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<document id="A65A255FFDD85606F0EF64D5F33ADDB7" ID-CLB-Dataset="8752" ID-DOI="10.11646/zootaxa.4899.1.16" ID-GBIF-Dataset="dde87a19-79a2-44a3-8533-b8970944cb99" ID-ISSN="1175-5326" ID-Zenodo-Dep="4456831" ID-ZooBank="27E67BBB-DFD0-4A96-8269-9E1CB6153B83" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609326131882" checkinUser="plazi" docAuthor="Salgado, Alexandre &amp; Ruiz, Gustavo R. S." docDate="2020" docId="03403F11FFA7FFBF538BF990092BFF5D" docLanguage="en" docName="zootaxa.4899.1.16.pdf" docOrigin="Zootaxa 4899 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Marma spelunca Salgado &amp; Ruiz 2020, sp. nov." docType="treatment" docVersion="8" lastPageNumber="350" masterDocId="FF794769FF9EFF80531CFFE40C1AFF88" masterDocTitle="Taxonomic revision of Marma Simon, 1902 (Araneae: Salticidae: Euophryini)" masterLastPageNumber="353" masterPageNumber="287" pageNumber="344" updateTime="1698848557116" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="14A1AE4CAE7A5952DFE2238ACF1D0FC4">
<mods:title id="3C01C3DE6A30D34467525AEE29776336">Taxonomic revision of Marma Simon, 1902 (Araneae: Salticidae: Euophryini)</mods:title>
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<mods:name id="244C3E6F0F354FAF2A2FCC0DAB2E7B37" type="personal">
<mods:role id="17C97F05D3ABC557E3105910FCBA20A7">
<mods:roleTerm id="D0E6B50BA519AA352C294195DD91B72A">Author</mods:roleTerm>
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<mods:namePart id="39AA052BDE8DB099FA6EBE0AF598B8B8">Salgado, Alexandre</mods:namePart>
<mods:nameIdentifier id="D6CA6516EA4BB87F25F8EDC6BD759D5B" type="ORCID">0000-0002-9562-403X</mods:nameIdentifier>
<mods:affiliation id="FE67CC6D571D4421D65E3A482D951D52">Programa de Pós-Graduação em Zoologia, Universidade Federal do Pará / Museu Paraense Emílio Goeldi, Belém, PA, Brazil Instituto de Ciências Biológicas, Universidade Federal do Pará, Rua Augusto Corrêa, 01, CEP 66075 - 110, Belém, PA, Brazil</mods:affiliation>
<mods:nameIdentifier id="31D5615712A6CF2914DD7C3860D29009" type="email">alexandre.salgado.souza@gmail.com</mods:nameIdentifier>
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<mods:namePart id="7ACB1D139AD92A7BF923ED0F51387D7B">Ruiz, Gustavo R. S.</mods:namePart>
<mods:nameIdentifier id="D1AADA9D2CE44255A59925FE72490DF8" type="ORCID">0000-0002-7326-7000</mods:nameIdentifier>
<mods:affiliation id="02E009E92520349764376BC259CED803">Instituto de Ciências Biológicas, Universidade Federal do Pará, Rua Augusto Corrêa, 01, CEP 66075 - 110, Belém, PA, Brazil</mods:affiliation>
<mods:nameIdentifier id="B9804952DB419555D0FC3DAB1E7CD3C3" type="email">gustavoruiz86@hotmail.com</mods:nameIdentifier>
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<mods:title id="F52B099BD8BCAE433AA4C459471D29A5">Zootaxa</mods:title>
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<mods:part id="BC41D3A5B28F0954A4DBD4CAD8FB5A00">
<mods:date id="3A775BC7A3A93264D4E570A7E4707DA4">2020</mods:date>
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<mods:number id="45260305EF22D3D4DDA802E555B4F722">2020-12-30</mods:number>
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<mods:number id="54EB09088FE4B376D6C2AB840F58CC20">4899</mods:number>
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<mods:classification id="36F92D93284869AF6914183EC98FDBE3">journal article</mods:classification>
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<treatment id="03403F11FFA7FFBF538BF990092BFF5D" ID-DOI="http://doi.org/10.5281/zenodo.4456907" ID-GBIF-Taxon="177010983" ID-Zenodo-Dep="4456907" LSID="urn:lsid:plazi:treatment:03403F11FFA7FFBF538BF990092BFF5D" httpUri="http://treatment.plazi.org/id/03403F11FFA7FFBF538BF990092BFF5D" lastPageId="63" lastPageNumber="350" pageId="57" pageNumber="344">
<subSubSection id="C3F3DD8CFFA7FFB9538BF9900DD3F906" box="[151,457,1652,1679]" pageId="57" pageNumber="344" type="nomenclature">
<paragraph id="8B568E07FFA7FFB9538BF9900DD3F906" blockId="57.[151,457,1652,1715]" box="[151,457,1652,1679]" pageId="57" pageNumber="344">
<heading id="D01E396BFFA7FFB9538BF9900DD3F906" bold="true" box="[151,457,1652,1679]" fontSize="11" level="1" pageId="57" pageNumber="344" reason="1">
<emphasis id="B99D5215FFA7FFB9538BF9900DD3F906" bold="true" box="[151,457,1652,1679]" pageId="57" pageNumber="344">
<taxonomicName id="4CE9F584FFA7FFB9538BF9900D7EF907" ID-CoL="98Z85" authority="Salgado &amp; Ruiz, 2020" authorityName="Salgado &amp; Ruiz" authorityYear="2020" box="[151,356,1652,1679]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="344" phylum="Arthropoda" rank="species" species="spelunca" status="sp. nov.">
<emphasis id="B99D5215FFA7FFB9538BF9900D7EF907" bold="true" box="[151,356,1652,1679]" italics="true" pageId="57" pageNumber="344">Marma spelunca</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2AEEF6EFFA7FFB95277F9900DD3F906" box="[363,457,1652,1678]" pageId="57" pageNumber="344" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3F3DD8CFFA7FFB9538BF97C0DB0F93B" box="[151,426,1688,1715]" pageId="57" pageNumber="344" type="description">
<paragraph id="8B568E07FFA7FFB9538BF97C0DB0F93B" blockId="57.[151,457,1652,1715]" box="[151,426,1688,1715]" pageId="57" pageNumber="344">
<figureCitation id="13D29282FFA7FFB9538BF97C0D21F93B" box="[151,315,1688,1715]" captionStart-0="FIGURE 53" captionStart-1="FIGURE 54" captionStart-2="FIGURE 55" captionStart-3="FIGURE 56" captionStartId-0="58.[151,250,1894,1918]" captionStartId-1="59.[151,250,1921,1945]" captionStartId-2="60.[151,250,1210,1234]" captionStartId-3="61.[151,250,1861,1885]" captionTargetBox-0="[151,1435,388,1871]" captionTargetBox-1="[156,1424,749,1894]" captionTargetBox-2="[157,1424,191,1176]" captionTargetBox-3="[177,1409,181,1837]" captionTargetId-0="figure-131@58.[151,1435,381,1871]" captionTargetId-1="figure-300@59.[151,1435,741,1897]" captionTargetId-2="figure-24@60.[151,1435,181,1186]" captionTargetId-3="figure-17@61.[177,1409,181,1837]" captionTargetPageId-0="58" captionTargetPageId-1="59" captionTargetPageId-2="60" captionTargetPageId-3="61" captionText-0="FIGURE 53. Male holotype of Marma spelunca sp. nov. (MPEG 34347). A–B habitus (A dorsal, B ventral); C–D left palp (ventral; D retrolateral); D–E type locality, Nova Kararaô cave opening (E view of the entrance; F live specimen, indicated by white arrow, walking on moss present on walls around opening)." captionText-1="FIGURE 54. Female paratype of Marma spelunca sp. nov. (MPEG 34348). A–B habitus (A dorsal, B ventral); C–D cleared epigyne/vulva (C ventral, D detail of right spermatheca, dorsal). Abbreviations: BG—Bennett’s gland; cd—copulatory duct; co—copulatory opening; fd—fertilization duct; PS—Primary spermatheca; SS—secondary spermatheca." captionText-2="FIGURE 55. Marma spelunca sp. nov. A–E left male palp (A ventral; B retrolateral; C proventral; D dorsal; E cleared bulb, ventral); F–G epigyne/vulva (F ventral, G cleared, ventral).Abbreviations: BG—Bennett’s gland; cc—cymbial conductor; cdcopulatory duct; co—copulatory opening; dh—distal hematodocha; e—embolus shaft; ed—embolic disc; fd—fertilization duct; PED—process on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermatheca." captionText-3="FIGURE 56. Male of Marma spelunca sp. nov. A–D left palp (A bulb, dorsal; B detail of fundus and subtegulum, dorsal; C dissected embolic disc and embolus, dorsal; D apical region of cymbium, ventral. Abbreviations: bh—basal hematodocha; cc—cymbial conductor; e—embolus shaft; fnd—fundus; PED—process on embolic disc; st—subtegulum. White arrows in C show spikes of embolus shaft and black arrow shows groove; white arrow in D shows depression of cymbial conductor." figureDoi-0="http://doi.org/10.5281/zenodo.4456969" figureDoi-1="http://doi.org/10.5281/zenodo.4456971" figureDoi-2="http://doi.org/10.5281/zenodo.4456973" figureDoi-3="http://doi.org/10.5281/zenodo.4456977" httpUri-0="https://zenodo.org/record/4456969/files/figure.png" httpUri-1="https://zenodo.org/record/4456971/files/figure.png" httpUri-2="https://zenodo.org/record/4456973/files/figure.png" httpUri-3="https://zenodo.org/record/4456977/files/figure.png" pageId="57" pageNumber="344">Figures 53–56</figureCitation>
,
<figureCitation id="13D29282FFA7FFB9525AF97C0D74F93B" box="[326,366,1688,1715]" captionStart="FIGURE 57" captionStartId="62.[151,250,1843,1867]" captionTargetBox="[159,1422,193,1813]" captionTargetId="figure-24@62.[151,1435,181,1819]" captionTargetPageId="62" captionText="FIGURE 57. Left male palp, ventral [A Marma baeri Simon, 1902; B M. linae sp. nov.; C M. pipa sp. nov.; D M. wesolowskae sp. nov.; E M. nigritarsis (Simon, 1900); F M. sinuosa sp. nov., G M. argentina (Mello-Leit„o, 1941), H M. rosea (Mello- Leit„o, 1941), I M. spelunca sp. nov.]. Colors represent homologous structures." figureDoi="http://doi.org/10.5281/zenodo.4456979" httpUri="https://zenodo.org/record/4456979/files/figure.png" pageId="57" pageNumber="344">57I</figureCitation>
,
<figureCitation id="13D29282FFA7FFB95266F97C0DB0F93B" box="[378,426,1688,1715]" captionStart="FIGURE 58" captionStartId="63.[151,250,1595,1619]" captionTargetBox="[156,1429,271,1549]" captionTargetId="figure-41@63.[151,1435,237,1571]" captionTargetPageId="63" captionText="FIGURE 58. Epigyne/vulva, A–C, E–K ventral, D dorsal [A Marma baeri Simon, 1902; B M. linae sp. nov.; C M. abaira sp. nov.; D M. femella (Caporiacco, 1955); E M. pipa sp. nov.; F M. wesolowskae sp. nov.; G M. nigritarsis (Simon, 1900); H M. sinuosa sp. nov.; I M. argentina (Mello-Leitão, 1941); J M. rosea (Mello-Leitão, 1941); K M. spelunca sp. nov.]. Colors represent homologous structures; arrow in D shows hypothetical position of secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456981" httpUri="https://zenodo.org/record/4456981/files/figure.png" pageId="57" pageNumber="344">58K</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3F3DD8CFFA7FFB9538BF9040D04F896" pageId="57" pageNumber="344" type="etymology">
<paragraph id="8B568E07FFA7FFB9538BF9040D04F896" blockId="57.[151,1437,1760,2039]" pageId="57" pageNumber="344">
<emphasis id="B99D5215FFA7FFB9538BF9040D04F972" bold="true" box="[151,286,1760,1786]" pageId="57" pageNumber="344">Etymology.</emphasis>
The specific epithet is a Latin noun meaning “cave”, in reference to the habitat where the specimens were found.
</paragraph>
</subSubSection>
<subSubSection id="C3F3DD8CFFA7FFBA53DBF8CC0E62FF71" lastPageId="58" lastPageNumber="345" pageId="57" pageNumber="344" type="diagnosis">
<paragraph id="8B568E07FFA7FFBA53DBF8CC0E62FF71" blockId="57.[151,1437,1760,2039]" lastBlockId="58.[151,1437,150,357]" lastPageId="58" lastPageNumber="345" pageId="57" pageNumber="344">
<emphasis id="B99D5215FFA7FFB953DBF8CC0D58F8CA" bold="true" box="[199,322,1832,1858]" pageId="57" pageNumber="344">Diagnosis.</emphasis>
<taxonomicName id="4CE9F584FFA7FFB95255F8CD0E12F8CA" authority="Salgado &amp; Ruiz, 2020" authorityName="Salgado &amp; Ruiz" authorityYear="2020" box="[329,520,1832,1858]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="344" phylum="Arthropoda" rank="species" species="spelunca" status="sp. nov.">
<emphasis id="B99D5215FFA7FFB95255F8CD0E12F8CA" box="[329,520,1832,1858]" italics="true" pageId="57" pageNumber="344">Marma spelunca</emphasis>
</taxonomicName>
<emphasis id="B99D5215FFA7FFB95113F8CC0E73F8CA" bold="true" box="[527,617,1832,1858]" pageId="57" pageNumber="344">
<taxonomicNameLabel id="A2AEEF6EFFA7FFB95113F8CC0E73F8CA" box="[527,617,1832,1858]" pageId="57" pageNumber="344" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
shares most similarities with
<taxonomicName id="4CE9F584FFA7FFB950A0F8CD0805F8CA" box="[956,1055,1833,1858]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="344" phylum="Arthropoda" rank="species" species="rosea">
<emphasis id="B99D5215FFA7FFB950A0F8CD0805F8CA" box="[956,1055,1833,1858]" italics="true" pageId="57" pageNumber="344">M. rosea</emphasis>
</taxonomicName>
, both with the same length of the process on the embolic disc and cymbium (proportionally to the bulb) (
<figureCitation id="13D29282FFA7FFB950B3F8A8081FF8EF" box="[943,1029,1868,1895]" captionStart="FIGURE 57" captionStartId="62.[151,250,1843,1867]" captionTargetBox="[159,1422,193,1813]" captionTargetId="figure-24@62.[151,1435,181,1819]" captionTargetPageId="62" captionText="FIGURE 57. Left male palp, ventral [A Marma baeri Simon, 1902; B M. linae sp. nov.; C M. pipa sp. nov.; D M. wesolowskae sp. nov.; E M. nigritarsis (Simon, 1900); F M. sinuosa sp. nov., G M. argentina (Mello-Leit„o, 1941), H M. rosea (Mello- Leit„o, 1941), I M. spelunca sp. nov.]. Colors represent homologous structures." figureDoi="http://doi.org/10.5281/zenodo.4456979" httpUri="https://zenodo.org/record/4456979/files/figure.png" pageId="57" pageNumber="344">Figs 57</figureCitation>
H–I), and similar pattern of epigynal structure (copulatory ducts with similar length, encircling primary spermathecae and almost touching each other;
<figureCitation id="13D29282FFA7FFB9538BF8700CF6F827" box="[151,236,1940,1967]" captionStart="FIGURE 58" captionStartId="63.[151,250,1595,1619]" captionTargetBox="[156,1429,271,1549]" captionTargetId="figure-41@63.[151,1435,237,1571]" captionTargetPageId="63" captionText="FIGURE 58. Epigyne/vulva, A–C, E–K ventral, D dorsal [A Marma baeri Simon, 1902; B M. linae sp. nov.; C M. abaira sp. nov.; D M. femella (Caporiacco, 1955); E M. pipa sp. nov.; F M. wesolowskae sp. nov.; G M. nigritarsis (Simon, 1900); H M. sinuosa sp. nov.; I M. argentina (Mello-Leitão, 1941); J M. rosea (Mello-Leitão, 1941); K M. spelunca sp. nov.]. Colors represent homologous structures; arrow in D shows hypothetical position of secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456981" httpUri="https://zenodo.org/record/4456981/files/figure.png" pageId="57" pageNumber="344">Figs 58</figureCitation>
J–K). However, males of
<taxonomicName id="4CE9F584FFA7FFB95111F8710E8DF826" authority="Salgado &amp; Ruiz, 2020" authorityName="Salgado &amp; Ruiz" authorityYear="2020" box="[525,663,1940,1966]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="344" phylum="Arthropoda" rank="species" species="spelunca" status="sp. nov.">
<emphasis id="B99D5215FFA7FFB95111F8710E8DF826" box="[525,663,1940,1966]" italics="true" pageId="57" pageNumber="344">M. spelunca</emphasis>
</taxonomicName>
<emphasis id="B99D5215FFA7FFB95183F8700EE3F826" bold="true" box="[671,761,1940,1966]" pageId="57" pageNumber="344">
<taxonomicNameLabel id="A2AEEF6EFFA7FFB95183F8700EE3F826" box="[671,761,1940,1966]" pageId="57" pageNumber="344" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished by having a symmetrical embolic disc (both edges with the same degree of curvature;
<figureCitation id="13D29282FFA7FFB951AEF85C0F06F85B" box="[690,796,1976,2003]" captionStart="FIGURE 53" captionStartId="58.[151,250,1894,1918]" captionTargetBox="[151,1435,388,1871]" captionTargetId="figure-131@58.[151,1435,381,1871]" captionTargetPageId="58" captionText="FIGURE 53. Male holotype of Marma spelunca sp. nov. (MPEG 34347). A–B habitus (A dorsal, B ventral); C–D left palp (ventral; D retrolateral); D–E type locality, Nova Kararaô cave opening (E view of the entrance; F live specimen, indicated by white arrow, walking on moss present on walls around opening)." figureDoi="http://doi.org/10.5281/zenodo.4456969" httpUri="https://zenodo.org/record/4456969/files/figure.png" pageId="57" pageNumber="344">Figs 53C</figureCitation>
,
<figureCitation id="13D29282FFA7FFB95034F85C0F66F85A" box="[808,892,1976,2003]" captionStart="FIGURE 55" captionStartId="60.[151,250,1210,1234]" captionTargetBox="[157,1424,191,1176]" captionTargetId="figure-24@60.[151,1435,181,1186]" captionTargetPageId="60" captionText="FIGURE 55. Marma spelunca sp. nov. A–E left male palp (A ventral; B retrolateral; C proventral; D dorsal; E cleared bulb, ventral); F–G epigyne/vulva (F ventral, G cleared, ventral).Abbreviations: BG—Bennett’s gland; cc—cymbial conductor; cdcopulatory duct; co—copulatory opening; dh—distal hematodocha; e—embolus shaft; ed—embolic disc; fd—fertilization duct; PED—process on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456973" httpUri="https://zenodo.org/record/4456973/files/figure.png" pageId="57" pageNumber="344">55A, C</figureCitation>
,
<figureCitation id="13D29282FFA7FFB95095F85C0FA8F85B" box="[905,946,1976,2003]" captionStart="FIGURE 57" captionStartId="62.[151,250,1843,1867]" captionTargetBox="[159,1422,193,1813]" captionTargetId="figure-24@62.[151,1435,181,1819]" captionTargetPageId="62" captionText="FIGURE 57. Left male palp, ventral [A Marma baeri Simon, 1902; B M. linae sp. nov.; C M. pipa sp. nov.; D M. wesolowskae sp. nov.; E M. nigritarsis (Simon, 1900); F M. sinuosa sp. nov., G M. argentina (Mello-Leit„o, 1941), H M. rosea (Mello- Leit„o, 1941), I M. spelunca sp. nov.]. Colors represent homologous structures." figureDoi="http://doi.org/10.5281/zenodo.4456979" httpUri="https://zenodo.org/record/4456979/files/figure.png" pageId="57" pageNumber="344">57I</figureCitation>
), whereas in
<taxonomicName id="4CE9F584FFA7FFB95754F85D08B1F85A" box="[1096,1195,1977,2002]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="344" phylum="Arthropoda" rank="species" species="rosea">
<emphasis id="B99D5215FFA7FFB95754F85D08B1F85A" box="[1096,1195,1977,2002]" italics="true" pageId="57" pageNumber="344">M. rosea</emphasis>
</taxonomicName>
, the retrolateral edge of the embolic disc has more accentuated curvature (
<figureCitation id="13D29282FFA7FFB951FFF8380F56F87E" box="[739,844,2012,2038]" captionStart="FIGURE 48" captionStartId="53.[151,250,1675,1699]" captionTargetBox="[151,1435,181,1651]" captionTargetId="figure-17@53.[151,1435,181,1651]" captionTargetPageId="53" captionText="FIGURE 48. Males of Marma rosea (Mello-Leit„o, 1941). A–D types of Paralophostica centralis [A habitus (Lectotype ♁, MZUSP E.740 C.1334), dorsal; B ventral; C left palp (paralectotype ♁, MZUSP E.740 C.1335), ventral; D retrolateral]; E–G specimen from São Geraldo do Araguaia/Pará/Brasil, MPEG 34360 (E habitus, dorsal; F ventral; G left palp, ventral). White arrows in F show black tarsi on legs I and II." figureDoi="http://doi.org/10.5281/zenodo.4456957" httpUri="https://zenodo.org/record/4456957/files/figure.png" pageId="57" pageNumber="344">Figs 48C</figureCitation>
,
<figureCitation id="13D29282FFA7FFB95044F8380FB1F87E" box="[856,939,2012,2039]" captionStart="FIGURE 50" captionStartId="55.[151,250,1179,1203]" captionTargetBox="[153,1409,200,1135]" captionTargetId="figure-17@55.[151,1435,181,1155]" captionTargetPageId="55" captionText="FIGURE 50. Marma rosea (Mello-Leit„o, 1941). A–E male paralectotype of Paralophostica centralis, E.740 C.1335 (A left male palp, ventral; B retrolateral; C proventral; D dorsal; E cleared bulb, ventral). F–G epigyne/vulva, holotype of Marma rosea, MLP 14965 (F ventral, G cleared, ventral).Abbreviations: BG—Bennett’s gland; cc—cymbial conductor; cd—copulatory duct; co—copulatory opening; dh—distal hematodocha; e—embolus shaft; ed—embolic disc; fd—fertilization duct; PED—process on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456961" httpUri="https://zenodo.org/record/4456961/files/figure.png" pageId="57" pageNumber="344">50A, C</figureCitation>
,
<figureCitation id="13D29282FFA7FFB950AAF8380FF0F87F" box="[950,1002,2012,2039]" captionStart="FIGURE 57" captionStartId="62.[151,250,1843,1867]" captionTargetBox="[159,1422,193,1813]" captionTargetId="figure-24@62.[151,1435,181,1819]" captionTargetPageId="62" captionText="FIGURE 57. Left male palp, ventral [A Marma baeri Simon, 1902; B M. linae sp. nov.; C M. pipa sp. nov.; D M. wesolowskae sp. nov.; E M. nigritarsis (Simon, 1900); F M. sinuosa sp. nov., G M. argentina (Mello-Leit„o, 1941), H M. rosea (Mello- Leit„o, 1941), I M. spelunca sp. nov.]. Colors represent homologous structures." figureDoi="http://doi.org/10.5281/zenodo.4456979" httpUri="https://zenodo.org/record/4456979/files/figure.png" pageId="57" pageNumber="344">57H</figureCitation>
). Females of
<taxonomicName id="4CE9F584FFA7FFB9579CF8390913F87E" authority="Salgado &amp; Ruiz, 2020" authorityName="Salgado &amp; Ruiz" authorityYear="2020" box="[1152,1289,2012,2038]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="344" phylum="Arthropoda" rank="species" species="spelunca" status="sp. nov.">
<emphasis id="B99D5215FFA7FFB9579CF8390913F87E" box="[1152,1289,2012,2038]" italics="true" pageId="57" pageNumber="344">M. spelunca</emphasis>
</taxonomicName>
<emphasis id="B99D5215FFA7FFB95613F8380972F87E" bold="true" box="[1295,1384,2012,2038]" pageId="57" pageNumber="344">
<taxonomicNameLabel id="A2AEEF6EFFA7FFB95613F8380972F87E" box="[1295,1384,2012,2038]" pageId="57" pageNumber="344" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
also differ from those of
<taxonomicName id="4CE9F584FFA4FFBA5266FF730DC6FF38" box="[378,476,151,176]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="345" phylum="Arthropoda" rank="species" species="rosea">
<emphasis id="B99D5215FFA4FFBA5266FF730DC6FF38" box="[378,476,151,176]" italics="true" pageId="58" pageNumber="345">M. rosea</emphasis>
</taxonomicName>
by having larger copulatory openings (
<figureCitation id="13D29282FFA4FFBA5088FF730FF0FF39" box="[916,1002,150,177]" captionStart="FIGURE 58" captionStartId="63.[151,250,1595,1619]" captionTargetBox="[156,1429,271,1549]" captionTargetId="figure-41@63.[151,1435,237,1571]" captionTargetPageId="63" captionText="FIGURE 58. Epigyne/vulva, A–C, E–K ventral, D dorsal [A Marma baeri Simon, 1902; B M. linae sp. nov.; C M. abaira sp. nov.; D M. femella (Caporiacco, 1955); E M. pipa sp. nov.; F M. wesolowskae sp. nov.; G M. nigritarsis (Simon, 1900); H M. sinuosa sp. nov.; I M. argentina (Mello-Leitão, 1941); J M. rosea (Mello-Leitão, 1941); K M. spelunca sp. nov.]. Colors represent homologous structures; arrow in D shows hypothetical position of secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456981" httpUri="https://zenodo.org/record/4456981/files/figure.png" pageId="58" pageNumber="345">Figs 58</figureCitation>
J–K). Further, the body of
<taxonomicName id="4CE9F584FFA4FFBA560FFF730987FF39" authority="Salgado &amp; Ruiz, 2020" authorityName="Salgado &amp; Ruiz" authorityYear="2020" box="[1299,1437,151,177]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="345" phylum="Arthropoda" rank="species" species="spelunca" status="sp. nov.">
<emphasis id="B99D5215FFA4FFBA560FFF730987FF39" box="[1299,1437,151,177]" italics="true" pageId="58" pageNumber="345">M. spelunca</emphasis>
</taxonomicName>
<emphasis id="B99D5215FFA4FFBA538BFF5F0CE8FF5D" bold="true" box="[151,242,187,213]" pageId="58" pageNumber="345">
<taxonomicNameLabel id="A2AEEF6EFFA4FFBA538BFF5F0CE8FF5D" box="[151,242,187,213]" pageId="58" pageNumber="345" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is larger (on average
<quantity id="4C1123E2FFA4FFBA52F7FF5E0E4AFF5D" box="[491,592,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="58" pageNumber="345" unit="mm" value="4.5">4.50 mm</quantity>
in total length, while specimens of
<taxonomicName id="4CE9F584FFA4FFBA50F4FF5F0851FF5C" box="[1000,1099,187,212]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="345" phylum="Arthropoda" rank="species" species="rosea">
<emphasis id="B99D5215FFA4FFBA50F4FF5F0851FF5C" box="[1000,1099,187,212]" italics="true" pageId="58" pageNumber="345">M. rosea</emphasis>
</taxonomicName>
are about 3.0 mm), being the largest species of
<taxonomicName id="4CE9F584FFA4FFBA5242FF3B0DAAFF70" authorityName="Simon" authorityYear="1902" box="[350,432,223,248]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="345" phylum="Arthropoda" rank="genus">
<emphasis id="B99D5215FFA4FFBA5242FF3B0DAAFF70" box="[350,432,223,248]" italics="true" pageId="58" pageNumber="345">Marma</emphasis>
</taxonomicName>
currently known.
</paragraph>
</subSubSection>
<subSubSection id="C3F3DD8CFFA4FFBC53DBFEE60EA0F8BF" lastPageId="60" lastPageNumber="347" pageId="58" pageNumber="345" type="description">
<paragraph id="8B568E07FFA4FFBA53DBFEE60943FEED" blockId="58.[151,1437,150,357]" pageId="58" pageNumber="345">
<emphasis id="B99D5215FFA4FFBA53DBFEE60EC7FE94" bold="true" box="[199,733,258,285]" pageId="58" pageNumber="345">
Description.
<emphasis id="B99D5215FFA4FFBA527DFEE60D86FE95" bold="true" box="[353,412,258,285]" italics="true" pageId="58" pageNumber="345">Male</emphasis>
(
<typeStatus id="545230A5FFA4FFBA52B0FEE60E06FE94" box="[428,540,258,284]" pageId="58" pageNumber="345" type="holotype">Holotype</typeStatus>
, MPEG 34347).
</emphasis>
Total length: 4.44. Carapace 2.51 long, 1.72 wide, 1.28 high. Ocular quadrangle 1.19 long. Anterior eye row 1.66 wide, posterior 1.29 wide. Legs 4312. Length of legs: I 5.44 (1.78 + 2.09 + 1.57); II 4.86 (1.56 + 1.79 + 1.51); III 5.69 (1.82 + 1.95 + 1.92); IV 5.77 (1.75 + 1.89 + 2.13).
</paragraph>
<caption id="DF96DE8FFFA4FFBA538BF8820F3DF84E" ID-DOI="http://doi.org/10.5281/zenodo.4456969" ID-Zenodo-Dep="4456969" httpUri="https://zenodo.org/record/4456969/files/figure.png" pageId="58" pageNumber="345" startId="58.[151,250,1894,1918]" targetBox="[151,1435,388,1871]" targetPageId="58">
<paragraph id="8B568E07FFA4FFBA538BF8820F3DF84E" blockId="58.[151,1437,1894,1990]" pageId="58" pageNumber="345">
<emphasis id="B99D5215FFA4FFBA538BF8820D38F8F6" bold="true" box="[151,290,1894,1918]" pageId="58" pageNumber="345">FIGURE 53.</emphasis>
Male holotype of
<taxonomicName id="4CE9F584FFA4FFBA52F8F8830E8AF8F6" authority="Salgado &amp; Ruiz, 2020" authorityName="Salgado &amp; Ruiz" authorityYear="2020" box="[484,656,1894,1918]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="345" phylum="Arthropoda" rank="species" species="spelunca" status="sp. nov.">
<emphasis id="B99D5215FFA4FFBA52F8F8830E8AF8F6" box="[484,656,1894,1918]" italics="true" pageId="58" pageNumber="345">Marma spelunca</emphasis>
</taxonomicName>
<emphasis id="B99D5215FFA4FFBA5185F8830EF1F8F7" bold="true" box="[665,747,1895,1919]" pageId="58" pageNumber="345">
<taxonomicNameLabel id="A2AEEF6EFFA4FFBA5185F8830EF1F8F7" box="[665,747,1895,1919]" pageId="58" pageNumber="345" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(MPEG 34347). A–B habitus (A dorsal, B ventral); C–D left palp (ventral; D retrolateral); D–E type locality, Nova Kararaô cave opening (E view of the entrance; F live specimen, indicated by white arrow, walking on moss present on walls around opening).
</paragraph>
</caption>
<paragraph id="8B568E07FFA5FFBB53DBFF730895FF71" blockId="59.[151,1437,151,717]" pageId="59" pageNumber="346">Leg macrosetae: Femur I d1-1-1, p1di, r0; II–IV d1-1-1, p1di, r1di. Patella I–II 0; III–IV p0, r1. Tibia I–II p0- 1-0, r0, v1r-2-2; III p1-1-0, r1-1-0, v1p-0-2; IV p0-1-1-0, r0-1-1-0, v1r-0-0-2. Metatarsus I p1-1, r0, v2-2; II p1-1, r1di, v2-2; III d1p-0-0, p1-0-2, r1-0-2, v2-0-2; IV d0 (or d1p-0-0), p1-1-2, r1-1-2, v1p-0-2.</paragraph>
<paragraph id="8B568E07FFA5FFBB53DBFEE70E7DFE7D" blockId="59.[151,1437,151,717]" pageId="59" pageNumber="346">
Color in alcohol (
<figureCitation id="13D29282FFA5FFBB5297FEE70DFAFE95" box="[395,480,258,285]" captionStart="FIGURE 53" captionStartId="58.[151,250,1894,1918]" captionTargetBox="[151,1435,388,1871]" captionTargetId="figure-131@58.[151,1435,381,1871]" captionTargetPageId="58" captionText="FIGURE 53. Male holotype of Marma spelunca sp. nov. (MPEG 34347). A–B habitus (A dorsal, B ventral); C–D left palp (ventral; D retrolateral); D–E type locality, Nova Kararaô cave opening (E view of the entrance; F live specimen, indicated by white arrow, walking on moss present on walls around opening)." figureDoi="http://doi.org/10.5281/zenodo.4456969" httpUri="https://zenodo.org/record/4456969/files/figure.png" pageId="59" pageNumber="346">Figs 53</figureCitation>
A–B): thoracic area with rhombus of setae; abdomen ventrally with three longitudinal dark brown stripes delimited by stripes of pale circles; lateral borders with pale circles; legs: femur I almost black (ventrally not totally colored), II with dark distal ring and proximal region with prolateral and retrolateral dark spot, III with proximal and distal dark ring, IV with dark distal ring and proximal region with dark proventral spot; tibia I with dark proximal ring and distal region with dark retrolateral spot, II with dark proximal ring and distal region with prolateral and retrolateral dark spot, III–IV with proximal and distal dark ring; tarsus I almost black, II with black tip, III–IV with dark proximal ring.
</paragraph>
<paragraph id="8B568E07FFA5FFBB53DBFE1B089BFDE9" blockId="59.[151,1437,151,717]" pageId="59" pageNumber="346">
Palp: RTA finger-shaped (
<figureCitation id="13D29282FFA5FFBB52F0FE1B0E43FD91" box="[492,601,510,537]" captionStart="FIGURE 53" captionStartId="58.[151,250,1894,1918]" captionTargetBox="[151,1435,388,1871]" captionTargetId="figure-131@58.[151,1435,381,1871]" captionTargetPageId="58" captionText="FIGURE 53. Male holotype of Marma spelunca sp. nov. (MPEG 34347). A–B habitus (A dorsal, B ventral); C–D left palp (ventral; D retrolateral); D–E type locality, Nova Kararaô cave opening (E view of the entrance; F live specimen, indicated by white arrow, walking on moss present on walls around opening)." figureDoi="http://doi.org/10.5281/zenodo.4456969" httpUri="https://zenodo.org/record/4456969/files/figure.png" pageId="59" pageNumber="346">Figs 53D</figureCitation>
,
<figureCitation id="13D29282FFA5FFBB5179FE1A0E82FD91" box="[613,664,510,537]" captionStart="FIGURE 55" captionStartId="60.[151,250,1210,1234]" captionTargetBox="[157,1424,191,1176]" captionTargetId="figure-24@60.[151,1435,181,1186]" captionTargetPageId="60" captionText="FIGURE 55. Marma spelunca sp. nov. A–E left male palp (A ventral; B retrolateral; C proventral; D dorsal; E cleared bulb, ventral); F–G epigyne/vulva (F ventral, G cleared, ventral).Abbreviations: BG—Bennett’s gland; cc—cymbial conductor; cdcopulatory duct; co—copulatory opening; dh—distal hematodocha; e—embolus shaft; ed—embolic disc; fd—fertilization duct; PED—process on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456973" httpUri="https://zenodo.org/record/4456973/files/figure.png" pageId="59" pageNumber="346">55B</figureCitation>
); embolic disc with curved and symmetrical edges (
<figureCitation id="13D29282FFA5FFBB57F8FE1B0955FD91" box="[1252,1359,510,537]" captionStart="FIGURE 53" captionStartId="58.[151,250,1894,1918]" captionTargetBox="[151,1435,388,1871]" captionTargetId="figure-131@58.[151,1435,381,1871]" captionTargetPageId="58" captionText="FIGURE 53. Male holotype of Marma spelunca sp. nov. (MPEG 34347). A–B habitus (A dorsal, B ventral); C–D left palp (ventral; D retrolateral); D–E type locality, Nova Kararaô cave opening (E view of the entrance; F live specimen, indicated by white arrow, walking on moss present on walls around opening)." figureDoi="http://doi.org/10.5281/zenodo.4456969" httpUri="https://zenodo.org/record/4456969/files/figure.png" pageId="59" pageNumber="346">Figs 53C</figureCitation>
,
<figureCitation id="13D29282FFA5FFBB5647FE1A098AFD91" box="[1371,1424,510,537]" captionStart="FIGURE 55" captionStartId="60.[151,250,1210,1234]" captionTargetBox="[157,1424,191,1176]" captionTargetId="figure-24@60.[151,1435,181,1186]" captionTargetPageId="60" captionText="FIGURE 55. Marma spelunca sp. nov. A–E left male palp (A ventral; B retrolateral; C proventral; D dorsal; E cleared bulb, ventral); F–G epigyne/vulva (F ventral, G cleared, ventral).Abbreviations: BG—Bennett’s gland; cc—cymbial conductor; cdcopulatory duct; co—copulatory opening; dh—distal hematodocha; e—embolus shaft; ed—embolic disc; fd—fertilization duct; PED—process on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456973" httpUri="https://zenodo.org/record/4456973/files/figure.png" pageId="59" pageNumber="346">55A</figureCitation>
); PED very long (about three times the length of the exposed portion of embolic disc), emerging from middle-distal part of embolic disc (
<figureCitation id="13D29282FFA5FFBB5299FDA30E0AFDE9" box="[389,528,582,609]" captionStart="FIGURE 55" captionStartId="60.[151,250,1210,1234]" captionTargetBox="[157,1424,191,1176]" captionTargetId="figure-24@60.[151,1435,181,1186]" captionTargetPageId="60" captionText="FIGURE 55. Marma spelunca sp. nov. A–E left male palp (A ventral; B retrolateral; C proventral; D dorsal; E cleared bulb, ventral); F–G epigyne/vulva (F ventral, G cleared, ventral).Abbreviations: BG—Bennett’s gland; cc—cymbial conductor; cdcopulatory duct; co—copulatory opening; dh—distal hematodocha; e—embolus shaft; ed—embolic disc; fd—fertilization duct; PED—process on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456973" httpUri="https://zenodo.org/record/4456973/files/figure.png" pageId="59" pageNumber="346">Figs 55A, E</figureCitation>
); embolus tip ends beyond tip of PED (
<figureCitation id="13D29282FFA5FFBB50D6FDA3082EFDE9" box="[970,1076,582,609]" captionStart="FIGURE 53" captionStartId="58.[151,250,1894,1918]" captionTargetBox="[151,1435,388,1871]" captionTargetId="figure-131@58.[151,1435,381,1871]" captionTargetPageId="58" captionText="FIGURE 53. Male holotype of Marma spelunca sp. nov. (MPEG 34347). A–B habitus (A dorsal, B ventral); C–D left palp (ventral; D retrolateral); D–E type locality, Nova Kararaô cave opening (E view of the entrance; F live specimen, indicated by white arrow, walking on moss present on walls around opening)." figureDoi="http://doi.org/10.5281/zenodo.4456969" httpUri="https://zenodo.org/record/4456969/files/figure.png" pageId="59" pageNumber="346">Figs 53C</figureCitation>
,
<figureCitation id="13D29282FFA5FFBB575CFDA2086EFDE9" box="[1088,1140,582,609]" captionStart="FIGURE 55" captionStartId="60.[151,250,1210,1234]" captionTargetBox="[157,1424,191,1176]" captionTargetId="figure-24@60.[151,1435,181,1186]" captionTargetPageId="60" captionText="FIGURE 55. Marma spelunca sp. nov. A–E left male palp (A ventral; B retrolateral; C proventral; D dorsal; E cleared bulb, ventral); F–G epigyne/vulva (F ventral, G cleared, ventral).Abbreviations: BG—Bennett’s gland; cc—cymbial conductor; cdcopulatory duct; co—copulatory opening; dh—distal hematodocha; e—embolus shaft; ed—embolic disc; fd—fertilization duct; PED—process on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456973" httpUri="https://zenodo.org/record/4456973/files/figure.png" pageId="59" pageNumber="346">55A</figureCitation>
).
</paragraph>
<paragraph id="8B568E07FFA5FFBB53DBFD8E08D2FD45" blockId="59.[151,1437,151,717]" pageId="59" pageNumber="346">
<emphasis id="B99D5215FFA5FFBB53DBFD8E0E43FD0C" bold="true" box="[199,601,618,645]" pageId="59" pageNumber="346">
<emphasis id="B99D5215FFA5FFBB53DBFD8E0D04FD0D" bold="true" box="[199,286,618,645]" italics="true" pageId="59" pageNumber="346">Female</emphasis>
(
<typeStatus id="545230A5FFA5FFBB5230FD8E0D86FD0C" box="[300,412,618,644]" pageId="59" pageNumber="346" type="paratype">Paratype</typeStatus>
, MPEG 34348).
</emphasis>
Total length: 4.89. Carapace 2.36 long, 1.67 wide, 1.27 high. Ocular quadrangle 1.20 long. Anterior eye row 1.56 wide, posterior 1.33 wide. Legs 3=412. Length of legs: I 4.20 (1.32 + 1.61 + 1.27); II 4.15 (1.41 + 1.52 + 1.22); III 5.55 (1.82 + 1.88 + 1.85); IV 5.55 (1.71 + 1.84 + 2.00).
</paragraph>
<caption id="DF96DE8FFFA5FFBB538BF86508DAF869" ID-DOI="http://doi.org/10.5281/zenodo.4456971" ID-Zenodo-Dep="4456971" httpUri="https://zenodo.org/record/4456971/files/figure.png" pageId="59" pageNumber="346" startId="59.[151,250,1921,1945]" targetBox="[156,1424,749,1894]" targetPageId="59">
<paragraph id="8B568E07FFA5FFBB538BF86508DAF869" blockId="59.[151,1436,1921,2017]" pageId="59" pageNumber="346">
<emphasis id="B99D5215FFA5FFBB538BF8650D3BF811" bold="true" box="[151,289,1921,1945]" pageId="59" pageNumber="346">FIGURE 54.</emphasis>
Female paratype of
<taxonomicName id="4CE9F584FFA5FFBB52EBF8660EB9F811" authority="Salgado &amp; Ruiz, 2020" authorityName="Salgado &amp; Ruiz" authorityYear="2020" box="[503,675,1921,1945]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="59" pageNumber="346" phylum="Arthropoda" rank="species" species="spelunca" status="sp. nov.">
<emphasis id="B99D5215FFA5FFBB52EBF8660EB9F811" box="[503,675,1921,1945]" italics="true" pageId="59" pageNumber="346">Marma spelunca</emphasis>
</taxonomicName>
<emphasis id="B99D5215FFA5FFBB51B7F8650EE7F811" bold="true" box="[683,765,1921,1945]" pageId="59" pageNumber="346">
<taxonomicNameLabel id="A2AEEF6EFFA5FFBB51B7F8650EE7F811" box="[683,765,1921,1945]" pageId="59" pageNumber="346" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(MPEG 34348). A–B habitus (A dorsal, B ventral); C–D cleared epigyne/vulva (C ventral, D detail of right spermatheca, dorsal). Abbreviations: BG—Bennett’s gland; cd—copulatory duct; co—copulatory opening; fd—fertilization duct; PS—Primary spermatheca; SS—secondary spermatheca.
</paragraph>
</caption>
<caption id="DF96DE8FFFA2FFBC538BFB5E0996FAB6" ID-DOI="http://doi.org/10.5281/zenodo.4456973" ID-Zenodo-Dep="4456973" httpUri="https://zenodo.org/record/4456973/files/figure.png" pageId="60" pageNumber="347" startId="60.[151,250,1210,1234]" targetBox="[157,1424,191,1176]" targetPageId="60">
<paragraph id="8B568E07FFA2FFBC538BFB5E0996FAB6" blockId="60.[151,1436,1210,1342]" pageId="60" pageNumber="347">
<emphasis id="B99D5215FFA2FFBC538BFB5E0D3BFB5B" bold="true" box="[151,289,1210,1235]" pageId="60" pageNumber="347">FIGURE 55.</emphasis>
<taxonomicName id="4CE9F584FFA2FFBC5234FB5F0DCEFB5A" authority="Salgado &amp; Ruiz, 2020" authorityName="Salgado &amp; Ruiz" authorityYear="2020" box="[296,468,1210,1234]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="60" pageNumber="347" phylum="Arthropoda" rank="species" species="spelunca" status="sp. nov.">
<emphasis id="B99D5215FFA2FFBC5234FB5F0DCEFB5A" box="[296,468,1210,1234]" italics="true" pageId="60" pageNumber="347">Marma spelunca</emphasis>
</taxonomicName>
<emphasis id="B99D5215FFA2FFBC52C0FB5F0E34FB5B" bold="true" box="[476,558,1211,1235]" pageId="60" pageNumber="347">
<taxonomicNameLabel id="A2AEEF6EFFA2FFBC52C0FB5F0E34FB5B" box="[476,558,1211,1235]" pageId="60" pageNumber="347" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A–E left male palp (A ventral; B retrolateral; C proventral; D dorsal; E cleared bulb, ventral); F–G epigyne/vulva (F ventral, G cleared, ventral). Abbreviations: BG—Bennett’s gland; cc—cymbial conductor; cdcopulatory duct; co—copulatory opening; dh—distal hematodocha; e—embolus shaft; ed—embolic disc; fd—fertilization duct; PED—process on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermatheca.
</paragraph>
</caption>
<paragraph id="8B568E07FFA2FFBC53DBFA890D51FA7B" blockId="60.[151,1437,1389,2029]" pageId="60" pageNumber="347">Leg macrosetae: Femur I d1-1-1, p1di, r0; II d1-1-1, p1di, r1di; III d1-1-1, p1di, r0; IV d1-1-1, p1di, r1di. Patella I–II 0; III–IV p0, r1. Tibia I–II p0-1-0, r0, v1r-2-2; III p1-1-0, r1-1-0, v1p-0-2; IV p1-1-1-0 (or p0-1-1-0), r1-1-1-0, v0-1p-0-2. Metatarsus I p1-1, r0, v2-2; II p1-1, r1di, v2-2; III d1p-0-0, p1-0-2, r1-0-2, v2-0-2; IV p1-1-2, r1-1-2, v0-1p-2.</paragraph>
<paragraph id="8B568E07FFA2FFBC53DBFA1908FFF92F" blockId="60.[151,1437,1389,2029]" pageId="60" pageNumber="347">Color in alcohol (Figs: 54A–B): carapace and abdomen as in male; legs: femur I with dark distal ring and with prolateral and retrolateral dark stripe, II with dark distal ring, with dark retrolateral stripe and proximal region with dark proventral spot, III with proximal and distal dark ring, IV with dark distal ring and proximal region with dark proventral spot; tibia I–II with dark proximal ring and distal region with prolateral and retrolateral dark spot, III–IV with proximal and distal dark ring; tarsus I–II without dark marks, III–IV with dark proximal ring.</paragraph>
<paragraph id="8B568E07FFA2FFBC53DBF9550EA0F8BF" blockId="60.[151,1437,1389,2029]" pageId="60" pageNumber="347">
Epigyne (
<figureCitation id="13D29282FFA2FFBC5228F9550D93F943" box="[308,393,1712,1739]" captionStart="FIGURE 54" captionStartId="59.[151,250,1921,1945]" captionTargetBox="[156,1424,749,1894]" captionTargetId="figure-300@59.[151,1435,741,1897]" captionTargetPageId="59" captionText="FIGURE 54. Female paratype of Marma spelunca sp. nov. (MPEG 34348). A–B habitus (A dorsal, B ventral); C–D cleared epigyne/vulva (C ventral, D detail of right spermatheca, dorsal). Abbreviations: BG—Bennett’s gland; cd—copulatory duct; co—copulatory opening; fd—fertilization duct; PS—Primary spermatheca; SS—secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456971" httpUri="https://zenodo.org/record/4456971/files/figure.png" pageId="60" pageNumber="347">Figs 54</figureCitation>
C–D, 55F–G): copulatory openings with large diameter (similar to diameter of primary spermathecae), very close to each other, placed more anteriorly than primary spermathecae; copulatory ducts long, encircling primary spermathecae; proximal copulatory duct longer than distal section; primary spermathecae slightly arched; secondary spermathecae well developed.
</paragraph>
</subSubSection>
<subSubSection id="C3F3DD8CFFA2FFBC53DBF8A50D6CF864" pageId="60" pageNumber="347" type="materials_examined">
<paragraph id="8B568E07FFA2FFBC53DBF8A50FE5F8F7" blockId="60.[151,1437,1389,2029]" pageId="60" pageNumber="347">
<emphasis id="B99D5215FFA2FFBC53DBF8A50D64F8D2" bold="true" box="[199,382,1856,1883]" pageId="60" pageNumber="347">Type material.</emphasis>
<materialsCitation id="3B81845AFFA2FFBC5291F8A50FE5F8F7" ID-GBIF-Occurrence="3020125451" collectingDate="2016-08-05" collectionCode="MPEG" collectorName="G. R. S. Ruiz &amp; R. Valente" country="Brazil" latitude="-3.1387308" location="Nova Kararao cave" longLatPrecision="1" longitude="-51.820145" municipality="Vitoria do Xingu" pageId="60" pageNumber="347" specimenCode="MPEG 34347" specimenCount="1" stateProvince="Para" typeStatus="holotype">
<typeStatus id="545230A5FFA2FFBC5291F8A50DEFF8D3" box="[397,501,1857,1883]" pageId="60" pageNumber="347" type="holotype">Holotype</typeStatus>
♁:
<emphasis id="B99D5215FFA2FFBC512EF8A40EBCF8D2" bold="true" box="[562,678,1856,1882]" pageId="60" pageNumber="347">
<collectingCountry id="F3FECE97FFA2FFBC512EF8A40EB8F8D2" box="[562,674,1856,1882]" name="Brazil" pageId="60" pageNumber="347">BRAZIL</collectingCountry>
:
</emphasis>
<emphasis id="B99D5215FFA2FFBC51AAF8A50EF4F8D3" box="[694,750,1857,1883]" italics="true" pageId="60" pageNumber="347">
<collectingRegion id="492D40E5FFA2FFBC51AAF8A50EF4F8D3" box="[694,750,1857,1883]" country="Brazil" name="Para" pageId="60" pageNumber="347">Pará</collectingRegion>
</emphasis>
:
<collectingMunicipality id="6B32147DFFA2FFBC5019F8A50FCDF8D3" box="[773,983,1857,1883]" pageId="60" pageNumber="347">Vitória do Xingú</collectingMunicipality>
,
<location id="8E36D8DCFFA2FFBC50F7F8A508CEF8D3" LSID="urn:lsid:plazi:treatment:03403F11FFA7FFBF538BF990092BFF5D:8E36D8DCFFA2FFBC50F7F8A508CEF8D3" box="[1003,1236,1857,1883]" country="Brazil" latitude="-3.1387308" longLatPrecision="1" longitude="-51.820145" municipality="Vitoria do Xingu" name="Nova Kararao cave" pageId="60" pageNumber="347" stateProvince="Para">Nova Kararaô cave</location>
[
<geoCoordinate id="EEDDE8C0FFA2FFBC57F7F8A5098DF8D3" box="[1259,1431,1857,1883]" degrees="03" direction="south" minutes="08" orientation="latitude" pageId="60" pageNumber="347" precision="1" seconds="19.43" value="-3.1387308">03°08’19.43”S</geoCoordinate>
,
<geoCoordinate id="EEDDE8C0FFA2FFBC538BF8800D55F8F7" box="[151,335,1892,1919]" degrees="51" direction="west" minutes="49" orientation="longitude" pageId="60" pageNumber="347" precision="1" seconds="12.53" value="-51.820145">51°49’12.53”W</geoCoordinate>
], leg.
<collectorName id="261CEBD1FFA2FFBC528DF8810E03F8F7" box="[401,537,1893,1919]" pageId="60" pageNumber="347">G.R.S. Ruiz</collectorName>
&amp;
<collectorName id="261CEBD1FFA2FFBC5120F8810EA9F8F7" box="[572,691,1893,1919]" pageId="60" pageNumber="347">R. Valente</collectorName>
,
<date id="FF57A8C7FFA2FFBC51A2F8800F59F8F7" box="[702,835,1892,1919]" pageId="60" pageNumber="347" value="2016-08-05">
<collectingDate id="EF13512FFFA2FFBC51A2F8800F59F8F7" box="[702,835,1892,1919]" pageId="60" pageNumber="347" value="2016-08-05">5.VIII.2016</collectingDate>
</date>
(
<specimenCode id="DB4F267CFFA2FFBC504DF8810FE9F8F7" box="[849,1011,1893,1919]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="60" pageNumber="347">MPEG 34347</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8B568E07FFA2FFBC53DBF86D0F3AF82B" blockId="60.[151,1437,1389,2029]" box="[199,800,1929,1955]" pageId="60" pageNumber="347">
<materialsCitation id="3B81845AFFA2FFBC53DBF86D0F06F82B" ID-GBIF-Occurrence="3020125331" box="[199,796,1929,1955]" collectingDate="2016-08-05" collectionCode="MPEG" collectorName="G. R. S. Ruiz &amp; R. Valente" country="Brazil" latitude="-3.1387308" location="Nova Kararao cave" longLatPrecision="1" longitude="-51.820145" municipality="Vitoria do Xingu" pageId="60" pageNumber="347" specimenCode="MPEG 34348" specimenCount="1" specimenCount-female="1" stateProvince="Para" typeStatus="paratype">
<typeStatus id="545230A5FFA2FFBC53DBF86D0D37F82B" box="[199,301,1929,1955]" pageId="60" pageNumber="347" type="paratype">Paratype</typeStatus>
: Same data as holotype,
<specimenCount id="9DEF458EFFA2FFBC515DF86D0E7EF82B" box="[577,612,1929,1955]" pageId="60" pageNumber="347" type="female">1♀</specimenCount>
(
<specimenCode id="DB4F267CFFA2FFBC516FF86D0F0EF82B" box="[627,788,1929,1955]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="60" pageNumber="347">MPEG 34348</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B568E07FFA2FFBC53DBF8480D6CF864" blockId="60.[151,1437,1389,2029]" pageId="60" pageNumber="347">
<materialsCitation id="3B81845AFFA2FFBC53DBF8480FCDF84F" ID-GBIF-Occurrence="3020125393" box="[199,983,1964,1993]" collectingDate="2016-08-05" collectionCode="MPEG" collectorName="G. R. S. Ruiz &amp; R. Valente" country="Brazil" latitude="-3.1387308" location="Nova Kararao cave" longLatPrecision="1" longitude="-51.820145" municipality="Vitoria do Xingu" pageId="60" pageNumber="347" specimenCode="MPEG 37193" specimenCount="1" stateProvince="Para">
<emphasis id="B99D5215FFA2FFBC53DBF8480DE2F84E" bold="true" box="[199,504,1964,1990]" pageId="60" pageNumber="347">Other material examined.</emphasis>
Same data as holotype, 3♁ (
<specimenCode id="DB4F267CFFA2FFBC502CF8490FD5F84F" box="[816,975,1965,1991]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="60" pageNumber="347">MPEG 37193</specimenCode>
)
</materialsCitation>
,
<materialsCitation id="3B81845AFFA2FFBC50FDF84A08A2F84F" ID-GBIF-Occurrence="3020125406" box="[993,1208,1965,1993]" collectionCode="MPEG" country="Brazil" pageId="60" pageNumber="347" specimenCode="MPEG 37194" specimenCount="1" stateProvince="Para">
4♁ (
<specimenCode id="DB4F267CFFA2FFBC570CF84908B5F84F" box="[1040,1199,1965,1991]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="60" pageNumber="347">MPEG 37194</specimenCode>
)
</materialsCitation>
,
<materialsCitation id="3B81845AFFA2FFBC57DDF84A0982F84F" ID-GBIF-Occurrence="3020125307" box="[1217,1432,1964,1993]" collectionCode="MPEG" country="Brazil" pageId="60" pageNumber="347" specimenCode="MPEG 37195" specimenCount="1" stateProvince="Para">
4♁ (
<specimenCode id="DB4F267CFFA2FFBC57EDF849098AF84F" box="[1265,1424,1964,1991]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="60" pageNumber="347">MPEG 37195</specimenCode>
)
</materialsCitation>
,
<materialsCitation id="3B81845AFFA2FFBC538BF8360D68F864" ID-GBIF-Occurrence="3020125445" box="[151,370,2001,2029]" collectionCode="MPEG" country="Brazil" pageId="60" pageNumber="347" specimenCode="MPEG 37196" specimenCount="1" stateProvince="Para">
2♁ (
<specimenCode id="DB4F267CFFA2FFBC53D5F8350D70F864" box="[201,362,2001,2028]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="60" pageNumber="347">MPEG 37196</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF96DE8FFFA3FFBD538BF8A10949F841" ID-DOI="http://doi.org/10.5281/zenodo.4456977" ID-Zenodo-Dep="4456977" httpUri="https://zenodo.org/record/4456977/files/figure.png" pageId="61" pageNumber="348" startId="61.[151,250,1861,1885]" targetBox="[177,1409,181,1837]" targetPageId="61">
<paragraph id="8B568E07FFA3FFBD538BF8A10949F841" blockId="61.[151,1437,1861,1994]" pageId="61" pageNumber="348">
<emphasis id="B99D5215FFA3FFBD538BF8A10D3BF8D5" bold="true" box="[151,289,1861,1885]" pageId="61" pageNumber="348">FIGURE 56.</emphasis>
Male of
<taxonomicName id="4CE9F584FFA3FFBD529DF8A20E37F8D5" authority="Salgado &amp; Ruiz, 2020" authorityName="Salgado &amp; Ruiz" authorityYear="2020" box="[385,557,1861,1885]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="61" pageNumber="348" phylum="Arthropoda" rank="species" species="spelunca" status="sp. nov.">
<emphasis id="B99D5215FFA3FFBD529DF8A20E37F8D5" box="[385,557,1861,1885]" italics="true" pageId="61" pageNumber="348">Marma spelunca</emphasis>
</taxonomicName>
<emphasis id="B99D5215FFA3FFBD5129F8A20E9DF8D6" bold="true" box="[565,647,1862,1886]" pageId="61" pageNumber="348">
<taxonomicNameLabel id="A2AEEF6EFFA3FFBD5129F8A20E9DF8D6" box="[565,647,1862,1886]" pageId="61" pageNumber="348" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A–D left palp (A bulb, dorsal; B detail of fundus and subtegulum, dorsal; C dissected embolic disc and embolus, dorsal; D apical region of cymbium, ventral. Abbreviations: bh—basal hematodocha; cc—cymbial conductor; e—embolus shaft; fnd—fundus; PED—process on embolic disc; st—subtegulum. White arrows in C show spikes of embolus shaft and black arrow shows groove; white arrow in D shows depression of cymbial conductor.
</paragraph>
</caption>
<caption id="DF96DE8FFFA0FFBE538BF8D70FA4F81B" ID-DOI="http://doi.org/10.5281/zenodo.4456979" ID-Zenodo-Dep="4456979" httpUri="https://zenodo.org/record/4456979/files/figure.png" pageId="62" pageNumber="349" startId="62.[151,250,1843,1867]" targetBox="[159,1422,193,1813]" targetPageId="62">
<paragraph id="8B568E07FFA0FFBE538BF8D70FA4F81B" blockId="62.[151,1436,1843,1940]" pageId="62" pageNumber="349">
<emphasis id="B99D5215FFA0FFBE538BF8D70D05F8C4" bold="true" box="[151,287,1843,1868]" pageId="62" pageNumber="349">FIGURE 57.</emphasis>
Left male palp, ventral [A
<taxonomicName id="4CE9F584FFA0FFBE5133F8D00F25F8C4" authority="Simon, 1902" authorityName="Simon" authorityYear="1902" box="[559,831,1843,1868]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="62" pageNumber="349" phylum="Arthropoda" rank="species" species="baeri">
<emphasis id="B99D5215FFA0FFBE5133F8D00EAFF8C3" box="[559,693,1843,1867]" italics="true" pageId="62" pageNumber="349">Marma baeri</emphasis>
<bibRefCitation id="EF78F3F6FFA0FFBE51A7F8D70F25F8C4" author="Simon, E." box="[699,831,1843,1868]" pageId="62" pageNumber="349" pagination="24 - 56" refId="ref29638" refString="Simon, E. (1902) Description d'arachnides nouveaux de la famille des Salticidae (Attidae) (suite). Annales de la Societe Entomologique de Belgique, 46, 24 - 56 + 363 - 406. https: // doi. org / 10.5962 / bhl. part. 19237" type="journal article" year="1902">Simon, 1902</bibRefCitation>
</taxonomicName>
; B
<taxonomicName id="4CE9F584FFA0FFBE5043F8D00FAEF8C3" authority="Salgado &amp; Ruiz, 2020" authorityName="Salgado &amp; Ruiz" authorityYear="2020" box="[863,948,1843,1867]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="62" pageNumber="349" phylum="Arthropoda" rank="species" species="linae" status="sp. nov.">
<emphasis id="B99D5215FFA0FFBE5043F8D00FAEF8C3" box="[863,948,1843,1867]" italics="true" pageId="62" pageNumber="349">M. linae</emphasis>
</taxonomicName>
<emphasis id="B99D5215FFA0FFBE50A5F8D00813F8C4" bold="true" box="[953,1033,1844,1868]" pageId="62" pageNumber="349">
<taxonomicNameLabel id="A2AEEF6EFFA0FFBE50A5F8D00813F8C4" box="[953,1033,1844,1868]" pageId="62" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; C
<taxonomicName id="4CE9F584FFA0FFBE5731F8D00861F8C3" authority="Salgado &amp; Ruiz, 2020" authorityName="Salgado &amp; Ruiz" authorityYear="2020" box="[1069,1147,1844,1867]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="62" pageNumber="349" phylum="Arthropoda" rank="species" species="pipa" status="sp. nov.">
<emphasis id="B99D5215FFA0FFBE5731F8D00861F8C3" box="[1069,1147,1844,1867]" italics="true" pageId="62" pageNumber="349">M. pipa</emphasis>
</taxonomicName>
<emphasis id="B99D5215FFA0FFBE579DF8D008CBF8C4" bold="true" box="[1153,1233,1844,1868]" pageId="62" pageNumber="349">
<taxonomicNameLabel id="A2AEEF6EFFA0FFBE579DF8D008CBF8C4" box="[1153,1233,1844,1868]" pageId="62" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; D
<taxonomicName id="4CE9F584FFA0FFBE57EBF8D00986F8C3" authority="Salgado &amp; Ruiz, 2020" authorityName="Salgado &amp; Ruiz" authorityYear="2020" box="[1271,1436,1843,1867]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="62" pageNumber="349" phylum="Arthropoda" rank="species" species="wesolowskae" status="sp. nov.">
<emphasis id="B99D5215FFA0FFBE57EBF8D00986F8C3" box="[1271,1436,1843,1867]" italics="true" pageId="62" pageNumber="349">M. wesolowskae</emphasis>
</taxonomicName>
<emphasis id="B99D5215FFA0FFBE538BF8BC0CF0F8F8" bold="true" box="[151,234,1880,1904]" pageId="62" pageNumber="349">
<taxonomicNameLabel id="A2AEEF6EFFA0FFBE538BF8BC0CF0F8F8" box="[151,234,1880,1904]" pageId="62" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; E
<taxonomicName id="4CE9F584FFA0FFBE520EF8BC0E5AF8F8" authority="(Simon, 1900)" baseAuthorityName="Simon" baseAuthorityYear="1900" box="[274,576,1879,1904]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="62" pageNumber="349" phylum="Arthropoda" rank="species" species="nigritarsis">
<emphasis id="B99D5215FFA0FFBE520EF8BC0DBAF8E7" box="[274,416,1880,1903]" italics="true" pageId="62" pageNumber="349">M. nigritarsis</emphasis>
(
<bibRefCitation id="EF78F3F6FFA0FFBE52ACF8B30E23F8F8" author="Simon, E." box="[432,569,1879,1904]" pageId="62" pageNumber="349" pagination="27 - 61" refId="ref29572" refString="Simon, E. (1900) Etudes arachnologiques. 30 e Memoire. XLVII. Descriptions d'especes nouvelles de la famille des Attidae. Annales de la Societe Entomologique de France, 69, 27 - 61." type="journal article" year="1900">Simon, 1900</bibRefCitation>
)
</taxonomicName>
; F
<taxonomicName id="4CE9F584FFA0FFBE517FF8BC0ECEF8E7" authority="Salgado &amp; Ruiz, 2020" authorityName="Salgado &amp; Ruiz" authorityYear="2020" box="[611,724,1880,1903]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="62" pageNumber="349" phylum="Arthropoda" rank="species" species="sinuosa" status="sp. nov.">
<emphasis id="B99D5215FFA0FFBE517FF8BC0ECEF8E7" box="[611,724,1880,1903]" italics="true" pageId="62" pageNumber="349">M. sinuosa</emphasis>
</taxonomicName>
<emphasis id="B99D5215FFA0FFBE51C1F8BC0F2AF8F8" bold="true" box="[733,816,1880,1904]" pageId="62" pageNumber="349">
<taxonomicNameLabel id="A2AEEF6EFFA0FFBE51C1F8BC0F2AF8F8" box="[733,816,1880,1904]" pageId="62" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, G
<taxonomicName id="4CE9F584FFA0FFBE5047F8BC0FFBF8E7" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1941" box="[859,993,1880,1903]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="62" pageNumber="349" phylum="Arthropoda" rank="species" species="argentina">
<emphasis id="B99D5215FFA0FFBE5047F8BC0FFBF8E7" box="[859,993,1880,1903]" italics="true" pageId="62" pageNumber="349">M. argentina</emphasis>
</taxonomicName>
(Mello-Leit„o, 1941), H
<taxonomicName id="4CE9F584FFA0FFBE57F7F8BC095CF8E7" box="[1259,1350,1880,1903]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="62" pageNumber="349" phylum="Arthropoda" rank="species" species="rosea">
<emphasis id="B99D5215FFA0FFBE57F7F8BC095CF8E7" box="[1259,1350,1880,1903]" italics="true" pageId="62" pageNumber="349">M. rosea</emphasis>
</taxonomicName>
(Mello-Leit„o, 1941), I
<taxonomicName id="4CE9F584FFA0FFBE5227F8980DADF81B" authority="Salgado &amp; Ruiz, 2020" authorityName="Salgado &amp; Ruiz" authorityYear="2020" box="[315,439,1915,1939]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="62" pageNumber="349" phylum="Arthropoda" rank="species" species="spelunca" status="sp. nov.">
<emphasis id="B99D5215FFA0FFBE5227F8980DADF81B" box="[315,439,1915,1939]" italics="true" pageId="62" pageNumber="349">M. spelunca</emphasis>
</taxonomicName>
<emphasis id="B99D5215FFA0FFBE52A1F8980E14F81C" bold="true" box="[445,526,1916,1940]" pageId="62" pageNumber="349">
<taxonomicNameLabel id="A2AEEF6EFFA0FFBE52A1F8980E14F81C" box="[445,526,1916,1940]" pageId="62" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
]. Colors represent homologous structures.
</paragraph>
</caption>
<subSubSection id="C3F3DD8CFFA1FFBF53DBFF72092BFF5D" pageId="63" pageNumber="350" type="distribution">
<paragraph id="8B568E07FFA1FFBF53DBFF720F74FF39" blockId="63.[199,1329,150,213]" box="[199,878,150,177]" pageId="63" pageNumber="350">
<emphasis id="B99D5215FFA1FFBF53DBFF720D7BFF38" bold="true" box="[199,353,150,176]" pageId="63" pageNumber="350">Distribution.</emphasis>
Known only from the
<typeStatus id="545230A5FFA1FFBF517CFF730E8AFF39" box="[608,656,151,177]" pageId="63" pageNumber="350">type</typeStatus>
locality (
<figureCitation id="13D29282FFA1FFBF51E7FF730F7BFF39" box="[763,865,150,177]" captionStart="FIGURE 59" captionStartId="64.[151,250,1837,1861]" captionTargetBox="[162,1435,181,1814]" captionTargetId="figure-24@64.[151,1435,181,1814]" captionTargetPageId="64" captionText="FIGURE 59. Maps of South America showing records of Marma species (A species previously described; B new species). Record from the state of Amazonas (Brazil) in Fig. 59A is taken from Zhang &amp; Maddison (2015)." figureDoi="http://doi.org/10.5281/zenodo.4456983" httpUri="https://zenodo.org/record/4456983/files/figure.png" pageId="63" pageNumber="350">Fig. 59B</figureCitation>
).
</paragraph>
<paragraph id="8B568E07FFA1FFBF53DBFF5F092BFF5D" blockId="63.[199,1329,150,213]" box="[199,1329,186,213]" pageId="63" pageNumber="350">
<emphasis id="B99D5215FFA1FFBF53DBFF5F0D9FFF5C" bold="true" box="[199,389,186,213]" pageId="63" pageNumber="350">Natural history.</emphasis>
The specimens were collected from the walls around a cave entrance (
<figureCitation id="13D29282FFA1FFBF5781FF5F08EBFF5D" box="[1181,1265,186,213]" captionStart="FIGURE 53" captionStartId="58.[151,250,1894,1918]" captionTargetBox="[151,1435,388,1871]" captionTargetId="figure-131@58.[151,1435,381,1871]" captionTargetPageId="58" captionText="FIGURE 53. Male holotype of Marma spelunca sp. nov. (MPEG 34347). A–B habitus (A dorsal, B ventral); C–D left palp (ventral; D retrolateral); D–E type locality, Nova Kararaô cave opening (E view of the entrance; F live specimen, indicated by white arrow, walking on moss present on walls around opening)." figureDoi="http://doi.org/10.5281/zenodo.4456969" httpUri="https://zenodo.org/record/4456969/files/figure.png" pageId="63" pageNumber="350">Figs 53</figureCitation>
E–F).
</paragraph>
</subSubSection>
</treatment>
</document>
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