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<document id="7415AEC6E965A228605B740E58443900" ID-CLB-Dataset="20928" ID-DOI="10.11646/zootaxa.5093.4.4" ID-GBIF-Dataset="d6297960-52dd-4b0a-a51b-05bda0e7d69f" ID-ISSN="1175-5326" ID-Zenodo-Dep="5912781" ID-ZooBank="4FE93B11-0714-400F-B9A6-41A54E2FCDC9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1643354449005" checkinUser="plazi" docAuthor="Spinelli, Gustavo R., Ronderos, Maria M. &amp; Grogan, William L." docDate="2022" docId="4D4A9349FFE539251AD2F8FAFD94FDA8" docLanguage="en" docName="zootaxa.5093.4.4.pdf" docOrigin="Zootaxa 5093 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macrurohelea bassoi Spinelli &amp; Ronderos &amp; Grogan 2022, n.sp." docType="treatment" docVersion="7" lastPageNumber="450" masterDocId="B173EB31FFE739201A45FFD5FFFBFF95" masterDocTitle="Five new species in the predaceous midge genus Macrurohelea Ingram &amp; Macfie from Argentina, and descriptions of the previously unknown males of M. kuscheli Wirth and M. monotheca Spinelli &amp; Grogan (Diptera: Ceratopogonidae)" masterLastPageNumber="464" masterPageNumber="445" pageNumber="447" updateTime="1699304617883" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="92917528C9434CA4C1C77E02196ABCA9">Five new species in the predaceous midge genus Macrurohelea Ingram &amp; Macfie from Argentina, and descriptions of the previously unknown males of M. kuscheli Wirth and M. monotheca Spinelli &amp; Grogan (Diptera: Ceratopogonidae)</mods:title>
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<mods:namePart id="0D7E6C033B955E2AB1BFAF1C2DCEF616">Spinelli, Gustavo R.</mods:namePart>
<mods:affiliation id="28E685545BC893EB0F57EF12E9519956">División Entomología, Museo de La Plata, Paseo del Bosque s / n, 1900 La Plata, Argentina. &amp; Instituto de Limnología “ Dr. Raúl A. Ringuelet ” (Consejo Nacional de Investigaciones Científicas y Técnicas de la Argentina - Universidad Nacional de La Plata), Boulevard 120 y 62 s / n, 1900 La Plata, Buenos Aires, Argentina.</mods:affiliation>
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<mods:namePart id="C8D515ACDC8BFB7ACEB52CA8BCBA51BE">Ronderos, Maria M.</mods:namePart>
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<mods:affiliation id="ECF23F904726BCBDB7033857C1948D6B">División Entomología, Museo de La Plata, Paseo del Bosque s / n, 1900 La Plata, Argentina. &amp; Centro de Estudios Parasitológicos y de Vectores (Consejo Nacional de Investigaciones Científicas y Técnicas de la Argentina - Universidad Nacional de La Plata), Boulevard 120 y 62 s / n, 1900 La Plata, Buenos Aires, Argentina. &amp; ronderos @ fcnym. unlp. edu. ar; https: // orcid. org / 0000 - 0001 - 8322 - 1471</mods:affiliation>
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<mods:namePart id="F97A3B526C87077993C8AD5838E59858">Grogan, William L.</mods:namePart>
<mods:nameIdentifier id="5070D04E50A52013343A7C57D2C939CE" type="ORCID">0000-0002-6475-8692</mods:nameIdentifier>
<mods:affiliation id="AAFDF09782587BB8CE03AEFD717169C2">Florida State Collection of Arthropods, Florida Department of Agriculture and Consumer Services, Gainesville, Florida 32614 - 7100 USA. &amp; William. Grogan @ fdacs. gov; https: // orcid. org / 0000 - 0002 - 6475 - 8692</mods:affiliation>
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<treatment id="4D4A9349FFE539251AD2F8FAFD94FDA8" ID-DOI="http://doi.org/10.5281/zenodo.5914093" ID-GBIF-Taxon="192700745" ID-Zenodo-Dep="5914093" LSID="urn:lsid:plazi:treatment:4D4A9349FFE539251AD2F8FAFD94FDA8" httpUri="http://treatment.plazi.org/id/4D4A9349FFE539251AD2F8FAFD94FDA8" lastPageId="5" lastPageNumber="450" pageId="2" pageNumber="447">
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<heading id="9E149533FFE539221AD2F8FAFE22F8DF" bold="true" box="[151,473,1839,1866]" fontSize="11" level="1" pageId="2" pageNumber="447" reason="1">
<emphasis id="F797FE4DFFE539221AD2F8FAFE22F8DF" bold="true" box="[151,473,1839,1866]" pageId="2" pageNumber="447">
<taxonomicName id="02E359DCFFE539221AD2F8FAFE6DF8DF" ID-CoL="9HVMC" authority="Spinelli &amp; Ronderos &amp; Grogan, 2022" authorityName="Spinelli &amp; Ronderos &amp; Grogan" authorityYear="2022" box="[151,406,1839,1866]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="bassoi" status="sp. nov.">
<emphasis id="F797FE4DFFE539221AD2F8FAFE6DF8DF" bold="true" box="[151,406,1839,1866]" italics="true" pageId="2" pageNumber="447">Macrurohelea bassoi</emphasis>
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<taxonomicNameLabel id="ECA44336FFE539221BDBF8E5FE22F8DF" box="[414,473,1840,1866]" pageId="2" pageNumber="447" rank="species">n.sp.</taxonomicNameLabel>
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<subSubSection id="8DF971D4FFE539221AD2F881FBCDF807" pageId="2" pageNumber="447" type="description">
<paragraph id="C55C225FFFE539221AD2F881FE9DF8FB" blockId="2.[151,1078,1839,1938]" box="[151,358,1876,1902]" pageId="2" pageNumber="447">
(
<figureCitation id="5DD83EDAFFE539221ADAF881FEF0F8FB" box="[159,267,1876,1902]" captionStart="FIGURES 17" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[214,1376,188,1920]" captionTargetId="figure-23@4.[210,1378,181,1920]" captionTargetPageId="4" captionText="FIGURES 17. Macrurohelea bassoi n. sp. Allotype female, 14. Holotype male, 57. 1, 5 head. 2 thorax. 3, 6 wing. 4 tip of abdomen. 7 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912783" httpUri="https://zenodo.org/record/5912783/files/figure.png" pageId="2" pageNumber="447">Figs. 17</figureCitation>
,
<figureCitation id="5DD83EDAFFE539221B52F881FEA5F8FB" box="[279,350,1876,1902]" captionStart="FIGURES 3742" captionStartId="16.[151,264,1913,1937]" captionTargetBox="[214,1370,182,1882]" captionTargetId="figure-23@16.[214,1373,181,1889]" captionTargetPageId="16" captionText="FIGURES 3742. Macrurohelea bassoi n. sp. Holotype male, 3738. Macrurohelea donatoi n. sp. Holotype male, 3940. Macrurohelea kuscheli Wirth, male, 4142. 37, 39, 41 parameres. 38, 40, 42 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912797" httpUri="https://zenodo.org/record/5912797/files/figure.png" pageId="2" pageNumber="447">3738</figureCitation>
)
</paragraph>
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Zoobank
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urn:lsid:zoobank.org:act:
<uuid id="B145188AFFE539221851F8ADFBCDF807" box="[532,1078,1912,1938]" pageId="2" pageNumber="447">CCBCE121-819C-47B1-97CF-827112AD687D</uuid>
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<subSubSection id="8DF971D4FFE539231AD2F815FDFFF9C4" lastPageId="3" lastPageNumber="448" pageId="2" pageNumber="447" type="diagnosis">
<paragraph id="C55C225FFFE539231AD2F815FB5FFED4" blockId="2.[151,1436,1984,2010]" lastBlockId="3.[151,1437,151,2013]" lastPageId="3" lastPageNumber="448" pageId="2" pageNumber="447">
<emphasis id="F797FE4DFFE539221AD2F815FEE9F84F" bold="true" box="[151,274,1984,2010]" pageId="2" pageNumber="447">Diagnosis.</emphasis>
Females: the only species with a dark brown thorax and contrasting yellowish legs and abdomen; halter pale, transparent; and abdominal segment 9 greatly elongate. Males: the only species with a crescent-shaped tergite 10; gonocoxite stout with pointed basomesal tubercle; gonostylus abruptly curved at mid-length, distal ½ highly modified with inner mesal hump and broad foot-shaped apex; parameres with distal halves closely approximated, nearly straight, basal arms tapered distally, recurved 120° with inner slender external extension, apices with broad bulbous tips; and distal portion of aedeagus tongue-shaped with tip slightly notched mesally.
</paragraph>
<paragraph id="C55C225FFFE439231A82FE9EFF2BFC30" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="448">
<emphasis id="F797FE4DFFE439231A82FE9EFEDEFEF0" bold="true" box="[199,293,331,357]" pageId="3" pageNumber="448">Female.</emphasis>
Head (
<figureCitation id="5DD83EDAFFE439231B3AFE9EFE3FFEF0" box="[383,452,331,357]" captionStart="FIGURES 17" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[214,1376,188,1920]" captionTargetId="figure-23@4.[210,1378,181,1920]" captionTargetPageId="4" captionText="FIGURES 17. Macrurohelea bassoi n. sp. Allotype female, 14. Holotype male, 57. 1, 5 head. 2 thorax. 3, 6 wing. 4 tip of abdomen. 7 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912783" httpUri="https://zenodo.org/record/5912783/files/figure.png" pageId="3" pageNumber="448">Fig. 1</figureCitation>
) dark brown. Eyes narrowly separated medially by the width of 2 ommatidia, with numerous short interommatidial spicules. Antennal flagellum uniformly golden brown; flagellomere 1 with pair of apical sensilla coeloconica; flagellomeres 18 short, vasiform, 913 more elongate, 13 longest; antennal ratio 1.061.20 (1.13, n=10). Palpus pale brown, segment 5 slightly infuscated on distal 1/2; segment 3 with very shallow sensory pit (imperceptible in some specimens); segment 4 short, segment 5 as long as segment 3; palpal ratio 2.002.80 (2.30, n=10). Mandible with 810 coarse medial teeth. Thorax (
<figureCitation id="5DD83EDAFFE4392319AFFE2AFBD4FD8C" box="[1002,1071,511,537]" captionStart="FIGURES 17" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[214,1376,188,1920]" captionTargetId="figure-23@4.[210,1378,181,1920]" captionTargetPageId="4" captionText="FIGURES 17. Macrurohelea bassoi n. sp. Allotype female, 14. Holotype male, 57. 1, 5 head. 2 thorax. 3, 6 wing. 4 tip of abdomen. 7 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912783" httpUri="https://zenodo.org/record/5912783/files/figure.png" pageId="3" pageNumber="448">Fig. 2</figureCitation>
) uniformly dark brown; scutum with 2 stout prealar setae, 1 postalar seta; scutellum with 3 stout setae and 4 slender setae. Legs yellowish, hind tibia slightly darker; tarsomeres 35 slightly infuscated; tarsomere 1 of fore, hind legs with ventral row of palisade setae; tarsomeres 4 cordiform; tarsomeres 5 with pair of small claws slightly curved at tip; hind tarsal ratio 2.00 (n=10). Wing (
<figureCitation id="5DD83EDAFFE439231AA1FD5AFEDCFD3C" box="[228,295,655,681]" captionStart="FIGURES 17" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[214,1376,188,1920]" captionTargetId="figure-23@4.[210,1378,181,1920]" captionTargetPageId="4" captionText="FIGURES 17. Macrurohelea bassoi n. sp. Allotype female, 14. Holotype male, 57. 1, 5 head. 2 thorax. 3, 6 wing. 4 tip of abdomen. 7 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912783" httpUri="https://zenodo.org/record/5912783/files/figure.png" pageId="3" pageNumber="448">Fig. 3</figureCitation>
) membrane hyaline, slightly infuscated with dense microtrichiae; veins pale yellowish; 2
<superScript id="32968F17FFE439231F50FD5BFADEFD09" attach="left" box="[1301,1317,654,668]" fontSize="6" pageId="3" pageNumber="448">nd</superScript>
radial cell 2.443.33 (2.91, n=10)x longer than 1st; cell r
<subScript id="5967201AFFE4392318D9FD10FD5FFD46" attach="left" box="[668,676,709,723]" fontSize="6" pageId="3" pageNumber="448">3</subScript>
without intercalary veins; r-m crossvein slightly longer than petiole of M; costa with fringe of stout macrotrichia; wing length 1.301.42 (1.38, n=10) mm, width 0.580.63 (0.60, n=10) mm; costal ratio 0.720.76 (0.73, n=10). Halter pale. Abdomen (
<figureCitation id="5DD83EDAFFE43923192AFD2EFC48FC80" box="[879,947,763,789]" captionStart="FIGURES 17" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[214,1376,188,1920]" captionTargetId="figure-23@4.[210,1378,181,1920]" captionTargetPageId="4" captionText="FIGURES 17. Macrurohelea bassoi n. sp. Allotype female, 14. Holotype male, 57. 1, 5 head. 2 thorax. 3, 6 wing. 4 tip of abdomen. 7 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912783" httpUri="https://zenodo.org/record/5912783/files/figure.png" pageId="3" pageNumber="448">Fig. 4</figureCitation>
) yellowish. Segment 9 greatly elongate, 10 elongate but much shorter than 9, both segments bent anteroventrally; sternite 8 lightly sclerotized with a short, rounded, posteromedian excavation. Two ovoid, slightly unequal-size spermathecae with moderately long, slender necks, measuring 5470 (65, n=7) by 4554 (48, n=7) µm, 5066 (58, n=7) by 4348 (45, n=7) µm, necks 9 µm long.
</paragraph>
<paragraph id="C55C225FFFE439231A82FC7AFDFFF9C4" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="448">
<emphasis id="F797FE4DFFE439231A82FC7AFEF0FC5C" bold="true" box="[199,267,943,969]" pageId="3" pageNumber="448">Male.</emphasis>
Similar to female with the following notable sexual differences. Head (
<figureCitation id="5DD83EDAFFE439231E6AFC7AFB88FC5C" box="[1071,1139,942,969]" captionStart="FIGURES 17" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[214,1376,188,1920]" captionTargetId="figure-23@4.[210,1378,181,1920]" captionTargetPageId="4" captionText="FIGURES 17. Macrurohelea bassoi n. sp. Allotype female, 14. Holotype male, 57. 1, 5 head. 2 thorax. 3, 6 wing. 4 tip of abdomen. 7 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912783" httpUri="https://zenodo.org/record/5912783/files/figure.png" pageId="3" pageNumber="448">Fig. 5</figureCitation>
). Eyes narrowly separated medially (visible in the male
<typeStatus id="1A589CFDFFE439231BADFC06FDB7FC78" box="[488,588,979,1005]" pageId="3" pageNumber="448" type="paratype">paratype</typeStatus>
; the top of the head is fractured in the
<typeStatus id="1A589CFDFFE439231E48FC06FB8FFC7B" box="[1037,1140,979,1006]" pageId="3" pageNumber="448" type="holotype">holotype</typeStatus>
). Antennal flagellum with flagellomeres distinctly separated; flagellomeres 28 vasiform, 910 slightly elongate, 1113 greatly elongate; antennal ratio 0.720.80 (0.76, n=3). Palpus similar to female, moderately short; segment 3 with few sensilla on surface; palpal ratio 2.703.00 (2.90, n=3). Thorax similar to female. Legs brown, tarsal claws small, nearly straight with bifid tips. Wing (
<figureCitation id="5DD83EDAFFE439231BCBFBB6FE29FBEB" box="[398,466,1123,1150]" captionStart="FIGURES 17" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[214,1376,188,1920]" captionTargetId="figure-23@4.[210,1378,181,1920]" captionTargetPageId="4" captionText="FIGURES 17. Macrurohelea bassoi n. sp. Allotype female, 14. Holotype male, 57. 1, 5 head. 2 thorax. 3, 6 wing. 4 tip of abdomen. 7 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912783" httpUri="https://zenodo.org/record/5912783/files/figure.png" pageId="3" pageNumber="448">Fig. 6</figureCitation>
) narrower than female, with darker, well defined posterior veins; 2
<superScript id="32968F17FFE439231E84FBB7FB2AFBE5" attach="left" box="[1217,1233,1122,1136]" fontSize="6" pageId="3" pageNumber="448">nd</superScript>
radial cell shorter than female, only 1.5x longer than 1
<superScript id="32968F17FFE43923187DFB52FDB8FB00" attach="none" box="[568,579,1159,1173]" fontSize="6" pageId="3" pageNumber="448">st</superScript>
; wing length 1.321.48 (1.39, n=3) mm, width 0.490.51 (0.50, n=3) mm; costal ratio 0.64 (n=3). Abdomen dark brown. Genitalia (
<figureCitation id="5DD83EDAFFE439231977FB7EFC81FB50" box="[818,890,1195,1221]" captionStart="FIGURES 17" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[214,1376,188,1920]" captionTargetId="figure-23@4.[210,1378,181,1920]" captionTargetPageId="4" captionText="FIGURES 17. Macrurohelea bassoi n. sp. Allotype female, 14. Holotype male, 57. 1, 5 head. 2 thorax. 3, 6 wing. 4 tip of abdomen. 7 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912783" httpUri="https://zenodo.org/record/5912783/files/figure.png" pageId="3" pageNumber="448">Fig. 7</figureCitation>
). Tergite 9 triangular, extending 0.70 length of gonocoxites, with blunt apex. Tergite 10 crescent-shaped, extending to almost the level of apex of gonocoxites; apicolateral process minute, with medium-size seta; cercus slender, finger-like, setose. Sternite 9 2.7x broader than long, posteromedian margin slightly convex. Gonocoxite stout, 1.45x longer than wide, with prominent basomesal tubercle with pointed tip; gonostylus stout, 0.85 length of gonocoxite, abruptly curved at mid-length, distal ½ highly modified with inner mesal hump and broad foot-shaped apex. Parameres (
<figureCitation id="5DD83EDAFFE439231E6CFA8AFB80FAEC" box="[1065,1147,1375,1401]" captionStart="FIGURES 3742" captionStartId="16.[151,264,1913,1937]" captionTargetBox="[214,1370,182,1882]" captionTargetId="figure-23@16.[214,1373,181,1889]" captionTargetPageId="16" captionText="FIGURES 3742. Macrurohelea bassoi n. sp. Holotype male, 3738. Macrurohelea donatoi n. sp. Holotype male, 3940. Macrurohelea kuscheli Wirth, male, 4142. 37, 39, 41 parameres. 38, 40, 42 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912797" httpUri="https://zenodo.org/record/5912797/files/figure.png" pageId="3" pageNumber="448">Fig. 37</figureCitation>
) separate; basal apodeme slender, arcuate, directed posterolaterally, anterior margin with heavily sclerotized colon-shaped process directed posteriorly; distal portion nearly straight, basal 2/3 broad, abruptly tapering distally, tip bulbous. Aedeagus (
<figureCitation id="5DD83EDAFFE439231F06FA72FA6FFA54" box="[1347,1428,1447,1473]" captionStart="FIGURES 3742" captionStartId="16.[151,264,1913,1937]" captionTargetBox="[214,1370,182,1882]" captionTargetId="figure-23@16.[214,1373,181,1889]" captionTargetPageId="16" captionText="FIGURES 3742. Macrurohelea bassoi n. sp. Holotype male, 3738. Macrurohelea donatoi n. sp. Holotype male, 3940. Macrurohelea kuscheli Wirth, male, 4142. 37, 39, 41 parameres. 38, 40, 42 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912797" httpUri="https://zenodo.org/record/5912797/files/figure.png" pageId="3" pageNumber="448">Fig. 38</figureCitation>
) elongate, tongue-shaped, slightly shorter than basal width; basal arms broad basally, widely spaced at bases, apex narrowed, recurved, heavily sclerotized; basal arch V-shaped extending 0.25 of total length (shorter and slightly concave in
<specimenCount id="D3E5E9D6FFE439231B50F9C6FE52F9B8" box="[277,425,1555,1581]" pageId="3" pageNumber="448" type="generic" typeStatus="paratype">one paratype</specimenCount>
); distolateral extension hyaline with lateral margins slightly divergent, with broad, rounded and slightly mesally notched tip.
</paragraph>
</subSubSection>
<subSubSection id="8DF971D4FFE439231A82F98FFB8FF9E0" box="[199,1140,1626,1653]" pageId="3" pageNumber="448" type="distribution">
<paragraph id="C55C225FFFE439231A82F98FFB8FF9E0" blockId="3.[151,1437,151,2013]" box="[199,1140,1626,1653]" pageId="3" pageNumber="448">
<emphasis id="F797FE4DFFE439231A82F98FFE9AF9E1" bold="true" box="[199,353,1626,1652]" pageId="3" pageNumber="448">Distribution.</emphasis>
<collectingCountry id="BDF462CFFFE439231B22F98EFE22F9E0" box="[359,473,1627,1653]" name="Argentina" pageId="3" pageNumber="448">Argentina</collectingCountry>
, in
<taxonomicName id="02E359DCFFE439231847F98EFD78F9E0" box="[514,643,1627,1653]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="448" phylum="Tracheophyta" rank="genus">
<emphasis id="F797FE4DFFE439231847F98EFD78F9E0" box="[514,643,1627,1653]" italics="true" pageId="3" pageNumber="448">Nothofagus</emphasis>
</taxonomicName>
forests of
<collectingRegion id="0727ECBDFFE4392318BFF98EFC94F9E0" box="[762,879,1627,1653]" country="Argentina" name="Rio Negro" pageId="3" pageNumber="448">Río Negro</collectingRegion>
and
<collectingRegion id="0727ECBDFFE4392319E3F98EFC03F9E0" box="[934,1016,1627,1653]" country="Argentina" name="Chubut" pageId="3" pageNumber="448">Chubut</collectingRegion>
provinces.
</paragraph>
</subSubSection>
<subSubSection id="8DF971D4FFE439231A82F9AAFE98F848" pageId="3" pageNumber="448" type="materials_examined">
<paragraph id="C55C225FFFE439231A82F9AAFD32F8E4" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="448">
<emphasis id="F797FE4DFFE439231A82F9AAFE86F90D" bold="true" box="[199,381,1662,1689]" pageId="3" pageNumber="448">Type material.</emphasis>
<materialsCitation id="758B2802FFE439231BCEF9AAFA62F928" ID-GBIF-Occurrence="3458732304" collectingDate="1999-11-29" collectionCode="MLPA" collectorName="G. Spinelli" country="Argentina" latitude="-41.964924" location="El Bolson" longLatPrecision="1" longitude="-71.53534" pageId="3" pageNumber="448" specimenCount="2" stateProvince="Rio Negro" typeStatus="holotype">
<typeStatus id="1A589CFDFFE439231BCEF9AAFE08F90C" box="[395,499,1663,1689]" pageId="3" pageNumber="448" type="holotype">Holotype</typeStatus>
male labeled “
<typeStatus id="1A589CFDFFE4392318F4F9AAFCE7F90C" box="[689,796,1663,1689]" pageId="3" pageNumber="448" type="holotype">Holotype</typeStatus>
<taxonomicName id="02E359DCFFE43923196FF9AAFA75F90C" authority="Spinelli, Ronderos &amp; Grogan" authorityName="Spinelli, Ronderos &amp; Grogan" box="[810,1422,1663,1689]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="448" phylum="Arthropoda" rank="species" species="bassoi">Macrurohelea bassoi Spinelli, Ronderos &amp; Grogan</taxonomicName>
”, “
<collectingCountry id="BDF462CFFFE439231AE4F976FEEDF928" box="[161,278,1699,1725]" name="Argentina" pageId="3" pageNumber="448">Argentina</collectingCountry>
,
<collectingRegion id="0727ECBDFFE439231B60F976FE59F928" box="[293,418,1699,1725]" country="Argentina" name="Rio Negro" pageId="3" pageNumber="448">Río Negro</collectingRegion>
,
<location id="C03C7484FFE439231BF7F976FDD3F928" LSID="urn:lsid:plazi:treatment:4D4A9349FFE539251AD2F8FAFD94FDA8:C03C7484FFE439231BF7F976FDD3F928" box="[434,552,1699,1725]" country="Argentina" latitude="-41.964924" longLatPrecision="1" longitude="-71.53534" name="El Bolson" pageId="3" pageNumber="448" stateProvince="Rio Negro">El Bolsón</location>
,
<geoCoordinate id="A0D74498FFE43923187DF977FD19F928" box="[568,738,1698,1725]" degrees="41" direction="south" minutes="57" orientation="latitude" pageId="3" pageNumber="448" precision="1" seconds="53.72" value="-41.964924">41°5753.72”S</geoCoordinate>
,
<geoCoordinate id="A0D74498FFE4392318B7F976FC6DF928" box="[754,918,1699,1725]" degrees="71" direction="west" minutes="32" orientation="longitude" pageId="3" pageNumber="448" precision="1" seconds="7.22" value="-71.53534">71°327.22”W</geoCoordinate>
,
<date id="B15D049FFFE4392319E3F976FBD7F928" box="[934,1068,1699,1725]" pageId="3" pageNumber="448" value="1999-11-29">
<collectingDate id="A119FD77FFE4392319E3F976FBD7F928" box="[934,1068,1699,1725]" pageId="3" pageNumber="448" value="1999-11-29">29-XI-1999</collectingDate>
</date>
,
<collectorName id="68164789FFE439231E79F976FB46F928" box="[1084,1213,1699,1725]" pageId="3" pageNumber="448">G. Spinelli</collectorName>
, at light” (
<collectionCode id="A3F2BA9AFFE439231F7AF976FA6AF928" box="[1343,1425,1699,1725]" pageId="3" pageNumber="448">MLPA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="758B2802FFE439231AD2F912FE24F8BC" ID-GBIF-Occurrence="3458732308" collectingDate="2007-02-18" collectionCode="MLPA" collectorName="A. Garre &amp; F. Montes de Oca" country="Argentina" elevation="962" latitude="-41.19775" location="rio Nireco" longLatPrecision="1" longitude="-71.32792" municipality="Parque Nacional Nahuel Huapi" pageId="3" pageNumber="448" specimenCount="21" specimenCount-female="19" specimenCount-male="2" stateProvince="Rio Negro" typeStatus="paratype">
<typeStatus id="1A589CFDFFE439231AD2F912FEFCF974" box="[151,263,1735,1761]" pageId="3" pageNumber="448" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="D3E5E9D6FFE439231B56F912FE96F974" box="[275,365,1735,1761]" pageId="3" pageNumber="448" type="male">2 males</specimenCount>
,
<specimenCount id="D3E5E9D6FFE439231B3CF912FE0CF974" box="[377,503,1735,1761]" pageId="3" pageNumber="448" type="female">19 females</specimenCount>
, as follows:
<typeStatus id="1A589CFDFFE4392318C3F912FD1BF974" box="[646,736,1735,1761]" pageId="3" pageNumber="448" type="allotype">allotype</typeStatus>
female and 15 other females with same data as holotype;
<collectingRegion id="0727ECBDFFE439231F31F912FF1BF890" country="Argentina" name="Rio Negro" pageId="3" pageNumber="448">Río Negro</collectingRegion>
,
<collectingMunicipality id="2538B825FFE439231AA8F93EFDAAF890" box="[237,593,1771,1797]" pageId="3" pageNumber="448">Parque Nacional Nahuel Huapi</collectingMunicipality>
,
<location id="C03C7484FFE43923181BF93EFD2EF892" LSID="urn:lsid:plazi:treatment:4D4A9349FFE539251AD2F8FAFD94FDA8:C03C7484FFE43923181BF93EFD2EF892" box="[606,725,1771,1799]" latitude="-41.19775" longLatPrecision="1" longitude="-71.32792" municipality="Parque Nacional Nahuel Huapi" name="rio Nireco" pageId="3" pageNumber="448" stateProvince="Rio Negro">río Ñireco</location>
,
<geoCoordinate id="A0D74498FFE4392318A7F93FFC80F890" box="[738,891,1770,1797]" degrees="41" direction="south" minutes="11" orientation="latitude" pageId="3" pageNumber="448" precision="1" seconds="51.9" value="-41.19775">41º1151.9S</geoCoordinate>
<geoCoordinate id="A0D74498FFE4392319C1F93FFBD1F890" box="[900,1066,1770,1797]" degrees="71" direction="west" minutes="19" orientation="longitude" pageId="3" pageNumber="448" precision="1" seconds="40.5" value="-71.32792">71º1940.5W</geoCoordinate>
,
<quantity id="021B8FBAFFE439231E72F93EFB79F890" box="[1079,1154,1771,1797]" metricMagnitude="2" metricUnit="m" metricValue="9.62" pageId="3" pageNumber="448" unit="m" value="962.0">
<elevation id="4ECEC56CFFE439231E72F93EFB79F890" box="[1079,1154,1771,1797]" metricMagnitude="2" metricUnit="m" metricValue="9.62" pageId="3" pageNumber="448" unit="m" value="962.0">962 m</elevation>
</quantity>
,
<date id="B15D049FFFE439231ECAF93EFAF0F890" box="[1167,1291,1771,1797]" pageId="3" pageNumber="448" value="2007-02-18">
<collectingDate id="A119FD77FFE439231ECAF93EFAF0F890" box="[1167,1291,1771,1797]" pageId="3" pageNumber="448" value="2007-02-18">18-II-2007</collectingDate>
</date>
,
<collectorName id="68164789FFE439231F53F93EFA80F890" box="[1302,1403,1771,1797]" pageId="3" pageNumber="448">A. Garré</collectorName>
<collectorName id="68164789FFE439231FCFF93EFEBEF8BC" pageId="3" pageNumber="448">F. Montes de Oca</collectorName>
,
<collectingMethod id="1CA25A48FFE439231B15F8DAFE24F8BC" box="[336,479,1807,1833]" pageId="3" pageNumber="448">Malaise trap</collectingMethod>
</materialsCitation>
,
<materialsCitation id="758B2802FFE439231BAEF8DAFE8EF8D8" ID-GBIF-Occurrence="3458732315" collectorName="L. Quate" country="Argentina" latitude="-42.86661" location="Parque Nacional Los Alerces" longLatPrecision="1" longitude="-71.60132" pageId="3" pageNumber="448" specimenCount="4" specimenCount-female="4" stateProvince="Chubut" typeStatus="paratype">
<specimenCount id="D3E5E9D6FFE439231BAEF8DAFDA0F8BC" box="[491,603,1807,1833]" pageId="3" pageNumber="448" type="female">4 females</specimenCount>
;
<collectingRegion id="0727ECBDFFE439231822F8DAFD47F8BC" box="[615,700,1807,1833]" country="Argentina" name="Chubut" pageId="3" pageNumber="448">Chubut</collectingRegion>
,
<location id="C03C7484FFE43923188DF8DAFBE9F8BC" LSID="urn:lsid:plazi:treatment:4D4A9349FFE539251AD2F8FAFD94FDA8:C03C7484FFE43923188DF8DAFBE9F8BC" box="[712,1042,1807,1833]" country="Argentina" latitude="-42.86661" longLatPrecision="1" longitude="-71.60132" name="Parque Nacional Los Alerces" pageId="3" pageNumber="448" stateProvince="Chubut">Parque Nacional Los Alerces</location>
,
<geoCoordinate id="A0D74498FFE439231E5BF8DBFB33F8BC" box="[1054,1224,1806,1833]" degrees="42" direction="south" minutes="51" orientation="latitude" pageId="3" pageNumber="448" precision="1" seconds="59.81" value="-42.86661">42°5159.81”S</geoCoordinate>
,
<geoCoordinate id="A0D74498FFE439231E91F8DAFA83F8BC" box="[1236,1400,1807,1833]" degrees="71" direction="west" minutes="36" orientation="longitude" pageId="3" pageNumber="448" precision="1" seconds="4.76" value="-71.60132">71°364.76”W</geoCoordinate>
,
<collectorName id="68164789FFE439231FC1F8DAFF26F8D8" pageId="3" pageNumber="448">L. Quate</collectorName>
,
<collectingMethod id="1CA25A48FFE439231AADF8E6FE8EF8D8" box="[232,373,1843,1869]" pageId="3" pageNumber="448">Malaise trap</collectingMethod>
</materialsCitation>
,
<materialsCitation id="758B2802FFE439231BC4F8E6FE0DF8E4" ID-GBIF-Occurrence="3458732317" collectingDate="2002-12-02" collectorName="G. Spinelli" country="Argentina" elevation="670" latitude="-45.780556" location="Galera" longLatPrecision="18" longitude="-71.73222" municipality="Puesto de Gendarmeria" pageId="3" pageNumber="448" specimenCount="1" specimenCount-male="1" stateProvince="Chubut" typeStatus="paratype">
<specimenCount id="D3E5E9D6FFE439231BC4F8E6FE2BF8D8" box="[385,464,1843,1869]" pageId="3" pageNumber="448" type="male">1 male</specimenCount>
;
<collectingRegion id="0727ECBDFFE439231B9EF8E6FDCBF8D8" box="[475,560,1843,1869]" country="Argentina" name="Chubut" pageId="3" pageNumber="448">Chubut</collectingRegion>
,
<collectingMunicipality id="2538B825FFE43923187EF8E6FCBBF8D8" box="[571,832,1843,1869]" pageId="3" pageNumber="448">Puesto de Gendarmería</collectingMunicipality>
“El Triana”, cerro
<location id="C03C7484FFE439231E50F8E6FB99F8D8" LSID="urn:lsid:plazi:treatment:4D4A9349FFE539251AD2F8FAFD94FDA8:C03C7484FFE439231E50F8E6FB99F8D8" box="[1045,1122,1843,1869]" country="Argentina" latitude="-45.780556" longLatPrecision="18" longitude="-71.73222" municipality="Puesto de Gendarmeria" name="Galera" pageId="3" pageNumber="448" stateProvince="Chubut">Galera</location>
,
<geoCoordinate id="A0D74498FFE439231E2BF8E7FB03F8D8" box="[1134,1272,1842,1869]" degrees="45" direction="south" minutes="46" orientation="latitude" pageId="3" pageNumber="448" precision="15" seconds="50" value="-45.780556">45°4650S</geoCoordinate>
,
<geoCoordinate id="A0D74498FFE439231F41F8E7FA63F8D8" box="[1284,1432,1842,1869]" degrees="71" direction="west" minutes="43" orientation="longitude" pageId="3" pageNumber="448" precision="15" seconds="56" value="-71.73222">71°4356W</geoCoordinate>
,
<quantity id="021B8FBAFFE439231AD2F882FF1BF8E4" box="[151,224,1879,1905]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="3" pageNumber="448" unit="m" value="670.0">
<elevation id="4ECEC56CFFE439231AD2F882FF1BF8E4" box="[151,224,1879,1905]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="3" pageNumber="448" unit="m" value="670.0">670 m</elevation>
</quantity>
,
<date id="B15D049FFFE439231AA9F882FE95F8E4" box="[236,366,1879,1905]" pageId="3" pageNumber="448" value="2002-12-02">
<collectingDate id="A119FD77FFE439231AA9F882FE95F8E4" box="[236,366,1879,1905]" pageId="3" pageNumber="448" value="2002-12-02">2-XII-2002</collectingDate>
</date>
,
<collectorName id="68164789FFE439231B3CF882FE0DF8E4" box="[377,502,1879,1905]" pageId="3" pageNumber="448">G. Spinelli</collectorName>
</materialsCitation>
,
<specimenCount id="D3E5E9D6FFE439231844F882FDB4F8E4" box="[513,591,1879,1905]" pageId="3" pageNumber="448" type="male">1 male</specimenCount>
, aerial net.
</paragraph>
<paragraph id="C55C225FFFE439231A82F8AFFE98F848" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="448">
<emphasis id="F797FE4DFFE439231A82F8AFFDD2F801" bold="true" box="[199,553,1914,1940]" pageId="3" pageNumber="448">Derivation of specific epithet.</emphasis>
We are pleased to name this new species after Dr. Néstor G. Basso of Centro Nacional Patagónico, Puerto Madryn,
<collectingCountry id="BDF462CFFFE439231808F84AFD44F82C" box="[589,703,1951,1977]" name="Argentina" pageId="3" pageNumber="448">Argentina</collectingCountry>
, for his friendship and companionship during several collecting trips in Patagonia.
</paragraph>
</subSubSection>
<caption id="919C72D7FFE339241AD2F842FDA0F846" ID-DOI="http://doi.org/10.5281/zenodo.5912783" ID-Zenodo-Dep="5912783" httpUri="https://zenodo.org/record/5912783/files/figure.png" pageId="4" pageNumber="449" startId="4.[151,264,1943,1967]" targetBox="[214,1376,188,1920]" targetPageId="4">
<paragraph id="C55C225FFFE339241AD2F842FDA0F846" blockId="4.[151,1436,1943,2004]" pageId="4" pageNumber="449">
<emphasis id="F797FE4DFFE339241AD2F842FEC0F83A" bold="true" box="[151,315,1943,1967]" pageId="4" pageNumber="449">FIGURES 17.</emphasis>
<taxonomicName id="02E359DCFFE339241B07F84DFDE2F825" authority="Spinelli &amp; Ronderos &amp; Grogan, 2022" authorityName="Spinelli &amp; Ronderos &amp; Grogan" authorityYear="2022" box="[322,537,1944,1968]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="449" phylum="Arthropoda" rank="species" species="bassoi" status="sp. nov.">
<emphasis id="F797FE4DFFE339241B07F84DFDE2F825" box="[322,537,1944,1968]" italics="true" pageId="4" pageNumber="449">Macrurohelea bassoi</emphasis>
</taxonomicName>
<emphasis id="F797FE4DFFE339241865F84DFDA1F825" bold="true" box="[544,602,1944,1968]" pageId="4" pageNumber="449">
<taxonomicNameLabel id="ECA44336FFE339241865F84DFDA1F825" box="[544,602,1944,1968]" pageId="4" pageNumber="449" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Allotype female, 14. Holotype male, 57. 1, 5 head. 2 thorax. 3, 6 wing. 4 tip of abdomen. 7 genitalia. Scale bars = 0.05 mm.
</paragraph>
</caption>
<subSubSection id="8DF971D4FFE239251A82FF43FD94FDA8" pageId="5" pageNumber="450" type="discussion">
<paragraph id="C55C225FFFE239251A82FF43FDBCFE44" blockId="5.[151,1437,150,573]" pageId="5" pageNumber="450">
<emphasis id="F797FE4DFFE239251A82FF43FEB7FF25" bold="true" box="[199,332,150,176]" pageId="5" pageNumber="450">Discussion.</emphasis>
This new species keys to couplets 8 (females) and 16 (males) of
<taxonomicName id="02E359DCFFE239251E68FF42FAAEFF24" authority="Grogan &amp; Wirth" authorityName="Grogan &amp; Wirth" box="[1069,1365,151,177]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="450" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="F797FE4DFFE239251E68FF42FBBFFF25" box="[1069,1092,151,176]" italics="true" pageId="5" pageNumber="450">M</emphasis>
.
<emphasis id="F797FE4DFFE239251E17FF42FB6EFF25" box="[1106,1173,151,176]" italics="true" pageId="5" pageNumber="450">irwini</emphasis>
Grogan &amp; Wirth
</taxonomicName>
in the key to Neotropical
<taxonomicName id="02E359DCFFE239251B32FF6EFDEDFF40" box="[375,534,187,213]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="450" phylum="Arthropoda" rank="genus">
<emphasis id="F797FE4DFFE239251B32FF6EFDEDFF40" box="[375,534,187,213]" italics="true" pageId="5" pageNumber="450">Macrurohelea</emphasis>
</taxonomicName>
by
<bibRefCitation id="A1725FAEFFE239251800FF6EFC8AFF40" author="Spinelli, G. R. &amp; Grogan, W. L. Jr." box="[581,881,187,213]" pageId="5" pageNumber="450" pagination="127 - 134" refId="ref14299" refString="Spinelli, G. R. &amp; Grogan, W. L. Jr. (1990) New species of predaceous midges of the tribe Ceratopogonini from subantarctic Argentina (Diptera: Ceratopogonidae). Proceedings of the Entomological Society of Washington, 92, 127 - 134." type="journal article" year="1990">Spinelli &amp; Grogan (1990)</bibRefCitation>
. Males of
<taxonomicName id="02E359DCFFE2392519ABFF6EFBA5FF40" authority="Spinelli &amp; Ronderos &amp; Grogan, 2022" authorityName="Spinelli &amp; Ronderos &amp; Grogan" authorityYear="2022" box="[1006,1118,187,213]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="450" phylum="Arthropoda" rank="species" species="bassoi" status="sp. nov.">
<emphasis id="F797FE4DFFE2392519ABFF6EFBFEFF41" box="[1006,1029,187,212]" italics="true" pageId="5" pageNumber="450">M</emphasis>
.
<emphasis id="F797FE4DFFE239251E53FF6EFBA5FF40" box="[1046,1118,187,213]" italics="true" pageId="5" pageNumber="450">bassoi</emphasis>
</taxonomicName>
<emphasis id="F797FE4DFFE239251E22FF6EFB53FF40" bold="true" box="[1127,1192,187,213]" pageId="5" pageNumber="450">
<taxonomicNameLabel id="ECA44336FFE239251E22FF6EFB53FF40" box="[1127,1192,187,213]" pageId="5" pageNumber="450" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differ from males of
<taxonomicName id="02E359DCFFE239251AD2FF0AFEFAFF6D" box="[151,257,223,249]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="450" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="F797FE4DFFE239251AD2FF0AFF55FF6D" box="[151,174,223,248]" italics="true" pageId="5" pageNumber="450">M</emphasis>
.
<emphasis id="F797FE4DFFE239251AFBFF0AFEFAFF6D" box="[190,257,223,248]" italics="true" pageId="5" pageNumber="450">irwini</emphasis>
</taxonomicName>
by the crescent-shaped tergite 10, the highly modified gonostylus that is abruptly curved at mid-length, the distal ½ is highly modified with an inner mesal hump and broad foot-shaped apex (gonostylus much shorter, narrower with a curved sharply pointed apex in
<taxonomicName id="02E359DCFFE2392518E0FEF2FCF7FED5" box="[677,780,295,321]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="450" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="F797FE4DFFE2392518E0FEF2FD47FED5" box="[677,700,295,320]" italics="true" pageId="5" pageNumber="450">M</emphasis>
.
<emphasis id="F797FE4DFFE23925188CFEF2FCF7FED5" box="[713,780,295,320]" italics="true" pageId="5" pageNumber="450">irwini</emphasis>
</taxonomicName>
); parameres separate with bulbous apices (parameres fused basally, slender, divergent distally with apices bent at 90º in
<taxonomicName id="02E359DCFFE239251907FE9EFC50FEF1" box="[834,939,331,357]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="450" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="F797FE4DFFE239251907FE9EFCA2FEF1" box="[834,857,331,356]" italics="true" pageId="5" pageNumber="450">M</emphasis>
.
<emphasis id="F797FE4DFFE23925192DFE9EFC50FEF1" box="[872,939,331,356]" italics="true" pageId="5" pageNumber="450">irwini</emphasis>
</taxonomicName>
); and, aedeagus slender, elongate with apex rounded and slightly notched mesally (aedeagus short, triangular with very long basal arms that are doubly recurved in
<taxonomicName id="02E359DCFFE239251AF1FE46FEE7FE39" box="[180,284,403,429]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="450" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="F797FE4DFFE239251AF1FE46FF30FE39" box="[180,203,403,428]" italics="true" pageId="5" pageNumber="450">M</emphasis>
.
<emphasis id="F797FE4DFFE239251A9CFE46FEE7FE39" box="[217,284,403,428]" italics="true" pageId="5" pageNumber="450">irwini</emphasis>
</taxonomicName>
). Females differ from those of
<taxonomicName id="02E359DCFFE23925183CFE46FD1AFE39" box="[633,737,403,429]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="450" phylum="Arthropoda" rank="species" species="irwini">
<emphasis id="F797FE4DFFE23925183CFE46FD6BFE39" box="[633,656,403,428]" italics="true" pageId="5" pageNumber="450">M</emphasis>
.
<emphasis id="F797FE4DFFE2392518DBFE46FD1AFE39" box="[670,737,403,428]" italics="true" pageId="5" pageNumber="450">irwini</emphasis>
</taxonomicName>
by the contrasting coloration between their dark brown thorax and their yellowish legs and abdomen.
</paragraph>
<paragraph id="C55C225FFFE239251AD2FE0EFD94FDA8" blockId="5.[151,1437,150,573]" pageId="5" pageNumber="450">
Females and males of this new species were collected on the same date at the type-locality. Although the two sexes differ in the coloration of their legs and abdomen, their wings are nearly identical with the usual sexual differences such as the shorter 2
<superScript id="32968F17FFE239251B39FDF7FE77FDA5" attach="left" box="[380,396,546,560]" fontSize="6" pageId="5" pageNumber="450">nd</superScript>
radial cell in males.
</paragraph>
</subSubSection>
</treatment>
</document>