treatments-xml/data/71/75/51/71755174FFAE0330CE30F918CBB9F892.xml

<document id="90DD57493670B2529D3C9CB8EC3372BC" ID-DOI="10.26879/1209" ID-ISSN="1094-8074" ID-Zenodo-Dep="11063294" ID-ZooBank="1726FDAE-2EE5-4145-A124-6D24287C0514" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1714014612185" checkinUser="felipe" docAuthor="Zazhigin, Vladimir S. &amp; Voyta, Leonid L." docDate="2022" docId="71755174FFAE0330CE30F918CBB9F892" docLanguage="en" docName="PalaeontolElectron.25.3.a29.pdf" docOrigin="Palaeontologia Electronica (a 29) 25 (3)" docSource="http://dx.doi.org/10.26879/1209" docStyle="DocumentStyle:322FC57CC99045555090E8A84CFB029A.4:PalaeontolElectron.2018-.journal_article" docStyleId="322FC57CC99045555090E8A84CFB029A" docStyleName="PalaeontolElectron.2018-.journal_article" docStyleVersion="4" docTitle="Ishimosorex ishimiensis Zazhigin &amp; Voyta 2022" docType="treatment" docVersion="1" lastPageNumber="10" masterDocId="8D4C290CFFAB0339CF1EFFE8C819FF81" masterDocTitle="New Neogene anourosoricin shrews from northern Asia" masterLastPageNumber="36" masterPageNumber="1" pageNumber="6" updateTime="1714663430529" updateUser="julia" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:title id="B4FC87AC957268017410962A1DEBC75A">New Neogene anourosoricin shrews from northern Asia</mods:title>
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<mods:namePart id="1ACC260EF0478AAEECC8BA8CD0D04555">Zazhigin, Vladimir S.</mods:namePart>
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<mods:namePart id="2C893011CE2FCA7EAE326E8BA4356112">Voyta, Leonid L.</mods:namePart>
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<mods:title id="62EECE90900482F20214AED69E02FEFA">Palaeontologia Electronica</mods:title>
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<mods:date id="F6077B3B682AFCC467F33635A3D21EC9">2022</mods:date>
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<subSubSection id="B1C6B3E9FFAE033CCE30F918CAB2F888" box="[302,683,1776,1801]" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="F963E062FFAE033CCE30F918CAB2F888" blockId="5.[302,683,1776,1833]" box="[302,683,1776,1801]" pageId="5" pageNumber="6">
<taxonomicName id="3EDC9BE1FFAE033CCE30F918CA51F888" authority="Zazhigin &amp; Voyta, 2022" authorityName="Zazhigin &amp; Voyta" authorityYear="2022" box="[302,584,1776,1801]" class="Mammalia" family="Soricidae" genus="Ishimosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="ishimiensis" status="sp. nov.">
<emphasis id="CBA83C70FFAE033CCE30F918CA51F888" box="[302,584,1776,1801]" italics="true" pageId="5" pageNumber="6">Ishimosorex ishimiensis</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
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<figureCitation id="61E7FCE7FFAE033CCEA5F8F8CA04F8A8" box="[443,541,1808,1833]" captionStart="FIGURE 2" captionStartId="5.[192,283,1464,1487]" captionTargetBox="[188,1437,164,1440]" captionTargetId="figure-53@5.[188,1437,164,1441]" captionTargetPageId="5" captionText="FIGURE 2. Teeth and bones of Ishimosorex ishimiensis gen. et sp. nov. from late Miocene Petropavlovsk 1A locality (A–E, G–K) and Borki 1A locality (F). A, GIN 952/1161, left I1 in lateral view (Figure 2A cf. Figure 7A); B, GIN 952/ 1160, right I1 in medial view; C, GIN 952/1152, left fragment of mandibular ramus in posterior view (image displays articular surface of the condylar process); D, ibid., diagrammatic image of mandibular ramus in medial view; E, GIN 952/1159, left i1 in lateral view; F, GIN 1115/1149, right i1 in medial view; G, GIN 952/1155, left M1 in occlusal view (G1, pure view; G2, augmented view) (Figure 2G cf. Figure 7H); H, GIN 952/1153, right dentary fragment with p4–m1 and anterior alveolus of m2; J, GIN 952/1158, right m1 in occlusal view; K, GIN 952/1154, left m2 in occlusal view. Star indicates reversed image. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11063298" httpUri="https://zenodo.org/record/11063298/files/figure.png" pageId="5" pageNumber="6">Figure 2</figureCitation>
</paragraph>
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zoobank.org/ 
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<paragraph id="F963E062FFAE033CCC56F918CC8EF8E8" blockId="5.[840,1441,1776,1897]" pageId="5" pageNumber="6">
<emphasis id="CBA83C70FFAE033CCC56F918CBF7F888" bold="true" box="[840,1006,1776,1801]" pageId="5" pageNumber="6">
<typeStatus id="26675EC0FFAE033CCC56F918CB9DF888" box="[840,900,1776,1801]" pageId="5" pageNumber="6">Type</typeStatus>
locality.
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<location id="FC03B6B9FFAE033CCCE9F918CC84F888" LSID="urn:lsid:plazi:treatment:71755174FFAE0330CE30F918CBB9F892:FC03B6B9FFAE033CCCE9F918CC84F888" box="[1015,1181,1776,1801]" country="Kazakhstan" county="Petropavlovsk town" latitude="54.9" longLatPrecision="1067" longitude="69.11667" municipality="Ishim River Valley" name="Petropavlovsk" pageId="5" pageNumber="6" stateProvince="Soltustik Qazaqstan Oblysy">Petropavlovsk</location>
1A, right slope of the 
<collectingMunicipality id="19077A18FFAE033CCC56F8F8CC3DF8A8" box="[840,1060,1808,1833]" pageId="5" pageNumber="6">Ishim River Valley</collectingMunicipality>
, within 
<collectingCounty id="100298EEFFAE033CCB9EF8F8CD70F8A8" box="[1152,1385,1808,1833]" pageId="5" pageNumber="6">Petropavlovsk town</collectingCounty>
(ca. 
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<geoCoordinate id="9CE886A5FFAE033CCCDEF8D8CC3AF8C8" box="[960,1059,1840,1865]" degrees="69" direction="east" minutes="07" orientation="longitude" pageId="5" pageNumber="6" precision="925" value="69.11667">E69°07'</geoCoordinate>
), 
<collectingRegion id="3B182E80FFAE033CCB58F8D8CD84F8C8" box="[1094,1437,1840,1865]" country="Kazakhstan" name="Soltustik Qazaqstan Oblysy" pageId="5" pageNumber="6">North Kazakhstan Region</collectingRegion>
, 
<collectingCountry id="81CBA0F2FFAE033CCC56F8B8CBCAF8E8" box="[840,979,1872,1897]" name="Kazakhstan" pageId="5" pageNumber="6">Kazakhstan</collectingCountry>
(
<figureCitation id="61E7FCE7FFAE033CCCFFF8B8CC5FF8E8" box="[993,1094,1872,1897]" captionStart="FIGURE 1" captionStartId="2.[194,285,1060,1083]" captionTargetBox="[196,1434,169,1028]" captionTargetId="figure-256@2.[193,1438,164,1031]" captionTargetPageId="2" captionText="FIGURE 1. Map of Asia (A) showing the location of Russian and Kazakh late Miocene and early Pliocene sites (B): ANR, Andreevka 1A, 2A; BRK, Borki 1A–C; BTK, Biteke; CHR, Cherlak 1A; ISK, Isakovka 1A, 2A; KRN, Krasnokutsk; KRT, Kartashovo; LZN, Lezhanka 1A, 2B; NIL, Nizhneil’inka; NST, Novaya Stanitsa 1A–B; PPL, Petropavlovsk 1A; PSH, Peshnevo 1A–B; PVL, Pavlodar 1A, 2 ‘Quarry’; SLT, Selety 1A (type locality of Paranourosorex seletiensis Storch and Zazhigin, 1996). Age of the deposits is marked by colours, which corresponds to ELMA Neogene zones ‘MN 10, 12–14’; the redeposited fossiliferous layers are marked by ‘?’. Map data from resource ESRI (http:// www.esri.com/) using SASPlanet software (v.160707.9476)." figureDoi="http://doi.org/10.5281/zenodo.11063296" httpUri="https://zenodo.org/record/11063296/files/figure.png" pageId="5" pageNumber="6">Figure 1</figureCitation>
: PPL).
</paragraph>
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<paragraph id="F963E062FFAD033FCFD8FBB0CB56FB73" blockId="6.[198,1433,1112,1266]" pageId="6" pageNumber="7">
<emphasis id="CBA83C70FFAD033FCFD8FBB0C925FBF1" bold="true" box="[198,316,1112,1136]" pageId="6" pageNumber="7">FIGURE 3.</emphasis>
The characters of the first lower incisors of 
<taxonomicName id="3EDC9BE1FFAD033FCC0FFBB1CBE5FBF1" authorityName="Rzebik-Kowalska" authorityYear="1975" box="[785,1020,1113,1136]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="gigas">
<emphasis id="CBA83C70FFAD033FCC0FFBB1CBE5FBF1" box="[785,1020,1113,1136]" italics="true" pageId="6" pageNumber="7">Paranourosorex gigas</emphasis>
</taxonomicName>
from ANR/1A (GIN 1112/1108; 
<emphasis id="CBA83C70FFAD033FCA53FBB1CD70FBF5" bold="true" box="[1357,1385,1113,1140]" pageId="6" pageNumber="7">
A 
<subScript id="6558E227FFAD033FCA40FB8ACD70FBF5" attach="left" box="[1374,1385,1122,1140]" fontSize="7" pageId="6" pageNumber="7">1</subScript>
</emphasis>
, lateral, 
<emphasis id="CBA83C70FFAD033FCFE2FB92C901FB17" bold="true" box="[252,280,1146,1174]" pageId="6" pageNumber="7">
A 
<subScript id="6558E227FFAD033FCE13FB6CC901FB17" attach="left" box="[269,280,1156,1174]" fontSize="7" pageId="6" pageNumber="7">2</subScript>
</emphasis>
, medial views), 
<taxonomicName id="3EDC9BE1FFAD033FCEDBFB92CADBFB10" authority="Zazhigin &amp; Voyta, 2022" authorityName="Zazhigin &amp; Voyta" authorityYear="2022" box="[453,706,1146,1169]" class="Mammalia" family="Soricidae" genus="Ishimosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="ishimiensis" status="gen. et sp. nov.">
<emphasis id="CBA83C70FFAD033FCEDBFB92CADBFB10" box="[453,706,1146,1169]" italics="true" pageId="6" pageNumber="7">Ishimosorex ishimiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D09B810BFFAD033FCDD4FB92CB76FB10" box="[714,879,1146,1169]" pageId="6" pageNumber="7" rank="species">gen. et sp. nov.</taxonomicNameLabel>
from BRK/1A (GIN 1115/1149, paratype; 
<emphasis id="CBA83C70FFAD033FCA38FB92CD5AFB17" bold="true" box="[1318,1347,1146,1174]" pageId="6" pageNumber="7">
B 
<subScript id="6558E227FFAD033FCA26FB6CCD5AFB17" attach="right" box="[1336,1347,1156,1174]" fontSize="7" pageId="6" pageNumber="7">1</subScript>
</emphasis>
, lateral, 
<emphasis id="CBA83C70FFAD033FCFD8FB74C8FBFB39" bold="true" box="[198,226,1180,1208]" pageId="6" pageNumber="7">
B 
<subScript id="6558E227FFAD033FCFC9FB4EC8FBFB39" attach="left" box="[215,226,1190,1208]" fontSize="7" pageId="6" pageNumber="7">2</subScript>
</emphasis>
, medial views), 
<taxonomicName id="3EDC9BE1FFAD033FCE93FB74CAC7FB32" authority="Spanish" authorityName="Spanish" box="[397,734,1180,1203]" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="endemica">
<emphasis id="CBA83C70FFAD033FCE93FB74CA9AFB32" box="[397,643,1180,1203]" italics="true" pageId="6" pageNumber="7">Crusafontina endemica</emphasis>
Spanish
</taxonomicName>
from Can Llobateres 1 locality (
<emphasis id="CBA83C70FFAD033FCB31FB74CC59FB32" bold="true" box="[1071,1088,1180,1203]" pageId="6" pageNumber="7">C</emphasis>
, CL1 2217, see 
<bibRefCitation id="9D4D9D93FFAD033FCBF1FB74C8EBFB54" author="Van Dam, J. A." pageId="6" pageNumber="7" pagination="741 - 764" refId="ref17856" refString="Van Dam, J. A. 2004. Anourosoricini (Mammalia: Soricidae) from the Mediterranean region: A pre-Quaternary example of recurrent climate-controlled north - south range shifting. Journal of Paleontology, 78: 741 - 764. https: // www. jstor. org / stable / 4094903" type="journal article" year="2004">van Dam, 2004: 746</bibRefCitation>
; in lateral view), 
<taxonomicName id="3EDC9BE1FFAD033FCEBBFB55CA01FB54" baseAuthorityName="Bachmayer and Wilson" baseAuthorityYear="1970" box="[421,536,1213,1237]" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="kormosi">
<emphasis id="CBA83C70FFAD033FCEBBFB55CA01FB54" box="[421,536,1213,1237]" italics="true" pageId="6" pageNumber="7">C. kormosi</emphasis>
</taxonomicName>
(
<emphasis id="CBA83C70FFAD033FCD39FB56CA21FB54" bold="true" box="[551,568,1214,1237]" pageId="6" pageNumber="7">D</emphasis>
, Polgárdi 4, in lateral view) and 
<taxonomicName id="3EDC9BE1FFAD033FCC8DFB56CC15FB54" box="[915,1036,1214,1237]" class="Mammalia" family="Soricidae" genus="Amblycoptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="oligodon">
<emphasis id="CBA83C70FFAD033FCC8DFB56CC15FB54" box="[915,1036,1214,1237]" italics="true" pageId="6" pageNumber="7">A. oligodon</emphasis>
</taxonomicName>
(
<emphasis id="CBA83C70FFAD033FCB05FB56CC32FB54" bold="true" box="[1051,1067,1214,1237]" pageId="6" pageNumber="7">E</emphasis>
, Polgárdi 4, in lateral view). Scale bars equal 1 mm; C, unscaled. Abbreviations see in Figure 1.
</paragraph>
</caption>
<paragraph id="F963E062FFAD033FCFDEFACACA41FA1A" blockId="6.[192,793,1314,1883]" pageId="6" pageNumber="7">
<emphasis id="CBA83C70FFAD033FCFDEFACAC974FABA" bold="true" box="[192,365,1314,1339]" pageId="6" pageNumber="7">
<typeStatus id="26675EC0FFAD033FCFDEFACAC8E5FABA" box="[192,252,1314,1339]" pageId="6" pageNumber="7">Type</typeStatus>
horizon.
</emphasis>
Stratotypic locality of Ishim Formation (
<emphasis id="CBA83C70FFAD033FCE1AFAAAC93FFADA" box="[260,294,1346,1371]" italics="true" pageId="6" pageNumber="7">ish</emphasis>
), late Miocene (MN 10,?9.7–8.7 Ma). Material was collected by VZ during fieldworks of 1964, 1976, 1980, 1982 and 1987.
</paragraph>
<paragraph id="F963E062FFAD033FCFDEFA4ACA1BF95A" blockId="6.[192,793,1314,1883]" pageId="6" pageNumber="7">
<emphasis id="CBA83C70FFAD033FCFDEFA4AC96EFA3A" bold="true" box="[192,375,1442,1467]" pageId="6" pageNumber="7">Type material.</emphasis>
<materialsCitation id="49B4EA3FFFAD033FCE9BFA4AC909FA7A" pageId="6" pageNumber="7">
<typeStatus id="26675EC0FFAD033FCE9BFA4AC9EAFA3A" box="[389,499,1442,1467]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
: GIN 952/1153 — right dentary fragment with p4–m1, the anterior alveolus of m2.
</materialsCitation>
<materialsCitation id="49B4EA3FFFAD033FCE03FA0AC9F9F9BA" pageId="6" pageNumber="7">
<typeStatus id="26675EC0FFAD033FCE03FA0AC980FA7A" box="[285,409,1506,1531]" pageId="6" pageNumber="7" type="paratype">Paratypes</typeStatus>
: (n = 7): GIN 952/1152 — left fragment of mandibular ramus with whole coronoid and condylar processes
</materialsCitation>
; 
<materialsCitation id="49B4EA3FFFAD033FCEF5F9CAC90EF9DA" pageId="6" pageNumber="7">GIN 952/1154 — isolated left m2;</materialsCitation>
<materialsCitation id="49B4EA3FFFAD033FCE00F9AACABFF9DA" box="[286,678,1602,1628]" pageId="6" pageNumber="7">GIN 952/1055 — isolated left M1;</materialsCitation>
<materialsCitation id="49B4EA3FFFAD033FCDB0F9AAC9E7F9FA" pageId="6" pageNumber="7">GIN 952/ 1158 — isolated right m1</materialsCitation>
; 
<materialsCitation id="49B4EA3FFFAD033FCD0EF98ACA80F91A" pageId="6" pageNumber="7">GIN 952/1159 — isolated damaged crown part of the left i1</materialsCitation>
; 
<materialsCitation id="49B4EA3FFFAD033FCDB4F96ACAC7F93A" pageId="6" pageNumber="7">GIN 952/ 1160 — isolated right I1 with damaged root;</materialsCitation>
<materialsCitation id="49B4EA3FFFAD033FCDF7F94ACA1BF95A" pageId="6" pageNumber="7">GIN 952/1161 — isolated left I1.</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="B1C6B3E9FFAD033FCFDEF90ACA7EF89A" pageId="6" pageNumber="7" type="description">
<paragraph id="F963E062FFAD033FCFDEF90ACA7EF89A" blockId="6.[192,793,1314,1883]" pageId="6" pageNumber="7">
<emphasis id="CBA83C70FFAD033FCFDEF90AC9BEF97A" bold="true" box="[192,423,1762,1787]" pageId="6" pageNumber="7">Location of types.</emphasis>
<typeStatus id="26675EC0FFAD033FCEAEF90ACA00F97A" box="[432,537,1762,1787]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
and 
<typeStatus id="26675EC0FFAD033FCD46F90ACAD4F97A" box="[600,717,1762,1787]" pageId="6" pageNumber="7" type="paratype">paratypes</typeStatus>
in the collection of GIN, 
<collectingRegion id="3B182E80FFAD033FCE8CF8EAC9EDF89A" box="[402,500,1794,1819]" country="Russia" name="Moscow" pageId="6" pageNumber="7">Moscow</collectingRegion>
(
<collectingCountry id="81CBA0F2FFAD033FCD1CF8EACA45F89A" box="[514,604,1794,1819]" name="Russia" pageId="6" pageNumber="7">Russia</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="B1C6B3E9FFAD033FCFDEF8CAC8E2F8DA" pageId="6" pageNumber="7" type="etymology">
<paragraph id="F963E062FFAD033FCFDEF8CAC8E2F8DA" blockId="6.[192,793,1314,1883]" pageId="6" pageNumber="7">
<emphasis id="CBA83C70FFAD033FCFDEF8CAC957F8BA" bold="true" box="[192,334,1826,1851]" pageId="6" pageNumber="7">Etymology.</emphasis>
As for genus name, after the Ishim river.
</paragraph>
</subSubSection>
<subSubSection id="B1C6B3E9FFAD033FCC56FACACCD3F99A" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="F963E062FFAD033FCC56FACACCD3F99A" blockId="6.[840,1441,1314,1915]" pageId="6" pageNumber="7">
<emphasis id="CBA83C70FFAD033FCC56FACACBADFABA" bold="true" box="[840,948,1314,1339]" pageId="6" pageNumber="7">Material.</emphasis>
<typeStatus id="26675EC0FFAD033FCCA5FACACBECFABA" box="[955,1013,1314,1339]" pageId="6" pageNumber="7">Type</typeStatus>
material and four remains from 
<typeStatus id="26675EC0FFAD033FCA70FACACDB9FABA" box="[1390,1440,1314,1339]" pageId="6" pageNumber="7">type</typeStatus>
locality (GIN 952/1157 — damaged right first lower molar; GIN 952/1400 — damaged left P4; GIN 952/ 1401 — damaged left P4; GIN 952/1402 — damaged left P4); and three rediposited remains (GIN 1115/1148 — worn left I1, crown part; GIN 1115/ 1149 — right i1; GIN 1115/1150 — left i1) from Borki 1A locality (Appendix 2: 1).
</paragraph>
</subSubSection>
<subSubSection id="B1C6B3E9FFAD033FCC56F9CACCCDF9BA" box="[840,1236,1570,1596]" pageId="6" pageNumber="7" type="description">
<paragraph id="F963E062FFAD033FCC56F9CACCCDF9BA" blockId="6.[840,1441,1314,1915]" box="[840,1236,1570,1596]" pageId="6" pageNumber="7">
<emphasis id="CBA83C70FFAD033FCC56F9CACC10F9BA" bold="true" box="[840,1033,1570,1595]" pageId="6" pageNumber="7">Measurements.</emphasis>
See Appendix 4.
</paragraph>
</subSubSection>
<subSubSection id="B1C6B3E9FFAD033FCC56F9AACC6DF9DA" box="[840,1140,1602,1627]" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="F963E062FFAD033FCC56F9AACC6DF9DA" blockId="6.[840,1441,1314,1915]" box="[840,1140,1602,1627]" pageId="6" pageNumber="7">
<emphasis id="CBA83C70FFAD033FCC56F9AACBD6F9DA" bold="true" box="[840,975,1602,1627]" pageId="6" pageNumber="7">Diagnosis.</emphasis>
As for genus.
</paragraph>
</subSubSection>
<subSubSection id="B1C6B3E9FFAD0330CC56F98AC9BBFB52" lastPageId="9" lastPageNumber="10" pageId="6" pageNumber="7" type="description">
<paragraph id="F963E062FFAD0331CC56F98AC911FD1E" blockId="6.[840,1441,1314,1915]" lastBlockId="8.[192,792,166,1887]" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
<emphasis id="CBA83C70FFAD033FCC56F98ACBFBF9FA" bold="true" box="[840,994,1634,1659]" pageId="6" pageNumber="7">Description.</emphasis>
I1 has an elongated massive crown; the large hatchet-like talon bears the sharp cutting edge and expressed narrow, deep notch; the apex of the incisor is not fissident. The buccal cingulum is wide and well expressed without a bulge and reaches half of the crown height. The root is notably shorter and more slender than the crown (
<figureCitation id="61E7FCE7FFAD033FCA6EF8CACBD9F8DA" captionStart="FIGURE 3" captionStartId="6.[198,289,1112,1135]" captionTargetBox="[203,1433,167,1084]" captionTargetId="figure-302@6.[190,1439,162,1096]" captionTargetPageId="6" captionText="FIGURE 3. The characters of the first lower incisors of Paranourosorex gigas from ANR/1A (GIN 1112/1108; A1, lateral, A2, medial views), Ishimosorex ishimiensis gen. et sp. nov. from BRK/1A (GIN 1115/1149, paratype; B1, lateral, B2, medial views), Crusafontina endemica Spanish from Can Llobateres 1 locality (C, CL1 2217, see van Dam, 2004: 746; in lateral view), C. kormosi (D, Polgárdi 4, in lateral view) and A. oligodon (E, Polgárdi 4, in lateral view). Scale bars equal 1 mm; C, unscaled. Abbreviations see in Figure 1." figureDoi="http://doi.org/10.5281/zenodo.11063300" httpUri="https://zenodo.org/record/11063300/files/figure.png" pageId="6" pageNumber="7">Figure 3A, B</figureCitation>
). Only three damaged fragments of the left P4 without lingual part are known. The buccal part of these teeth has a relatively short postparacrista; the small parastyle is weakly separated from the paracone base. M1 has a subquadrate occlusal shape with an approximately equal length of the lingual and buccal sides (BL/LL ratio; see Appendix 4); the mesostyle is well-developed and buccally protruded outward of the M1 base level; the buccal crests (preparacrista + ectoloph + postmetacrista) are represented as an expressed W-shaped line; the metaloph is short and separated from the metacone base; the hypocone is rounded distinct cusp and shifted to the lingual margin of the tooth; the weak ridge lays across the hypocone tip obliquely. The posterior emargination of M1 is relatively deep; the hypoconal flange is narrow and elongated backward (
<figureCitation id="61E7FCE7FFAC033ECEDFF8ECCA58F89F" box="[449,577,1796,1822]" captionStart="FIGURE 2" captionStartId="5.[192,283,1464,1487]" captionTargetBox="[188,1437,164,1440]" captionTargetId="figure-53@5.[188,1437,164,1441]" captionTargetPageId="5" captionText="FIGURE 2. Teeth and bones of Ishimosorex ishimiensis gen. et sp. nov. from late Miocene Petropavlovsk 1A locality (A–E, G–K) and Borki 1A locality (F). A, GIN 952/1161, left I1 in lateral view (Figure 2A cf. Figure 7A); B, GIN 952/ 1160, right I1 in medial view; C, GIN 952/1152, left fragment of mandibular ramus in posterior view (image displays articular surface of the condylar process); D, ibid., diagrammatic image of mandibular ramus in medial view; E, GIN 952/1159, left i1 in lateral view; F, GIN 1115/1149, right i1 in medial view; G, GIN 952/1155, left M1 in occlusal view (G1, pure view; G2, augmented view) (Figure 2G cf. Figure 7H); H, GIN 952/1153, right dentary fragment with p4–m1 and anterior alveolus of m2; J, GIN 952/1158, right m1 in occlusal view; K, GIN 952/1154, left m2 in occlusal view. Star indicates reversed image. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11063298" httpUri="https://zenodo.org/record/11063298/files/figure.png" pageId="7" pageNumber="8">Figure 2G</figureCitation>
). The relative distance between the hypocone and the posterior margin of the flange is distinctly larger than the other taxa have (see comparisons below). The crown of i1 is two times longer than the root (along the lateral side). The medial groove of the root is long and overreaches the crown-root junction. The cutting edge of i1 is bicuspidate; the basal (first) cuspule is slender without any bulge-like expression (opposite state to 
<taxonomicName id="3EDC9BE1FFAC033ECB68FA2CCD14FA5C" authorityName="Gibert" authorityYear="1974" box="[1142,1293,1476,1501]" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFAC033ECB68FA2CCD14FA5C" box="[1142,1293,1476,1501]" italics="true" pageId="7" pageNumber="8">Crusafontina</emphasis>
</taxonomicName>
); the distal cuspule is also weak expressed; an ectocingulid is absent (
<figureCitation id="61E7FCE7FFAC033ECCB0F9ECCC4DF99C" box="[942,1108,1540,1565]" captionStart="FIGURE 2" captionStartId="5.[192,283,1464,1487]" captionTargetBox="[188,1437,164,1440]" captionTargetId="figure-53@5.[188,1437,164,1441]" captionTargetPageId="5" captionText="FIGURE 2. Teeth and bones of Ishimosorex ishimiensis gen. et sp. nov. from late Miocene Petropavlovsk 1A locality (A–E, G–K) and Borki 1A locality (F). A, GIN 952/1161, left I1 in lateral view (Figure 2A cf. Figure 7A); B, GIN 952/ 1160, right I1 in medial view; C, GIN 952/1152, left fragment of mandibular ramus in posterior view (image displays articular surface of the condylar process); D, ibid., diagrammatic image of mandibular ramus in medial view; E, GIN 952/1159, left i1 in lateral view; F, GIN 1115/1149, right i1 in medial view; G, GIN 952/1155, left M1 in occlusal view (G1, pure view; G2, augmented view) (Figure 2G cf. Figure 7H); H, GIN 952/1153, right dentary fragment with p4–m1 and anterior alveolus of m2; J, GIN 952/1158, right m1 in occlusal view; K, GIN 952/1154, left m2 in occlusal view. Star indicates reversed image. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11063298" httpUri="https://zenodo.org/record/11063298/files/figure.png" pageId="7" pageNumber="8">Figure 2E, F</figureCitation>
). p4 has a massive and rounded crown; the central (single) wear facet is a rounded or ‘spot-shaped’ (opposite a ‘commashaped’ in 
<emphasis id="CBA83C70FFAC033ECCD5F98CCC40F9FC" box="[971,1113,1636,1661]" italics="true" pageId="7" pageNumber="8">Crisafontina</emphasis>
; see 
<bibRefCitation id="9D4D9D93FFAC033ECB83F98CCD8CF9FC" author="Van Dam, J. A." box="[1181,1429,1636,1661]" pageId="7" pageNumber="8" pagination="741 - 764" refId="ref17856" refString="Van Dam, J. A. 2004. Anourosoricini (Mammalia: Soricidae) from the Mediterranean region: A pre-Quaternary example of recurrent climate-controlled north - south range shifting. Journal of Paleontology, 78: 741 - 764. https: // www. jstor. org / stable / 4094903" type="journal article" year="2004">van Dam, 2004: 744</bibRefCitation>
). The ecto- and entocingulids are well developed; the ectocindulid continuously passed to postcingulid (
<figureCitation id="61E7FCE7FFAC033ECC8DF92CCC0AF95C" box="[915,1043,1732,1757]" captionStart="FIGURE 4" captionStartId="7.[196,287,1008,1031]" captionTargetBox="[199,1432,178,975]" captionTargetId="figure-328@7.[190,1439,168,984]" captionTargetPageId="7" captionText="FIGURE 4. Schematic image of the fourth lower premolar of Ishimosorex ishimiensis gen. et sp. nov. (A, GIN 952/ 1153, PPL/1A), Crusafontina kormosi (B, Polgárdi 4), Paranourosorex intermedius sp. nov. (C, GIN 948/1051, NST/ 1A), Paranourosorex gigas (D, GIN 1118/1011, PSH/1B), Anourosorex squamipes (E, ZIN 98253, Recent), Crusafontina fastigata (F, AG5A from Los Aguanaces 5A, Teruel Basin, Spain by van Dam, 2004: figs. 4-1), Amblycoptus oligodon (G, Polgárdi 4) and Amblycoptus jessiae (H, KS 3157 from Las Casiones, Teruel Basin, Spain by van Dam, 2004: figs. 5-11). Abbreviations: a, concavity; add, additional crown elements (1, crest; 2, cusplet); ccr, central crest; eccd, ectocingulid; encd, entocingulid; pcd, postcingulid; plb, posterolingual basin (1, shallow, weak; 2, expressed); wen, wrinkled enamel; wft, wear facet (1, spot-like; 2, comma-like). Star indicates reversed image. Scale bars are 1 mm; F, H unscaled." figureDoi="http://doi.org/10.5281/zenodo.11063302" httpUri="https://zenodo.org/record/11063302/files/figure.png" pageId="7" pageNumber="8">Figure 4A</figureCitation>
). p4 bears a shallow posterolingual basin and extremely weak the central crest, which doesn’t reach the postcingulid. The lower molars are graded in size: the first molar is the largest, the second is moderately smaller than the first (
<figureCitation id="61E7FCE7FFAC033ECC50F88CCBE8F8FC" box="[846,1009,1892,1917]" captionStart="FIGURE 2" captionStartId="5.[192,283,1464,1487]" captionTargetBox="[188,1437,164,1440]" captionTargetId="figure-53@5.[188,1437,164,1441]" captionTargetPageId="5" captionText="FIGURE 2. Teeth and bones of Ishimosorex ishimiensis gen. et sp. nov. from late Miocene Petropavlovsk 1A locality (A–E, G–K) and Borki 1A locality (F). A, GIN 952/1161, left I1 in lateral view (Figure 2A cf. Figure 7A); B, GIN 952/ 1160, right I1 in medial view; C, GIN 952/1152, left fragment of mandibular ramus in posterior view (image displays articular surface of the condylar process); D, ibid., diagrammatic image of mandibular ramus in medial view; E, GIN 952/1159, left i1 in lateral view; F, GIN 1115/1149, right i1 in medial view; G, GIN 952/1155, left M1 in occlusal view (G1, pure view; G2, augmented view) (Figure 2G cf. Figure 7H); H, GIN 952/1153, right dentary fragment with p4–m1 and anterior alveolus of m2; J, GIN 952/1158, right m1 in occlusal view; K, GIN 952/1154, left m2 in occlusal view. Star indicates reversed image. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11063298" httpUri="https://zenodo.org/record/11063298/files/figure.png" pageId="7" pageNumber="8">Figure 2H–K</figureCitation>
); the third molar is unknown. The hypoconid of m1 and m2 is protruded buccally and overhangs the base of the crown; the entostylid is presented as the small distinct bulge and does not reach the entoconid level. The entocristid is short and steeply descends to the metaconid base. The talonid basin opens posterolingually between the entoconid and entostylid through the posterolateral groove and lingually between the end of the entocristid and the metaconid base through the transverse groove. The narrow and well-distinguished ectocingulid does not reach the anterior side of the tooth (
<figureCitation id="61E7FCE7FFA30331CE15FDEEC9BEFD9E" box="[267,423,518,543]" captionStart="FIGURE 2" captionStartId="5.[192,283,1464,1487]" captionTargetBox="[188,1437,164,1440]" captionTargetId="figure-53@5.[188,1437,164,1441]" captionTargetPageId="5" captionText="FIGURE 2. Teeth and bones of Ishimosorex ishimiensis gen. et sp. nov. from late Miocene Petropavlovsk 1A locality (A–E, G–K) and Borki 1A locality (F). A, GIN 952/1161, left I1 in lateral view (Figure 2A cf. Figure 7A); B, GIN 952/ 1160, right I1 in medial view; C, GIN 952/1152, left fragment of mandibular ramus in posterior view (image displays articular surface of the condylar process); D, ibid., diagrammatic image of mandibular ramus in medial view; E, GIN 952/1159, left i1 in lateral view; F, GIN 1115/1149, right i1 in medial view; G, GIN 952/1155, left M1 in occlusal view (G1, pure view; G2, augmented view) (Figure 2G cf. Figure 7H); H, GIN 952/1153, right dentary fragment with p4–m1 and anterior alveolus of m2; J, GIN 952/1158, right m1 in occlusal view; K, GIN 952/1154, left m2 in occlusal view. Star indicates reversed image. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11063298" httpUri="https://zenodo.org/record/11063298/files/figure.png" pageId="8" pageNumber="9">Figure 2J, K</figureCitation>
). The buccal side of m1 has a slightly wrinkled enamel surface. The ectocingulid of m2 is well developed along the tooth base with the plate-like extension in the first third (paraconid level).
</paragraph>
<caption id="ADA3B0EAFFAC033ECFDAFC18C98AFB73" ID-DOI="http://doi.org/10.5281/zenodo.11063302" ID-Zenodo-Dep="11063302" httpUri="https://zenodo.org/record/11063302/files/figure.png" pageId="7" pageNumber="8" startId="7.[196,287,1008,1031]" targetBox="[199,1432,178,975]" targetPageId="7" targetType="figure">
<paragraph id="F963E062FFAC033ECFDAFC18C98AFB73" blockId="7.[196,1439,1008,1266]" pageId="7" pageNumber="8">
<emphasis id="CBA83C70FFAC033ECFDAFC18C925FB89" bold="true" box="[196,316,1008,1032]" pageId="7" pageNumber="8">FIGURE 4.</emphasis>
Schematic image of the fourth lower premolar of 
<taxonomicName id="3EDC9BE1FFAC033ECC44FC19CC40FB89" authority="Zazhigin &amp; Voyta, 2022" authorityName="Zazhigin &amp; Voyta" authorityYear="2022" box="[858,1113,1009,1032]" class="Mammalia" family="Soricidae" genus="Ishimosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="ishimiensis" status="gen. et sp. nov.">
<emphasis id="CBA83C70FFAC033ECC44FC19CC40FB89" box="[858,1113,1009,1032]" italics="true" pageId="7" pageNumber="8">Ishimosorex ishimiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D09B810BFFAC033ECB7CFC19CD15FB89" box="[1122,1292,1009,1032]" pageId="7" pageNumber="8" rank="species">gen. et sp. nov.</taxonomicNameLabel>
(
<emphasis id="CBA83C70FFAC033ECA05FC19CD35FB89" bold="true" box="[1307,1324,1009,1032]" pageId="7" pageNumber="8">A</emphasis>
, GIN 952/ 1153, PPL/1A), 
<taxonomicName id="3EDC9BE1FFAC033ECE70FBE6CA4BFBA4" baseAuthorityName="Bachmayer and Wilson" baseAuthorityYear="1970" box="[366,594,1038,1061]" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="kormosi">
<emphasis id="CBA83C70FFAC033ECE70FBE6CA4BFBA4" box="[366,594,1038,1061]" italics="true" pageId="7" pageNumber="8">Crusafontina kormosi</emphasis>
</taxonomicName>
(
<emphasis id="CBA83C70FFAC033ECD7DFBE6CA6DFBA4" bold="true" box="[611,628,1038,1061]" pageId="7" pageNumber="8">B</emphasis>
, Polgárdi 4), 
<taxonomicName id="3EDC9BE1FFAC033ECC19FBE6CC2FFBA4" authority="Zazhigin &amp; Voyta, 2022" authorityName="Zazhigin &amp; Voyta" authorityYear="2022" box="[775,1078,1038,1061]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="intermedius" status="sp. nov.">
<emphasis id="CBA83C70FFAC033ECC19FBE6CC2FFBA4" box="[775,1078,1038,1061]" italics="true" pageId="7" pageNumber="8">Paranourosorex intermedius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D09B810BFFAC033ECB20FBE6CC8AFBA4" box="[1086,1171,1038,1061]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
(
<emphasis id="CBA83C70FFAC033ECBBCFBE6CCAAFBA4" bold="true" box="[1186,1203,1038,1061]" pageId="7" pageNumber="8">C</emphasis>
, GIN 948/1051, NST/ 1A), 
<taxonomicName id="3EDC9BE1FFAC033ECFE9FBC3C9F8FBC3" authorityName="Rzebik-Kowalska" authorityYear="1975" box="[247,481,1067,1090]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="gigas">
<emphasis id="CBA83C70FFAC033ECFE9FBC3C9F8FBC3" box="[247,481,1067,1090]" italics="true" pageId="7" pageNumber="8">Paranourosorex gigas</emphasis>
</taxonomicName>
(
<emphasis id="CBA83C70FFAC033ECEEFFBC3CA1BFBC3" bold="true" box="[497,514,1067,1090]" pageId="7" pageNumber="8">D</emphasis>
, GIN 1118/1011, PSH/1B), 
<taxonomicName id="3EDC9BE1FFAC033ECC38FBC3CC30FBC3" authorityName="Milne-Edwards" authorityYear="1870" box="[806,1065,1067,1090]" class="Mammalia" family="Soricidae" genus="Anourosorex" kingdom="Animalia" order="Soricomorpha" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="squamipes">
<emphasis id="CBA83C70FFAC033ECC38FBC3CC30FBC3" box="[806,1065,1067,1090]" italics="true" pageId="7" pageNumber="8">Anourosorex squamipes</emphasis>
</taxonomicName>
(
<emphasis id="CBA83C70FFAC033ECB26FBC3CC51FBC3" bold="true" box="[1080,1096,1067,1090]" pageId="7" pageNumber="8">E</emphasis>
, ZIN 98253, Recent), 
<taxonomicName id="3EDC9BE1FFAC033ECA2BFBC3C954FBDE" authorityName="van Dam" authorityYear="2004" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="fastigata">
<emphasis id="CBA83C70FFAC033ECA2BFBC3C954FBDE" italics="true" pageId="7" pageNumber="8">Crusafontina fastigata</emphasis>
</taxonomicName>
(
<emphasis id="CBA83C70FFAC033ECE43FBA1C970FBE1" bold="true" box="[349,361,1097,1120]" pageId="7" pageNumber="8">F</emphasis>
, AG5A from Los Aguanaces 5A, Teruel Basin, Spain by 
<bibRefCitation id="9D4D9D93FFAC033ECCD2FBA1CC6AFBE1" author="Van Dam, J. A." box="[972,1139,1097,1120]" pageId="7" pageNumber="8" pagination="741 - 764" refId="ref17856" refString="Van Dam, J. A. 2004. Anourosoricini (Mammalia: Soricidae) from the Mediterranean region: A pre-Quaternary example of recurrent climate-controlled north - south range shifting. Journal of Paleontology, 78: 741 - 764. https: // www. jstor. org / stable / 4094903" type="journal article" year="2004">van Dam, 2004</bibRefCitation>
: figs. 4-1), 
<taxonomicName id="3EDC9BE1FFAC033ECBF3FBA1C91EFBFC" class="Mammalia" family="Soricidae" genus="Amblycoptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="oligodon">
<emphasis id="CBA83C70FFAC033ECBF3FBA1C91EFBFC" italics="true" pageId="7" pageNumber="8">Amblycoptus oligodon</emphasis>
</taxonomicName>
(
<emphasis id="CBA83C70FFAC033ECE06FB8EC931FBFC" bold="true" box="[280,296,1126,1149]" pageId="7" pageNumber="8">G</emphasis>
, Polgárdi 4) and 
<taxonomicName id="3EDC9BE1FFAC033ECEF9FB8ECADDFBFC" box="[487,708,1126,1149]" class="Mammalia" family="Soricidae" genus="Amblycoptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="jessiae">
<emphasis id="CBA83C70FFAC033ECEF9FB8ECADDFBFC" box="[487,708,1126,1149]" italics="true" pageId="7" pageNumber="8">Amblycoptus jessiae</emphasis>
</taxonomicName>
(
<emphasis id="CBA83C70FFAC033ECDCBFB8ECAFFFBFC" bold="true" box="[725,742,1126,1149]" pageId="7" pageNumber="8">H</emphasis>
, KS 3157 from Las Casiones, Teruel Basin, Spain by 
<bibRefCitation id="9D4D9D93FFAC033ECA2BFB8EC8E4FB1B" author="Van Dam, J. A." pageId="7" pageNumber="8" pagination="741 - 764" refId="ref17856" refString="Van Dam, J. A. 2004. Anourosoricini (Mammalia: Soricidae) from the Mediterranean region: A pre-Quaternary example of recurrent climate-controlled north - south range shifting. Journal of Paleontology, 78: 741 - 764. https: // www. jstor. org / stable / 4094903" type="journal article" year="2004">van Dam, 2004</bibRefCitation>
: figs. 5-11). Abbreviations: a, concavity; add, additional crown elements (1, crest; 2, cusplet); ccr, central crest; eccd, ectocingulid; encd, entocingulid; pcd, postcingulid; plb, posterolingual basin (1, shallow, weak; 2, expressed); wen, wrinkled enamel; wft, wear facet (1, spot-like; 2, comma-like). Star indicates reversed image. Scale bars are 1 mm; F, H unscaled.
</paragraph>
</caption>
<paragraph id="F963E062FFA30331CFE4FD4EC9CBFC5E" blockId="8.[192,792,166,1887]" pageId="8" pageNumber="9">
The horizontal ramus of the lower jaw is narrow; the small mental foramen is situated slightly backwards of the m1 protoconid level without groove. The mandibular ramus is relatively low; its anterior border is notably tilted backwards. The internal fossa of the temporal muscle ('internal fossa') is moderately developed. The condylar process bears the widely divided upper and lower articular facets and a moderately broad interarticular area (
<figureCitation id="61E7FCE7FFA30331CE36FC2EC9DCFC5E" box="[296,453,966,992]" captionStart="FIGURE 2" captionStartId="5.[192,283,1464,1487]" captionTargetBox="[188,1437,164,1440]" captionTargetId="figure-53@5.[188,1437,164,1441]" captionTargetPageId="5" captionText="FIGURE 2. Teeth and bones of Ishimosorex ishimiensis gen. et sp. nov. from late Miocene Petropavlovsk 1A locality (A–E, G–K) and Borki 1A locality (F). A, GIN 952/1161, left I1 in lateral view (Figure 2A cf. Figure 7A); B, GIN 952/ 1160, right I1 in medial view; C, GIN 952/1152, left fragment of mandibular ramus in posterior view (image displays articular surface of the condylar process); D, ibid., diagrammatic image of mandibular ramus in medial view; E, GIN 952/1159, left i1 in lateral view; F, GIN 1115/1149, right i1 in medial view; G, GIN 952/1155, left M1 in occlusal view (G1, pure view; G2, augmented view) (Figure 2G cf. Figure 7H); H, GIN 952/1153, right dentary fragment with p4–m1 and anterior alveolus of m2; J, GIN 952/1158, right m1 in occlusal view; K, GIN 952/1154, left m2 in occlusal view. Star indicates reversed image. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11063298" httpUri="https://zenodo.org/record/11063298/files/figure.png" pageId="8" pageNumber="9">Figure 2C, D</figureCitation>
).
</paragraph>
<paragraph id="F963E062FFA30331CFDEFC0ECCE0FE5E" blockId="8.[192,792,166,1887]" lastBlockId="8.[840,1441,166,1887]" pageId="8" pageNumber="9">
<emphasis id="CBA83C70FFA30331CFDEFC0ECA09FC7E" bold="true" box="[192,528,998,1023]" pageId="8" pageNumber="9">Association of fragments.</emphasis>
The studied remains from PPL/1A were each matched using the size recovery approach and several RSTs, 
<taxonomicName id="3EDC9BE1FFA30331CD8BFBCEC8F2FBDE" authorityName="Milne-Edwards" authorityYear="1870" class="Mammalia" family="Soricidae" genus="Anourosorex" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="squamipes">
<emphasis id="CBA83C70FFA30331CD8BFBCEC8F2FBDE" italics="true" pageId="8" pageNumber="9">A. squamipes</emphasis>
</taxonomicName>
and two 
<taxonomicName id="3EDC9BE1FFA30331CE4BFBAECA0BFBDE" authorityName="Rzebik-Kowalska" authorityYear="1975" box="[341,530,1094,1119]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFA30331CE4BFBAECA0BFBDE" box="[341,530,1094,1119]" italics="true" pageId="8" pageNumber="9">Paranourosorex</emphasis>
</taxonomicName>
samples (Appendix 5: tables 5.1 and 5.2) for the following pairs of dental and mandibular elements and measurements: m1/ M1: measurements L(m1)/BL(M1); m1/I1: measurements L(m1)/H(I1); I1/i1: measurements L(I1)/ L(i1); m1/mandibular ramus: measuremets L(m1)/ COR and L(m1)/MRWc. Most of the matched pairs displayed compliance; e.g., the matched m1 and M1 corresponded to each other in size based on the RST ratio. Noncompliance (pink blocks in Table 5.2 of Appendix 5) was revealed for matched upper and lower incisors from PPL/1A (and BRK/1A), i.e., the observed dimension of the lower incisor was less than the supposed dimension calculated based on the 
<taxonomicName id="3EDC9BE1FFA30331CE7FF9EEC9E1F99E" authorityName="Zazhigin &amp; Voyta" authorityYear="2022" box="[353,504,1542,1567]" class="Mammalia" family="Soricidae" genus="Anourosorex" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFA30331CE7FF9EEC9E1F99E" box="[353,504,1542,1567]" italics="true" pageId="8" pageNumber="9">Anourosorex</emphasis>
</taxonomicName>
ratio (Appendix 5: Table 5.1). This noncompliance between calculated and observed incisor sizes could be a specific feature of the particular species, which can be explained in terms allometric heterochrony (sense Mitteroecker et al., 2005: 250). This feature (the ratio between upper and lower incisors) in 
<taxonomicName id="3EDC9BE1FFA30331CD0BF92EC8F9F97E" authority="Zazhigin &amp; Voyta, 2022" authorityName="Zazhigin &amp; Voyta" authorityYear="2022" class="Mammalia" family="Soricidae" genus="Ishimosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="ishimiensis" status="gen. et sp. nov.">
<emphasis id="CBA83C70FFA30331CD0BF92EC8F9F97E" italics="true" pageId="8" pageNumber="9">Ishimosorex ishimiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D09B810BFFA30331CFF3F90EC9AAF97E" box="[237,435,1766,1791]" pageId="8" pageNumber="9" rank="species">gen. et sp. nov.</taxonomicNameLabel>
differs from 
<taxonomicName id="3EDC9BE1FFA30331CD48F90ECAF4F97E" authorityName="Zazhigin &amp; Voyta" authorityYear="2022" box="[598,749,1766,1791]" class="Mammalia" family="Soricidae" genus="Anourosorex" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFA30331CD48F90ECAF4F97E" box="[598,749,1766,1791]" italics="true" pageId="8" pageNumber="9">Anourosorex</emphasis>
</taxonomicName>
. In addition, the short lower incisor of 
<taxonomicName id="3EDC9BE1FFA30331CD96F8EEC95BF8BE" authority="Zazhigin &amp; Voyta, 2022" authorityName="Zazhigin &amp; Voyta" authorityYear="2022" class="Mammalia" family="Soricidae" genus="Ishimosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="ishimiensis" status="gen. et sp. nov.">
<emphasis id="CBA83C70FFA30331CD96F8EEC95BF8BE" italics="true" pageId="8" pageNumber="9">Ishimosorex ishimiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D09B810BFFA30331CE52F8CECA11F8BE" box="[332,520,1830,1855]" pageId="8" pageNumber="9" rank="species">gen. et sp. nov.</taxonomicNameLabel>
can probably be compensated through a relatively long mandible protraction during a chewing cycle (for details, see below remark on a phenomenon of ‘cutting edges straightening’). Proof of this is the best value for the ratio L(m1)/L(i1); i.e., the 
<taxonomicName id="3EDC9BE1FFA30331CBBBFEEECD25FE9E" authorityName="Zazhigin &amp; Voyta" authorityYear="2022" box="[1189,1340,262,287]" class="Mammalia" family="Soricidae" genus="Anourosorex" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFA30331CBBBFEEECD25FE9E" box="[1189,1340,262,287]" italics="true" pageId="8" pageNumber="9">Anourosorex</emphasis>
</taxonomicName>
ratio for L(m1)/L(i1) corresponds more to the observed ratio in 
<taxonomicName id="3EDC9BE1FFA30331CC78FEAECC98FEDE" authority="Zazhigin &amp; Voyta, 2022" authorityName="Zazhigin &amp; Voyta" authorityYear="2022" box="[870,1153,326,351]" class="Mammalia" family="Soricidae" genus="Ishimosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="ishimiensis" status="gen. et sp. nov.">
<emphasis id="CBA83C70FFA30331CC78FEAECC98FEDE" box="[870,1153,326,351]" italics="true" pageId="8" pageNumber="9">Ishimosorex ishimiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D09B810BFFA30331CB94FEAECD5AFEDE" box="[1162,1347,326,351]" pageId="8" pageNumber="9" rank="species">gen. et sp. nov.</taxonomicNameLabel>
than to the ratio L(I1)/L(i1). Thus, the studied remains from PPL/1A can be treated here as belonging to 
<taxonomicName id="3EDC9BE1FFA30331CA70FE6ECC21FE3E" authority="Zazhigin &amp; Voyta, 2022" authorityName="Zazhigin &amp; Voyta" authorityYear="2022" class="Mammalia" family="Soricidae" genus="Ishimosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="ishimiensis" status="gen. et sp. nov.">
<emphasis id="CBA83C70FFA30331CA70FE6ECC21FE3E" italics="true" pageId="8" pageNumber="9">Ishimosorex ishimiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D09B810BFFA30331CB20FE4ECCE8FE3E" box="[1086,1265,422,447]" pageId="8" pageNumber="9" rank="species">gen. et sp. nov.</taxonomicNameLabel>
based on metric matching (Appendix 5: Table 5.2).
</paragraph>
<paragraph id="F963E062FFA30331CC56FE0ECBA2FAFE" blockId="8.[840,1441,166,1887]" pageId="8" pageNumber="9">
<emphasis id="CBA83C70FFA30331CC56FE0ECBE0FD81" bold="true" box="[840,1017,486,512]" pageId="8" pageNumber="9">Comparisons.</emphasis>
<taxonomicName id="3EDC9BE1FFA30331CB1FFE0ECD03FE7E" authority="Zazhigin &amp; Voyta, 2022" authorityName="Zazhigin &amp; Voyta" authorityYear="2022" box="[1025,1306,486,511]" class="Mammalia" family="Soricidae" genus="Ishimosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="ishimiensis" status="gen. et sp. nov.">
<emphasis id="CBA83C70FFA30331CB1FFE0ECD03FE7E" box="[1025,1306,486,511]" italics="true" pageId="8" pageNumber="9">Ishimosorex ishimiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D09B810BFFA30331CA3DFE0ECB60FD9E" pageId="8" pageNumber="9" rank="species">gen. et sp. nov.</taxonomicNameLabel>
(L(p4) = 
<quantity id="3E244D87FFA30331CCFAFDEECC56FD9E" box="[996,1103,518,543]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="8" pageNumber="9" unit="mm" value="1.75">1.75 mm</quantity>
; L(m1) = 
<quantity id="3E244D87FFA30331CBA3FDEECD73FD9E" box="[1213,1386,518,543]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" metricValueMax="2.73" metricValueMin="2.47" pageId="8" pageNumber="9" unit="mm" value="2.6" valueMax="2.73" valueMin="2.47">2.47–2.73 mm</quantity>
, see Appendix 4) differs in larger size from 
<taxonomicName id="3EDC9BE1FFA30331CA14FDCECD86FDBE" box="[1290,1439,550,575]" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="endemica">
<emphasis id="CBA83C70FFA30331CA14FDCECD86FDBE" box="[1290,1439,550,575]" italics="true" pageId="8" pageNumber="9">C. endemica</emphasis>
</taxonomicName>
(L(m1) = 
<quantity id="3E244D87FFA30331CCABFDAECC7DFDDE" box="[949,1124,582,607]" metricMagnitude="-3" metricUnit="m" metricValue="2.01" metricValueMax="2.12" metricValueMin="1.9" pageId="8" pageNumber="9" unit="mm" value="2.01" valueMax="2.12" valueMin="1.9">1.90–2.12 mm</quantity>
, see 
<bibRefCitation id="9D4D9D93FFA30331CBBCFDAECD45FDDE" author="Van Dam, J. A." box="[1186,1372,582,607]" pageId="8" pageNumber="9" pagination="741 - 764" refId="ref17856" refString="Van Dam, J. A. 2004. Anourosoricini (Mammalia: Soricidae) from the Mediterranean region: A pre-Quaternary example of recurrent climate-controlled north - south range shifting. Journal of Paleontology, 78: 741 - 764. https: // www. jstor. org / stable / 4094903" type="journal article" year="2004">van Dam, 2004</bibRefCitation>
: table 4), 
<taxonomicName id="3EDC9BE1FFA30331CC67FD8ECB95FD1E" authority="(Mayr and Fahlbusch, 1975)" baseAuthorityName="Mayr and Fahlbusch" baseAuthorityYear="1975" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="exculta">
<emphasis id="CBA83C70FFA30331CC67FD8ECC6EFDFE" box="[889,1143,614,639]" italics="true" pageId="8" pageNumber="9">Crusafontina exculta</emphasis>
(Mayr and Fahlbusch, 1975)
</taxonomicName>
(L(m1) = 
<quantity id="3E244D87FFA30331CB1DFD6ECCABFD1E" box="[1027,1202,646,671]" metricMagnitude="-3" metricUnit="m" metricValue="2.01" metricValueMax="2.21" metricValueMin="1.81" pageId="8" pageNumber="9" unit="mm" value="2.01" valueMax="2.21" valueMin="1.81">1.81–2.21 mm</quantity>
, see Prieto and van Dam, 2012), 
<taxonomicName id="3EDC9BE1FFA30331CCFCFD4ECDB9FD3E" authority="van Dam, 2004" authorityName="van Dam" authorityYear="2004" box="[994,1440,678,703]" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="fastigata">
<emphasis id="CBA83C70FFA30331CCFCFD4ECCFDFD3E" box="[994,1252,678,703]" italics="true" pageId="8" pageNumber="9">Crusafontina fastigata</emphasis>
<bibRefCitation id="9D4D9D93FFA30331CBF2FD4ECDB9FD3E" author="Van Dam, J. A." box="[1260,1440,678,703]" pageId="8" pageNumber="9" pagination="741 - 764" refId="ref17856" refString="Van Dam, J. A. 2004. Anourosoricini (Mammalia: Soricidae) from the Mediterranean region: A pre-Quaternary example of recurrent climate-controlled north - south range shifting. Journal of Paleontology, 78: 741 - 764. https: // www. jstor. org / stable / 4094903" type="journal article" year="2004">van Dam, 2004</bibRefCitation>
</taxonomicName>
(L(p4) = 
<quantity id="3E244D87FFA30331CCA2FD2ECC6BFD5E" box="[956,1138,710,735]" metricMagnitude="-3" metricUnit="m" metricValue="1.435" metricValueMax="1.44" metricValueMin="1.43" pageId="8" pageNumber="9" unit="mm" value="1.435" valueMax="1.44" valueMin="1.43">1.43–1.44 mm</quantity>
, ibid.) and 
<taxonomicName id="3EDC9BE1FFA30331CA17FD2ECD3AFD7E" authority="(Hutchinson and Bown, 1980)" baseAuthorityName="Hutchinson and Bown" baseAuthorityYear="1980" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="minima">
<emphasis id="CBA83C70FFA30331CA17FD2ECB87FD7E" italics="true" pageId="8" pageNumber="9">Crusafontina minima</emphasis>
(Hutchinson and Bown, 1980)
</taxonomicName>
(L(m1) = 
<quantity id="3E244D87FFA30331CC56FCEECBEEFC9E" box="[840,1015,774,799]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="1.93" metricValueMin="1.77" pageId="8" pageNumber="9" unit="mm" value="1.85" valueMax="1.93" valueMin="1.77">1.77–1.93 mm</quantity>
, see Bown, 1980: table VI). 
<taxonomicName id="3EDC9BE1FFA30331CA57FCEECC05FCBE" authority="Zazhigin &amp; Voyta, 2022" authorityName="Zazhigin &amp; Voyta" authorityYear="2022" class="Mammalia" family="Soricidae" genus="Ishimosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="ishimiensis" status="gen. et sp. nov.">
<emphasis id="CBA83C70FFA30331CA57FCEECC05FCBE" italics="true" pageId="8" pageNumber="9">Ishimosorex ishimiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D09B810BFFA30331CB30FCCECD18FCBE" box="[1070,1281,806,831]" pageId="8" pageNumber="9" rank="species">gen. et sp. nov.</taxonomicNameLabel>
differs from 
<taxonomicName id="3EDC9BE1FFA30331CC56FCAECD23FCDE" authority="van Dam, 2004" authorityName="van Dam" authorityYear="2004" box="[840,1338,838,863]" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="vandeweerdi">
<emphasis id="CBA83C70FFA30331CC56FCAECC67FCDE" box="[840,1150,838,863]" italics="true" pageId="8" pageNumber="9">Crusafontina vandeweerdi</emphasis>
<bibRefCitation id="9D4D9D93FFA30331CB9BFCAECD23FCDE" author="Van Dam, J. A." box="[1157,1338,838,863]" pageId="8" pageNumber="9" pagination="741 - 764" refId="ref17856" refString="Van Dam, J. A. 2004. Anourosoricini (Mammalia: Soricidae) from the Mediterranean region: A pre-Quaternary example of recurrent climate-controlled north - south range shifting. Journal of Paleontology, 78: 741 - 764. https: // www. jstor. org / stable / 4094903" type="journal article" year="2004">van Dam, 2004</bibRefCitation>
</taxonomicName>
(L(p4) = 
<quantity id="3E244D87FFA30331CC56FC8ECBA1FCFE" box="[840,952,870,895]" metricMagnitude="-3" metricUnit="m" metricValue="2.24" pageId="8" pageNumber="9" unit="mm" value="2.24">2.24 mm</quantity>
, 
<bibRefCitation id="9D4D9D93FFA30331CCD6FC8ECC92FCFE" author="Van Dam, J. A." box="[968,1163,870,895]" pageId="8" pageNumber="9" pagination="741 - 764" refId="ref17856" refString="Van Dam, J. A. 2004. Anourosoricini (Mammalia: Soricidae) from the Mediterranean region: A pre-Quaternary example of recurrent climate-controlled north - south range shifting. Journal of Paleontology, 78: 741 - 764. https: // www. jstor. org / stable / 4094903" type="journal article" year="2004">van Dam, 2004</bibRefCitation>
: table 4) in its smaller size. 
<taxonomicName id="3EDC9BE1FFA30331CC8CFC6ECCAFFC1E" authority="Zazhigin &amp; Voyta, 2022" authorityName="Zazhigin &amp; Voyta" authorityYear="2022" box="[914,1206,902,927]" class="Mammalia" family="Soricidae" genus="Ishimosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="ishimiensis" status="gen. et sp. nov.">
<emphasis id="CBA83C70FFA30331CC8CFC6ECCAFFC1E" box="[914,1206,902,927]" italics="true" pageId="8" pageNumber="9">Ishimosorex ishimiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D09B810BFFA30331CBD7FC6ECDB8FC1E" box="[1225,1441,902,927]" pageId="8" pageNumber="9" rank="species">gen. et sp. nov.</taxonomicNameLabel>
shows the similar teeth size to 
<taxonomicName id="3EDC9BE1FFA30331CBD1FC4ECD07FC5E" authority="(Bachmayer and Wilson, 1970)" baseAuthorityName="Bachmayer and Wilson" baseAuthorityYear="1970" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="kormosi">
<emphasis id="CBA83C70FFA30331CBD1FC4ECB99FC5E" italics="true" pageId="8" pageNumber="9">Crusafontina kormosi</emphasis>
(Bachmayer and Wilson, 1970)
</taxonomicName>
(L(m1) = 
<quantity id="3E244D87FFA30331CC56FC0ECBE0FC7E" box="[840,1017,998,1023]" metricMagnitude="-3" metricUnit="m" metricValue="2.61" metricValueMax="2.84" metricValueMin="2.38" pageId="8" pageNumber="9" unit="mm" value="2.61" valueMax="2.84" valueMin="2.38">2.38–2.84 mm</quantity>
, see Mészáros, 1998: table 3) and 
<taxonomicName id="3EDC9BE1FFA30331CC56FBEECDB8FB9E" authority="(Hutchinson and Bown, 1980)" baseAuthorityName="Hutchinson and Bown" baseAuthorityYear="1980" box="[840,1441,1030,1055]" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="magna">
<emphasis id="CBA83C70FFA30331CC56FBEECC21FB9E" box="[840,1080,1030,1055]" italics="true" pageId="8" pageNumber="9">Crusafontina magna</emphasis>
(Hutchinson and Bown, 1980)
</taxonomicName>
(L(m1) = 
<quantity id="3E244D87FFA30331CCDCFBCECC6FFBBE" box="[962,1142,1062,1087]" metricMagnitude="-3" metricUnit="m" metricValue="2.41" metricValueMax="2.52" metricValueMin="2.3" pageId="8" pageNumber="9" unit="mm" value="2.41" valueMax="2.52" valueMin="2.3">2.30–2.52 mm</quantity>
, Hutchinson and Bown, 1980: table V) but differs from both species in the well-developed mesostyle of M1, which contrary to 
<taxonomicName id="3EDC9BE1FFA30331CC56FB6ECBD1FB1E" baseAuthorityName="Bachmayer and Wilson" baseAuthorityYear="1970" box="[840,968,1158,1183]" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="kormosi">
<emphasis id="CBA83C70FFA30331CC56FB6ECBD1FB1E" box="[840,968,1158,1183]" italics="true" pageId="8" pageNumber="9">C. kormosi</emphasis>
</taxonomicName>
and 
<taxonomicName id="3EDC9BE1FFA30331CB1BFB6ECC60FB1E" baseAuthorityName="Hutchinson and Bown" baseAuthorityYear="1980" box="[1029,1145,1158,1183]" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="magna">
<emphasis id="CBA83C70FFA30331CB1BFB6ECC60FB1E" box="[1029,1145,1158,1183]" italics="true" pageId="8" pageNumber="9">C. magna</emphasis>
</taxonomicName>
shows buccal protrusion of parastyle and metastyle tips (this is also a general difference from 
<taxonomicName id="3EDC9BE1FFA30331CB4BFB2ECD37FB5E" box="[1109,1326,1222,1247]" class="Mammalia" family="Soricidae" genus="Anourosorex" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="CBA83C70FFA30331CB4BFB2ECCF5FB5E" box="[1109,1260,1222,1247]" italics="true" pageId="8" pageNumber="9">Anourosorex</emphasis>
spp.
</taxonomicName>
); in the extremely weak central crest and the spot-shaped wear facet of the p4 opposite well-developed central crest and comma-shaped wear facet of the p4 of 
<taxonomicName id="3EDC9BE1FFA30331CC6EFAAECBE3FADE" baseAuthorityName="Bachmayer and Wilson" baseAuthorityYear="1970" box="[880,1018,1350,1375]" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="kormosi">
<emphasis id="CBA83C70FFA30331CC6EFAAECBE3FADE" box="[880,1018,1350,1375]" italics="true" pageId="8" pageNumber="9">C. kormosi</emphasis>
</taxonomicName>
(
<figureCitation id="61E7FCE7FFA30331CB0CFAAECC66FADE" box="[1042,1151,1350,1375]" captionStart="FIGURE 4" captionStartId="7.[196,287,1008,1031]" captionTargetBox="[199,1432,178,975]" captionTargetId="figure-328@7.[190,1439,168,984]" captionTargetPageId="7" captionText="FIGURE 4. Schematic image of the fourth lower premolar of Ishimosorex ishimiensis gen. et sp. nov. (A, GIN 952/ 1153, PPL/1A), Crusafontina kormosi (B, Polgárdi 4), Paranourosorex intermedius sp. nov. (C, GIN 948/1051, NST/ 1A), Paranourosorex gigas (D, GIN 1118/1011, PSH/1B), Anourosorex squamipes (E, ZIN 98253, Recent), Crusafontina fastigata (F, AG5A from Los Aguanaces 5A, Teruel Basin, Spain by van Dam, 2004: figs. 4-1), Amblycoptus oligodon (G, Polgárdi 4) and Amblycoptus jessiae (H, KS 3157 from Las Casiones, Teruel Basin, Spain by van Dam, 2004: figs. 5-11). Abbreviations: a, concavity; add, additional crown elements (1, crest; 2, cusplet); ccr, central crest; eccd, ectocingulid; encd, entocingulid; pcd, postcingulid; plb, posterolingual basin (1, shallow, weak; 2, expressed); wen, wrinkled enamel; wft, wear facet (1, spot-like; 2, comma-like). Star indicates reversed image. Scale bars are 1 mm; F, H unscaled." figureDoi="http://doi.org/10.5281/zenodo.11063302" httpUri="https://zenodo.org/record/11063302/files/figure.png" pageId="8" pageNumber="9">Figure 4</figureCitation>
); p4 of 
<taxonomicName id="3EDC9BE1FFA30331CBE2FAAECD62FADE" baseAuthorityName="Hutchinson and Bown" baseAuthorityYear="1980" box="[1276,1403,1350,1375]" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="magna">
<emphasis id="CBA83C70FFA30331CBE2FAAECD62FADE" box="[1276,1403,1350,1375]" italics="true" pageId="8" pageNumber="9">C. magna</emphasis>
</taxonomicName>
is unknown.
</paragraph>
<paragraph id="F963E062FFA30331CC9CFA6ECDB8F89E" blockId="8.[840,1441,166,1887]" pageId="8" pageNumber="9">
<taxonomicName id="3EDC9BE1FFA30331CC9CFA6ECC86FA1E" authority="Zazhigin &amp; Voyta, 2022" authorityName="Zazhigin &amp; Voyta" authorityYear="2022" box="[898,1183,1414,1439]" class="Mammalia" family="Soricidae" genus="Ishimosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="ishimiensis" status="gen. et sp. nov.">
<emphasis id="CBA83C70FFA30331CC9CFA6ECC86FA1E" box="[898,1183,1414,1439]" italics="true" pageId="8" pageNumber="9">Ishimosorex ishimiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D09B810BFFA30331CBB2FA6ECD69FA1E" box="[1196,1392,1414,1439]" pageId="8" pageNumber="9" rank="species">gen. et sp. nov.</taxonomicNameLabel>
differs from 
<taxonomicName id="3EDC9BE1FFA30331CCA6FA4ECC49FA3E" authorityName="Storch and Zazhigin" authorityYear="1996" box="[952,1104,1446,1471]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="seletiensis">
<emphasis id="CBA83C70FFA30331CCA6FA4ECC49FA3E" box="[952,1104,1446,1471]" italics="true" pageId="8" pageNumber="9">P. seletiensis</emphasis>
</taxonomicName>
in a slightly larger size; from 
<taxonomicName id="3EDC9BE1FFA30331CC56FA2ECC80FA5E" authority="Zazhigin &amp; Voyta, 2022" authorityName="Zazhigin &amp; Voyta" authorityYear="2022" box="[840,1177,1478,1503]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="intermedius" status="sp. nov.">
<emphasis id="CBA83C70FFA30331CC56FA2ECC80FA5E" box="[840,1177,1478,1503]" italics="true" pageId="8" pageNumber="9">Paranourosorex intermedius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D09B810BFFA30331CBBDFA2ECD1BFA5E" box="[1187,1282,1478,1503]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="3EDC9BE1FFA30331CA5FFA2ECDB9FA5E" authorityName="Rzebik-Kowalska" authorityYear="1975" box="[1345,1440,1478,1503]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="gigas">
<emphasis id="CBA83C70FFA30331CA5FFA2ECDB9FA5E" box="[1345,1440,1478,1503]" italics="true" pageId="8" pageNumber="9">P. gigas</emphasis>
</taxonomicName>
in a distinctly smaller size (
<figureCitation id="61E7FCE7FFA30331CB91FA0ECCE1FA7E" box="[1167,1272,1510,1535]" captionStart="FIGURE 5" captionStartId="9.[194,285,954,977]" captionTargetBox="[247,1385,168,925]" captionTargetId="figure-340@9.[246,1389,168,929]" captionTargetPageId="9" captionText="FIGURE 5. Bivariate plot of linear differences between five Asian species, Paranourosorex seletiensis Storch and Zazhigin, 1996 (PSt.), Paranourosorex gigas Rzebik-Kowalska, 1975 (P. gigas), Paranourosorex intermedius sp. nov. (PIm.) and Ishimosorex ishimiensis gen. et sp. nov. (PPl) based on the second lower molar measurements (L vs TRL), mm). Studied remains from Asian Neogene localities are marked by colours, which corresponds to ELMA (see Figure 1); numbers are given in the Appendix 2." figureDoi="http://doi.org/10.5281/zenodo.11063304" httpUri="https://zenodo.org/record/11063304/files/figure.png" pageId="8" pageNumber="9">Figure 5</figureCitation>
). 
<taxonomicName id="3EDC9BE1FFA30331CA0EFA0ECBD3F99E" authority="Zazhigin &amp; Voyta, 2022" authorityName="Zazhigin &amp; Voyta" authorityYear="2022" class="Mammalia" family="Soricidae" genus="Ishimosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="ishimiensis" status="gen. et sp. nov.">
<emphasis id="CBA83C70FFA30331CA0EFA0ECBD3F99E" italics="true" pageId="8" pageNumber="9">Ishimosorex ishimiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D09B810BFFA30331CCC6F9EECC86F99E" box="[984,1183,1542,1567]" pageId="8" pageNumber="9" rank="species">gen. et sp. nov.</taxonomicNameLabel>
differs from 
<taxonomicName id="3EDC9BE1FFA30331CA5DF9EECBA3F9BE" baseAuthorityName="Schlosser" baseAuthorityYear="1924" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="inexspectatus">
<emphasis id="CBA83C70FFA30331CA5DF9EECBA3F9BE" italics="true" pageId="8" pageNumber="9">P. inexspectatus</emphasis>
</taxonomicName>
in a slightly smaller size (Appendix 4). In addition, 
<taxonomicName id="3EDC9BE1FFA30331CCA6F9AECCCCF9DE" authority="Zazhigin &amp; Voyta, 2022" authorityName="Zazhigin &amp; Voyta" authorityYear="2022" box="[952,1237,1606,1631]" class="Mammalia" family="Soricidae" genus="Ishimosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="ishimiensis" status="gen. et sp. nov.">
<emphasis id="CBA83C70FFA30331CCA6F9AECCCCF9DE" box="[952,1237,1606,1631]" italics="true" pageId="8" pageNumber="9">Ishimosorex ishimiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D09B810BFFA30331CBFEF9AECDB8F9DE" box="[1248,1441,1606,1631]" pageId="8" pageNumber="9" rank="species">gen. et sp. nov.</taxonomicNameLabel>
morphometrically differs from the known 
<taxonomicName id="3EDC9BE1FFA30331CA2CF98ECC1DF91E" pageId="8" pageNumber="9">
<emphasis id="CBA83C70FFA30331CA2CF98ECBB8F91E" italics="true" pageId="8" pageNumber="9">Paranourosorex</emphasis>
species
</taxonomicName>
in longer trigonid of m1 regarding the tooth length (TRL = 62.5% of m1 length) than show 
<taxonomicName id="3EDC9BE1FFA30331CC8BF92ECC2CF95E" authorityName="Storch and Zazhigin" authorityYear="1996" box="[917,1077,1734,1759]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="seletiensis">
<emphasis id="CBA83C70FFA30331CC8BF92ECC2CF95E" box="[917,1077,1734,1759]" italics="true" pageId="8" pageNumber="9">P. seletiensis</emphasis>
</taxonomicName>
(TRL = 57% of m1 length), 
<taxonomicName id="3EDC9BE1FFA30331CC56F90ECC83F97E" authority="Zazhigin &amp; Voyta, 2022" authorityName="Zazhigin &amp; Voyta" authorityYear="2022" box="[840,1178,1766,1791]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="intermedius" status="sp. nov.">
<emphasis id="CBA83C70FFA30331CC56F90ECC83F97E" box="[840,1178,1766,1791]" italics="true" pageId="8" pageNumber="9">Paranourosorex intermedius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D09B810BFFA30331CBBAF90ECD19F97E" box="[1188,1280,1766,1791]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
(TRL = 56% of m1 length) or 
<taxonomicName id="3EDC9BE1FFA30331CB19F8EECC7DF89E" authorityName="Rzebik-Kowalska" authorityYear="1975" box="[1031,1124,1798,1823]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="gigas">
<emphasis id="CBA83C70FFA30331CB19F8EECC7DF89E" box="[1031,1124,1798,1823]" italics="true" pageId="8" pageNumber="9">P. gigas</emphasis>
</taxonomicName>
(TRL = 54% of m1 length).
</paragraph>
<paragraph id="F963E062FFA30330CC9CF8CEC9BBFB52" blockId="8.[840,1441,166,1887]" lastBlockId="9.[192,793,1146,1907]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
The differences of 
<taxonomicName id="3EDC9BE1FFA30331CB62F8CECDB9F8BE" authority="Zazhigin &amp; Voyta, 2022" authorityName="Zazhigin &amp; Voyta" authorityYear="2022" box="[1148,1440,1830,1855]" class="Mammalia" family="Soricidae" genus="Ishimosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="ishimiensis" status="gen. et sp. nov.">
<emphasis id="CBA83C70FFA30331CB62F8CECDB9F8BE" box="[1148,1440,1830,1855]" italics="true" pageId="8" pageNumber="9">Ishimosorex ishimiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D09B810BFFA30331CC56F8AECC19F8DE" box="[840,1024,1862,1887]" pageId="8" pageNumber="9" rank="species">gen. et sp. nov.</taxonomicNameLabel>
from species of 
<taxonomicName id="3EDC9BE1FFA30331CBD8F8AECD47F8DE" authorityName="Kormos" authorityYear="1926" box="[1222,1374,1862,1887]" class="Mammalia" family="Soricidae" genus="Amblycoptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFA30331CBD8F8AECD47F8DE" box="[1222,1374,1862,1887]" italics="true" pageId="8" pageNumber="9">Amblycoptus</emphasis>
</taxonomicName>
, 
<taxonomicName id="3EDC9BE1FFA30330CA70F8AEC919FB12" authorityName="Meszaros" authorityYear="1997" class="Mammalia" family="Soricidae" genus="Kordosia" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="9" lastPageNumber="10" order="Soricomorpha" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFA30330CA70F8AEC919FB12" italics="true" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">Kordosia</emphasis>
</taxonomicName>
and 
<taxonomicName id="3EDC9BE1FFA20330CE23FB92C9CDFB12" authorityName="van Dam" authorityYear="2010" box="[317,468,1146,1171]" class="Mammalia" family="Soricidae" genus="Darocasorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFA20330CE23FB92C9CDFB12" box="[317,468,1146,1171]" italics="true" pageId="9" pageNumber="10">Darocasorex</emphasis>
</taxonomicName>
were described in the new genus comparisons (see above the Differential Diagnosis section).
</paragraph>
</subSubSection>
<caption id="ADA3B0EAFFA20330CFDCFC52CA6BFBC6" ID-DOI="http://doi.org/10.5281/zenodo.11063304" ID-Zenodo-Dep="11063304" httpUri="https://zenodo.org/record/11063304/files/figure.png" pageId="9" pageNumber="10" startId="9.[194,285,954,977]" targetBox="[247,1385,168,925]" targetPageId="9" targetType="figure">
<paragraph id="F963E062FFA20330CFDCFC52CA6BFBC6" blockId="9.[194,1437,954,1095]" pageId="9" pageNumber="10">
<emphasis id="CBA83C70FFA20330CFDCFC52C922FC53" bold="true" box="[194,315,954,978]" pageId="9" pageNumber="10">FIGURE 5.</emphasis>
Bivariate plot of linear differences between five Asian species, 
<taxonomicName id="3EDC9BE1FFA20330CCE6FC53C946FC6E" authority="Storch and Zazhigin, 1996" authorityName="Storch and Zazhigin" authorityYear="1996" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="seletiensis">
<emphasis id="CBA83C70FFA20330CCE6FC53CD05FC53" box="[1016,1308,955,978]" italics="true" pageId="9" pageNumber="10">Paranourosorex seletiensis</emphasis>
<bibRefCitation id="9D4D9D93FFA20330CA38FC52C946FC6E" author="Storch, G. &amp; Zazhigin, V. S." pageId="9" pageNumber="10" pagination="257 - 268" refId="ref17571" refString="Storch, G. and Zazhigin, V. S. 1996. Taxonomy and phylogeny of the Paranourosorex lineage, Neogene of Eurasia (Mammalia: Soricidae: Anourooricini). Palaontologische Zeitschrift, 70: 257 - 268. https: // doi. org / 10.1007 / BF 02988282" type="journal article" year="1996">Storch and Zazhigin, 1996</bibRefCitation>
</taxonomicName>
(PSt.), 
<taxonomicName id="3EDC9BE1FFA20330CEAEFC30CB85FC6E" authority="Rzebik-Kowalska, 1975" authorityName="Rzebik-Kowalska" authorityYear="1975" box="[432,924,984,1007]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="gigas">
<emphasis id="CBA83C70FFA20330CEAEFC30CA82FC6E" box="[432,667,984,1007]" italics="true" pageId="9" pageNumber="10">Paranourosorex gigas</emphasis>
Rzebik-Kowalska, 1975
</taxonomicName>
(
<taxonomicName id="3EDC9BE1FFA20330CCB5FC30CC1CFC6E" authorityName="Rzebik-Kowalska" authorityYear="1975" box="[939,1029,984,1007]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="gigas">P. gigas</taxonomicName>
), 
<taxonomicName id="3EDC9BE1FFA20330CB0BFC30CD5AFC6E" authority="Zazhigin &amp; Voyta, 2022" authorityName="Zazhigin &amp; Voyta" authorityYear="2022" box="[1045,1347,984,1007]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="intermedius" status="sp. nov.">
<emphasis id="CBA83C70FFA20330CB0BFC30CD5AFC6E" box="[1045,1347,984,1007]" italics="true" pageId="9" pageNumber="10">Paranourosorex intermedius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D09B810BFFA20330CA54FC30CD84FC6E" box="[1354,1437,984,1007]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
(PIm.) and 
<taxonomicName id="3EDC9BE1FFA20330CE26FC1DCA2DFB8D" authority="Zazhigin &amp; Voyta, 2022" authorityName="Zazhigin &amp; Voyta" authorityYear="2022" box="[312,564,1013,1036]" class="Mammalia" family="Soricidae" genus="Ishimosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="ishimiensis" status="gen. et sp. nov.">
<emphasis id="CBA83C70FFA20330CE26FC1DCA2DFB8D" box="[312,564,1013,1036]" italics="true" pageId="9" pageNumber="10">Ishimosorex ishimiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D09B810BFFA20330CD24FC1DCAC5FB8D" box="[570,732,1013,1036]" pageId="9" pageNumber="10" rank="species">gen. et sp. nov.</taxonomicNameLabel>
(PPl) based on the second lower molar measurements (L vs TRL), mm). Studied remains from Asian Neogene localities are marked by colours, which corresponds to ELMA (see Figure 1); numbers are given in the Appendix 2.
</paragraph>
</caption>
<subSubSection id="B1C6B3E9FFA20330CFDEFB32CBB9F892" pageId="9" pageNumber="10" type="discussion">
<paragraph id="F963E062FFA20330CFDEFB32CD1CF9B2" blockId="9.[192,793,1146,1907]" lastBlockId="9.[840,1440,1146,1811]" pageId="9" pageNumber="10">
<emphasis id="CBA83C70FFA20330CFDEFB32C921FB72" bold="true" box="[192,312,1242,1267]" pageId="9" pageNumber="10">Remarks.</emphasis>
<taxonomicName id="3EDC9BE1FFA20330CE5EFB32CA43FB72" authority="Zazhigin &amp; Voyta, 2022" authorityName="Zazhigin &amp; Voyta" authorityYear="2022" box="[320,602,1242,1267]" class="Mammalia" family="Soricidae" genus="Ishimosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="ishimiensis" status="gen. et sp. nov.">
<emphasis id="CBA83C70FFA20330CE5EFB32CA43FB72" box="[320,602,1242,1267]" italics="true" pageId="9" pageNumber="10">Ishimosorex ishimiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D09B810BFFA20330CD7CFB32CB00FB72" box="[610,793,1242,1267]" pageId="9" pageNumber="10" rank="species">gen. et sp. nov.</taxonomicNameLabel>
shows an intermediate combination of 'omnivorous' features of 
<taxonomicName id="3EDC9BE1FFA20330CE55FAF2CA10FAB2" authorityName="Rzebik-Kowalska" authorityYear="1975" box="[331,521,1306,1331]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFA20330CE55FAF2CA10FAB2" box="[331,521,1306,1331]" italics="true" pageId="9" pageNumber="10">Paranourosorex</emphasis>
</taxonomicName>
and ‘carnivorous’ features of 
<taxonomicName id="3EDC9BE1FFA20330CE33FAD2C9DDFAD2" authorityName="Gibert" authorityYear="1974" box="[301,452,1338,1363]" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFA20330CE33FAD2C9DDFAD2" box="[301,452,1338,1363]" italics="true" pageId="9" pageNumber="10">Crusafontina</emphasis>
</taxonomicName>
in the dentition. M1 bears expressed W-shaped buccal crests and a buccally protruding well-developed mesostyle similar to 
<taxonomicName id="3EDC9BE1FFA20330CFDEFA72C967FA32" authorityName="Rzebik-Kowalska" authorityYear="1975" box="[192,382,1434,1459]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFA20330CFDEFA72C967FA32" box="[192,382,1434,1459]" italics="true" pageId="9" pageNumber="10">Paranourosorex</emphasis>
</taxonomicName>
conditions. In addition, both taxa have a moderately developed parastyle of M1, the bulbous and anteriorly inflated p4 without sharp crests and wide lower molars (buccal shifting of the protoconid and hypoconid). These features are intended for tearing and crushing, i.e., they can be determined as ‘omnivore-like features.’ In contrast, 
<taxonomicName id="3EDC9BE1FFA20330CFDEF992C94EF912" authorityName="Gibert" authorityYear="1974" box="[192,343,1658,1683]" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFA20330CFDEF992C94EF912" box="[192,343,1658,1683]" italics="true" pageId="9" pageNumber="10">Crusafontina</emphasis>
</taxonomicName>
(American and European species, except 
<taxonomicName id="3EDC9BE1FFA20330CE00F972C9C6F932" authorityName="van Dam" authorityYear="2004" box="[286,479,1690,1715]" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="vandeweerdi">
<emphasis id="CBA83C70FFA20330CE00F972C9C6F932" box="[286,479,1690,1715]" italics="true" pageId="9" pageNumber="10">C. vandeweerdi</emphasis>
</taxonomicName>
), 
<taxonomicName id="3EDC9BE1FFA20330CEE1F972CA81F932" authorityName="Kormos" authorityYear="1926" box="[511,664,1690,1715]" class="Mammalia" family="Soricidae" genus="Amblycoptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFA20330CEE1F972CA81F932" box="[511,664,1690,1715]" italics="true" pageId="9" pageNumber="10">Amblycoptus</emphasis>
</taxonomicName>
, 
<taxonomicName id="3EDC9BE1FFA20330CDB1F972CB01F932" authorityName="Meszaros" authorityYear="1997" box="[687,792,1690,1715]" class="Mammalia" family="Soricidae" genus="Kordosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFA20330CDB1F972CB01F932" box="[687,792,1690,1715]" italics="true" pageId="9" pageNumber="10">Kordosia</emphasis>
</taxonomicName>
and 
<taxonomicName id="3EDC9BE1FFA20330CE18F952C984F952" authorityName="Zazhigin &amp; Voyta" authorityYear="2022" box="[262,413,1722,1747]" class="Mammalia" family="Soricidae" genus="Anourosorex" kingdom="Animalia" order="Soricomorpha" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFA20330CE18F952C984F952" box="[262,413,1722,1747]" italics="true" pageId="9" pageNumber="10">Anourosorex</emphasis>
</taxonomicName>
show different degrees of straightening of the buccal crests and increasing (from 
<taxonomicName id="3EDC9BE1FFA20330CE14F912C9B8F892" authorityName="Gibert" authorityYear="1974" box="[266,417,1786,1811]" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFA20330CE14F912C9B8F892" box="[266,417,1786,1811]" italics="true" pageId="9" pageNumber="10">Crusafontina</emphasis>
</taxonomicName>
to 
<taxonomicName id="3EDC9BE1FFA20330CECEF912CA20F892" authorityName="Meszaros" authorityYear="1997" box="[464,569,1786,1811]" class="Mammalia" family="Soricidae" genus="Kordosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFA20330CECEF912CA20F892" box="[464,569,1786,1811]" italics="true" pageId="9" pageNumber="10">Kordosia</emphasis>
</taxonomicName>
) parastyle size of M1. The change in the parastyle together with the trapeziform M1 outline of these taxa probably are related to carnivorous adaptations such as the lengthening of cutting edges; e.g., to the sharp and well-developed central crest of p4 of American and European 
<taxonomicName id="3EDC9BE1FFA20330CCDBFB52CC45FB52" authorityName="Gibert" authorityYear="1974" box="[965,1116,1210,1235]" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFA20330CCDBFB52CC45FB52" box="[965,1116,1210,1235]" italics="true" pageId="9" pageNumber="10">Crusafontina</emphasis>
</taxonomicName>
; the central position of the protoconid of m1 and the corresponding lengthening of the preprotocristid of 
<taxonomicName id="3EDC9BE1FFA20330CBBDFB12CD23FA92" authorityName="Gibert" authorityYear="1974" box="[1187,1338,1274,1299]" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFA20330CBBDFB12CD23FA92" box="[1187,1338,1274,1299]" italics="true" pageId="9" pageNumber="10">Crusafontina</emphasis>
</taxonomicName>
, 
<taxonomicName id="3EDC9BE1FFA20330CA53FB12CB8FFAB2" authorityName="Kormos" authorityYear="1926" class="Mammalia" family="Soricidae" genus="Amblycoptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFA20330CA53FB12CB8FFAB2" italics="true" pageId="9" pageNumber="10">Amblycoptus</emphasis>
</taxonomicName>
, 
<taxonomicName id="3EDC9BE1FFA20330CCAFFAF2CC03FAB2" authorityName="Meszaros" authorityYear="1997" box="[945,1050,1306,1331]" class="Mammalia" family="Soricidae" genus="Kordosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFA20330CCAFFAF2CC03FAB2" box="[945,1050,1306,1331]" italics="true" pageId="9" pageNumber="10">Kordosia</emphasis>
</taxonomicName>
and 
<taxonomicName id="3EDC9BE1FFA20330CB73FAF2CD1DFAB2" authorityName="Zazhigin &amp; Voyta" authorityYear="2022" box="[1133,1284,1306,1331]" class="Mammalia" family="Soricidae" genus="Anourosorex" kingdom="Animalia" order="Soricomorpha" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFA20330CB73FAF2CD1DFAB2" box="[1133,1284,1306,1331]" italics="true" pageId="9" pageNumber="10">Anourosorex</emphasis>
</taxonomicName>
to different degrees. Thus, some characteristics of 
<taxonomicName id="3EDC9BE1FFA20330CA25FAD2CC10FAF2" authority="Zazhigin &amp; Voyta, 2022" authorityName="Zazhigin &amp; Voyta" authorityYear="2022" class="Mammalia" family="Soricidae" genus="Ishimosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="ishimiensis" status="sp. nov.">
<emphasis id="CBA83C70FFA20330CA25FAD2CC10FAF2" italics="true" pageId="9" pageNumber="10">Ishimosorex ishimiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D09B810BFFA20330CB0BFAB2CC6CFAF2" box="[1045,1141,1370,1395]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
together with 
<taxonomicName id="3EDC9BE1FFA20330CA36FAB2CB8EFA12" authorityName="van Dam" authorityYear="2004" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="vandeweerdi">
<emphasis id="CBA83C70FFA20330CA36FAB2CB8EFA12" italics="true" pageId="9" pageNumber="10">C. vandeweerdi</emphasis>
</taxonomicName>
can be determined as ‘carnivore-like’ compared with pronounced carnivorous anourosoricins. The deep notch in the talon of I1 and two sharp, slender denticles of the cutting edge of i1 are similar to the American and European species of the more carnivorous 
<taxonomicName id="3EDC9BE1FFA20330CB79F9F2CCE7F9B2" authorityName="Gibert" authorityYear="1974" box="[1127,1278,1562,1587]" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFA20330CB79F9F2CCE7F9B2" box="[1127,1278,1562,1587]" italics="true" pageId="9" pageNumber="10">Crusafontina</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="F963E062FFA20330CC56F9D2CBB9F892" blockId="9.[840,1440,1146,1811]" pageId="9" pageNumber="10">
<emphasis id="CBA83C70FFA20330CC56F9D2CD20F9D2" bold="true" box="[840,1337,1594,1620]" pageId="9" pageNumber="10">Stratigraphic and geographic range.</emphasis>
Known from the 
<typeStatus id="26675EC0FFA20330CCAFF9B2CBFAF9F2" box="[945,995,1626,1651]" pageId="9" pageNumber="10">type</typeStatus>
locality of the species (
<collectingCounty id="100298EEFFA20330CBE8F9B2CDB9F9F2" box="[1270,1440,1626,1651]" pageId="9" pageNumber="10">Petropavlovsk</collectingCounty>
1A, MN 10, 
<collectingMunicipality id="19077A18FFA20330CCF4F992CCADF912" box="[1002,1204,1658,1683]" pageId="9" pageNumber="10">Ishim Formation</collectingMunicipality>
; 
<collectingRegion id="3B182E80FFA20330CBD8F992CB8EF932" country="Kazakhstan" name="Soltustik Qazaqstan Oblysy" pageId="9" pageNumber="10">North Kazakhstan Region</collectingRegion>
) and 
<location id="FC03B6B9FFA20330CCEFF972CC3CF932" LSID="urn:lsid:plazi:treatment:71755174FFAE0330CE30F918CBB9F892:FC03B6B9FFA20330CCEFF972CC3CF932" box="[1009,1061,1690,1715]" county="Petropavlovsk" municipality="Ishim Formation" name="Borki" pageId="9" pageNumber="10" stateProvince="Soltustik Qazaqstan Oblysy">Borki</location>
1A locality (MN 13, 
<location id="FC03B6B9FFA20330CA58F972CC37F952" LSID="urn:lsid:plazi:treatment:71755174FFAE0330CE30F918CBB9F892:FC03B6B9FFA20330CA58F972CC37F952" county="Petropavlovsk" municipality="Ishim Formation" name="Novaya Stanitsa Formation" pageId="9" pageNumber="10" stateProvince="Soltustik Qazaqstan Oblysy">Novaya Stanitsa Formation</location>
; 
<collectingRegion id="3B182E80FFA20330CB23F952CD61F952" box="[1085,1400,1722,1747]" country="Kazakhstan" name="Soltustik Qazaqstan Oblysy" pageId="9" pageNumber="10">North Kazakhstan Region</collectingRegion>
) to which the material was redeposited from 
<location id="FC03B6B9FFA20330CA37F932CB85F892" LSID="urn:lsid:plazi:treatment:71755174FFAE0330CE30F918CBB9F892:FC03B6B9FFA20330CA37F932CB85F892" county="Petropavlovsk" municipality="Ishim Formation" name="Ishim Formation" pageId="9" pageNumber="10" stateProvince="Soltustik Qazaqstan Oblysy">Ishim Formation</location>
.
</paragraph>
</subSubSection>
</treatment>
</document>