521 lines
76 KiB
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521 lines
76 KiB
XML
<document id="395D04CAD747CE6F716789C7B71656E6" ID-DOI="10.11646/zootaxa.1074.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5050779" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625108368558" checkinUser="felipe" docAuthor="Ponder, W. F., Clark, S. A., Eberhard, S. & Studdert, J. B." docDate="2005" docId="03AB87BA2D23A22BE41FFCB87AB0F9F2" docLanguage="en" docName="zt01074p066.pdf" docOrigin="Zootaxa 1074 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1074.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Pseudotricula progenitor Ponder & Clark & Eberhard & Studdert 2005, n.sp." docType="treatment" docVersion="4" lastPageNumber="40" masterDocId="FF92FFC22D02A20CE517FFCC7E3DFF83" masterDocTitle="A radiation of hydrobiid snails in the caves and streams at Precipitous Bluff southwest Tasmania, Australia (Mollusca: Caenogastropoda: Rissooidea: Hydrobiidae s. l.)," masterLastPageNumber="66" masterPageNumber="1" pageNumber="34" updateTime="1698981223375" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="EEBD4DED988A0D982FE869CFC7DD8AC4">A radiation of hydrobiid snails in the caves and streams at Precipitous Bluff southwest Tasmania, Australia (Mollusca: Caenogastropoda: Rissooidea: Hydrobiidae s. l.),</mods:title>
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<mods:name id="695C1E0610E59FD514AEE40AF548A0C5" type="personal">
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<mods:namePart id="205300882978E3562643C7E5C430C83B">Ponder, W. F.</mods:namePart>
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<mods:affiliation id="4C310DE008F28772F93E3D8D033BA1E2">Australian Museum, 6 College Street, Sydney, NSW, 2010, Australia. Email: winstonp @ austmus. gov. au Previously Australian Museum, now Department of Biological Sciences, Biodiversity and Systematics. University of Alabama, Box 870345, Tuscaloosa, AL 35487, United States of America Department of Conservation and Land Management, Science Division, PO Box 51, Wanneroo, WA 6065, Australia</mods:affiliation>
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<mods:namePart id="3D638791C6921CCE51C99DF0D864C609">Clark, S. A.</mods:namePart>
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<mods:namePart id="144B54CAE261531D21A0D956B588E568">Eberhard, S.</mods:namePart>
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<mods:namePart id="8123EEF5138CE951205F3BC4297E63BC">Studdert, J. B.</mods:namePart>
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<mods:affiliation id="08CA24044E02358F47D63FB8B0E6ECBC">Australian Museum, 6 College Street, Sydney, NSW, 2010, Australia. Email: winstonp @ austmus. gov. au Previously Australian Museum, now Department of Biological Sciences, Biodiversity and Systematics. University of Alabama, Box 870345, Tuscaloosa, AL 35487, United States of America Department of Conservation and Land Management, Science Division, PO Box 51, Wanneroo, WA 6065, Australia</mods:affiliation>
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<mods:title id="F7777FDD358801D0B0086DCF4869A251">Zootaxa</mods:title>
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<mods:date id="FC20E83FF61657278CA806C71FC5C16D">2005</mods:date>
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<mods:number id="E992E822F4BF0972C53C814F60764134">1074</mods:number>
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<heading id="D0F581C02D23A22DE41FFCB87C45FC0D" bold="true" box="[264,632,884,910]" fontSize="11" level="1" pageId="33" pageNumber="34" reason="1">
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<taxonomicName id="4C024D2F2D23A22DE41FFCB87C08FC0D" authority="Ponder & Clark & Eberhard & Studdert, 2005" authorityName="Ponder & Clark & Eberhard & Studdert" authorityYear="2005" box="[264,565,884,910]" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="progenitor" status="sp. nov.">
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<emphasis id="B976EABE2D23A22DE41FFCB87C08FC0D" bold="true" box="[264,565,884,910]" italics="true" pageId="33" pageNumber="34">Pseudotricula progenitor</emphasis>
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<paragraph id="8BBD36AC2D23A22DE41FFC5B7A2FFC32" blockId="33.[264,1042,884,945]" box="[264,1042,919,945]" pageId="33" pageNumber="34">
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<figureCitation id="13392A292D23A22DE41FFC5B7F8CFC32" box="[264,433,919,945]" captionStart="FIGURE 6" captionStartId="16.[264,357,1641,1662]" captionTargetBox="[302,1285,329,1616]" captionTargetId="figure-146@16.[302,1285,329,1633]" captionTargetPageId="16" captionText="FIGURE 6. Protoconchs and teleoconch microsculpture of Pseudotricula and Nanocochlea species. A–D, Pseudotricula expandolabra, A, B, D, C.201493, paratype, main streamway, Bauhaus, C, C.201495, Persephone Streamway, Bauhaus; E–G, Pseudotricula conica, E, F, C.201464, Cane Toad Abuse Streamway, Damper Cave, G, Persephone Pot, Bauhaus; Honey and Cream Streamway, Damper Cave; H, Nanocochlea exigua, C.201475; I, J, Nanocochlea stylesae, C.201476, paratypes, Malani Creek; K–M, Nanocochlea damperensis, K, L, C.203685, paratypes, first crossing, Damper Creek, M, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; N, O, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, B, E, F, H, K–N, protoconch; C, D, G, I, J, O, microsculpture. Scales: A, C–E, J, K, M, O, 200 m; B, F–I, L, N, 100 m." figureDoi="http://doi.org/10.5281/zenodo.5050797" httpUri="https://zenodo.org/record/5050797/files/figure.png" pageId="33" pageNumber="34">Figures 6N, O</figureCitation>
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;
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<figureCitation id="13392A292D23A22DE4A9FC5B7FD8FC32" box="[446,485,919,945]" captionStart="FIGURE 8" captionStartId="20.[264,357,1612,1633]" captionTargetBox="[341,1270,339,1585]" captionTargetId="figure-146@20.[299,1287,324,1608]" captionTargetPageId="20" captionText="FIGURE 8. Pallial cavities of male specimens of Pseudotricula and Nanocochlea species. A, Pseudotricula expandolabra, C.201493, paratype, Persephone Streamway, Bauhaus; B, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; C, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; D, E, Pseudotricula conica, D, C.203680, main streamway near entrance, Damper Cave, E, Cane Toad Abuse Streamway, Damper Cave; F, Nanocochlea exigua, C.203677, paratype, Cane Toad Abuse Streamway, Damper Cave; G, Nanocochlea damperensis, C.203685, paratype, first crossing, Damper Creek; H, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. Scales: A–D, F–H, 500 m, E, 1 mm. Abbreviations: a, anus; ct, ctenidium; ev, efferent vein; os, osphradium; pc, pericardium; pe, pallial edge; ppw, posterior pallial wall; r, rectum; rg, renal gland. The stippled area represents the hypobranchial gland." figureDoi="http://doi.org/10.5281/zenodo.5050803" httpUri="https://zenodo.org/record/5050803/files/figure.png" pageId="33" pageNumber="34">8H</figureCitation>
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;
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<figureCitation id="13392A292D23A22DE4E6FC5B7C12FC32" box="[497,559,919,945]" captionStart="FIGURE 9" captionStartId="21.[264,357,1723,1744]" captionTargetBox="[350,1309,380,1631]" captionTargetId="figure-97@21.[299,1342,370,1653]" captionTargetPageId="21" captionText="FIGURE 9. Head, penis and stomach of Pseudotricula species. A, D, Pseudotricula conica, C.203680, main streamway near entrance, Damper Cave; C, Pseudotricula expandolabra, C.201493, paratype, Persephone Streamway, Bauhaus; B, E, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, B, head and penis; C–E, stomach. Scales: A–E, 500 m. Abbreviations: dgo, opening from stomach to digestive gland; ey, eye; int, intestine; oes, oesophagus, p, penis; pg, penial duck; sn, snout; ss, style sack; st, stomach; tn, tentacle." figureDoi="http://doi.org/10.5281/zenodo.5050807" httpUri="https://zenodo.org/record/5050807/files/figure.png" pageId="33" pageNumber="34">9B,E</figureCitation>
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;
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<figureCitation id="13392A292D23A22DE72BFC5B7CB4FC32" box="[572,649,919,945]" captionStart="FIGURE 14" captionStartId="35.[264,357,1682,1703]" captionTargetBox="[353,1231,287,1666]" captionTargetId="figure-112@35.[336,1250,272,1680]" captionTargetPageId="35" captionText="FIGURE 14. Shells of Nanocochlea species and Pseudotricula progenitor. A C, Nanocochlea exigua, A, holotype, C.439393, Cane Toad Abuse Streamway, Damper Cave; B, C, C.201465, main streamway, Damper Cave; D, Nanocochlea stylesae, holotype, C.439398, Malani Creek; E–G, Nanocochlea damperensis, E, C.203685, paratype, first crossing, Damper Creek; F, holotype, C.439399, first crossing, Damper Creek, F, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; H, I, Pseudotricula progenitor, H, paratype, C.439397, Persephone Streamway, Bauhaus; I, holotype, C.439396, Persephone Streamway, Bauhaus. Scales: 500 m." figureDoi="http://doi.org/10.5281/zenodo.5050821" httpUri="https://zenodo.org/record/5050821/files/figure.png" pageId="33" pageNumber="34">14H, I</figureCitation>
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;
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<figureCitation id="13392A292D23A22DE781FC5B7CDBFC32" box="[662,742,919,945]" captionStart="FIGURE 15" captionStartId="36.[264,357,1703,1724]" captionTargetBox="[304,1298,334,1666]" captionTargetId="figure-104@36.[284,1304,324,1686]" captionTargetPageId="36" captionText="FIGURE 15. Shells of Nanocochlea species and Pseudotricula progenitor. A C, Nanocochlea exigua, A, C.203674, outside Damper Cave, Damper Creek B, C.166855, Persephone Pot, Bauhaus, C, C.201465, main streamway, Damper Cave; D–F, Nanocochlea stylesae, D, C.201476, paratype, Malani Creek, E, F, C.203686, first crossing, Damper Creek; GI, Nanocochlea damperensis, G,H, C.303685, seepages at Bauhaus entrance, I, C.203685, paratype, first crossing, Damper Creek; J–L, Pseudotricula progenitor, J, L, C.201463, paratype, Persephone streamway, Bauhaus, K, C.201454, Persephone, Bauhaus. Scales: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5050823" httpUri="https://zenodo.org/record/5050823/files/figure.png" pageId="33" pageNumber="34">15J, K</figureCitation>
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;
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<figureCitation id="13392A292D23A22DE7E5FC5B7D75FC32" box="[754,840,919,945]" captionStart="FIGURE 16" captionStartId="37.[264,357,1113,1134]" captionTargetBox="[305,1279,351,1060]" captionTargetId="figure-243@37.[300,1287,333,1089]" captionTargetPageId="37" captionText="FIGURE 16. Opercula of Nanocochlea species and Pseudotricula progenitor. A C, Nanocochlea exigua, C.203677, paratypes, Cane Toad Abuse Streamway, Damper Cave; D F, Nanocochlea damperensis; C–E, C.203685, paratypes, first crossing, Damper Creek; F, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; G, H, Pseudotricula progenitor, C.201821, main streamway, Bauhaus; A, D, G, outer side; B, C, E, F, H, inner side; Scales: A, B, D, 200 m, C, E–H, 100 m." figureDoi="http://doi.org/10.5281/zenodo.5052059" httpUri="https://zenodo.org/record/5052059/files/figure.png" pageId="33" pageNumber="34">16G, H</figureCitation>
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;
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<figureCitation id="13392A292D23A22DE643FC5B7D97FC32" box="[852,938,919,945]" captionStart="FIGURE 17" captionStartId="38.[264,357,1649,1670]" captionTargetBox="[317,1274,286,1638]" captionTargetId="figure-154@38.[313,1274,286,1643]" captionTargetPageId="38" captionText="FIGURE 17. Radulae of Nanocochlea species Pseudotricula progenitor. A C, Nanocochlea exigua, A, C.201475, Honey and Cream Streamway, Damper Cave, B, D, C.201465, main streamway, Damper Cave; D– E, Nanocochlea stylesae, C.201476, paratypes, Malani Creek; G–K, Nanocochlea damperensis, G, H, C.203685, paratypes, first crossing, Damper Creek, I–K, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; L–O, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, E, central and lateral teeth; B, F, I, M, central, lateral, inner and outer marginal teeth; C, D, J, N central teeth; G, L, lateral teeth; H, central lateral inner and outer marginal teeth; K, outer marginal teeth. Scales: A, G–H, K, L, 10 m; B, I, M, 20 m; C, D, J, N, 5 m." figureDoi="http://doi.org/10.5281/zenodo.5050829" httpUri="https://zenodo.org/record/5050829/files/figure.png" pageId="33" pageNumber="34">17L–O</figureCitation>
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;
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<figureCitation id="13392A292D23A22DE6A1FC5B7A33FC32" box="[950,1038,919,945]" captionStart="FIGURE 18" captionStartId="39.[264,357,1046,1067]" captionTargetBox="[308,1274,317,1017]" captionTargetId="figure-219@39.[254,1333,301,1043]" captionTargetPageId="39" captionText="FIGURE 18. Female reproductive system of Nanocochlea species and Pseudotricula progenitor. A, Nanocochlea exigua, C.203677, paratype, Cane Toad Abuse Streamway, Damper Cave; B, Nanocochlea damperensis, C.203685, paratype, first crossing, Damper Creek; C, D, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. Scales: 500 m. Abbreviations: ag, albumen gland; bc, bursa copulatrix; bd, bursal duct; cg, capsule gland; co, coiled oviduct; po, genital opening; pw, pallial wall; sr, seminal receptacle; vc, ventral channel." figureDoi="http://doi.org/10.5281/zenodo.5050831" httpUri="https://zenodo.org/record/5050831/files/figure.png" pageId="33" pageNumber="34">18C, D</figureCitation>
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.
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</paragraph>
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<paragraph id="8BBD36AC2D23A22DE41FFC2B7F94FB83" blockId="33.[264,1323,999,1145]" box="[264,425,999,1024]" pageId="33" pageNumber="34">
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<emphasis id="B976EABE2D23A22DE41FFC2B7F94FB83" box="[264,425,999,1024]" italics="true" pageId="33" pageNumber="34">Type material</emphasis>
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</paragraph>
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<paragraph id="8BBD36AC2D23A22DE42FFBC37AC7FBAA" blockId="33.[264,1323,999,1145]" box="[312,1274,1039,1065]" pageId="33" pageNumber="34">
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<materialsCitation id="3B6A3CF12D23A22DE42FFBC37AC7FBAA" box="[312,1274,1039,1065]" collectingDate="1991-12-23" collectionCode="AMS" pageId="33" pageNumber="34" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54B9880E2D23A22DE42FFBC37F94FBAA" box="[312,425,1039,1065]" pageId="33" pageNumber="34" type="holotype">Holotype</typeStatus>
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:
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<collectionCode id="ED13AE692D23A22DE4A3FBC37FCEFBAA" box="[436,499,1039,1065]" pageId="33" pageNumber="34">AMS</collectionCode>
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C.439396, Bauhaus, Persephone streamway, stn 8,
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<date id="FFBC106C2D23A22DE146FBC37ACBFBAA" box="[1105,1270,1039,1065]" pageId="33" pageNumber="34" value="1991-12-23">
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<collectingDate id="EFF8E9842D23A22DE146FBC37ACBFBAA" box="[1105,1270,1039,1065]" pageId="33" pageNumber="34" value="1991-12-23">23 DEC 1991</collectingDate>
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</date>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BBD36AC2D23A22DE42FFBFB7CF2FBFA" blockId="33.[264,1323,999,1145]" pageId="33" pageNumber="34">
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<materialsCitation id="3B6A3CF12D23A22DE42FFBFB7A35FBD2" box="[312,1032,1079,1105]" collectionCode="AMS" pageId="33" pageNumber="34" specimenCount="1" typeStatus="paratype">
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<typeStatus id="54B9880E2D23A22DE42FFBFB7F93FBD2" box="[312,430,1079,1105]" pageId="33" pageNumber="34" type="paratype">Paratypes</typeStatus>
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:
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<collectionCode id="ED13AE692D23A22DE4AEFBFB7FC4FBD2" box="[441,505,1079,1105]" pageId="33" pageNumber="34">AMS</collectionCode>
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C.201463, same data, 30+ (11 dry, 20+ wet)
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</materialsCitation>
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;
|
||
<materialsCitation id="3B6A3CF12D23A22DE103FBFB7CF7FBFA" collectionCode="QVM, AMS" pageId="33" pageNumber="34" specimenCount="1" typeStatus="paratype">
|
||
<collectionCode id="ED13AE692D23A22DE103FBFB7A66FBD2" box="[1044,1115,1079,1105]" country="Australia" httpUri="http://grbio.org/cool/2s8k-2m7w" name="Queen Victoria Museum" pageId="33" pageNumber="34">QVM</collectionCode>
|
||
, 9:20540 (5 wet);
|
||
<collectionCode id="ED13AE692D23A22DE41FFB937F7AFBFA" box="[264,327,1119,1145]" pageId="33" pageNumber="34">AMS</collectionCode>
|
||
C.439397, same data (
|
||
<figureCitation id="13392A292D23A22DE743FB937CFCFBFA" box="[596,705,1119,1145]" captionStart="FIGURE 14" captionStartId="35.[264,357,1682,1703]" captionTargetBox="[353,1231,287,1666]" captionTargetId="figure-112@35.[336,1250,272,1680]" captionTargetPageId="35" captionText="FIGURE 14. Shells of Nanocochlea species and Pseudotricula progenitor. A C, Nanocochlea exigua, A, holotype, C.439393, Cane Toad Abuse Streamway, Damper Cave; B, C, C.201465, main streamway, Damper Cave; D, Nanocochlea stylesae, holotype, C.439398, Malani Creek; E–G, Nanocochlea damperensis, E, C.203685, paratype, first crossing, Damper Creek; F, holotype, C.439399, first crossing, Damper Creek, F, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; H, I, Pseudotricula progenitor, H, paratype, C.439397, Persephone Streamway, Bauhaus; I, holotype, C.439396, Persephone Streamway, Bauhaus. Scales: 500 m." figureDoi="http://doi.org/10.5281/zenodo.5050821" httpUri="https://zenodo.org/record/5050821/files/figure.png" pageId="33" pageNumber="34">Fig. 14H</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBD36AC2D23A22DE41FFB637C95FB4B" blockId="33.[264,1322,1199,1545]" box="[264,680,1199,1224]" pageId="33" pageNumber="34">
|
||
<emphasis id="B976EABE2D23A22DE41FFB637C95FB4B" box="[264,680,1199,1224]" italics="true" pageId="33" pageNumber="34">Other material examined (all AMS)</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BBD36AC2D23A22DE42FFB1B7DB5FA3A" blockId="33.[264,1322,1199,1545]" pageId="33" pageNumber="34">
|
||
<emphasis id="B976EABE2D23A22DE42FFB1B7FA3FB73" box="[312,414,1239,1264]" italics="true" pageId="33" pageNumber="34">Bauhaus</emphasis>
|
||
: C.438369, stn PB61 (5 dry); C.201821, main streamway, stn PB61A (10 dry, 20+ wet); C.166856, Persephone Pot, stn PB178R (5 dry, 25 wet); C.201812, same locality, middle and upper streamway, stn PB172a.2 (20+ dry, 20+ wet); C.438370, C.201816, same data (20+ dry, 20+ wet); same locality, PB172A (20+ dry); C.201454, Persephone, stn 7 (4 dry, 20+ wet); C.201468, Persephone streamway, stn 9 (15 dry, 20+ wet); C.201484, same locality, stn 10 (6 dry, 20+ wet).
|
||
</paragraph>
|
||
<paragraph id="8BBD36AC2D23A22DE42FFA0B7D54F98A" blockId="33.[264,1322,1199,1545]" pageId="33" pageNumber="34">
|
||
<emphasis id="B976EABE2D23A22DE42FFA0B7FE1FA63" box="[312,476,1479,1504]" italics="true" pageId="33" pageNumber="34">Damper Cave</emphasis>
|
||
: C.203682, Main streamway near entrance, stn PB11A (3 dry, 5 wet); C.201458, main streamway, stn 6 (14 dry, 20+ wet).
|
||
</paragraph>
|
||
<paragraph id="8BBD36AC2D23A22DE41FF9F37FB8F9DB" blockId="33.[264,1323,1599,1745]" box="[264,389,1599,1624]" pageId="33" pageNumber="34">
|
||
<emphasis id="B976EABE2D23A22DE41FF9F37FB8F9DB" box="[264,389,1599,1624]" italics="true" pageId="33" pageNumber="34">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BBD36AC2D23A22DE42FF9AB7F8EF952" blockId="33.[264,1323,1599,1745]" pageId="33" pageNumber="34">
|
||
<emphasis id="B976EABE2D23A22DE42FF9AB7F8AF903" box="[312,439,1639,1664]" italics="true" pageId="33" pageNumber="34">Progenitor</emphasis>
|
||
Latin, founder of a family, ancestor. Refers to the apparently basal position (and plesiomorphic nature) of this species in relation to the other taxa included in
|
||
<taxonomicName id="4C024D2F2D23A22DE41FF97B7F91F953" authorityName="Ponder" authorityYear="1992" box="[264,428,1719,1744]" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="33" pageNumber="34" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B976EABE2D23A22DE41FF97B7F91F953" box="[264,428,1719,1744]" italics="true" pageId="33" pageNumber="34">Pseudotricula</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBD36AC2D23A22DE41FF8CB7FAFF8A3" blockId="33.[264,1322,1799,1905]" box="[264,402,1799,1824]" pageId="33" pageNumber="34">
|
||
<emphasis id="B976EABE2D23A22DE41FF8CB7FAFF8A3" box="[264,402,1799,1824]" italics="true" pageId="33" pageNumber="34">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BBD36AC2D23A22EE42FF8E27A50FDA5" blockId="33.[264,1322,1799,1905]" lastBlockId="34.[264,1323,284,590]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
|
||
<emphasis id="B976EABE2D23A22DE42FF8E27F4BF8CB" bold="true" box="[312,374,1838,1864]" pageId="33" pageNumber="34">Shell</emphasis>
|
||
(
|
||
<figureCitation id="13392A292D23A22DE492F8E37C38F8CA" box="[389,517,1839,1865]" captionStart="FIGURE 6" captionStartId="16.[264,357,1641,1662]" captionTargetBox="[302,1285,329,1616]" captionTargetId="figure-146@16.[302,1285,329,1633]" captionTargetPageId="16" captionText="FIGURE 6. Protoconchs and teleoconch microsculpture of Pseudotricula and Nanocochlea species. A–D, Pseudotricula expandolabra, A, B, D, C.201493, paratype, main streamway, Bauhaus, C, C.201495, Persephone Streamway, Bauhaus; E–G, Pseudotricula conica, E, F, C.201464, Cane Toad Abuse Streamway, Damper Cave, G, Persephone Pot, Bauhaus; Honey and Cream Streamway, Damper Cave; H, Nanocochlea exigua, C.201475; I, J, Nanocochlea stylesae, C.201476, paratypes, Malani Creek; K–M, Nanocochlea damperensis, K, L, C.203685, paratypes, first crossing, Damper Creek, M, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; N, O, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, B, E, F, H, K–N, protoconch; C, D, G, I, J, O, microsculpture. Scales: A, C–E, J, K, M, O, 200 m; B, F–I, L, N, 100 m." figureDoi="http://doi.org/10.5281/zenodo.5050797" httpUri="https://zenodo.org/record/5050797/files/figure.png" pageId="33" pageNumber="34">Fig. 6N, O</figureCitation>
|
||
;
|
||
<figureCitation id="13392A292D23A22DE706F8E37C63F8CA" box="[529,606,1839,1865]" captionStart="FIGURE 14" captionStartId="35.[264,357,1682,1703]" captionTargetBox="[353,1231,287,1666]" captionTargetId="figure-112@35.[336,1250,272,1680]" captionTargetPageId="35" captionText="FIGURE 14. Shells of Nanocochlea species and Pseudotricula progenitor. A C, Nanocochlea exigua, A, holotype, C.439393, Cane Toad Abuse Streamway, Damper Cave; B, C, C.201465, main streamway, Damper Cave; D, Nanocochlea stylesae, holotype, C.439398, Malani Creek; E–G, Nanocochlea damperensis, E, C.203685, paratype, first crossing, Damper Creek; F, holotype, C.439399, first crossing, Damper Creek, F, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; H, I, Pseudotricula progenitor, H, paratype, C.439397, Persephone Streamway, Bauhaus; I, holotype, C.439396, Persephone Streamway, Bauhaus. Scales: 500 m." figureDoi="http://doi.org/10.5281/zenodo.5050821" httpUri="https://zenodo.org/record/5050821/files/figure.png" pageId="33" pageNumber="34">14H, I</figureCitation>
|
||
;
|
||
<figureCitation id="13392A292D23A22DE77DF8E37C87F8CA" box="[618,698,1839,1865]" captionStart="FIGURE 15" captionStartId="36.[264,357,1703,1724]" captionTargetBox="[304,1298,334,1666]" captionTargetId="figure-104@36.[284,1304,324,1686]" captionTargetPageId="36" captionText="FIGURE 15. Shells of Nanocochlea species and Pseudotricula progenitor. A C, Nanocochlea exigua, A, C.203674, outside Damper Cave, Damper Creek B, C.166855, Persephone Pot, Bauhaus, C, C.201465, main streamway, Damper Cave; D–F, Nanocochlea stylesae, D, C.201476, paratype, Malani Creek, E, F, C.203686, first crossing, Damper Creek; GI, Nanocochlea damperensis, G,H, C.303685, seepages at Bauhaus entrance, I, C.203685, paratype, first crossing, Damper Creek; J–L, Pseudotricula progenitor, J, L, C.201463, paratype, Persephone streamway, Bauhaus, K, C.201454, Persephone, Bauhaus. Scales: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5050823" httpUri="https://zenodo.org/record/5050823/files/figure.png" pageId="33" pageNumber="34">15J, K</figureCitation>
|
||
). Length up to
|
||
<quantity id="4CFA9B492D23A22DE67DF8E37DFBF8CA" box="[874,966,1839,1865]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="33" pageNumber="34" unit="mm" value="2.8">2.8 mm</quantity>
|
||
; elongateconic (SW/SL 0.50– 0.64, mean 0.56, n = 22); spire tall, straight to slightly convex in outline; subsutural indentation on last two whorls; periphery of last whorl evenly rounded; suture simple. Protoconch (
|
||
<figureCitation id="13392A292D20A22EE488FE887FC1FEDD" box="[415,508,324,350]" captionStart="FIGURE 6" captionStartId="16.[264,357,1641,1662]" captionTargetBox="[302,1285,329,1616]" captionTargetId="figure-146@16.[302,1285,329,1633]" captionTargetPageId="16" captionText="FIGURE 6. Protoconchs and teleoconch microsculpture of Pseudotricula and Nanocochlea species. A–D, Pseudotricula expandolabra, A, B, D, C.201493, paratype, main streamway, Bauhaus, C, C.201495, Persephone Streamway, Bauhaus; E–G, Pseudotricula conica, E, F, C.201464, Cane Toad Abuse Streamway, Damper Cave, G, Persephone Pot, Bauhaus; Honey and Cream Streamway, Damper Cave; H, Nanocochlea exigua, C.201475; I, J, Nanocochlea stylesae, C.201476, paratypes, Malani Creek; K–M, Nanocochlea damperensis, K, L, C.203685, paratypes, first crossing, Damper Creek, M, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; N, O, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, B, E, F, H, K–N, protoconch; C, D, G, I, J, O, microsculpture. Scales: A, C–E, J, K, M, O, 200 m; B, F–I, L, N, 100 m." figureDoi="http://doi.org/10.5281/zenodo.5050797" httpUri="https://zenodo.org/record/5050797/files/figure.png" pageId="34" pageNumber="35">Fig. 6N</figureCitation>
|
||
) as for genus, microsculpture largely obscured in available material. Teleoconch up to 4.0 whorls in adult; umbilicus absent in juveniles; aperture oval to pearshaped; small, shorter than spire (AL/SL 0.34–0.49, mean 0.40, n = 22); outer lip orthocline to opisthocline, weakly thickened in adult, straight, usually with slight reflection; posterior notch absent; inner lip thin to moderately thickened and of narrow to medium width, in partial contact or narrowly separated from parietal wall.
|
||
</paragraph>
|
||
<paragraph id="8BBD36AC2D20A22EE42FFDF87C5CFDCD" blockId="34.[264,1323,284,590]" box="[312,609,564,590]" pageId="34" pageNumber="35">
|
||
<emphasis id="B976EABE2D20A22EE42FFDF87FFFFDCE" box="[312,450,564,589]" italics="true" pageId="34" pageNumber="35">Dimensions</emphasis>
|
||
. See
|
||
<tableCitation id="C68003172D20A22EE715FDF87C66FDCD" box="[514,603,564,590]" captionStart="TABLE 9" captionStartId="34.[264,352,645,669]" captionText="TABLE 9. Shell dimensions and teleoconch whorl counts of Pseudotricula progenitor n.sp." pageId="34" pageNumber="35">Table 9</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF7D66242D20A22EE41FFD497AE6FD1D" box="[264,1243,644,670]" pageId="34" pageNumber="35" startId="34.[264,352,645,669]">
|
||
<paragraph id="8BBD36AC2D20A22EE41FFD497AE6FD1D" blockId="34.[264,1243,644,670]" box="[264,1243,644,670]" pageId="34" pageNumber="35">
|
||
<emphasis id="B976EABE2D20A22EE41FFD497F47FD1E" bold="true" box="[264,378,645,669]" pageId="34" pageNumber="35">TABLE 9.</emphasis>
|
||
Shell dimensions and teleoconch whorl counts of
|
||
<taxonomicName id="4C024D2F2D20A22EE683FD4A7A9FFD1E" authority="Ponder & Clark & Eberhard & Studdert, 2005" authorityName="Ponder & Clark & Eberhard & Studdert" authorityYear="2005" box="[916,1186,646,669]" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="progenitor" status="sp. nov.">
|
||
<emphasis id="B976EABE2D20A22EE683FD4A7A9FFD1E" box="[916,1186,646,669]" italics="true" pageId="34" pageNumber="35">Pseudotricula progenitor</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24557C52D20A22EE1BEFD4A7AE6FD1D" box="[1193,1243,644,670]" pageId="34" pageNumber="35" rank="species">n.sp.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBD36AC2D20A22EE4ECFD0F7ADBFB80" pageId="34" pageNumber="35">
|
||
<table id="F902C40C2D205DF3E407FD0F7ADAFB80" box="[272,1255,707,1027]" colsContinueFrom="31.[272,1255,1258,1901]" colsContinueIn="42.[272,1266,1542,1852]" gridcols="8" gridrows="8" pageId="34" pageNumber="35">
|
||
<tr id="353234EE2D205DF3E407FD0F7ADAFD5B" box="[272,1255,707,728]" gridrow="0" pageId="34" pageNumber="35" rowspan-0="1">
|
||
<th id="76E35D922D205DF3E4ECFD0F7C18FD5B" box="[507,549,707,728]" gridcol="1" gridrow="0" pageId="34" pageNumber="35">
|
||
<emphasis id="B976EABE2D20A22EE4ECFD0F7C25FD5B" bold="true" box="[507,536,707,728]" pageId="34" pageNumber="35">SL</emphasis>
|
||
</th>
|
||
<th id="76E35D922D205DF3E767FD0F7CA6FD5B" box="[624,667,707,728]" gridcol="2" gridrow="0" pageId="34" pageNumber="35">
|
||
<emphasis id="B976EABE2D20A22EE766FD0F7CABFD5B" bold="true" box="[625,662,707,728]" pageId="34" pageNumber="35">SW</emphasis>
|
||
</th>
|
||
<th id="76E35D922D205DF3E7F1FD0F7D2DFD5B" box="[742,784,707,728]" gridcol="3" gridrow="0" pageId="34" pageNumber="35">
|
||
<emphasis id="B976EABE2D20A22EE7F1FD0F7D35FD5B" bold="true" box="[742,776,707,728]" pageId="34" pageNumber="35">AL</emphasis>
|
||
</th>
|
||
<th id="76E35D922D205DF3E64BFD0F7DBBFD5B" box="[860,902,707,728]" gridcol="4" gridrow="0" pageId="34" pageNumber="35">
|
||
<emphasis id="B976EABE2D20A22EE64BFD0F7DBEFD5B" bold="true" box="[860,899,707,728]" pageId="34" pageNumber="35">AW</emphasis>
|
||
</th>
|
||
<th id="76E35D922D205DF3E6C6FD0F7DC6FD5B" box="[977,1019,707,728]" gridcol="5" gridrow="0" pageId="34" pageNumber="35">
|
||
<emphasis id="B976EABE2D20A22EE6C6FD0F7DC4FD5B" bold="true" box="[977,1017,707,728]" pageId="34" pageNumber="35">BW</emphasis>
|
||
</th>
|
||
<th id="76E35D922D205DF3E150FD0F7A4CFD5B" box="[1095,1137,707,728]" gridcol="6" gridrow="0" pageId="34" pageNumber="35">
|
||
<emphasis id="B976EABE2D20A22EE150FD0F7A57FD5B" bold="true" box="[1095,1130,707,728]" pageId="34" pageNumber="35">CV</emphasis>
|
||
</th>
|
||
<th id="76E35D922D205DF3E1ABFD0F7ADAFD5B" box="[1212,1255,707,728]" gridcol="7" gridrow="0" pageId="34" pageNumber="35">
|
||
<emphasis id="B976EABE2D20A22EE1ABFD0F7AD9FD5B" bold="true" box="[1212,1252,707,728]" pageId="34" pageNumber="35">TW</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="353234EE2D205DF3E407FD227ADAFC80" box="[272,1255,750,771]" gridrow="1" pageId="34" pageNumber="35">
|
||
<th id="76E35D922D205DF3E407FD227F8FFC80" box="[272,434,750,771]" gridcol="0" gridrow="1" pageId="34" pageNumber="35">Holotype</th>
|
||
<td id="76E35D922D205DF3E4ECFD227C18FC80" box="[507,549,750,771]" gridcol="1" gridrow="1" pageId="34" pageNumber="35">2.64</td>
|
||
<td id="76E35D922D205DF3E767FD227CA6FC80" box="[624,667,750,771]" gridcol="2" gridrow="1" pageId="34" pageNumber="35">1.35</td>
|
||
<td id="76E35D922D205DF3E7F1FD227D2DFC80" box="[742,784,750,771]" gridcol="3" gridrow="1" pageId="34" pageNumber="35">0.95</td>
|
||
<td id="76E35D922D205DF3E64BFD227DBBFC80" box="[860,902,750,771]" gridcol="4" gridrow="1" pageId="34" pageNumber="35">0.86</td>
|
||
<td id="76E35D922D205DF3E6C6FD227DC6FC80" box="[977,1019,750,771]" gridcol="5" gridrow="1" pageId="34" pageNumber="35">1.77</td>
|
||
<td id="76E35D922D205DF3E150FD227A4CFC80" box="[1095,1137,750,771]" gridcol="6" gridrow="1" pageId="34" pageNumber="35">0.13</td>
|
||
<td id="76E35D922D205DF3E1ABFD227ADAFC80" box="[1212,1255,750,771]" gridcol="7" gridrow="1" pageId="34" pageNumber="35">3.6</td>
|
||
</tr>
|
||
<tr id="353234EE2D205DF3E407FCD57ADAFCAD" box="[272,1255,793,814]" gridrow="2" pageId="34" pageNumber="35">
|
||
<th id="76E35D922D205DF3E407FCD57F8FFCAD" box="[272,434,793,814]" gridcol="0" gridrow="2" pageId="34" pageNumber="35">Figured paratype</th>
|
||
<td id="76E35D922D205DF3E4ECFCD57C18FCAD" box="[507,549,793,814]" gridcol="1" gridrow="2" pageId="34" pageNumber="35">2.78</td>
|
||
<td id="76E35D922D205DF3E767FCD57CA6FCAD" box="[624,667,793,814]" gridcol="2" gridrow="2" pageId="34" pageNumber="35">1.39</td>
|
||
<td id="76E35D922D205DF3E7F1FCD57D2DFCAD" box="[742,784,793,814]" gridcol="3" gridrow="2" pageId="34" pageNumber="35">0.98</td>
|
||
<td id="76E35D922D205DF3E64BFCD57DBBFCAD" box="[860,902,793,814]" gridcol="4" gridrow="2" pageId="34" pageNumber="35">0.83</td>
|
||
<td id="76E35D922D205DF3E6C6FCD57DC6FCAD" box="[977,1019,793,814]" gridcol="5" gridrow="2" pageId="34" pageNumber="35">1.65</td>
|
||
<td id="76E35D922D205DF3E150FCD57A4CFCAD" box="[1095,1137,793,814]" gridcol="6" gridrow="2" pageId="34" pageNumber="35">0.11</td>
|
||
<td id="76E35D922D205DF3E1ABFCD57ADAFCAD" box="[1212,1255,793,814]" gridcol="7" gridrow="2" pageId="34" pageNumber="35">4.0</td>
|
||
</tr>
|
||
<tr id="353234EE2D205DF3E407FC8F7ADAFCDB" box="[272,1255,835,856]" gridrow="3" pageId="34" pageNumber="35" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
|
||
<th id="76E35D922D205DF3E407FC8F7F8FFCDB" box="[272,434,835,856]" gridcol="0" gridrow="3" pageId="34" pageNumber="35">C.201468 (20)</th>
|
||
</tr>
|
||
<tr id="353234EE2D205DF3E407FCA27ADAFC00" box="[272,1255,878,899]" gridrow="4" pageId="34" pageNumber="35">
|
||
<th id="76E35D922D205DF3E407FCA27F8FFC00" box="[272,434,878,899]" gridcol="0" gridrow="4" pageId="34" pageNumber="35">Minimum</th>
|
||
<td id="76E35D922D205DF3E4ECFCA27C18FC00" box="[507,549,878,899]" gridcol="1" gridrow="4" pageId="34" pageNumber="35">1.91</td>
|
||
<td id="76E35D922D205DF3E767FCA27CA6FC00" box="[624,667,878,899]" gridcol="2" gridrow="4" pageId="34" pageNumber="35">1.22</td>
|
||
<td id="76E35D922D205DF3E7F1FCA27D2DFC00" box="[742,784,878,899]" gridcol="3" gridrow="4" pageId="34" pageNumber="35">0.89</td>
|
||
<td id="76E35D922D205DF3E64BFCA27DBBFC00" box="[860,902,878,899]" gridcol="4" gridrow="4" pageId="34" pageNumber="35">0.80</td>
|
||
<td id="76E35D922D205DF3E6C6FCA27DC6FC00" box="[977,1019,878,899]" gridcol="5" gridrow="4" pageId="34" pageNumber="35">1.39</td>
|
||
<td id="76E35D922D205DF3E150FCA27A4CFC00" box="[1095,1137,878,899]" gridcol="6" gridrow="4" pageId="34" pageNumber="35">0.06</td>
|
||
<td id="76E35D922D205DF3E1ABFCA27ADAFC00" box="[1212,1255,878,899]" gridcol="7" gridrow="4" pageId="34" pageNumber="35">2.75</td>
|
||
</tr>
|
||
<tr id="353234EE2D205DF3E407FC547ADAFC2E" box="[272,1255,920,941]" gridrow="5" pageId="34" pageNumber="35">
|
||
<th id="76E35D922D205DF3E407FC547F8FFC2E" box="[272,434,920,941]" gridcol="0" gridrow="5" pageId="34" pageNumber="35">Maximum</th>
|
||
<td id="76E35D922D205DF3E4ECFC547C18FC2E" box="[507,549,920,941]" gridcol="1" gridrow="5" pageId="34" pageNumber="35">2.82</td>
|
||
<td id="76E35D922D205DF3E767FC547CA6FC2E" box="[624,667,920,941]" gridcol="2" gridrow="5" pageId="34" pageNumber="35">1.56</td>
|
||
<td id="76E35D922D205DF3E7F1FC547D2DFC2E" box="[742,784,920,941]" gridcol="3" gridrow="5" pageId="34" pageNumber="35">1.09</td>
|
||
<td id="76E35D922D205DF3E64BFC547DBBFC2E" box="[860,902,920,941]" gridcol="4" gridrow="5" pageId="34" pageNumber="35">0.95</td>
|
||
<td id="76E35D922D205DF3E6C6FC547DC6FC2E" box="[977,1019,920,941]" gridcol="5" gridrow="5" pageId="34" pageNumber="35">1.77</td>
|
||
<td id="76E35D922D205DF3E150FC547A4CFC2E" box="[1095,1137,920,941]" gridcol="6" gridrow="5" pageId="34" pageNumber="35">0.16</td>
|
||
<td id="76E35D922D205DF3E1ABFC547ADAFC2E" box="[1212,1255,920,941]" gridcol="7" gridrow="5" pageId="34" pageNumber="35">3.75</td>
|
||
</tr>
|
||
<tr id="353234EE2D205DF3E407FC0F7ADAFC5B" box="[272,1255,963,984]" gridrow="6" pageId="34" pageNumber="35">
|
||
<th id="76E35D922D205DF3E407FC0F7F8FFC5B" box="[272,434,963,984]" gridcol="0" gridrow="6" pageId="34" pageNumber="35">Mean</th>
|
||
<td id="76E35D922D205DF3E4ECFC0F7C18FC5B" box="[507,549,963,984]" gridcol="1" gridrow="6" pageId="34" pageNumber="35">2.47</td>
|
||
<td id="76E35D922D205DF3E767FC0F7CA6FC5B" box="[624,667,963,984]" gridcol="2" gridrow="6" pageId="34" pageNumber="35">1.40</td>
|
||
<td id="76E35D922D205DF3E7F1FC0F7D2DFC5B" box="[742,784,963,984]" gridcol="3" gridrow="6" pageId="34" pageNumber="35">0.98</td>
|
||
<td id="76E35D922D205DF3E64BFC0F7DBBFC5B" box="[860,902,963,984]" gridcol="4" gridrow="6" pageId="34" pageNumber="35">0.87</td>
|
||
<td id="76E35D922D205DF3E6C6FC0F7DC6FC5B" box="[977,1019,963,984]" gridcol="5" gridrow="6" pageId="34" pageNumber="35">1.64</td>
|
||
<td id="76E35D922D205DF3E150FC0F7A4CFC5B" box="[1095,1137,963,984]" gridcol="6" gridrow="6" pageId="34" pageNumber="35">0.11</td>
|
||
<td id="76E35D922D205DF3E1ABFC0F7ADAFC5B" box="[1212,1255,963,984]" gridcol="7" gridrow="6" pageId="34" pageNumber="35">3.38</td>
|
||
</tr>
|
||
<tr id="353234EE2D205DF3E407FC227ADAFB80" box="[272,1255,1006,1027]" gridrow="7" pageId="34" pageNumber="35">
|
||
<th id="76E35D922D205DF3E407FC227F8FFB80" box="[272,434,1006,1027]" gridcol="0" gridrow="7" pageId="34" pageNumber="35">Standard Dev.</th>
|
||
<td id="76E35D922D205DF3E4ECFC227C18FB80" box="[507,549,1006,1027]" gridcol="1" gridrow="7" pageId="34" pageNumber="35">0.25</td>
|
||
<td id="76E35D922D205DF3E767FC227CA6FB80" box="[624,667,1006,1027]" gridcol="2" gridrow="7" pageId="34" pageNumber="35">0.08</td>
|
||
<td id="76E35D922D205DF3E7F1FC227D2DFB80" box="[742,784,1006,1027]" gridcol="3" gridrow="7" pageId="34" pageNumber="35">0.05</td>
|
||
<td id="76E35D922D205DF3E64BFC227DBBFB80" box="[860,902,1006,1027]" gridcol="4" gridrow="7" pageId="34" pageNumber="35">0.03</td>
|
||
<td id="76E35D922D205DF3E6C6FC227DC6FB80" box="[977,1019,1006,1027]" gridcol="5" gridrow="7" pageId="34" pageNumber="35">0.10</td>
|
||
<td id="76E35D922D205DF3E150FC227A4CFB80" box="[1095,1137,1006,1027]" gridcol="6" gridrow="7" pageId="34" pageNumber="35">0.02</td>
|
||
<td id="76E35D922D205DF3E1ABFC227ADAFB80" box="[1212,1255,1006,1027]" gridcol="7" gridrow="7" pageId="34" pageNumber="35">0.30</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BBD36AC2D20A22EE42FFBA97A5FFBFC" blockId="34.[264,1323,1125,1872]" box="[312,1122,1125,1151]" pageId="34" pageNumber="35">
|
||
<emphasis id="B976EABE2D20A22EE42FFBA97FFAFBFC" bold="true" box="[312,455,1125,1151]" pageId="34" pageNumber="35">Operculum</emphasis>
|
||
(
|
||
<figureCitation id="13392A292D20A22EE4C1FBA97C60FBFC" box="[470,605,1125,1151]" captionStart="FIGURE 16" captionStartId="37.[264,357,1113,1134]" captionTargetBox="[305,1279,351,1060]" captionTargetId="figure-243@37.[300,1287,333,1089]" captionTargetPageId="37" captionText="FIGURE 16. Opercula of Nanocochlea species and Pseudotricula progenitor. A C, Nanocochlea exigua, C.203677, paratypes, Cane Toad Abuse Streamway, Damper Cave; D F, Nanocochlea damperensis; C–E, C.203685, paratypes, first crossing, Damper Creek; F, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; G, H, Pseudotricula progenitor, C.201821, main streamway, Bauhaus; A, D, G, outer side; B, C, E, F, H, inner side; Scales: A, B, D, 200 m, C, E–H, 100 m." figureDoi="http://doi.org/10.5281/zenodo.5052059" httpUri="https://zenodo.org/record/5052059/files/figure.png" pageId="34" pageNumber="35">Fig. 16G,H</figureCitation>
|
||
). Inner surface with or without white smear.
|
||
</paragraph>
|
||
<paragraph id="8BBD36AC2D20A22EE42FFB417C15FB2B" blockId="34.[264,1323,1125,1872]" box="[312,552,1165,1192]" pageId="34" pageNumber="35">
|
||
<emphasis id="B976EABE2D20A22EE42FFB417F4FFB24" bold="true" box="[312,370,1165,1191]" pageId="34" pageNumber="35">Eyes</emphasis>
|
||
. Unpigmented.
|
||
</paragraph>
|
||
<paragraph id="8BBD36AC2D20A22EE42FFB797F92FACB" blockId="34.[264,1323,1125,1872]" pageId="34" pageNumber="35">
|
||
<emphasis id="B976EABE2D20A22EE42FFB797FE0FB4C" bold="true" box="[312,477,1205,1231]" pageId="34" pageNumber="35">Pallial cavity</emphasis>
|
||
(
|
||
<figureCitation id="13392A292D20A22EE4E7FB7A7C6CFB53" box="[496,593,1206,1232]" captionStart="FIGURE 8" captionStartId="20.[264,357,1612,1633]" captionTargetBox="[341,1270,339,1585]" captionTargetId="figure-146@20.[299,1287,324,1608]" captionTargetPageId="20" captionText="FIGURE 8. Pallial cavities of male specimens of Pseudotricula and Nanocochlea species. A, Pseudotricula expandolabra, C.201493, paratype, Persephone Streamway, Bauhaus; B, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; C, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; D, E, Pseudotricula conica, D, C.203680, main streamway near entrance, Damper Cave, E, Cane Toad Abuse Streamway, Damper Cave; F, Nanocochlea exigua, C.203677, paratype, Cane Toad Abuse Streamway, Damper Cave; G, Nanocochlea damperensis, C.203685, paratype, first crossing, Damper Creek; H, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. Scales: A–D, F–H, 500 m, E, 1 mm. Abbreviations: a, anus; ct, ctenidium; ev, efferent vein; os, osphradium; pc, pericardium; pe, pallial edge; ppw, posterior pallial wall; r, rectum; rg, renal gland. The stippled area represents the hypobranchial gland." figureDoi="http://doi.org/10.5281/zenodo.5050803" httpUri="https://zenodo.org/record/5050803/files/figure.png" pageId="34" pageNumber="35">Fig. 8H</figureCitation>
|
||
). Ctenidium narrow; 9–10 very small filaments; osphradium between posterior end and middle of ctenidium; hypobranchial gland thick to moderately developed; renal organ extends forward ca. ½ into pallial cavity; pericardium more than ½ in pallial roof.
|
||
</paragraph>
|
||
<paragraph id="8BBD36AC2D20A22EE42FFA997DB5F9BB" blockId="34.[264,1323,1125,1872]" pageId="34" pageNumber="35">
|
||
<emphasis id="B976EABE2D20A22EE42FFA997FAEFAEC" bold="true" box="[312,403,1365,1391]" pageId="34" pageNumber="35">Radula</emphasis>
|
||
(
|
||
<figureCitation id="13392A292D20A22EE4B2FA9A7C0FFAF3" box="[421,562,1366,1392]" captionStart="FIGURE 17" captionStartId="38.[264,357,1649,1670]" captionTargetBox="[317,1274,286,1638]" captionTargetId="figure-154@38.[313,1274,286,1643]" captionTargetPageId="38" captionText="FIGURE 17. Radulae of Nanocochlea species Pseudotricula progenitor. A C, Nanocochlea exigua, A, C.201475, Honey and Cream Streamway, Damper Cave, B, D, C.201465, main streamway, Damper Cave; D– E, Nanocochlea stylesae, C.201476, paratypes, Malani Creek; G–K, Nanocochlea damperensis, G, H, C.203685, paratypes, first crossing, Damper Creek, I–K, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; L–O, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, E, central and lateral teeth; B, F, I, M, central, lateral, inner and outer marginal teeth; C, D, J, N central teeth; G, L, lateral teeth; H, central lateral inner and outer marginal teeth; K, outer marginal teeth. Scales: A, G–H, K, L, 10 m; B, I, M, 20 m; C, D, J, N, 5 m." figureDoi="http://doi.org/10.5281/zenodo.5050829" httpUri="https://zenodo.org/record/5050829/files/figure.png" pageId="34" pageNumber="35">Fig. 17L–O</figureCitation>
|
||
).
|
||
<emphasis id="B976EABE2D20A22EE75DFA9A7CDBFAEC" box="[586,742,1366,1391]" italics="true" pageId="34" pageNumber="35">Central teeth</emphasis>
|
||
: dorsal edge with moderate to deep indentation; 5–6 lateral cusps, median cusp narrow, sharply pointed, about twice as long as adjacent cusps.
|
||
<emphasis id="B976EABE2D20A22EE44FFA6A7FD2FA3C" box="[344,495,1446,1471]" italics="true" pageId="34" pageNumber="35">Lateral teeth</emphasis>
|
||
: dorsal edge with shallow indentation; with 5–6 cusps on outer and 4– 5 on inner side; median cusp narrow, sharply pointed, less than twice as long as adjacent cusps; ratio of cutting edge to shaft about ¼; basal projection bluntly pointed.
|
||
<emphasis id="B976EABE2D20A22EE1AAFA3A7F7CF9B4" italics="true" pageId="34" pageNumber="35">Marginal teeth</emphasis>
|
||
: Inner with 25–29 cusps; outer with 22–28 cusps.
|
||
</paragraph>
|
||
<paragraph id="8BBD36AC2D20A22EE42FF9897D39F90B" blockId="34.[264,1323,1125,1872]" pageId="34" pageNumber="35">
|
||
<emphasis id="B976EABE2D20A22EE42FF9897F9EF9DC" bold="true" box="[312,419,1605,1631]" pageId="34" pageNumber="35">Stomach</emphasis>
|
||
(
|
||
<figureCitation id="13392A292D20A22EE4A4F98A7C30F9E3" box="[435,525,1606,1632]" captionStart="FIGURE 9" captionStartId="21.[264,357,1723,1744]" captionTargetBox="[350,1309,380,1631]" captionTargetId="figure-97@21.[299,1342,370,1653]" captionTargetPageId="21" captionText="FIGURE 9. Head, penis and stomach of Pseudotricula species. A, D, Pseudotricula conica, C.203680, main streamway near entrance, Damper Cave; C, Pseudotricula expandolabra, C.201493, paratype, Persephone Streamway, Bauhaus; B, E, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, B, head and penis; C–E, stomach. Scales: A–E, 500 m. Abbreviations: dgo, opening from stomach to digestive gland; ey, eye; int, intestine; oes, oesophagus, p, penis; pg, penial duck; sn, snout; ss, style sack; st, stomach; tn, tentacle." figureDoi="http://doi.org/10.5281/zenodo.5050807" httpUri="https://zenodo.org/record/5050807/files/figure.png" pageId="34" pageNumber="35">Fig. 9E</figureCitation>
|
||
). Stomach with posterior chamber and anterior chamber about equal in size or posterior chamber a little smaller.
|
||
</paragraph>
|
||
<paragraph id="8BBD36AC2D20A22EE42FF9597A76F8D3" blockId="34.[264,1323,1125,1872]" pageId="34" pageNumber="35">
|
||
<emphasis id="B976EABE2D20A22EE42FF9597C05F92C" bold="true" box="[312,568,1685,1711]" pageId="34" pageNumber="35">Male genital system.</emphasis>
|
||
Testis of 0.9–1.5 whorls; prostate gland oval to kidneyshaped; compressed in section. Pallial vas deferens straight. Penis (
|
||
<figureCitation id="13392A292D20A22EE6AAF9727A2AF95B" box="[957,1047,1726,1752]" captionStart="FIGURE 9" captionStartId="21.[264,357,1723,1744]" captionTargetBox="[350,1309,380,1631]" captionTargetId="figure-97@21.[299,1342,370,1653]" captionTargetPageId="21" captionText="FIGURE 9. Head, penis and stomach of Pseudotricula species. A, D, Pseudotricula conica, C.203680, main streamway near entrance, Damper Cave; C, Pseudotricula expandolabra, C.201493, paratype, Persephone Streamway, Bauhaus; B, E, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, B, head and penis; C–E, stomach. Scales: A–E, 500 m. Abbreviations: dgo, opening from stomach to digestive gland; ey, eye; int, intestine; oes, oesophagus, p, penis; pg, penial duck; sn, snout; ss, style sack; st, stomach; tn, tentacle." figureDoi="http://doi.org/10.5281/zenodo.5050807" httpUri="https://zenodo.org/record/5050807/files/figure.png" pageId="34" pageNumber="35">Fig. 9B</figureCitation>
|
||
) with weak swelling in middistal portion; distal end long, papillalike; medial section tapering to parallel sided; of medium length; penial duct in medial section of penis strongly undulating; base of penis moderately wide; with moderate folds; penial duct strongly undulating.
|
||
</paragraph>
|
||
<caption id="DF7D66242D21A22FE41FF95E7C45F8D9" ID-DOI="http://doi.org/10.5281/zenodo.5050821" ID-Zenodo-Dep="5050821" httpUri="https://zenodo.org/record/5050821/files/figure.png" pageId="35" pageNumber="36" startId="35.[264,357,1682,1703]" targetBox="[353,1231,287,1666]" targetPageId="35">
|
||
<paragraph id="8BBD36AC2D21A22FE41FF95E7C45F8D9" blockId="35.[264,1323,1682,1882]" pageId="35" pageNumber="36">
|
||
<emphasis id="B976EABE2D21A22FE41FF95E7FB1F924" bold="true" box="[264,396,1682,1703]" pageId="35" pageNumber="36">FIGURE 14.</emphasis>
|
||
Shells of
|
||
<taxonomicName id="4C024D2F2D21A22FE4E3F95E7CFDF924" box="[500,704,1682,1703]" class="Gastropoda" family="Hydrobiidae" genus="Nanocochlea" kingdom="Animalia" order="Littorinimorpha" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="B976EABE2D21A22FE4E3F95E7C4EF924" box="[500,627,1682,1703]" italics="true" pageId="35" pageNumber="36">Nanocochlea</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C024D2F2D21A22FE7E3F95E7DDAF924" authorityName="Ponder & Clark & Eberhard & Studdert" authorityYear="2005" box="[756,999,1682,1703]" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="progenitor">
|
||
<emphasis id="B976EABE2D21A22FE7E3F95E7DDAF924" box="[756,999,1682,1703]" italics="true" pageId="35" pageNumber="36">Pseudotricula progenitor</emphasis>
|
||
</taxonomicName>
|
||
. A C,
|
||
<taxonomicName id="4C024D2F2D21A22FE129F95E7B38F924" authorityName="Ponder & Clark & Eberhard & Studdert" authorityYear="2005" box="[1086,1285,1682,1703]" class="Gastropoda" family="Hydrobiidae" genus="Nanocochlea" kingdom="Animalia" order="Littorinimorpha" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="exigua">
|
||
<emphasis id="B976EABE2D21A22FE129F95E7B38F924" box="[1086,1285,1682,1703]" italics="true" pageId="35" pageNumber="36">Nanocochlea exigua</emphasis>
|
||
</taxonomicName>
|
||
, A, holotype, C.439393, Cane Toad Abuse Streamway, Damper Cave; B, C, C.201465, main streamway, Damper Cave; D,
|
||
<taxonomicName id="4C024D2F2D21A22FE479F9037C78F967" authorityName="Ponder & Clark & Eberhard & Studdert" authorityYear="2005" box="[366,581,1743,1764]" class="Gastropoda" family="Hydrobiidae" genus="Nanocochlea" kingdom="Animalia" order="Littorinimorpha" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="stylesae">
|
||
<emphasis id="B976EABE2D21A22FE479F9037C78F967" box="[366,581,1743,1764]" italics="true" pageId="35" pageNumber="36">Nanocochlea stylesae</emphasis>
|
||
</taxonomicName>
|
||
, holotype, C.439398, Malani Creek; E–G,
|
||
<taxonomicName id="4C024D2F2D21A22FE115F9037B39F967" authorityName="Ponder & Clark & Eberhard & Studdert" authorityYear="2005" box="[1026,1284,1743,1764]" class="Gastropoda" family="Hydrobiidae" genus="Nanocochlea" kingdom="Animalia" order="Littorinimorpha" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="damperensis">
|
||
<emphasis id="B976EABE2D21A22FE115F9037B39F967" box="[1026,1284,1743,1764]" italics="true" pageId="35" pageNumber="36">Nanocochlea damperensis</emphasis>
|
||
</taxonomicName>
|
||
, E, C.203685, paratype, first crossing, Damper Creek; F, holotype, C.439399, first crossing, Damper Creek, F, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; H, I,
|
||
<taxonomicName id="4C024D2F2D21A22FE1B3F8C67F50F8BE" authorityName="Ponder & Clark & Eberhard & Studdert" authorityYear="2005" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="progenitor">
|
||
<emphasis id="B976EABE2D21A22FE1B3F8C67F50F8BE" italics="true" pageId="35" pageNumber="36">Pseudotricula progenitor</emphasis>
|
||
</taxonomicName>
|
||
, H, paratype, C.439397, Persephone Streamway, Bauhaus; I, holotype, C.439396, Persephone Streamway, Bauhaus. Scales: 500 m.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7D66242D26A228E41FF96B7A08F8D6" ID-DOI="http://doi.org/10.5281/zenodo.5050823" ID-Zenodo-Dep="5050823" httpUri="https://zenodo.org/record/5050823/files/figure.png" pageId="36" pageNumber="37" startId="36.[264,357,1703,1724]" targetBox="[304,1298,334,1666]" targetPageId="36">
|
||
<paragraph id="8BBD36AC2D26A228E41FF96B7A08F8D6" blockId="36.[264,1323,1703,1877]" pageId="36" pageNumber="37">
|
||
<emphasis id="B976EABE2D26A228E41FF96B7FB1F93F" bold="true" box="[264,396,1703,1724]" pageId="36" pageNumber="37">FIGURE 15.</emphasis>
|
||
Shells of
|
||
<taxonomicName id="4C024D2F2D26A228E4E3F9647CFDF93E" box="[500,704,1704,1725]" class="Gastropoda" family="Hydrobiidae" genus="Nanocochlea" kingdom="Animalia" order="Littorinimorpha" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="B976EABE2D26A228E4E3F9647C4EF93E" box="[500,627,1704,1725]" italics="true" pageId="36" pageNumber="37">Nanocochlea</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C024D2F2D26A228E7E3F9647DDBF93E" authorityName="Ponder & Clark & Eberhard & Studdert" authorityYear="2005" box="[756,998,1704,1725]" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="progenitor">
|
||
<emphasis id="B976EABE2D26A228E7E3F9647DDBF93E" box="[756,998,1704,1725]" italics="true" pageId="36" pageNumber="37">Pseudotricula progenitor</emphasis>
|
||
</taxonomicName>
|
||
. A C,
|
||
<taxonomicName id="4C024D2F2D26A228E129F9647B38F93E" authorityName="Ponder & Clark & Eberhard & Studdert" authorityYear="2005" box="[1086,1285,1704,1725]" class="Gastropoda" family="Hydrobiidae" genus="Nanocochlea" kingdom="Animalia" order="Littorinimorpha" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="exigua">
|
||
<emphasis id="B976EABE2D26A228E129F9647B38F93E" box="[1086,1285,1704,1725]" italics="true" pageId="36" pageNumber="37">Nanocochlea exigua</emphasis>
|
||
</taxonomicName>
|
||
, A, C.203674, outside Damper Cave, Damper Creek B, C.166855, Persephone Pot, Bauhaus, C, C.201465, main streamway, Damper Cave; D–F,
|
||
<taxonomicName id="4C024D2F2D26A228E755F9287D2EF97A" authorityName="Ponder & Clark & Eberhard & Studdert" authorityYear="2005" box="[578,787,1764,1785]" class="Gastropoda" family="Hydrobiidae" genus="Nanocochlea" kingdom="Animalia" order="Littorinimorpha" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="stylesae">
|
||
<emphasis id="B976EABE2D26A228E755F9287D2EF97A" box="[578,787,1764,1785]" italics="true" pageId="36" pageNumber="37">Nanocochlea stylesae</emphasis>
|
||
</taxonomicName>
|
||
, D, C.201476, paratype, Malani Creek, E, F, C.203686, first crossing, Damper Creek; GI,
|
||
<taxonomicName id="4C024D2F2D26A228E743F8CF7D6DF89B" authorityName="Ponder & Clark & Eberhard & Studdert" authorityYear="2005" box="[596,848,1795,1816]" class="Gastropoda" family="Hydrobiidae" genus="Nanocochlea" kingdom="Animalia" order="Littorinimorpha" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="damperensis">
|
||
<emphasis id="B976EABE2D26A228E743F8CF7D6DF89B" box="[596,848,1795,1816]" italics="true" pageId="36" pageNumber="37">Nanocochlea damperensis</emphasis>
|
||
</taxonomicName>
|
||
, G,H, C.303685, seepages at Bauhaus entrance, I, C.203685, paratype, first crossing, Damper Creek; J–L,
|
||
<taxonomicName id="4C024D2F2D26A228E623F8EE7A1AF8B4" authorityName="Ponder & Clark & Eberhard & Studdert" authorityYear="2005" box="[820,1063,1826,1847]" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="progenitor">
|
||
<emphasis id="B976EABE2D26A228E623F8EE7A1AF8B4" box="[820,1063,1826,1847]" italics="true" pageId="36" pageNumber="37">Pseudotricula progenitor</emphasis>
|
||
</taxonomicName>
|
||
, J, L, C.201463, paratype, Persephone streamway, Bauhaus, K, C.201454, Persephone, Bauhaus. Scales: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7D66242D27A229E41FFB957A3FFB70" ID-DOI="http://doi.org/10.5281/zenodo.5052059" ID-Zenodo-Dep="5052059" httpUri="https://zenodo.org/record/5052059/files/figure.png" pageId="37" pageNumber="38" startId="37.[264,357,1113,1134]" targetBox="[305,1279,351,1060]" targetPageId="37">
|
||
<paragraph id="8BBD36AC2D27A229E41FFB957A3FFB70" blockId="37.[264,1322,1113,1267]" pageId="37" pageNumber="38">
|
||
<emphasis id="B976EABE2D27A229E41FFB957FB1FBED" bold="true" box="[264,396,1113,1134]" pageId="37" pageNumber="38">FIGURE 16.</emphasis>
|
||
Opercula of
|
||
<taxonomicName id="4C024D2F2D27A229E705FB957CE3FBED" box="[530,734,1113,1134]" class="Gastropoda" family="Hydrobiidae" genus="Nanocochlea" kingdom="Animalia" order="Littorinimorpha" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="B976EABE2D27A229E705FB957CADFBED" box="[530,656,1113,1134]" italics="true" pageId="37" pageNumber="38">Nanocochlea</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C024D2F2D27A229E605FB957A38FBED" authorityName="Ponder & Clark & Eberhard & Studdert" authorityYear="2005" box="[786,1029,1113,1134]" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="progenitor">
|
||
<emphasis id="B976EABE2D27A229E605FB957A38FBED" box="[786,1029,1113,1134]" italics="true" pageId="37" pageNumber="38">Pseudotricula progenitor</emphasis>
|
||
</taxonomicName>
|
||
. A C,
|
||
<taxonomicName id="4C024D2F2D27A229E14AFB957B19FBED" authorityName="Ponder & Clark & Eberhard & Studdert" authorityYear="2005" box="[1117,1316,1113,1134]" class="Gastropoda" family="Hydrobiidae" genus="Nanocochlea" kingdom="Animalia" order="Littorinimorpha" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="exigua">
|
||
<emphasis id="B976EABE2D27A229E14AFB957B19FBED" box="[1117,1316,1113,1134]" italics="true" pageId="37" pageNumber="38">Nanocochlea exigua</emphasis>
|
||
</taxonomicName>
|
||
, C.203677, paratypes, Cane Toad Abuse Streamway, Damper Cave; D F,
|
||
<taxonomicName id="4C024D2F2D27A229E6FDFBB77AD4FB13" authorityName="Ponder & Clark & Eberhard & Studdert" authorityYear="2005" box="[1002,1257,1147,1168]" class="Gastropoda" family="Hydrobiidae" genus="Nanocochlea" kingdom="Animalia" order="Littorinimorpha" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="damperensis">
|
||
<emphasis id="B976EABE2D27A229E6FDFBB77AD4FB13" box="[1002,1257,1147,1168]" italics="true" pageId="37" pageNumber="38">Nanocochlea damperensis</emphasis>
|
||
</taxonomicName>
|
||
; C–E, C.203685, paratypes, first crossing, Damper Creek; F, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; G, H,
|
||
<taxonomicName id="4C024D2F2D27A229E765FB707D5FFB52" authorityName="Ponder & Clark & Eberhard & Studdert" authorityYear="2005" box="[626,866,1212,1233]" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="progenitor">
|
||
<emphasis id="B976EABE2D27A229E765FB707D5FFB52" box="[626,866,1212,1233]" italics="true" pageId="37" pageNumber="38">Pseudotricula progenitor</emphasis>
|
||
</taxonomicName>
|
||
, C.201821, main streamway, Bauhaus; A, D, G, outer side; B, C, E, F, H, inner side; Scales: A, B, D, 200 m, C, E–H, 100 m.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBD36AC2D27A229E42FFA827F8DF953" blockId="37.[264,1323,1358,1744]" pageId="37" pageNumber="38">
|
||
<emphasis id="B976EABE2D27A229E42FFA827C6CFAEB" bold="true" box="[312,593,1358,1384]" pageId="37" pageNumber="38">Female genital system</emphasis>
|
||
(
|
||
<figureCitation id="13392A292D27A229E772FA827CC6FAEB" box="[613,763,1358,1384]" captionStart="FIGURE 18" captionStartId="39.[264,357,1046,1067]" captionTargetBox="[308,1274,317,1017]" captionTargetId="figure-219@39.[254,1333,301,1043]" captionTargetPageId="39" captionText="FIGURE 18. Female reproductive system of Nanocochlea species and Pseudotricula progenitor. A, Nanocochlea exigua, C.203677, paratype, Cane Toad Abuse Streamway, Damper Cave; B, Nanocochlea damperensis, C.203685, paratype, first crossing, Damper Creek; C, D, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. Scales: 500 m. Abbreviations: ag, albumen gland; bc, bursa copulatrix; bd, bursal duct; cg, capsule gland; co, coiled oviduct; po, genital opening; pw, pallial wall; sr, seminal receptacle; vc, ventral channel." figureDoi="http://doi.org/10.5281/zenodo.5050831" httpUri="https://zenodo.org/record/5050831/files/figure.png" pageId="37" pageNumber="38">Fig. 18C, D</figureCitation>
|
||
). Ovary of 0.5–0.7 whorls; oviduct extends to posterior edge of bursa copulatrix; joins bursal duct in front of posterior pallial wall at junction of albumen and capsule glands to half way between posterior pallial wall and capsule gland. Bursa copulatrix large, extending to posterior pallial wall; globular to pyriform; with bursal duct arising from middle of anterior edge of bursa, straight or with undulations. Seminal receptacle at middle of inner wall of bursa copulatrix; pyriform. More than half of albumen gland in front of posterior pallial wall; capsule gland about same length as albumen gland; compressed oval in section; anterior end blunt; ventral channel with distinct muscular vestibule; genital opening overlapping anterior end of capsule gland.
|
||
</paragraph>
|
||
<paragraph id="8BBD36AC2D27A22BE41FF8CB7F53FAB3" blockId="37.[264,1228,1799,1864]" lastBlockId="39.[264,1323,1303,1649]" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38">
|
||
<emphasis id="B976EABE2D27A229E41FF8CB7C18F8A3" box="[264,549,1799,1824]" italics="true" pageId="37" pageNumber="38">Distribution and habitat</emphasis>
|
||
Bauhaus and Damper Cave in low energy small streams with mixed substrate.
|
||
<emphasis id="B976EABE2D25A22BE41FFADB7F53FAB3" box="[264,366,1303,1328]" italics="true" pageId="39" pageNumber="40">Remarks</emphasis>
|
||
</paragraph>
|
||
<caption id="DF7D66242D24A22AE41FF9BD7D59F8E2" ID-DOI="http://doi.org/10.5281/zenodo.5050829" ID-Zenodo-Dep="5050829" httpUri="https://zenodo.org/record/5050829/files/figure.png" pageId="38" pageNumber="39" startId="38.[264,357,1649,1670]" targetBox="[317,1274,286,1638]" targetPageId="38">
|
||
<paragraph id="8BBD36AC2D24A22AE41FF9BD7D59F8E2" blockId="38.[264,1323,1649,1889]" pageId="38" pageNumber="39">
|
||
<emphasis id="B976EABE2D24A22AE41FF9BD7FB3F905" bold="true" box="[264,398,1649,1670]" pageId="38" pageNumber="39">FIGURE 17.</emphasis>
|
||
Radulae of
|
||
<taxonomicName id="4C024D2F2D24A22AE718F9BE7CE1F904" box="[527,732,1650,1671]" class="Gastropoda" family="Hydrobiidae" genus="Nanocochlea" kingdom="Animalia" order="Littorinimorpha" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="B976EABE2D24A22AE718F9BE7CB3F904" box="[527,654,1650,1671]" italics="true" pageId="38" pageNumber="39">Nanocochlea</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
<taxonomicName id="4C024D2F2D24A22AE7F0F9BE7DE1F904" authorityName="Ponder & Clark & Eberhard & Studdert" authorityYear="2005" box="[743,988,1650,1671]" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="progenitor">
|
||
<emphasis id="B976EABE2D24A22AE7F0F9BE7DE1F904" box="[743,988,1650,1671]" italics="true" pageId="38" pageNumber="39">Pseudotricula progenitor</emphasis>
|
||
</taxonomicName>
|
||
. A C,
|
||
<taxonomicName id="4C024D2F2D24A22AE12DF9BE7B3EF904" authorityName="Ponder & Clark & Eberhard & Studdert" authorityYear="2005" box="[1082,1283,1650,1671]" class="Gastropoda" family="Hydrobiidae" genus="Nanocochlea" kingdom="Animalia" order="Littorinimorpha" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="exigua">
|
||
<emphasis id="B976EABE2D24A22AE12DF9BE7B3EF904" box="[1082,1283,1650,1671]" italics="true" pageId="38" pageNumber="39">Nanocochlea exigua</emphasis>
|
||
</taxonomicName>
|
||
, A, C.201475, Honey and Cream Streamway, Damper Cave, B, D, C.201465, main streamway, Damper Cave; D– E,
|
||
<taxonomicName id="4C024D2F2D24A22AE43DF97D7C3FF945" authorityName="Ponder & Clark & Eberhard & Studdert" authorityYear="2005" box="[298,514,1713,1734]" class="Gastropoda" family="Hydrobiidae" genus="Nanocochlea" kingdom="Animalia" order="Littorinimorpha" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="stylesae">
|
||
<emphasis id="B976EABE2D24A22AE43DF97D7C3FF945" box="[298,514,1713,1734]" italics="true" pageId="38" pageNumber="39">Nanocochlea stylesae</emphasis>
|
||
</taxonomicName>
|
||
, C.201476, paratypes, Malani Creek; G–K,
|
||
<taxonomicName id="4C024D2F2D24A22AE6CEF97D7AE0F945" authorityName="Ponder & Clark & Eberhard & Studdert" authorityYear="2005" box="[985,1245,1713,1734]" class="Gastropoda" family="Hydrobiidae" genus="Nanocochlea" kingdom="Animalia" order="Littorinimorpha" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="damperensis">
|
||
<emphasis id="B976EABE2D24A22AE6CEF97D7AE0F945" box="[985,1245,1713,1734]" italics="true" pageId="38" pageNumber="39">Nanocochlea damperensis</emphasis>
|
||
</taxonomicName>
|
||
, G, H, C.203685, paratypes, first crossing, Damper Creek, I–K, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; L–O,
|
||
<taxonomicName id="4C024D2F2D24A22AE61CF93D7A3FF885" authorityName="Ponder & Clark & Eberhard & Studdert" authorityYear="2005" box="[779,1026,1777,1798]" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="progenitor">
|
||
<emphasis id="B976EABE2D24A22AE61CF93D7A3FF885" box="[779,1026,1777,1798]" italics="true" pageId="38" pageNumber="39">Pseudotricula progenitor</emphasis>
|
||
</taxonomicName>
|
||
, C.201821, main streamway, Bauhaus. A, E, central and lateral teeth; B, F, I, M, central, lateral, inner and outer marginal teeth; C, D, J, N central teeth; G, L, lateral teeth; H, central lateral inner and outer marginal teeth; K, outer marginal teeth. Scales: A, G–H, K, L, 10 m; B, I, M, 20 m; C, D, J, N, 5 m.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7D66242D25A22BE41FFBDA7F9FFB3D" ID-DOI="http://doi.org/10.5281/zenodo.5050831" ID-Zenodo-Dep="5050831" httpUri="https://zenodo.org/record/5050831/files/figure.png" pageId="39" pageNumber="40" startId="39.[264,357,1046,1067]" targetBox="[308,1274,317,1017]" targetPageId="39">
|
||
<paragraph id="8BBD36AC2D25A22BE41FFBDA7F9FFB3D" blockId="39.[264,1323,1046,1214]" pageId="39" pageNumber="40">
|
||
<emphasis id="B976EABE2D25A22BE41FFBDA7FAFFBA8" bold="true" box="[264,402,1046,1067]" pageId="39" pageNumber="40">FIGURE 18.</emphasis>
|
||
Female reproductive system of
|
||
<taxonomicName id="4C024D2F2D25A22BE7E5FBDB7DF8FBAF" box="[754,965,1047,1068]" class="Gastropoda" family="Hydrobiidae" genus="Nanocochlea" kingdom="Animalia" order="Littorinimorpha" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="B976EABE2D25A22BE7E5FBDB7D4DFBAF" box="[754,880,1047,1068]" italics="true" pageId="39" pageNumber="40">Nanocochlea</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C024D2F2D25A22BE112FBDB7AC3FBAF" authorityName="Ponder & Clark & Eberhard & Studdert" authorityYear="2005" box="[1029,1278,1047,1068]" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="progenitor">
|
||
<emphasis id="B976EABE2D25A22BE112FBDB7AC3FBAF" box="[1029,1278,1047,1068]" italics="true" pageId="39" pageNumber="40">Pseudotricula progenitor</emphasis>
|
||
</taxonomicName>
|
||
. A,
|
||
<taxonomicName id="4C024D2F2D25A22BE41FFBF87FECFBCA" authorityName="Ponder & Clark & Eberhard & Studdert" authorityYear="2005" box="[264,465,1076,1097]" class="Gastropoda" family="Hydrobiidae" genus="Nanocochlea" kingdom="Animalia" order="Littorinimorpha" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="exigua">
|
||
<emphasis id="B976EABE2D25A22BE41FFBF87FECFBCA" box="[264,465,1076,1097]" italics="true" pageId="39" pageNumber="40">Nanocochlea exigua</emphasis>
|
||
</taxonomicName>
|
||
, C.203677, paratype, Cane Toad Abuse Streamway, Damper Cave; B,
|
||
<taxonomicName id="4C024D2F2D25A22BE1BBFBF87FBDFBE5" authorityName="Ponder & Clark & Eberhard & Studdert" authorityYear="2005" class="Gastropoda" family="Hydrobiidae" genus="Nanocochlea" kingdom="Animalia" order="Littorinimorpha" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="damperensis">
|
||
<emphasis id="B976EABE2D25A22BE1BBFBF87FBDFBE5" italics="true" pageId="39" pageNumber="40">Nanocochlea damperensis</emphasis>
|
||
</taxonomicName>
|
||
, C.203685, paratype, first crossing, Damper Creek; C, D,
|
||
<taxonomicName id="4C024D2F2D25A22BE6D2FB9D7A85FBE5" authorityName="Ponder & Clark & Eberhard & Studdert" authorityYear="2005" box="[965,1208,1105,1126]" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="progenitor">
|
||
<emphasis id="B976EABE2D25A22BE6D2FB9D7A85FBE5" box="[965,1208,1105,1126]" italics="true" pageId="39" pageNumber="40">Pseudotricula progenitor</emphasis>
|
||
</taxonomicName>
|
||
, C.201821, main streamway, Bauhaus. Scales: 500 m. Abbreviations: ag, albumen gland; bc, bursa copulatrix; bd, bursal duct; cg, capsule gland; co, coiled oviduct; po, genital opening; pw, pallial wall; sr, seminal receptacle; vc, ventral channel.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBD36AC2D25A22BE42FFAF37AB0F9F2" blockId="39.[264,1323,1303,1649]" pageId="39" pageNumber="40">
|
||
<taxonomicName id="4C024D2F2D25A22BE42FFAF37C5EFADB" authorityName="Ponder & Clark & Eberhard & Studdert" authorityYear="2005" box="[312,611,1343,1368]" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="progenitor">
|
||
<emphasis id="B976EABE2D25A22BE42FFAF37C5EFADB" box="[312,611,1343,1368]" italics="true" pageId="39" pageNumber="40">Pseudotricula progenitor</emphasis>
|
||
</taxonomicName>
|
||
shares characters of both
|
||
<taxonomicName id="4C024D2F2D25A22BE6B3FAF37A02FADB" authorityName="Ponder and Clark" authorityYear="1993" box="[932,1087,1343,1368]" class="Gastropoda" family="Hydrobiidae" genus="Nanocochlea" kingdom="Animalia" order="Littorinimorpha" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B976EABE2D25A22BE6B3FAF37A02FADB" box="[932,1087,1343,1368]" italics="true" pageId="39" pageNumber="40">Nanocochlea</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C024D2F2D25A22BE168FAF37B1EFADB" authorityName="Ponder" authorityYear="1992" box="[1151,1315,1343,1368]" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B976EABE2D25A22BE168FAF37B1EFADB" box="[1151,1315,1343,1368]" italics="true" pageId="39" pageNumber="40">Pseudotricula</emphasis>
|
||
</taxonomicName>
|
||
. The smallsized, tallspired shell is typical of
|
||
<taxonomicName id="4C024D2F2D25A22BE62FFAAB7DEEFA03" authorityName="Ponder and Clark" authorityYear="1993" box="[824,979,1383,1408]" class="Gastropoda" family="Hydrobiidae" genus="Nanocochlea" kingdom="Animalia" order="Littorinimorpha" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B976EABE2D25A22BE62FFAAB7DEEFA03" box="[824,979,1383,1408]" italics="true" pageId="39" pageNumber="40">Nanocochlea</emphasis>
|
||
</taxonomicName>
|
||
but the (narrowly) reflected outer lip, straight spire outline and subsutural indentation are characters found in typical
|
||
<taxonomicName id="4C024D2F2D25A22BE41FFA7B7F91FA53" authorityName="Ponder" authorityYear="1992" box="[264,428,1463,1488]" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B976EABE2D25A22BE41FFA7B7F91FA53" box="[264,428,1463,1488]" italics="true" pageId="39" pageNumber="40">Pseudotricula</emphasis>
|
||
</taxonomicName>
|
||
taxa. The protoconch microsculpture, while not able to be examined in detail, appears to be of the plesiomorphic
|
||
<typeStatus id="54B9880E2D25A22BE7E2FA137D1AFA7A" box="[757,807,1503,1529]" pageId="39" pageNumber="40">type</typeStatus>
|
||
(shared by
|
||
<taxonomicName id="4C024D2F2D25A22BE6A5FA137A70FA7B" authorityName="Ponder and Clark" authorityYear="1993" box="[946,1101,1503,1528]" class="Gastropoda" family="Hydrobiidae" genus="Nanocochlea" kingdom="Animalia" order="Littorinimorpha" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B976EABE2D25A22BE6A5FA137A70FA7B" box="[946,1101,1503,1528]" italics="true" pageId="39" pageNumber="40">Nanocochlea</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C024D2F2D25A22BE191FA137B17FA7B" authorityName="Ponder" authorityYear="1992" box="[1158,1322,1503,1528]" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus" sensu="stricto">
|
||
<emphasis id="B976EABE2D25A22BE191FA137B17FA7B" box="[1158,1322,1503,1528]" italics="true" pageId="39" pageNumber="40">Pseudotricula</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24557C52D25A22BE41FF9CB7F10F9A2" box="[264,301,1543,1569]" pageId="39" pageNumber="40" sensu="stricto">s.s.</taxonomicNameLabel>
|
||
), as are the radular characters, which are like those of
|
||
<taxonomicName id="4C024D2F2D25A22BE13CF9CB7AFBF9A3" authorityName="Ponder and Clark" authorityYear="1993" box="[1067,1222,1543,1568]" class="Gastropoda" family="Hydrobiidae" genus="Nanocochlea" kingdom="Animalia" order="Littorinimorpha" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B976EABE2D25A22BE13CF9CB7AFBF9A3" box="[1067,1222,1543,1568]" italics="true" pageId="39" pageNumber="40">Nanocochlea</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C024D2F2D25A22BE002F9CB7F9EF9CB" authorityName="Ponder & Clark & Eberhard & Studdert" authorityYear="2005" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="arthurclarkei">
|
||
<emphasis id="B976EABE2D25A22BE002F9CB7F9EF9CB" italics="true" pageId="39" pageNumber="40">P. arthurclarkei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C024D2F2D25A22BE4F2F9E37C69F9CB" authorityName="Ponder & Clark & Eberhard & Studdert" authorityYear="2005" box="[485,596,1583,1608]" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="conica">
|
||
<emphasis id="B976EABE2D25A22BE4F2F9E37C69F9CB" box="[485,596,1583,1608]" italics="true" pageId="39" pageNumber="40">P. conica</emphasis>
|
||
</taxonomicName>
|
||
. It is probable that this species most closely resembles the ancestral
|
||
<taxonomicName id="4C024D2F2D25A22BE46FF99B7C21F9F3" authorityName="Ponder" authorityYear="1992" box="[376,540,1623,1648]" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B976EABE2D25A22BE46FF99B7C21F9F3" box="[376,540,1623,1648]" italics="true" pageId="39" pageNumber="40">Pseudotricula</emphasis>
|
||
</taxonomicName>
|
||
that gave rise to the Precipitous Bluff cave radiation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |