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<document ID-DOI="http://doi.org/10.5281/zenodo.6449337" ID-ISSN="2073-2333" ID-Zenodo-Dep="6449337" LSID="urn:lsid:zoobank.org:act:52CDB184-2A52-4A84-99B9-7620B52FE2A4" approvalRequired="5" approvalRequired_for_originalDoi="1" approvalRequired_for_taxonomicNames="3" approvalRequired_for_textStreams="1" checkinTime="1649700634309" checkinUser="felipe" docAuthor="Guglya, Yu. A." docDate="2016" docId="03A187A0FFEE483A2987F997FDBAB8D9" docLanguage="en" docName="VestnikZoologii.50.5.407-414.pdf.imf" docOrigin="Vestnik Zoologii (Vestn. Zool.) 50 (5)" docSource="http://dx.doi.org/10.1515/vzoo-2016-0047" docStyle="DocumentStyle:9FE6554C034D0A4D8EA9CEAE255F4368.2:VestnikZoologii.2014-2019.journal_article" docStyleId="9FE6554C034D0A4D8EA9CEAE255F4368" docStyleName="VestnikZoologii.2014-2019.journal_article" docStyleVersion="2" docTitle="Melanagromyza pratensis Guglya Results &amp; Melanagromyza &amp; Guglya &amp; e &amp; Head &amp; Orbit 2016, sp. n." docType="treatment" docUuid="52CDB184-2A52-4A84-99B9-7620B52FE2A4" docUuidSource="ZooBank" docVersion="3" lastPageId="6" lastPageNumber="413" masterDocId="FF98FFD8FFEA483C2921FF83FFBBBA73" masterDocTitle="Four New Species Of The Genus Melanagromyza (Diptera, Agromyzidae) From Ukraine" masterLastPageNumber="414" masterPageNumber="407" pageId="4" pageNumber="411" updateTime="1649701288189" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>Four New Species Of The Genus Melanagromyza (Diptera, Agromyzidae) From Ukraine</mods:title>
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<mods:namePart>Guglya, Yu. A.</mods:namePart>
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<mods:title>Vestnik Zoologii</mods:title>
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<mods:date>2016</mods:date>
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<mods:title>Vestn. Zool.</mods:title>
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<mods:number>2016-10-01</mods:number>
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<mods:number>50</mods:number>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6449337" ID-Zenodo-Dep="6449337" LSID="urn:lsid:zoobank.org:act:52CDB184-2A52-4A84-99B9-7620B52FE2A4" httpUri="http://treatment.plazi.org/id/03A187A0FFEE483A2987F997FDBAB8D9" lastPageId="6" lastPageNumber="413" pageId="4" pageNumber="411">
<subSubSection box="[166,787,1556,1584]" pageId="4" pageNumber="411" type="nomenclature">
<paragraph blockId="4.[166,787,1556,1584]" box="[166,787,1556,1584]" pageId="4" pageNumber="411">
<heading box="[166,787,1556,1584]" fontSize="36" level="2" pageId="4" pageNumber="411" reason="3">
<taxonomicName authority="Guglya" authorityName="Guglya Results &amp; Melanagromyza &amp; Guglya &amp; e &amp; Head &amp; Orbit" authorityYear="2016" box="[166,571,1556,1584]" class="Insecta" family="Agromyzidae" genus="Melanagromyza" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="411" phylum="Arthropoda" rank="species" species="pratensis" status="sp. n.">
<emphasis bold="true" box="[166,478,1556,1583]" italics="true" pageId="4" pageNumber="411">Melanagromyza pratensis</emphasis>
Guglya
</taxonomicName>
,
<taxonomicNameLabel box="[581,642,1557,1584]" pageId="4" pageNumber="411" rank="species">
<emphasis bold="true" box="[581,635,1557,1583]" pageId="4" pageNumber="411">sp. n</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation box="[656,779,1557,1584]" captionStart="Figs 3038" captionStartId="5.[190,229,1758,1780]" captionTargetBox="[299,1156,895,1722]" captionTargetId="figure-306@5.[267,1169,874,1740]" captionTargetPageId="5" captionText="Figs 3038. M. pratensis sp. n., holotype: 30, 31 — male head; 32 — phallus, ventral view; 33 — same, lateral view; 34 — male wing; 35 — epandrium-hypandrium complex; 36 — ejaculatory apodeme; paratype (female terminalia): 37 — spermathecae, VII tergit, VII sternit; 38 — ventral seminal receptacle, egg guide, left blade, proctiger and cerci. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6449448" httpUri="https://zenodo.org/record/6449448/files/figure.png" pageId="4" pageNumber="411">figs 3038</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="413" pageId="4" pageNumber="411" type="materials_examined">
<paragraph blockId="4.[166,1225,1612,1769]" pageId="4" pageNumber="411">
<emphasis bold="true" box="[219,363,1614,1636]" pageId="4" pageNumber="411">Type material</emphasis>
.
<materialsCitation collectingDate="2013-06-13" country="Ukraine" latitude="51.033333" location="Vakalivshchyna" longLatPrecision="1093" longitude="34.916668" pageId="4" pageNumber="411" specimenCount="1" stateProvince="Sumy" typeStatus="holotype">
<typeStatus box="[376,466,1614,1636]" pageId="4" pageNumber="411">Holotype</typeStatus>
Ơ,
<collectingCountry box="[504,585,1614,1636]" name="Ukraine" pageId="4" pageNumber="411">Ukraine</collectingCountry>
:
<collectingRegion box="[597,731,1614,1636]" country="Ukraine" name="Sumy" pageId="4" pageNumber="411">Sumy Region</collectingRegion>
:
<location LSID="urn:lsid:plazi:treatment:03A187A0FFEE483A2987F997FDBAB8D9:8ED7606DFFEE48382BC7F9CDFCC4BC17" box="[742,895,1614,1636]" country="Ukraine" latitude="51.033333" longLatPrecision="1093" longitude="34.916668" name="Vakalivshchyna" pageId="4" pageNumber="411" stateProvince="Sumy">Vakalivshchyna</location>
(
<geoCoordinate box="[910,999,1614,1636]" degrees="51" direction="north" minutes="02" orientation="latitude" pageId="4" pageNumber="411" precision="925" value="51.033333">51°02ʹ N</geoCoordinate>
,
<geoCoordinate box="[1012,1096,1614,1636]" degrees="34" direction="east" minutes="55" orientation="longitude" pageId="4" pageNumber="411" precision="925" value="34.916668">34°55ʹ E</geoCoordinate>
),
<date box="[1116,1221,1614,1636]" pageId="4" pageNumber="411" value="2013-06-13">
<collectingDate box="[1116,1221,1614,1636]" pageId="4" pageNumber="411" value="2013-06-13">13.06.2013</collectingDate>
</date>
, 10.0011.00, meadow,
<collectedFrom box="[384,567,1641,1663]" pageId="4" pageNumber="411">
on
<taxonomicName box="[414,567,1641,1662]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="411" phylum="Tracheophyta" rank="species" species="jacea">
<emphasis box="[414,567,1641,1662]" italics="true" pageId="4" pageNumber="411">Centaurea jacea</emphasis>
</taxonomicName>
</collectedFrom>
L. (No Н-89) (Guglya) (dissected)
</materialsCitation>
.
<materialsCitation country="Ukraine" location="Sumy Region" pageId="4" pageNumber="411" specimenCount="4" specimenCount-female="4" stateProvince="Sumy" typeStatus="paratype">
<typeStatus box="[905,1001,1641,1663]" pageId="4" pageNumber="411">Paratypes</typeStatus>
:
<collectingRegion box="[1008,1139,1641,1663]" country="Ukraine" name="Sumy" pageId="4" pageNumber="411">Sumy Region</collectingRegion>
: 1 Ơ,
<specimenCount box="[1187,1219,1639,1663]" pageId="4" pageNumber="411" type="female">4 ♀</specimenCount>
, labels as in
<typeStatus box="[276,360,1667,1689]" pageId="4" pageNumber="411">holotype</typeStatus>
(No Н-90, H-94) (Guglya) (dissected)
</materialsCitation>
;
<materialsCitation country="Ukraine" location="Sumy Region" pageId="4" pageNumber="411" specimenCount="1" specimenCount-female="1" stateProvince="Sumy Region" typeStatus="paratype">
<specimenCount box="[742,774,1666,1689]" pageId="4" pageNumber="411" type="female">1 ♀</specimenCount>
, same locality and date, 11.00, motley grass on a slope (No Н-95) (Guglya) (dissected)
</materialsCitation>
;
<materialsCitation collectingDate="2011-11-06" country="Ukraine" latitude="50.166668" location="Rubizhne" longLatPrecision="1099" longitude="36.783333" pageId="4" pageNumber="411" specimenCount="1" stateProvince="Kharkiv" typeStatus="paratype">
<collectingRegion box="[556,711,1694,1716]" country="Ukraine" name="Kharkiv" pageId="4" pageNumber="411">Kharkiv Region</collectingRegion>
: 1 Ơ, near
<location LSID="urn:lsid:plazi:treatment:03A187A0FFEE483A2987F997FDBAB8D9:8ED7606DFFEE48382A0CF91DFC32BCC7" box="[813,905,1694,1716]" country="Ukraine" latitude="50.166668" longLatPrecision="1099" longitude="36.783333" name="Rubizhne" pageId="4" pageNumber="411" stateProvince="Kharkiv">Rubizhne</location>
(
<geoCoordinate box="[919,1006,1694,1716]" degrees="50" direction="north" minutes="10" orientation="latitude" pageId="4" pageNumber="411" precision="925" value="50.166668">50°10ʹ N</geoCoordinate>
,
<geoCoordinate box="[1016,1098,1694,1716]" degrees="36" direction="east" minutes="47" orientation="longitude" pageId="4" pageNumber="411" precision="925" value="36.783333">36°47ʹ E</geoCoordinate>
),
<date box="[1116,1221,1694,1716]" pageId="4" pageNumber="411" value="2011-11-06">
<collectingDate box="[1116,1221,1694,1716]" pageId="4" pageNumber="411" value="2011-11-06">11.06.2011</collectingDate>
</date>
, 16.00, dry gully (No Н-96) (Guglya) (dissected); 1 Ơ
</materialsCitation>
,
<materialsCitation collectingDate="2011-11-06" country="Ukraine" latitude="50.166668" location="Rubizhne" longLatPrecision="1099" longitude="36.783333" pageId="4" pageNumber="411" specimenCount="6" specimenCount-female="6" stateProvince="Kharkiv Region" typeStatus="paratype">
<specimenCount box="[669,701,1719,1743]" pageId="4" pageNumber="411" type="female">6 ♀</specimenCount>
, same locality and date, 13.00, edge of deciduous forest (No Н-97, H-103) (Guglya) (dissected)
</materialsCitation>
.
</paragraph>
<paragraph blockId="4.[166,1225,1802,1861]" lastBlockId="5.[174,1233,175,874]" lastPageId="5" lastPageNumber="412" pageId="4" pageNumber="411">
D e s c r i p t i o n. Head (
<figureCitation box="[498,618,1802,1829]" captionStart="Figs 3038" captionStartId="5.[190,229,1758,1780]" captionTargetBox="[299,1156,895,1722]" captionTargetId="figure-306@5.[267,1169,874,1740]" captionTargetPageId="5" captionText="Figs 3038. M. pratensis sp. n., holotype: 30, 31 — male head; 32 — phallus, ventral view; 33 — same, lateral view; 34 — male wing; 35 — epandrium-hypandrium complex; 36 — ejaculatory apodeme; paratype (female terminalia): 37 — spermathecae, VII tergit, VII sternit; 38 — ventral seminal receptacle, egg guide, left blade, proctiger and cerci. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6449448" httpUri="https://zenodo.org/record/6449448/files/figure.png" pageId="4" pageNumber="411">
fig. 30
<emphasis bold="true" box="[572,587,1802,1828]" pageId="4" pageNumber="411"></emphasis>
31
</figureCitation>
). Orbit projected above eye in profile only at level of orb s; 3 orb s, 3 fr s; frorb sta elongated in several rows, reclinate nearest to eye and proclinate nearest to frons; orbit wide and shining (frontal view); frontal carina absent; lunule slightly shining, high, rounded dorsally, with deep furrow; ocellar triangle as matt as frons, wide, with distinct contours; vibrissa and sbvb s are of the same length; maximum high of eye 4.8× as maximum high of gena; gena slightly acute ventrally; 1st flagellomere rounded (lateral view).
</paragraph>
<paragraph blockId="5.[174,1233,175,874]" pageId="5" pageNumber="412">
Wing (
<figureCitation box="[308,384,303,330]" captionStart="Figs 3038" captionStartId="5.[190,229,1758,1780]" captionTargetBox="[299,1156,895,1722]" captionTargetId="figure-306@5.[267,1169,874,1740]" captionTargetPageId="5" captionText="Figs 3038. M. pratensis sp. n., holotype: 30, 31 — male head; 32 — phallus, ventral view; 33 — same, lateral view; 34 — male wing; 35 — epandrium-hypandrium complex; 36 — ejaculatory apodeme; paratype (female terminalia): 37 — spermathecae, VII tergit, VII sternit; 38 — ventral seminal receptacle, egg guide, left blade, proctiger and cerci. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6449448" httpUri="https://zenodo.org/record/6449448/files/figure.png" pageId="5" pageNumber="412">fig. 34</figureCitation>
) slightly yellowish, with all veins brown; costa reaching M
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; last section of CuA
<subScript attach="left" box="[261,269,354,368]" fontSize="6" pageId="5" pageNumber="412">1</subScript>
0.85× as long as penultimate; costal sections
<quantity box="[809,884,335,362]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" metricValueMax="10.16" metricValueMin="5.08" pageId="5" pageNumber="412" unit="in" value="3.0" valueMax="4.0" valueMin="2.0">24 in</quantity>
the ratio of 3.87: 1.06: 1.00; calypter and fringe white, margin yellow. Wing length
<quantity box="[815,904,367,394]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="5" pageNumber="412" unit="mm" value="2.7">2.7 mm</quantity>
in both sexes.
</paragraph>
<paragraph blockId="5.[174,1233,175,874]" pageId="5" pageNumber="412">Mesonotum and scutellum slightly shining, blackish-grey (dorsal view); abdomen strongly shining, yellowish-green; 2 dc; ac in 8 irregular rows at level of 2nd dc; halters and legs dark brown.</paragraph>
<paragraph blockId="5.[174,1233,175,874]" pageId="5" pageNumber="412">
Male terminalia (
<figureCitation box="[432,648,495,522]" captionStart="Figs 3038" captionStartId="5.[190,229,1758,1780]" captionTargetBox="[299,1156,895,1722]" captionTargetId="figure-306@5.[267,1169,874,1740]" captionTargetPageId="5" captionText="Figs 3038. M. pratensis sp. n., holotype: 30, 31 — male head; 32 — phallus, ventral view; 33 — same, lateral view; 34 — male wing; 35 — epandrium-hypandrium complex; 36 — ejaculatory apodeme; paratype (female terminalia): 37 — spermathecae, VII tergit, VII sternit; 38 — ventral seminal receptacle, egg guide, left blade, proctiger and cerci. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6449448" httpUri="https://zenodo.org/record/6449448/files/figure.png" pageId="5" pageNumber="412">figs 32, 33, 35, 36</figureCitation>
). Epandrium covered with long hairs; inner surface of surstylus covered only with numerous short and small prensisetae; hypandrium V-shaped, with obtuse apex (ventral view); ejaculatory apodeme slightly longer than phallus; phallus
<quantity box="[313,416,591,618]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="5" pageNumber="412" unit="mm" value="0.21">0.21 mm</quantity>
long; distiphallus complex elongated, oval and narrowing posteriorly (ventral view); mesophallus long, flattened apically; basiphallus typical for the genus.
</paragraph>
<paragraph blockId="5.[174,1233,175,874]" pageId="5" pageNumber="412">
Female terminalia (
<figureCitation box="[454,579,655,682]" captionStart="Figs 3038" captionStartId="5.[190,229,1758,1780]" captionTargetBox="[299,1156,895,1722]" captionTargetId="figure-306@5.[267,1169,874,1740]" captionTargetPageId="5" captionText="Figs 3038. M. pratensis sp. n., holotype: 30, 31 — male head; 32 — phallus, ventral view; 33 — same, lateral view; 34 — male wing; 35 — epandrium-hypandrium complex; 36 — ejaculatory apodeme; paratype (female terminalia): 37 — spermathecae, VII tergit, VII sternit; 38 — ventral seminal receptacle, egg guide, left blade, proctiger and cerci. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6449448" httpUri="https://zenodo.org/record/6449448/files/figure.png" pageId="5" pageNumber="412">
figs 37
<emphasis bold="true" box="[532,547,655,681]" pageId="5" pageNumber="412"></emphasis>
38
</figureCitation>
). Both spermathecae oval, of different size, dark brown, flattened basally; both spermathecae with collars; egg guide relatively wide, 4.5× as long as maximum width, acute apically; middle and ventral part of medial membrane covered with numerous small brown teeth; ventral seminal receptacle, left blade, proctiger and cerci as on
<figureCitation box="[211,289,783,810]" captionStart="Figs 3038" captionStartId="5.[190,229,1758,1780]" captionTargetBox="[299,1156,895,1722]" captionTargetId="figure-306@5.[267,1169,874,1740]" captionTargetPageId="5" captionText="Figs 3038. M. pratensis sp. n., holotype: 30, 31 — male head; 32 — phallus, ventral view; 33 — same, lateral view; 34 — male wing; 35 — epandrium-hypandrium complex; 36 — ejaculatory apodeme; paratype (female terminalia): 37 — spermathecae, VII tergit, VII sternit; 38 — ventral seminal receptacle, egg guide, left blade, proctiger and cerci. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6449448" httpUri="https://zenodo.org/record/6449448/files/figure.png" pageId="5" pageNumber="412">fig. 38.</figureCitation>
</paragraph>
<paragraph blockId="5.[174,1233,175,874]" lastBlockId="6.[166,1225,175,682]" lastPageId="6" lastPageNumber="413" pageId="5" pageNumber="412">
B i o n o m i c s. One generation was recorded in June. Host plant unknown, but probably the flies are associated with
<taxonomicName box="[619,808,848,874]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="412" phylum="Tracheophyta" rank="species" species="jacea">
<emphasis box="[619,808,848,874]" italics="true" pageId="5" pageNumber="412">Centaurea jacea</emphasis>
</taxonomicName>
L., as
<specimenCount box="[886,1048,847,874]" pageId="5" pageNumber="412" type="generic">six specimens</specimenCount>
were caught in thicket of this plant in a small meadow, where no other possible host plants were present. Other specimens were collected on motley grass meadows with
<taxonomicName box="[913,1100,208,234]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="413" phylum="Tracheophyta" rank="species" species="jacea">
<emphasis box="[913,1100,208,234]" italics="true" pageId="6" pageNumber="413">Centaurea jacea</emphasis>
</taxonomicName>
L., too.
</paragraph>
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<paragraph blockId="5.[189,1248,1758,1860]" pageId="5" pageNumber="412">
Figs 3038.
<taxonomicName authority="Guglya Results &amp; Melanagromyza &amp; Guglya &amp; e &amp; Head &amp; Orbit, 2016" authorityName="Guglya Results &amp; Melanagromyza &amp; Guglya &amp; e &amp; Head &amp; Orbit" authorityYear="2016" box="[305,424,1759,1780]" class="Insecta" family="Agromyzidae" genus="Melanagromyza" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="412" phylum="Arthropoda" rank="species" species="pratensis" status="sp. n.">
<emphasis box="[305,424,1759,1780]" italics="true" pageId="5" pageNumber="412">M. pratensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[430,484,1758,1780]" pageId="5" pageNumber="412" rank="species">sp. n.</taxonomicNameLabel>
, holotype: 30, 31 — male head; 32 — phallus, ventral view; 33 — same, lateral view; 34 — male wing; 35 — epandrium-hypandrium complex; 36 — ejaculatory apodeme; paratype (female terminalia): 37 — spermathecae, VII tergit, VII sternit; 38 — ventral seminal receptacle, egg guide, left blade, proctiger and cerci. Scale bar 0.1 mm.
</paragraph>
</caption>
<paragraph blockId="6.[166,1225,175,682]" pageId="6" pageNumber="413">
D i a g n o s i s. The new species is superficially similar, in having proclinate and reclinate frorb sta together, wide lunule with slight furrow and white calypter, to
<taxonomicName authority="Hering, 1933" authorityName="Hering" authorityYear="1933" class="Insecta" family="Agromyzidae" genus="Melanagromyza" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="413" phylum="Arthropoda" rank="species" species="dettmeri">
<emphasis box="[1085,1224,271,297]" italics="true" pageId="6" pageNumber="413">M. dettmeri</emphasis>
Hering, 1933
</taxonomicName>
, running to that species in the key by
<bibRefCitation author="Papp, L. &amp; Cerny, M." box="[759,989,303,330]" pageId="6" pageNumber="413" pagination="1 - 416" refId="ref5650" refString="Papp, L., Cerny, M. 2015. Agromyzidae (Diptera) of Hungary. Vol. 1. Agromyzinae. Pars Ltd., Nagykovacsi, 1 - 416." type="book chapter" year="2015">Papp, Černý (2015)</bibRefCitation>
, but differing by the orbit slightly projecting above eye only at level of orb s (lateral view); last section of CuA
<subScript attach="left" box="[1193,1201,354,368]" fontSize="6" pageId="6" pageNumber="413">1</subScript>
&gt; 0.75× (0.85×) as long as penultimate; margin yellow; mesonotum and scutellum slightly shining, blackish-grey (dorsal view); abdomen strongly shining, yellowish-green; inner surface of surstylus covered only with numerous short and small spines; setae on cerci short, whereas in
<taxonomicName authorityName="Hering" authorityYear="1933" box="[376,514,463,489]" class="Insecta" family="Agromyzidae" genus="Melanagromyza" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="413" phylum="Arthropoda" rank="species" species="dettmeri">
<emphasis box="[376,514,463,489]" italics="true" pageId="6" pageNumber="413">M. dettmeri</emphasis>
</taxonomicName>
orbit projecting as narrow ring above eye (lateral view); last section of CuA
<subScript attach="left" box="[346,354,514,528]" fontSize="6" pageId="6" pageNumber="413">1</subScript>
&lt;0.75× (0.67×) as long as penultimate; margin pale brown; mesonotum “matt, faintly greenish or coppery, abdomen distinctly shining, variably coppery-greenish or bluish”; “inner surface of surstylus with group of denser and long setae on ventral margin and group numerous spines on inner surface” (
<bibRefCitation author="Papp, L. &amp; Cerny, M." box="[762,983,591,618]" pageId="6" pageNumber="413" pagination="1 - 416" refId="ref5650" refString="Papp, L., Cerny, M. 2015. Agromyzidae (Diptera) of Hungary. Vol. 1. Agromyzinae. Pars Ltd., Nagykovacsi, 1 - 416." type="book chapter" year="2015">Papp, Černý, 2015</bibRefCitation>
); setae on cerci long.
</paragraph>
<paragraph blockId="6.[166,1225,175,682]" pageId="6" pageNumber="413">E t y m o l o g y. The name reflects the locality (a thicket of meadow plants), where all the specimens were collected.</paragraph>
</subSubSection>
</treatment>
</document>