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<document ID-CLB-Dataset="81246" ID-DOI="http://dx.doi.org/10.3897/aiep.52.84796" ID-GBIF-Dataset="ac6076b5-ee3c-47ab-8434-824ffe683a20" ID-Pensoft-Pub="1734-1515-2-135" ID-Pensoft-UUID="304432CC04565BA19606EE4ED73F459E" ID-ZooBank="D009F127C4FE4E7DBDCFD2BB549F6B79" ModsDocID="1734-1515-52-2-135" checkinTime="1656145909991" checkinUser="pensoft" docAuthor="Murugan, Arumugam, Mahadevan, Gopalan &amp; Fricke, Ronald" docDate="2022" docId="715213C776D5544C9956E514238204BA" docLanguage="en" docName="ActIchthPisc 52(2): 135-139" docOrigin="Acta Ichthyologica et Piscatoria 52 (2)" docPubDate="2022-06-24" docSource="http://dx.doi.org/10.3897/aiep.52.84796" docTitle="Oxycheilinus arenatus" docType="treatment" docVersion="6" id="304432CC04565BA19606EE4ED73F459E" lastPageNumber="135" masterDocId="304432CC04565BA19606EE4ED73F459E" masterDocTitle="First record of the speckled Maori wrasse, Oxycheilinus arenatus (Valenciennes, 1840) (Actinopterygii: Perciformes: Labridae), from Indian coastal waters" masterLastPageNumber="139" masterPageNumber="135" pageNumber="135" updateTime="1732862269424" updateUser="ExternalLinkService">
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<mods:title id="A3B75654727EF9879B40668863B4FCBB">First record of the speckled Maori wrasse, Oxycheilinus arenatus (Valenciennes, 1840) (Actinopterygii: Perciformes: Labridae), from Indian coastal waters</mods:title>
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<mods:namePart id="02CE7784861D9910268D18D0D32526B3">Murugan, Arumugam</mods:namePart>
<mods:affiliation id="D72C666B6E9B3FB5834F301BF45727D4">Centre for Marine Living Resources and Ecology, Ministry of Earth Sciences, Atal Bhavan, Puthuvype, Kochi, India</mods:affiliation>
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<mods:namePart id="25251D1DDE89FE741F73C00E54D1D0DD">Mahadevan, Gopalan</mods:namePart>
<mods:nameIdentifier id="00476491F2A3D0F12BD8489BE0C27394" type="ORCID">https://orcid.org/0000-0002-2654-4087</mods:nameIdentifier>
<mods:affiliation id="43B3A444ED77460EB1E0E1BBA7D69D48">Centre of Advanced Study in Marine Biology, Annamalai University, Parangipettai, Tamil Nadu, India</mods:affiliation>
<mods:nameIdentifier id="2E1537C33AF2A222D735A02F4A427B47" type="email">marinemahadevan@gmail.com</mods:nameIdentifier>
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<mods:namePart id="C0217EF8B531C6D9011B9F28D07AC232">Fricke, Ronald</mods:namePart>
<mods:affiliation id="18552A63D76194C20987E2656D81A7F4">Staatliches Museum fuer Naturkunde in Stuttgart, Stuttgart, Germany</mods:affiliation>
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<mods:title id="2CA58ADA164C464004A73D5714DE5B95">Acta Ichthyologica et Piscatoria</mods:title>
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<mods:date id="6C3344341D5D7DAA51501B38414B19C8">2022</mods:date>
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<mods:number id="D70EF91702F091190F1D765F55E08C1F">2022-06-24</mods:number>
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<mods:number id="D50C75873D035C6D1CF06552E3BFADA6">52</mods:number>
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<mods:start id="9F02CBF34C518085047DE5F86228CAB9">135</mods:start>
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<treatment id="715213C776D5544C9956E514238204BA" ID-GBIF-Taxon="196283062" LSID="urn:lsid:plazi:treatment:715213C776D5544C9956E514238204BA" httpUri="http://treatment.plazi.org/id/715213C776D5544C9956E514238204BA" lastPageNumber="135" pageId="0" pageNumber="135" scope_family="Labridae" scope_order="Perciformes">
<subSubSection id="A35B333EC70D0BFACD3202049D2DB4A7" pageId="0" pageNumber="135" type="nomenclature">
<paragraph id="3AB33D93B946FE637CD562A39210D4D0" pageId="0" pageNumber="135">
<taxonomicName id="B53487FA2D45585DDD786587B19D962F" ID-CoL="4BDM7" LSID="715213C7-76D5-544C-9956-E514238204BA" authority="(Valenciennes, 1840)" baseAuthorityName="Valenciennes" baseAuthorityYear="1840" family="Labridae" genus="Oxycheilinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oxycheilinus arenatus" order="Perciformes" pageId="0" pageNumber="135" phylum="Chordata" rank="species" species="arenatus">Oxycheilinus arenatus (Valenciennes, 1840)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="3DDB28ACD9B26BC004AAECC0A2BB1723" pageId="0" pageNumber="135" type="description">
<paragraph id="32ED3CD631D656CCFE66DD22BC3995EC" pageId="0" pageNumber="135">
<figureCitation id="D83F71E3217DBA1BB1B37671EF89BF8C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geographical distribution of Oxycheilinus arenatus in Indian Ocean, (A) New record from Tuticorin, India and (B) Previous records." figureDoi="10.3897/aiep.52.84796.figure1" httpUri="https://binary.pensoft.net/fig/705744" pageId="0" pageNumber="135">Figs 1</figureCitation>
<figureCitation id="A4A69D58BE0FFD5DC001A2D90433AD8E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Freshly preserved Oxycheilinus arenatus (A) (CASMBAURM / 232116489) and (B) X-ray of the adult Speckled Maori wrasse with 22 vertebrae (10 abdominal + 12 caudal)." figureDoi="10.3897/aiep.52.84796.figure2" httpUri="https://binary.pensoft.net/fig/705745" pageId="0" pageNumber="135">, 2</figureCitation>
<figureCitation id="F3981A1DEE8EAE639568F9F37BC81300" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Oxycheilinus arenatus (A) collected from Tuticorin, India (CASMBAURM / 232116487); (B) Head; (C) Dorsal fin and (D) Caudal fin." figureDoi="10.3897/aiep.52.84796.figure3" httpUri="https://binary.pensoft.net/fig/705746" pageId="0" pageNumber="135">, 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="1625FE748DD389165B9F44037C82CEF6" pageId="0" pageNumber="135" type="reference_group">
<paragraph id="41AF6400820EEF68067DBD2DCC24F08D" pageId="0" pageNumber="135">
<taxonomicName id="1F4EBEE9793B48AABDB1E82DB9ADA1D3" authorityName="Valenciennes" authorityYear="1840" family="Labridae" genus="Cheilinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheilinus arenatus" order="Perciformes" pageId="0" pageNumber="135" phylum="Chordata" rank="species" species="arenatus">Cheilinus arenatus</taxonomicName>
Valenciennes, 1840: 101, pl. 397 (
<normalizedToken id="90F8E373E8E6800BAAAB30D290EBCEDF" originalValue="Réunion">Reunion</normalizedToken>
, western Mascarenes, southwestern Indian Ocean).
</paragraph>
<paragraph id="F45C5F40B5CED25CD11E92450DBD8C50" pageId="0" pageNumber="135">
<taxonomicName id="5F73B15215864AFC343A1357351C4C57" authorityName="Bleeker" authorityYear="1853" family="Labridae" genus="Cheilinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheilinus notophthalmus" order="Perciformes" pageId="0" pageNumber="135" phylum="Chordata" rank="species" species="notophthalmus">Cheilinus notophthalmus</taxonomicName>
Bleeker, 1853: 493 (Jakarta, Java, Indonesia).
</paragraph>
</subSubSection>
<subSubSection id="16956047AF8F65B3366B316E65947950" pageId="0" pageNumber="135" type="description">
<paragraph id="B7AA74A9B027B5680ED337E0BA503FC9" pageId="0" pageNumber="135">Description.</paragraph>
<paragraph id="8A661CC2975B9BA5E3D606C8CF9FCA41" pageId="0" pageNumber="135">
Apart from the presence of the generic characters mentioned above, the specimens are diagnosed by the following characters: Dorsal fin IX, 10-11; anal fin III, 8-9; pelvic-fin rays I, 5; pectoral-fin rays of left and right sides 11/11~12/12 (2 uppermost rays unbranched); caudal-fin rays 13-14 (upper and lower 2 rays unbranched); pored lateral-line scales 22 (13 on anterior lateral line, 9 on posterior lateral line); vertical scale rows 2 + 7; number of scales on black streak 19; pored scales on black streak 8; predorsal scales 6; gill rakers 5 + 6 = 11; branchiostegal rays 5; vertebrae 10 + 12 = 22 (Fig.
<figureCitation id="D11F7350777DD8EDBF7B10E29DD38DEB" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Freshly preserved Oxycheilinus arenatus (A) (CASMBAURM / 232116489) and (B) X-ray of the adult Speckled Maori wrasse with 22 vertebrae (10 abdominal + 12 caudal)." figureDoi="10.3897/aiep.52.84796.figure2" httpUri="https://binary.pensoft.net/fig/705745" pageId="0" pageNumber="135">2</figureCitation>
). Body elongate, laterally compressed. Snout short, profile moderately rounded. Eye small; pupil slightly elongate. Interorbital space slightly convex, naked. Head and body covered with scales. Predorsal scales reaching anterior end of orbit. Opercular margins scaleless. Last scale of anterior lateral line located just below base of 4th dorsal-fin soft ray. First scale of posterior lateral line located just above base of 2nd anal-fin soft ray. Head laterally compressed, mouth terminal, gape oblique; posterior margin of lower lip extending beyond vertical through that of upper lip; posterior margin of maxilla extending beyond vertical through anterior margin of orbit; teeth in jaws affixed to outer edge of bony ridge; 2 pairs of large, slender, curved canine teeth anteriorly in each jaw; 16-18 small conical teeth fixed posteriorly on each bony plate behind upper- and lower-jaw canine teeth. Tongue slender, rounded, its upper surface covered with small papillae. Gill rakers thick, short, compressed; rakers on upper limb slightly shorter than those on lower limb; gill membranes free from isthmus. Posterior end of pectoral fin not reaching anus. Pelvic-fin origin and dorsal-fin origin almost on same vertical. Caudal fin truncate, with central part of medium edge slightly rounded, and upper and lower rays elongate. 9th dorsal-fin spine and 7th soft dorsal-fin ray longest; 3rd anal-fin spine and 4th soft anal-fin ray longest; 1st pelvic-fin soft ray longest (Table
<tableCitation id="444BC9547C40C5BAB5DF5D1ECD1BEE00" captionStart="Table 1" captionStartId="T1" captionText="Table 1. A comparison of morphometric characters of Oxycheilinus arenatus collected from Indian coastal waters and from Amami-oshima island, Ryukyu Islands, Japan." httpUri="http://table.plazi.org/id/7C8FAC0F4104FD6FD990FFF5C4B9D906" pageId="0" pageNumber="135" tableUuid="7C8FAC0F4104FD6FD990FFF5C4B9D906">1</tableCitation>
).
</paragraph>
<caption id="7C8FAC0F4104FD6FD990FFF5C4B9D906" ID-Table-UUID="7C8FAC0F4104FD6FD990FFF5C4B9D906" httpUri="http://table.plazi.org/id/7C8FAC0F4104FD6FD990FFF5C4B9D906" pageId="0" pageNumber="135" start="Table 1" startId="T1">
<paragraph id="5C581243C55AF2DAC21297A133F7A944" pageId="0" pageNumber="135">
<emphasis id="D9B4260586B5331FD918CD013F8D1CDD" bold="true" pageId="0" pageNumber="135">Table 1.</emphasis>
A comparison of morphometric characters of
<taxonomicName id="2018DE6D488C61642C71061C5CE89439" baseAuthorityName="Valenciennes" baseAuthorityYear="1840" family="Labridae" genus="Oxycheilinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oxycheilinus arenatus" order="Perciformes" pageId="0" pageNumber="135" phylum="Chordata" rank="species" species="arenatus">
<emphasis id="E9364EB68EDFEA51C3E258A75EFE0978" italics="true" pageId="0" pageNumber="135">Oxycheilinus arenatus</emphasis>
</taxonomicName>
collected from Indian coastal waters and from Amami-oshima island, Ryukyu Islands, Japan.
</paragraph>
</caption>
<paragraph id="53F3A84AED6C35FDAD08F74E2140091A" pageId="0" pageNumber="135">
<table id="091C1F27BD90F882DFD100D6FA786B56" pageId="0" pageNumber="135">
<tr id="8AEAAFD2E02CAFA62F9BFAE5E4451863" pageId="0" pageNumber="135">
<th id="022687EFBEDA51EF4254406940449B44" colspan="1" pageId="0" pageNumber="135" rowspan="3">Character</th>
<th id="3F9097DDD2D3DC6653F8FCB3A7CD3676" colspan="4" pageId="0" pageNumber="135" rowspan="1">
This study (
<emphasis id="9F168545B5B38EC9A71CF0B80BC4099D" italics="true" pageId="0" pageNumber="135">n</emphasis>
= 3)
</th>
<th id="B3BAEFEF9C0EB073E0F09D0762A36964" colspan="2" pageId="0" pageNumber="135" rowspan="1">
<bibRefCitation id="A5A25507BA00D70DCE7B28773AE3CEF0" author="Dewa, Y" journalOrPublisher="] Ichthy, Natural History of Fishes of Japan" pageId="0" pageNumber="135" pagination="5 - 7" refId="B4" refString="Dewa, Y, Maekawa, T, Motomura, H, 2021. [Second Japanese record of Oxycheilinus arenatus (Labridae) from Amami-oshima island, Ryukyu Islands, Japan.] Ichthy, Natural History of Fishes of Japan 8: 5 - 7" title="[Second Japanese record of Oxycheilinus arenatus (Labridae) from Amami-oshima island, Ryukyu Islands, Japan." volume="8" year="2021">Dewa et al. 2021</bibRefCitation>
(
<emphasis id="D4FA7A20FC895E458217684DB22F1587" italics="true" pageId="0" pageNumber="135">n</emphasis>
= 1)
</th>
</tr>
<tr id="15FF59B287D4C1C8C1A2A533D2C34268" pageId="0" pageNumber="135">
<th id="44E9480A7EF0AA198C2A0993707FE5EB" colspan="2" pageId="0" pageNumber="135" rowspan="1">Absolute value [mm]</th>
<th id="62842CEEF58F43FDD423DF387F7F2067" colspan="2" pageId="0" pageNumber="135" rowspan="1">%SL</th>
<th id="E49E3B6390E3E99A694AE351865A7873" colspan="1" pageId="0" pageNumber="135" rowspan="1">Abs. value [mm]</th>
<th id="67BA865C48A83B8573E101223E3EFB80" colspan="1" pageId="0" pageNumber="135" rowspan="1">%SL</th>
</tr>
<tr id="79912E3AEDE28746D0DEB50FAB9C94E9" pageId="0" pageNumber="135">
<th id="C656A87198FFC02F152732CCFA1222E6" colspan="1" pageId="0" pageNumber="135" rowspan="1">Range</th>
<th id="8CDFCFB4E5415E7AFC1A2AD62ECF940D" colspan="1" pageId="0" pageNumber="135" rowspan="1">Mean</th>
<th id="E006006AEBB530098BDAB48B5493434E" colspan="1" pageId="0" pageNumber="135" rowspan="1">Range</th>
<th id="E5F6826F441E8543482B737B1D0EDB8C" colspan="1" pageId="0" pageNumber="135" rowspan="1">Mean</th>
<th id="F835E64EEE832BE762DB87C713434748" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</th>
<th id="41A0C972EC6058D5F7003ABA632893F9" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</th>
</tr>
<tr id="39A5AA0D7C99F9170283A1DAEAB43C0C" pageId="0" pageNumber="135">
<td id="267310AB61B4E1BCFFC2A429C109FDAA" colspan="1" pageId="0" pageNumber="135" rowspan="1">Total length (TL)</td>
<td id="A1F25AA955FDFD49B74AE652A0743FB4" colspan="1" pageId="0" pageNumber="135" rowspan="1">177-195</td>
<td id="F7A9DE0DD591E4C31FE623FAA8680998" colspan="1" pageId="0" pageNumber="135" rowspan="1">186</td>
<td id="13FBF47BB27649B2E98D6578BCBABDF4" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="A755B8397B41358A9DE852975FB61B28" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="122EE38898EE6A5D177B9C3C44DA8041" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="A9054FBE2CE91FD16EAA180D025117EF" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
</tr>
<tr id="E1B406E92467EEAD6224B26637785669" pageId="0" pageNumber="135">
<td id="05BFB7ECAA9CFC7DAE74CC4B37002A67" colspan="1" pageId="0" pageNumber="135" rowspan="1">Standard length (SL)</td>
<td id="1785A24D1F67601FBEB0F41FC3885CA5" colspan="1" pageId="0" pageNumber="135" rowspan="1">142-157</td>
<td id="B269BC3DFC7047479E9D293CB5AFDC95" colspan="1" pageId="0" pageNumber="135" rowspan="1">151</td>
<td id="D062EA859EF19C311CB403D67308241E" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="898E9F53C08FCA2798D42BABF56C6FAB" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="14AA1A3085FDC68912AD816FC28642DE" colspan="1" pageId="0" pageNumber="135" rowspan="1">128.7</td>
<td id="8990D0836E59B372A90C40009BB09598" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
</tr>
<tr id="29203B701A754848E1A523C8DE641043" pageId="0" pageNumber="135">
<td id="383664FBD780ABA7BF1057D685E6F18A" colspan="1" pageId="0" pageNumber="135" rowspan="1">Head length</td>
<td id="CD8A13B914BA9197E0F4819B7EE492D9" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="4CA3617B702E3FF3A8B9C5598635046B" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="AA38DEB6D2A39265A32606A21BF3E4B0" colspan="1" pageId="0" pageNumber="135" rowspan="1">37.32-38.85</td>
<td id="00AB21A0272B366E45C7F9822C956A4F" colspan="1" pageId="0" pageNumber="135" rowspan="1">38.02</td>
<td id="6D70C53A970F1F57650BD225F1C49320" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="AA9569C2D708A102A663822635208DD0" colspan="1" pageId="0" pageNumber="135" rowspan="1">39.2</td>
</tr>
<tr id="59BE3D4CFEEBB7F297E67BC98F1DBF0A" pageId="0" pageNumber="135">
<td id="803BF06D9950C40F0539E4D274AE3669" colspan="1" pageId="0" pageNumber="135" rowspan="1">Head depth</td>
<td id="4625C04714C862F297BEFBBEE6770281" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="A48C6C6184BA25BECE4733855CAC96BE" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="51378407ABEF95D9DEDA333D1ADC60F3" colspan="1" pageId="0" pageNumber="135" rowspan="1">28.16-29.29</td>
<td id="E07874349F645B07234078336F45BFD5" colspan="1" pageId="0" pageNumber="135" rowspan="1">28.64</td>
<td id="C8112434EBFB5CB3475DD25A8187C6BB" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="A83CBC820E0A67728EA054273CBC9E4F" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
</tr>
<tr id="023A689735F6370A2C029830914EDD2F" pageId="0" pageNumber="135">
<td id="4E1381AAC948C484D6597032C31EE686" colspan="1" pageId="0" pageNumber="135" rowspan="1">Snout length</td>
<td id="3362958CCBFB5F1379042A52B8CA3823" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="834D2A2F61DE65E41F2D9AAF492A94AA" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="7410F381E45B700BFE2500A5544821D3" colspan="1" pageId="0" pageNumber="135" rowspan="1">14.08-15.28</td>
<td id="88C36E9E9CF64F33D1A3B05C0D81E5CF" colspan="1" pageId="0" pageNumber="135" rowspan="1">14.65</td>
<td id="82287A8D114AA0C692D922783C798A54" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="AFBC940CF170909AF1ACAD07631EEEEA" colspan="1" pageId="0" pageNumber="135" rowspan="1">14.1</td>
</tr>
<tr id="FA5BCF182605077BDD96B205A6EADF69" pageId="0" pageNumber="135">
<td id="0C2686322F34BF09D9E18111140B36CD" colspan="1" pageId="0" pageNumber="135" rowspan="1">Snout to end of preoperculum</td>
<td id="0B7BFC9E028C9BBDCF4D8333513DD1D5" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="793EDE0DD9813D298B6688BF235FF80A" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="E4B29B79EED2EBA523F83ABC302A4D1F" colspan="1" pageId="0" pageNumber="135" rowspan="1">27.46-28.02</td>
<td id="519E221CFD48EA0A037BC265477FEFD4" colspan="1" pageId="0" pageNumber="135" rowspan="1">27.73</td>
<td id="5F9B4ECE6E78B9397962B29A1B4026C5" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="7BA2CD328C870A7B47A62062B5A3DECF" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
</tr>
<tr id="A745C782B60DF501CCFCA0D22E6D49EF" pageId="0" pageNumber="135">
<td id="AB3CE8D887A88219198593F3179B20D5" colspan="1" pageId="0" pageNumber="135" rowspan="1">Orbit diameter</td>
<td id="DD13B94B2ACBCAD071EBBD8CDA466485" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="FA2E48447DE4B1198344ADD5C4E100B1" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="FC826ED095E1B5C4D026E585DC175AC9" colspan="1" pageId="0" pageNumber="135" rowspan="1">7.04-7.64</td>
<td id="379BB0E72DAD8BBCADB9F1BF97E5ACCB" colspan="1" pageId="0" pageNumber="135" rowspan="1">7.30</td>
<td id="7ED9E9DFF439D0767DCCF5CD96B7065D" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="900A7D8A4E16C07BB12239AEF8A6F1C8" colspan="1" pageId="0" pageNumber="135" rowspan="1">9.9</td>
</tr>
<tr id="7FB3BCB60BC7D666838C5DE3851C05BE" pageId="0" pageNumber="135">
<td id="6EC38905CA4501E95182C608F5842022" colspan="1" pageId="0" pageNumber="135" rowspan="1">Upper-jaw length</td>
<td id="637509642F5D8B8CDD3FA51CBFB7A130" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="B99A6D86F7E7C947D7AFD78D8363C1B0" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="40A35437D9FC7F30879C6064CD920A27" colspan="1" pageId="0" pageNumber="135" rowspan="1">10.56-12.10</td>
<td id="C80F656991A626CE77F5F0263E090424" colspan="1" pageId="0" pageNumber="135" rowspan="1">11.31</td>
<td id="8D1D16440AB15EE5AC1655503A337A1C" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="B6E45AAE7EA374A8F76A177A41C2FD7A" colspan="1" pageId="0" pageNumber="135" rowspan="1">12.9</td>
</tr>
<tr id="D03C3D0289F14EC3BBDCC784A944F5D5" pageId="0" pageNumber="135">
<td id="F909F1F8AFFB01A180DBC065F74B2F61" colspan="1" pageId="0" pageNumber="135" rowspan="1">Lower-jaw length</td>
<td id="5AD903F2817AFBD18AFB2D7160F8BA29" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="46F29E279D46088DDA642C3AEAF9EE92" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="462FD5690FBB5DA3BCA9C95E84429732" colspan="1" pageId="0" pageNumber="135" rowspan="1">11.97-12.73</td>
<td id="4D520984291E895F6CAD68DC0830931B" colspan="1" pageId="0" pageNumber="135" rowspan="1">12.42</td>
<td id="3AA91F61BFBC5C2B11B1E3D275C1BEF4" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="BC33617EF6B0D8F8CCC146ADD21596A7" colspan="1" pageId="0" pageNumber="135" rowspan="1">13.3</td>
</tr>
<tr id="BF9149610E8ACEC169FFA4441C78CC76" pageId="0" pageNumber="135">
<td id="977E52037944FCA3B998E34B728605AD" colspan="1" pageId="0" pageNumber="135" rowspan="1">Caudal-peduncle depth</td>
<td id="14A1387F672D895E8A5BC52B034A0855" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="E9DBBD6346D7264ABE39A628B857E89C" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="D67E0E723E186D06F738A05A8C392FC7" colspan="1" pageId="0" pageNumber="135" rowspan="1">16.90-18.44</td>
<td id="3AFD7DCD76AFE664E983CBF05DD189F7" colspan="1" pageId="0" pageNumber="135" rowspan="1">17.54</td>
<td id="EF0E10B33527B07075B950B3B065430E" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="6A4ABD6114F9E8894C9EDDCA307BBE6A" colspan="1" pageId="0" pageNumber="135" rowspan="1">14.4</td>
</tr>
<tr id="92BCE4BC711AD5DA03CBE63005D62068" pageId="0" pageNumber="135">
<td id="CEF5B1D0F4664FC8310AB165F31D93C9" colspan="1" pageId="0" pageNumber="135" rowspan="1">Caudal-peduncle length</td>
<td id="4B988E5189980310586CE93B0406F06E" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="BE6AC89966CE07A93758CEDD13120C10" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="535EDCE8BCB4B0B03D12AA5EC02FA8E3" colspan="1" pageId="0" pageNumber="135" rowspan="1">12.67-14.01</td>
<td id="DA2C90D0A5DE9EE8CF83597789D3C94A" colspan="1" pageId="0" pageNumber="135" rowspan="1">13.37</td>
<td id="BD6335184EF6E59A51391B469A5C2B02" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="C611EF7FF4BA80DDBBEED82B83D0CF79" colspan="1" pageId="0" pageNumber="135" rowspan="1">12.9</td>
</tr>
<tr id="3163E32840EA7393090E2E1A9443C233" pageId="0" pageNumber="135">
<td id="27E36B9DC3B4A29F90047B05A587C166" colspan="1" pageId="0" pageNumber="135" rowspan="1">Pectoral-fin length</td>
<td id="383363ABC330EA25F849F8254ADE083D" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="11659E905EFCF8D95B81BA5907DF3D6B" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="5B5A4F2F121234A9BA2114D5CBC673EB" colspan="1" pageId="0" pageNumber="135" rowspan="1">14.78-16.56</td>
<td id="35BD164EF15A4B4FF35D6CD244CC3F32" colspan="1" pageId="0" pageNumber="135" rowspan="1">15.64</td>
<td id="988CD609A1A0188C076B060FBF5D1720" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="8B3F360BE3E73B208955B1442D31AB69" colspan="1" pageId="0" pageNumber="135" rowspan="1">16.4</td>
</tr>
<tr id="EA05F92AC34808C4CBD98ED4A0AD5164" pageId="0" pageNumber="135">
<td id="3A9FE27CE3CD90B08322496601096A5E" colspan="1" pageId="0" pageNumber="135" rowspan="1">Pelvic-fin length</td>
<td id="E4D3ED0D2DFD03794B5A453CFC1A7BE4" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="FC8806B1170C264FC24D425542D3B6A4" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="ADF853BA11DB6EE7A8AAD8C8EDC7D3E4" colspan="1" pageId="0" pageNumber="135" rowspan="1">12.67-14.64</td>
<td id="46923E7C49420494CDA807E87F448498" colspan="1" pageId="0" pageNumber="135" rowspan="1">13.84</td>
<td id="C7753ADEA04F2A6C5DF8CA0158D428A7" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="E8A03378197385658D6E3FE1A23C27E5" colspan="1" pageId="0" pageNumber="135" rowspan="1">11.3</td>
</tr>
<tr id="763C0AF131C648D378B8906F904A11C9" pageId="0" pageNumber="135">
<td id="FBA3799771DD92B98B0709F0B1FC9BF9" colspan="1" pageId="0" pageNumber="135" rowspan="1">Dorsal-fin length</td>
<td id="C202D47471DE28C8A9976BE66ADCB313" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="42BEF84CD2C8560CE567818109A31470" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="4FF5A8473B1BE08F79935E23FA6552B6" colspan="1" pageId="0" pageNumber="135" rowspan="1">4.90-5.70</td>
<td id="5FCB2D46395FED896B1F9E6C6BF3BA6F" colspan="1" pageId="0" pageNumber="135" rowspan="1">5.30</td>
<td id="9956E80B0B2431DD353609C26028470B" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="7C51D5C2AB8DFBE68B0AD16A874EBBF7" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
</tr>
<tr id="3EFDBB9B387B7AF45EF9738076D700B7" pageId="0" pageNumber="135">
<td id="47CBCB6BB1DB27EF0C9AF970C4061227" colspan="1" pageId="0" pageNumber="135" rowspan="1">Dorsal-fin base length</td>
<td id="7AB5189D52C359EEB9C5E2D74B603651" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="6C6521D74852E0D4C0D48ECA75E5A37C" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="EF8B09235683F7363C9D5EF39D7A766F" colspan="1" pageId="0" pageNumber="135" rowspan="1">50.70-53.50</td>
<td id="DB0FD18C0F66FC179CE82119C9FEA6F7" colspan="1" pageId="0" pageNumber="135" rowspan="1">52.17</td>
<td id="59F11E609D722C40806884EAE0F6A533" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="90017DA7C5E00242EF6E851F09C56D43" colspan="1" pageId="0" pageNumber="135" rowspan="1">53.6</td>
</tr>
<tr id="E0D43C72B442FB6D78E4B45C78DA2A42" pageId="0" pageNumber="135">
<td id="B094210C54CB3F40A1494F5F0C60F4C7" colspan="1" pageId="0" pageNumber="135" rowspan="1">Anal-fin base length</td>
<td id="088B2C3F7EF30E2389B7CF5A2536D436" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="238EB02BF66EEB73B32E7893EF4FEDB3" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="88ADA06754A82E2048AA0AA4DCC7B67E" colspan="1" pageId="0" pageNumber="135" rowspan="1">4.90-5.70</td>
<td id="04C5FC64413B027F68CB48F46F4012FB" colspan="1" pageId="0" pageNumber="135" rowspan="1">5.30</td>
<td id="85D29A4EFC02EEF97E028B408B5686BB" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="953265EE1FD29CC4D298E7577A3EFD48" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
</tr>
<tr id="F02C5C2FB0FAFCD13402CE5A6DE33E85" pageId="0" pageNumber="135">
<td id="885F2960FEFEE08FA74A291DD412D4DF" colspan="1" pageId="0" pageNumber="135" rowspan="1">Pelvic-fin base length</td>
<td id="080C554F9D48C543E6901885058A06AF" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="D03B55C9BAB7DFFE9DE47FFAE5F68779" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="4949B53A6C436F8123FA6947DA37B0F6" colspan="1" pageId="0" pageNumber="135" rowspan="1">4.20-5.00</td>
<td id="4855B858132AF897B977228B12806710" colspan="1" pageId="0" pageNumber="135" rowspan="1">4.66</td>
<td id="3E8F9320E34F58A82FE4AFFE8616452F" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="E1794A9FB2C7899A377CCCC526D856A1" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
</tr>
<tr id="FD86D638A7C68942795A3F11C2C766DB" pageId="0" pageNumber="135">
<td id="2EAFF0670E4F15F811D6B535C234E9E3" colspan="1" pageId="0" pageNumber="135" rowspan="1">Anal-fin length</td>
<td id="4CEB4C409B70D905F4FFBA83715E983A" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="74A39215A7E7270F457E9415CC921CEE" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="B5889B107871B5BBAC6AFECDD61E9FCE" colspan="1" pageId="0" pageNumber="135" rowspan="1">26.76-26.82</td>
<td id="9C6CE0A97E2F85104EC3D93455B588D7" colspan="1" pageId="0" pageNumber="135" rowspan="1">26.67</td>
<td id="FF08FA3F95CAC26E139FBD0F80FC9551" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="AABB504BB05F9A1B8C426DB575723A44" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
</tr>
<tr id="D9BFE1DEFD4EC3B4F5B0C99B9BE65B25" pageId="0" pageNumber="135">
<td id="9C681672A068150D79709E9B062F8EF3" colspan="1" pageId="0" pageNumber="135" rowspan="1">Anal-fin width</td>
<td id="B6481B423C2E8E41098359D197CE892A" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="C96C38991C14210AF1D70D3552B8BF2E" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="0C0AE0EA96F5ECA07FB6AE4671770E1D" colspan="1" pageId="0" pageNumber="135" rowspan="1">35.21-35.32</td>
<td id="2F38F88C562FC6AFEDEBC251104AF83C" colspan="1" pageId="0" pageNumber="135" rowspan="1">35.35</td>
<td id="8FA8FA738A17B12A1AAEE64E07090C55" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="43702B46B8B54C046A8AC14B3DE92CC2" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
</tr>
<tr id="1A5001A2BFFE6927A1C7F9D33768B704" pageId="0" pageNumber="135">
<td id="3011476AD8FDA8186C1EB7ECC1FB0017" colspan="1" pageId="0" pageNumber="135" rowspan="1">Opercular length</td>
<td id="42E83D82E599662FC0A81679C44C34E6" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="99F0384516F05F574E63A823DF207190" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="EA2CC096067412B1A3B8DD253024CD62" colspan="1" pageId="0" pageNumber="135" rowspan="1">9.80-11.46</td>
<td id="52A8426BB0405D92D1D1A88BD9562CB8" colspan="1" pageId="0" pageNumber="135" rowspan="1">10.66</td>
<td id="3AA4B71E5951C6BBE2BE3CCA10B3E522" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="96E586EB50A2A2BD97AB18C4645EA387" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
</tr>
<tr id="DD891270A64691ED8AA1E7928F7B8F21" pageId="0" pageNumber="135">
<td id="BA14F20D680FD75BFDADE6AEB4615786" colspan="1" pageId="0" pageNumber="135" rowspan="1">Interorbital width</td>
<td id="657FFE6BAF5CAFA557499F5B10E5B726" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="607BA2ADD0091DA88A6AC16249E0F6E5" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="3045CDF044B89A5E6EA9D67F3B322ABB" colspan="1" pageId="0" pageNumber="135" rowspan="1">14.08-15.28</td>
<td id="A8AF172888DEB46E5B3E5714D1E3328C" colspan="1" pageId="0" pageNumber="135" rowspan="1">14.66</td>
<td id="403C4B81ED44F33F2DEA6200D62A0593" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="E574ACD897F9409044F0FABAB36F721B" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
</tr>
<tr id="CED59AE44CDAB2E4D9BE39F73C974663" pageId="0" pageNumber="135">
<td id="C8FB3F2B502CD24D97F34E90233ACADF" colspan="1" pageId="0" pageNumber="135" rowspan="1">Predorsal length</td>
<td id="98E818E4B0F6585942D08564587C9136" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="228AF34B8D09FB35B94DD06DD6B0DEBB" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="25E6E6226CF10B47AF5F6E4A6314642D" colspan="1" pageId="0" pageNumber="135" rowspan="1">45.19-47.75</td>
<td id="8F827D56AA3789CE012164E8AF2B8807" colspan="1" pageId="0" pageNumber="135" rowspan="1">46.53</td>
<td id="8AF809FCEEB2E6539AF87B21E19743B6" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="C0CE6C323FCBBFF18FDC3E5BF67B7761" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
</tr>
<tr id="6AFC5A84B4F2DEBD1D5A6AA097BDA339" pageId="0" pageNumber="135">
<td id="35B648EEECAD18DF0B08BCAC05B1286E" colspan="1" pageId="0" pageNumber="135" rowspan="1">Prepectoral length</td>
<td id="346B96928FFCFE210973CED7690D8F37" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="8D0F96BEB7F8F3FD72C022F96F28D29A" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="DE1A5EE1D5F0E37A158FE49B61E65DDD" colspan="1" pageId="0" pageNumber="135" rowspan="1">38.02-41.54</td>
<td id="CDD70080D261B95580CF400E04D63AC3" colspan="1" pageId="0" pageNumber="135" rowspan="1">39.74</td>
<td id="A3ECFB497390C0559DA0644C6158EB0C" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="483540E2C0A64CBD426866B153E38AD6" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
</tr>
<tr id="845D1906028C4735D0B148EB1CB424D1" pageId="0" pageNumber="135">
<td id="EFD3921237F9E4B6DD8C52C26E7F946A" colspan="1" pageId="0" pageNumber="135" rowspan="1">Prepelvic length</td>
<td id="7FF7EF184E54402F5A78CA8375ECED5A" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="50ABB20A35789A07B310D8351CFF2347" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="DBA675B2F06DB8B6DA0C08EB5DAE9755" colspan="1" pageId="0" pageNumber="135" rowspan="1">45.07-47.88</td>
<td id="7F28C86619A144FF4E4378D3CF3F44E3" colspan="1" pageId="0" pageNumber="135" rowspan="1">46.49</td>
<td id="E6F79DC2ED4903A0B6C623D5BA4781C9" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="A4BBBAD7D0D27D74D3433DDFE746A9D4" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
</tr>
<tr id="0734B58C7AA9FE5B243FBD390C4E985F" pageId="0" pageNumber="135">
<td id="09C6BD2A51C1D272DCE8727F362D8635" colspan="1" pageId="0" pageNumber="135" rowspan="1">Preanal length</td>
<td id="08696754587C50586673230DCA06BD27" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="BDF429D9FB088EA1FB5E1C08C44E95FF" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="F54AFF24654299A8A7089D53AA9660C3" colspan="1" pageId="0" pageNumber="135" rowspan="1">69.71-72.21</td>
<td id="3FF1F03C98A89E5D44604D103DB12A58" colspan="1" pageId="0" pageNumber="135" rowspan="1">70.76</td>
<td id="F700E49C56FBC5CFECBE9D8059F5CE15" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="C58725471A946C80A612B0629BC564D0" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
</tr>
<tr id="FDD375E5C18EF7CB05FCCC2D4EA2EE5A" pageId="0" pageNumber="135">
<td id="052283DD4543CAE7ED86E8B7E557059D" colspan="1" pageId="0" pageNumber="135" rowspan="1">1st anal-fin spine length</td>
<td id="33C5B0A4D3C3C44E77863228BA208AB5" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="A1B0E9BAE5443B9FB1E4A2531DCC71BE" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="7A00561573B0AF2CC052F9CECD355035" colspan="1" pageId="0" pageNumber="135" rowspan="1">4.90-5.70</td>
<td id="99EE1DEE4ED02FD7952D7366BCA5D61C" colspan="1" pageId="0" pageNumber="135" rowspan="1">5.33</td>
<td id="7239772BA3B820FC838885C0C8EA1B4C" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="5C126DEA326E11738BB0E84E46408C9A" colspan="1" pageId="0" pageNumber="135" rowspan="1">5.1</td>
</tr>
<tr id="D4EB95CD590549F2699ED9E995524FBC" pageId="0" pageNumber="135">
<td id="5EDFEE280F8B2C611B1B58A7DEE7281A" colspan="1" pageId="0" pageNumber="135" rowspan="1">2nd anal-fin spine length</td>
<td id="157AC8212C8FA6E4C9A193D9D71741F3" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="611A6B61CE23CC9FC22E622FF113CC79" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="953C8AF28F9A1D8F6B98EF77BCC8F6E0" colspan="1" pageId="0" pageNumber="135" rowspan="1">7.04-7.64</td>
<td id="B0605959486425EEFA17267E4005B776" colspan="1" pageId="0" pageNumber="135" rowspan="1">7.30</td>
<td id="E66130206CC7E502F51394FBFCE7BF3B" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="18654C3ECD5EDA448270A893FF981F9C" colspan="1" pageId="0" pageNumber="135" rowspan="1">10.4</td>
</tr>
<tr id="DFC5162F5E9CAF4AD0DB2D51BDC77341" pageId="0" pageNumber="135">
<td id="5B96C99CE5B033E6E8E0E737CA56CE11" colspan="1" pageId="0" pageNumber="135" rowspan="1">3rd anal-fin spine length</td>
<td id="82A415E3E498C94BA2C4F9B8602053DF" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="561C7DE7E33A68447D6491318EDB185F" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="F32CD2F23FC825CB98A7EBCA6573D65A" colspan="1" pageId="0" pageNumber="135" rowspan="1">9.80-11.97</td>
<td id="2C48239304D9787A2435704CCD6E1F38" colspan="1" pageId="0" pageNumber="135" rowspan="1">10.62</td>
<td id="04FDDCC7DFE488B8297FFACAEB26DF64" colspan="1" isEmpty="true" pageId="0" pageNumber="135" rowspan="1">-</td>
<td id="255DA8C8371F8492B659E838BEEE771E" colspan="1" pageId="0" pageNumber="135" rowspan="1">11.2</td>
</tr>
</table>
</paragraph>
<caption id="13C160A2BB13A670E1D426373DF2A00F" doi="10.3897/aiep.52.84796.figure2" httpUri="https://binary.pensoft.net/fig/705745" pageId="0" pageNumber="135" start="Figure 2" startId="F2">
<paragraph id="1C85CC6189E4479F13A5B34B737332A9" pageId="0" pageNumber="135">
<emphasis id="6645CC3EA12B5452AB3A3E2CF5091BCC" bold="true" pageId="0" pageNumber="135">Figure 2.</emphasis>
Freshly preserved
<taxonomicName id="A4F9375B3C2786CAC9A85BE76A8495DD" baseAuthorityName="Valenciennes" baseAuthorityYear="1840" family="Labridae" genus="Oxycheilinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oxycheilinus arenatus" order="Perciformes" pageId="0" pageNumber="135" phylum="Chordata" rank="species" species="arenatus">
<emphasis id="6B954D0B3BF8A979BA7FDEA58B3DF11F" italics="true" pageId="0" pageNumber="135">Oxycheilinus arenatus</emphasis>
</taxonomicName>
(
<emphasis id="E4E94451086536E4C169AD9F33E2B47F" bold="true" pageId="0" pageNumber="135">A</emphasis>
) (CASMBAURM/232116489) and (
<emphasis id="3867B1696AB4994B686F01FE120E5FF9" bold="true" pageId="0" pageNumber="135">B</emphasis>
) X-ray of the adult Speckled Maori wrasse with 22 vertebrae (10 abdominal +12 caudal).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="BFFE71B507A6BA3168FD3FA0A9144C11" pageId="0" pageNumber="135" type="color">
<paragraph id="558D8703796A99AB85DFB7AE887784A2" pageId="0" pageNumber="135">Color.</paragraph>
<paragraph id="F994A870AFEB58AF3691EED0B7F3291D" pageId="0" pageNumber="135">
Upper part of body light red, abdomen white. Sides with mottled pattern reaching from upper side of head to upper back, consisting of small, scattered orange or dark brown spots at low density. Dark vertical black band at center of body extending from posterior margin of eye to caudal-fin base. Small white spots scattered at high density above vertical band. Large black spot on membranes of first four dorsal-fin spines. Membranes between first four dorsal-fin soft rays otherwise pale green, with white lines and irregularly arranged white spots. Membranes between 4-10 dorsal-fin soft rays whitish translucent. Pectoral-fin base bright yellow, fin membranes transparent. Pelvic-fin base and anal-fin membranes white, fins mottled with light red. Medium caudal-fin membranes white-translucent, mottled with light red, upper and lower lobes yellow (Fig.
<figureCitation id="450684C0CC061EA73B654FCD28B27786" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Oxycheilinus arenatus (A) collected from Tuticorin, India (CASMBAURM / 232116487); (B) Head; (C) Dorsal fin and (D) Caudal fin." figureDoi="10.3897/aiep.52.84796.figure3" httpUri="https://binary.pensoft.net/fig/705746" pageId="0" pageNumber="135">3</figureCitation>
).
</paragraph>
<caption id="B4213EF8B6F6F518B3D24FF4F6299869" doi="10.3897/aiep.52.84796.figure3" httpUri="https://binary.pensoft.net/fig/705746" pageId="0" pageNumber="135" start="Figure 3" startId="F3">
<paragraph id="DED2125129940282C4187FCA72B41C6F" pageId="0" pageNumber="135">
<emphasis id="EC4B9F281D25D9E92BD529C1E0D76E7F" bold="true" pageId="0" pageNumber="135">Figure 3.</emphasis>
<taxonomicName id="454E66A8CE233FD3C9A760D47E6902BB" baseAuthorityName="Valenciennes" baseAuthorityYear="1840" family="Labridae" genus="Oxycheilinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oxycheilinus arenatus" order="Perciformes" pageId="0" pageNumber="135" phylum="Chordata" rank="species" species="arenatus">
<emphasis id="D4057137E766D276AC679C11C7DDDAB6" italics="true" pageId="0" pageNumber="135">Oxycheilinus arenatus</emphasis>
</taxonomicName>
(
<emphasis id="5B2F7A41645BCBEA4D845D9CF85A0D95" bold="true" pageId="0" pageNumber="135">A</emphasis>
) collected from Tuticorin, India (CASMBAURM/232116487); (
<emphasis id="D283E848E8E2F5A0E4878B0E4A798280" bold="true" pageId="0" pageNumber="135">B</emphasis>
) Head; (
<emphasis id="B0EA85703CE7E9AF7B1D7FB39EF9FA08" bold="true" pageId="0" pageNumber="135">C</emphasis>
) Dorsal fin and (
<emphasis id="40562060CACCB1B32C481373752E9E74" bold="true" pageId="0" pageNumber="135">D</emphasis>
) Caudal fin.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="078F39EAFB1B4A59EBBD73EFF7E7E52C" pageId="0" pageNumber="135" type="remarks">
<paragraph id="0152CBE1E2520BB687A465E19E2B349E" pageId="0" pageNumber="135">Remarks.</paragraph>
<paragraph id="381B0406A8160DE97369C397813551FF" pageId="0" pageNumber="135">
<taxonomicName id="010E38CC85FE8ECB2ACAC4D7AFC32FAF" baseAuthorityName="Valenciennes" baseAuthorityYear="1840" family="Labridae" genus="Oxycheilinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oxycheilinus arenatus" order="Perciformes" pageId="0" pageNumber="135" phylum="Chordata" rank="species" species="arenatus">
<emphasis id="EAE2E189CDAB1E0F5FCE54C2BAE0219F" italics="true" pageId="0" pageNumber="135">Oxycheilinus arenatus</emphasis>
</taxonomicName>
(Valenciennes, 1840), commonly known as speckled Maori wrasse, is known from the Red Sea and the Indo-West Pacific: East Africa, Seychelles, Madagascar, and Mascarenes east to Marshall Islands and Samoa, north to Ryukyu Islands (Japan). Generally, from the conservation point of view, they are considered 'Least
<normalizedToken id="B60709F3BF79A3B8C47168E706BB2A77" originalValue="concern’">concern'</normalizedToken>
(
<bibRefCitation id="D1AF7D1256863701A82342A16789CD92" DOI="https://doi.org/10.2305/IUCN.UK.2010-4.RLTS.T187718A8611549.en" author="Liu, M" journalOrPublisher="Journal of the Ocean Science Foundation" pageId="0" pageNumber="135" refId="B21" refString="Liu, M, To, A, 2010. Oxycheilinus arenatus, thin-line Maori wrasse. The IUCN Red List of Threatened Species 2010: e.T187718A8611549. https://doi.org/10.2305/IUCN.UK.2010-4.RLTS.T187718A8611549.en" title="Oxycheilinus arenatus, thin-line Maori wrasse. The IUCN Red List of Threatened Species 2010: e. T 187718 A 8611549." url="https://doi.org/10.2305/IUCN.UK.2010-4.RLTS.T187718A8611549.en" year="2010">Liu and To 2010</bibRefCitation>
). In the Red Sea and Indian Ocean, the species was previously recorded from La
<normalizedToken id="F0DF8E4584AF87256BEBA3A688C3992A" originalValue="Réunion">Reunion</normalizedToken>
(
<bibRefCitation id="2714CB0FB3F3427C29DCAF02DC41B9A6" DOI="https://doi.org/10.5962/bhl.title.7339" author="Valenciennes, A" journalOrPublisher="Zoological Studies (Taipei, Taiwan)" pageId="0" pageNumber="135" refId="B30" refString="Valenciennes, A, 1839. [Effectively published Jan. 1840.] Suite du livre seizieme. Labroides. Pp. 1-307. In: Cuvier G, Valenciennes A (Eds) Histoire naturelle des poissons. Tome 14. Levrault, Paris. Pp. i-xxii + 2 pp. + 1-464 + 4 pp., pls. 389-420. https://doi.org/10.5962/bhl.title.7339" title="[Effectively published Jan. 1840.] Suite du livre seizieme. Labroides. Pp. 1 - 307. In: Cuvier G, Valenciennes A (Eds) Histoire naturelle des poissons. Tome 14. Levrault, Paris. Pp. i-xxii + 2 pp. + 1 - 464 + 4 pp., pls. 389 - 420." url="https://doi.org/10.5962/bhl.title.7339" year="1839">Valenciennes 1839</bibRefCitation>
, as
<taxonomicName id="5ED1269AA62B87CB8B0BDC876E49A974" authorityName="Valenciennes" authorityYear="1840" family="Labridae" genus="Cheilinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheilinus arenatus" order="Perciformes" pageId="0" pageNumber="135" phylum="Chordata" rank="species" species="arenatus">
<emphasis id="D68E079C6F489F33EF8DC4BB00688F77" italics="true" pageId="0" pageNumber="135">Cheilinus arenatus</emphasis>
</taxonomicName>
;
<bibRefCitation id="FFB430B7DCDC09D9318049DC4CD02446" author="Fricke, R" journalOrPublisher="] Ichthy, Natural History of Fishes of Japan" pageId="0" pageNumber="135" refId="B7" refString="Fricke, R, Mulochau, T, Durville, P, Chabanet, P, Tessier, E, Letourneur, Y, 2009. Annotated checklist of the fish species (Pisces) of La Reunion, including a Red List of threatened and declining species. Stuttgarter Beitraege zur Naturkunde A. Neue Serie.2: 1-168." title="Annotated checklist of the fish species (Pisces) of La Reunion, including a Red List of threatened and declining species. Stuttgarter Beitraege zur Naturkunde A. Neue Serie. 2: 1 - 168." year="2009">Fricke et al. 2009</bibRefCitation>
), Chagos Archipelago (
<bibRefCitation id="E11C8DACB215FD99A6E65D16597A48C9" DOI="https://doi.org/10.5962/bhl.title.52237" author="Winterbottom, R" journalOrPublisher="Bulletin of Marine Science" pageId="0" pageNumber="135" refId="B34" refString="Winterbottom, R, Emery, AR, Holm, E, 1989. An annotated checklist of the fishes of the Chagos Archipelago, central Indian Ocean. Royal Ontario Museum Life Science Contributions, 145: 1-226, pls. 1-8. https://doi.org/10.5962/bhl.title.52237" title="An annotated checklist of the fishes of the Chagos Archipelago, central Indian Ocean. Royal Ontario Museum Life Science Contributions, 145: 1 - 226, pls. 1 - 8." url="https://doi.org/10.5962/bhl.title.52237" year="1989">Winterbottom et al. 1989</bibRefCitation>
, as
<taxonomicName id="C6B9C4DE7158E7F669AFA8F568455FB7" authorityName="Valenciennes" authorityYear="1840" family="Labridae" genus="Cheilinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheilinus arenatus" order="Perciformes" pageId="0" pageNumber="135" phylum="Chordata" rank="species" species="arenatus">
<emphasis id="390F5F9CAB7A307673BC8B92D00A8227" italics="true" pageId="0" pageNumber="135">Cheilinus arenatus</emphasis>
</taxonomicName>
), Mauritius (
<bibRefCitation id="E3F4C7B98892CA06E4E3142FDB72933F" DOI="https://doi.org/10.5962/bhl.title.8809" author="Guenther, ACLG" journalOrPublisher="Journal of the Ocean Science Foundation" pageId="0" pageNumber="135" refId="B19" refString="Guenther, ACLG, 1862. Catalogue of the fishes in the British Museum. Catalogue of the Acanthopterygii, Pharyngognathi and Anacanthini in the collection of the British Museum. Volume 4. British Museum, London, i-xxi + 1-534. https://doi.org/10.5962/bhl.title.8809" title="Catalogue of the fishes in the British Museum. Catalogue of the Acanthopterygii, Pharyngognathi and Anacanthini in the collection of the British Museum. Volume 4. British Museum, London, i-xxi + 1 - 534." url="https://doi.org/10.5962/bhl.title.8809" year="1862">
<normalizedToken id="B17BD9FAEE71AD1E298F6969446B7189" originalValue="Günther">Guenther</normalizedToken>
1862
</bibRefCitation>
, as
<taxonomicName id="4391DFA22459E463ED584E72762C8F9A" authorityName="Valenciennes" authorityYear="1840" family="Labridae" genus="Cheilinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheilinus arenatus" order="Perciformes" pageId="0" pageNumber="135" phylum="Chordata" rank="species" species="arenatus">
<emphasis id="D467D938998986339AF97FFB71FA50FF" italics="true" pageId="0" pageNumber="135">Cheilinus arenatus</emphasis>
</taxonomicName>
), Seychelles (
<bibRefCitation id="FF003780623494ADF5DE36F66AD5C170" author="Smith, JLB" journalOrPublisher="Zoological Studies (Taipei, Taiwan)" pageId="0" pageNumber="135" refId="B28" refString="Smith, JLB, Smith, MM, 1963. The fishes of Seychelles. 1st edition. Department of Ichthyology, Rhodes University, Grahamstown, i-viii + 1-215, pls. 1-98." title="The fishes of Seychelles. 1 st edition. Department of Ichthyology, Rhodes University, Grahamstown, i-viii + 1 - 215, pls. 1 - 98." year="1963">Smith and Smith 1963</bibRefCitation>
, as
<taxonomicName id="151176879ED0B4B452306BD4E2BE33A8" authorityName="Valenciennes" authorityYear="1840" family="Labridae" genus="Cheilinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheilinus arenatus" order="Perciformes" pageId="0" pageNumber="135" phylum="Chordata" rank="species" species="arenatus">
<emphasis id="F8ACE83739D43AAFFCE5ACB8A1115851" italics="true" pageId="0" pageNumber="135">Cheilinus arenatus</emphasis>
</taxonomicName>
), Maldives (
<bibRefCitation id="323407BF50C1A8D0B1210C6C9D127B4B" author="Randall, JE" journalOrPublisher="Smith Institute of Ichthyology" pageId="0" pageNumber="135" pagination="1 - 47" refId="B24" refString="Randall, JE, Anderson, RC, 1993. Annotated checklist of the epipelagic and shore fishes of the Maldive Islands. Ichthyological Bulletin of the J. L. B. Smith Institute of Ichthyology 59: 1 - 47" title="Annotated checklist of the epipelagic and shore fishes of the Maldive Islands. Ichthyological Bulletin of the J. L. B." volume="59" year="1993">Randall and Anderson 1993</bibRefCitation>
, as
<taxonomicName id="5E9ED58CA11A776537D88D136909D5FA" authorityName="Valenciennes" authorityYear="1840" family="Labridae" genus="Cheilinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheilinus arenatus" order="Perciformes" pageId="0" pageNumber="135" phylum="Chordata" rank="species" species="arenatus">
<emphasis id="67ADC4F65F5AD81CF8119D0CC513FFD7" italics="true" pageId="0" pageNumber="135">Cheilinus arenatus</emphasis>
</taxonomicName>
;
<bibRefCitation id="B7C888E56CC1F9B3348558E1AFB66489" author="Anderson, RC" pageId="0" pageNumber="135" refId="B1" refString="Anderson, RC, Randall, JE, Kuiter, RH, 1998. Additions to the fish fauna of the Maldive Islands. Part 2: New records of fishes from the Maldive Islands, with notes on other species. Ichthyological Bulletin of the J. L. B. Smith Institute of Ichthyology 67: 20-32, pls. 1-4." title="Additions to the fish fauna of the Maldive Islands. Part 2: New records of fishes from the Maldive Islands, with notes on other species. Ichthyological Bulletin of the J. L. B. Smith Institute of Ichthyology 67: 20 - 32, pls. 1 - 4." year="1998">Anderson et al. 1998</bibRefCitation>
), Mozambique (
<bibRefCitation id="81DB07DDBE7F96AA22FD9DD1C77DCB0C" DOI="https://doi.org/10.1071/MF01125" author="Gell, FR" journalOrPublisher="Marine and Freshwater Research" pageId="0" pageNumber="135" pagination="115 - 121" refId="B15" refString="Gell, FR, Whittington, MW, 2002. Diversity of fishes in seagrass beds in the Quirimba Archipelago, northern Mozambique. Marine and Freshwater Research 53 (2): 115 - 121, DOI: https://doi.org/10.1071/MF01125" title="Diversity of fishes in seagrass beds in the Quirimba Archipelago, northern Mozambique." url="https://doi.org/10.1071/MF01125" volume="53" year="2002">Gell and Whittington 2002</bibRefCitation>
), Red Sea (
<bibRefCitation id="0FDB54F9B8C457D179F7B4640D2E2BD5" author="Randall, JE" journalOrPublisher="Zoological Studies (Taipei, Taiwan)" pageId="0" pageNumber="135" pagination="135 - 139" refId="B25" refString="Randall, JE, Khalaf, M, 2003. Redescription of the labrid fish Oxycheilinus orientalis (Guenther), a senior synonym of O. rhodochrous (Guenther), and the first record from the Red Sea. Zoological Studies (Taipei, Taiwan) 42 (1): 135 - 139" title="Redescription of the labrid fish Oxycheilinus orientalis (Guenther), a senior synonym of O. rhodochrous (Guenther), and the first record from the Red Sea." volume="42" year="2003">Randall and Khalaf 2003</bibRefCitation>
;
<bibRefCitation id="DBAF38CDDBEAF6B2B4209D05C5A16185" DOI="https://doi.org/10.11646/zootaxa.4509.1.1" author="Golani, D" journalOrPublisher="Zootaxa" pageId="0" pageNumber="135" pagination="1 - 215" refId="B17" refString="Golani, D, Fricke, R, 2018. Checklist of the Red Sea fishes with delineation of the Gulf of Suez, Gulf of Aqaba, endemism and Lessepsian migrants. Zootaxa 4509 (1): 1 - 215, DOI: https://doi.org/10.11646/zootaxa.4509.1.1" title="Checklist of the Red Sea fishes with delineation of the Gulf of Suez, Gulf of Aqaba, endemism and Lessepsian migrants." url="https://doi.org/10.11646/zootaxa.4509.1.1" volume="4509" year="2018">Golani and Fricke 2018</bibRefCitation>
), Madagascar (
<bibRefCitation id="F234305C462C74000FD4D84E4853E888" author="Fricke, R" journalOrPublisher="] Ichthy, Natural History of Fishes of Japan" pageId="0" pageNumber="135" refId="B6" refString="Fricke, R, 1999. Fishes of the Mascarene Islands (Reunion, Mauritius, Rodriguez). An annotated checklist with descriptions of new species. Koeltz Scientific Books, Koenigstein, i-viii + 1-759." title="Fishes of the Mascarene Islands (Reunion, Mauritius, Rodriguez). An annotated checklist with descriptions of new species. Koeltz Scientific Books, Koenigstein, i-viii + 1 - 759." year="1999">Fricke 1999</bibRefCitation>
;
<bibRefCitation id="B4D8E9493E433AEB4618112D1D8DCA9E" author="Fricke, R" journalOrPublisher="FishTaxa: Journal of Fish Taxonomy" pageId="0" pageNumber="135" pagination="1 - 432" refId="B8" refString="Fricke, R, Mahafina, J, Behivoke, F, Jaonalison, H, Leopold, M, Ponton, D, 2018. Annotated checklist of the fishes of Madagascar, southwestern Indian Ocean, with 158 new records. FishTaxa: Journal of Fish Taxonomy 3 (1): 1 - 432" title="Annotated checklist of the fishes of Madagascar, southwestern Indian Ocean, with 158 new records." volume="3" year="2018">Fricke et al. 2018</bibRefCitation>
). The current findings represent a range extension of ca. 700 km to the northeast of the closest previously known region (Maldives), and the first record of this species from India.
<taxonomicName id="B2F7328305DF7FD92F6E7A300289EA0D" baseAuthorityName="Valenciennes" baseAuthorityYear="1840" family="Labridae" genus="Oxycheilinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oxycheilinus arenatus" order="Perciformes" pageId="0" pageNumber="135" phylum="Chordata" rank="species" species="arenatus">
<emphasis id="8ABA12F35D1692F6F62C16DBE6510C3F" italics="true" pageId="0" pageNumber="135">Oxycheilinus arenatus</emphasis>
</taxonomicName>
is widespread in the Indo-West Pacific and was probably previously overlooked in Indian waters. Apparently, the species is very rare in India.
</paragraph>
<paragraph id="5A14392FFB5BBC861732DF202DBBB8CA" pageId="0" pageNumber="135">
This species is found in caves of steep outer reef drop-offs from 25 to at least 46 m with rich invertebrate growth such as large gorgonians or soft corals (
<bibRefCitation id="641A8CFF8B18CC325784CE0621E5253E" author="Froese, R" journalOrPublisher="FishTaxa: Journal of Fish Taxonomy" pageId="0" pageNumber="135" publicationUrl="https://www.fishbase.se/search.php" refId="B11" refString="Froese, R, Pauly, D, 2022. (2022) FishBase. [Version 02/2022] https://www.fishbase.se/search.php" title="(2022) FishBase. [Version 02 / 2022]" url="https://www.fishbase.se/search.php" year="2022">Froese and Pauly 2022</bibRefCitation>
). It is naturally rare and has been rarely documented due to its cryptic occurrence in deep reefs. The lower extent of the depth distribution range is unknown. The collecting depth of the Indian specimens below 40 m well agrees with the previously known depth range.
</paragraph>
</subSubSection>
</treatment>
</document>
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