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<document id="9DB6F123FCE7545E00032259243CC2C4" ID-CLB-Dataset="22660" ID-DOI="10.11646/zootaxa.4419.1.1" ID-GBIF-Dataset="f0d6e3e8-7d48-4440-aad9-88eb43736798" ID-ISSN="1175-5326" ID-Zenodo-Dep="3769633" ID-ZooBank="03CAFD21-185F-4C86-ACC3-8CEB61E7F7DD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587999891170" checkinUser="plazi" docAuthor="Martino, Emanuela Di &amp; Taylor, Paul D." docDate="2018" docId="CF6D87AAE87AD27BFF7DF9330EA9F945" docLanguage="en" docName="zootaxa.4419.1.1.pdf" docOrigin="Zootaxa 4419 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Psammocleidochasma Winston &amp; Vieira 2013" docType="treatment" docVersion="8" lastPageNumber="54" masterDocId="3354FFD2E84ED24EFFEAFF8E0D46FF89" masterDocTitle="Early Pleistocene and Holocene bryozoans from Indonesia" masterLastPageNumber="70" masterPageNumber="1" pageNumber="53" updateTime="1698836999326" updateUser="ExternalLinkService">
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<mods:title id="33778CA6D9F229AD4736DAD2F7C72995">Early Pleistocene and Holocene bryozoans from Indonesia</mods:title>
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<mods:namePart id="2171EF3D5B6CE4169BD07681A15945AF">Martino, Emanuela Di</mods:namePart>
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<treatment id="CF6D87AAE87AD27BFF7DF9330EA9F945" ID-DOI="http://doi.org/10.5281/zenodo.3799606" ID-GBIF-Taxon="163752417" ID-Zenodo-Dep="3799606" LSID="urn:lsid:plazi:treatment:CF6D87AAE87AD27BFF7DF9330EA9F945" httpUri="http://treatment.plazi.org/id/CF6D87AAE87AD27BFF7DF9330EA9F945" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
<subSubSection id="0FDE6537E87AD27AFF7DF9330C88F95E" box="[151,462,1725,1751]" pageId="52" pageNumber="53" type="nomenclature">
<paragraph id="477B36BCE87AD27AFF7DF9330C88F95E" blockId="52.[151,462,1725,1785]" box="[151,462,1725,1751]" pageId="52" pageNumber="53">
<heading id="1C3381D0E87AD27AFF7DF9330C88F95E" bold="true" box="[151,462,1725,1751]" fontSize="11" level="1" pageId="52" pageNumber="53" reason="1">
<emphasis id="75B0EAAEE87AD27AFF7DF9330C88F95E" bold="true" box="[151,462,1725,1751]" pageId="52" pageNumber="53">
<taxonomicName id="80C44D3FE87AD27AFF7DF9330CE2F95E" ID-CoL="84SV9" authorityName="Winston &amp; Vieira" authorityYear="2013" box="[151,420,1725,1751]" class="Gymnolaemata" family="Phidoloporidae" genus="Psammocleidochasma" kingdom="Animalia" order="Cheilostomatida" pageId="52" pageNumber="53" phylum="Bryozoa" rank="genus">
<emphasis id="75B0EAAEE87AD27AFF7DF9330CE2F95E" bold="true" box="[151,420,1725,1751]" italics="true" pageId="52" pageNumber="53">Psammocleidochasma</emphasis>
</taxonomicName>
sp.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0FDE6537E87AD27AFF7DF96E0CF4F970" box="[151,434,1760,1785]" pageId="52" pageNumber="53" type="description">
<paragraph id="477B36BCE87AD27AFF7DF96E0CF4F970" blockId="52.[151,462,1725,1785]" box="[151,434,1760,1785]" pageId="52" pageNumber="53">
(
<figureCitation id="DFFF2A39E87AD27AFF75F96E0C7BF970" box="[159,317,1760,1785]" captionStart="FIGURES 158, 159" captionStartId="53.[151,264,1292,1315]" captionTargetBox="[151,1436,687,1283]" captionTargetId="figure@53.[151,1436,687,1283]" captionTargetPageId="53" captionText="FIGURES 158, 159. Psammocleidochasma sp., RGM.1350589, early Pleistocene, Java. 158. View of the colony encrusting a shell fragment. 159. Close-up of zooids showing oral and frontal tubercles, marginal areolar pores, cleithridiate orifice, oral spine bases usually covered by distal calcification (see arrow), and pore-chamber windows. Scale bars: Fig. 158 = 400 µm; Fig. 159 = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.3769729" httpUri="https://zenodo.org/record/3769729/files/figure.png" pageId="52" pageNumber="53">Figs 158–159</figureCitation>
;
<tableCitation id="0A460307E87AD27AFEA2F96E0CECF970" box="[328,426,1760,1785]" captionStart="TABLE 35" captionStartId="53.[151,239,1446,1469]" captionText="TABLE 35. Measurements (in µm) of Psammocleidochasma sp." pageId="52" pageNumber="53">Table 35</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0FDE6537E87AD27AFF7DF8AD0E6AF8B2" box="[151,812,1827,1852]" pageId="52" pageNumber="53" type="materials_examined">
<paragraph id="477B36BCE87AD27AFF7DF8AD0E6AF8B2" blockId="52.[151,1437,1827,2032]" box="[151,812,1827,1852]" pageId="52" pageNumber="53">
<emphasis id="75B0EAAEE87AD27AFF7DF8AD0C2EF8B5" bold="true" box="[151,360,1827,1852]" pageId="52" pageNumber="53">Figured material.</emphasis>
<collectionCode id="21D5AE79E87AD27AFE85F8AD0CF4F8B5" box="[367,434,1827,1852]" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="52" pageNumber="53" type="Museum">RGM</collectionCode>
.1350589, early Pleistocene,
<collectingRegion id="8500F85EE87AD27AFD18F8AA0E61F8B2" box="[754,807,1828,1851]" country="Indonesia" name="Jawa Timur" pageId="52" pageNumber="53">Java</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="0FDE6537E87AD27BFF2DF8C90ED8FE94" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53" type="description">
<paragraph id="477B36BCE87AD27BFF2DF8C90ED8FE94" blockId="52.[151,1437,1827,2032]" lastBlockId="53.[151,1436,151,645]" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
<emphasis id="75B0EAAEE87AD27AFF2DF8C90C1FF8E9" bold="true" box="[199,345,1863,1888]" pageId="52" pageNumber="53">Description.</emphasis>
Colony encrusting a shell fragment, multiserial, unilaminar. Autozooids distinct by shallow interzooidal furrows, hexagonal, longer than wide (mean L/W = 1.31). Pore-chamber windows visible along inner walls of broken zooids. Frontal shield convex, finely granular, nodular, imperforate except for a pair of small (about
<quantity id="803C9B59E87AD27AFF0CF83D0C76F845" box="[230,304,1971,1996]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="52" pageNumber="53" unit="mm" value="20.0">20 µm</quantity>
long), elliptical, rimmed, marginal areolar pores, leant against the zooidal margin slightly below the orifice; a few additional marginal pores may occur proximally. Orifice placed terminally, cleithridiate, anter deeply rounded separated from the slightly narrower, shallowly concave sinus by pair of small, rounded, downwardly directed condyles. An uncertain number of oral spine bases obscured by distal calcification. Peristome slightly raised, thick, made of 3–5 projecting solid tubercles. Two additional tubercles may develop proximally on the frontal shield with ontogeny. Avicularia absent. Ooecia not observed.
</paragraph>
</subSubSection>
<subSubSection id="0FDE6537E87BD27BFF2DFEA90EA9F945" pageId="53" pageNumber="54" type="discussion">
<paragraph id="477B36BCE87BD27BFF2DFEA90F43FD0C" blockId="53.[151,1436,151,645]" pageId="53" pageNumber="54">
<emphasis id="75B0EAAEE87BD27BFF2DFEA90C7DFEC9" bold="true" box="[199,315,295,320]" pageId="53" pageNumber="54">Remarks.</emphasis>
A single specimen of
<taxonomicName id="80C44D3FE87BD27BFDD2FEA90E75FEC9" authorityName="Winston &amp; Vieira" authorityYear="2013" box="[568,819,295,320]" class="Gymnolaemata" family="Phidoloporidae" genus="Psammocleidochasma" kingdom="Animalia" order="Cheilostomatida" pageId="53" pageNumber="54" phylum="Bryozoa" rank="genus">
<emphasis id="75B0EAAEE87BD27BFDD2FEA90E75FEC9" box="[568,819,295,320]" italics="true" pageId="53" pageNumber="54">Psammocleidochasma</emphasis>
</taxonomicName>
sp. was found in our samples, encrusting both sides of a small shell fragment. The genus
<taxonomicName id="80C44D3FE87BD27BFDADFEC20E04FEEC" authorityName="Winston &amp; Vieira" authorityYear="2013" box="[583,834,332,357]" class="Gymnolaemata" family="Phidoloporidae" genus="Psammocleidochasma" kingdom="Animalia" order="Cheilostomatida" pageId="53" pageNumber="54" phylum="Bryozoa" rank="genus">
<emphasis id="75B0EAAEE87BD27BFDADFEC20E04FEEC" box="[583,834,332,357]" italics="true" pageId="53" pageNumber="54">Psammocleidochasma</emphasis>
</taxonomicName>
was introduced by
<bibRefCitation id="23554B4DE87BD27BFBD7FEC2082AFEED" author="Winston, J. E. &amp; Vieira, L. M." box="[1085,1388,332,357]" pageId="53" pageNumber="54" pagination="101 - 146" refId="ref43733" refString="Winston, J. E. &amp; Vieira, L. M. (2013) Systematics of interstitial encrusting bryozoans from southeastern Brazil. Zootaxa, 3710 (2), 101 - 146. https: // doi. org / 10.11646 / zootaxa. 3710.2.1" type="journal article" year="2013">Winston &amp; Vieira (2013)</bibRefCitation>
for ‘
<taxonomicName id="80C44D3FE87BD27BFF4AFEE10C06FE01" authorityName="Harmer" authorityYear="1957" box="[160,320,367,392]" class="Gymnolaemata" family="Cleidochasmatidae" genus="Cleidochasma" kingdom="Animalia" order="Cheilostomatida" pageId="53" pageNumber="54" phylum="Bryozoa" rank="genus">
<emphasis id="75B0EAAEE87BD27BFF4AFEE10C06FE01" box="[160,320,367,392]" italics="true" pageId="53" pageNumber="54">Cleidochasma</emphasis>
</taxonomicName>
’-like species with uni- to pluriserial small-sized colonies encrusting sand grains, and was characterized by its relatively small autozooids, tatiform ancestrula, lack of avicularia, thickened calcification of the ovicells, and solid tubercles encircling the orifice. The Florida species
<taxonomicName id="80C44D3FE87BD27BFC10FE370D9EFE7C" authority="(Winston &amp; Hakansson 1986)" baseAuthorityName="Winston &amp; Hakansson" baseAuthorityYear="1986" class="Gymnolaemata" family="Phidoloporidae" genus="Psammocleidochasma" kingdom="Animalia" order="Cheilostomatida" pageId="53" pageNumber="54" phylum="Bryozoa" rank="species" species="angustum">
<emphasis id="75B0EAAEE87BD27BFC10FE3709C0FE59" box="[1018,1158,441,464]" italics="true" pageId="53" pageNumber="54">P. angustum</emphasis>
(
<bibRefCitation id="23554B4DE87BD27BFB7DFE390D96FE7C" author="Winston, J. E. &amp; Hakansson, E." pageId="53" pageNumber="54" pagination="1 - 98" refId="ref43664" refString="Winston, J. E. &amp; Hakansson, E. (1986) The interstitial fauna of the Capron Shoals, Florida. American Museum Novitates, 2865, 1 - 98." type="journal article" year="1986">Winston &amp; Håkansson 1986</bibRefCitation>
)
</taxonomicName>
differs from the Javanese species in having four tubercles arranged distolaterally and proximolaterally around the peristome, while
<taxonomicName id="80C44D3FE87BD27BFE0BFD8F0EE6FD91" authority="Winston &amp; Vieira, 2013" authorityName="Winston &amp; Vieira" authorityYear="2013" box="[481,928,511,536]" class="Gymnolaemata" family="Phidoloporidae" genus="Psammocleidochasma" kingdom="Animalia" order="Cheilostomatida" pageId="53" pageNumber="54" phylum="Bryozoa" rank="species" species="tridentatum">
<emphasis id="75B0EAAEE87BD27BFE0BFD8F0FC5FD91" box="[481,643,511,536]" italics="true" pageId="53" pageNumber="54">P. tridentatum</emphasis>
<bibRefCitation id="23554B4DE87BD27BFD66FE710EE6FD91" author="Winston, J. E. &amp; Vieira, L. M." box="[652,928,511,536]" pageId="53" pageNumber="54" pagination="101 - 146" refId="ref43733" refString="Winston, J. E. &amp; Vieira, L. M. (2013) Systematics of interstitial encrusting bryozoans from southeastern Brazil. Zootaxa, 3710 (2), 101 - 146. https: // doi. org / 10.11646 / zootaxa. 3710.2.1" type="journal article" year="2013">Winston &amp; Vieira, 2013</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="3FD3762CE87BD27BFC00FE710968FD91" box="[1002,1070,511,536]" name="Brazil" pageId="53" pageNumber="54">Brazil</collectingCountry>
differs in having three massive peristomial tubercles, one proximal and two lateral, as well as uniserial colonies with ovicellate zooids occurring precociously. Although probably new, we prefer to keep the species in open nomenclature because the single specimen available lacks ooecia.
</paragraph>
<caption id="13BB6634E87BD27BFF7DFA820C6BFAF6" ID-DOI="http://doi.org/10.5281/zenodo.3769729" ID-Zenodo-Dep="3769729" httpUri="https://zenodo.org/record/3769729/files/figure.png" pageId="53" pageNumber="54" startId="53.[151,264,1292,1315]" targetBox="[151,1436,687,1283]" targetPageId="53">
<paragraph id="477B36BCE87BD27BFF7DFA820C6BFAF6" blockId="53.[151,1436,1292,1407]" pageId="53" pageNumber="54">
<emphasis id="75B0EAAEE87BD27BFF7DFA820C28FAAB" bold="true" box="[151,366,1292,1315]" pageId="53" pageNumber="54">FIGURES 158, 159.</emphasis>
<taxonomicName id="80C44D3FE87BD27BFE9FFA830F1EFAAA" authorityName="Winston &amp; Vieira" authorityYear="2013" box="[373,600,1293,1315]" class="Gymnolaemata" family="Phidoloporidae" genus="Psammocleidochasma" kingdom="Animalia" order="Cheilostomatida" pageId="53" pageNumber="54" phylum="Bryozoa" rank="genus">
<emphasis id="75B0EAAEE87BD27BFE9FFA830F1EFAAA" box="[373,600,1293,1315]" italics="true" pageId="53" pageNumber="54">Psammocleidochasma</emphasis>
</taxonomicName>
sp., RGM.1350589, early Pleistocene, Java. 158. View of the colony encrusting a shell fragment. 159. Close-up of zooids showing oral and frontal tubercles, marginal areolar pores, cleithridiate orifice, oral spine bases usually covered by distal calcification (see arrow), and pore-chamber windows. Scale bars: Fig. 158 = 400 µm; Fig. 159 = 200 µm.
</paragraph>
</caption>
<caption id="13BB6634E87BD27BFF7DFA280E01FA34" box="[151,839,1446,1469]" pageId="53" pageNumber="54">
<paragraph id="477B36BCE87BD27BFF7DFA280E01FA34" blockId="53.[151,839,1446,1469]" box="[151,839,1446,1469]" pageId="53" pageNumber="54">
<emphasis id="75B0EAAEE87BD27BFF7DFA280C51FA34" bold="true" box="[151,279,1446,1469]" pageId="53" pageNumber="54">TABLE 35.</emphasis>
Measurements (in µm) of
<taxonomicName id="80C44D3FE87BD27BFDDEFA280E64FA34" authorityName="Winston &amp; Vieira" authorityYear="2013" box="[564,802,1446,1469]" class="Gymnolaemata" family="Phidoloporidae" genus="Psammocleidochasma" kingdom="Animalia" order="Cheilostomatida" pageId="53" pageNumber="54" phylum="Bryozoa" rank="genus">
<emphasis id="75B0EAAEE87BD27BFDDEFA280E64FA34" box="[564,802,1446,1469]" italics="true" pageId="53" pageNumber="54">Psammocleidochasma</emphasis>
</taxonomicName>
sp.
</paragraph>
</caption>
<paragraph id="477B36BCE87BD27BFE0BFA540865F91D" pageId="53" pageNumber="54">
<table id="35C4C41CE87B2DB1FF75FA570869F91D" box="[159,1327,1497,1684]" colsContinueFrom="46.[159,1326,1428,1615]" gridcols="5" gridrows="5" pageId="53" pageNumber="54" rowsContinueFrom="52.[159,1322,1281,1469]">
<tr id="F9F434FEE87B2DB1FF75FA570869FA66" box="[159,1327,1497,1519]" gridrow="0" pageId="53" pageNumber="54" rowspan-0="1">
<th id="BA255D82E87B2DB1FE0BFA570FEBFA66" box="[481,685,1497,1519]" gridcol="1" gridrow="0" pageId="53" pageNumber="54">N (colonies, zooids)</th>
<th id="BA255D82E87B2DB1FCD9FA570E2BFA66" box="[819,877,1497,1519]" gridcol="2" gridrow="0" pageId="53" pageNumber="54">Mean</th>
<th id="BA255D82E87B2DB1FBEFFA570963FA66" box="[1029,1061,1497,1519]" gridcol="3" gridrow="0" pageId="53" pageNumber="54">SD</th>
<th id="BA255D82E87B2DB1FB3CFA570869FA66" box="[1238,1327,1497,1519]" gridcol="4" gridrow="0" pageId="53" pageNumber="54">Range</th>
</tr>
<tr id="F9F434FEE87B2DB1FF75F98C0869F991" box="[159,1327,1538,1560]" gridrow="1" pageId="53" pageNumber="54">
<th id="BA255D82E87B2DB1FF75F98C0C6AF991" box="[159,300,1538,1560]" gridcol="0" gridrow="1" pageId="53" pageNumber="54">Zooid length</th>
<td id="BA255D82E87B2DB1FE0BF98C0FEBF991" box="[481,685,1538,1560]" gridcol="1" gridrow="1" pageId="53" pageNumber="54">1, 12</td>
<td id="BA255D82E87B2DB1FCD9F98C0E2BF991" box="[819,877,1538,1560]" gridcol="2" gridrow="1" pageId="53" pageNumber="54">414</td>
<td id="BA255D82E87B2DB1FBEFF98C0963F991" box="[1029,1061,1538,1560]" gridcol="3" gridrow="1" pageId="53" pageNumber="54">34</td>
<td id="BA255D82E87B2DB1FB3CF98C0869F991" box="[1238,1327,1538,1560]" gridcol="4" gridrow="1" pageId="53" pageNumber="54">351–478</td>
</tr>
<tr id="F9F434FEE87B2DB1FF75F9A20869F9CB" box="[159,1327,1580,1602]" gridrow="2" pageId="53" pageNumber="54">
<th id="BA255D82E87B2DB1FF75F9A20C6AF9CB" box="[159,300,1580,1602]" gridcol="0" gridrow="2" pageId="53" pageNumber="54">Zooid width</th>
<td id="BA255D82E87B2DB1FE0BF9A20FEBF9CB" box="[481,685,1580,1602]" gridcol="1" gridrow="2" pageId="53" pageNumber="54">1, 12</td>
<td id="BA255D82E87B2DB1FCD9F9A20E2BF9CB" box="[819,877,1580,1602]" gridcol="2" gridrow="2" pageId="53" pageNumber="54">315</td>
<td id="BA255D82E87B2DB1FBEFF9A20963F9CB" box="[1029,1061,1580,1602]" gridcol="3" gridrow="2" pageId="53" pageNumber="54">38</td>
<td id="BA255D82E87B2DB1FB3CF9A20869F9CB" box="[1238,1327,1580,1602]" gridcol="4" gridrow="2" pageId="53" pageNumber="54">237–388</td>
</tr>
<tr id="F9F434FEE87B2DB1FF75F9DB0869F9E2" box="[159,1327,1621,1643]" gridrow="3" pageId="53" pageNumber="54">
<th id="BA255D82E87B2DB1FF75F9DB0C6AF9E2" box="[159,300,1621,1643]" gridcol="0" gridrow="3" pageId="53" pageNumber="54">Orifice length</th>
<td id="BA255D82E87B2DB1FE0BF9DB0FEBF9E2" box="[481,685,1621,1643]" gridcol="1" gridrow="3" pageId="53" pageNumber="54">1, 6</td>
<td id="BA255D82E87B2DB1FCD9F9DB0E2BF9E2" box="[819,877,1621,1643]" gridcol="2" gridrow="3" pageId="53" pageNumber="54">100</td>
<td id="BA255D82E87B2DB1FBEFF9DB0963F9E2" box="[1029,1061,1621,1643]" gridcol="3" gridrow="3" pageId="53" pageNumber="54">2</td>
<td id="BA255D82E87B2DB1FB3CF9DB0869F9E2" box="[1238,1327,1621,1643]" gridcol="4" gridrow="3" pageId="53" pageNumber="54">97–101</td>
</tr>
<tr id="F9F434FEE87B2DB1FF75F9F00869F91D" box="[159,1327,1662,1684]" gridrow="4" pageId="53" pageNumber="54">
<th id="BA255D82E87B2DB1FF75F9F00C6AF91D" box="[159,300,1662,1684]" gridcol="0" gridrow="4" pageId="53" pageNumber="54">Orifice width</th>
<td id="BA255D82E87B2DB1FE0BF9F00FEBF91D" box="[481,685,1662,1684]" gridcol="1" gridrow="4" pageId="53" pageNumber="54">1, 6</td>
<td id="BA255D82E87B2DB1FCD9F9F00E2BF91D" box="[819,877,1662,1684]" gridcol="2" gridrow="4" pageId="53" pageNumber="54">97</td>
<td id="BA255D82E87B2DB1FBEFF9F00963F91D" box="[1029,1061,1662,1684]" gridcol="3" gridrow="4" pageId="53" pageNumber="54">4</td>
<td id="BA255D82E87B2DB1FB3CF9F00869F91D" box="[1238,1327,1662,1684]" gridcol="4" gridrow="4" pageId="53" pageNumber="54">90–101</td>
</tr>
</table>
</paragraph>
<paragraph id="477B36BCE87BD27BFF7DF93B0EA9F945" blockId="53.[151,1007,1717,1740]" box="[151,1007,1717,1740]" pageId="53" pageNumber="54">N, Number of colonies and number of zooids measured; SD, standard deviation.</paragraph>
</subSubSection>
</treatment>
</document>
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