325 lines
38 KiB
XML
325 lines
38 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/vz.74.e109088" ID-Pensoft-Pub="2625-8498-74-169" ID-Pensoft-UUID="C126D0A1493E5126BE9F97ABDD4605C9" ID-ZooBank="D85DA495456848ECB0AA2A9B48F817EA" ModsDocID="2625-8498-74-169" checkinTime="1709890918002" checkinUser="pensoft" docAuthor="Wang, Kai, Deepak, V., Das, Abhijit, Grismer, L. Lee, Liu, Shuo & Che, Jing" docDate="2024" docId="BF744154BDC55F078993E3E216081DC3" docLanguage="en" docName="VertZool 74: 169-192" docOrigin="Vertebrate Zoology 74" docPubDate="2024-03-07" docSource="http://dx.doi.org/10.3897/vz.74.e109088" docTitle="Calotes maria Gray 1845" docType="treatment" docVersion="1" id="C126D0A1493E5126BE9F97ABDD4605C9" lastPageNumber="169" masterDocId="C126D0A1493E5126BE9F97ABDD4605C9" masterDocTitle="Systematic revision of the Calotes jerdoni complex (Reptilia: Squamata: Agamidae) in the Pan-Himalaya" masterLastPageNumber="192" masterPageNumber="169" pageNumber="169" updateTime="1709890918002" updateUser="pensoft">
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<mods:title id="C8442C7BEFF6A3D07EA5B7A2F150E4BE">Systematic revision of the Calotes jerdoni complex (Reptilia: Squamata: Agamidae) in the Pan-Himalaya</mods:title>
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<mods:namePart id="F25D59BA5C40CD944B064A5BC0EC317C">Wang, Kai</mods:namePart>
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<mods:affiliation id="546E9869681D99DAD65B07F84D362CE1">State Key Laboratory of Genetic Resources and Evolution and Yunnan Key Laboratory of Biodiversity and Ecological Conservation of Gaoligong Mountain, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan, 650223, China & Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences, Yezin, Nay Pyi Taw, 05282, Myanmar</mods:affiliation>
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<mods:nameIdentifier id="F2EB8AD9C9D08BF9DADE960C07F192E1" type="email">wangkai@mail.kiz.ac.cn</mods:nameIdentifier>
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<mods:namePart id="D85FE6F5B497E380F8C718837C6A89CF">Deepak, V.</mods:namePart>
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<mods:affiliation id="8751E828DA0C773150D5FB866FD54BAF">Senckenberg Dresden, Koenigsbruecker Landstrasse 159, 01109 Dresden, Germany & Science Group, The Natural History Museum, London SW 7 5 BD, UK</mods:affiliation>
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<mods:namePart id="1C724E084C6AE2F5E17014781167F71C">Das, Abhijit</mods:namePart>
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<mods:affiliation id="72C9109228BDE1A515ECAC59E786BCE2">Wildlife Institute of India, Chandrabani, Dehradun, 248002, India</mods:affiliation>
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<mods:namePart id="803E4EC264DE287727AC760993898383">Grismer, L. Lee</mods:namePart>
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<mods:affiliation id="EE09D83329E555A42D52EA0E2C8B23EC">Herpetology Laboratory, Department of Biology, La Sierra University, 4500 Riverwalk Parkway, Riverside, California, 92515, USA & Department of Herpetology, San Diego Natural History Museum, PO Box 121390, San Diego, California, 92112, USA & Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu, Malaysia</mods:affiliation>
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<mods:namePart id="CCA9AC39916E514B7D991623B4CA1387">Liu, Shuo</mods:namePart>
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<mods:affiliation id="C7793E9998BDA04112EC70764E0174AC">State Key Laboratory of Genetic Resources and Evolution and Yunnan Key Laboratory of Biodiversity and Ecological Conservation of Gaoligong Mountain, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan, 650223, China</mods:affiliation>
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<mods:namePart id="EAF9DB94EC3F9C85608E2065F51F32F9">Che, Jing</mods:namePart>
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<mods:affiliation id="3853866F8DE612A7042F0C7019658CB3">State Key Laboratory of Genetic Resources and Evolution and Yunnan Key Laboratory of Biodiversity and Ecological Conservation of Gaoligong Mountain, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan, 650223, China & Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences, Yezin, Nay Pyi Taw, 05282, Myanmar</mods:affiliation>
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<mods:title id="394BC75BE4E3EA5EA37E2FAEA3179780">Vertebrate Zoology</mods:title>
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<mods:date id="2D48B067D5DBFA722D683CCAFF032DD6">2024</mods:date>
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<mods:number id="DBFE7E0E501FC66ECA6731B9AF97E424">2024-03-07</mods:number>
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<mods:number id="00121DBDC806B70DD0240FE0C5AA5E58">74</mods:number>
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<mods:start id="DB24953D4D2E3AB3211EE7F43A0286B6">169</mods:start>
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<treatment id="BF744154BDC55F078993E3E216081DC3" LSID="urn:lsid:plazi:treatment:BF744154BDC55F078993E3E216081DC3" httpUri="http://treatment.plazi.org/id/BF744154BDC55F078993E3E216081DC3" lastPageNumber="169" pageId="0" pageNumber="169">
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<subSubSection id="A1E36B409746DC49A637BBE9C2915461" pageId="0" pageNumber="169" type="nomenclature">
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<paragraph id="0EA8C10CE7C5CEB08E4273D0035B5C12" pageId="0" pageNumber="169">
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<taxonomicName id="71FB77312C6973C131E8BE16DF981CCB" ID-CoL="87TN2" LSID="BF744154-BDC5-5F07-8993-E3E216081DC3" authority="Gray, 1845" authorityName="Gray" authorityYear="1845" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calotes maria" order="Squamata" pageId="0" pageNumber="169" phylum="Chordata" rank="species" species="maria">Calotes maria Gray, 1845</taxonomicName>
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</paragraph>
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<subSubSection id="F740289D5BED9D4158418624D16577C6" pageId="0" pageNumber="169" type="description">
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<paragraph id="8F30FBA0FA78D1AEB8508837EC163C8A" pageId="0" pageNumber="169">
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<figureCitation id="A6DF1C3B0DFAED264E2C7C63BF4736AC" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Morphological comparison among C. jerdoni (A) (NHMUK 1946.8.11.54), C. iadina sp. nov. (B) (CAS 219992), C. yunnanensis (C) (KIZ 014148), and C. medogensis (D) (KIZ 019546), and C. maria (NHMUK 1946.8.11.24), showing characteristics of the gular scales (1) and axillary scales (2). Photos by V. Deepak and Kai Wang." figureDoi="10.3897/vz.74.e109088.figure4" httpUri="https://binary.pensoft.net/fig/997305" pageId="0" pageNumber="169">Figures 4</figureCitation>
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<figureCitation id="4A72271F833E8D0869000778A76128DA" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lectotype of Calotes maria (NHMUK 1946.8.11.24), showing dorsal (A), ventral (B), lateral head (C), ventral head (D), dorsal head (E), and axillary region (F). Photos by V. Deepak." figureDoi="10.3897/vz.74.e109088.figure5" httpUri="https://binary.pensoft.net/fig/997306" pageId="0" pageNumber="169">, 5</figureCitation>
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<figureCitation id="733854B5B7FBC2008A1131396F5D9518" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Non-vouchered individuals of Calotes maria in life, showing an adult (A) from Royal Manas National Park and a juvenile (B) from Garo Hill, Meghalaya. Photos by Abhijit Das." figureDoi="10.3897/vz.74.e109088.figure6" httpUri="https://binary.pensoft.net/fig/997307" pageId="0" pageNumber="169">, 6; Appendix VI</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="525754A27A47E4676C711FE615336A0D" pageId="0" pageNumber="169" type="synonymy">
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<paragraph id="89B40BFE34C21DE544C9119354343796" pageId="0" pageNumber="169">Synonymy.</paragraph>
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<paragraph id="76EE816E25ED35DAE1628DA8DDA6846A" pageId="0" pageNumber="169">
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<taxonomicName id="22A9772D8027EDE9E51067C306AD4ECA" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calotes platyceps" order="Squamata" pageId="0" pageNumber="169" phylum="Chordata" rank="species" species="platyceps">
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<emphasis id="B4AA31672898C36C57B5246DCF93B323" italics="true" pageId="0" pageNumber="169">Calotes platyceps</emphasis>
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</taxonomicName>
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Blyth, 1852: 354
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</paragraph>
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||
</subSubSection>
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<subSubSection id="9709A3AD91CDC3ABD9C8FD232131E3EF" pageId="0" pageNumber="169" type="lectotype">
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<paragraph id="CB56A615CED08B17C2A44210407B5A21" pageId="0" pageNumber="169">Lectotype.</paragraph>
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<paragraph id="94DE3682CB0A0D372BF1E38B7A5C2688" pageId="0" pageNumber="169">
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NHMUK 1946.8.11.24, adult male from
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<normalizedToken id="90667E29B1E012C547984B13270ADF18" originalValue="“Afghanistan”">"Afghanistan"</normalizedToken>
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(see remarks on its type locality below); designated by
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<bibRefCitation id="0E25EBF3CA3F60B52987C912E6427D3D" author="Boulenger, GA" journalOrPublisher="British Museum (Natural History), London" pageId="0" pageNumber="169" refId="B8" refString="Boulenger, GA, 1885. Catalogue of the Lizards in the British Museum (Natural History), Volume I: Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. British Museum (Natural History), London" title="Catalogue of the Lizards in the British Museum (Natural History), Volume I: Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae." year="1885">Boulenger (1885)</bibRefCitation>
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||
.
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</paragraph>
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||
<caption id="387CDFC76BAE1F6F595BEED4B2EEEC5E" doi="10.3897/vz.74.e109088.figure5" httpUri="https://binary.pensoft.net/fig/997306" pageId="0" pageNumber="169" start="Figure 5" startId="F5">
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<paragraph id="7EEBCE93C31DA7F6DA8C8E9E308F57CB" pageId="0" pageNumber="169">
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<emphasis id="43FE0E00BBC077E25BE4D523A8FA3AC1" bold="true" pageId="0" pageNumber="169">Figure 5.</emphasis>
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Lectotype of
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<taxonomicName id="24D785D7B29C37A3142C026DEF2B1E52" authorityName="Gray" authorityYear="1845" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calotes maria" order="Squamata" pageId="0" pageNumber="169" phylum="Chordata" rank="species" species="maria">
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<emphasis id="75C891709E77A9B414989B764C8353F5" italics="true" pageId="0" pageNumber="169">Calotes maria</emphasis>
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</taxonomicName>
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||
(NHMUK 1946.8.11.24), showing dorsal (
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<emphasis id="D39C8A9F92D5C5E205E8F32704D09D45" bold="true" pageId="0" pageNumber="169">A</emphasis>
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), ventral (
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||
<emphasis id="F4AF5D7416ADDFBBD5571696A79149A1" bold="true" pageId="0" pageNumber="169">B</emphasis>
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), lateral head (
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||
<emphasis id="1ED4DFC26D7DFAAB6D0FB1E4E7200D51" bold="true" pageId="0" pageNumber="169">C</emphasis>
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), ventral head (
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<emphasis id="4954100381D1BA72608FB03D528E1821" bold="true" pageId="0" pageNumber="169">D</emphasis>
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), dorsal head (
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<emphasis id="E0361F9AF67DEB8385ACE22E69C4BFFC" bold="true" pageId="0" pageNumber="169">E</emphasis>
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), and axillary region (
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<emphasis id="0E24462608C94B9A56C9B8BDCA3C3694" bold="true" pageId="0" pageNumber="169">F</emphasis>
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). Photos by V. Deepak.
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</paragraph>
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</caption>
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||
</subSubSection>
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<subSubSection id="927F66D2BE15532A7C0B838150FEB9A6" pageId="0" pageNumber="169" type="additional examined specimens">
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<paragraph id="954BDC66F30DAA751665C914E5F17C4D" pageId="0" pageNumber="169">Additional examined specimens.</paragraph>
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<paragraph id="AEFAD02EE09EB6642F81149E46F91D1A" pageId="0" pageNumber="169">
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NHMUK 1870.11.29.30a, adult male; 1870.11.29.30b, adult female; both from Khasia (Khasi Hill), India; NHMUK 1860.3.19.1020, adult female from
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<normalizedToken id="A24703B35D28EB52F4E65DC29E905B50" originalValue="“Afghanistan”">"Afghanistan"</normalizedToken>
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||
.
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||
</paragraph>
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||
<caption id="B9F9EECDCF335EC024CB480B0B041811" doi="10.3897/vz.74.e109088.figure6" httpUri="https://binary.pensoft.net/fig/997307" pageId="0" pageNumber="169" start="Figure 6" startId="F6">
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<paragraph id="662E36FC313D528541E17E52D37779FA" pageId="0" pageNumber="169">
|
||
<emphasis id="EB447EDA2AFC409DC7C122782CA22313" bold="true" pageId="0" pageNumber="169">Figure 6.</emphasis>
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Non-vouchered individuals of
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||
<taxonomicName id="897563DCFBF1BA46E83379BACA3DF2D9" authorityName="Gray" authorityYear="1845" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calotes maria" order="Squamata" pageId="0" pageNumber="169" phylum="Chordata" rank="species" species="maria">
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<emphasis id="9C4DAB2AD6FAD93911B7127E25AE3CCC" italics="true" pageId="0" pageNumber="169">Calotes maria</emphasis>
|
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</taxonomicName>
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in life, showing an adult (
|
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<emphasis id="3CC412A1DD6183F51DADBEAC92620061" bold="true" pageId="0" pageNumber="169">A</emphasis>
|
||
) from Royal Manas National Park and a juvenile (
|
||
<emphasis id="7FC9810E56234EC3AC1AEC74D820819E" bold="true" pageId="0" pageNumber="169">B</emphasis>
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) from Garo Hill, Meghalaya. Photos by Abhijit Das.
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</paragraph>
|
||
</caption>
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||
</subSubSection>
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||
<subSubSection id="92A0C82D8554CC5B9C1CE62E51671BE7" pageId="0" pageNumber="169" type="remarks">
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<paragraph id="1171BC56E7BA3778AFD14D6F4EC2B15D" pageId="0" pageNumber="169">Remarks on its type locality and type specimens.</paragraph>
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||
<paragraph id="CD576D3E6E39FF26291C92D0DB7B5949" pageId="0" pageNumber="169">
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On its original description,
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||
<bibRefCitation id="509393C97531CE9C1A47564D7A25E45E" author="Gray, JE" journalOrPublisher="Trustees of die British Museum / Edward Newman, London" pageId="0" pageNumber="169" refId="B18" refString="Gray, JE, 1845. Catalogue of the specimens of lizards in the collection of the British Museum. Trustees of die British Museum/Edward Newman, London" title="Catalogue of the specimens of lizards in the collection of the British Museum." year="1845">Gray (1845)</bibRefCitation>
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described the species based on four specimens: three individuals (a juvenile, a subadult, and an adult) from "Affghanistan and a single half-grown individual from Khassia Hill". Later
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<bibRefCitation id="880BC47615825990E8B5C74DA678392F" author="Boulenger, GA" journalOrPublisher="British Museum (Natural History), London" pageId="0" pageNumber="169" refId="B8" refString="Boulenger, GA, 1885. Catalogue of the Lizards in the British Museum (Natural History), Volume I: Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. British Museum (Natural History), London" title="Catalogue of the Lizards in the British Museum (Natural History), Volume I: Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae." year="1885">Boulenger (1885)</bibRefCitation>
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designated a male from Affghanistan as the
|
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<normalizedToken id="BA8BF6F2FF240D013EF4CD0DE68DF144" originalValue="“type”">"type"</normalizedToken>
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||
(= lectotype) of
|
||
<taxonomicName id="A4974482DBFAFC4B006B183EC8E8050B" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. maria" pageId="0" pageNumber="169" phylum="Chordata" rank="species" species="maria">
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||
<emphasis id="07D26F2C09306BD2E08D8FA1B4B5DB58" italics="true" pageId="0" pageNumber="169">C. maria</emphasis>
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</taxonomicName>
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(NHMUK 1946.8.11.24), and he identified two of the young male syntypes of
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<taxonomicName id="24DA501A9BB81B330A001DE7CCBFCE44" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. maria" pageId="0" pageNumber="169" phylum="Chordata" rank="species" species="maria">
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<emphasis id="BF7182C4EA2E10CEB6096BF296C86F4E" italics="true" pageId="0" pageNumber="169">C. maria</emphasis>
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</taxonomicName>
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as
|
||
<taxonomicName id="AF70EE21F89C2454FA952B3D2CE3D299" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. jerdoni" pageId="0" pageNumber="169" phylum="Chordata" rank="species" species="jerdoni">
|
||
<emphasis id="1DB6B35087F68BCA7B43F187379552BF" italics="true" pageId="0" pageNumber="169">C. jerdoni</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="C30F96325E6DB65748F05742E52386F9" pageId="0" pageNumber="169">
|
||
Based on
|
||
<bibRefCitation id="18D7886CB06D78241A87B022A5A54DBA" author="Blyth, E" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="169" refId="B6" refString="Blyth, E, 1853. Report of the curator, zoological department. Journal of the Asiatic Society of Bengal: 411-413." title="Report of the curator, zoological department. Journal of the Asiatic Society of Bengal: 411 - 413." year="1853">Blyth (1853)</bibRefCitation>
|
||
, the initial syntypes of
|
||
<taxonomicName id="B9BF5BA6E3362BAFCD54E4ACAE5E11B7" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. maria" pageId="0" pageNumber="169" phylum="Chordata" rank="species" species="maria">
|
||
<emphasis id="6F30D0D91764B8E4EDC9F9B3DEBA2EFE" italics="true" pageId="0" pageNumber="169">C. maria</emphasis>
|
||
</taxonomicName>
|
||
, along with other species of the genus
|
||
<taxonomicName id="2A3E5B25A838384A827426885E5C6C0C" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calotes" order="Squamata" pageId="0" pageNumber="169" phylum="Chordata" rank="genus">
|
||
<emphasis id="5A5A798BD1F7B426D3DC2353CDC79444" italics="true" pageId="0" pageNumber="169">Calotes</emphasis>
|
||
</taxonomicName>
|
||
, were collected by Dr. Griffith and later handled by the East India Company before donating them to the Natural History Museum.
|
||
<bibRefCitation id="5A0C6E856F0488D38317B4BD41369216" author="Blyth, E" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="169" refId="B6" refString="Blyth, E, 1853. Report of the curator, zoological department. Journal of the Asiatic Society of Bengal: 411-413." title="Report of the curator, zoological department. Journal of the Asiatic Society of Bengal: 411 - 413." year="1853">Blyth (1853)</bibRefCitation>
|
||
suspected that those specimens from
|
||
<normalizedToken id="99339A804D6136156F9508ABAAF3A370" originalValue="“Affghanistan”">"Affghanistan"</normalizedToken>
|
||
(= Afghanistan) and "Khassia Hills" (= Khasi Hill) were mixed during the handling by the East India Company. As the tropical species
|
||
<taxonomicName id="D6B25392CD092E42129715E22A7E7165" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. maria" pageId="0" pageNumber="169" phylum="Chordata" rank="species" species="maria">
|
||
<emphasis id="7AD8DEAA64479349DF9A75EA1742D33C" italics="true" pageId="0" pageNumber="169">C. maria</emphasis>
|
||
</taxonomicName>
|
||
certainly cannot survive in desert or cold highlands of Afghanistan, and given Afghanistan is zoogeographically distinct from northeast India, the type locality of
|
||
<taxonomicName id="E32AFA26AFFF7B297D2EC91EEC400429" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. maria" pageId="0" pageNumber="169" phylum="Chordata" rank="species" species="maria">
|
||
<emphasis id="25B24EFFADA97CCDA61A51D40D23F3D6" italics="true" pageId="0" pageNumber="169">C. maria</emphasis>
|
||
</taxonomicName>
|
||
was certainly mistaken as Afghanistan and should be replaced by "Khassia Hills", which refers to the Khasi Hill in Meghalaya State, Northeast India.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="771CC28F4FA2E2BB975AE3045FEAF17B" pageId="0" pageNumber="169" type="remarks">
|
||
<paragraph id="F622F480EDB6B18677A0A18B30C66C0C" pageId="0" pageNumber="169">
|
||
Remarks on the
|
||
<normalizedToken id="945E976B523E19F4B766C367AD70B436" originalValue="“synonym”">"synonym"</normalizedToken>
|
||
<taxonomicName id="C39C45800DFC8484D4A4C2E4E16E0B11" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. platyceps" pageId="0" pageNumber="169" phylum="Chordata" rank="species" species="platyceps">
|
||
<emphasis id="763D9A1EC8619412BE1FE9DC9579AC46" italics="true" pageId="0" pageNumber="169">C. platyceps</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="FDDA637C20DFCCD2A07D999DD25823D9" pageId="0" pageNumber="169">
|
||
First described in 1852,
|
||
<taxonomicName id="13DC0FE485CAB437A11024F9BB174F22" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. platyceps" pageId="0" pageNumber="169" phylum="Chordata" rank="species" species="platyceps">
|
||
<emphasis id="32452F22EE99BCDD26A4BBC0703898D2" italics="true" pageId="0" pageNumber="169">C. platyceps</emphasis>
|
||
</taxonomicName>
|
||
was described in a short, rather simple description without a clear statement regarding its type specimen. Later, based on specimens collected from Khasi hills, Jerdon (1870) identified his specimens as
|
||
<taxonomicName id="3961DAC02AAC5CE83C2FEE191B0C9346" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. platyceps" pageId="0" pageNumber="169" phylum="Chordata" rank="species" species="platyceps">
|
||
<emphasis id="DA57E88B603786DD79656A19D0FB3BEB" italics="true" pageId="0" pageNumber="169">C. platyceps</emphasis>
|
||
</taxonomicName>
|
||
and added a diagnosis of
|
||
<taxonomicName id="39BAD717A610F87B21891689A7CD882B" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. platyceps" pageId="0" pageNumber="169" phylum="Chordata" rank="species" species="platyceps">
|
||
<emphasis id="F8B38FCFEFBE83AE69EDD6250A3C0369" italics="true" pageId="0" pageNumber="169">C. platyceps</emphasis>
|
||
</taxonomicName>
|
||
against
|
||
<taxonomicName id="528FE2367CB246A6B234F6AA6AB638E9" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. maria" pageId="0" pageNumber="169" phylum="Chordata" rank="species" species="maria">
|
||
<emphasis id="802907539245A7A995C658C415A1202B" italics="true" pageId="0" pageNumber="169">C. maria</emphasis>
|
||
</taxonomicName>
|
||
, including a smaller body size, larger and fewer gular and body scales, and a narrower distance between supratympanic crest and the upper edge of tympanum. While the diagnosis of "
|
||
<taxonomicName id="085D66E20B054745789C1E742DF5C668" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. platyceps" pageId="0" pageNumber="169" phylum="Chordata" rank="species" species="platyceps">
|
||
<emphasis id="17AB1D1BDB933BAA0A54AD64BC9061D8" italics="true" pageId="0" pageNumber="169">C. platyceps</emphasis>
|
||
</taxonomicName>
|
||
" described by Jerdon (1870) matches the diagnosis of
|
||
<taxonomicName id="19EA70E5D2F9A1EEA115D79CA46F6E9C" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. jerdoni" pageId="0" pageNumber="169" phylum="Chordata" rank="species" species="jerdoni">
|
||
<emphasis id="19791F139D9F936DC52540F16DD4B8DF" italics="true" pageId="0" pageNumber="169">C. jerdoni</emphasis>
|
||
</taxonomicName>
|
||
, they were not based on the type specimen of
|
||
<taxonomicName id="14B24D92D2B8E294236904BF2A9BB8BE" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. platyceps" pageId="0" pageNumber="169" phylum="Chordata" rank="species" species="platyceps">
|
||
<emphasis id="135A5C5B7114F8EAC5413DBD8C3D3D6C" italics="true" pageId="0" pageNumber="169">C. platyceps</emphasis>
|
||
</taxonomicName>
|
||
. Subsequently,
|
||
<bibRefCitation id="47C8138FA6D34CAA2D80D469B6433B64" author="Boulenger, GA" journalOrPublisher="British Museum (Natural History), London" pageId="0" pageNumber="169" refId="B8" refString="Boulenger, GA, 1885. Catalogue of the Lizards in the British Museum (Natural History), Volume I: Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. British Museum (Natural History), London" title="Catalogue of the Lizards in the British Museum (Natural History), Volume I: Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae." year="1885">Boulenger (1885)</bibRefCitation>
|
||
without explanation, considered
|
||
<taxonomicName id="957A63B8A2A1691520325C245F5460A8" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. platyceps" pageId="0" pageNumber="169" phylum="Chordata" rank="species" species="platyceps">
|
||
<emphasis id="0A5A395664690B181D6EDD8DA06A4686" italics="true" pageId="0" pageNumber="169">C. platyceps</emphasis>
|
||
</taxonomicName>
|
||
a junior synonym of both
|
||
<taxonomicName id="7EB7A56CD25B671598260A0D471BFA15" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. maria" pageId="0" pageNumber="169" phylum="Chordata" rank="species" species="maria">
|
||
<emphasis id="E7279A3F26F5F9EADE9507F2DC29CA67" italics="true" pageId="0" pageNumber="169">C. maria</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="55B08C24CE531882161DF3739417EE29" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. jerdoni" pageId="0" pageNumber="169" phylum="Chordata" rank="species" species="jerdoni">
|
||
<emphasis id="E96847846FC4259E24808E40365FA2C5" italics="true" pageId="0" pageNumber="169">C. jerdoni</emphasis>
|
||
</taxonomicName>
|
||
-a taxonomic treatment followed by subsequent authors (
|
||
<bibRefCitation id="B9F05E14C5862D8A08913258AEB09A33" author="Boulenger, GA" journalOrPublisher="Taylor & Francis, London" pageId="0" pageNumber="169" refId="B9" refString="Boulenger, GA, 1890. The Fauna of British India, Including Ceylon and Burma. Reptilia and Batrachia. Taylor & Francis, London" title="The Fauna of British India, Including Ceylon and Burma. Reptilia and Batrachia." year="1890">Boulenger 1890</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="B0356DE6B0F65FDF10F514F0DF943352" author="Smith, MA" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="169" refId="B48" refString="Smith, MA, 1935. The fauna of British India, including Ceylon and Burma. Reptiles and Amphibia, Vol. II. Sauria. Taylor and Francis, London" title="The fauna of British India, including Ceylon and Burma. Reptiles and Amphibia, Vol. II. Sauria." year="1935">Smith 1935</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="29149F9D7DDE3EF6C0603B5A1B616529" author="Che, J" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="169" refId="B11" refString="Che, J, Jiang, K, Yan, F, Zhang, YP, 2020. Amphibians and Reptiles in Tibet--Diversity and Evolution. Science Press, Beijing" title="Amphibians and Reptiles in Tibet- - Diversity and Evolution." year="2020">Che et al. 2020</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="E07539EA1DF182E493C4D283BB3D3451" author="Uetz, P" journalOrPublisher="Turkish Journal of Zoology" pageId="0" pageNumber="169" publicationUrl="http://www.reptile-database.org" refId="B56" refString="Uetz, P, Freed, P, Aguilar, R, Reyes, F, Hosek, J, 2023. The Reptile Database., http://www.reptile-database.org [ [accessed 11 June 2023] ]" title="The Reptile Database" url="http://www.reptile-database.org" year="2023">Uetz et al. 2023</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="A5812219E3EEBF5B1E799DD729C44CAB" pageId="0" pageNumber="169">
|
||
Although the presence of a supratympanic crest on each side of the head aligns
|
||
<taxonomicName id="52E4DC70EB2C646B351DAA81091782BD" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. platyceps" pageId="0" pageNumber="169" phylum="Chordata" rank="species" species="platyceps">
|
||
<emphasis id="928E0D62070CCFB8182A746D92430F7A" italics="true" pageId="0" pageNumber="169">C. platyceps</emphasis>
|
||
</taxonomicName>
|
||
with the
|
||
<taxonomicName id="35A97A6DCDF65F25D090EBB0D14BFDF0" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. jerdoni" pageId="0" pageNumber="169" phylum="Chordata" rank="species" species="jerdoni">
|
||
<emphasis id="662D8FBBF2BF5E7B43AB6987F555EF84" italics="true" pageId="0" pageNumber="169">C. jerdoni</emphasis>
|
||
</taxonomicName>
|
||
complex, based on the limited information in the original morphological description, it is impossible to confirm whether
|
||
<taxonomicName id="D6E8FEE8ECFFF2050A8A17DDCAF25AEC" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. platyceps" pageId="0" pageNumber="169" phylum="Chordata" rank="species" species="platyceps">
|
||
<emphasis id="315D1F50F79D0F52F4AA62FE7CA083C4" italics="true" pageId="0" pageNumber="169">C. platyceps</emphasis>
|
||
</taxonomicName>
|
||
represents a junior synonym of
|
||
<taxonomicName id="4505CA3A001847423A0E7CAC4775FCED" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. maria" pageId="0" pageNumber="169" phylum="Chordata" rank="species" species="maria">
|
||
<emphasis id="8FF6BB8AEF941000676CA91AC9C2A17A" italics="true" pageId="0" pageNumber="169">C. maria</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="9539A3C54C60C48340609F167117E1FC" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. jerdoni" pageId="0" pageNumber="169" phylum="Chordata" rank="species" species="jerdoni">
|
||
<emphasis id="ED10B5A1A8FCCBB6961E9D1BE83F694C" italics="true" pageId="0" pageNumber="169">C. jerdoni</emphasis>
|
||
</taxonomicName>
|
||
. As the type specimen of
|
||
<taxonomicName id="C28113028985F3F7251CDB170256C3A8" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. platyceps" pageId="0" pageNumber="169" phylum="Chordata" rank="species" species="platyceps">
|
||
<emphasis id="7DB15A6BAD9472F0ECA7F59D42A932C0" italics="true" pageId="0" pageNumber="169">C. platyceps</emphasis>
|
||
</taxonomicName>
|
||
cannot be located, and since the original description of the species provides little useful information on its taxonomic identity, we treat the name
|
||
<taxonomicName id="ED94058A900C026F23452055564D1CC0" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. platyceps" pageId="0" pageNumber="169" phylum="Chordata" rank="species" species="platyceps">
|
||
<emphasis id="7C854921B28778504D787806D0BBBCB7" italics="true" pageId="0" pageNumber="169">C. platyceps</emphasis>
|
||
</taxonomicName>
|
||
as a nomen dubium. As such, the name is nomenclaturally still available, but it has no bearing on the taxonomic and nomenclatural decisions made here.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="268CF72D316A8274E760AEEEFE8322E2" pageId="0" pageNumber="169" type="diagnosis">
|
||
<paragraph id="7DD04221508A527BA7584BCB5218940D" pageId="0" pageNumber="169">Diagnosis.</paragraph>
|
||
<paragraph id="65BE0E1B24012E54ADD2A3547409E5BF" pageId="0" pageNumber="169">
|
||
<taxonomicName id="F8CAE0F2AA4152D65C5A1D8BCC7B1844" authorityName="Gray" authorityYear="1845" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calotes maria" order="Squamata" pageId="0" pageNumber="169" phylum="Chordata" rank="species" species="maria">
|
||
<emphasis id="0E2A198E586CB4D8C27BA19BCCB4698A" italics="true" pageId="0" pageNumber="169">Calotes maria</emphasis>
|
||
</taxonomicName>
|
||
can be diagnosed from its congeners by a combination of following morphological characters: (1) body size large, SVL 101-117.5 mm; (2) tail slender, long, TAL 318.3-323.9% SVL; (3) posterior-most conical scales of parallel ridges of head distinctively elongated into spine shape, about same length as longest nuchal crest; (4) inferior row of conical/spinous scales 4 scales away from superior tympanum; (5) dorsal head scales posterolateral of parietal bearing longitudinal keels; (6) mental smaller than first pair of chin shields; (7) gular scales count 27-31, homogeneous in posteromedial region, much larger than ventrals, mucronate with distinct, moderate-lengthed tips; (8) body scales small, ABR 58-67; (9) distinct gular pouch absent, transverse gular fold absent; (10) shoulder fold absent; (11) neck scales oriented posterosuperiorly; (12) axillary scales oriented posterosuperiorly at an angle less than 60°; (13) nuchal crests long, elongated, lanceolate-shaped, differentiated from dorsal crest, TNC 17.6-24.0% HL; (14) middorsal crest scale count 47-52; (15) ventral scale count 63-76, (17) F4S 23-28, T4S 28-32; (18) dorsal background coloration
|
||
<normalizedToken id="B5CD8E6E26806015FE09FF60102619C6" originalValue="Pratt’s">Pratt's</normalizedToken>
|
||
<normalizedToken id="CA94A38984A224FD468111D333F4556B" originalValue="Payne’s">Payne's</normalizedToken>
|
||
Gray (Color 293) to Sky Blue (Color 192) under long-term preservation, ventral Paris White (Color 139); and (19) short, Pale Pinkish Buff (Color 3) to Yellow Ocher (Color 14), oblique stripes present on dorsum, similar colored patches present on tail base.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="910E32DF3323AF585A14E77201D0D003" pageId="0" pageNumber="169" type="redescription">
|
||
<paragraph id="AF00E7960E8221A0B9677018A8B4E6A6" pageId="0" pageNumber="169">Redescription of the lectotype.</paragraph>
|
||
<paragraph id="B9343F9304C353DB75F6D0F7EEDA97AE" pageId="0" pageNumber="169">
|
||
Large sized
|
||
<taxonomicName id="F0A5337D6D7EC94650BC00E65C67DB54" class="Reptilia" family="Agamidae" genus="Calotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calotes" order="Squamata" pageId="0" pageNumber="169" phylum="Chordata" rank="genus">
|
||
<emphasis id="9EF20FF1B5899BFE11F89FAF98FE8EE0" italics="true" pageId="0" pageNumber="169">Calotes</emphasis>
|
||
</taxonomicName>
|
||
, SVL 101.0 mm; body not compressed dorsally, lachrymiform in cross-section; tail swollen at base, long, TAL 322 mm, 318.8% SVL. Limbs slender, forelimb 52.7 mm, 52.2% SVL; hindlimb 69.8 mm, 69.1% SVL. Rostral rectangular, bordering four small scales excluding supralabials; supralabials 10/11, feebly keeled posteriorly; nasal elongated oval, bordering second supralabial, 6/7 scales away from orbit circle; loreals distinctively keeled, each bearing single keel; canthal ridge distinct, canthus rostralis scales 10/10, elongated except anterior most two, supraciliaries only slightly overlapping, overlapping length less than half of scale length; suborbital scale rows 3/3, about equal size, each scale bearing single, distinct, inferior-located lateral keel. Distinctively protruding, enlarged keeled scales each bearing multiple keels aligned in two parallel lateral ridges: superior row continuing from posterior most supraciliary to superior squamosal head, consisting 8 raised scales, anterior-most 2 and 4 scales conical on left and right, respectively, remaining 6 and 4 much more elongated into spikes; inferior row two small scales away from posterior mid orbit to inferior squamosal head, consisting 12/10 scales, anterior-most 5 sub-pyramidal or conical, remaining posterior scales elongated into spikes. Inferior one of parallel ridges four small scale rows away from anterior tympanum. Tympanum exposed, round, TD 10.5% HL; no post orbital spikes, posterosuperior tympanic and temporal spikes present, which are located on parallel ridges. Dorsal head scales heterogeneous in size and shape, slightly larger above eyes, distinctively smaller on temporal, moderately keeled, each bearing single keel; enlarged scales arranged in Y-shape three scales posterior to rostral along longitudinal midline, with posterior most scale on stem of
|
||
<normalizedToken id="80E502B0D1C89D3E95485E72DCAF3297" originalValue="“Y”">"Y"</normalizedToken>
|
||
figure largest; 17 scales transversely across dorsal head between and excluding supraciliary at widest point; parietal scale elongated, narrow, parietal eye present; scales posterolateral of parietal bearing longitudinal keels.
|
||
</paragraph>
|
||
<paragraph id="0AEA0D1F016628E9AD7B44471E749402" pageId="0" pageNumber="169">Except scales anterior to shoulder, all dorsal body scales broad triangular shaped, regularly arranged, imbricate, mostly homogeneous, weakly keeled; neck scales oriented posterosuperiorly; axillary scales oriented posterosuperiorly; remaining lateral and superior body scales oriented posterosuperiorly, inferolateral body scales more distinctively keeled, oriented straight posteriorly or inferoposteriorly; midbody scale row around body 67. Nuchal crest 13, distinctively elongated into spikes, much taller than remaining dorsal crests, tallest one 5.6 mm in height, 19.5% HL; dorsal crests much lower, curving posteriorly; mid-dorsal crest scale count 52 anterior to cloaca; shoulder fold absent. Scales of dorsal limbs more or less homogeneous, keeled. Dorsal tail scales homogeneous, feebly keeled or smooth anteriorly, more distinctively keeled posteriorly.</paragraph>
|
||
<paragraph id="23F3AFF8F67D2DFE6C9212BA7729E4CF" pageId="0" pageNumber="169">Mental spear-shaped, enclosed by and smaller than first pair of chin shields; chin shields feebly keeled, larger than nearby gulars, 5/6, anterior most 3/2 in contact with infralabials on each side, remaining ones separated by one or two rows of small scales; infralabials 10/10, more distinctively keeled than supralabials; gular scales all distinctively keeled; anterior gulars smaller, juxtaposed, various in shape and size; remaining posterior gular scales larger, similar size to dorsals, rhomboid shaped, imbricated, mucronate with moderate-length tip on posterior end of each gular; post mental gular scales 30 along medial line. Ventral body cut open from chest to pelvis, ventral scales imbricated, slightly smaller than larger gulars, distinctively keeled, carinated, ventral scale count 76 along medial axis from neck to cloaca. Ventral limb scales homogeneous in size, more distinctively keeled than dorsal limb scales; subdigital lamellae 27/28 under finger IV, 32/32 under toe IV. Tail scales distinctively keeled, carinated (ventral tail cut open at base).</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |