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<document id="A33A51AFB5964C45188F82D3CD8585AD" ID-CLB-Dataset="4384" ID-DOI="10.1111/j.1096-3642.2007.00312.x" ID-GBIF-Dataset="1339ad9b-f051-4055-ad11-05e6d4245392" ID-ISSN="0024-4082" ID-Zenodo-Dep="5431970" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630429432031" checkinUser="felipe" docAuthor="Reisz, Robert R., Müller, Johannes, Tsuji, Linda &amp; Scott, Diane" docDate="2007" docId="03824254FFD4FFD027C03A169755FAAB" docLanguage="en" docName="j.1096-3642.2007.00312.x.pdf" docOrigin="Zoological Journal of the Linnean Society 151 (1)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2007.00312.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Belebey vegrandis" docType="treatment" docVersion="7" lastPageNumber="192" masterDocId="FFBB3A2CFFD5FFD12444385B9362FF8D" masterDocTitle="The cranial osteology of Belebey vegrandis (Parareptilia: Bolosauridae), from the Middle Permian of Russia, and its bearing on reptilian evolution" masterLastPageNumber="214" masterPageNumber="191" pageNumber="192" updateTime="1699273722102" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="F093DB5D7C6C3A6E10CAB5762188CA46">The cranial osteology of Belebey vegrandis (Parareptilia: Bolosauridae), from the Middle Permian of Russia, and its bearing on reptilian evolution</mods:title>
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<mods:namePart id="E84F5F54ECE4D70E00D996F56ECAC772">Reisz, Robert R.</mods:namePart>
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<mods:namePart id="E084A6F2E0E4B249D6B881433D96FDFB">Müller, Johannes</mods:namePart>
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<mods:namePart id="4E0478869CB15F770874B607C5B27406">Tsuji, Linda</mods:namePart>
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<mods:namePart id="883BEF0BAEF25079E7ABA3A06E6B3030">Scott, Diane</mods:namePart>
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<treatment id="03824254FFD4FFD027C03A169755FAAB" ID-DOI="http://doi.org/10.5281/zenodo.5489406" ID-GBIF-Taxon="185436240" ID-Zenodo-Dep="5489406" LSID="urn:lsid:plazi:treatment:03824254FFD4FFD027C03A169755FAAB" httpUri="http://treatment.plazi.org/id/03824254FFD4FFD027C03A169755FAAB" lastPageNumber="192" pageId="1" pageNumber="192">
<subSubSection id="C331A0C9FFD4FFD027C03A16963AFDE9" box="[900,1368,588,612]" pageId="1" pageNumber="192" type="nomenclature">
<paragraph id="8B94F342FFD4FFD027C03A16963AFDE9" blockId="1.[900,1368,588,612]" box="[900,1368,588,612]" pageId="1" pageNumber="192">
<heading id="D0DC442EFFD4FFD027C03A16963AFDE9" box="[900,1368,588,612]" centered="true" fontSize="9" level="2" pageId="1" pageNumber="192" reason="2">
<taxonomicName id="4C2B88C1FFD4FFD027C03A169716FDEE" baseAuthorityName="Ivakhnenko" baseAuthorityYear="1973" box="[900,1140,589,612]" class="Reptilia" family="Bolosauridae" genus="Belebey" higherTaxonomySource="GBIF" kingdom="Animalia" order="Bolosauria" pageId="1" pageNumber="199" phylum="Chordata" rank="species" species="vegrandis">
<emphasis id="B95F2F50FFD4FFD027C03A169716FDEE" box="[900,1140,589,612]" italics="true" pageId="1" pageNumber="192">BELEBEY VEGRANDIS</emphasis>
</taxonomicName>
<bibRefCitation id="EFBA8EB3FFD4FFD020383A0B963AFDE9" author="Ivakhnenko MF" box="[1148,1368,588,612]" pageId="1" pageNumber="192" pagination="131 - 134" refId="ref11265" refString="Ivakhnenko MF. 1973. New Cisuralian cotylosaurs. Paleontologicheskii Zhurnal 1973: 131 - 134." type="journal article" year="1973">IVAKHNENKO, 1973</bibRefCitation>
</heading>
</paragraph>
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<subSubSection id="C331A0C9FFD4FFD0277E3A269755FAAB" pageId="1" pageNumber="192" type="materials_examined">
<paragraph id="8B94F342FFD4FFD0277E3A26967CFD5D" blockId="1.[826,1441,637,720]" pageId="1" pageNumber="192">
<materialsCitation id="3B43F91FFFD4FFD0277E3A26967CFD5D" ID-GBIF-Occurrence="3341489303" collectionCode="PIN" country="Russia" location="Palaeontological Institute" municipality="Russian Academy of Science" pageId="1" pageNumber="192" specimenCount="1" stateProvince="Moscow" typeStatus="holotype">
<emphasis id="B95F2F50FFD4FFD0277E3A2690C7FD1F" box="[826,933,637,658]" italics="true" pageId="1" pageNumber="192">
<typeStatus id="54904DE0FFD4FFD0277E3A2690FDFD1F" box="[826,927,637,658]" pageId="1" pageNumber="192" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="ED3A6B87FFD4FFD027F53A269082FD1F" box="[945,992,637,658]" country="Russia" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="1" pageNumber="192">PIN</collectionCode>
(
<location id="8EF4A599FFD4FFD027BE3A26964CFD1E" LSID="urn:lsid:plazi:treatment:03824254FFD4FFD027C03A169755FAAB:8EF4A599FFD4FFD027BE3A26964CFD1E" box="[1018,1326,637,659]" country="Russia" municipality="Russian Academy of Science" name="Palaeontological Institute" pageId="1" pageNumber="192" stateProvince="Moscow">Palaeontological Institute</location>
,
<collectingMunicipality id="6BF06938FFD4FFD021013A269778FD3C" pageId="1" pageNumber="192">Russian Academy of Science</collectingMunicipality>
,
<collectingRegion id="49EF3DA0FFD4FFD020603AC7971FFD3C" box="[1060,1149,668,689]" country="Russia" name="Moscow" pageId="1" pageNumber="192">Moscow</collectingRegion>
,
<collectingCountry id="F33CB3D2FFD4FFD020C23AC097B6FD3C" box="[1158,1236,667,689]" name="Russia" pageId="1" pageNumber="192">Russia</collectingCountry>
) 104/50 maxillary bone with partly preserved palatine bone.
</materialsCitation>
</paragraph>
<paragraph id="8B94F342FFD4FFD0277E3AB39646FC3B" blockId="1.[826,1442,744,950]" pageId="1" pageNumber="192">
<emphasis id="B95F2F50FFD4FFD0277E3AB3977EFD73" box="[826,1052,744,766]" italics="true" pageId="1" pageNumber="192">Referred specimens:</emphasis>
<materialsCitation id="3B43F91FFFD4FFD0206D3AB39642FC3B" ID-GBIF-Occurrence="3341489301" country="Russia" location="Saratov" pageId="1" pageNumber="192" specimenCount="1" stateProvince="Saratov">
SGU (
<collectingRegion id="49EF3DA0FFD4FFD0202A3AB397AAFD70" box="[1134,1224,744,765]" country="Russia" name="Saratov" pageId="1" pageNumber="192">Saratov</collectingRegion>
Geological University,
<collectingRegion id="49EF3DA0FFD4FFD027303B5C90AEFC91" box="[884,972,775,796]" country="Russia" name="Saratov" pageId="1" pageNumber="192">Saratov</collectingRegion>
,
<collectingCountry id="F33CB3D2FFD4FFD0279A3B5C974EFC90" box="[990,1068,775,797]" name="Russia" pageId="1" pageNumber="192">Russia</collectingCountry>
) 104/B-2020, a small complete skull that has been disarticulated (
<figureCitation id="1310EFC7FFD4FFD020AC3B7E9633FCB6" box="[1256,1361,805,827]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStart-5="Figure 6" captionStartId-0="2.[142,220,1261,1280]" captionStartId-1="3.[162,240,1029,1048]" captionStartId-2="3.[162,240,1662,1681]" captionStartId-3="4.[142,220,852,871]" captionStartId-4="4.[142,220,1825,1844]" captionStartId-5="5.[162,240,1099,1118]" captionTargetBox-0="[302,1262,192,1229]" captionTargetBox-1="[322,1282,194,997]" captionTargetBox-2="[166,774,1145,1629]" captionTargetBox-3="[302,1262,195,820]" captionTargetBox-4="[302,1262,984,1792]" captionTargetBox-5="[166,774,197,1066]" captionTargetId-0="figure-373@2.[302,1262,192,1229]" captionTargetId-1="figure-282@3.[322,1282,193,997]" captionTargetId-2="figure-347@3.[166,774,1143,1630]" captionTargetId-3="figure-54@4.[302,1262,195,820]" captionTargetId-4="figure-84@4.[302,1262,984,1792]" captionTargetId-5="figure-736@5.[166,775,195,1066]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="4" captionTargetPageId-5="5" captionText-0="Figure 1. Skull of Belebey vegrandis, SGU 104/B-2020: antorbital region and palate in lateral view, and skull table and temporal region in dorsal view. These are parts of the same skull, prepared and separated from each other. Scale bar = 1 cm; see Appendix 1 for abbreviations list." captionText-1="Figure 2. Skull of Belebey vegrandis, SGU 104/B-2020: antorbital region of the skull roof and the palate in both dorsal and ventral views. Scale bar = 1 cm; see Appendix 1 for abbreviations list." captionText-2="Figure 3. Skull of Belebey vegrandis, SGU 104/B-2020: photograph of the antorbital area of the skull in anterior view, showing details of the palate, the medial process of the prefrontal, and the palatineprefrontal buttress. The dorsal surface of the maxilla, where the largest maxillary teeth are attached, is pockmarked. Scale bar = 1 cm; see Appendix 1 for abbreviations list." captionText-3="Figure 4. Belebey vegrandis, SGU 104/B-2020: left mandible in medial and lateral views. Scale bar = 1 cm; see Appendix 1 for abbreviations list." captionText-4="Figure 5. Belebey vegrandis, SGU 104/B-2020: right mandible in medial, lateral and occlusal views. Scale bar = 1 cm; see Appendix 1 for abbreviations list." captionText-5="Figure 6. Belebey vegrandis, SGU 104/B-2020: photograph of left and right mandibular rami in occlusal view. Scale bar = 1 cm." figureDoi-0="http://doi.org/10.5281/zenodo.5431972" figureDoi-1="http://doi.org/10.5281/zenodo.5431974" figureDoi-2="http://doi.org/10.5281/zenodo.5431976" figureDoi-3="http://doi.org/10.5281/zenodo.5431978" figureDoi-4="http://doi.org/10.5281/zenodo.5431980" figureDoi-5="http://doi.org/10.5281/zenodo.5431982" httpUri-0="https://zenodo.org/record/5431972/files/figure.png" httpUri-1="https://zenodo.org/record/5431974/files/figure.png" httpUri-2="https://zenodo.org/record/5431976/files/figure.png" httpUri-3="https://zenodo.org/record/5431978/files/figure.png" httpUri-4="https://zenodo.org/record/5431980/files/figure.png" httpUri-5="https://zenodo.org/record/5431982/files/figure.png" pageId="1" pageNumber="192">Figs 16</figureCitation>
); SGU 104/B-2021, a slightly larger, complete skull (
<figureCitation id="1310EFC7FFD4FFD021113B1F96FFFCD4" box="[1365,1437,836,858]" captionStart="Figure 7" captionStartId="6.[142,220,1302,1321]" captionTargetBox="[302,1262,197,1270]" captionTargetId="figure-333@6.[302,1262,195,1270]" captionTargetPageId="6" captionText="Figure 7. Belebey vegrandis, SGU 104/B-2021: skull and mandible in dorsal and lateral views. Scale bar = 1 cm; see Appendix 1 for abbreviations list." figureDoi="http://doi.org/10.5281/zenodo.5431986" httpUri="https://zenodo.org/record/5431986/files/figure.png" pageId="1" pageNumber="192">Figs 7</figureCitation>
,
<figureCitation id="1310EFC7FFD4FFD0277E3B38902AFCF4" box="[826,840,867,889]" captionStart="Figure 8" captionStartId="7.[162,240,1025,1044]" captionTargetBox="[166,774,193,992]" captionTargetId="figure-758@7.[166,775,191,992]" captionTargetPageId="7" captionText="Figure 8. Belebey vegrandis, SGU 104/B-2021: photograph of skull and mandible in palatal view. The skull was not available for illustration in this view. Arrow points to ventral location of the mandibular foramen (mf). Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5431988" httpUri="https://zenodo.org/record/5431988/files/figure.png" pageId="1" pageNumber="192">8</figureCitation>
); SGU 104/B-2022, a large, partial skull, preorbital region not preserved. This large individual was available for study but could not be illustrated
</materialsCitation>
.
</paragraph>
<paragraph id="8B94F342FFD4FFD0277E3B9490EDFBAC" blockId="1.[826,1443,974,1057]" pageId="1" pageNumber="192">
<emphasis id="B95F2F50FFD4FFD0277E3B94974EFC6E" box="[826,1068,974,996]" italics="true" pageId="1" pageNumber="192">Horizon and locality:</emphasis>
Upper Kazanian, Middle Permian;
<materialsCitation id="3B43F91FFFD4FFD027C63BB690E9FBAC" ID-GBIF-Occurrence="3341489302" country="Russia" location="Bashkortostan" pageId="1" pageNumber="192" specimenCount="1" stateProvince="Bashkortostan">
<collectingRegion id="49EF3DA0FFD4FFD027C63BB69748FB8E" box="[898,1066,1005,1027]" country="Russia" name="Bashkortostan" pageId="1" pageNumber="192">Bashkortostan</collectingRegion>
and
<collectingRegion id="49EF3DA0FFD4FFD020263BB697B3FB8E" box="[1122,1233,1005,1027]" country="Russia" name="Orenburg" pageId="1" pageNumber="192">Orenburg</collectingRegion>
regions, Southern
<collectingCountry id="F33CB3D2FFD4FFD0277E3C5090E9FBAC" box="[826,907,1035,1057]" name="Russia" pageId="1" pageNumber="192">Russia</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8B94F342FFD4FFD0277E3C619755FAAB" blockId="1.[826,1442,1081,1318]" pageId="1" pageNumber="192">
<emphasis id="B95F2F50FFD4FFD0277E3C6190D3FBC2" box="[826,945,1082,1103]" italics="true" pageId="1" pageNumber="192">Diagnosis:</emphasis>
Bolosaurid parareptiles characterized by the presence of strongly procumbent incisiform dentition anteriorly, and linguo-labially elongated, bulbous, posterior cheek teeth with strongly developed longitudinal striations. Non-dental potential autapomorphies cannot be recognized because most of the other bolosaurid taxa are known from fragmentary cranial and mandibular elements.
</paragraph>
</subSubSection>
</treatment>
</document>