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<document ID-DOI="10.1080/00222933.2018.1524032" ID-GBIF-Dataset="034ebcd9-efa7-416a-822c-b6aee0e57a1c" ID-ISSN="1464-5262" ID-Zenodo-Dep="3670229" checkinTime="1581941790231" checkinUser="valdenar" docAuthor="DeVries, Thomas J." docDate="2019" docId="03BB2338E36DE21DFE78A3085BDEFEC2" docLanguage="en" docName="JNATHIST.53.25–26.1533–1584.pdf.imf" docOrigin="Journal of Natural History 53 (25)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Turritella saposa Olssoni 1944" docType="treatment" docVersion="19" lastPageNumber="1570" masterDocId="FF825B40E348E23BFFA1A1435A0CFFCC" masterDocTitle="Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru)" masterLastPageNumber="1584" masterPageNumber="1533" pageNumber="1569" updateTime="1636522412784" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>DeVries, Thomas J.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="pubDate">
<mods:number>2018-12-17</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>53</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>25</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1533</mods:start>
<mods:end>1584</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1080/00222933.2018.1524032</mods:identifier>
<mods:identifier type="GBIF-Dataset">034ebcd9-efa7-416a-822c-b6aee0e57a1c</mods:identifier>
<mods:identifier type="ISSN">1464-5262</mods:identifier>
<mods:identifier type="Zenodo-Dep">3670229</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3671272" ID-GBIF-Taxon="162146474" ID-Zenodo-Dep="3671272" LSID="urn:lsid:plazi:treatment:03BB2338E36DE21DFE78A3085BDEFEC2" httpUri="http://treatment.plazi.org/id/03BB2338E36DE21DFE78A3085BDEFEC2" lastPageId="38" lastPageNumber="1570" pageId="37" pageNumber="1569">
<subSubSection box="[473,841,587,612]" pageId="37" pageNumber="1569" type="nomenclature">
<paragraph blockId="37.[473,841,587,647]" box="[473,841,587,612]" pageId="37" pageNumber="1569">
<heading box="[473,841,587,612]" centered="true" fontSize="11" level="2" pageId="37" pageNumber="1569" reason="2">
<taxonomicName authority="Olssoni 1944" authorityName="Olssoni" authorityYear="1944" box="[473,841,587,612]" class="Gastropoda" family="Turritellidae" genus="Turritella" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="37" pageNumber="1569" phylum="Mollusca" rank="species" species="saposa">
<emphasis bold="true" box="[473,681,587,612]" italics="true" pageId="37" pageNumber="1569">Turritella saposa</emphasis>
Olsson, 1944
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[585,730,622,647]" pageId="37" pageNumber="1569" type="description">
<paragraph blockId="37.[473,841,587,647]" box="[585,730,622,647]" pageId="37" pageNumber="1569">
<heading box="[585,730,622,647]" centered="true" fontSize="11" level="2" pageId="37" pageNumber="1569" reason="5">
(
<figureCitation box="[593,692,622,647]" captionStart="Figure 5" captionStartId="14.[160,225,1068,1091]" captionTargetBox="[164,1150,144,1035]" captionTargetId="figure@14.[162,1153,142,1037]" captionTargetPageId="14" captionText="Figure 5. Gastropods from the Caballas Formation. (a–e) Caballasea segmentata gen. nov., sp. nov. B8769. (a) UWBM 107607, paratype, L 12.4 mm, apertural view. (b) UWBM 107607, abapertural view. (c) UWBM 107606, holotype, L 12.6 mm, abapertural view. (d) MUSM INV 262, paratype, L 8.3 mm, oblique view of parietal callus and columella. (e) UWBM 107608, paratype, L 15.3 mm, abapertural view. (f, g) Turritella capistrata sp. nov. (f) MUSM INV 259, paratype, B8772, L 16.3 mm, lateral view. (g) UWBM 107601, holotype, B8770, L 14.4 mm, apertural view. (h) Turritella saposa Olsson, 1944, UWBM 107602, B8770, L 15.5 mm. (i) Cristispira caballasensis sp. nov., UWBM 107605, holotype, B8769, L 5.1 mm, fragment. (j, k, m, n) Rhombopsis longinquus sp. nov. (j) UWBM 107630, holotype, B8772, L 18.3 mm, apertural view, outer lip missing. (k) MUSM INV 269, paratype, B8769, L 19.9 mm, abapertural view. (m) UWBM 107631, paratype, B8772, L 22.5 mm, abapertural view. (n) UWBM 107632, paratype, B8769, L 16.4 mm, lateral view of spire and columella. (l, o–q) Tristeirostoma saxaquadrata sp. nov. B8771. (l) UWBM 107636, paratype, L 10.0 mm, spire. (o) UWBM 107,635, holotype, L 7.6 mm, abapertural view. (p) UWBM 107635, lateral view of columella. (q) UWBM 107635, apertural/lateral view." figureDoi="http://doi.org/10.5281/zenodo.3670237" httpUri="https://zenodo.org/record/3670237/files/figure.png" pageId="37" pageNumber="1569">Figure 5</figureCitation>
(h))
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="1569" type="reference_group">
<paragraph blockId="37.[160,863,674,733]" box="[160,863,674,699]" pageId="37" pageNumber="1569">
<taxonomicName authority="Olsson 1944" authorityName="Olssoni" authorityYear="1944" box="[160,489,675,699]" class="Gastropoda" family="Turritellidae" genus="Turritella" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="37" pageNumber="1569" phylum="Mollusca" rank="species" species="saposa">
<emphasis box="[160,341,675,699]" italics="true" pageId="37" pageNumber="1569">Turritella saposa</emphasis>
<bibRefCitation author="Olsson AA" box="[350,489,675,699]" pageId="37" pageNumber="1569" pagination="1 - 147" refId="ref25769" refString="Olsson AA. 1944. Contributions to the Tertiary paleontology of northern Peru. Part VII. The Cretaceous of the Paita region. Bulletins Am Paleontol. 19 (68): 1 - 147." type="journal article" year="1944">Olsson 1944</bibRefCitation>
</taxonomicName>
, 71, pl. 10, figs. 4, 6, 7, 9, 11, 12.
</paragraph>
<paragraph blockId="37.[160,863,674,733]" box="[160,810,709,733]" pageId="37" pageNumber="1569">
<taxonomicName authority="Pilsbry 1944" authorityName="Pilsbryi" authorityYear="1944" box="[160,577,709,733]" class="Gastropoda" family="Turritellidae" genus="Turritella" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="37" pageNumber="1569" phylum="Mollusca" rank="subSpecies" species="saposa" subSpecies="austera">
<emphasis box="[160,431,709,733]" italics="true" pageId="37" pageNumber="1569">Turritella saposa austera</emphasis>
<bibRefCitation author="Pilsbry HA" box="[439,577,709,733]" pageId="37" pageNumber="1569" pagination="137 - 153" refId="ref25981" refString="Pilsbry HA. 1944. Molluscan fossils from the Rio Pachitea and vicinity in eastern Peru. Proc Acad Nat Sci Philadelphia. 96: 137 - 153." type="journal article" year="1944">Pilsbry 1944</bibRefCitation>
</taxonomicName>
, 139, pl. 9, figs. 5
<emphasis box="[777,790,709,733]" italics="true" pageId="37" pageNumber="1569">–</emphasis>
7.
</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="1569" type="diagnosis">
<paragraph blockId="37.[160,1156,783,878]" box="[160,276,783,809]" pageId="37" pageNumber="1569">
<emphasis bold="true" box="[160,276,783,809]" italics="true" pageId="37" pageNumber="1569">Diagnosis</emphasis>
</paragraph>
<paragraph blockId="37.[160,1156,783,878]" pageId="37" pageNumber="1569">Whorls with three strong primary spiral cords, nearly evenly spaced between sutures. Basal primary spiral cord bracketed by strong secondary spiral cords.</paragraph>
<paragraph blockId="37.[160,1156,927,1161]" box="[160,410,927,953]" pageId="37" pageNumber="1569">
<emphasis bold="true" box="[160,410,927,953]" italics="true" pageId="37" pageNumber="1569">Emended description</emphasis>
</paragraph>
<paragraph blockId="37.[160,1156,927,1161]" pageId="37" pageNumber="1569">
Estimated shell length to 30 mm, pleural angle about 20°. Protoconch unknown. Spire with estimated 10 whorls. Whorls straight-sided to convex, imbricate, spiral formula from later teleoconch whorls only D1u1C
<quantity box="[567,587,1033,1059]" metricMagnitude="3" metricUnit="kg" metricValue="1.0" pageId="37" pageNumber="1569" unit="t" value="1.0">1t</quantity>
1B1s1A1, becoming v1D1u1C
<quantity box="[900,919,1033,1059]" metricMagnitude="3" metricUnit="kg" metricValue="1.0" pageId="37" pageNumber="1569" unit="t" value="1.0">1t</quantity>
1B1s1s2A1; subsutural spiral cords, if present, not preserved. Spacing formula C30B55A86 (n = 1). Lateral growth line weakly prosocline, nearly orthocyrt. Lateral sinus very shallow, apex centred on spiral
<emphasis box="[229,235,1137,1161]" italics="true" pageId="37" pageNumber="1569">‘</emphasis>
B
<emphasis box="[249,255,1137,1161]" italics="true" pageId="37" pageNumber="1569">’</emphasis>
. Base not preserved. Allmon formula?-3-S-B-P.
</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="1569" type="discussion">
<paragraph blockId="37.[160,1156,1210,1617]" box="[160,262,1210,1236]" pageId="37" pageNumber="1569">
<emphasis bold="true" box="[160,262,1210,1236]" italics="true" pageId="37" pageNumber="1569">Remarks</emphasis>
</paragraph>
<paragraph blockId="37.[160,1156,1210,1617]" pageId="37" pageNumber="1569">
In the Caballas Formation, specimens of
<taxonomicName authorityName="Olssoni" authorityYear="1944" box="[618,798,1247,1271]" class="Gastropoda" family="Turritellidae" genus="Turritella" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="37" pageNumber="1569" phylum="Mollusca" rank="species" species="saposa">
<emphasis box="[618,798,1247,1271]" italics="true" pageId="37" pageNumber="1569">Turritella saposa</emphasis>
</taxonomicName>
occur together with specimens of
<taxonomicName authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[190,324,1281,1305]" class="Gastropoda" family="Turritellidae" genus="Turritella" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="37" pageNumber="1569" phylum="Mollusca" rank="species" species="capistrata" status="sp. nov.">
<emphasis box="[190,324,1281,1305]" italics="true" pageId="37" pageNumber="1569">T. capistrata</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[333,421,1281,1305]" pageId="37" pageNumber="1569" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[478,734,1281,1305]" class="Spirochaetes" family="Spirochaetaceae" genus="Cristispira" kingdom="Bacteria" order="Spirochaetales" pageId="37" pageNumber="1569" phylum="Spirochaetes" rank="species" species="caballasensis" status="sp. nov.">
<emphasis box="[478,734,1281,1305]" italics="true" pageId="37" pageNumber="1569">Cristispira caballasensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[742,830,1281,1305]" pageId="37" pageNumber="1569" rank="species">sp. nov.</taxonomicNameLabel>
<materialsCitation ID-GBIF-Occurrence="2565890655" location="In" municipality="northern Perui" pageId="37" pageNumber="1569" specimenCount="1">
<location LSID="urn:lsid:plazi:treatment:03BB2338E36DE21DFE78A3085BDEFEC2:8ECDC4F5E36DE21EFCE4A4425956FAD5" box="[837,858,1281,1305]" municipality="northern Perui" name="In" pageId="37" pageNumber="1569">In</location>
<collectingMunicipality box="[866,1029,1281,1305]" pageId="37" pageNumber="1569">northern Peru,</collectingMunicipality>
specimens of
<taxonomicName authorityName="Olssoni" authorityYear="1944" box="[190,291,1316,1340]" class="Gastropoda" family="Turritellidae" genus="Turritella" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="37" pageNumber="1569" phylum="Mollusca" rank="species" species="saposa">
<emphasis box="[190,205,1316,1340]" italics="true" pageId="37" pageNumber="1569">T</emphasis>
<emphasis box="[205,209,1316,1340]" italics="true" pageId="37" pageNumber="1569">.</emphasis>
<materialsCitation ID-GBIF-Occurrence="2565890649" collectorName="Jaillard" country="Haiti" location="upper Campanian" pageId="37" pageNumber="1569" specimenCount="1">
<emphasis box="[215,291,1316,1340]" italics="true" pageId="37" pageNumber="1569">saposa</emphasis>
occur alone in the
<location LSID="urn:lsid:plazi:treatment:03BB2338E36DE21DFE78A3085BDEFEC2:8ECDC4F5E36DE21EFE5FA46758CBFAF0" box="[510,711,1316,1340]" country="Haiti" name="upper Campanian" pageId="37" pageNumber="1569">upper Campanian</location>
lower
<collectingCountry box="[789,877,1316,1340]" name="Haiti" pageId="37" pageNumber="1569">Tortuga</collectingCountry>
Formation (
<collectorName box="[1013,1093,1316,1340]" pageId="37" pageNumber="1569">Jaillard</collectorName>
et al. 2005)
</materialsCitation>
</taxonomicName>
</materialsCitation>
.
<taxonomicName authority="Olssoni 1944" authorityName="Olssoni" authorityYear="1944" box="[240,599,1351,1375]" class="Gastropoda" family="Turritellidae" genus="Turritella" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="37" pageNumber="1569" phylum="Mollusca" rank="species" species="desolata">
<emphasis box="[240,440,1351,1375]" italics="true" pageId="37" pageNumber="1569">Turritella desolata</emphasis>
Olsson, 1944
</taxonomicName>
, which closely resembles
<taxonomicName authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[909,1047,1351,1375]" class="Gastropoda" family="Turritellidae" genus="Turritella" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="37" pageNumber="1569" phylum="Mollusca" rank="species" species="capistrata" status="sp. nov.">
<emphasis box="[909,1047,1351,1375]" italics="true" pageId="37" pageNumber="1569">T. capistrata</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1059,1153,1351,1375]" pageId="37" pageNumber="1569" rank="species">sp. nov.</taxonomicNameLabel>
<materialsCitation ID-GBIF-Occurrence="2565890727" collectorName="Formation" country="Haiti" location="Maastrichtian" pageId="37" pageNumber="1569" specimenCount="1">
, occurs alone in the
<location LSID="urn:lsid:plazi:treatment:03BB2338E36DE21DFE78A3085BDEFEC2:8ECDC4F5E36DE21EFE28A42A582CFA4D" box="[393,544,1385,1409]" country="Haiti" name="Maastrichtian" pageId="37" pageNumber="1569">Maastrichtian</location>
<emphasis box="[555,560,1385,1409]" italics="true" pageId="37" pageNumber="1569">‘</emphasis>
Baculites
<emphasis box="[659,665,1385,1409]" italics="true" pageId="37" pageNumber="1569">’</emphasis>
beds of the
<collectingCountry box="[819,907,1385,1409]" name="Haiti" pageId="37" pageNumber="1569">Tortuga</collectingCountry>
<collectorName box="[917,1036,1385,1409]" pageId="37" pageNumber="1569">Formation</collectorName>
</materialsCitation>
.
<taxonomicName authority="(Olssoni 1944)" baseAuthorityName="Olssoni" baseAuthorityYear="1944" class="Spirochaetes" family="Spirochaetaceae" genus="Cristispira" kingdom="Bacteria" order="Spirochaetales" pageId="37" pageNumber="1569" phylum="Spirochaetes" rank="species" species="prechira">
<emphasis italics="true" pageId="37" pageNumber="1569">Cristispira prechira</emphasis>
(Olsson, 1944)
</taxonomicName>
, which is similar to
<taxonomicName authorityName="DeVries" authorityYear="2019" box="[662,836,1420,1444]" class="Spirochaetes" family="Spirochaetaceae" genus="Cristispira" kingdom="Bacteria" order="Spirochaetales" pageId="37" pageNumber="1569" phylum="Spirochaetes" rank="species" species="caballasensis">
<emphasis box="[662,683,1420,1444]" italics="true" pageId="37" pageNumber="1569">C.</emphasis>
<materialsCitation ID-GBIF-Occurrence="2565890698" box="[694,836,1420,1444]" collectorName="Jaillard" country="Haiti" location="upper Campanian La Mesa Formation" pageId="37" pageNumber="1569" specimenCount="1">
<emphasis box="[694,836,1420,1444]" italics="true" pageId="37" pageNumber="1569">caballasensis</emphasis>
</materialsCitation>
</taxonomicName>
, occurs alone in the
<location LSID="urn:lsid:plazi:treatment:03BB2338E36DE21DFE78A3085BDEFEC2:8ECDC4F5E36DE21EFBE1A4CF580DFA0B" country="Haiti" name="upper Campanian La Mesa Formation" pageId="37" pageNumber="1569">upper Campanian La Mesa Formation</location>
(
<collectorName box="[528,608,1455,1479]" pageId="37" pageNumber="1569">Jaillard</collectorName>
et al. 2005).
</paragraph>
<paragraph blockId="37.[160,1156,1210,1617]" pageId="37" pageNumber="1569">
Specimens of
<taxonomicName authority="Pilsbryi 1944" authorityName="Pilsbryi" authorityYear="1944" box="[364,809,1489,1513]" class="Gastropoda" family="Turritellidae" genus="Turritella" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="37" pageNumber="1569" phylum="Mollusca" rank="subSpecies" species="saposa" subSpecies="austera">
<emphasis box="[364,646,1489,1513]" italics="true" pageId="37" pageNumber="1569">Turritella saposa austera</emphasis>
Pilsbry, 1944
</taxonomicName>
were described from shaley limestone beds near
<collectingCountry box="[411,447,1524,1548]" name="Iceland" pageId="37" pageNumber="1569">Isla</collectingCountry>
Chonta, along the Rio Pachitea in eastern
<collectingCountry box="[968,1019,1524,1548]" name="Peru" pageId="37" pageNumber="1569">Peru</collectingCountry>
(
<bibRefCitation author="Singewald JT Jr." pageId="37" pageNumber="1569" pagination="447 - 464" refId="ref26385" refString="Singewald JT Jr. 1926. Geology of the Pichis and Pachitea Riversi Peru. Bull Geol Soc Am. 39: 447 - 464." type="journal article" year="1926">Singewald 1926</bibRefCitation>
;
<bibRefCitation author="Pilsbry HA" box="[228,365,1559,1583]" pageId="37" pageNumber="1569" pagination="137 - 153" refId="ref25981" refString="Pilsbry HA. 1944. Molluscan fossils from the Rio Pachitea and vicinity in eastern Peru. Proc Acad Nat Sci Philadelphia. 96: 137 - 153." type="journal article" year="1944">Pilsbry 1944</bibRefCitation>
), in beds now recognised as the Upper Cretaceous Chonta Formation (CGS Consultores Associados S.A 1997).
</paragraph>
</subSubSection>
<subSubSection lastPageId="38" lastPageNumber="1570" pageId="37" pageNumber="1569" type="materials_examined">
<paragraph blockId="37.[160,1077,1667,1727]" box="[160,259,1667,1693]" pageId="37" pageNumber="1569">
<emphasis bold="true" box="[160,259,1667,1693]" italics="true" pageId="37" pageNumber="1569">Material</emphasis>
</paragraph>
<paragraph blockId="37.[160,1077,1667,1727]" box="[160,1077,1703,1727]" pageId="37" pageNumber="1569">
<materialsCitation ID-GBIF-Occurrence="2565890730" box="[160,1077,1703,1727]" pageId="37" pageNumber="1569">
<collectionCode box="[160,236,1703,1727]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="37" pageNumber="1569" type="Museum">UWBM</collectionCode>
107602, B8770, L (15.5), W (10.4);
<collectionCode box="[629,705,1703,1727]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="37" pageNumber="1569" type="Museum">UWBM</collectionCode>
107640, B8770, L (15.9), W (6.2).
</materialsCitation>
</paragraph>
<paragraph blockId="38.[160,1155,140,270]" box="[160,292,140,166]" pageId="38" pageNumber="1570">
<emphasis bold="true" box="[160,292,140,166]" italics="true" pageId="38" pageNumber="1570">Occurrence</emphasis>
</paragraph>
<paragraph blockId="38.[160,1155,140,270]" pageId="38" pageNumber="1570">
Upper Cretaceous,
<collectingCountry box="[399,487,177,201]" name="Haiti" pageId="38" pageNumber="1570">Tortuga</collectingCountry>
Formation, Talara Basin, northern Peru, and Chonta Formation, eastern
<collectingCountry box="[386,439,212,236]" name="Peru" pageId="38" pageNumber="1570">Peru</collectingCountry>
; Lower Paleogene, Cuenca Member, Caballas Formation, East Pisco Basin, southern
<collectingCountry box="[409,463,246,270]" name="Peru" pageId="38" pageNumber="1570">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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