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<document id="E0C228874C19AC0EAA2FDF3B4A6B361D" ID-DOI="10.1080/00222933.2012.759285" ID-ISSN="1464-5262" ID-Zenodo-Dep="5197572" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628882145230" checkinUser="felipe" docAuthor="Park, Kyu-Tek, Bae, Yang-Seop, Kim, Sora &amp; Heppner, John B." docDate="2013" docId="BD468B24D060303954DF4DFFFC48FBAE" docLanguage="en" docName="JNatHist.47.35-36.2289-2304.pdf" docOrigin="Journal of Natural History 47 (35 - 36)" docSource="http://dx.doi.org/10.1080/00222933.2012.759285" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.6:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="6" docTitle="Torodora cambodiana Park &amp; Bae &amp; Kim &amp; Heppner 2013, sp. nov." docType="treatment" docVersion="4" lastPageNumber="2301" masterDocId="417FF35CD06D303755764E11FFFEFFC7" masterDocTitle="Genus Torodora Meyrick in Cambodia (Lepidoptera: Lecithoceridae: Torodorinae), with descriptions of three new species" masterLastPageNumber="2304" masterPageNumber="2289" pageNumber="2300" updateTime="1699054287737" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="013AFC4A98A6C4F078FD03CDCC00F3B1">Genus Torodora Meyrick in Cambodia (Lepidoptera: Lecithoceridae: Torodorinae), with descriptions of three new species</mods:title>
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<mods:namePart id="A8FEE900FB40AF97258BE674BC20CD00">Park, Kyu-Tek</mods:namePart>
<mods:affiliation id="94C26E687987F518DA6E85F5A0F104CA">The Korean Academy of Science and Technology, Seungnam, Korea; &amp; McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, FL, USA</mods:affiliation>
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<mods:namePart id="25769C3991887BD9D01F4DDC7408F835">Bae, Yang-Seop</mods:namePart>
<mods:affiliation id="318969FE852C5CD49DCC296B0A7F0E3B">Division of Life Sciences, College of Natural Sciences, University of Incheon, Incheon, Korea;</mods:affiliation>
</mods:name>
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<mods:namePart id="CF5B51C68B2AE58077F02A888EFE6E66">Kim, Sora</mods:namePart>
<mods:affiliation id="08EDEB10DE7D04F56312C3F609624CE7">Division of Entomology, School of Agricultural Biotechnology, Research Institute for Agriculture and Life Sciences, Seoul National University, Seoul, Korea;</mods:affiliation>
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<mods:namePart id="8720FFFFCC6A36EC9E144135E9634A3A">Heppner, John B.</mods:namePart>
<mods:affiliation id="DEA29B3E638BC08B41109A9BD01BF328">McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, FL, USA</mods:affiliation>
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<mods:title id="C9C0D8ADC6B2EE1A17059CE749A6EB2D">Journal of Natural History</mods:title>
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<mods:date id="979A1F500E4259799D9E139AA58C2618">2013</mods:date>
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<mods:number id="35EE95F6A9E46BAF6075735E926F777B">2013-08-06</mods:number>
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<mods:number id="BEE55BC9CE62367751179F6F7CFD9ED8">47</mods:number>
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<mods:number id="4BD83DBBFCEB48D35B1DB260EDCB6144">35 - 36</mods:number>
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<treatment id="BD468B24D060303954DF4DFFFC48FBAE" ID-DOI="http://doi.org/10.5281/zenodo.5197568" ID-Zenodo-Dep="5197568" LSID="urn:lsid:zoobank.org:act:D9D3137E-2A02-4CF1-8083-1ED87C93C1C9" httpUri="http://treatment.plazi.org/id/BD468B24D060303954DF4DFFFC48FBAE" lastPageId="14" lastPageNumber="2301" pageId="13" pageNumber="2300">
<subSubSection id="7DF569B9D060303A54DF4DFFFCC9FBC1" box="[425,823,1006,1030]" pageId="13" pageNumber="2300" type="nomenclature">
<paragraph id="35503A32D060303A54DF4DFFFCC9FBC1" blockId="13.[425,823,1006,1062]" box="[425,823,1006,1030]" pageId="13" pageNumber="2300">
<heading id="6E188D5ED060303A54DF4DFFFCC9FBC1" box="[425,823,1006,1030]" centered="true" fontSize="10" level="2" pageId="13" pageNumber="2300" reason="2">
<taxonomicName id="F2EF41B1D060303A54DF4DFFFD29FBC1" ID-CoL="9FNT2" authority="Park" authorityName="Park &amp; Bae &amp; Kim &amp; Heppner" authorityYear="2013" box="[425,727,1006,1030]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="2300" phylum="Arthropoda" rank="species" species="cambodiana" status="sp. nov.">
<emphasis id="079BE620D060303A54DF4DFFFD66FBC1" bold="true" box="[425,664,1006,1030]" italics="true" pageId="13" pageNumber="2300">Torodora cambodiana</emphasis>
Park
</taxonomicName>
,
<taxonomicNameLabel id="1CA85B5BD060303A57944DFEFCC9FBC1" box="[738,823,1007,1030]" pageId="13" pageNumber="2300" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
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<subSubSection id="7DF569B9D060303A548D4A1FFE58FB42" pageId="13" pageNumber="2300" type="description">
<paragraph id="35503A32D060303A548D4A1FFD1AFBE1" blockId="13.[425,823,1006,1062]" box="[507,740,1038,1062]" pageId="13" pageNumber="2300">
(
<figureCitation id="ADD426B7D060303A57744A1FFD78FBE1" box="[514,646,1038,1062]" captionStart="Figure 2" captionStartId="6.[211,278,1409,1431]" captionTargetBox="[213,1172,167,1376]" captionTargetId="figure-5@6.[212,1172,167,1376]" captionTargetPageId="6" captionText="Figure 2. Adults of Torodora spp. (AC) Torodora parotidosa (Wu); (B) head part in dorsal view; (C) labial palpus; (DF) T. osamensis Park, sp. nov.; (E) head part in dorsal view; (F) labial palpus; (GI) T. occidentalis Park, sp. nov.; (H) head part in dorsal view; (I) labial palpus; (JL) T. aritai Park, sp. nov.; (K) head part in dorsal view; (L) labial palpus." figureDoi="http://doi.org/10.5281/zenodo.5197576" httpUri="https://zenodo.org/record/5197576/files/figure.png" pageId="13" pageNumber="2300">Figs 2MO</figureCitation>
;
<figureCitation id="ADD426B7D060303A57E44A1FFD22FBE1" box="[658,732,1038,1062]" captionStart="Figure 3" captionStartId="8.[211,278,1613,1635]" captionTargetBox="[213,1172,165,1580]" captionTargetId="figure-5@8.[212,1172,162,1580]" captionTargetPageId="8" captionText="Figure 3. Male genitalia of Torodora spp. (AC) Torodora parotidosa (Wu); (B) aedeagus; (C) abdominal segments; (DF) T. osamensis Park, sp. nov.; (E) aedeagus; (F) abdominal segments; (GI) T. occidentalis Park, sp. nov.; (H) aedeagus; (I) abdominal segments; (JL) T. aritai Park, sp. nov.; (K) aedeagus; (L) abdominal segments." figureDoi="http://doi.org/10.5281/zenodo.5197580" httpUri="https://zenodo.org/record/5197580/files/figure.png" pageId="13" pageNumber="2300">3MO</figureCitation>
)
</paragraph>
<paragraph id="35503A32D060303A55F94A5FFE58FB42" blockId="13.[143,1104,1101,1158]" pageId="13" pageNumber="2300">
<materialsCitation id="8587306FD060303A55F94A5FFE58FB42" collectingDate="2010-05-04" country="Cambodia" location="Pursat" pageId="13" pageNumber="2300" specimenCount="1" specimenCount-male="1" stateProvince="Pursat" typeStatus="holotype">
<emphasis id="079BE620D060303A55F94A5FFF08FBA1" box="[143,246,1102,1126]" italics="true" pageId="13" pageNumber="2300">
<typeStatus id="EA548490D060303A55F94A5FFF08FBA1" box="[143,246,1102,1126]" pageId="13" pageNumber="2300" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="23E9F1BBD060303A54704A5FFEE5FBA1" box="[262,283,1102,1126]" pageId="13" pageNumber="2300" type="male"></specimenCount>
,
<collectingCountry id="4DF87AA2D060303A545C4A5CFE5AFBA2" box="[298,420,1101,1125]" name="Cambodia" pageId="13" pageNumber="2300">Cambodia</collectingCountry>
, Prov.
<collectingRegion id="F72BF4D0D060303A54854A5FFDC2FBA2" box="[499,572,1102,1125]" country="Cambodia" name="Pursat" pageId="13" pageNumber="2300">Pursat</collectingRegion>
(Osam),
<date id="41511CF2D060303A57D14A5CFCD0FBA2" box="[679,814,1101,1126]" pageId="13" pageNumber="2300" value="2010-05-04">
<collectingDate id="5115E51AD060303A57D14A5CFCD0FBA2" box="[679,814,1101,1126]" pageId="13" pageNumber="2300" value="2010-05-04">4 May 2010</collectingDate>
</date>
(Bae, Chae and Vi), gen. slide no. CIS-6069
<emphasis id="079BE620D060303A542B4A7FFE97FB41" bold="true" box="[349,361,1134,1158]" italics="true" pageId="13" pageNumber="2300">/</emphasis>
Park.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="7DF569B9D060303A55F94AC0FC28FA28" pageId="13" pageNumber="2300" type="diagnosis">
<paragraph id="35503A32D060303A55F94AC0FF05FB2F" blockId="13.[143,1104,1233,1519]" box="[143,251,1233,1256]" pageId="13" pageNumber="2300">
<heading id="6E188D5ED060303A55F94AC0FF05FB2F" box="[143,251,1233,1256]" fontSize="10" level="3" pageId="13" pageNumber="2300" reason="5">
<emphasis id="079BE620D060303A55F94AC0FF05FB2F" box="[143,251,1233,1256]" italics="true" pageId="13" pageNumber="2300">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="35503A32D060303A55F94AE9FC28FA28" blockId="13.[143,1104,1233,1519]" pageId="13" pageNumber="2300">
The wing pattern of this species is similar to
<taxonomicName id="F2EF41B1D060303A57DE4AE8FC66FAD7" authority="(Meyrick)" baseAuthorityName="Meyrick" box="[680,920,1272,1296]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="2300" phylum="Arthropoda" rank="species" species="acrifera">
<emphasis id="079BE620D060303A57DE4AE8FCE3FAD7" box="[680,797,1272,1296]" italics="true" pageId="13" pageNumber="2300">T. acrifera</emphasis>
(Meyrick)
</taxonomicName>
described from
<collectingCountry id="4DF87AA2D060303A55F94B09FF29FAF7" box="[143,215,1304,1328]" name="Nepal" pageId="13" pageNumber="2300">Nepal</collectingCountry>
,
<taxonomicName id="F2EF41B1D060303A55954B08FE84FAF7" authority="Wu" authorityName="Wu" authorityYear="1997" box="[227,378,1304,1328]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="2300" phylum="Arthropoda" rank="species" species="sirtalis">
<emphasis id="079BE620D060303A55954B08FEB4FAF7" box="[227,330,1304,1328]" italics="true" pageId="13" pageNumber="2300">T. sirtalis</emphasis>
Wu
</taxonomicName>
described from
<collectingCountry id="4DF87AA2D060303A57474B09FD87FAF7" box="[561,633,1304,1328]" name="China" pageId="13" pageNumber="2300">China</collectingCountry>
,
<taxonomicName id="F2EF41B1D060303A57F24B08FCC0FAF7" authority="Park" authorityName="Park" box="[644,830,1304,1328]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="2300" phylum="Arthropoda" rank="species" species="rectlinea">
<emphasis id="079BE620D060303A57F24B08FCFEFAF7" box="[644,768,1304,1328]" italics="true" pageId="13" pageNumber="2300">T. rectlinea</emphasis>
Park
</taxonomicName>
described from
<collectingCountry id="4DF87AA2D060303A56834B09FBB5FAF7" box="[1013,1099,1304,1328]" name="Taiwan" pageId="13" pageNumber="2300">Taiwan</collectingCountry>
, and
<taxonomicName id="F2EF41B1D060303A55B64B28FE93FA97" authority="Park" authorityName="Park" authorityYear="2006" box="[192,365,1336,1360]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="2300" phylum="Arthropoda" rank="species" species="nabiella">
<emphasis id="079BE620D060303A55B64B28FECFFA97" box="[192,305,1336,1360]" italics="true" pageId="13" pageNumber="2300">T. nabiella</emphasis>
Park
</taxonomicName>
described from
<collectingCountry id="4DF87AA2D060303A57694B29FD7CFA97" box="[543,642,1336,1360]" name="Vietnam" pageId="13" pageNumber="2300">Vietnam</collectingCountry>
, but they can be easily distinguished from each other by the different shapes of their male genitalia. The male genitalia of the new species are allied to those of
<taxonomicName id="F2EF41B1D060303A54A84B68FD81FA57" authority="Park" authorityName="Park" authorityYear="2002" box="[478,639,1400,1424]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="2300" phylum="Arthropoda" rank="species" species="loeica">
<emphasis id="079BE620D060303A54A84B68FDC0FA57" box="[478,574,1400,1424]" italics="true" pageId="13" pageNumber="2300">T. loeica</emphasis>
Park
</taxonomicName>
, but can be distinguished by the slender uncus and the shorter cucullus. The gnathos of the male genitalia resembles that of
<taxonomicName id="F2EF41B1D060303A55F94BA9FED8FA08" authority="Wu" authorityName="Wu" authorityYear="1997" box="[143,294,1464,1488]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="2300" phylum="Arthropoda" rank="species" species="sirtalis">
<emphasis id="079BE620D060303A55F94BA9FF09FA17" box="[143,247,1464,1488]" italics="true" pageId="13" pageNumber="2300">T. sirtalis</emphasis>
Wu
</taxonomicName>
or
<taxonomicName id="F2EF41B1D060303A543A4BA9FDB9FA08" authority="(Meyrick)" baseAuthorityName="Meyrick" box="[332,583,1463,1487]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="2300" phylum="Arthropoda" rank="species" species="octavana">
<emphasis id="079BE620D060303A543A4BA9FE32FA08" box="[332,460,1464,1487]" italics="true" pageId="13" pageNumber="2300">T. octavana</emphasis>
(Meyrick)
</taxonomicName>
, with a small apical tip of the median process; however, the forewing patterns can be easily distinguished from each other.
</paragraph>
</subSubSection>
<subSubSection id="7DF569B9D060303955F9482AFC35FC82" lastPageId="14" lastPageNumber="2301" pageId="13" pageNumber="2300" type="description">
<paragraph id="35503A32D060303A55F9482AFEF0F995" blockId="13.[143,1104,1595,1691]" box="[143,270,1595,1618]" pageId="13" pageNumber="2300">
<heading id="6E188D5ED060303A55F9482AFEF0F995" box="[143,270,1595,1618]" fontSize="10" level="3" pageId="13" pageNumber="2300" reason="5">
<emphasis id="079BE620D060303A55F9482AFEF0F995" box="[143,270,1595,1618]" italics="true" pageId="13" pageNumber="2300">Description</emphasis>
</heading>
</paragraph>
<paragraph id="35503A32D060303955F94873FD25FDD9" blockId="13.[143,1104,1595,1691]" lastBlockId="14.[211,1172,167,542]" lastPageId="14" lastPageNumber="2301" pageId="13" pageNumber="2300">
<emphasis id="079BE620D060303A55F94873FF33F9BD" box="[143,205,1634,1658]" italics="true" pageId="13" pageNumber="2300">Adult</emphasis>
(
<figureCitation id="ADD426B7D060303A55974873FE7EF9BD" box="[225,384,1634,1658]" captionStart="Figure 2" captionStartId="6.[211,278,1409,1431]" captionTargetBox="[213,1172,167,1376]" captionTargetId="figure-5@6.[212,1172,167,1376]" captionTargetPageId="6" captionText="Figure 2. Adults of Torodora spp. (AC) Torodora parotidosa (Wu); (B) head part in dorsal view; (C) labial palpus; (DF) T. osamensis Park, sp. nov.; (E) head part in dorsal view; (F) labial palpus; (GI) T. occidentalis Park, sp. nov.; (H) head part in dorsal view; (I) labial palpus; (JL) T. aritai Park, sp. nov.; (K) head part in dorsal view; (L) labial palpus." figureDoi="http://doi.org/10.5281/zenodo.5197576" httpUri="https://zenodo.org/record/5197576/files/figure.png" pageId="13" pageNumber="2300">Figure 2MO</figureCitation>
). Wingspan
<quantity id="F21797D7D060303A576E4873FD93F9BD" box="[536,621,1634,1658]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="13" pageNumber="2300" unit="mm" value="16.0">16 mm</quantity>
. Head and thorax dark fuscous dorsally. Basal segment of antenna elongate, greyish orange speckled with dark fuscous scales; flagellum orange grey. Second segment of labial palpus long, thickened with rough scales, dark fuscous in basal half and paler towards apex on outer surface, orange white on inner surface; third segment slender, longer than second. Forewing dark fuscous in basal two-fifths of length, greyish orange medially, fuscous beyond postmedian line; blackish discal stigma at end of cell and a slightly larger one below it; apex obtuse; termen nearly straight, blackish scales along margin; venation with R
<subScript id="A96B3877D063303956954F41FC12FEA6" attach="left" box="[995,1004,336,353]" fontSize="7" pageId="14" pageNumber="2301">2</subScript>
closer to R
<subScript id="A96B3877D063303951074F41FB6CFEA6" attach="left" box="[1137,1170,336,354]" fontSize="7" pageId="14" pageNumber="2301">3+4</subScript>
than R
<subScript id="A96B3877D063303954534F61FED0FE46" attach="left" box="[293,302,368,385]" fontSize="7" pageId="14" pageNumber="2301">1</subScript>
; R
<subScript id="A96B3877D063303954204F61FEA1FE46" attach="left" box="[342,351,368,385]" fontSize="7" pageId="14" pageNumber="2301">3</subScript>
and R
<subScript id="A96B3877D063303954C24F61FE43FE46" attach="left" box="[436,445,368,385]" fontSize="7" pageId="14" pageNumber="2301">4</subScript>
stalked for basal thre-fifths; R
<subScript id="A96B3877D0633039565E4F61FCCFFE46" attach="left" box="[808,817,368,385]" fontSize="7" pageId="14" pageNumber="2301">4</subScript>
and R
<subScript id="A96B3877D063303956F04F61FC71FE46" attach="left" box="[902,911,368,385]" fontSize="7" pageId="14" pageNumber="2301">5</subScript>
stalked beyond threequarters; R
<subScript id="A96B3877D063303954224F81FEA3FE66" attach="left" box="[340,349,400,417]" fontSize="7" pageId="14" pageNumber="2301">5</subScript>
to termen; M
<subScript id="A96B3877D0633039548D4F81FDFAFE66" attach="left" box="[507,516,400,417]" fontSize="7" pageId="14" pageNumber="2301">2</subScript>
close to M
<subScript id="A96B3877D063303957F54F81FD72FE66" attach="left" box="[643,652,400,417]" fontSize="7" pageId="14" pageNumber="2301">3</subScript>
at base; M
<subScript id="A96B3877D0633039567D4F81FCEAFE66" attach="left" box="[779,788,400,417]" fontSize="7" pageId="14" pageNumber="2301">3</subScript>
and CuA
<subScript id="A96B3877D063303956F24F81FC73FE66" attach="left" box="[900,909,400,417]" fontSize="7" pageId="14" pageNumber="2301">1</subScript>
stalked for basal third. Hindwing broader than forewing, pale brownish grey; apex more or less obtuse; termen sinuate; fringe same colour throughout, with narrow orange-white basal line; venation with M
<subScript id="A96B3877D0633039545F4FE1FECCFDC6" attach="left" box="[297,306,496,513]" fontSize="7" pageId="14" pageNumber="2301">3</subScript>
and CuA
<subScript id="A96B3877D063303954DF4FE1FE4CFDC6" attach="left" box="[425,434,496,513]" fontSize="7" pageId="14" pageNumber="2301">1</subScript>
short stalked. Hind tibia with dark-brown scales dorsally, with white-orange apex. The female was not found.
</paragraph>
<paragraph id="35503A32D063303955A54C5FFC35FC82" blockId="14.[211,1172,590,837]" pageId="14" pageNumber="2301">
<emphasis id="079BE620D063303955A54C5FFE8AFDA1" box="[211,372,590,614]" italics="true" pageId="14" pageNumber="2301">Male genitalia</emphasis>
(
<figureCitation id="ADD426B7D063303954F34C5FFDDFFDA1" box="[389,545,590,614]" captionStart="Figure 3" captionStartId="8.[211,278,1613,1635]" captionTargetBox="[213,1172,165,1580]" captionTargetId="figure-5@8.[212,1172,162,1580]" captionTargetPageId="8" captionText="Figure 3. Male genitalia of Torodora spp. (AC) Torodora parotidosa (Wu); (B) aedeagus; (C) abdominal segments; (DF) T. osamensis Park, sp. nov.; (E) aedeagus; (F) abdominal segments; (GI) T. occidentalis Park, sp. nov.; (H) aedeagus; (I) abdominal segments; (JL) T. aritai Park, sp. nov.; (K) aedeagus; (L) abdominal segments." figureDoi="http://doi.org/10.5281/zenodo.5197580" httpUri="https://zenodo.org/record/5197580/files/figure.png" pageId="14" pageNumber="2301">Figure 3MO</figureCitation>
). Similar to those of
<taxonomicName id="F2EF41B1D063303956604C5EFBECFDA1" authority="(Meyrick)" baseAuthorityName="Meyrick" box="[790,1042,590,614]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="2301" phylum="Arthropoda" rank="species" species="octavana">
<emphasis id="079BE620D063303956604C5EFC68FDA1" box="[790,918,591,614]" italics="true" pageId="14" pageNumber="2301">T. octavana</emphasis>
(Meyrick)
</taxonomicName>
, but distinguished by the aedeagus: more slender and without a bundle of needle-like cornuti. Uncus slender, heavily sclerotized with acute apex. Median process of gnathos long, stout, nearly straight, bent at extremity. Valva foot-shaped; costa strongly angled at basal quarter; cucullus nearly quadrate, densely setose; outer margin gently rounded. Juxta shield-shaped; caudal margin slightly concave. Aedeagus as long as valva, gently curved and narrowed towards apex; cornuti consist of two narrow sclerotized plates ventrally and dorsally, and a broad plate bearing spinules at middle.
</paragraph>
</subSubSection>
<subSubSection id="7DF569B9D063303955A54D89FE0DFC10" box="[211,499,920,983]" pageId="14" pageNumber="2301" type="distribution">
<paragraph id="35503A32D063303955A54D89FEA6FC77" blockId="14.[211,499,920,983]" box="[211,344,920,944]" pageId="14" pageNumber="2301">
<heading id="6E188D5ED063303955A54D89FEA6FC77" box="[211,344,920,944]" fontSize="10" level="3" pageId="14" pageNumber="2301" reason="5">
<emphasis id="079BE620D063303955A54D89FEA6FC77" box="[211,344,920,944]" italics="true" pageId="14" pageNumber="2301">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="35503A32D063303955A54DAEFE0DFC10" blockId="14.[211,499,920,983]" box="[211,499,959,983]" pageId="14" pageNumber="2301">
<collectingCountry id="4DF87AA2D063303955A54DAEFEB4FC10" box="[211,330,959,983]" name="Cambodia" pageId="14" pageNumber="2301">Cambodia</collectingCountry>
(
<collectingRegion id="F72BF4D0D0633039542E4DAEFE19FC10" box="[344,487,959,983]" country="Cambodia" name="Pursat" pageId="14" pageNumber="2301">Pursat Prov.</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="7DF569B9D063303955A54A3BFC48FBAE" box="[211,950,1066,1130]" pageId="14" pageNumber="2301" type="etymology">
<paragraph id="35503A32D063303955A54A3BFEB5FB85" blockId="14.[211,950,1066,1130]" box="[211,331,1066,1090]" pageId="14" pageNumber="2301">
<heading id="6E188D5ED063303955A54A3BFEB5FB85" box="[211,331,1066,1090]" fontSize="10" level="3" pageId="14" pageNumber="2301" reason="5">
<emphasis id="079BE620D063303955A54A3BFEB5FB85" box="[211,331,1066,1090]" italics="true" pageId="14" pageNumber="2301">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="35503A32D063303955A54A40FC48FBAE" blockId="14.[211,950,1066,1130]" box="[211,950,1105,1130]" pageId="14" pageNumber="2301">
The species name is derived from the country name of
<collectingCountry id="4DF87AA2D063303956414A40FC4CFBAE" box="[823,946,1105,1129]" name="Cambodia" pageId="14" pageNumber="2301">Cambodia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>