344 lines
38 KiB
XML
344 lines
38 KiB
XML
<document ID-DOI="10.11646/zootaxa.5264.1.4" ID-GBIF-Dataset="860be0e8-ff7b-408d-8daa-4fa16dece7b6" ID-ISSN="1175-5326" ID-Zenodo-Dep="7836172" ID-ZooBank="47E141C8-F69A-422E-B7A7-E1AF4AD201BE" IM.bibliography_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" approvalRequired="6" approvalRequired_for_taxonomicNames="3" approvalRequired_for_textStreams="3" checkinTime="1681726002599" checkinUser="plazi" docAuthor="Nozaki, Tsubasa" docDate="2023" docId="03AB87F4FFB58B2FFF21FD09FBB333F0" docLanguage="en" docName="zootaxa.5264.1.4.pdf" docOrigin="Zootaxa 5264 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5264.1.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Andrikothelyna Pace 2000" docType="treatment" docVersion="4" lastPageNumber="66" masterDocId="FF92FF8CFFB48B2DFFB6FFCEFFF93055" masterDocTitle="Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan" masterLastPageNumber="76" masterPageNumber="64" pageNumber="65" updateTime="1681818565220" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Nozaki, Tsubasa</mods:namePart>
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<mods:affiliation>Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka 819 - 0395, Japan & The Kyushu University Museum, Fukuoka 812 - 8581, Japan</mods:affiliation>
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<mods:title>Zootaxa</mods:title>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:number>2023-04-12</mods:number>
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<mods:detail type="volume">
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<mods:number>5264</mods:number>
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<mods:start>64</mods:start>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5264.1.4</mods:url>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7836174" ID-Zenodo-Dep="7836174" LSID="urn:lsid:plazi:treatment:03AB87F4FFB58B2FFF21FD09FBB333F0" httpUri="http://treatment.plazi.org/id/03AB87F4FFB58B2FFF21FD09FBB333F0" lastPageId="2" lastPageNumber="66" pageId="1" pageNumber="65">
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<subSubSection box="[151,567,711,738]" pageId="1" pageNumber="65" type="nomenclature">
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<paragraph blockId="1.[151,567,711,738]" box="[151,567,711,738]" pageId="1" pageNumber="65">
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<heading bold="true" box="[151,567,711,738]" fontSize="11" level="1" pageId="1" pageNumber="65" reason="1">
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<emphasis bold="true" box="[151,567,711,738]" pageId="1" pageNumber="65">
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Genus
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<taxonomicName ID-CoL="8XJKK" authority="Pace, 2000" authorityName="Pace" authorityYear="2000" box="[238,567,711,738]" class="Insecta" family="Staphylinidae" genus="Andrikothelyna" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
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<emphasis bold="true" box="[238,427,711,738]" italics="true" pageId="1" pageNumber="65">Andrikothelyna</emphasis>
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<bibRefCitation author="Pace, R." box="[435,567,711,737]" pageId="1" pageNumber="65" pagination="109 - 163" refId="ref7639" refString="Pace, R. (2000) Aleocharinae di Papua-Nuova Guinea (Coleoptera, Staphylinidae). Staphylinidae). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Entomologie 70, 109 - 163." type="journal article" year="2000">Pace, 2000</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<paragraph blockId="1.[151,1436,775,892]" box="[151,818,775,800]" pageId="1" pageNumber="65">
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<treatmentCitation author="Pace, R." box="[151,478,776,800]" page="124" pageId="1" pageNumber="65" year="2000">
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<taxonomicName ID-CoL="8XJKK" authority="Pace, 2000: 124" authorityName="Pace" authorityPageNumber="124" authorityYear="2000" box="[151,478,776,800]" class="Insecta" family="Staphylinidae" genus="Andrikothelyna" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
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<emphasis box="[151,307,776,800]" italics="true" pageId="1" pageNumber="65">Andrikothelyna</emphasis>
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<bibRefCitation author="Pace, R." box="[314,478,776,800]" pageId="1" pageNumber="65" pagination="109 - 163" refId="ref7639" refString="Pace, R. (2000) Aleocharinae di Papua-Nuova Guinea (Coleoptera, Staphylinidae). Staphylinidae). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Entomologie 70, 109 - 163." type="journal article" year="2000">Pace, 2000: 124</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, figs 54–58 (original description)
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection box="[824,1353,776,800]" pageId="1" pageNumber="65" type="type_taxon">
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<paragraph blockId="1.[151,1436,775,892]" box="[824,1353,776,800]" pageId="1" pageNumber="65">
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[
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<typeStatus box="[830,876,776,800]" pageId="1" pageNumber="65">type</typeStatus>
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species:
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<taxonomicName authority="Pace, 2000" authorityName="Pace" authorityYear="2000" box="[969,1343,776,800]" class="Insecta" family="Staphylinidae" genus="Andrikothelyna" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="papuana">
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<emphasis box="[969,1220,776,800]" italics="true" pageId="1" pageNumber="65">Andrikothelyna papuana</emphasis>
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<bibRefCitation author="Pace, R." box="[1226,1343,776,800]" pageId="1" pageNumber="65" pagination="109 - 163" refId="ref7639" refString="Pace, R. (2000) Aleocharinae di Papua-Nuova Guinea (Coleoptera, Staphylinidae). Staphylinidae). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Entomologie 70, 109 - 163." type="journal article" year="2000">Pace, 2000</bibRefCitation>
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</taxonomicName>
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].
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,819,806,830]" pageId="1" pageNumber="65" type="reference_group">
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<paragraph blockId="1.[151,1436,775,892]" box="[151,819,806,830]" pageId="1" pageNumber="65">
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<treatmentCitationGroup box="[151,819,806,830]" pageId="1" pageNumber="65">
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<treatmentCitation author="Pace, R." box="[151,459,806,830]" page="35" pageId="1" pageNumber="65" year="2013">
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<taxonomicName ID-CoL="8XPXL" authority="Pace, 2013: 35" authorityName="Pace" authorityPageNumber="35" authorityYear="2013" box="[151,459,806,830]" class="Insecta" family="Staphylinidae" genus="Speiraphallusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
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<emphasis box="[151,304,806,830]" italics="true" pageId="1" pageNumber="65">Speiraphallusa</emphasis>
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<bibRefCitation author="Pace, R." box="[310,459,806,830]" pageId="1" pageNumber="65" pagination="33 - 63" refId="ref7746" refString="Pace, R. (2013) New distributional data, New Species and Three New Genera of Aleocharinae from Malaysia, Vietnam and Taiwan (Coleoptera: Staphylinidae). Tropical Zoology, 26 (1), 33 - 63. https: // doi. org / 10.1080 / 03946975.2013.775832" type="journal article" year="2013">Pace, 2013: 35</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, figs 1, 12–17 (original description)
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="65" type="multiple">
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<paragraph blockId="1.[151,1436,775,892]" pageId="1" pageNumber="65">
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[
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<typeStatus box="[830,876,807,831]" pageId="1" pageNumber="65">type</typeStatus>
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species:
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<taxonomicName authority="Pace, 2013" authorityName="Pace" authorityYear="2013" box="[967,1324,806,830]" class="Insecta" family="Staphylinidae" genus="Speiraphallusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="species" species="orientis">
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<emphasis box="[967,1203,806,830]" italics="true" pageId="1" pageNumber="65">Speiraphallusa orientis</emphasis>
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<bibRefCitation author="Pace, R." box="[1208,1324,806,830]" pageId="1" pageNumber="65" pagination="33 - 63" refId="ref7746" refString="Pace, R. (2013) New distributional data, New Species and Three New Genera of Aleocharinae from Malaysia, Vietnam and Taiwan (Coleoptera: Staphylinidae). Tropical Zoology, 26 (1), 33 - 63. https: // doi. org / 10.1080 / 03946975.2013.775832" type="journal article" year="2013">Pace, 2013</bibRefCitation>
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</taxonomicName>
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]; SantiagoJiménez and Santiago-Navarro 2016: 49 (key to genera of
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<taxonomicName authorityName="Mulsant & Rey" authorityYear="1871" box="[810,908,837,861]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="tribe" tribe="Placusini">Placusini</taxonomicName>
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);
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<treatmentCitation author="Pires-Silva, M. C." box="[926,1150,837,861]" page="469" pageId="1" pageNumber="65" year="2023">
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<bibRefCitation author="Pires-Silva, M. C." box="[926,1150,837,861]" pageId="1" pageNumber="65" pagination="46 - 7477" refId="ref7799" refString="Pires-Silva, M. C. (2023) Euryachen seuzei, a new genus and species of Placusini (Coleoptera: Staphylinidae: Aleocharinae) from Brazil. Zootaxa, 5230 (4), 46 - 7477. https: // doi. org / 10.11646 / zootaxa. 5230.4.4" type="journal article" year="2023">Pires-Silva 2023: 469</bibRefCitation>
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</treatmentCitation>
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(modified key to genera of
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<taxonomicName authorityName="Mulsant & Rey" authorityYear="1871" box="[199,297,868,892]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="tribe" tribe="Placusini">Placusini</taxonomicName>
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).
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<emphasis bold="true" box="[313,411,867,892]" pageId="1" pageNumber="65">Syn. nov.</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="65" type="diagnosis">
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<paragraph blockId="1.[151,1437,933,2040]" pageId="1" pageNumber="65">
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<emphasis bold="true" box="[151,274,933,959]" pageId="1" pageNumber="65">Diagnosis.</emphasis>
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<taxonomicName authorityName="Pace" authorityYear="2000" box="[280,453,934,960]" class="Insecta" family="Staphylinidae" genus="Andrikothelyna" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
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<emphasis box="[280,453,934,960]" italics="true" pageId="1" pageNumber="65">Andrikothelyna</emphasis>
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</taxonomicName>
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may be distinguished from other genera based on the following characteristics: strongly convex body; posterior angle of head round and neck almost absent; two-segmented labial palpi; short and unilobed ligula; two medial setae distant from each other; strongly transverse mentum; tarsal formula 4-4-5; extremely long segment I of meso- and metatarsi (fig.
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<bibRefCitation author="Pace, R." box="[652,771,1042,1068]" pageId="1" pageNumber="65" pagination="33 - 63" refId="ref7746" refString="Pace, R. (2013) New distributional data, New Species and Three New Genera of Aleocharinae from Malaysia, Vietnam and Taiwan (Coleoptera: Staphylinidae). Tropical Zoology, 26 (1), 33 - 63. https: // doi. org / 10.1080 / 03946975.2013.775832" type="journal article" year="2013">Pace 2013</bibRefCitation>
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; elytra with a pair of a longitudinal row of weak or distinct granules along the suture.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="65" type="description">
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<paragraph blockId="1.[151,1437,933,2040]" pageId="1" pageNumber="65">
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<emphasis bold="true" box="[199,444,1113,1140]" pageId="1" pageNumber="65">Redescription. Body</emphasis>
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fusiform (
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<figureCitation box="[569,673,1114,1140]" captionStart="FIGURES 1–8" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[214,1377,193,1897]" captionTargetId="figure-32@3.[206,1381,181,1920]" captionTargetPageId="3" captionText="FIGURES 1–8. Habitus of Andrikothelyna spp. 1, 5―Holotype of Andrikothelyna rubiginosa sp. nov., female; 2, 6―Holotype of Andrikothelyna limbata sp. nov., male (Kyushu); 3, 7―Andrikothelyna limbata sp. nov. male (Ishigaki Is.); 4, 8―Holotype of Andrikothelyna naomichii sp. nov. male." figureDoi="http://doi.org/10.5281/zenodo.7836176" httpUri="https://zenodo.org/record/7836176/files/figure.png" pageId="1" pageNumber="65">Figs 1–8</figureCitation>
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); dorsal surface microreticulation indistinct but that on abdominal tergite II distinct; pronotum, elytra, and abdominal tergites covered with rough reticulation sometimes forming small granules around pores.
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</paragraph>
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<paragraph blockId="1.[151,1437,933,2040]" pageId="1" pageNumber="65">
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<emphasis bold="true" box="[199,263,1222,1248]" pageId="1" pageNumber="65">Head</emphasis>
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transverse; almost all microsetae directed anterolaterally; infraorbital carina well developed; posterior angle round; neck almost absent. Antennae moderately long; segments IV–XI with dense microsetae.
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</paragraph>
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<paragraph blockId="1.[151,1437,933,2040]" pageId="1" pageNumber="65">
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<emphasis bold="true" box="[199,349,1294,1320]" pageId="1" pageNumber="65">Mouthparts.</emphasis>
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Labrum (
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<figureCitation box="[458,525,1294,1320]" captionStart="FIGURES 9–15" captionStartId="4.[151,264,1965,1989]" captionTargetBox="[178,1408,532,1923]" captionTargetId="figure-128@4.[166,1421,520,1942]" captionTargetPageId="4" captionText="FIGURES 9–15. Mouth parts of Andrikothelyna rubiginosa sp. nov. 9―Labrum; 10―epipharynx; 11―left mandible; 12― right mandible; 13―maxilla; 14―labium; 15―mentum." figureDoi="http://doi.org/10.5281/zenodo.7836180" httpUri="https://zenodo.org/record/7836180/files/figure.png" pageId="1" pageNumber="65">Fig. 9</figureCitation>
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). Anterior margin weakly concave, surface with about 17 setae; epipharynx with 2 pairs of small sensillae on anterior margin, and 3 pairs of micro setae on lateral margin; basal region with transverse row of sensory pores. Mandible (
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<figureCitation box="[536,666,1366,1392]" captionStart="FIGURES 9–15" captionStartId="4.[151,264,1965,1989]" captionTargetBox="[178,1408,532,1923]" captionTargetId="figure-128@4.[166,1421,520,1942]" captionTargetPageId="4" captionText="FIGURES 9–15. Mouth parts of Andrikothelyna rubiginosa sp. nov. 9―Labrum; 10―epipharynx; 11―left mandible; 12― right mandible; 13―maxilla; 14―labium; 15―mentum." figureDoi="http://doi.org/10.5281/zenodo.7836180" httpUri="https://zenodo.org/record/7836180/files/figure.png" pageId="1" pageNumber="65">Figs 11–12</figureCitation>
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) short, right with large subapical teeth; prostheca not fringed with tooth or setulae, basal area forming tongue like lobe; spinules of molar dorsal area not in distinct row; ventral molar area without denticles. Maxilla (
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<figureCitation box="[519,603,1438,1464]" captionStart="FIGURES 9–15" captionStartId="4.[151,264,1965,1989]" captionTargetBox="[178,1408,532,1923]" captionTargetId="figure-128@4.[166,1421,520,1942]" captionTargetPageId="4" captionText="FIGURES 9–15. Mouth parts of Andrikothelyna rubiginosa sp. nov. 9―Labrum; 10―epipharynx; 11―left mandible; 12― right mandible; 13―maxilla; 14―labium; 15―mentum." figureDoi="http://doi.org/10.5281/zenodo.7836180" httpUri="https://zenodo.org/record/7836180/files/figure.png" pageId="1" pageNumber="65">Fig. 13</figureCitation>
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), palpi without pseudosegment; apex of galea moderately brushy; lacinia widened apicad, spines on adoral margin distributed at only apical half. Labium (
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<figureCitation box="[1055,1137,1474,1500]" captionStart="FIGURES 9–15" captionStartId="4.[151,264,1965,1989]" captionTargetBox="[178,1408,532,1923]" captionTargetId="figure-128@4.[166,1421,520,1942]" captionTargetPageId="4" captionText="FIGURES 9–15. Mouth parts of Andrikothelyna rubiginosa sp. nov. 9―Labrum; 10―epipharynx; 11―left mandible; 12― right mandible; 13―maxilla; 14―labium; 15―mentum." figureDoi="http://doi.org/10.5281/zenodo.7836180" httpUri="https://zenodo.org/record/7836180/files/figure.png" pageId="1" pageNumber="65">Fig. 14</figureCitation>
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), palpi with two segments, apical segment longer than basal one; apex of ligula unilobed and rounded. Prementum with two medial setae with pores distant from each other, with many pseudopores, one real pore, and one setalpore on each side; lateral margin of hypopharynx with setae entirely; apodeme with two medial projections. Mentum (
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<figureCitation box="[1100,1181,1582,1609]" captionStart="FIGURES 9–15" captionStartId="4.[151,264,1965,1989]" captionTargetBox="[178,1408,532,1923]" captionTargetId="figure-128@4.[166,1421,520,1942]" captionTargetPageId="4" captionText="FIGURES 9–15. Mouth parts of Andrikothelyna rubiginosa sp. nov. 9―Labrum; 10―epipharynx; 11―left mandible; 12― right mandible; 13―maxilla; 14―labium; 15―mentum." figureDoi="http://doi.org/10.5281/zenodo.7836180" httpUri="https://zenodo.org/record/7836180/files/figure.png" pageId="1" pageNumber="65">Fig. 15</figureCitation>
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) strongly transverse, 3 times as wide as long, with about 10 setae.
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</paragraph>
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<paragraph blockId="1.[151,1437,933,2040]" pageId="1" pageNumber="65">
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<emphasis bold="true" box="[199,295,1654,1680]" pageId="1" pageNumber="65">Thorax.</emphasis>
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Pronotum strongly transverse, postero-lateral angle gently sinuate, microsetae directed posteriorly. Elytra transverse; surface along suture and lateral margin with longitudinal row of strong to weak granules. Mesoventral processes (
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<figureCitation box="[425,508,1726,1752]" captionStart="FIGURES 16–19" captionStartId="6.[151,264,1900,1924]" captionTargetBox="[197,1383,622,1849]" captionTargetId="figure-191@6.[151,1436,597,1877]" captionTargetPageId="6" captionText="FIGURES 16–19. Andrikothelyna rubiginosa sp. nov. female. 16―Meso- and metaventrite; 17―abdominal tergite VIII; 18― abdominal sternite VIII; 19―spermatheca." figureDoi="http://doi.org/10.5281/zenodo.7836182" httpUri="https://zenodo.org/record/7836182/files/figure.png" pageId="1" pageNumber="65">Fig. 16</figureCitation>
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) without medial carina, apex round or truncated, extending middle of mesocoxal cavities; mesocoxal cavities mostly separated; apex of metaventral process round.
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</paragraph>
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<paragraph blockId="1.[151,1437,933,2040]" box="[199,1227,1797,1824]" pageId="1" pageNumber="65">
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<emphasis bold="true" box="[199,316,1797,1823]" pageId="1" pageNumber="65">Abdomen</emphasis>
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elongate, slightly narrowed posteriad; posterior margin of tergite VIII with teeth.
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</paragraph>
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<paragraph blockId="1.[151,1437,933,2040]" box="[199,1008,1834,1861]" pageId="1" pageNumber="65">
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<emphasis bold="true" box="[199,262,1834,1860]" pageId="1" pageNumber="65">Legs.</emphasis>
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Tarsal formula 4-4-5; segment I of meso- and metatarsi very long.
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</paragraph>
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<paragraph blockId="1.[151,1437,933,2040]" box="[199,1005,1870,1896]" pageId="1" pageNumber="65">
|
||
<emphasis bold="true" box="[199,267,1870,1896]" pageId="1" pageNumber="65">Male.</emphasis>
|
||
Median lobe of aedeagus narrow; apical lobe of paramerite short.
|
||
</paragraph>
|
||
<paragraph blockId="1.[151,1437,933,2040]" box="[199,645,1906,1932]" pageId="1" pageNumber="65">
|
||
<emphasis bold="true" box="[199,293,1906,1932]" pageId="1" pageNumber="65">Female.</emphasis>
|
||
Spermatheca short and simple.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="2" lastPageNumber="66" pageId="1" pageNumber="65" type="discussion">
|
||
<paragraph blockId="1.[151,1437,933,2040]" pageId="1" pageNumber="65">
|
||
<emphasis bold="true" box="[199,315,1942,1968]" pageId="1" pageNumber="65">Remarks.</emphasis>
|
||
<taxonomicName authorityName="Pace" authorityYear="2013" box="[322,492,1942,1968]" class="Insecta" family="Staphylinidae" genus="Speiraphallusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[322,492,1942,1968]" italics="true" pageId="1" pageNumber="65">Speiraphallusa</emphasis>
|
||
</taxonomicName>
|
||
completely matches
|
||
<taxonomicName authorityName="Pace" authorityYear="2000" box="[727,900,1942,1968]" class="Insecta" family="Staphylinidae" genus="Andrikothelyna" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[727,900,1942,1968]" italics="true" pageId="1" pageNumber="65">Andrikothelyna</emphasis>
|
||
</taxonomicName>
|
||
in character states which are essential for genus classification such as mouth parts, tarsi, and granules on elytra. Thus, it is reasonable to consider
|
||
<taxonomicName authorityName="Pace" authorityYear="2013" box="[1237,1407,1978,2004]" class="Insecta" family="Staphylinidae" genus="Speiraphallusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1237,1407,1978,2004]" italics="true" pageId="1" pageNumber="65">Speiraphallusa</emphasis>
|
||
</taxonomicName>
|
||
as a junior synonym of
|
||
<taxonomicName authorityName="Pace" authorityYear="2000" box="[384,557,2014,2040]" class="Insecta" family="Staphylinidae" genus="Andrikothelyna" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="65" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[384,557,2014,2040]" italics="true" pageId="1" pageNumber="65">Andrikothelyna</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,933]" pageId="2" pageNumber="66">
|
||
Despite examining only males,
|
||
<bibRefCitation author="Pace, R." box="[557,693,151,177]" pageId="2" pageNumber="66" pagination="33 - 63" refId="ref7746" refString="Pace, R. (2013) New distributional data, New Species and Three New Genera of Aleocharinae from Malaysia, Vietnam and Taiwan (Coleoptera: Staphylinidae). Tropical Zoology, 26 (1), 33 - 63. https: // doi. org / 10.1080 / 03946975.2013.775832" type="journal article" year="2013">Pace (2013)</bibRefCitation>
|
||
described the rows of granules of elytra, along the suture and the lateral margin of elytra, as a characteristic of the males of
|
||
<taxonomicName authorityName="Pace" authorityYear="2013" box="[828,998,187,213]" class="Insecta" family="Staphylinidae" genus="Speiraphallusa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[828,998,187,213]" italics="true" pageId="2" pageNumber="66">Speiraphallusa</emphasis>
|
||
</taxonomicName>
|
||
. However, no sex-specific differences were present in the elytra in all the new species studied in this paper. Distinct sexual dimorphism was observed on the tergite VII and sternite VIII of
|
||
<taxonomicName authority="Nozaki, 2023" authorityName="Nozaki" authorityYear="2023" box="[538,803,259,285]" class="Insecta" family="Staphylinidae" genus="Andrikothelyna" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="66" phylum="Arthropoda" rank="species" species="limbata" status="sp. nov.">
|
||
<emphasis box="[538,803,259,285]" italics="true" pageId="2" pageNumber="66">Andrikothelyna limbata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[810,899,259,285]" pageId="2" pageNumber="66">
|
||
<taxonomicNameLabel box="[810,899,259,285]" pageId="2" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName authority="Nozaki, 2023" authorityName="Nozaki" authorityYear="2023" box="[954,1245,259,285]" class="Insecta" family="Staphylinidae" genus="Andrikothelyna" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="66" phylum="Arthropoda" rank="species" species="naomichii" status="sp. nov.">
|
||
<emphasis box="[954,1245,259,285]" italics="true" pageId="2" pageNumber="66">Andrikothelyna naomichii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[1252,1342,259,285]" pageId="2" pageNumber="66">
|
||
<taxonomicNameLabel box="[1252,1342,259,285]" pageId="2" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,933]" pageId="2" pageNumber="66">
|
||
In the original description of the genus
|
||
<taxonomicName authorityName="Pace" authorityYear="2013" box="[667,837,295,321]" class="Insecta" family="Staphylinidae" genus="Speiraphallusa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[667,837,295,321]" italics="true" pageId="2" pageNumber="66">Speiraphallusa</emphasis>
|
||
</taxonomicName>
|
||
, the head was described as “neck moderate” (
|
||
<bibRefCitation author="Pace, R." pageId="2" pageNumber="66" pagination="33 - 63" refId="ref7746" refString="Pace, R. (2013) New distributional data, New Species and Three New Genera of Aleocharinae from Malaysia, Vietnam and Taiwan (Coleoptera: Staphylinidae). Tropical Zoology, 26 (1), 33 - 63. https: // doi. org / 10.1080 / 03946975.2013.775832" type="journal article" year="2013">Pace 2013</bibRefCitation>
|
||
), following which,
|
||
<bibRefCitation author="Santiago-Jimenez, Q. J. & Santiago-Navarro, R." box="[432,950,331,357]" pageId="2" pageNumber="66" pagination="45 - 57" refId="ref7851" refString="Santiago-Jimenez, Q. J. & Santiago-Navarro, R. (2016) A new genus and species of Placusini from a high mountain in Mexico. ZooKeys, 640, 45 - 57. https: // doi. org / 10.3897 / zookeys. 640.10911" type="journal article" year="2016">Santiago-Jiménez & Santiago-Navarro (2016)</bibRefCitation>
|
||
recognized the state “with neck” as one of the characteristics of the genus in key to the genera of the tribe
|
||
<taxonomicName authorityName="Mulsant & Rey" authorityYear="1871" box="[867,970,367,393]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="66" phylum="Arthropoda" rank="tribe" tribe="Placusini">Placusini</taxonomicName>
|
||
(p. 49). However, the neck was absent in all the new species examined in this paper. Perhaps the presence of the “neck” was incorrectly recognized by
|
||
<bibRefCitation author="Pace, R." pageId="2" pageNumber="66" pagination="33 - 63" refId="ref7746" refString="Pace, R. (2013) New distributional data, New Species and Three New Genera of Aleocharinae from Malaysia, Vietnam and Taiwan (Coleoptera: Staphylinidae). Tropical Zoology, 26 (1), 33 - 63. https: // doi. org / 10.1080 / 03946975.2013.775832" type="journal article" year="2013">Pace (2013)</bibRefCitation>
|
||
when he viewed the specimen in the dorsal view.
|
||
<bibRefCitation author="Pires-Silva, M. C." box="[787,992,439,465]" pageId="2" pageNumber="66" pagination="46 - 7477" refId="ref7799" refString="Pires-Silva, M. C. (2023) Euryachen seuzei, a new genus and species of Placusini (Coleoptera: Staphylinidae: Aleocharinae) from Brazil. Zootaxa, 5230 (4), 46 - 7477. https: // doi. org / 10.11646 / zootaxa. 5230.4.4" type="journal article" year="2023">Pires-Silva (2023)</bibRefCitation>
|
||
mentioned the diagnostic characteristic of
|
||
<taxonomicName authorityName="Pace" authorityYear="2013" box="[181,351,475,501]" class="Insecta" family="Staphylinidae" genus="Speiraphallusa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[181,351,475,501]" italics="true" pageId="2" pageNumber="66">Speiraphallusa</emphasis>
|
||
</taxonomicName>
|
||
in the key to genera of
|
||
<taxonomicName authorityName="Mulsant & Rey" authorityYear="1871" box="[609,712,475,501]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="66" phylum="Arthropoda" rank="tribe" tribe="Placusini">Placusini</taxonomicName>
|
||
(p. 469) as “tergite VIII transverse, not modified”, but all species of this genus have teeth on posterior margin of the tergite VIII.
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,933]" pageId="2" pageNumber="66">
|
||
In
|
||
<taxonomicName authority="(Pace 2013)" authorityName="Nozaki" authorityYear="2023" baseAuthorityName="Pace" baseAuthorityYear="2013" box="[228,631,547,573]" class="Insecta" family="Staphylinidae" genus="Andrikothelyna" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="66" phylum="Arthropoda" rank="species" species="orientis" status="comb. nov.">
|
||
<emphasis box="[228,493,547,573]" italics="true" pageId="2" pageNumber="66">Andrikothelyna orientis</emphasis>
|
||
(
|
||
<bibRefCitation author="Pace, R." box="[505,624,547,573]" pageId="2" pageNumber="66" pagination="33 - 63" refId="ref7746" refString="Pace, R. (2013) New distributional data, New Species and Three New Genera of Aleocharinae from Malaysia, Vietnam and Taiwan (Coleoptera: Staphylinidae). Tropical Zoology, 26 (1), 33 - 63. https: // doi. org / 10.1080 / 03946975.2013.775832" type="journal article" year="2013">Pace 2013</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[637,764,547,573]" pageId="2" pageNumber="66">
|
||
<taxonomicNameLabel box="[637,764,547,573]" pageId="2" pageNumber="66" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
only a flagellum of the median lobe of the aedeagus appears to be coiled (as shown in fig.
|
||
<quantity box="[483,541,583,609]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="2" pageNumber="66" unit="in" value="12.0">12 in</quantity>
|
||
<bibRefCitation author="Pace, R." box="[547,667,583,609]" pageId="2" pageNumber="66" pagination="33 - 63" refId="ref7746" refString="Pace, R. (2013) New distributional data, New Species and Three New Genera of Aleocharinae from Malaysia, Vietnam and Taiwan (Coleoptera: Staphylinidae). Tropical Zoology, 26 (1), 33 - 63. https: // doi. org / 10.1080 / 03946975.2013.775832" type="journal article" year="2013">Pace 2013</bibRefCitation>
|
||
), but in
|
||
<taxonomicName authority="Nozaki, 2023" authorityName="Nozaki" authorityYear="2023" box="[759,876,583,609]" class="Insecta" family="Staphylinidae" genus="Andrikothelyna" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="66" phylum="Arthropoda" rank="species" species="limbata" status="sp. nov.">
|
||
<emphasis box="[759,776,583,609]" italics="true" pageId="2" pageNumber="66">A</emphasis>
|
||
.
|
||
<emphasis box="[790,876,583,609]" italics="true" pageId="2" pageNumber="66">limbata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[883,972,583,609]" pageId="2" pageNumber="66">
|
||
<taxonomicNameLabel box="[883,972,583,609]" pageId="2" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, which has a similar shape of the median lobe to that of
|
||
<taxonomicName authorityName="Nozaki" authorityYear="2023" baseAuthorityName="Pace" baseAuthorityYear="2013" box="[311,427,619,645]" class="Insecta" family="Staphylinidae" genus="Andrikothelyna" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="66" phylum="Arthropoda" rank="species" species="orientis" status="comb. nov.">
|
||
<emphasis box="[311,328,619,645]" italics="true" pageId="2" pageNumber="66">A</emphasis>
|
||
.
|
||
<emphasis box="[341,427,619,644]" italics="true" pageId="2" pageNumber="66">orientis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[433,561,619,645]" pageId="2" pageNumber="66">
|
||
<taxonomicNameLabel box="[433,561,619,645]" pageId="2" pageNumber="66" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
not only flagellum but also bulb is spiraled (
|
||
<figureCitation box="[1055,1185,618,645]" captionStart="FIGURES 20–28" captionStartId="8.[151,264,1885,1909]" captionTargetBox="[157,1429,326,1833]" captionTargetId="figure-86@8.[151,1436,309,1861]" captionTargetPageId="8" captionText="FIGURES 20–28. Andrikothelyna limbata sp. nov. 20―abdominal tergite VIII, male; 21―abdominal sternite VIII male; 22― abdominal sternite VIII female; 23–25―median lobe of male aedeagus: 23―lateral view; 24―ventral view; 25―lateral view; 26―paramere of male aedeagus; 27―female spermatheca (Kyushu); 28―female spermatheca (Ishigaki Is.)." figureDoi="http://doi.org/10.5281/zenodo.7836189" httpUri="https://zenodo.org/record/7836189/files/figure.png" pageId="2" pageNumber="66">Figs 23–25</figureCitation>
|
||
). Therefore, it is likely that the figure shown in
|
||
<bibRefCitation author="Pace, R." box="[420,553,655,681]" pageId="2" pageNumber="66" pagination="33 - 63" refId="ref7746" refString="Pace, R. (2013) New distributional data, New Species and Three New Genera of Aleocharinae from Malaysia, Vietnam and Taiwan (Coleoptera: Staphylinidae). Tropical Zoology, 26 (1), 33 - 63. https: // doi. org / 10.1080 / 03946975.2013.775832" type="journal article" year="2013">Pace (2013)</bibRefCitation>
|
||
is drawn based on a misunderstanding of the morphology and the bulb is coiled or spiraled in
|
||
<taxonomicName authorityName="Nozaki" authorityYear="2023" baseAuthorityName="Pace" baseAuthorityYear="2013" box="[307,424,691,717]" class="Insecta" family="Staphylinidae" genus="Andrikothelyna" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="66" phylum="Arthropoda" rank="species" species="orientis" status="comb. nov.">
|
||
<emphasis box="[307,324,691,717]" italics="true" pageId="2" pageNumber="66">A</emphasis>
|
||
.
|
||
<emphasis box="[338,424,691,716]" italics="true" pageId="2" pageNumber="66">orientis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[431,560,691,717]" pageId="2" pageNumber="66">
|
||
<taxonomicNameLabel box="[431,560,691,717]" pageId="2" pageNumber="66" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
The morphology of the flagellum and the bulb itself being spiral shaped in
|
||
<taxonomicName authority="Nozaki, 2023" authorityName="Nozaki" authorityYear="2023" class="Insecta" family="Staphylinidae" genus="Andrikothelyna" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="66" phylum="Arthropoda" rank="species" species="limbata" status="sp. nov.">
|
||
<emphasis box="[1412,1429,691,717]" italics="true" pageId="2" pageNumber="66">A</emphasis>
|
||
.
|
||
<emphasis box="[151,237,727,753]" italics="true" pageId="2" pageNumber="66">limbata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[243,333,727,753]" pageId="2" pageNumber="66">
|
||
<taxonomicNameLabel box="[243,333,727,753]" pageId="2" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, possibly also in
|
||
<taxonomicName authorityName="Nozaki" authorityYear="2023" baseAuthorityName="Pace" baseAuthorityYear="2013" box="[526,643,727,753]" class="Insecta" family="Staphylinidae" genus="Andrikothelyna" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="66" phylum="Arthropoda" rank="species" species="orientis" status="comb. nov.">
|
||
<emphasis box="[526,543,727,753]" italics="true" pageId="2" pageNumber="66">A</emphasis>
|
||
.
|
||
<emphasis box="[557,643,727,752]" italics="true" pageId="2" pageNumber="66">orientis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[649,777,727,753]" pageId="2" pageNumber="66">
|
||
<taxonomicNameLabel box="[649,777,727,753]" pageId="2" pageNumber="66" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, is unique, and no similar morphology is known within the subfamily.
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,933]" box="[199,906,799,825]" pageId="2" pageNumber="66">
|
||
The genus is recorded newly in
|
||
<collectingCountry box="[556,638,799,825]" name="Taiwan" pageId="2" pageNumber="66">Taiwan</collectingCountry>
|
||
and
|
||
<collectingCountry box="[693,757,799,825]" name="Japan" pageId="2" pageNumber="66">Japan</collectingCountry>
|
||
in this study.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[199,921,834,861]" pageId="2" pageNumber="66" type="distribution">
|
||
<paragraph blockId="2.[151,1437,151,933]" box="[199,921,834,861]" pageId="2" pageNumber="66">
|
||
<emphasis bold="true" box="[199,353,834,860]" pageId="2" pageNumber="66">Distribution.</emphasis>
|
||
<collectingCountry box="[359,426,835,861]" name="Japan" pageId="2" pageNumber="66">Japan</collectingCountry>
|
||
,
|
||
<collectingCountry box="[437,522,835,861]" name="Taiwan" pageId="2" pageNumber="66">Taiwan</collectingCountry>
|
||
,
|
||
<collectingCountry box="[533,638,835,861]" name="Malaysia" pageId="2" pageNumber="66">Malaysia</collectingCountry>
|
||
, and
|
||
<collectingCountry box="[697,917,835,861]" name="Papua New Guinea" pageId="2" pageNumber="66">Papua New Guinea</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="2" pageNumber="66" type="discussion">
|
||
<paragraph blockId="2.[151,1437,151,933]" pageId="2" pageNumber="66">
|
||
<emphasis bold="true" box="[199,331,870,896]" pageId="2" pageNumber="66">Bionomics.</emphasis>
|
||
The following three new species were collected from dead vines or dead branches on living trunks.
|
||
<taxonomicName box="[151,367,907,933]" class="Magnoliopsida" family="Malvaceae" genus="Talipariti" kingdom="Plantae" order="Malvales" pageId="2" pageNumber="66" phylum="Tracheophyta" rank="species" species="tiliaceum">
|
||
<emphasis box="[151,367,907,933]" italics="true" pageId="2" pageNumber="66">Talipariti tiliaceum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName box="[422,649,907,933]" class="Magnoliopsida" family="Lauraceae" genus="Machilus" kingdom="Plantae" order="Laurales" pageId="2" pageNumber="66" phylum="Tracheophyta" rank="species" species="thunbergii">
|
||
<emphasis box="[422,649,907,933]" italics="true" pageId="2" pageNumber="66">Machilus thunbergii</emphasis>
|
||
</taxonomicName>
|
||
are recorded as the plant species below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |