https://zenodo.org/record/4402195/files/figure.png" pageId="56" pageNumber="57">Fig. 116</figureCitation>
|
||
). However they differ sharply from each other by: a. visibly thinned and elongated (in
|
||
<taxonomicName id="4C77C3B8FFCAD732FDAFFCF52FF2DF31" authorityName="Tomosvary" authorityYear="1882" box="[555,695,788,814]" class="Chilopoda" family="Scolopendridae" genus="Edentistoma" kingdom="Animalia" order="Scolopendromorpha" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFCAD732FDAFFCF52FF2DF31" box="[555,695,788,814]" italics="true" pageId="56" pageNumber="57">Edentistoma</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="134CA4BEFFCAD732FD42FCF52E61DF30" box="[710,804,788,815]" captionStart="FIGURES 113–118" captionStartId="54.[151,264,1746,1770]" captionTargetBox="[167,1419,198,1704]" captionTargetId="figure-19@54.[151,1436,181,1722]" captionTargetPageId="54" captionText="FIGURES 113–118. Edentistoma octosulcatum Tömösváry, 1882; Voucher ZMUT_Chi01 113 left ultimate leg laterally; ZMB 3618 114 forcipular segment ventrally; Voucher ZMUT_Chi01 115 right pleuron of LBS 7 laterally 116 LBS 13–21+ right ultimate leg dorsally 117 LBS 20–21+ right ultimate leg ventrally 118 left ultimate leg medially; (20)—leg 20, (bl)—low blade-like lobe in place of forcipular tooth-plate, (bs)—basal suture of blade-like lobe, (cp)—spineless coxopleural process, (csp)—spineless “corner spine(s)” of ultimate prefemur, (cx)—coxopleuron, (im)—intersclerite membrane, (lsc)—sternal median longitudinal sulcus, (pf)—relatively short ultimate prefemur, (pts)—pretarsus, (rpr)—rudiment of trochantero-prefemoral process, (s)—open spiracle without atrium, (sh)—spiracle hump, (t1)—tarsus 1, (t2)—tarsus 2, (tk)—tergal longitudinal keels, (tr)—visibly elongated tarsungula, (ust)—ultimate sternite, (ut)—ultimate tergite." figureDoi="http://doi.org/10.5281/zenodo.4402195" httpUri="https://zenodo.org/record/4402195/files/figure.png" pageId="56" pageNumber="57">Fig. 114</figureCitation>
|
||
) vs. much shortened and enlarged (in
|
||
<taxonomicName id="4C77C3B8FFCAD732FB55FCF42824DF30" authorityName="Attems" authorityYear="1934" box="[1233,1377,789,815]" class="Chilopoda" family="Scolopendridae" genus="Sterropristes" kingdom="Animalia" order="Scolopendromorpha" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFCAD732FB55FCF42824DF30" box="[1233,1377,789,815]" italics="true" pageId="56" pageNumber="57">Sterropristes</emphasis>
|
||
</taxonomicName>
|
||
; Fig. 120) forcipular tarsungula, b. presence (in
|
||
<taxonomicName id="4C77C3B8FFCAD732FDEAFCD92FBFDF4D" authorityName="Tomosvary" authorityYear="1882" box="[622,762,824,850]" class="Chilopoda" family="Scolopendridae" genus="Edentistoma" kingdom="Animalia" order="Scolopendromorpha" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFCAD732FDEAFCD92FBFDF4D" box="[622,762,824,850]" italics="true" pageId="56" pageNumber="57">Edentistoma</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="134CA4BEFFCAD732FC8CFCD92E21DF4C" box="[776,868,824,851]" captionStart="FIGURES 113–118" captionStartId="54.[151,264,1746,1770]" captionTargetBox="[167,1419,198,1704]" captionTargetId="figure-19@54.[151,1436,181,1722]" captionTargetPageId="54" captionText="FIGURES 113–118. Edentistoma octosulcatum Tömösváry, 1882; Voucher ZMUT_Chi01 113 left ultimate leg laterally; ZMB 3618 114 forcipular segment ventrally; Voucher ZMUT_Chi01 115 right pleuron of LBS 7 laterally 116 LBS 13–21+ right ultimate leg dorsally 117 LBS 20–21+ right ultimate leg ventrally 118 left ultimate leg medially; (20)—leg 20, (bl)—low blade-like lobe in place of forcipular tooth-plate, (bs)—basal suture of blade-like lobe, (cp)—spineless coxopleural process, (csp)—spineless “corner spine(s)” of ultimate prefemur, (cx)—coxopleuron, (im)—intersclerite membrane, (lsc)—sternal median longitudinal sulcus, (pf)—relatively short ultimate prefemur, (pts)—pretarsus, (rpr)—rudiment of trochantero-prefemoral process, (s)—open spiracle without atrium, (sh)—spiracle hump, (t1)—tarsus 1, (t2)—tarsus 2, (tk)—tergal longitudinal keels, (tr)—visibly elongated tarsungula, (ust)—ultimate sternite, (ut)—ultimate tergite." figureDoi="http://doi.org/10.5281/zenodo.4402195" httpUri="https://zenodo.org/record/4402195/files/figure.png" pageId="56" pageNumber="57">Fig. 116</figureCitation>
|
||
) vs. absence of tergal keels, c. spiracles without (in
|
||
<taxonomicName id="4C77C3B8FFCAD732FF13FCBD2C66DF69" authorityName="Tomosvary" authorityYear="1882" box="[151,291,860,886]" class="Chilopoda" family="Scolopendridae" genus="Edentistoma" kingdom="Animalia" order="Scolopendromorpha" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFCAD732FF13FCBD2C66DF69" box="[151,291,860,886]" italics="true" pageId="56" pageNumber="57">Edentistoma</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="134CA4BEFFCAD732FEB6FCBD2CCADF68" box="[306,399,860,887]" captionStart="FIGURES 113–118" captionStartId="54.[151,264,1746,1770]" captionTargetBox="[167,1419,198,1704]" captionTargetId="figure-19@54.[151,1436,181,1722]" captionTargetPageId="54" captionText="FIGURES 113–118. Edentistoma octosulcatum Tömösváry, 1882; Voucher ZMUT_Chi01 113 left ultimate leg laterally; ZMB 3618 114 forcipular segment ventrally; Voucher ZMUT_Chi01 115 right pleuron of LBS 7 laterally 116 LBS 13–21+ right ultimate leg dorsally 117 LBS 20–21+ right ultimate leg ventrally 118 left ultimate leg medially; (20)—leg 20, (bl)—low blade-like lobe in place of forcipular tooth-plate, (bs)—basal suture of blade-like lobe, (cp)—spineless coxopleural process, (csp)—spineless “corner spine(s)” of ultimate prefemur, (cx)—coxopleuron, (im)—intersclerite membrane, (lsc)—sternal median longitudinal sulcus, (pf)—relatively short ultimate prefemur, (pts)—pretarsus, (rpr)—rudiment of trochantero-prefemoral process, (s)—open spiracle without atrium, (sh)—spiracle hump, (t1)—tarsus 1, (t2)—tarsus 2, (tk)—tergal longitudinal keels, (tr)—visibly elongated tarsungula, (ust)—ultimate sternite, (ut)—ultimate tergite." figureDoi="http://doi.org/10.5281/zenodo.4402195" httpUri="https://zenodo.org/record/4402195/files/figure.png" pageId="56" pageNumber="57">Fig. 115</figureCitation>
|
||
) vs. with a well-developed atrium (Fig. 119). On the other hand, we also note shared characters in the structure of the forcipular segment of
|
||
<taxonomicName id="4C77C3B8FFCAD732FD29FC612E09DF85" authorityName="Pocock" authorityYear="1898" box="[685,844,896,922]" class="Chilopoda" family="Scolopendridae" genus="Ethmostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFCAD732FD29FC612E09DF85" box="[685,844,896,922]" italics="true" pageId="56" pageNumber="57">Ethmostigmus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C77C3B8FFCAD732FC05FC602954DF84" authorityName="Attems" authorityYear="1934" box="[897,1041,897,923]" class="Chilopoda" family="Scolopendridae" genus="Sterropristes" kingdom="Animalia" order="Scolopendromorpha" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFCAD732FC05FC602954DF84" box="[897,1041,897,923]" italics="true" pageId="56" pageNumber="57">Sterropristes</emphasis>
|
||
</taxonomicName>
|
||
(compare
|
||
<figureCitation id="134CA4BEFFCAD732FB02FC6129A3DF84" box="[1158,1254,896,923]" captionStart="FIGURES 102–107" captionStartId="50.[151,264,1685,1709]" captionTargetBox="[274,1314,198,1647]" captionTargetId="figure-19@50.[257,1331,181,1661]" captionTargetPageId="50" captionText="FIGURES 102–107. Ethmostigmus rubripes spinosus (Newport, 1845); Juv. Rc 7174 102 forcipular segment ventrally; Ethmostigmus rubripes platycephalus (Newport, 1845); Ad. Rc 7514 103 left pleuron of LBS 3 laterally; Ad. Rc 7513 104 LBS 21 ventrally; Sad. СDT 105 coxopleural processes + prefemora of ultimate legs ventrally; Rhysida longipes (Newport, 1845); Ad. Rc 6683 106 anterior margin of forcipular coxosternite ventrally 107 coxopleural processes, legs 20 + ultimate legs laterally; (ap)—apical spine, (bs)—basal suture of forcipular tooth-plate, (cp)—coxopleural process, (csp)—corner spine of ultimate prefemur, (cx)—coxopleuron, (im)—intersclerite membrane, (lc)—lateral spines of coxopleural process, (pf)—ultimate prefemur, (pr)—trochantero-prefemoral process, (pts)—ultimate pretarsus, (rpf)—abnormally small (regenerated) right ultimate prefemur, (rpr)—rudiment of trochantero-prefemoral process, (s)—open spiracle without atrium, (scs)— spine at place of corner spine of ultimate prefemur, (su)—spines of ultimate prefemur, (t1)—ultimate tarsus 1, (tp)—forcipular tooth-plates, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402191" httpUri="https://zenodo.org/record/4402191/files/figure.png" pageId="56" pageNumber="57">Figs 102</figureCitation>
|
||
and 121)—both of them have very similar shape of the tooth-plates and lack a well-developed forcipular trochantero-prefemoral process. However, molecular phylogenetics indicates that “monophyly (non-partitioned analyses) versus paraphyly (partitioned analyses) of
|
||
<taxonomicName id="4C77C3B8FFCAD732FE2AFC0C2F7BD818" authorityName="Attems" authorityYear="1934" box="[430,574,1005,1031]" class="Chilopoda" family="Scolopendridae" genus="Sterropristes" kingdom="Animalia" order="Scolopendromorpha" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFCAD732FE2AFC0C2F7BD818" box="[430,574,1005,1031]" italics="true" pageId="56" pageNumber="57">Sterropristes</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="4C77C3B8FFCAD732FDD8FC0D2FBED819" authorityName="Pocock" authorityYear="1898" box="[604,763,1004,1030]" class="Chilopoda" family="Scolopendridae" genus="Ethmostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFCAD732FDD8FC0D2FBED819" box="[604,763,1004,1030]" italics="true" pageId="56" pageNumber="57">Ethmostigmus</emphasis>
|
||
</taxonomicName>
|
||
remains ambiguous” (
|
||
<bibRefCitation id="EFE6C5CAFFCAD732FC73FC0D2851D818" author="Siriwut, W. & Edgecombe, G. D. & Sutcharit, C. & Tongkerd, P. & Panha, S." box="[1015,1300,1004,1031]" pageId="56" pageNumber="57" pagination="1005 - 1049" refId="ref51057" refString="Siriwut, W., Edgecombe, G. D., Sutcharit, C., Tongkerd, P. & Panha, S. (2018) Systematic revision and phylogenetic reassessment of the centipede genera Rhysida Wood, 1862 and Alluropus Silvestri, 1912 (Chilopoda: Scolopendromorpha) in Southeast Asia, with further discussion of the subfamily Otostigminae. Invertebrate Systematics, 32, 1005 - 1049. https: // doi. org / 10.1071 / IS 17081" type="journal article" year="2018">
|
||
Siriwut
|
||
<emphasis id="B9036429FFCAD732FBD6FC0C29C6D819" box="[1106,1155,1004,1030]" italics="true" pageId="56" pageNumber="57">et al</emphasis>
|
||
. 2018: 1043
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC8B83BFFCAD732FF43FBF0283DD86D" blockId="56.[151,1437,356,1139]" pageId="56" pageNumber="57">
|
||
It should be noted that
|
||
<bibRefCitation id="EFE6C5CAFFCAD732FE42FBF02FE3D834" author="Bonato, L. & Chagas-Junior, A. & Edgecombe, G. D. & Lewis, J. G. E. & Minelli, A. & Pereira, L. A. & Shelley, R. M. & Stoev, P. & Zapparoli, M." box="[454,678,1040,1067]" pageId="56" pageNumber="57" refId="ref45701" refString="Bonato, L., Chagas-Junior, A., Edgecombe, G. D., Lewis, J. G. E., Minelli, A., Pereira, L. A., Shelley, R. M., Stoev, P. & Zapparoli, M. (2016) ChiloBase 2.0 - A World Catalogue of Centipedes (Chilopoda). Available from: http: // chilobase. biologia. unipd. it (accessed 2 August 2016)" type="url" year="2016">
|
||
Bonato
|
||
<emphasis id="B9036429FFCAD732FD99FBF02F10D835" box="[541,597,1040,1066]" italics="true" pageId="56" pageNumber="57">et al.</emphasis>
|
||
(2016)
|
||
</bibRefCitation>
|
||
are obviously mistaken for not mentioning
|
||
<taxonomicName id="4C77C3B8FFCAD732FB08FBF0285FD835" baseAuthorityName="Verhoeff" baseAuthorityYear="1937" box="[1164,1306,1040,1067]" class="Chilopoda" family="Scolopendridae" genus="Sterropristes" kingdom="Animalia" order="Scolopendromorpha" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="metallicus">
|
||
<emphasis id="B9036429FFCAD732FB08FBF0285FD835" box="[1164,1306,1040,1067]" italics="true" pageId="56" pageNumber="57">S. metallicus</emphasis>
|
||
</taxonomicName>
|
||
as the third valid species of
|
||
<taxonomicName id="4C77C3B8FFCAD732FEC8FBD42C99D850" authorityName="Attems" authorityYear="1934" box="[332,476,1077,1103]" class="Chilopoda" family="Scolopendridae" genus="Sterropristes" kingdom="Animalia" order="Scolopendromorpha" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFCAD732FEC8FBD42C99D850" box="[332,476,1077,1103]" italics="true" pageId="56" pageNumber="57">Sterropristes</emphasis>
|
||
</taxonomicName>
|
||
; its validity was confirmed by the molecular study of
|
||
<bibRefCitation id="EFE6C5CAFFCAD732FBBFFBD52820D850" author="Siriwut, W. & Edgecombe, G. D. & Sutcharit, C. & Tongkerd, P. & Panha, S." box="[1083,1381,1076,1103]" pageId="56" pageNumber="57" pagination="1005 - 1049" refId="ref51057" refString="Siriwut, W., Edgecombe, G. D., Sutcharit, C., Tongkerd, P. & Panha, S. (2018) Systematic revision and phylogenetic reassessment of the centipede genera Rhysida Wood, 1862 and Alluropus Silvestri, 1912 (Chilopoda: Scolopendromorpha) in Southeast Asia, with further discussion of the subfamily Otostigminae. Invertebrate Systematics, 32, 1005 - 1049. https: // doi. org / 10.1071 / IS 17081" type="journal article" year="2018">
|
||
Siriwut
|
||
<emphasis id="B9036429FFCAD732FB10FBD42983D851" box="[1172,1222,1076,1102]" italics="true" pageId="56" pageNumber="57">et al</emphasis>
|
||
. (2018: 1043)
|
||
</bibRefCitation>
|
||
who wrote: “the mainland and insular species,
|
||
<taxonomicName id="4C77C3B8FFCAD732FDE8FBB82FBED86D" baseAuthorityName="Verhoeff" baseAuthorityYear="1937" box="[620,763,1112,1139]" class="Chilopoda" family="Scolopendridae" genus="Sterropristes" kingdom="Animalia" order="Scolopendromorpha" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="metallicus">
|
||
<emphasis id="B9036429FFCAD732FDE8FBB82FBED86D" box="[620,763,1112,1139]" italics="true" pageId="56" pageNumber="57">S. metallicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C77C3B8FFCAD732FCB6FBB82EFDD86D" authorityName="Muadsub & Panha" authorityYear="2012" box="[818,952,1112,1139]" class="Chilopoda" family="Scolopendridae" genus="Sterropristes" kingdom="Animalia" order="Scolopendromorpha" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="violaceus">
|
||
<emphasis id="B9036429FFCAD732FCB6FBB82EFDD86D" box="[818,952,1112,1139]" italics="true" pageId="56" pageNumber="57">S. violaceus</emphasis>
|
||
</taxonomicName>
|
||
, have been verified as distinct species”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |