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<document id="D5209A8610AE0DB84AE3BB0DF6BBAFCF" ID-CLB-Dataset="28065" ID-DOI="10.11646/zootaxa.4504.4.5" ID-GBIF-Dataset="343a6a67-e8bf-467c-9818-de9621bbe955" ID-ISSN="1175-5326" ID-Zenodo-Dep="2606614" ID-ZooBank="08E1A79F-BAA4-4EAB-9636-FE44BF7EB7F3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553585132792" checkinUser="plazi" docAuthor="Jäger, Peter" docDate="2018" docId="03AB87D4E7616F6D12F9F8F6C8797A20" docLanguage="en" docName="zootaxa.4504.4.5.pdf" docOrigin="Zootaxa 4504 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Systaria elberti" docType="treatment" docVersion="9" lastPageNumber="539" masterDocId="FF92FFACE76F6F62126EFFC4C9517912" masterDocTitle="On the genus Systaria (Araneae: Clubionidae) in Southeast Asia: new species from caves and forests" masterLastPageNumber="544" masterPageNumber="524" pageNumber="538" updateTime="1698712839155" updateUser="ExternalLinkService">
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<mods:title id="EB98431080415A4DD0A29CAEC7AB0174">On the genus Systaria (Araneae: Clubionidae) in Southeast Asia: new species from caves and forests</mods:title>
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<mods:namePart id="EE0A84333EAD5A44C6CEE6A24D603D49">Jäger, Peter</mods:namePart>
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<treatment id="03AB87D4E7616F6D12F9F8F6C8797A20" ID-DOI="http://doi.org/10.5281/zenodo.5988020" ID-GBIF-Taxon="154495205" ID-Zenodo-Dep="5988020" LSID="urn:lsid:plazi:treatment:03AB87D4E7616F6D12F9F8F6C8797A20" httpUri="http://treatment.plazi.org/id/03AB87D4E7616F6D12F9F8F6C8797A20" lastPageId="15" lastPageNumber="539" pageId="14" pageNumber="538">
<subSubSection id="C3186549E7616F6C12F9F8F6C8AC7E5E" box="[151,509,1842,1868]" pageId="14" pageNumber="538" type="nomenclature">
<paragraph id="8BBD36C2E7616F6C12F9F8F6C8AC7E5E" blockId="14.[151,509,1842,1901]" box="[151,509,1842,1868]" pageId="14" pageNumber="538">
<heading id="D0F581AEE7616F6C12F9F8F6C8AC7E5E" bold="true" box="[151,509,1842,1868]" fontSize="11" level="1" pageId="14" pageNumber="538" reason="1">
<taxonomicName id="4C024D41E7616F6C12F9F8F6C8AC7E5E" authority="(Strand 1913)" baseAuthorityName="Strand" baseAuthorityYear="1913" box="[151,509,1842,1868]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="538" phylum="Arthropoda" rank="species" species="elberti">
<emphasis id="B976EAD0E7616F6C12F9F8F6C8AC7E5E" bold="true" box="[151,509,1842,1868]" pageId="14" pageNumber="538">
<emphasis id="B976EAD0E7616F6C12F9F8F6C81D7E5E" bold="true" box="[151,332,1842,1868]" italics="true" pageId="14" pageNumber="538">Systaria elberti</emphasis>
(
<bibRefCitation id="EF934B33E7616F6C1335F8F6C8A47E5E" author="Strand, E." box="[347,501,1842,1868]" pageId="14" pageNumber="538" pagination="113 - 123" refId="ref12465" refString="Strand, E. (1913) Neue indoaustralische und polynesische Spinnen des Senckenbergischen Museums. Archiv fur Naturgeschichte, 79, 113 - 123." type="journal article" year="1913">Strand 1913</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3186549E7616F6C12F9F890C8397E7F" box="[151,360,1876,1901]" pageId="14" pageNumber="538" type="description">
<paragraph id="8BBD36C2E7616F6C12F9F890C8397E7F" blockId="14.[151,509,1842,1901]" box="[151,360,1876,1901]" pageId="14" pageNumber="538">
<figureCitation id="13392A47E7616F6C12F9F890C8467E7F" box="[151,279,1876,1901]" captionStart="FIGURES 4044" captionStartId="17.[151,263,1890,1913]" captionTargetBox="[152,1436,203,1859]" captionTargetId="figure@17.[151,1436,203,1859]" captionTargetPageId="17" captionText="FIGURES 4044. Systaria spp., copulatory organs. 4043 Systaria elberti (Strand 1913) from Indonesia, Lombok Island (40 41 holotype male, 4243 female), 44 Systaria insulana (Rainbow 1809), male from New Hebrides. 4041, 44 Left palp (40 ventral, 41, 44 retrolateral). 42 Epigyne, ventral. 43 Vulva, dorsal. For explanations of a, b see diagnosis (44 modified from Platnick &amp; Bonaldo 1995, not to scale). Abbreviation: CO—copulatory opening." httpUri="https://zenodo.org/record/2606656/files/figure.png" pageId="14" pageNumber="538">Figs 4043</figureCitation>
,
<figureCitation id="13392A47E7616F6C134CF890C8397E7F" box="[290,360,1876,1901]" captionStart-0="FIGURE 45" captionStart-1="FIGURE 46" captionStartId-0="18.[151,250,1002,1024]" captionStartId-1="18.[151,250,1733,1755]" captionTargetBox-0="[150,1436,194,981]" captionTargetBox-1="[150,1436,1106,1712]" captionTargetId-0="figure@18.[150,1436,193,981]" captionTargetId-1="figure@18.[150,1436,1105,1712]" captionTargetPageId-0="18" captionTargetPageId-1="18" captionText-0="FIGURE 45. Records of Systaria spp. in SE Asia described in this paper. Triangles—species recorded from caves, circlesspecies recorded from forest." captionText-1="FIGURE 46. All known records of the genus Systaria (World Spider Catalog 2018) including undescribed species mentioned in Deeleman-Reinhold (2001) according to habitats. Triangles—species recorded from caves, circles—species recorded from forest. See discussion." httpUri-0="https://zenodo.org/record/2606658/files/figure.png" httpUri-1="https://zenodo.org/record/2606660/files/figure.png" pageId="14" pageNumber="538">4546</figureCitation>
</paragraph>
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<subSubSection id="C3186549E7616F6D12F9F858CAEE79BC" lastPageId="15" lastPageNumber="539" pageId="14" pageNumber="538" type="reference_group">
<paragraph id="8BBD36C2E7616F6C12F9F858CA457EE2" blockId="14.[151,1436,1948,2032]" pageId="14" pageNumber="538">
<treatmentCitationGroup id="AB1211ECE7616F6C12F9F858CA457EE2" pageId="14" pageNumber="538">
<taxonomicName id="4C024D41E7616F6C12F9F858C8A57EA0" authority="Strand 1913: 119" authorityName="Strand" authorityPageNumber="119" authorityYear="1913" box="[151,500,1948,1970]" class="Arachnida" family="Clubionidae" genus="Clubiona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="538" phylum="Arthropoda" rank="species" species="elberti">
<emphasis id="B976EAD0E7616F6C12F9F858C86F7EA0" box="[151,318,1948,1970]" italics="true" pageId="14" pageNumber="538">Clubiona elberti</emphasis>
<treatmentCitation id="0AA310D3E7616F6C132BF858C8A57EA0" author="Strand, E." box="[325,500,1948,1970]" page="119" pageId="14" pageNumber="538" year="1913">
<bibRefCitation id="EF934B33E7616F6C132BF858C8957EA0" author="Strand, E." box="[325,452,1948,1970]" pageId="14" pageNumber="538" pagination="113 - 123" refId="ref12465" refString="Strand, E. (1913) Neue indoaustralische und polynesische Spinnen des Senckenbergischen Museums. Archiv fur Naturgeschichte, 79, 113 - 123." type="journal article" year="1913">Strand 1913</bibRefCitation>
: 119
</treatmentCitation>
</taxonomicName>
(description of male;
<typeStatus id="54B98860E7616F6C10B8F858CA7E7EA0" box="[726,815,1948,1970]" pageId="14" pageNumber="538" type="holotype">holotype</typeStatus>
male,
<specimenCount id="9D04FD4BE7616F6C111AF858CA867EA0" box="[884,983,1948,1970]" pageId="14" pageNumber="538" type="female">2 females</specimenCount>
[not mentioned in the original paper, but on the label as questionable, one described as
<taxonomicName id="4C024D41E7616F6C1014F878CB847EC3" authorityName="Strand" authorityYear="1913" box="[634,725,1979,2001]" class="Arachnida" family="Clubionidae" genus="Clubiona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="538" phylum="Arthropoda" rank="species" species="elberti">
<emphasis id="B976EAD0E7616F6C1014F878CB847EC3" box="[634,725,1979,2001]" italics="true" pageId="14" pageNumber="538">S. elberti</emphasis>
</taxonomicName>
by
<treatmentCitation id="0AA310D3E7616F6C1092F87FCD677EC3" author="Deeleman-Reinhold, C. L." box="[764,1078,1979,2001]" page="212" pageId="14" pageNumber="538" year="2001">
<bibRefCitation id="EF934B33E7616F6C1092F87FCD547EC3" author="Deeleman-Reinhold, C. L." box="[764,1029,1979,2001]" pageId="14" pageNumber="538" refId="ref11533" refString="Deeleman-Reinhold, C. L. (2001) Forest spiders of South East Asia: with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae). Brill, Leiden, 591 pp." type="book" year="2001">Deeleman-Reinhold 2001</bibRefCitation>
: 212
</treatmentCitation>
, one belonging to
<taxonomicName id="4C024D41E7616F6C169CF87FCC377EC3" ID-CoL="H6N" box="[1266,1382,1979,2001]" class="Arachnida" family="Theridiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="538" phylum="Arthropoda" rank="family">Theridiidae</taxonomicName>
{sub
<taxonomicName id="4C024D41E7616F6C12A9F81ECBDE7EE2" ID-CoL="6MF" authority="in Deeleman-Reinhold 2001: 209" authorityName="in Deeleman-Reinhold" authorityPageNumber="209" authorityYear="2001" box="[199,655,2010,2032]" class="Arachnida" family="Araneidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="538" phylum="Arthropoda" rank="family">
Araneidae in
<treatmentCitation id="0AA310D3E7616F6C1321F81ECBDE7EE2" author="Deeleman-Reinhold, C. L." box="[335,655,2010,2032]" page="209" pageId="14" pageNumber="538" year="2001">
<bibRefCitation id="EF934B33E7616F6C1321F81ECB067EE2" author="Deeleman-Reinhold, C. L." box="[335,599,2010,2032]" pageId="14" pageNumber="538" refId="ref11533" refString="Deeleman-Reinhold, C. L. (2001) Forest spiders of South East Asia: with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae). Brill, Leiden, 591 pp." type="book" year="2001">Deeleman-Reinhold 2001</bibRefCitation>
: 209
</treatmentCitation>
</taxonomicName>
}]; examined)
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</paragraph>
<paragraph id="8BBD36C2E7606F6D12F9FF5DCAEE79BC" blockId="15.[151,959,152,174]" box="[151,959,152,174]" pageId="15" pageNumber="539">
<treatmentCitationGroup id="AB1211ECE7606F6D12F9FF5DCAEE79BC" box="[151,959,152,174]" pageId="15" pageNumber="539">
<taxonomicName id="4C024D41E7606F6D12F9FF5DC86379BC" baseAuthorityName="Strand" baseAuthorityYear="1913" box="[151,306,152,174]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="539" phylum="Arthropoda" rank="species" species="elberti">
<emphasis id="B976EAD0E7606F6D12F9FF5DC86379BC" box="[151,306,152,174]" italics="true" pageId="15" pageNumber="539">Systaria elberti</emphasis>
</taxonomicName>
;
<treatmentCitation id="0AA310D3E7606F6D132EFF5CCB2979BC" author="Deeleman-Reinhold, C. L." box="[320,632,152,174]" page="209" pageId="15" pageNumber="539" year="2001">
<bibRefCitation id="EF934B33E7606F6D132EFF5CCB1879BC" author="Deeleman-Reinhold, C. L." box="[320,585,152,174]" pageId="15" pageNumber="539" refId="ref11533" refString="Deeleman-Reinhold, C. L. (2001) Forest spiders of South East Asia: with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae). Brill, Leiden, 591 pp." type="book" year="2001">Deeleman-Reinhold 2001</bibRefCitation>
: 209
</treatmentCitation>
(transfer, description of female)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3186549E7606F6D12F9FF1EC8077805" pageId="15" pageNumber="539" type="description">
<paragraph id="8BBD36C2E7606F6D12F9FF1EC8077805" blockId="15.[151,1436,218,819]" pageId="15" pageNumber="539">
<emphasis id="B976EAD0E7606F6D12F9FF1EC98979E1" bold="true" box="[151,216,218,243]" pageId="15" pageNumber="539">Note:</emphasis>
<bibRefCitation id="EF934B33E7606F6D128EFF1EC82B79E1" author="Strand, E." box="[224,378,218,243]" pageId="15" pageNumber="539" pagination="113 - 123" refId="ref12465" refString="Strand, E. (1913) Neue indoaustralische und polynesische Spinnen des Senckenbergischen Museums. Archiv fur Naturgeschichte, 79, 113 - 123." type="journal article" year="1913">Strand (1913)</bibRefCitation>
did not describe the female as mentioned in the
<bibRefCitation id="EF934B33E7606F6D11CEFF1ECDBB79E1" author="World Spider Catalog" box="[928,1258,218,243]" pageId="15" pageNumber="539" refId="ref12552" refString="World Spider Catalog (2018) World Spider Catalog. Uersion 19.0. Natural History Museum Bern. Available from: http: // wsc. nmbe. ch (accessed 30 May 2018) https: // doi. org / 10.24436 / 2" type="book" year="2018">World Spider Catalog (2018)</bibRefCitation>
, but Deeleman-
<bibRefCitation id="EF934B33E7606F6D12F9FF3AC8037805" author="Deeleman-Reinhold, C. L." box="[151,338,254,279]" pageId="15" pageNumber="539" refId="ref11533" refString="Deeleman-Reinhold, C. L. (2001) Forest spiders of South East Asia: with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae). Brill, Leiden, 591 pp." type="book" year="2001">Reinhold (2001)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3186549E7606F6D12A9FEE6CBEF7B6D" pageId="15" pageNumber="539" type="diagnosis">
<paragraph id="8BBD36C2E7606F6D12A9FEE6CBEF7B6D" blockId="15.[151,1436,218,819]" pageId="15" pageNumber="539">
<emphasis id="B976EAD0E7606F6D12A9FEE6C8E97829" bold="true" box="[199,440,290,315]" pageId="15" pageNumber="539">Amended diagnosis.</emphasis>
Small to medium sized spiders with body length of males: 9.4, of females: 10.1. Males are similar to
<taxonomicName id="4C024D41E7606F6D135DFE83C8E1784D" baseAuthorityName="Rainbow" baseAuthorityYear="1901" box="[307,432,326,351]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="539" phylum="Arthropoda" rank="species" species="insulana">
<emphasis id="B976EAD0E7606F6D135DFE83C8E1784D" box="[307,432,326,351]" italics="true" pageId="15" pageNumber="539">S. insulana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C024D41E7606F6D1389FE83CB35784C" authority="Jäger, 2018" authorityName="Jäger" authorityYear="2018" box="[487,612,327,350]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="539" phylum="Arthropoda" rank="species" species="princesa" status="sp. nov.">
<emphasis id="B976EAD0E7606F6D1389FE83CB35784C" box="[487,612,327,350]" italics="true" pageId="15" pageNumber="539">S. princesa</emphasis>
</taxonomicName>
<emphasis id="B976EAD0E7606F6D1002FE82CB8F784D" bold="true" box="[620,734,326,351]" pageId="15" pageNumber="539">
<taxonomicNameLabel id="A24557ABE7606F6D1002FE82CB8F784D" box="[620,734,326,351]" pageId="15" pageNumber="539" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
in having a similar conformation of the palp including arising point and shape of embolus and the simple, stretched RTA (
<figureCitation id="13392A47E7606F6D115DFEAECAE77891" box="[819,950,362,387]" captionStart="FIGURES 4044" captionStartId="17.[151,263,1890,1913]" captionTargetBox="[152,1436,203,1859]" captionTargetId="figure@17.[151,1436,203,1859]" captionTargetPageId="17" captionText="FIGURES 4044. Systaria spp., copulatory organs. 4043 Systaria elberti (Strand 1913) from Indonesia, Lombok Island (40 41 holotype male, 4243 female), 44 Systaria insulana (Rainbow 1809), male from New Hebrides. 4041, 44 Left palp (40 ventral, 41, 44 retrolateral). 42 Epigyne, ventral. 43 Vulva, dorsal. For explanations of a, b see diagnosis (44 modified from Platnick &amp; Bonaldo 1995, not to scale). Abbreviation: CO—copulatory opening." httpUri="https://zenodo.org/record/2606656/files/figure.png" pageId="15" pageNumber="539">Figs 4041</figureCitation>
), but may be distinguished from the former by the male RTA with base relatively very wide (=a) and very short distal part (=b, best seen in retrolateral view,
<figureCitation id="13392A47E7606F6D12F9FE76C9BF78D9" box="[151,238,434,459]" captionStart="FIGURES 4044" captionStartId="17.[151,263,1890,1913]" captionTargetBox="[152,1436,203,1859]" captionTargetId="figure@17.[151,1436,203,1859]" captionTargetPageId="17" captionText="FIGURES 4044. Systaria spp., copulatory organs. 4043 Systaria elberti (Strand 1913) from Indonesia, Lombok Island (40 41 holotype male, 4243 female), 44 Systaria insulana (Rainbow 1809), male from New Hebrides. 4041, 44 Left palp (40 ventral, 41, 44 retrolateral). 42 Epigyne, ventral. 43 Vulva, dorsal. For explanations of a, b see diagnosis (44 modified from Platnick &amp; Bonaldo 1995, not to scale). Abbreviation: CO—copulatory opening." httpUri="https://zenodo.org/record/2606656/files/figure.png" pageId="15" pageNumber="539">Fig. 41</figureCitation>
; with a/b=1.4; 1.0 in
<taxonomicName id="4C024D41E7606F6D1399FE77CB2878D9" baseAuthorityName="Rainbow" baseAuthorityYear="1901" box="[503,633,434,459]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="539" phylum="Arthropoda" rank="species" species="insulana">
<emphasis id="B976EAD0E7606F6D1399FE77CB2878D9" box="[503,633,434,459]" italics="true" pageId="15" pageNumber="539">S. insulana</emphasis>
</taxonomicName>
,
<figureCitation id="13392A47E7606F6D10E2FE76CBB378D9" box="[652,738,434,459]" captionStart="FIGURES 4044" captionStartId="17.[151,263,1890,1913]" captionTargetBox="[152,1436,203,1859]" captionTargetId="figure@17.[151,1436,203,1859]" captionTargetPageId="17" captionText="FIGURES 4044. Systaria spp., copulatory organs. 4043 Systaria elberti (Strand 1913) from Indonesia, Lombok Island (40 41 holotype male, 4243 female), 44 Systaria insulana (Rainbow 1809), male from New Hebrides. 4041, 44 Left palp (40 ventral, 41, 44 retrolateral). 42 Epigyne, ventral. 43 Vulva, dorsal. For explanations of a, b see diagnosis (44 modified from Platnick &amp; Bonaldo 1995, not to scale). Abbreviation: CO—copulatory opening." httpUri="https://zenodo.org/record/2606656/files/figure.png" pageId="15" pageNumber="539">Fig. 44</figureCitation>
) and from the latter by the relatively shorter RTA base in comparison to the maximal cymbium width in retrolateral view, i.e. a &lt;width (a&gt; width in
<taxonomicName id="4C024D41E7606F6D16F8FE13CC4378FC" authority="Jäger, 2018" authorityName="Jäger" authorityYear="2018" box="[1174,1298,471,494]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="539" phylum="Arthropoda" rank="species" species="princesa" status="sp. nov.">
<emphasis id="B976EAD0E7606F6D16F8FE13CC4378FC" box="[1174,1298,471,494]" italics="true" pageId="15" pageNumber="539">S. princesa</emphasis>
</taxonomicName>
<emphasis id="B976EAD0E7606F6D1777FE12CCDD78FD" bold="true" box="[1305,1420,470,495]" pageId="15" pageNumber="539">
<taxonomicNameLabel id="A24557ABE7606F6D1777FE12CCDD78FD" box="[1305,1420,470,495]" pageId="15" pageNumber="539" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
). Females are similar to those of
<taxonomicName id="4C024D41E7606F6D106AFE3FCB267B01" box="[516,631,506,531]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="539" phylum="Arthropoda" rank="species" species="dentata">
<emphasis id="B976EAD0E7606F6D106AFE3FCB267B01" box="[516,631,506,531]" italics="true" pageId="15" pageNumber="539">S. dentata</emphasis>
</taxonomicName>
in having similarly bent copulatory ducts leading to connecting tubes between bursae and spermathecae as well as relatively small bursae, but may be distinguished by copulatory ducts roughly right-angled (U-shaped in
<taxonomicName id="4C024D41E7606F6D1044FD87CBCC7B49" box="[554,669,578,603]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="539" phylum="Arthropoda" rank="species" species="dentata">
<emphasis id="B976EAD0E7606F6D1044FD87CBCC7B49" box="[554,669,578,603]" italics="true" pageId="15" pageNumber="539">S. dentata</emphasis>
</taxonomicName>
) and copulatory openings surrounded by strongly sclerotised rims opened medially (with closed rims in
<taxonomicName id="4C024D41E7606F6D1050FDA3CBFF7B6D" box="[574,686,614,639]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="539" phylum="Arthropoda" rank="species" species="dentata">
<emphasis id="B976EAD0E7606F6D1050FDA3CBFF7B6D" box="[574,686,614,639]" italics="true" pageId="15" pageNumber="539">S. dentata</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3186549E7606F6D12A9FD4EC8B97BD5" pageId="15" pageNumber="539" type="description">
<paragraph id="8BBD36C2E7606F6D12A9FD4EC8B97BD5" blockId="15.[151,1436,218,819]" pageId="15" pageNumber="539">
<emphasis id="B976EAD0E7606F6D12A9FD4EC8087BB1" bold="true" box="[199,345,650,675]" pageId="15" pageNumber="539">Description.</emphasis>
See
<bibRefCitation id="EF934B33E7606F6D13FBFD4ECB9F7BB1" author="Deeleman-Reinhold, C. L." box="[405,718,650,675]" pageId="15" pageNumber="539" refId="ref11533" refString="Deeleman-Reinhold, C. L. (2001) Forest spiders of South East Asia: with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae). Brill, Leiden, 591 pp." type="book" year="2001">Deeleman-Reinhold (2001)</bibRefCitation>
. One pair of slit sensilla and only few sclerotised spheres are present in the female epigyne.
</paragraph>
</subSubSection>
<subSubSection id="C3186549E7606F6D12A9FD16C8797A20" pageId="15" pageNumber="539" type="distribution">
<paragraph id="8BBD36C2E7606F6D12A9FD16CADB7BF9" blockId="15.[151,1436,218,819]" box="[199,906,722,747]" pageId="15" pageNumber="539">
<emphasis id="B976EAD0E7606F6D12A9FD16C8317BF9" bold="true" box="[199,352,722,747]" pageId="15" pageNumber="539">Distribution.</emphasis>
Known only from the
<typeStatus id="54B98860E7606F6D100EFD17CBC17BF9" box="[608,656,723,747]" pageId="15" pageNumber="539">type</typeStatus>
locality (
<figureCitation id="13392A47E7606F6D1094FD16CA2C7BF9" box="[762,893,722,747]" captionStart-0="FIGURE 45" captionStart-1="FIGURE 46" captionStartId-0="18.[151,250,1002,1024]" captionStartId-1="18.[151,250,1733,1755]" captionTargetBox-0="[150,1436,194,981]" captionTargetBox-1="[150,1436,1106,1712]" captionTargetId-0="figure@18.[150,1436,193,981]" captionTargetId-1="figure@18.[150,1436,1105,1712]" captionTargetPageId-0="18" captionTargetPageId-1="18" captionText-0="FIGURE 45. Records of Systaria spp. in SE Asia described in this paper. Triangles—species recorded from caves, circlesspecies recorded from forest." captionText-1="FIGURE 46. All known records of the genus Systaria (World Spider Catalog 2018) including undescribed species mentioned in Deeleman-Reinhold (2001) according to habitats. Triangles—species recorded from caves, circles—species recorded from forest. See discussion." httpUri-0="https://zenodo.org/record/2606658/files/figure.png" httpUri-1="https://zenodo.org/record/2606660/files/figure.png" pageId="15" pageNumber="539">Figs 4546</figureCitation>
).
</paragraph>
<paragraph id="8BBD36C2E7606F6D12A9FD32C8797A20" blockId="15.[151,1436,218,819]" pageId="15" pageNumber="539">
<emphasis id="B976EAD0E7606F6D12A9FD32C8D77A1D" bold="true" box="[199,390,758,783]" pageId="15" pageNumber="539">Natural history.</emphasis>
The label suggests that specimens were found in the forest in an altitude between
<date id="FFBC1002E7606F6D175DFD33C99E7A21" pageId="15" pageNumber="539" value="1000" valueMax="1500">1000 and 1500</date>
metres.
</paragraph>
</subSubSection>
</treatment>
</document>