317 lines
27 KiB
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317 lines
27 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.898.46465" ID-GBIF-Dataset="4c647ed2-eacc-4651-8be2-4e96182a995a" ID-PMC="PMC6926426" ID-Pensoft-Pub="1313-2970-898-83" ID-Pensoft-UUID="A8C1E3B7F0A6517B9C01D45772231286" ID-PubMed="31875088" ID-ZooBank="E9639F08370640CF87BCA3E8D4933AB4" ModsDocID="1313-2970-898-83" checkinTime="1575997094558" checkinUser="pensoft" docAuthor="Kaminski, Marcin J., Kanda, Kojun & Smith, Aaron D." docDate="2019" docId="71528F9C04E25B6698BAB8ACFE354A77" docLanguage="en" docName="ZooKeys 898: 83-102" docOrigin="ZooKeys 898" docSource="http://dx.doi.org/10.3897/zookeys.898.46465" docTitle="Machleida devia Peringuey 1899" docType="treatment" docVersion="4" id="A8C1E3B7F0A6517B9C01D45772231286" lastPageNumber="83" masterDocId="A8C1E3B7F0A6517B9C01D45772231286" masterDocTitle="Taxonomic revision of the genus Machleida Fahraeus, 1870 (Tenebrionidae, Pimeliinae, Asidini)" masterLastPageNumber="102" masterPageNumber="83" pageNumber="83" updateTime="1668168227144" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of the genus Machleida Fahraeus, 1870 (Tenebrionidae, Pimeliinae, Asidini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kaminski, Marcin J.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kanda, Kojun</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Smith, Aaron D.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>898</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>83</mods:start>
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<mods:end>102</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.898.46465</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.898.46465</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-898-83</mods:identifier>
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<mods:identifier type="ZooBank">E9639F08370640CF87BCA3E8D4933AB4</mods:identifier>
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<mods:identifier type="Pensoft-UUID">A8C1E3B7F0A6517B9C01D45772231286</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="160689339" LSID="urn:lsid:plazi:treatment:71528F9C04E25B6698BAB8ACFE354A77" httpUri="http://treatment.plazi.org/id/71528F9C04E25B6698BAB8ACFE354A77" lastPageNumber="83" pageId="0" pageNumber="83">
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<subSubSection pageId="0" pageNumber="83" type="nomenclature">
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<paragraph pageId="0" pageNumber="83">
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<taxonomicName LSID="71528F9C-04E2-5B66-98BA-B8ACFE354A77" authority="Peringuey, 1899" authorityName="Peringuey" authorityYear="1899" class="Insecta" family="Tenebrionidae" genus="Machleida" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Machleida devia" order="Coleoptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="devia">
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Machleida devia (
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<normalizedToken originalValue="Péringuey">Peringuey</normalizedToken>
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, 1899)
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</taxonomicName>
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Selected diagnostic characters proposed for Machleida and its species. A, B Pronotal disc with specific surface modifications C ventral aspect of postoral regions D antenna E, F dorsal aspect of head. Illustrated species: A, D Machleida zofiae B M. nodulosa C M. tarskii E M. devia F M. flagstaffensis." figureDoi="10.3897/zookeys.898.46465.figure1" httpUri="https://binary.pensoft.net/fig/363751" pageId="0" pageNumber="83" tableDoi="10.3897/zookeys.898.46465.figure1">Figs 1E</figureCitation>
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,
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus images of the studied species. A Machleida banachi B M. devia C M. flagstaffensis D M. nodulosa E M. tarskii F M. zofiae." figureDoi="10.3897/zookeys.898.46465.figure2" httpUri="https://binary.pensoft.net/fig/363752" pageId="0" pageNumber="83" tableDoi="10.3897/zookeys.898.46465.figure2">2B</figureCitation>
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,
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphological details of selected Machleida species. A, B Lateral aspect of elytron C scutellum and D, E elytral disc. Illustrated species: A Machleida tarskii B M. banachi C M. zofiae D M. devia E M. nodulosa." figureDoi="10.3897/zookeys.898.46465.figure3" httpUri="https://binary.pensoft.net/fig/363753" pageId="0" pageNumber="83" tableDoi="10.3897/zookeys.898.46465.figure3">3D</figureCitation>
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,
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Female and male terminalia of Machleida. A, B Ovipositor and genital tubes of Machleida flagstaffensis C aedeagus of M. devia." figureDoi="10.3897/zookeys.898.46465.figure4" httpUri="https://binary.pensoft.net/fig/363754" pageId="0" pageNumber="83" tableDoi="10.3897/zookeys.898.46465.figure4">4C</figureCitation>
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,
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Distribution of species of the genus Machleida." figureDoi="10.3897/zookeys.898.46465.figure5" httpUri="https://binary.pensoft.net/fig/363755" pageId="0" pageNumber="83" tableDoi="10.3897/zookeys.898.46465.figure5">5</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="83" type="reference_group">
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<paragraph pageId="0" pageNumber="83">
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<taxonomicName class="Insecta" family="Tenebrionidae" genus="Asida" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Asida devia" order="Coleoptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="devia">Asida devia</taxonomicName>
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<normalizedToken originalValue="Péringuey">Peringuey</normalizedToken>
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, 1899: 258 [transferred to
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<taxonomicName class="Insecta" family="Tenebrionidae" genus="Machleida" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Machleida" order="Coleoptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">Machleida</taxonomicName>
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by
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<bibRefCitation DOI="https://doi.org/10.1002/mmnd.192419240602" author="Wilke, S" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="0" pageNumber="83" pagination="517 - 549" refId="B27" refString="Wilke, S, 1925. Dersudafrikanische Formenkreis der Asidinen (Col. Tenebr.). Deutsche Entomologische Zeitschrift 1924: 517 - 549" title="Dersudafrikanische Formenkreis der Asidinen (Col. Tenebr.)." url="https://doi.org/10.1002/mmnd.192419240602" volume="1924" year="1925">Wilke (1925</bibRefCitation>
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: 536)]
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</paragraph>
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<paragraph pageId="0" pageNumber="83">
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=
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<taxonomicName class="Insecta" family="Tenebrionidae" genus="Machleida" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Machleida capillosa" order="Coleoptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="capillosa">Machleida capillosa</taxonomicName>
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Wilke, 1925: 536 [syn. nov.]
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="83" type="materials_examined">
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<paragraph pageId="0" pageNumber="83">Material studied.</paragraph>
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<paragraph pageId="0" pageNumber="83">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="83">Syntype</emphasis>
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of
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<taxonomicName class="Insecta" family="Tenebrionidae" genus="Machleida" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Machleida capillosa" order="Coleoptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="capillosa">
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<emphasis italics="true" pageId="0" pageNumber="83">Machleida capillosa</emphasis>
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</taxonomicName>
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(Berlin Mus.): "Natal Mus., / Maritzburg. / 1913-330", "
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<taxonomicName lsidName="capillosa" pageId="0" pageNumber="83" rank="species" species="capillosa">capillosa</taxonomicName>
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/ sp. n.". Two specimens (Transvaal Mus.): Tugela River / nr Kranskop / Lawrence & / Haacke; single specimen (Transvaal Mus.): "S. Afr.: Zululand / Hluhluwe Game Res. /
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<geoCoordinate degrees="28.05" direction="south" orientation="latitude" precision="555" value="-28.05">
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28.05
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<normalizedToken originalValue="S–">S-</normalizedToken>
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</geoCoordinate>
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<geoCoordinate degrees="32.04" direction="east" orientation="longitude" precision="555" value="32.04">32.04E</geoCoordinate>
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", "20.111992: E-Y: 2840 / fung. Trunk & litter / leg. Endrody - Younga".
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="83" type="notes">
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<paragraph pageId="0" pageNumber="83">Notes.</paragraph>
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<paragraph pageId="0" pageNumber="83">
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During the preparation phase for this study the holotype of
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<taxonomicName class="Insecta" family="Tenebrionidae" genus="Asida" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Asida devia" order="Coleoptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="devia">
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<emphasis italics="true" pageId="0" pageNumber="83">Asida devia</emphasis>
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</taxonomicName>
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was not found in any of the queried collections, i.e. Iziko Museum of South Africa in Cape Town and the institutions listed in the Material and methods section. However, the original description indicates several unique morphological features of this species (i.e. large body size, presence of densely distributed microtubercles on the elytral disc), which were used to differentiate it from other congeners.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="83" type="redescription">
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<paragraph pageId="0" pageNumber="83">Redescription.</paragraph>
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<paragraph pageId="0" pageNumber="83">
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Length 13.0-14.0 mm, width of elytra 8.5-9.5 mm. Integument black, often densely coated with debris.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="83">Head</emphasis>
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: frons with longitudinal median depression, densely punctate (~0.2 diameters apart), each puncture with short yellowish rectangular, flattened scale-like seta; frontoclypeal suture medially indistinguishable, weakly indented at margins, with pair of lateral depressions; apical clypeal margin broadly and shallowly emarginate; clypeus projected toward front of body (
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Selected diagnostic characters proposed for Machleida and its species. A, B Pronotal disc with specific surface modifications C ventral aspect of postoral regions D antenna E, F dorsal aspect of head. Illustrated species: A, D Machleida zofiae B M. nodulosa C M. tarskii E M. devia F M. flagstaffensis." figureDoi="10.3897/zookeys.898.46465.figure1" httpUri="https://binary.pensoft.net/fig/363751" pageId="0" pageNumber="83" tableDoi="10.3897/zookeys.898.46465.figure1">Fig. 1E</figureCitation>
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); apical margin of labrum strongly and sharply emarginate, densely punctate (~0.2 diameters apart), each puncture with short yellowish acuminate seta. Eye elongate oval, length approximately 8
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<normalizedToken originalValue="×">x</normalizedToken>
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width, weakly emarginate around epistomal base. Mentum with rounded base, not fully filling buccal cavity; anterior margin weakly medially emarginate; densely punctate, each puncture with single rectangular flattened scale-like seta. Submentum semicircular, concave medially, densely punctate. Antenna moderately clothed in erect acuminate yellowish setae; length of antennomeres 10+11 equal to 0.8 of antennomere 3 length; length of antenna equal to 0.75 of pronotal length.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="83">Prothorax</emphasis>
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: pronotal lateral margin sinuate, slightly raised. Pronotum widest below middle. Disc with two median carinae merging in middle; lateral tubercles confluent with median carinae, forming convexities situated above half pronotal length; surface densely punctate (~0.2 diameters apart), each puncture with short yellowish rectangular, flattened scale-like seta; anterior margin strongly emarginate, anterior apices strongly produced; base bisinuate. Hypomeron with shallow antennal sulcus, densely punctate (~0.2 diameters apart), each puncture with short yellowish rectangular, flattened scale-like seta. Prosternal process strongly convex, without median sulcus (ventral view).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="83">Pterothorax</emphasis>
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: scutellum without median groove. Elytra widest behind middle, clothed with extremely short yellowish rectangular, flattened scale-like setae and microtubercles (
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphological details of selected Machleida species. A, B Lateral aspect of elytron C scutellum and D, E elytral disc. Illustrated species: A Machleida tarskii B M. banachi C M. zofiae D M. devia E M. nodulosa." figureDoi="10.3897/zookeys.898.46465.figure3" httpUri="https://binary.pensoft.net/fig/363753" pageId="0" pageNumber="83" tableDoi="10.3897/zookeys.898.46465.figure3">Fig. 3D</figureCitation>
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); marginal costae present, tuberculate, apex of each tubercle densely covered with acuminate setae, divided near humera, with marginal branch extending to approximately middle of 4th abdominal ventrite, dorsal branch extending to base of 3rd abdominal ventrite, terminal tubercles transverse; disc rugose, without any trace of intervals and rows, covered with microtubercles; ventral portion of elytra basally impunctate. Elytral slope steep (falling at angle of 75°). Epipleura clearly distinguishable. Mesanepisternum, mesepimeron, and metepimeron impunctate or sparsely punctate. Meso- and metaventrite densely punctate and covered with acuminate setae. Lateral regions of metaventrite (between coxae) extremely short.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="83">Legs</emphasis>
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: apex of profemora with small denticle on outer margin. Femora and tibia densely punctate and setose. Tarsi cylindrical, not flattened.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="83">Abdomen</emphasis>
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: ventrites 1-3 moderately punctate and weakly rugulose; ventrites 4 and 5 densely punctate and setose; ventrite 5 without submarginal sulcus.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="83">Terminalia</emphasis>
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: aedeagus bipartite, with apical part slightly shorter than basal portion (
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Female and male terminalia of Machleida. A, B Ovipositor and genital tubes of Machleida flagstaffensis C aedeagus of M. devia." figureDoi="10.3897/zookeys.898.46465.figure4" httpUri="https://binary.pensoft.net/fig/363754" pageId="0" pageNumber="83" tableDoi="10.3897/zookeys.898.46465.figure4">Fig. 4C</figureCitation>
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). Female specimens were not dissected due to scarcity of available materials.
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</paragraph>
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<caption doi="10.3897/zookeys.898.46465.figure1" httpUri="https://binary.pensoft.net/fig/363751" pageId="0" pageNumber="83" start="Figure 1" startId="F1">
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<paragraph pageId="0" pageNumber="83">
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<emphasis bold="true" pageId="0" pageNumber="83">Figure 1.</emphasis>
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Selected diagnostic characters proposed for
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<taxonomicName class="Insecta" family="Tenebrionidae" genus="Machleida" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Machleida" order="Coleoptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="83">Machleida</emphasis>
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</taxonomicName>
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and its species.
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<emphasis bold="true" pageId="0" pageNumber="83">A, B</emphasis>
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Pronotal disc with specific surface modifications
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<emphasis bold="true" pageId="0" pageNumber="83">C</emphasis>
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ventral aspect of postoral regions
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<emphasis bold="true" pageId="0" pageNumber="83">D</emphasis>
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antenna
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<emphasis bold="true" pageId="0" pageNumber="83">E, F</emphasis>
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dorsal aspect of head. Illustrated species:
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<emphasis bold="true" pageId="0" pageNumber="83">A, D</emphasis>
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<taxonomicName class="Insecta" family="Tenebrionidae" genus="Machleida" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Machleida zofiae" order="Coleoptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="zofiae">
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<emphasis italics="true" pageId="0" pageNumber="83">Machleida zofiae</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="83">B</emphasis>
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<taxonomicName lsidName="M. nodulosa" pageId="0" pageNumber="83" rank="species" species="nodulosa">
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<emphasis italics="true" pageId="0" pageNumber="83">M. nodulosa</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="83">C</emphasis>
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<taxonomicName lsidName="M. tarskii" pageId="0" pageNumber="83" rank="species" species="tarskii">
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<emphasis italics="true" pageId="0" pageNumber="83">M. tarskii</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="83">E</emphasis>
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<taxonomicName lsidName="M. devia" pageId="0" pageNumber="83" rank="species" species="devia">
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<emphasis italics="true" pageId="0" pageNumber="83">M. devia</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="83">F</emphasis>
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<taxonomicName lsidName="M. flagstaffensis" pageId="0" pageNumber="83" rank="species" species="flagstaffensis">
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<emphasis italics="true" pageId="0" pageNumber="83">M. flagstaffensis</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</caption>
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<caption doi="10.3897/zookeys.898.46465.figure2" httpUri="https://binary.pensoft.net/fig/363752" pageId="0" pageNumber="83" start="Figure 2" startId="F2">
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<paragraph pageId="0" pageNumber="83">
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<emphasis bold="true" pageId="0" pageNumber="83">Figure 2.</emphasis>
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Habitus images of the studied species.
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<emphasis bold="true" pageId="0" pageNumber="83">A</emphasis>
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<taxonomicName class="Insecta" family="Tenebrionidae" genus="Machleida" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Machleida banachi" order="Coleoptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="banachi">
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<emphasis italics="true" pageId="0" pageNumber="83">Machleida banachi</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="83">B</emphasis>
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<taxonomicName lsidName="M. devia" pageId="0" pageNumber="83" rank="species" species="devia">
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<emphasis italics="true" pageId="0" pageNumber="83">M. devia</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="83">C</emphasis>
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<taxonomicName lsidName="M. flagstaffensis" pageId="0" pageNumber="83" rank="species" species="flagstaffensis">
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<emphasis italics="true" pageId="0" pageNumber="83">M. flagstaffensis</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="83">D</emphasis>
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<taxonomicName lsidName="M. nodulosa" pageId="0" pageNumber="83" rank="species" species="nodulosa">
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<emphasis italics="true" pageId="0" pageNumber="83">M. nodulosa</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="83">E</emphasis>
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<taxonomicName lsidName="M. tarskii" pageId="0" pageNumber="83" rank="species" species="tarskii">
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<emphasis italics="true" pageId="0" pageNumber="83">M. tarskii</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="83">F</emphasis>
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<taxonomicName lsidName="M. zofiae" pageId="0" pageNumber="83" rank="species" species="zofiae">
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<emphasis italics="true" pageId="0" pageNumber="83">M. zofiae</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</caption>
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<caption doi="10.3897/zookeys.898.46465.figure3" httpUri="https://binary.pensoft.net/fig/363753" pageId="0" pageNumber="83" start="Figure 3" startId="F3">
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<paragraph pageId="0" pageNumber="83">
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<emphasis bold="true" pageId="0" pageNumber="83">Figure 3.</emphasis>
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Morphological details of selected
|
||
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Machleida" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Machleida" order="Coleoptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="83">Machleida</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
<emphasis bold="true" pageId="0" pageNumber="83">A, B</emphasis>
|
||
Lateral aspect of elytron
|
||
<emphasis bold="true" pageId="0" pageNumber="83">C</emphasis>
|
||
scutellum and
|
||
<emphasis bold="true" pageId="0" pageNumber="83">D, E</emphasis>
|
||
elytral disc. Illustrated species:
|
||
<emphasis bold="true" pageId="0" pageNumber="83">A</emphasis>
|
||
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Machleida" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Machleida tarskii" order="Coleoptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="tarskii">
|
||
<emphasis italics="true" pageId="0" pageNumber="83">Machleida tarskii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="83">B</emphasis>
|
||
<taxonomicName lsidName="M. banachi" pageId="0" pageNumber="83" rank="species" species="banachi">
|
||
<emphasis italics="true" pageId="0" pageNumber="83">M. banachi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="83">C</emphasis>
|
||
<taxonomicName lsidName="M. zofiae" pageId="0" pageNumber="83" rank="species" species="zofiae">
|
||
<emphasis italics="true" pageId="0" pageNumber="83">M. zofiae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="83">D</emphasis>
|
||
<taxonomicName lsidName="M. devia" pageId="0" pageNumber="83" rank="species" species="devia">
|
||
<emphasis italics="true" pageId="0" pageNumber="83">M. devia</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="83">E</emphasis>
|
||
<taxonomicName lsidName="M. nodulosa" pageId="0" pageNumber="83" rank="species" species="nodulosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="83">M. nodulosa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.898.46465.figure4" httpUri="https://binary.pensoft.net/fig/363754" pageId="0" pageNumber="83" start="Figure 4" startId="F4">
|
||
<paragraph pageId="0" pageNumber="83">
|
||
<emphasis bold="true" pageId="0" pageNumber="83">Figure 4.</emphasis>
|
||
Female and male terminalia of
|
||
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Machleida" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Machleida" order="Coleoptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="83">Machleida</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis bold="true" pageId="0" pageNumber="83">A, B</emphasis>
|
||
Ovipositor and genital tubes of
|
||
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Machleida" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Machleida flagstaffensis" order="Coleoptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="flagstaffensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="83">Machleida flagstaffensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="83">C</emphasis>
|
||
aedeagus of
|
||
<taxonomicName lsidName="M. devia" pageId="0" pageNumber="83" rank="species" species="devia">
|
||
<emphasis italics="true" pageId="0" pageNumber="83">M. devia</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="83" type="notes on synonymy">
|
||
<paragraph pageId="0" pageNumber="83">Notes on synonymy.</paragraph>
|
||
<paragraph pageId="0" pageNumber="83">
|
||
Type investigation of
|
||
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Machleida" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Machleida capillosa" order="Coleoptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="capillosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="83">Machleida capillosa</emphasis>
|
||
</taxonomicName>
|
||
did not reveal any characters to support its taxonomic distinctiveness from
|
||
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Asida" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Asida devia" order="Coleoptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="devia">
|
||
<emphasis italics="true" pageId="0" pageNumber="83">Asida devia</emphasis>
|
||
</taxonomicName>
|
||
(e.g. both share a unique structure of elytral disc - surface densely covered with microtubercles) (
|
||
<bibRefCitation author="Peringuey, LA" journalOrPublisher="Annals of the South Africa Museum" pageId="0" pageNumber="83" pagination="240 - 330" refId="B15" refString="Peringuey, LA, 1899. Fifth contribution to the South-African coleopterous fauna. Annals of the South Africa Museum 1: 240 - 330" title="Fifth contribution to the South-African coleopterous fauna." volume="1" year="1899">
|
||
<normalizedToken originalValue="Péringuey">Peringuey</normalizedToken>
|
||
1899
|
||
</bibRefCitation>
|
||
). Furthermore, both species have the same
|
||
<emphasis italics="true" pageId="0" pageNumber="83">locus typicus</emphasis>
|
||
(
|
||
<normalizedToken originalValue="“Maritzburg”">"Maritzburg"</normalizedToken>
|
||
, South Africa). As a result, M.
|
||
<taxonomicName lsidName="capillosa" pageId="0" pageNumber="83" rank="species" species="capillosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="83">capillosa</emphasis>
|
||
</taxonomicName>
|
||
is considered here as a junior subjective synonym of
|
||
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Asida" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Asida devia" order="Coleoptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="devia">
|
||
<emphasis italics="true" pageId="0" pageNumber="83">Asida devia</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="83" type="distribution">
|
||
<paragraph pageId="0" pageNumber="83">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="83">
|
||
Representatives of this species have been collected in the following ecoregions of South Africa (
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Distribution of species of the genus Machleida." figureDoi="10.3897/zookeys.898.46465.figure5" httpUri="https://binary.pensoft.net/fig/363755" pageId="0" pageNumber="83" tableDoi="10.3897/zookeys.898.46465.figure5">Fig. 5</figureCitation>
|
||
): Drakensberg Montane Woodlands and Grasslands, Maputaland-Pondoland bushland and thickets.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.898.46465.figure5" httpUri="https://binary.pensoft.net/fig/363755" pageId="0" pageNumber="83" start="Figure 5" startId="F5">
|
||
<paragraph pageId="0" pageNumber="83">
|
||
<emphasis bold="true" pageId="0" pageNumber="83">Figure 5.</emphasis>
|
||
Distribution of species of the genus
|
||
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Machleida" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Machleida" order="Coleoptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="83">Machleida</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |