treatments-xml/data/BD/CE/6B/BDCE6B4D4C1A5EED95E15FEC2B8DB383.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.997.54262" ID-GBIF-Dataset="e36130a3-a361-4d38-a1b3-5dcbbf8ad6f8" ID-PMC="PMC7710693" ID-Pensoft-Pub="1313-2970-997-47" ID-Pensoft-UUID="ADBB6106EE0C532E9AE1C4BAA5E4371F" ID-PubMed="33335440" ID-ZooBank="FFF388DB25544773ACB027189DCCACC9" ModsDocID="1313-2970-997-47" checkinTime="1606399186851" checkinUser="pensoft" docAuthor="Quintero-Gutierrez, Edwin Javier, Sandmann, Dorothee, Combita-Heredia, Orlando, Klarner, Bernhard, Widyastuti, Rahayu &amp; Scheu, Stefan" docDate="2020" docId="BDCE6B4D4C1A5EED95E15FEC2B8DB383" docLanguage="en" docName="ZooKeys 997: 47-68" docOrigin="ZooKeys 997" docSource="http://dx.doi.org/10.3897/zookeys.997.54262" docTitle="Krantzolaspina angustatus Quintero-Gutiérrez &amp; Sandmann &amp; Cómbita-Heredia &amp; Klarner &amp; Widyastuti &amp; Scheu 2020, comb. nov." docType="treatment" docVersion="4" id="ADBB6106EE0C532E9AE1C4BAA5E4371F" lastPageNumber="47" masterDocId="ADBB6106EE0C532E9AE1C4BAA5E4371F" masterDocTitle="Review of the mite genus Krantzolaspina Datta &amp; Bhattacharjee (Mesostigmata, Parholaspididae) with re-description of K. angustatus comb. nov. (Ishikawa) from Indonesia" masterLastPageNumber="68" masterPageNumber="47" pageNumber="47" updateTime="1668169969321" updateUser="ExternalLinkService">
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<mods:title>Review of the mite genus Krantzolaspina Datta &amp; Bhattacharjee (Mesostigmata, Parholaspididae) with re-description of K. angustatus comb. nov. (Ishikawa) from Indonesia</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Quintero-Gutierrez, Edwin Javier</mods:namePart>
<mods:affiliation>Instituto Colombiano Agropecuario (ICA), Subgerencia de Analisis y Diagnostico, Grupo Red de Analisis y Diagnostico Fitosanitario, Laboratorio de Analisis y Diagnostico Fitosanitario de Manizales, Caldas, Carrera 30 # 65 - 15, Manizales, Colombia</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-7906-1962</mods:nameIdentifier>
<mods:nameIdentifier type="email">ejquinterog@gmail.com</mods:nameIdentifier>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sandmann, Dorothee</mods:namePart>
<mods:affiliation>University of Goettingen, J. F. Blumenbach Institute of Zoology and Anthropology, Untere Karspuele 2, 37073 Goettingen, Germany</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-7003-6952</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Combita-Heredia, Orlando</mods:namePart>
<mods:affiliation>Acarology Laboratory, Ohio State University, 1315 Kinnear Rd., Columbus, OH 43212, USA &amp; Centro de Inversitgacion en Acarologia CIACARI Bogota, Colombia</mods:affiliation>
</mods:name>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Klarner, Bernhard</mods:namePart>
<mods:affiliation>University of Goettingen, J. F. Blumenbach Institute of Zoology and Anthropology, Untere Karspuele 2, 37073 Goettingen, Germany</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-1867-4911</mods:nameIdentifier>
</mods:name>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Widyastuti, Rahayu</mods:namePart>
<mods:affiliation>Institut Pertanian Bogor, Department of Soil Science and Land Resources, Damarga Campus, Bogor 16680, Indonesia</mods:affiliation>
</mods:name>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Scheu, Stefan</mods:namePart>
<mods:affiliation>University of Goettingen, J. F. Blumenbach Institute of Zoology and Anthropology, Untere Karspuele 2, 37073 Goettingen, Germany &amp; University of Goettingen, Centre of Biodiversity and Sustainable Land Use, Buesgenweg 1, 37077 Goettingen, Germany</mods:affiliation>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>997</mods:number>
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<mods:start>47</mods:start>
<mods:end>68</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.997.54262</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.997.54262</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-997-47</mods:identifier>
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<treatment ID-GBIF-Taxon="170531164" LSID="urn:lsid:plazi:treatment:BDCE6B4D4C1A5EED95E15FEC2B8DB383" httpUri="http://treatment.plazi.org/id/BDCE6B4D4C1A5EED95E15FEC2B8DB383" lastPageNumber="47" pageId="0" pageNumber="47">
<subSubSection pageId="0" pageNumber="47" type="nomenclature">
<paragraph pageId="0" pageNumber="47">
<taxonomicName LSID="BDCE6B4D-4C1A-5EED-95E1-5FEC2B8DB383" authority="(Ishikawa, 1987)" authorityName="Quintero-Gutiérrez &amp; Sandmann &amp; Cómbita-Heredia &amp; Klarner &amp; Widyastuti &amp; Scheu" authorityYear="2020" baseAuthorityName="Ishikawa" baseAuthorityYear="1987" class="Arachnida" family="Parholaspididae" genus="Krantzolaspina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Krantzolaspina angustatus" order="Mesostigmata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="angustatus" status="comb. nov.">Krantzolaspina angustatus (Ishikawa, 1987)</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="47">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="reference_group">
<paragraph pageId="0" pageNumber="47">
<taxonomicName authorityName="Ishikawa" authorityYear="1987" class="Arachnida" family="Parholaspididae" genus="Proparholaspulus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Proparholaspulus angustatus" order="Mesostigmata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="angustatus">Proparholaspulus angustatus</taxonomicName>
- Ishikawa, 1987a: 82; 
<bibRefCitation author="Kadkhodae, FE" journalOrPublisher="Journal of Crop Protection" pageId="0" pageNumber="47" pagination="127 - 138" refId="B41" refString="Kadkhodae, FE, Nemati, A, Kocheili, F, 2013. Some mesostigmatid mites from Iran with their world distribution. Journal of Crop Protection 2: 127 - 138" title="Some mesostigmatid mites from Iran with their world distribution." volume="2" year="2013">Kadkhodae et al., 2013</bibRefCitation>
: 131.
</paragraph>
<paragraph pageId="0" pageNumber="47">
<taxonomicName authorityName="Hajizadeh, Mortazavi, Balooch-Shahriari &amp; Castilho" authorityYear="2017" class="Arachnida" family="Parholaspididae" genus="Indutolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Indutolaelaps jiroftensis" order="Mesostigmata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="jiroftensis">Indutolaelaps jiroftensis</taxonomicName>
<bibRefCitation author="Hajizadeh, J" journalOrPublisher="Linzer" pageId="0" pageNumber="47" pagination="669 - 676" refId="B24" refString="Hajizadeh, J, Mortazavi, S, Balloch-Shahriari, N, Castilho, RC, 2017. A new species of the genus Indutolaelaps (Mesostigmata: Leptolaelapidae) from Iran. Linzer 49: 669 - 676" title="A new species of the genus Indutolaelaps (Mesostigmata: Leptolaelapidae) from Iran." volume="49" year="2017">Hajizadeh et al., 2017</bibRefCitation>
<emphasis bold="true" pageId="0" pageNumber="47">syn. nov.</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="diagnosis">
<paragraph pageId="0" pageNumber="47">Diagnosis</paragraph>
<paragraph pageId="0" pageNumber="47">
<emphasis bold="true" pageId="0" pageNumber="47">
. 
<emphasis italics="true" pageId="0" pageNumber="47">Female.</emphasis>
</emphasis>
Dorsal shield entire, mostly reticulate (except the anteromedial region which is smooth), partially covering the idiosoma, broad anteriorly and posteriorly narrowing; shield bearing 32 pairs of setae, most setae moderately long and smooth, except 
<emphasis italics="true" pageId="0" pageNumber="47">j1</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">2</emphasis>
and 
<emphasis italics="true" pageId="0" pageNumber="47">z1</emphasis>
which are slightly shorter and setae 
<emphasis italics="true" pageId="0" pageNumber="47">S5</emphasis>
and 
<emphasis italics="true" pageId="0" pageNumber="47">Z5</emphasis>
slightly pilose. Unsclerotised lateral cuticle of the idiosoma with a total of 15 pairs of smooth setae of similar length, except five pairs of 
<emphasis italics="true" pageId="0" pageNumber="47">UR</emphasis>
setae slightly pilose, the pairs of setae 
<emphasis italics="true" pageId="0" pageNumber="47">r6</emphasis>
and 
<emphasis italics="true" pageId="0" pageNumber="47">R1</emphasis>
slightly shorter than 
<emphasis italics="true" pageId="0" pageNumber="47">r5</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">7</emphasis>
plus seven pairs of 
<emphasis italics="true" pageId="0" pageNumber="47">UR</emphasis>
setae. In the idiosoma ventre, all setae aciculate and smooth, except 
<emphasis italics="true" pageId="0" pageNumber="47">JV4</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">5</emphasis>
and 
<emphasis italics="true" pageId="0" pageNumber="47">ZV4</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">5</emphasis>
which are pilose. Presternal area with two pairs of free and presternal platelets. Sternometasternal shield mostly reticulate and covered by semi-rounded cells in the lateral margins, bearing four pairs of setae. Peritrematal shield anteriorly free, fused to the sternometasternal shield between coxae I-III, posteriorly fused with the parapodal shield and extended beyond posterior margin of the coxae IV, region of peritrematal + parapodal shield reticulate and covered by some semi-rounded cells; peritreme extending between coxae I-II at level of 
<emphasis italics="true" pageId="0" pageNumber="47">st1</emphasis>
. Genitiventrianal shield longer than wide, reticulate, flask-shaped and bearing four pairs of setae 
<emphasis italics="true" pageId="0" pageNumber="47">st5</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="47">JV1-3</emphasis>
, as well as three circumanal setae 
<emphasis italics="true" pageId="0" pageNumber="47">pa</emphasis>
and 
<emphasis italics="true" pageId="0" pageNumber="47">po.</emphasis>
Soft opisthogastric ventral cuticle with six pairs of setae 
<emphasis italics="true" pageId="0" pageNumber="47">JV4</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">5</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="47">ZV2</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">5</emphasis>
. Metapodal platelets free, small and elliptical in shape. Deutosternal groove with six transverse rows, surface of the gnathosoma with pointed spine-like process similar to the ventral surface of the palp trochanter. Epistome with median projection bifurcate or trifurcate distally, lateral edges finely and irregularly serrate. Cheliceral digit movable and fixed with four and two teeth, respectively, base of movable digit with a plumose arthrodial brush and a narrow fringed arthrodial corona. Palp tarsal claw three-tined. Pretarsus I reduced or absent and pretarsi II-IV with pretarsi well-developed.
</paragraph>
<paragraph pageId="0" pageNumber="47">
<emphasis bold="true" pageId="0" pageNumber="47">
<emphasis italics="true" pageId="0" pageNumber="47">Male</emphasis>
.
</emphasis>
Unknown.
</paragraph>
<paragraph pageId="0" pageNumber="47">
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">Female</emphasis>
(Figs 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Krantzolaspina angustatus comb. nov., adult female. Dorsal idiosoma." figureDoi="10.3897/zookeys.997.54262.figure1" httpUri="https://binary.pensoft.net/fig/479381" pageId="0" pageNumber="47">1</figureCitation>
- 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Krantzolaspina angustatus comb. nov., adult female. A-D Legs I-IV, respectively. Coxae I-IV in the Fig. 2." figureDoi="10.3897/zookeys.997.54262.figure6" httpUri="https://binary.pensoft.net/fig/479386" pageId="0" pageNumber="47">6</figureCitation>
) (
<emphasis italics="true" pageId="0" pageNumber="47">n = 6</emphasis>
). 
<emphasis italics="true" pageId="0" pageNumber="47">Idiosomal dorsum</emphasis>
(Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Krantzolaspina angustatus comb. nov., adult female. Dorsal idiosoma." figureDoi="10.3897/zookeys.997.54262.figure1" httpUri="https://binary.pensoft.net/fig/479381" pageId="0" pageNumber="47">1</figureCitation>
). 541-611 
<normalizedToken originalValue="μm">μm</normalizedToken>
long, 287-372 
<normalizedToken originalValue="μm">μm</normalizedToken>
wide. 
<emphasis bold="true" pageId="0" pageNumber="47">Dorsal shield</emphasis>
514-536 
<normalizedToken originalValue="μm">μm</normalizedToken>
long, 219-239 
<normalizedToken originalValue="μm">μm</normalizedToken>
wide (at level between 
<emphasis italics="true" pageId="0" pageNumber="47">r2</emphasis>
-3), entire and broad anteriorly with the anterior margin slightly straight and posteriorly narrowing, covering partially the idiosoma with the lateral and posterior margins of soft integument broadly exposed, most surface of the shield reticulate, but smooth on the dorsocentral region between the bases of 
<emphasis italics="true" pageId="0" pageNumber="47">z1</emphasis>
and 
<emphasis italics="true" pageId="0" pageNumber="47">J2</emphasis>
. Dorsal shield bearing 32 pairs of setae of similar length and shape, most setae relatively long (36-46 
<normalizedToken originalValue="μm">μm</normalizedToken>
) and smooth, except 
<emphasis italics="true" pageId="0" pageNumber="47">j1</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">2</emphasis>
and 
<emphasis italics="true" pageId="0" pageNumber="47">z1</emphasis>
slightly shorter (31-35 
<normalizedToken originalValue="μm">μm</normalizedToken>
) and the setae 
<emphasis italics="true" pageId="0" pageNumber="47">S5</emphasis>
and 
<emphasis italics="true" pageId="0" pageNumber="47">Z5</emphasis>
slightly pilose (Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Krantzolaspina angustatus comb. nov., adult female. Dorsal idiosoma." figureDoi="10.3897/zookeys.997.54262.figure1" httpUri="https://binary.pensoft.net/fig/479381" pageId="0" pageNumber="47">1</figureCitation>
). 
<emphasis bold="true" pageId="0" pageNumber="47">Podonotal region</emphasis>
with 17 pairs of setae 
<emphasis italics="true" pageId="0" pageNumber="47">j1</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">6</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="47">z1</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">2</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="47">z4</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">6</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="47">s2</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="47">s6</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="47">r2</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">5</emphasis>
and 10 pairs of poroids, including three pairs of glands 
<emphasis italics="true" pageId="0" pageNumber="47">gd1</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">2</emphasis>
and 
<emphasis italics="true" pageId="0" pageNumber="47">gd4</emphasis>
. 
<emphasis bold="true" pageId="0" pageNumber="47">Opisthonotal region</emphasis>
with 15 pairs of setae 
<emphasis italics="true" pageId="0" pageNumber="47">J1</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">5</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="47">Z1</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">5</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="47">S1-5</emphasis>
and 10 pairs of poroids, including two glands 
<emphasis italics="true" pageId="0" pageNumber="47">gd8</emphasis>
and 
<emphasis italics="true" pageId="0" pageNumber="47">gd9</emphasis>
. 
<emphasis bold="true" pageId="0" pageNumber="47">Unsclerotised lateral cuticle</emphasis>
bearing a total of 15 pairs of smooth setae of similar length (38-44 
<normalizedToken originalValue="μm">μm</normalizedToken>
) except five pairs of 
<emphasis italics="true" pageId="0" pageNumber="47">UR</emphasis>
setae slightly pilose (Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Krantzolaspina angustatus comb. nov., adult female. Dorsal idiosoma." figureDoi="10.3897/zookeys.997.54262.figure1" httpUri="https://binary.pensoft.net/fig/479381" pageId="0" pageNumber="47">1</figureCitation>
); the pairs of setae 
<emphasis italics="true" pageId="0" pageNumber="47">r6</emphasis>
and 
<emphasis italics="true" pageId="0" pageNumber="47">R1</emphasis>
slightly shorter than 
<emphasis italics="true" pageId="0" pageNumber="47">r5</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">7</emphasis>
and the six pairs of 
<emphasis italics="true" pageId="0" pageNumber="47">UR</emphasis>
setae (two of them on the ventral side); a pair of lyrifissures (
<emphasis italics="true" pageId="0" pageNumber="47">Rp</emphasis>
) present between 
<emphasis italics="true" pageId="0" pageNumber="47">R3</emphasis>
and 
<emphasis italics="true" pageId="0" pageNumber="47">R4.</emphasis>
</paragraph>
<caption doi="10.3897/zookeys.997.54262.figure1" httpUri="https://binary.pensoft.net/fig/479381" pageId="0" pageNumber="47" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="47">
<emphasis bold="true" pageId="0" pageNumber="47">Figure 1.</emphasis>
<taxonomicName authorityName="Quintero-Gutiérrez &amp; Sandmann &amp; Cómbita-Heredia &amp; Klarner &amp; Widyastuti &amp; Scheu" authorityYear="2020" baseAuthorityName="Ishikawa" baseAuthorityYear="1987" class="Arachnida" family="Parholaspididae" genus="Krantzolaspina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Krantzolaspina angustatus" order="Mesostigmata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="angustatus">
<emphasis italics="true" pageId="0" pageNumber="47">Krantzolaspina angustatus</emphasis>
</taxonomicName>
comb. nov., adult female. Dorsal idiosoma.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="47">
<emphasis italics="true" pageId="0" pageNumber="47">Idiosomal venter</emphasis>
(Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Krantzolaspina angustatus comb. nov., adult female. Ventral idiosoma." figureDoi="10.3897/zookeys.997.54262.figure2" httpUri="https://binary.pensoft.net/fig/479382" pageId="0" pageNumber="47">2</figureCitation>
, 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Krantzolaspina angustatus comb. nov., adult female. Ventral idiosoma showing the sternometasternal, genitiventrianal and parapodal shield. A Holotype of Krantzolaspina angustatus B Paratype of Indutolaelaps jiroftensis syn. nov., photos by Raphael Castilho C New material from Indonesia." figureDoi="10.3897/zookeys.997.54262.figure3" httpUri="https://binary.pensoft.net/fig/479383" pageId="0" pageNumber="47">3A-C</figureCitation>
). All setae aciculate and smooth, except 
<emphasis italics="true" pageId="0" pageNumber="47">Jv4</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">5</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="47">Zv4</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">5</emphasis>
which are pilose (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Krantzolaspina angustatus comb. nov., adult female. Ventral idiosoma." figureDoi="10.3897/zookeys.997.54262.figure2" httpUri="https://binary.pensoft.net/fig/479382" pageId="0" pageNumber="47">2</figureCitation>
). 
<emphasis bold="true" pageId="0" pageNumber="47">Tritosternum</emphasis>
(Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Krantzolaspina angustatus comb. nov., adult female. Ventral idiosoma." figureDoi="10.3897/zookeys.997.54262.figure2" httpUri="https://binary.pensoft.net/fig/479382" pageId="0" pageNumber="47">2</figureCitation>
) with columnar base and pair of free pilose laciniae. 
<emphasis bold="true" pageId="0" pageNumber="47">Presternal area</emphasis>
bearing two pairs of free, well-sclerotised, transversely aligned and presternal platelets 
<emphasis italics="true" pageId="0" pageNumber="47">ppl</emphasis>
(Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Krantzolaspina angustatus comb. nov., adult female. Ventral idiosoma." figureDoi="10.3897/zookeys.997.54262.figure2" httpUri="https://binary.pensoft.net/fig/479382" pageId="0" pageNumber="47">2</figureCitation>
, 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Krantzolaspina angustatus comb. nov., adult female. Ventral idiosoma showing the sternometasternal, genitiventrianal and parapodal shield. A Holotype of Krantzolaspina angustatus B Paratype of Indutolaelaps jiroftensis syn. nov., photos by Raphael Castilho C New material from Indonesia." figureDoi="10.3897/zookeys.997.54262.figure3" httpUri="https://binary.pensoft.net/fig/479383" pageId="0" pageNumber="47">3A-C</figureCitation>
), with transversal lineae over surface. 
<emphasis bold="true" pageId="0" pageNumber="47">Sternometasternal shield</emphasis>
(Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Krantzolaspina angustatus comb. nov., adult female. Ventral idiosoma." figureDoi="10.3897/zookeys.997.54262.figure2" httpUri="https://binary.pensoft.net/fig/479382" pageId="0" pageNumber="47">2</figureCitation>
, 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Krantzolaspina angustatus comb. nov., adult female. Ventral idiosoma showing the sternometasternal, genitiventrianal and parapodal shield. A Holotype of Krantzolaspina angustatus B Paratype of Indutolaelaps jiroftensis syn. nov., photos by Raphael Castilho C New material from Indonesia." figureDoi="10.3897/zookeys.997.54262.figure3" httpUri="https://binary.pensoft.net/fig/479383" pageId="0" pageNumber="47">3A-C</figureCitation>
) 202-209 
<normalizedToken originalValue="μm">μm</normalizedToken>
long, 104-110 
<normalizedToken originalValue="μm">μm</normalizedToken>
wide (at level of 
<emphasis italics="true" pageId="0" pageNumber="47">st2</emphasis>
) well-defined, reticulate and covered by semi-rounded cells in the lateral margins, posteriomedial area smooth beyond to the setae 
<emphasis italics="true" pageId="0" pageNumber="47">st3</emphasis>
; anterior and posterior shield margin concave, bearing four pairs of setae 
<emphasis italics="true" pageId="0" pageNumber="47">st1</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">st4</emphasis>
(36-41) and three pairs 
<emphasis italics="true" pageId="0" pageNumber="47">iv1</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">3</emphasis>
of slit-like poroids; 
<emphasis italics="true" pageId="0" pageNumber="47">iv1</emphasis>
larger than 
<emphasis italics="true" pageId="0" pageNumber="47">iv2</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">3</emphasis>
, the latter rounded. 
<emphasis bold="true" pageId="0" pageNumber="47">Peritrematal shield</emphasis>
(Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Krantzolaspina angustatus comb. nov., adult female. Ventral idiosoma." figureDoi="10.3897/zookeys.997.54262.figure2" httpUri="https://binary.pensoft.net/fig/479382" pageId="0" pageNumber="47">2</figureCitation>
) broad, anteriorly free, fused to the exopodal shield (distinctly more sclerotised), to sternometasternal shield between the coxae I-III and posteriorly with the well-developed parapodal shield; shield extended beyond the posterior margin of the coxa IV, this area reticulate and covered by some semi-rounded cells; two pairs of poroids (
<emphasis italics="true" pageId="0" pageNumber="47">id7</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="47">ip</emphasis>
) and one pair of gland pores (
<emphasis italics="true" pageId="0" pageNumber="47">gdp</emphasis>
) discernible. Peritreme extending anteriorly between coxae I-II, at level of seta 
<emphasis italics="true" pageId="0" pageNumber="47">st1</emphasis>
. 
<emphasis bold="true" pageId="0" pageNumber="47">Genitiventrianal shield</emphasis>
(Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Krantzolaspina angustatus comb. nov., adult female. Ventral idiosoma." figureDoi="10.3897/zookeys.997.54262.figure2" httpUri="https://binary.pensoft.net/fig/479382" pageId="0" pageNumber="47">2</figureCitation>
, 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Krantzolaspina angustatus comb. nov., adult female. Ventral idiosoma showing the sternometasternal, genitiventrianal and parapodal shield. A Holotype of Krantzolaspina angustatus B Paratype of Indutolaelaps jiroftensis syn. nov., photos by Raphael Castilho C New material from Indonesia." figureDoi="10.3897/zookeys.997.54262.figure3" httpUri="https://binary.pensoft.net/fig/479383" pageId="0" pageNumber="47">3A-C</figureCitation>
) 246-264 
<normalizedToken originalValue="μm">μm</normalizedToken>
long 
<normalizedToken originalValue="×">x</normalizedToken>
127-135 
<normalizedToken originalValue="μm">μm</normalizedToken>
wide, reticulate and flask-shaped; shield with four pairs of setae 
<emphasis italics="true" pageId="0" pageNumber="47">st5</emphasis>
35-37 (slit-like poroids 
<emphasis italics="true" pageId="0" pageNumber="47">iv5</emphasis>
on unsclerotised cuticle and posterolaterad of 
<emphasis italics="true" pageId="0" pageNumber="47">st5</emphasis>
) 
<emphasis italics="true" pageId="0" pageNumber="47">ZV1</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="47">JV1</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">2</emphasis>
39-42 additionally to the circumanal setae 
<emphasis italics="true" pageId="0" pageNumber="47">pa</emphasis>
and 
<emphasis italics="true" pageId="0" pageNumber="47">po</emphasis>
, paranal setae 
<emphasis italics="true" pageId="0" pageNumber="47">po</emphasis>
(27-29) aligned with anterior margin of anal opening, postanal seta shorter 
<emphasis italics="true" pageId="0" pageNumber="47">po</emphasis>
(15-17); gland opening 
<emphasis italics="true" pageId="0" pageNumber="47">gv3</emphasis>
on posterolaterad shield margins, at level slightly anterior to paranal setae, cribrum well-developed. Soft opisthogastric cuticle with seven pairs of setae 
<emphasis italics="true" pageId="0" pageNumber="47">JV3</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">5</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="47">ZV2</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">5</emphasis>
(39-40) and three poroids, including one (
<emphasis italics="true" pageId="0" pageNumber="47">ivp</emphasis>
). 
<emphasis bold="true" pageId="0" pageNumber="47">Metapodal</emphasis>
(Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Krantzolaspina angustatus comb. nov., adult female. Ventral idiosoma." figureDoi="10.3897/zookeys.997.54262.figure2" httpUri="https://binary.pensoft.net/fig/479382" pageId="0" pageNumber="47">2</figureCitation>
; 
<emphasis italics="true" pageId="0" pageNumber="47">met</emphasis>
) platelets free, small and suboval.
</paragraph>
<caption doi="10.3897/zookeys.997.54262.figure2" httpUri="https://binary.pensoft.net/fig/479382" pageId="0" pageNumber="47" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="47">
<emphasis bold="true" pageId="0" pageNumber="47">Figure 2.</emphasis>
<taxonomicName authorityName="Quintero-Gutiérrez &amp; Sandmann &amp; Cómbita-Heredia &amp; Klarner &amp; Widyastuti &amp; Scheu" authorityYear="2020" baseAuthorityName="Ishikawa" baseAuthorityYear="1987" class="Arachnida" family="Parholaspididae" genus="Krantzolaspina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Krantzolaspina angustatus" order="Mesostigmata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="angustatus">
<emphasis italics="true" pageId="0" pageNumber="47">Krantzolaspina angustatus</emphasis>
</taxonomicName>
comb. nov., adult female. Ventral idiosoma.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.997.54262.figure3" httpUri="https://binary.pensoft.net/fig/479383" pageId="0" pageNumber="47" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="47">
<emphasis bold="true" pageId="0" pageNumber="47">Figure 3.</emphasis>
<taxonomicName authorityName="Quintero-Gutiérrez &amp; Sandmann &amp; Cómbita-Heredia &amp; Klarner &amp; Widyastuti &amp; Scheu" authorityYear="2020" baseAuthorityName="Ishikawa" baseAuthorityYear="1987" class="Arachnida" family="Parholaspididae" genus="Krantzolaspina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Krantzolaspina angustatus" order="Mesostigmata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="angustatus">
<emphasis italics="true" pageId="0" pageNumber="47">Krantzolaspina angustatus</emphasis>
</taxonomicName>
comb. nov., adult female. Ventral idiosoma showing the sternometasternal, genitiventrianal and parapodal shield. 
<emphasis bold="true" pageId="0" pageNumber="47">A</emphasis>
Holotype of 
<taxonomicName authorityName="Quintero-Gutiérrez &amp; Sandmann &amp; Cómbita-Heredia &amp; Klarner &amp; Widyastuti &amp; Scheu" authorityYear="2020" baseAuthorityName="Ishikawa" baseAuthorityYear="1987" class="Arachnida" family="Parholaspididae" genus="Krantzolaspina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Krantzolaspina angustatus" order="Mesostigmata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="angustatus">
<emphasis italics="true" pageId="0" pageNumber="47">Krantzolaspina angustatus</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="47">B</emphasis>
Paratype of 
<taxonomicName authorityName="Hajizadeh, Mortazavi, Balooch-Shahriari &amp; Castilho" authorityYear="2017" class="Arachnida" family="Leptolaelapidae" genus="Indutolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Indutolaelaps jiroftensis" order="Mesostigmata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="jiroftensis">
<emphasis italics="true" pageId="0" pageNumber="47">Indutolaelaps jiroftensis</emphasis>
</taxonomicName>
syn. nov., photos by Raphael Castilho 
<emphasis bold="true" pageId="0" pageNumber="47">C</emphasis>
New material from Indonesia.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="47">
<emphasis italics="true" pageId="0" pageNumber="47">Gnathosoma</emphasis>
(Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Krantzolaspina angustatus comb. nov., adult female, Gnathosomal structures. Holotype (Left); Indutolaelaps jiroftensis syn. nov. (Middle), photos by Raphael Castilho; New material from Indonesia (Right). A-C Subcapitulum D-F Epistome G-I Chelicera J-L Palp trochanter with a ventral spine-like process." figureDoi="10.3897/zookeys.997.54262.figure4" httpUri="https://binary.pensoft.net/fig/479384" pageId="0" pageNumber="47">4A-L</figureCitation>
). 
<emphasis bold="true" pageId="0" pageNumber="47">Subcapitulum</emphasis>
(Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Krantzolaspina angustatus comb. nov., adult female, Gnathosomal structures. Holotype (Left); Indutolaelaps jiroftensis syn. nov. (Middle), photos by Raphael Castilho; New material from Indonesia (Right). A-C Subcapitulum D-F Epistome G-I Chelicera J-L Palp trochanter with a ventral spine-like process." figureDoi="10.3897/zookeys.997.54262.figure4" httpUri="https://binary.pensoft.net/fig/479384" pageId="0" pageNumber="47">4A-C</figureCitation>
): corniculi well sclerotised, elongated and horn-like slightly shorter than cheliceral movable digit. Internal malae bifurcate, totally separated from each other, densely fimbriated on outer margin and with apices slightly shorter than corniculi. Deutosternal groove with six transverse rows denticles, each bearing 8-21 denticles and distal row smooth, with lateral ridges each side of the 2nd and 3rd row; subcapitular setae smooth and aciculate, 
<emphasis italics="true" pageId="0" pageNumber="47">h1</emphasis>
51-54 and 
<emphasis italics="true" pageId="0" pageNumber="47">h3</emphasis>
40-43 longer than 
<emphasis italics="true" pageId="0" pageNumber="47">h2</emphasis>
34-37 and 
<emphasis italics="true" pageId="0" pageNumber="47">pc</emphasis>
17-20. Surface posterolaterad to seta 
<emphasis italics="true" pageId="0" pageNumber="47">pc</emphasis>
with a minute or a small spine-like process (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Krantzolaspina angustatus comb. nov., adult female, Gnathosomal structures. Holotype (Left); Indutolaelaps jiroftensis syn. nov. (Middle), photos by Raphael Castilho; New material from Indonesia (Right). A-C Subcapitulum D-F Epistome G-I Chelicera J-L Palp trochanter with a ventral spine-like process." figureDoi="10.3897/zookeys.997.54262.figure4" httpUri="https://binary.pensoft.net/fig/479384" pageId="0" pageNumber="47">4A-C</figureCitation>
). Epistome (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Krantzolaspina angustatus comb. nov., adult female, Gnathosomal structures. Holotype (Left); Indutolaelaps jiroftensis syn. nov. (Middle), photos by Raphael Castilho; New material from Indonesia (Right). A-C Subcapitulum D-F Epistome G-I Chelicera J-L Palp trochanter with a ventral spine-like process." figureDoi="10.3897/zookeys.997.54262.figure4" httpUri="https://binary.pensoft.net/fig/479384" pageId="0" pageNumber="47">4D-F</figureCitation>
) with a median projection bifurcate or trifurcate distally, lateral edges finely and irregularly serrate. 
<emphasis bold="true" pageId="0" pageNumber="47">Cheliceral</emphasis>
(Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Krantzolaspina angustatus comb. nov., adult female, Gnathosomal structures. Holotype (Left); Indutolaelaps jiroftensis syn. nov. (Middle), photos by Raphael Castilho; New material from Indonesia (Right). A-C Subcapitulum D-F Epistome G-I Chelicera J-L Palp trochanter with a ventral spine-like process." figureDoi="10.3897/zookeys.997.54262.figure4" httpUri="https://binary.pensoft.net/fig/479384" pageId="0" pageNumber="47">4G-I</figureCitation>
) fixed digit (106-111) with four teeth (most proximal small and most distal inserted subapically) plus one distal hook-like tooth and a setiform 
<emphasis italics="true" pageId="0" pageNumber="47">pilus dentilus</emphasis>
; dorsal and anti-axial lyrifissures present as well as dorsal setae smooth; movable digit (97-103) with two teeth directed backwards and one distal hook-like tooth, base of the movable digit with a plumose arthrodial brush (44-48) (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Krantzolaspina angustatus comb. nov., adult female, Gnathosomal structures. Holotype (Left); Indutolaelaps jiroftensis syn. nov. (Middle), photos by Raphael Castilho; New material from Indonesia (Right). A-C Subcapitulum D-F Epistome G-I Chelicera J-L Palp trochanter with a ventral spine-like process." figureDoi="10.3897/zookeys.997.54262.figure4" httpUri="https://binary.pensoft.net/fig/479384" pageId="0" pageNumber="47">4I</figureCitation>
, see arrow) much shorter than movable digit and a narrow fringed arthrodial corona which is only discernible ventrally. 
<emphasis bold="true" pageId="0" pageNumber="47">Palp</emphasis>
(Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Krantzolaspina angustatus comb. nov., adult female. Palp, note the ventral spine-like process in palptrochanter." figureDoi="10.3897/zookeys.997.54262.figure5" httpUri="https://binary.pensoft.net/fig/479385" pageId="0" pageNumber="47">5</figureCitation>
) with normal chaetotaxy for 
<taxonomicName authorityName="Evans" authorityYear="1956" family="Parholaspididae" lsidName="" pageId="0" pageNumber="47" rank="family">Parholaspididae</taxonomicName>
, with 2-5-6-14-15 setae on trochanter-femur-genu-tibia-tarsus; palptrochanter almost twice longer than palpfemur, with a small pointed spine-like process on the ventral surface (Figs 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Krantzolaspina angustatus comb. nov., adult female, Gnathosomal structures. Holotype (Left); Indutolaelaps jiroftensis syn. nov. (Middle), photos by Raphael Castilho; New material from Indonesia (Right). A-C Subcapitulum D-F Epistome G-I Chelicera J-L Palp trochanter with a ventral spine-like process." figureDoi="10.3897/zookeys.997.54262.figure4" httpUri="https://binary.pensoft.net/fig/479384" pageId="0" pageNumber="47">4J-L</figureCitation>
, 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Krantzolaspina angustatus comb. nov., adult female. Palp, note the ventral spine-like process in palptrochanter." figureDoi="10.3897/zookeys.997.54262.figure5" httpUri="https://binary.pensoft.net/fig/479385" pageId="0" pageNumber="47">5</figureCitation>
; see arrow) and setae 
<emphasis italics="true" pageId="0" pageNumber="47">v1</emphasis>
-2 thickened and long, in contrast to the setae of the other palp segments; palpfemur 
<emphasis italics="true" pageId="0" pageNumber="47">al</emphasis>
thickened, palpgenu with setae 
<emphasis italics="true" pageId="0" pageNumber="47">al1</emphasis>
-2 thickened and spatulate distally. Palp tarsal claw three-tined, medial tine conspicuous distally spatulate.
</paragraph>
<caption doi="10.3897/zookeys.997.54262.figure4" httpUri="https://binary.pensoft.net/fig/479384" pageId="0" pageNumber="47" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="47">
<emphasis bold="true" pageId="0" pageNumber="47">Figure 4.</emphasis>
<taxonomicName authorityName="Quintero-Gutiérrez &amp; Sandmann &amp; Cómbita-Heredia &amp; Klarner &amp; Widyastuti &amp; Scheu" authorityYear="2020" baseAuthorityName="Ishikawa" baseAuthorityYear="1987" class="Arachnida" family="Parholaspididae" genus="Krantzolaspina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Krantzolaspina angustatus" order="Mesostigmata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="angustatus">
<emphasis italics="true" pageId="0" pageNumber="47">Krantzolaspina angustatus</emphasis>
</taxonomicName>
comb. nov., adult female, Gnathosomal structures. Holotype (Left); 
<taxonomicName authorityName="Hajizadeh, Mortazavi, Balooch-Shahriari &amp; Castilho" authorityYear="2017" class="Arachnida" family="Leptolaelapidae" genus="Indutolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Indutolaelaps jiroftensis" order="Mesostigmata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="jiroftensis">
<emphasis italics="true" pageId="0" pageNumber="47">Indutolaelaps jiroftensis</emphasis>
</taxonomicName>
syn. nov. (Middle), photos by Raphael Castilho; New material from Indonesia (Right). 
<emphasis bold="true" pageId="0" pageNumber="47">A-C</emphasis>
Subcapitulum 
<emphasis bold="true" pageId="0" pageNumber="47">D-F</emphasis>
Epistome 
<emphasis bold="true" pageId="0" pageNumber="47">G-I</emphasis>
Chelicera 
<emphasis bold="true" pageId="0" pageNumber="47">J-L</emphasis>
Palp trochanter with a ventral spine-like process.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.997.54262.figure5" httpUri="https://binary.pensoft.net/fig/479385" pageId="0" pageNumber="47" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="47">
<emphasis bold="true" pageId="0" pageNumber="47">Figure 5.</emphasis>
<taxonomicName authorityName="Quintero-Gutiérrez &amp; Sandmann &amp; Cómbita-Heredia &amp; Klarner &amp; Widyastuti &amp; Scheu" authorityYear="2020" baseAuthorityName="Ishikawa" baseAuthorityYear="1987" class="Arachnida" family="Parholaspididae" genus="Krantzolaspina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Krantzolaspina angustatus" order="Mesostigmata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="angustatus">
<emphasis italics="true" pageId="0" pageNumber="47">Krantzolaspina angustatus</emphasis>
</taxonomicName>
comb. nov., adult female. Palp, note the ventral spine-like process in palptrochanter.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="47">
<emphasis italics="true" pageId="0" pageNumber="47">Legs</emphasis>
(Fig. 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Krantzolaspina angustatus comb. nov., adult female. A-D Legs I-IV, respectively. Coxae I-IV in the Fig. 2." figureDoi="10.3897/zookeys.997.54262.figure6" httpUri="https://binary.pensoft.net/fig/479386" pageId="0" pageNumber="47">6A-D</figureCitation>
) lengths (in 
<normalizedToken originalValue="μm">μm</normalizedToken>
): I (Fig. 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Krantzolaspina angustatus comb. nov., adult female. A-D Legs I-IV, respectively. Coxae I-IV in the Fig. 2." figureDoi="10.3897/zookeys.997.54262.figure6" httpUri="https://binary.pensoft.net/fig/479386" pageId="0" pageNumber="47">6A</figureCitation>
): 573-625, II (Fig. 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Krantzolaspina angustatus comb. nov., adult female. A-D Legs I-IV, respectively. Coxae I-IV in the Fig. 2." figureDoi="10.3897/zookeys.997.54262.figure6" httpUri="https://binary.pensoft.net/fig/479386" pageId="0" pageNumber="47">6B</figureCitation>
) 461-495, III (Fig. 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Krantzolaspina angustatus comb. nov., adult female. A-D Legs I-IV, respectively. Coxae I-IV in the Fig. 2." figureDoi="10.3897/zookeys.997.54262.figure6" httpUri="https://binary.pensoft.net/fig/479386" pageId="0" pageNumber="47">6C</figureCitation>
) 396-421, IV (Fig. 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Krantzolaspina angustatus comb. nov., adult female. A-D Legs I-IV, respectively. Coxae I-IV in the Fig. 2." figureDoi="10.3897/zookeys.997.54262.figure6" httpUri="https://binary.pensoft.net/fig/479386" pageId="0" pageNumber="47">6D</figureCitation>
) 550-575. The leg chaetotaxy/setation-Coxae: I 2 (0, 0/1, 0/1, 0); II: 2 (0, 0/1, 0/1, 0); III: 2 (0, 0/1, 0/1, 0); IV: 1 (0, 0/0, 0/1, 0); trochanters: I: 6 (1, 0/1,1/2, 1); II: 5 (1, 0/1, 0/2, 1); III: 5 (1, 1/1, 0/2, 0); IV: 5 (1, 1/1, 0/2, 0); femora: I: 13 (2, 3/1, 2/3, 2); II: 11 (2, 3/1, 2/2, 1); III: 6 (1, 2/1, 1/0, 1): IV: 6 (1, 2/1, 1/0, 1); genua: I: 12 (2, 3/2, 2/1, 2); II: 10(2, 2/1, 2/1, 2); III: 8 (2, 2/1, 2/1, 1); IV: 8 (2, 2/1, 2/0, 1); tibia: I: 12 (2, 3/2, 2/1, 2); II: 10 (2, 2/1, 2/1, 2); III: 8 (2, 1/1, 2/1, 1); IV: 8 (2, 1/1, 2/1, 1); tarsi I: not counted, II: 18; III: 18; IV: 18. Pretarsus I reduced or absent and pretarsi II-IV well-developed, including a pair of claws and a pulvillus. Legs with all setae aciculate and smooth, most setae are relatively long, except leg III which has comparatively shorter setae, tarsi I-IV which have longest and thicker setae than leg segments especially tarsus IV and a set of very short setae as follows: 
<emphasis italics="true" pageId="0" pageNumber="47">ad</emphasis>
in trochanter I, 
<emphasis italics="true" pageId="0" pageNumber="47">av</emphasis>
and 
<emphasis italics="true" pageId="0" pageNumber="47">ad2</emphasis>
and 
<emphasis italics="true" pageId="0" pageNumber="47">al2</emphasis>
in femur I, 
<emphasis italics="true" pageId="0" pageNumber="47">ad2</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">3</emphasis>
in femur II, 
<emphasis italics="true" pageId="0" pageNumber="47">al1</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">2</emphasis>
in genu II, 
<emphasis italics="true" pageId="0" pageNumber="47">pl</emphasis>
and 
<emphasis italics="true" pageId="0" pageNumber="47">pd</emphasis>
in femur IV and 
<emphasis italics="true" pageId="0" pageNumber="47">pl</emphasis>
in genu IV.
</paragraph>
<caption doi="10.3897/zookeys.997.54262.figure6" httpUri="https://binary.pensoft.net/fig/479386" pageId="0" pageNumber="47" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="47">
<emphasis bold="true" pageId="0" pageNumber="47">Figure 6.</emphasis>
<taxonomicName authorityName="Quintero-Gutiérrez &amp; Sandmann &amp; Cómbita-Heredia &amp; Klarner &amp; Widyastuti &amp; Scheu" authorityYear="2020" baseAuthorityName="Ishikawa" baseAuthorityYear="1987" class="Arachnida" family="Parholaspididae" genus="Krantzolaspina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Krantzolaspina angustatus" order="Mesostigmata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="angustatus">
<emphasis italics="true" pageId="0" pageNumber="47">Krantzolaspina angustatus</emphasis>
</taxonomicName>
comb. nov., adult female. 
<emphasis bold="true" pageId="0" pageNumber="47">A-D</emphasis>
Legs I-IV, respectively. Coxae I-IV in the Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Krantzolaspina angustatus comb. nov., adult female. Ventral idiosoma." figureDoi="10.3897/zookeys.997.54262.figure2" httpUri="https://binary.pensoft.net/fig/479382" pageId="0" pageNumber="47">2</figureCitation>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="47">
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="materials_examined">
<paragraph pageId="0" pageNumber="47">Material examined and depository.</paragraph>
<paragraph pageId="0" pageNumber="47">
• 1♀ Holotype at NSMT (NSMT-Ac 9805) collected in Philippines, Palawan Islands, Olanguan Valley, between Puerto Princesa and Roxas, on litter of tropical forest. •16♀ vouchers at ESALQ collected in Iran, Kerman Province on soil and litter at the base of 
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Medicago" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Medicago sativa" order="Fabales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="sativa">
<emphasis italics="true" pageId="0" pageNumber="47">Medicago sativa</emphasis>
</taxonomicName>
(
<taxonomicName family="Fabaceae" lsidName="" pageId="0" pageNumber="47" rank="family">Fabaceae</taxonomicName>
) and 
<taxonomicName class="Magnoliopsida" family="Rutaceae" genus="Citrus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Citrus sinensis" order="Sapindales" pageId="0" pageNumber="47" phylum="Tracheophyta" rank="species" species="sinensis">
<emphasis italics="true" pageId="0" pageNumber="47">Citrus sinensis</emphasis>
</taxonomicName>
(
<taxonomicName family="Rutaceae" lsidName="" pageId="0" pageNumber="47" rank="family">Rutaceae</taxonomicName>
). New material from Indonesia, Sumatra, Jambi Province, Bukit Duabelas region, upper soil layer (0-3 cm) of oil palm plantation, research site BO2b, 
<geoCoordinate degrees="2" direction="south" minutes="04" orientation="latitude" precision="1" seconds="32.0" value="-2.0755556">2°04'32.0&quot;S</geoCoordinate>
, 
<geoCoordinate degrees="102" direction="east" minutes="47" orientation="longitude" precision="1" seconds="30.7" value="102.79186">102°47'30.7&quot;E</geoCoordinate>
, 83.74 m a.s.l. 6♀ on slides as follows: • 1♀ deposited at LIPI (OSAL 00124840); • 1♀ deposited at SMNG (OSAL 00124839 SMNG 2020/62099). • 2♀ deposited at OSAL (OSAL 00124841, 00124842). • 2♀ deposited at ANIC (OSAL 00124843, 00124844). 10♀ in alcohol, as follows: • 2♀ deposited at LIPI, • 3♀ deposited at SMNG, • 2♀ deposited at OSAL and • 3♀ deposited at ANIC. All specimens were collected in November 2013 by B. Klarner. Additional photos of the species are deposited in Ecotaxonomy database at www.ecotaxonomy.org (ECOTAX_ID: 434549).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="differential diagnosis">
<paragraph pageId="0" pageNumber="47">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="47">
<taxonomicName lsidName="K. angustatus" pageId="0" pageNumber="47" rank="species" species="angustatus">
<emphasis italics="true" pageId="0" pageNumber="47">K. angustatus</emphasis>
</taxonomicName>
comb. nov. significantly differs from 
<taxonomicName lsidName="K. rebatii" pageId="0" pageNumber="47" rank="species" species="rebatii">
<emphasis italics="true" pageId="0" pageNumber="47">K. rebatii</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="K. solimani" pageId="0" pageNumber="47" rank="species" species="solimani">
<emphasis italics="true" pageId="0" pageNumber="47">K. solimani</emphasis>
</taxonomicName>
comb. nov. in the following combination of characters: 
<taxonomicName lsidName="K. angustatus" pageId="0" pageNumber="47" rank="species" species="angustatus">
<emphasis italics="true" pageId="0" pageNumber="47">K. angustatus</emphasis>
</taxonomicName>
has 32 pairs of smooth dorsal setae (except 
<emphasis italics="true" pageId="0" pageNumber="47">S5</emphasis>
and 
<emphasis italics="true" pageId="0" pageNumber="47">Z5</emphasis>
slightly pilose), while 
<taxonomicName lsidName="K. rebatii" pageId="0" pageNumber="47" rank="species" species="rebatii">
<emphasis italics="true" pageId="0" pageNumber="47">K. rebatii</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="K. solimani" pageId="0" pageNumber="47" rank="species" species="solimani">
<emphasis italics="true" pageId="0" pageNumber="47">K. solimani</emphasis>
</taxonomicName>
have 36 pairs of smooth setae, respectively; 
<taxonomicName lsidName="K. angustatus" pageId="0" pageNumber="47" rank="species" species="angustatus">
<emphasis italics="true" pageId="0" pageNumber="47">K. angustatus</emphasis>
</taxonomicName>
has 15 pairs of setae 
<emphasis italics="true" pageId="0" pageNumber="47">r</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">R</emphasis>
and 
<emphasis italics="true" pageId="0" pageNumber="47">UR</emphasis>
-series in the unsclerotised lateral cuticle, whereas 
<taxonomicName lsidName="K. solimani" pageId="0" pageNumber="47" rank="species" species="solimani">
<emphasis italics="true" pageId="0" pageNumber="47">K. solimani</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="K. rebatii" pageId="0" pageNumber="47" rank="species" species="rebatii">
<emphasis italics="true" pageId="0" pageNumber="47">K. rebatii</emphasis>
</taxonomicName>
have 13 and 10, respectively; in 
<taxonomicName lsidName="K. angustatus" pageId="0" pageNumber="47" rank="species" species="angustatus">
<emphasis italics="true" pageId="0" pageNumber="47">K. angustatus</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="K. solimani" pageId="0" pageNumber="47" rank="species" species="solimani">
<emphasis italics="true" pageId="0" pageNumber="47">K. solimani</emphasis>
</taxonomicName>
, the peritreme is extended anteriorly between coxae I-II at the level of 
<emphasis italics="true" pageId="0" pageNumber="47">st1</emphasis>
, while in 
<taxonomicName lsidName="K. rebatii" pageId="0" pageNumber="47" rank="species" species="rebatii">
<emphasis italics="true" pageId="0" pageNumber="47">K. rebatii</emphasis>
</taxonomicName>
, it is extended beyond coxa I; in 
<taxonomicName lsidName="K. angustatus" pageId="0" pageNumber="47" rank="species" species="angustatus">
<emphasis italics="true" pageId="0" pageNumber="47">K. angustatus</emphasis>
</taxonomicName>
, a spine-like process is developed in the subcapitulum, while this is absent in 
<taxonomicName lsidName="K. rebatii" pageId="0" pageNumber="47" rank="species" species="rebatii">
<emphasis italics="true" pageId="0" pageNumber="47">K. rebatii</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="K. solimani" pageId="0" pageNumber="47" rank="species" species="solimani">
<emphasis italics="true" pageId="0" pageNumber="47">K. solimani</emphasis>
</taxonomicName>
. Further, the arthrodial process in 
<taxonomicName lsidName="K. rebatii" pageId="0" pageNumber="47" rank="species" species="rebatii">
<emphasis italics="true" pageId="0" pageNumber="47">K. rebatii</emphasis>
</taxonomicName>
has a pair of long unequal arthrodial brushes, whereas the arthrodial process in 
<taxonomicName lsidName="K. solimani" pageId="0" pageNumber="47" rank="species" species="solimani">
<emphasis italics="true" pageId="0" pageNumber="47">K. solimani</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="K. angustatus" pageId="0" pageNumber="47" rank="species" species="angustatus">
<emphasis italics="true" pageId="0" pageNumber="47">K. angustatus</emphasis>
</taxonomicName>
has a relatively long arthrodial brush and a narrow fringed arthrodial corona. Additionally, in 
<taxonomicName lsidName="K. angustatus" pageId="0" pageNumber="47" rank="species" species="angustatus">
<emphasis italics="true" pageId="0" pageNumber="47">K. angustatus</emphasis>
</taxonomicName>
, small metapodal platelets elliptical in shape are present, while in 
<taxonomicName lsidName="K. solimani" pageId="0" pageNumber="47" rank="species" species="solimani">
<emphasis italics="true" pageId="0" pageNumber="47">K. solimani</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="K. rebatii" pageId="0" pageNumber="47" rank="species" species="rebatii">
<emphasis italics="true" pageId="0" pageNumber="47">K. rebatii</emphasis>
</taxonomicName>
, they are absent. Lastly, Table 
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Characteristics of the females of Krantzolaspina angustatus comb. nov., and some ambiguous or unknown data of K. rebatii and K. solimani comb. nov." httpUri="http://table.plazi.org/id/2D29BDD143D70A25C6721FC4BB62CB5E" pageId="0" pageNumber="47" tableUuid="2D29BDD143D70A25C6721FC4BB62CB5E">1</tableCitation>
provides uncertain/unclear or unknown characters of these species.
</paragraph>
<caption ID-Table-UUID="2D29BDD143D70A25C6721FC4BB62CB5E" httpUri="http://table.plazi.org/id/2D29BDD143D70A25C6721FC4BB62CB5E" pageId="0" pageNumber="47" start="Table 1" startId="T1">
<paragraph pageId="0" pageNumber="47">
<emphasis bold="true" pageId="0" pageNumber="47">Table 1.</emphasis>
Characteristics of the females of 
<taxonomicName authorityName="Quintero-Gutiérrez &amp; Sandmann &amp; Cómbita-Heredia &amp; Klarner &amp; Widyastuti &amp; Scheu" authorityYear="2020" baseAuthorityName="Ishikawa" baseAuthorityYear="1987" class="Arachnida" family="Parholaspididae" genus="Krantzolaspina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Krantzolaspina angustatus" order="Mesostigmata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="angustatus">
<emphasis italics="true" pageId="0" pageNumber="47">Krantzolaspina angustatus</emphasis>
</taxonomicName>
comb. nov., and some ambiguous or unknown data of 
<taxonomicName lsidName="K. rebatii" pageId="0" pageNumber="47" rank="species" species="rebatii">
<emphasis italics="true" pageId="0" pageNumber="47">K. rebatii</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="K. solimani" pageId="0" pageNumber="47" rank="species" species="solimani">
<emphasis italics="true" pageId="0" pageNumber="47">K. solimani</emphasis>
</taxonomicName>
comb. nov.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="47">
<table pageId="0" pageNumber="47">
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="47">Characters</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="47">
<taxonomicName lsidName="K. angustatus" pageId="0" pageNumber="47" rank="species" species="angustatus">
<emphasis italics="true" pageId="0" pageNumber="47">K. angustatus</emphasis>
</taxonomicName>
(Ishikawa, 1987) comb. nov.
</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
* 
<emphasis bold="true" pageId="0" pageNumber="47">
<taxonomicName lsidName="K. rebatii" pageId="0" pageNumber="47" rank="species" species="rebatii">
<emphasis italics="true" pageId="0" pageNumber="47">K. rebatii</emphasis>
</taxonomicName>
Datta &amp; Bhattacharjee, 1989
</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
* 
<emphasis bold="true" pageId="0" pageNumber="47">
<taxonomicName lsidName="K. solimani" pageId="0" pageNumber="47" rank="species" species="solimani">
<emphasis italics="true" pageId="0" pageNumber="47">K. solimani</emphasis>
</taxonomicName>
(Metwali, 1983) comb. nov.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">Dorsal shield ornamentation</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
mostly reticulate (smooth on the dorso-central region around bases of 
<emphasis italics="true" pageId="0" pageNumber="47">z1</emphasis>
and 
<emphasis italics="true" pageId="0" pageNumber="47">J2</emphasis>
)
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">completely smooth?</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
with tetra- and pentagonal reticulation (except on dorsocentral region of setae 
<emphasis italics="true" pageId="0" pageNumber="47">j-J</emphasis>
, faintly ornamented)
</td>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">Sternometasternal / ventrianal shield ornamentation</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">completely reticulate and with the lateral margins covered by some semi-rounded cells</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">mostly smooth but with lateral margins pointed?/ anterior margin pointed, elsewhere smooth?</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">mostly with tetra and pentagonal reticulation</td>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">Parapodal-peritremal shield ornamentation</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">slightly reticulate + semi-rounded cells posterior to the coxa IV</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">completely smooth?</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">completely smooth?</td>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="47">1</emphasis>
Setae of opisthogastric cuticle (
<emphasis italics="true" pageId="0" pageNumber="47">JV</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">ZV</emphasis>
)
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
Seven pairs (
<emphasis italics="true" pageId="0" pageNumber="47">JV3</emphasis>
-5, 
<emphasis italics="true" pageId="0" pageNumber="47">ZV2</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">5</emphasis>
); 
<emphasis italics="true" pageId="0" pageNumber="47">JV4</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">5</emphasis>
and 
<emphasis italics="true" pageId="0" pageNumber="47">ZV4</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">5</emphasis>
pilose
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
Six pairs (
<emphasis italics="true" pageId="0" pageNumber="47">JV4</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">5</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="47">ZV2</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">5</emphasis>
)?
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
Six pairs (
<emphasis italics="true" pageId="0" pageNumber="47">JV4</emphasis>
-5, 
<emphasis italics="true" pageId="0" pageNumber="47">ZV2</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">5</emphasis>
)
</td>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">Deutosternum: no. rows</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">Nine</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">three?</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">?</td>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">Leg chetotaxy</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="47" rowspan="2">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="47" rowspan="2">-</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="6">?</td>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">(Coxa - Tibia)</td>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">I</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">2, 6, 13, 12, 12, not counted</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">?,?, 10, 11, 11,?</td>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">II</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">2, 5, 11, 10, 10, 18</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">?, 4, 10, 9,?,?</td>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">III</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">2, 5, 6, 8, 8, 18</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">?, 3, 5, 8, 7,?</td>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">IV</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">1, 5, 6, 8, 8, 18</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">?,?, 4, 6, 7,?</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="47">
<tableNote pageId="0" pageNumber="47">
? indicates unknown or uncertain/unclear data. 
<emphasis bold="true" pageId="0" pageNumber="47">1</emphasis>
All setae are smooth and moderately long, except when mentioned otherwise. * holotype not found, presumed lost; characters presented are based on the original description.
</tableNote>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="remarks">
<paragraph pageId="0" pageNumber="47">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="47">
<taxonomicName authorityName="Datta &amp; Bhattacharjee" authorityYear="1988" class="Arachnida" family="Parholaspididae" genus="Krantzolaspina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Krantzolaspina" order="Mesostigmata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="47">Krantzolaspina</emphasis>
</taxonomicName>
is a well-defined genus by the unique combination of characters stated above (see diagnosis of the genus). However, a number of characters are also present in other dermanyssine families, for example, well-developed arthrodial brush(es) is/are also present in macrochelid mites, a sternometasternal shield bearing 
<emphasis italics="true" pageId="0" pageNumber="47">st1</emphasis>
- 
<emphasis italics="true" pageId="0" pageNumber="47">st4</emphasis>
is shared with species of 
<taxonomicName authorityName="Berlese" authorityYear="1913" family="Pachylaelapidae" lsidName="" pageId="0" pageNumber="47" rank="family">Pachylaelapidae</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3776.1.1" author="Masan, P" journalOrPublisher="Zootaxa" pageId="0" pageNumber="47" pagination="1 - 66" refId="B66" refString="Masan, P, Halliday, B, 2014. Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776: 1 - 66, DOI: https://doi.org/10.11646/zootaxa.3776.1.1" title="Review of the mite family Pachylaelapidae (Acari: Mesostigmata)." url="https://doi.org/10.11646/zootaxa.3776.1.1" volume="3776" year="2014">
<normalizedToken originalValue="Mašán">Masan</normalizedToken>
and Halliday 2014
</bibRefCitation>
) and 
<taxonomicName authorityName="Ryke" authorityYear="1962" family="Ologamasidae" lsidName="" pageId="0" pageNumber="47" rank="family">Ologamasidae</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4197.1.1" author="Castilho, RC" journalOrPublisher="Zootaxa" pageId="0" pageNumber="47" pagination="1 - 147" refId="B12" refString="Castilho, RC, Silva, ES, DeMoraes, GJ, Halliday, B, 2016. Catalogue of the family Ologamasidae Ryke (Acari: Mesostigmata). Zootaxa 4197: 1 - 147, DOI: https://doi.org/10.11646/zootaxa.4197.1.1" title="Catalogue of the family Ologamasidae Ryke (Acari: Mesostigmata)." url="https://doi.org/10.11646/zootaxa.4197.1.1" volume="4197" year="2016">Castilho et al. 2016</bibRefCitation>
) and the fusion of genital + ventral + anal shields forming a genitiventrianal shield is shared with two genera of 
<taxonomicName authorityName="A.Berlese" authorityYear="1892" family="Laelapidae" lsidName="" pageId="0" pageNumber="47" rank="family">Laelapidae</taxonomicName>
(
<taxonomicName class="Arachnida" family="Laelapidae" genus="Ololaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ololaelaps" order="Mesostigmata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="47">Ololaelaps</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Karg" authorityYear="1978" class="Arachnida" family="Laelapidae" genus="Oloopticus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Oloopticus" order="Mesostigmata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="47">Oloopticus</emphasis>
</taxonomicName>
) (
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.853.29407" author="Beaulieu, F" journalOrPublisher="Acta Arachnologica Sinica" pageId="0" pageNumber="47" refId="B5" refString="Beaulieu, F, Quintero-Gutierrez, EJ, Sandmann, D, Klarner, B, Widyastuti, R, Combita-heredia, O, Scheu, S, 2019. https://doi.org/10.3897/zookeys.853.29407" url="https://doi.org/10.3897/zookeys.853.29407" year="2019">Beaulieu et al. 2019</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="47">
Although the genitiventrianal shield is a diagnostic character for 
<taxonomicName authorityName="Datta &amp; Bhattacharjee" authorityYear="1988" class="Arachnida" family="Parholaspididae" genus="Krantzolaspina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Krantzolaspina" order="Mesostigmata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="47">Krantzolaspina</emphasis>
</taxonomicName>
, it is not an exclusive character as it also occurs in other genera of 
<taxonomicName authorityName="Evans" authorityYear="1956" family="Parholaspididae" lsidName="" pageId="0" pageNumber="47" rank="family">Parholaspididae</taxonomicName>
, such as 
<taxonomicName authorityName="Berlese" authorityYear="1904" class="Arachnida" family="Parholaspididae" genus="Holaspulus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Holaspulus" order="Mesostigmata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="47">Holaspulus</emphasis>
</taxonomicName>
, some species of 
<taxonomicName authorityName="Berlese" authorityYear="1917" class="Arachnida" family="Parholaspididae" genus="Holaspina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Holaspina" order="Mesostigmata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="47">Holaspina</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Ishikawa" authorityYear="1980" class="Arachnida" family="Parholaspididae" genus="Proparholaspulus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Proparholaspulus" order="Mesostigmata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="47">Proparholaspulus</emphasis>
</taxonomicName>
, as well as in other families, such as 
<taxonomicName authorityName="Karg" authorityYear="1978" family="Leptolaelapidae" lsidName="" pageId="0" pageNumber="47" rank="family">Leptolaelapidae</taxonomicName>
genus 
<taxonomicName authorityName="Karg" authorityYear="1997" class="Arachnida" family="Leptolaelapidae" genus="Indutolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Indutolaelaps" order="Mesostigmata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="47">Indutolaelaps</emphasis>
</taxonomicName>
Karg, 1997. In addition, this feature occurs in some species of 
<taxonomicName authorityName="A.Berlese" authorityYear="1892" family="Laelapidae" lsidName="" pageId="0" pageNumber="47" rank="family">Laelapidae</taxonomicName>
(see above), but in those species, the genitiventrianal shield is hyper-developed posteriorly and occupies most of the opisthogaster and is often named the hologastric shield (
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.853.29407" author="Beaulieu, F" journalOrPublisher="Acta Arachnologica Sinica" pageId="0" pageNumber="47" refId="B5" refString="Beaulieu, F, Quintero-Gutierrez, EJ, Sandmann, D, Klarner, B, Widyastuti, R, Combita-heredia, O, Scheu, S, 2019. https://doi.org/10.3897/zookeys.853.29407" url="https://doi.org/10.3897/zookeys.853.29407" year="2019">Beaulieu et al. 2019</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="47">
One particular feature of 
<taxonomicName authorityName="Datta &amp; Bhattacharjee" authorityYear="1988" class="Arachnida" family="Parholaspididae" genus="Krantzolaspina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Krantzolaspina" order="Mesostigmata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="47">Krantzolaspina</emphasis>
</taxonomicName>
is the pointed spur-like process present on the palp trochanter and this character is important for recognising the genus. However, a similar process is present in species (and all post-embryonic stages) of the monotypic family 
<taxonomicName family="Megalolaelapidae" lsidName="" pageId="0" pageNumber="47" rank="family">Megalolaelapidae</taxonomicName>
(
<taxonomicName authorityName="Berlese" authorityYear="1892" class="Arachnida" family="Megalolaelapidae" genus="Megalolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Megalolaelaps" order="Mesostigmata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="47">Megalolaelaps</emphasis>
</taxonomicName>
), in which the palp trochanter typically bears a large anteroventral horn-like projection. Unfortunately, the function of these processes of the palp trochanter in 
<taxonomicName authorityName="Datta &amp; Bhattacharjee" authorityYear="1988" class="Arachnida" family="Parholaspididae" genus="Krantzolaspina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Krantzolaspina" order="Mesostigmata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="47">Krantzolaspina</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Berlese" authorityYear="1892" class="Arachnida" family="Megalolaelapidae" genus="Megalolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Megalolaelaps" order="Mesostigmata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="47">Megalolaelaps</emphasis>
</taxonomicName>
is unknown (
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3776.1.1" author="Masan, P" journalOrPublisher="Zootaxa" pageId="0" pageNumber="47" pagination="1 - 66" refId="B66" refString="Masan, P, Halliday, B, 2014. Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa 3776: 1 - 66, DOI: https://doi.org/10.11646/zootaxa.3776.1.1" title="Review of the mite family Pachylaelapidae (Acari: Mesostigmata)." url="https://doi.org/10.11646/zootaxa.3776.1.1" volume="3776" year="2014">
<normalizedToken originalValue="Mašán">Masan</normalizedToken>
and Halliday 2014
</bibRefCitation>
, 
<bibRefCitation DOI="https://doi.org/10.11158/saa.23.6.7" author="Combita-Heredia, O" journalOrPublisher="Systematic &amp; Applied Acarology" pageId="0" pageNumber="47" pagination="1102 - 1124" refId="B14" refString="Combita-Heredia, O, Quintero-Gutierrez, EJ, Klompen, H, 2018. Ontogeny of Megalolaelaps colossus sp. nov. (Acari: Megalolaelapidae), an enigmatic symbiont of dung beetles (Coleoptera: Scarabaeidae) in Colombia. Systematic &amp; Applied Acarology 23: 1102 - 1124, DOI: https://doi.org/10.11158/saa.23.6.7" title="Ontogeny of Megalolaelaps colossus sp. nov. (Acari: Megalolaelapidae), an enigmatic symbiont of dung beetles (Coleoptera: Scarabaeidae) in Colombia." url="https://doi.org/10.11158/saa.23.6.7" volume="23" year="2018">
<normalizedToken originalValue="Cómbita-Heredia">Combita-Heredia</normalizedToken>
et al. 2018
</bibRefCitation>
). It should be noted that members of 
<taxonomicName family="Megalolaelapidae" lsidName="" pageId="0" pageNumber="47" rank="family">Megalolaelapidae</taxonomicName>
are more similar morphologically to 
<taxonomicName authorityName="Berlese" authorityYear="1913" family="Pachylaelapidae" lsidName="" pageId="0" pageNumber="47" rank="family">Pachylaelapidae</taxonomicName>
and to 
<taxonomicName authorityName="Vitzthum" authorityYear="1930" family="Macrochelidae" lsidName="" pageId="0" pageNumber="47" rank="family">Macrochelidae</taxonomicName>
of the genus 
<taxonomicName authorityName="Oudemans" authorityYear="1902" class="Arachnida" family="Macrochelidae" genus="Neopodocinum" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neopodocinum" order="Mesostigmata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="47">Neopodocinum</emphasis>
</taxonomicName>
Oudemans (
<taxonomicName authorityName="Vitzthum" authorityYear="1930" family="Macrochelidae" lsidName="" pageId="0" pageNumber="47" rank="family">Macrochelidae</taxonomicName>
) than to 
<taxonomicName authorityName="Evans" authorityYear="1956" family="Parholaspididae" lsidName="" pageId="0" pageNumber="47" rank="family">Parholaspididae</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.11158/saa.23.6.7" author="Combita-Heredia, O" journalOrPublisher="Systematic &amp; Applied Acarology" pageId="0" pageNumber="47" pagination="1102 - 1124" refId="B14" refString="Combita-Heredia, O, Quintero-Gutierrez, EJ, Klompen, H, 2018. Ontogeny of Megalolaelaps colossus sp. nov. (Acari: Megalolaelapidae), an enigmatic symbiont of dung beetles (Coleoptera: Scarabaeidae) in Colombia. Systematic &amp; Applied Acarology 23: 1102 - 1124, DOI: https://doi.org/10.11158/saa.23.6.7" title="Ontogeny of Megalolaelaps colossus sp. nov. (Acari: Megalolaelapidae), an enigmatic symbiont of dung beetles (Coleoptera: Scarabaeidae) in Colombia." url="https://doi.org/10.11158/saa.23.6.7" volume="23" year="2018">
<normalizedToken originalValue="Cómbita-Heredia">Combita-Heredia</normalizedToken>
et al. 2018
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>